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<document id="2C0E7EA0E6796AF5B395612B16258464" ID-DOI="10.5281/zenodo.282252" ID-GBIF-Dataset="85464cf2-2a3c-43d4-8393-6e2c40ed058f" ID-ISSN="1175-5326" ID-Zenodo-Dep="282252" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460524004077" checkinUser="plazi" docAuthor="Zhao, Shuang, Hájek, Jiří, Jia, Fenglong &amp; Pang, Hong" docDate="2012" docId="03DAEC011425FF83FF7C071318E7FE92" docLanguage="en" docName="zt03478p212.pdf" docOrigin="Zootaxa 3478" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Neptosternus hydaticoides Regimbart 1877" docType="treatment" docVersion="8" lastPageNumber="210" masterDocId="FFE394791427FF86FFEB01051C29FF8E" masterDocTitle="A taxonomic review of the genus Neptosternus Sharp of China with the description of a new species (Coleoptera: Dytiscidae: Laccophilinae)" masterLastPageNumber="212" masterPageNumber="205" pageNumber="207" updateTime="1698314870484" updateUser="plazi">
<mods:mods id="9F77A83EA6BDCE45231C8F012CE20E3B" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="6716138062D5B282B422836BC6884D71">
<mods:title id="0A72984BB67D8E5749BE0FDEE1F196E0">A taxonomic review of the genus Neptosternus Sharp of China with the description of a new species (Coleoptera: Dytiscidae: Laccophilinae)</mods:title>
</mods:titleInfo>
<mods:name id="8DE748464B94D4711576E83975C0B2F2" type="personal">
<mods:role id="3C661A24660803E7F3A8BA10CC412827">
<mods:roleTerm id="DDC3E98DB949660F68676F1963B937EF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FAB0BFB69EEF4919958F7DDCFB81B151">Zhao, Shuang</mods:namePart>
</mods:name>
<mods:name id="18C708ED518651161FFFA026756BEB28" type="personal">
<mods:role id="0A2BD3BC819F4B4EFF4102EEDA3AEDFA">
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</mods:role>
<mods:namePart id="3D76FFE6EFBAB1039768FABD426D2963">Hájek, Jiří</mods:namePart>
</mods:name>
<mods:name id="93C6A99625B5C5C16157B933F1376EF4" type="personal">
<mods:role id="DE0B174E9F6A47A44B6017773667A3DC">
<mods:roleTerm id="E07E939A08B28FE2B85605039998B209">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CD309E2BDC99BA356021D069B2C76B55">Jia, Fenglong</mods:namePart>
</mods:name>
<mods:name id="9AF312D9800F62030B8BA5BC738D3DB9" type="personal">
<mods:role id="F9B2E7C75D2F91E9913B5CF84EB6B812">
<mods:roleTerm id="83130171AA8E1DC38666F265F64B37E7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0780E3C1CD35C1972AEA8F119F591E83">Pang, Hong</mods:namePart>
</mods:name>
<mods:typeOfResource id="00F1177BAF358A0595C5DBEB99FA7A4D">text</mods:typeOfResource>
<mods:relatedItem id="460DF38AF7F83D170C3B762393702740" type="host">
<mods:titleInfo id="117E1A0CCEC5B49FBB00BFBE075407E5">
<mods:title id="52946127B3145D7CBE46F3A03C2D1E2A">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="058D0FFC01A5596A828D586DA6F3FA01">
<mods:date id="03225042F07E13400D7BA9FDB84229AB">2012</mods:date>
<mods:detail id="502D757027416FBAAB3A554EE55815FF" type="volume">
<mods:number id="7782426D9C7C2741D206EF343619FB26">3478</mods:number>
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<mods:extent id="E15BA72B0BE13F33CAF96221C0A37CA1" unit="page">
<mods:start id="6CEE87B82C920BB4D440ABD3424BA97B">205</mods:start>
<mods:end id="CA01FE9BC9E99D08FE3CA7C074FC240E">212</mods:end>
</mods:extent>
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<mods:classification id="31A234850464FDF0FA404F86D74686BB">journal article</mods:classification>
<mods:identifier id="265057B74B9D329D6EEF0ECA5CB60D73" type="DOI">10.5281/zenodo.282252</mods:identifier>
<mods:identifier id="D53A75A015887A232B72442CE32501FC" type="GBIF-Dataset">85464cf2-2a3c-43d4-8393-6e2c40ed058f</mods:identifier>
<mods:identifier id="8B68F993F248EE8ABFF9CD5B59256748" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="72E75B9A5F5741E6974D886638BFE650" type="Zenodo-Dep">282252</mods:identifier>
</mods:mods>
<treatment id="03DAEC011425FF83FF7C071318E7FE92" ID-DOI="http://doi.org/10.5281/zenodo.6170225" ID-GBIF-Taxon="119526133" ID-Zenodo-Dep="6170225" LSID="urn:lsid:plazi:treatment:03DAEC011425FF83FF7C071318E7FE92" httpUri="http://treatment.plazi.org/id/03DAEC011425FF83FF7C071318E7FE92" lastPageId="5" lastPageNumber="210" pageId="2" pageNumber="207">
<subSubSection id="C3690E9C1425FF84FF7C07131D69F97C" pageId="2" pageNumber="207" type="nomenclature">
<paragraph id="8BCC5D171425FF84FF7C07131E97F9BE" blockId="2.[151,702,1558,1616]" box="[151,702,1558,1584]" pageId="2" pageNumber="207">
<heading id="D084EA7B1425FF84FF7C07131E97F9BE" bold="true" box="[151,702,1558,1584]" fontSize="11" level="1" pageId="2" pageNumber="207" reason="1">
<taxonomicName id="4C7326941425FF84FF7C07131E97F9BE" ID-CoL="473X9" authority="Regimbart, 1877" authorityName="Regimbart" authorityYear="1877" box="[151,702,1558,1584]" class="Insecta" family="Dytiscidae" genus="Neptosternus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="207" phylum="Arthropoda" rank="species" species="hydaticoides">
<emphasis id="B90781051425FF84FF7C07131E97F9BE" bold="true" box="[151,702,1558,1584]" pageId="2" pageNumber="207">
<emphasis id="B90781051425FF84FF7C07131DFDF9BE" bold="true" box="[151,468,1558,1584]" italics="true" pageId="2" pageNumber="207">Neptosternus hydaticoides</emphasis>
(Régimbart, 1877)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BCC5D171425FF84FF7C073D1D36F9DE" blockId="2.[151,702,1558,1616]" box="[151,287,1592,1616]" pageId="2" pageNumber="207">(Figs. 5, 11)</paragraph>
<paragraph id="8BCC5D171425FF84FF7C078419B5F95A" blockId="2.[151,1436,1665,1809]" pageId="2" pageNumber="207">
<taxonomicName id="4C7326941425FF84FF7C07841E64F918" authority="Regimbart, 1877" authorityName="Regimbart" authorityYear="1877" box="[151,589,1665,1687]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="207" phylum="Arthropoda" rank="species" species="hydaticoides">
<emphasis id="B90781051425FF84FF7C07841DB0F919" box="[151,409,1665,1687]" italics="true" pageId="2" pageNumber="207">Laccophilus hydaticoides</emphasis>
Régimbart, 1877
</taxonomicName>
: LXXIX (original description,
<collectingCountry id="F3641D871425FF84FC6C07841FD4F919" box="[903,1021,1665,1687]" name="Philippines" pageId="2" pageNumber="207">Philippines</collectingCountry>
: Luzon).
<taxonomicName id="4C7326941425FF84FF7C07A41EAEF93B" authority="Regimbart 1899: 270" authorityName="Regimbart" authorityPageNumber="270" authorityYear="1899" box="[151,647,1695,1718]" class="Insecta" family="Dytiscidae" genus="Neptosternus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="207" phylum="Arthropoda" rank="species" species="hydaticoides">
<emphasis id="B90781051425FF84FF7C07A41D8AF93B" box="[151,419,1695,1718]" italics="true" pageId="2" pageNumber="207">Neptosternus hydaticoides</emphasis>
: Régimbart 1899: 270
</taxonomicName>
(new combination); Hendrich &amp; Balke 1997: 58 (redescription).
<taxonomicName id="4C7326941425FF84FF7C07BA1E5BF95A" authority="Guignot, 1956: 452" authorityName="Guignot" authorityPageNumber="452" authorityYear="1956" box="[151,626,1726,1748]" class="Insecta" family="Dytiscidae" genus="Neptosternus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="207" phylum="Arthropoda" rank="species" species="tenasserimus">
<emphasis id="B90781051425FF84FF7C07BA1D8EF95A" box="[151,423,1727,1748]" italics="true" pageId="2" pageNumber="207">Neptosternus tenasserimus</emphasis>
Guignot, 1956: 452
</taxonomicName>
(original description;
<collectingCountry id="F3641D871425FF84FCB807BA1F90F95A" box="[851,953,1727,1748]" name="Myanmar" pageId="2" pageNumber="207">Myanmar</collectingCountry>
: Tenasserim); Hendrich &amp; Balke 1997: 57 (new
</paragraph>
<paragraph id="8BCC5D171425FF84FF2C07D81D69F97C" blockId="2.[151,1436,1665,1809]" box="[199,320,1757,1778]" pageId="2" pageNumber="207">synonymy).</paragraph>
</subSubSection>
<subSubSection id="C3690E9C1425FF84FF7C07F9180BF89F" box="[151,1058,1787,1809]" pageId="2" pageNumber="207" type="reference_group">
<paragraph id="8BCC5D171425FF84FF7C07F9180BF89F" blockId="2.[151,1436,1665,1809]" box="[151,1058,1787,1809]" pageId="2" pageNumber="207">
<treatmentCitationGroup id="AB637A391425FF84FF7C07F9180BF89F" box="[151,1058,1787,1809]" pageId="2" pageNumber="207">
<taxonomicName id="4C7326941425FF84FF7C07F91E12F89F" ID-CoL="8LF86" authority="Bian &amp; Ji, 2009: 36" authorityName="Bian &amp; Ji" authorityPageNumber="36" authorityYear="2009" box="[151,571,1787,1809]" class="Insecta" family="Dytiscidae" genus="Neptosternus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="207" phylum="Arthropoda" rank="species" species="yanbini">
<emphasis id="B90781051425FF84FF7C07F91D46F89F" box="[151,367,1787,1809]" italics="true" pageId="2" pageNumber="207">Neptosternus yanbini</emphasis>
<treatmentCitation id="0AD27B061425FF84FE9E07FE1E12F89F" author="Bian" box="[373,571,1787,1809]" page="36" pageId="2" pageNumber="207" year="2009">
<bibRefCitation id="EFE220E61425FF84FE9E07FE1E31F89F" author="Bian" box="[373,536,1787,1809]" pageId="2" pageNumber="207" refString="Bian, D. &amp; Ji, L. (2009) Two new species of Dytiscidae from Hainan, China. The Coleopterists Bulletin, 63, 35 - 40." type="journal article" year="2009">Bian &amp; Ji, 2009</bibRefCitation>
: 36
</treatmentCitation>
</taxonomicName>
(original description;
<collectingCountry id="F3641D871425FF84FCF607FE1F74F89F" box="[797,861,1787,1809]" name="China" pageId="2" pageNumber="207">China</collectingCountry>
: Hainan),
<emphasis id="B90781051425FF84FC2E07F9180BF89F" bold="true" box="[965,1058,1788,1809]" pageId="2" pageNumber="207">syn. nov.</emphasis>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3690E9C1425FF85FF7C063B1847F925" lastPageId="3" lastPageNumber="208" pageId="2" pageNumber="207" type="materials_examined">
<paragraph id="8BCC5D171425FF84FF7C063B1DE6F84C" blockId="2.[151,1437,1853,1986]" pageId="2" pageNumber="207">
<emphasis id="B90781051425FF84FF7C063B1D83F8D8" bold="true" box="[151,426,1853,1878]" pageId="2" pageNumber="207">
<typeStatus id="54C8E3B51425FF84FF7C063B1CFBF8D8" box="[151,210,1854,1878]" pageId="2" pageNumber="207">Type</typeStatus>
material studied.
</emphasis>
<taxonomicName id="4C7326941425FF84FE5E063A1F5BF8D8" authority="Bian &amp; Ji, 2009" authorityName="Bian &amp; Ji" authorityYear="2009" box="[437,882,1854,1878]" class="Insecta" family="Dytiscidae" genus="Neptosternus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="207" phylum="Arthropoda" rank="species" species="yanbini">
<emphasis id="B90781051425FF84FE5E063A1E8EF8D8" box="[437,679,1854,1878]" italics="true" pageId="2" pageNumber="207">Neptosternus yanbini</emphasis>
<bibRefCitation id="EFE220E61425FF84FD59063B1F5BF8D8" author="Bian" box="[690,882,1854,1878]" pageId="2" pageNumber="207" refString="Bian, D. &amp; Ji, L. (2009) Two new species of Dytiscidae from Hainan, China. The Coleopterists Bulletin, 63, 35 - 40." type="journal article" year="2009">Bian &amp; Ji, 2009</bibRefCitation>
</taxonomicName>
:
<typeStatus id="54C8E3B51425FF84FC6A063B1FC0F8D8" box="[897,1001,1854,1878]" pageId="2" pageNumber="207" type="holotype">Holotype</typeStatus>
3 (
<collectionCode id="ED62C5D21425FF84FBF7063B184EF8D8" box="[1052,1127,1854,1878]" httpUri="http://grbio.org/cool/3ndm-kie9" name="Chinese Academy of Sciences, Shenyang" pageId="2" pageNumber="207">CASS</collectionCode>
), labelled: “
<collectingCountry id="F3641D871425FF84FB1F063B1914F8D8" box="[1268,1341,1854,1878]" name="China" pageId="2" pageNumber="207">China</collectingCountry>
: Hainan Province, Changjiang County, Bawangling Natural Reserve,
<quantity id="4C8BF0F21425FF84FCBC06671F8DF8F4" box="[855,932,1890,1914]" metricMagnitude="2" metricUnit="m" metricValue="6.09" pageId="2" pageNumber="207" unit="m" value="609.0">609 m</quantity>
,
<date id="FFCD7BD71425FF84FC580667186DF8F4" box="[947,1092,1890,1914]" pageId="2" pageNumber="207" value="2007-03-13">13. iii. 2007</date>
, Yanbin Liu leg.”, “
<typeStatus id="54C8E3B51425FF84FAD9066719B4F8F4" box="[1330,1437,1890,1914]" pageId="2" pageNumber="207" type="holotype">Holotype</typeStatus>
<taxonomicName id="4C7326941425FF84FF7C06821E2BF810" authority="Bian et Ji" authorityName="Bian et Ji" box="[151,514,1926,1950]" class="Insecta" family="Dytiscidae" genus="Neptosternus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="207" phylum="Arthropoda" rank="species" species="yanbini">Neptosternus yanbini Bian et Ji</taxonomicName>
”.
<typeStatus id="54C8E3B51425FF84FDF006821EA5F811" box="[539,652,1927,1951]" pageId="2" pageNumber="207" type="paratype">Paratypes</typeStatus>
: 6 exs., same label data as the
<typeStatus id="54C8E3B51425FF84FC190683187EF810" box="[1010,1111,1926,1950]" pageId="2" pageNumber="207" type="holotype">holotype</typeStatus>
, but “
<typeStatus id="54C8E3B51425FF84FB7006831929F810" box="[1179,1280,1926,1950]" pageId="2" pageNumber="207" type="paratype">Paratype</typeStatus>
<taxonomicName id="4C7326941425FF84FAE306821D74F84C" authority="Bian et Ji" authorityName="Bian et Ji" class="Insecta" family="Dytiscidae" genus="Neptosternus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="207" phylum="Arthropoda" rank="species" species="yanbini">Neptosternus yanbini Bian et Ji</taxonomicName>
” (
<collectionCode id="ED62C5D21425FF84FE9C06AF1DEAF84C" box="[375,451,1962,1986]" httpUri="http://grbio.org/cool/3ndm-kie9" name="Chinese Academy of Sciences, Shenyang" pageId="2" pageNumber="207">CASS</collectionCode>
).
</paragraph>
<paragraph id="8BCC5D171424FF85FF7C04C01D78FA74" blockId="3.[151,1435,1477,1530]" pageId="3" pageNumber="208">
<emphasis id="B90781051424FF85FF7C04C01D12FA55" bold="true" box="[151,315,1477,1500]" pageId="3" pageNumber="208">FIGURES 3–9.</emphasis>
Habitus of
<taxonomicName id="4C7326941424FF85FE5804C21E11FA52" box="[435,568,1479,1500]" class="Insecta" family="Dytiscidae" genus="Neptosternus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="208" phylum="Arthropoda" rank="genus">
<emphasis id="B90781051424FF85FE5804C21E11FA52" box="[435,568,1479,1500]" italics="true" pageId="3" pageNumber="208">Neptosternus</emphasis>
</taxonomicName>
. 3–4.
<taxonomicName id="4C7326941424FF85FD9204C21EDEFA52" box="[633,759,1478,1500]" class="Insecta" family="Dytiscidae" genus="Neptosternus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="208" phylum="Arthropoda" rank="species" species="wewalkai">
<emphasis id="B90781051424FF85FD9204C21EDEFA52" box="[633,759,1478,1500]" italics="true" pageId="3" pageNumber="208">N. wewalkai</emphasis>
</taxonomicName>
; 5.
<taxonomicName id="4C7326941424FF85FCF404C21F94FA52" box="[799,957,1478,1500]" class="Insecta" family="Dytiscidae" genus="Neptosternus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="208" phylum="Arthropoda" rank="species" species="hydaticoides">
<emphasis id="B90781051424FF85FCF404C21F94FA52" box="[799,957,1478,1500]" italics="true" pageId="3" pageNumber="208">N. hydaticoides</emphasis>
</taxonomicName>
; 6.
<taxonomicName id="4C7326941424FF85FC0F04C21870FA52" box="[996,1113,1479,1500]" class="Insecta" family="Dytiscidae" genus="Neptosternus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="208" phylum="Arthropoda" rank="species" species="coomani">
<emphasis id="B90781051424FF85FC0F04C21870FA52" box="[996,1113,1479,1500]" italics="true" pageId="3" pageNumber="208">N. coomani</emphasis>
</taxonomicName>
; 7.
<taxonomicName id="4C7326941424FF85FB6904C218C1FA52" box="[1154,1256,1478,1500]" class="Insecta" family="Dytiscidae" genus="Neptosternus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="208" phylum="Arthropoda" rank="species" species="strnadi">
<emphasis id="B90781051424FF85FB6904C218C1FA52" box="[1154,1256,1478,1500]" italics="true" pageId="3" pageNumber="208">N. strnadi</emphasis>
</taxonomicName>
; 8.
<taxonomicName id="4C7326941424FF85FAE404C219BCFA52" box="[1295,1429,1478,1500]" class="Insecta" family="Dytiscidae" genus="Neptosternus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="208" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="B90781051424FF85FAE404C219BCFA52" box="[1295,1429,1478,1500]" italics="true" pageId="3" pageNumber="208">N. maculatus</emphasis>
</taxonomicName>
. Scale bar =
<quantity id="4C8BF0F21424FF85FEE404E01D64FA74" box="[271,333,1509,1530]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="208" unit="mm" value="1.0">1 mm</quantity>
.
</paragraph>
<paragraph id="8BCC5D171424FF85FF2C07231847F925" blockId="3.[151,1436,1574,2031]" pageId="3" pageNumber="208">
<emphasis id="B90781051424FF85FF2C07231E1DF9B1" bold="true" box="[199,564,1574,1599]" pageId="3" pageNumber="208">Additional material examined.</emphasis>
1Ƥ,
<collectingCountry id="F3641D871424FF85FD8707221E9BF9B1" box="[620,690,1575,1599]" name="China" pageId="3" pageNumber="208">China</collectingCountry>
, Guangdong Province, Dinghu Mountain,
<date id="FFCD7BD71424FF85FB640722193BF9B1" box="[1167,1298,1575,1599]" pageId="3" pageNumber="208" value="1964-07-09">9. vii. 1964</date>
, Entian. Wu leg.; 1Ƥ, same data, but BangFa Lin leg.; 2ƤƤ,
<collectingCountry id="F3641D871424FF85FD25074E1F3AF9ED" box="[718,787,1611,1635]" name="China" pageId="3" pageNumber="208">China</collectingCountry>
, Guangdong Province, Qujiang County, Guitou, autumn 1942, Mengyi Mo leg. [identified as
<taxonomicName id="4C7326941424FF85FDAB07751F14F909" box="[576,829,1648,1671]" class="Insecta" family="Dytiscidae" genus="Neptosternus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="208" phylum="Arthropoda" rank="species" species="coomani">
<emphasis id="B90781051424FF85FDAB07751F14F909" box="[576,829,1648,1671]" italics="true" pageId="3" pageNumber="208">Neptosternus coomani</emphasis>
</taxonomicName>
by H. Zeng]; 13 exs., Guangdong Province, Zhuhai, Zhuhai Campus of Sun Yat-sen University,
<date id="FFCD7BD71424FF85FD9507961F3EF925" box="[638,791,1683,1707]" pageId="3" pageNumber="208" value="2011-07-05" valueMax="2011-07-08" valueMin="2011-07-05">5-8. vii. 2011</date>
, Fenglong Jia leg. (all
<collectionCode id="ED62C5D21424FF85FBFE07961848F925" LSID="urn:lsid:biocol.org:col:13728" box="[1045,1121,1683,1707]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13728" name="National Sun Yat-Sen University, Department of Biological Sciences" pageId="3" pageNumber="208">SYSU</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C3690E9C1424FF83FF2C07B21F99FF5A" lastPageId="5" lastPageNumber="210" pageId="3" pageNumber="208" type="discussion">
<paragraph id="8BCC5D171424FF85FF2C07B21EACF8D1" blockId="3.[151,1436,1574,2031]" pageId="3" pageNumber="208">
<emphasis id="B90781051424FF85FF2C07B21D19F941" bold="true" box="[199,304,1719,1743]" pageId="3" pageNumber="208">Remark.</emphasis>
<bibRefCitation id="EFE220E61424FF85FED307B21DD3F941" author="Bian" box="[312,506,1719,1743]" pageId="3" pageNumber="208" refString="Bian, D. &amp; Ji, L. (2009) Two new species of Dytiscidae from Hainan, China. The Coleopterists Bulletin, 63, 35 - 40." type="journal article" year="2009">Bian &amp; Ji (2009)</bibRefCitation>
described a new species
<taxonomicName id="4C7326941424FF85FCF607BD1824F941" box="[797,1037,1719,1743]" class="Insecta" family="Dytiscidae" genus="Neptosternus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="208" phylum="Arthropoda" rank="species" species="yanbini">
<emphasis id="B90781051424FF85FCF607BD1824F941" box="[797,1037,1719,1743]" italics="true" pageId="3" pageNumber="208">Neptosternus yanbini</emphasis>
</taxonomicName>
from Hainan,
<collectingCountry id="F3641D871424FF85FB5E07B218D5F941" box="[1205,1276,1719,1743]" name="China" pageId="3" pageNumber="208">China</collectingCountry>
. Based on the original description, the species was only compared with
<taxonomicName id="4C7326941424FF85FCC207D91FAEF97D" box="[809,903,1756,1779]" class="Insecta" family="Dytiscidae" genus="Neptosternus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="208" phylum="Arthropoda" rank="species" species="pocsi">
<emphasis id="B90781051424FF85FCC207D91FAEF97D" box="[809,903,1756,1779]" italics="true" pageId="3" pageNumber="208">N. pocsi</emphasis>
</taxonomicName>
, a species occurring in
<collectingCountry id="F3641D871424FF85FB7307DE18DEF97D" box="[1176,1271,1755,1779]" name="Vietnam" pageId="3" pageNumber="208">Vietnam</collectingCountry>
and
<collectingCountry id="F3641D871424FF85FADA07DE19BEF97D" box="[1329,1431,1755,1779]" name="Thailand" pageId="3" pageNumber="208">Thailand</collectingCountry>
, from which
<taxonomicName id="4C7326941424FF85FECE06051DB5F899" box="[293,412,1791,1815]" class="Insecta" family="Dytiscidae" genus="Neptosternus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="208" phylum="Arthropoda" rank="species" species="yanbini">
<emphasis id="B90781051424FF85FECE06051DB5F899" box="[293,412,1791,1815]" italics="true" pageId="3" pageNumber="208">N. yanbini</emphasis>
</taxonomicName>
differs in its narrower habitus and the male genitalia. However, examination of the
<typeStatus id="54C8E3B51424FF85FA87060519B5F896" box="[1388,1436,1792,1816]" pageId="3" pageNumber="208">type</typeStatus>
series of
<taxonomicName id="4C7326941424FF85FEE806211D52F8B5" box="[259,379,1827,1851]" class="Insecta" family="Dytiscidae" genus="Neptosternus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="208" phylum="Arthropoda" rank="species" species="yanbini">
<emphasis id="B90781051424FF85FEE806211D52F8B5" box="[259,379,1827,1851]" italics="true" pageId="3" pageNumber="208">N. yanbini</emphasis>
</taxonomicName>
by the senior author revealed no significant difference between
<taxonomicName id="4C7326941424FF85FB86062118CFF8B5" box="[1133,1254,1827,1851]" class="Insecta" family="Dytiscidae" genus="Neptosternus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="208" phylum="Arthropoda" rank="species" species="yanbini">
<emphasis id="B90781051424FF85FB86062118CFF8B5" box="[1133,1254,1827,1851]" italics="true" pageId="3" pageNumber="208">N. yanbini</emphasis>
</taxonomicName>
and the widely distributed Oriental species
<taxonomicName id="4C7326941424FF85FE3B064D1E57F8D1" box="[464,638,1863,1887]" class="Insecta" family="Dytiscidae" genus="Neptosternus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="208" phylum="Arthropoda" rank="species" species="hydaticoides">
<emphasis id="B90781051424FF85FE3B064D1E57F8D1" box="[464,638,1863,1887]" italics="true" pageId="3" pageNumber="208">N. hydaticoides</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BCC5D171424FF85FF2E066E1824F861" blockId="3.[151,1436,1574,2031]" pageId="3" pageNumber="208">
<bibRefCitation id="EFE220E61424FF85FF2E066E1DAFF80D" author="Bian" box="[197,390,1899,1923]" pageId="3" pageNumber="208" refString="Bian, D. &amp; Ji, L. (2009) Two new species of Dytiscidae from Hainan, China. The Coleopterists Bulletin, 63, 35 - 40." type="journal article" year="2009">Bian &amp; Ji (2009)</bibRefCitation>
reported a rather small body size for their specimens of
<taxonomicName id="4C7326941424FF85FBE0066918A8F80D" box="[1035,1153,1899,1923]" class="Insecta" family="Dytiscidae" genus="Neptosternus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="208" phylum="Arthropoda" rank="species" species="yanbini">
<emphasis id="B90781051424FF85FBE0066918A8F80D" box="[1035,1153,1899,1923]" italics="true" pageId="3" pageNumber="208">N. yanbini</emphasis>
</taxonomicName>
(TL:
<quantity id="4C8BF0F21424FF85FB2F066E1967F80D" box="[1220,1358,1899,1923]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="2.9" metricValueMin="2.6" pageId="3" pageNumber="208" unit="mm" value="2.75" valueMax="2.9" valueMin="2.6">2.6–2.9 mm</quantity>
) while Balke &amp; Hendrich (1997) gave
<quantity id="4C8BF0F21424FF85FE17068A1EB6F829" box="[508,671,1935,1959]" metricMagnitude="-3" metricUnit="m" metricValue="3.275" metricValueMax="3.3" metricValueMin="3.25" pageId="3" pageNumber="208" unit="mm" value="3.275" valueMax="3.3" valueMin="3.25">3.25–3.30 mm</quantity>
for their material of
<taxonomicName id="4C7326941424FF85FC6006951813F829" box="[907,1082,1935,1959]" class="Insecta" family="Dytiscidae" genus="Neptosternus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="208" phylum="Arthropoda" rank="species" species="hydaticoides">
<emphasis id="B90781051424FF85FC6006951813F829" box="[907,1082,1935,1959]" italics="true" pageId="3" pageNumber="208">N. hydaticoides</emphasis>
</taxonomicName>
. We can report a TL of
<quantity id="4C8BF0F21424FF85FAA3068A1CECF845" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="3.3" metricValueMin="2.7" pageId="3" pageNumber="208" unit="mm" value="3.0" valueMax="3.3" valueMin="2.7">2.7–3.3 mm</quantity>
in the extensive material of
<taxonomicName id="4C7326941424FF85FDE106B11E90F845" box="[522,697,1971,1995]" class="Insecta" family="Dytiscidae" genus="Neptosternus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="208" phylum="Arthropoda" rank="species" species="hydaticoides">
<emphasis id="B90781051424FF85FDE106B11E90F845" box="[522,697,1971,1995]" italics="true" pageId="3" pageNumber="208">N. hydaticoides</emphasis>
</taxonomicName>
available to us, from the whole area of its distribution. Thus, we attribute the slightly smaller body size of
<taxonomicName id="4C7326941424FF85FD8006DD1EF6F861" box="[619,735,2007,2031]" class="Insecta" family="Dytiscidae" genus="Neptosternus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="208" phylum="Arthropoda" rank="species" species="yanbini">
<emphasis id="B90781051424FF85FD8006DD1EF6F861" box="[619,735,2007,2031]" italics="true" pageId="3" pageNumber="208">N. yanbini</emphasis>
</taxonomicName>
to intraspecific variability.
</paragraph>
<caption id="DF0C0D9F1423FF82FF7C068C1F65F830" httpUri="https://zenodo.org/record/282254/files/figure.png" pageId="4" pageNumber="209" targetBox="[208,1366,208,1868]" targetPageId="4">
<paragraph id="8BCC5D171423FF82FF7C068C1F65F830" blockId="4.[151,1437,1929,1982]" pageId="4" pageNumber="209">
<emphasis id="B90781051423FF82FF7C068C1D63F811" bold="true" box="[151,330,1929,1952]" pageId="4" pageNumber="209">FIGURES 9–13.</emphasis>
Median lobe of aedeagus in ventral view. 9—
<taxonomicName id="4C7326941423FF82FCD9068E180DF82E" box="[818,1060,1931,1952]" class="Insecta" family="Dytiscidae" genus="Neptosternus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="209" phylum="Arthropoda" rank="species" species="punctatus" status="sp. nov.">
<emphasis id="B90781051423FF82FCD9068E180DF82E" box="[818,1060,1931,1952]" italics="true" pageId="4" pageNumber="209">Neptosternus punctatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2343C7E1423FF82FBC7068F18A9F811" box="[1068,1152,1930,1951]" pageId="4" pageNumber="209" rank="species">
<emphasis id="B90781051423FF82FBC7068F18A9F811" bold="true" box="[1068,1152,1930,1951]" pageId="4" pageNumber="209">sp. nov.</emphasis>
</taxonomicNameLabel>
; 10—
<taxonomicName id="4C7326941423FF82FB28068E196AF811" box="[1219,1347,1929,1952]" class="Insecta" family="Dytiscidae" genus="Neptosternus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="209" phylum="Arthropoda" rank="species" species="wewalkai">
<emphasis id="B90781051423FF82FB28068E196AF811" box="[1219,1347,1929,1952]" italics="true" pageId="4" pageNumber="209">N. wewalkai</emphasis>
</taxonomicName>
; 11—
<taxonomicName id="4C7326941423FF82FA6F068E1D30F830" class="Insecta" family="Dytiscidae" genus="Neptosternus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="209" phylum="Arthropoda" rank="species" species="hydaticoides">
<emphasis id="B90781051423FF82FA6F068E1D30F830" italics="true" pageId="4" pageNumber="209">N. hydaticoides</emphasis>
</taxonomicName>
; 12—
<taxonomicName id="4C7326941423FF82FEB206AC1DE4F830" box="[345,461,1961,1982]" class="Insecta" family="Dytiscidae" genus="Neptosternus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="209" phylum="Arthropoda" rank="species" species="coomani">
<emphasis id="B90781051423FF82FEB206AC1DE4F830" box="[345,461,1961,1982]" italics="true" pageId="4" pageNumber="209">N. coomani</emphasis>
</taxonomicName>
; 13—
<taxonomicName id="4C7326941423FF82FDE606AC1E5AF830" box="[525,627,1960,1982]" class="Insecta" family="Dytiscidae" genus="Neptosternus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="209" phylum="Arthropoda" rank="species" species="strnadi">
<emphasis id="B90781051423FF82FDE606AC1E5AF830" box="[525,627,1960,1982]" italics="true" pageId="4" pageNumber="209">N. strnadi</emphasis>
</taxonomicName>
. Scale bar = 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BCC5D171422FF83FF2E01921F99FF5A" blockId="5.[151,1436,151,285]" pageId="5" pageNumber="210">
This opinion has been confirmed by a recent study by D. Bian (personal communication). Therefore we consider
<taxonomicName id="4C7326941422FF83FF1401B81D5AFF5A" box="[255,371,188,212]" class="Insecta" family="Dytiscidae" genus="Neptosternus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="210" phylum="Arthropoda" rank="species" species="yanbini">
<emphasis id="B90781051422FF83FF1401B81D5AFF5A" box="[255,371,188,212]" italics="true" pageId="5" pageNumber="210">N. yanbini</emphasis>
</taxonomicName>
as a junior subjective synonym of
<taxonomicName id="4C7326941422FF83FD1101B81F81FF5A" box="[762,936,188,212]" class="Insecta" family="Dytiscidae" genus="Neptosternus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="210" phylum="Arthropoda" rank="species" species="hydaticoides">
<emphasis id="B90781051422FF83FD1101B81F81FF5A" box="[762,936,188,212]" italics="true" pageId="5" pageNumber="210">N. hydaticoides</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3690E9C1422FF83FF2C01DA18E7FE92" pageId="5" pageNumber="210" type="distribution">
<paragraph id="8BCC5D171422FF83FF2C01DA18E7FE92" blockId="5.[151,1436,151,285]" pageId="5" pageNumber="210">
<emphasis id="B90781051422FF83FF2C01DA1D49FF76" bold="true" box="[199,352,223,248]" pageId="5" pageNumber="210">Distribution.</emphasis>
<taxonomicName id="4C7326941422FF83FE8C01E41EA6FF76" box="[359,655,224,248]" class="Insecta" family="Dytiscidae" genus="Neptosternus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="210" phylum="Arthropoda" rank="species" species="hydaticoides">
<emphasis id="B90781051422FF83FE8C01E41EA6FF76" box="[359,655,224,248]" italics="true" pageId="5" pageNumber="210">Neptosternus hydaticoides</emphasis>
</taxonomicName>
is recorded from
<collectingCountry id="F3641D871422FF83FCBC01DA1FB3FF79" box="[855,922,223,247]" name="China" pageId="5" pageNumber="210">China</collectingCountry>
(Guangdong, Guangxi, Hainan,
<collectingCountry id="F3641D871422FF83FAE301E519A6FF79" box="[1288,1423,223,248]" name="Hong Kong" pageId="5" pageNumber="210">Hong Kong</collectingCountry>
), through
<collectingCountry id="F3641D871422FF83FF1E00001D4DFE93" box="[245,356,261,285]" name="Myanmar" pageId="5" pageNumber="210">Myanmar</collectingCountry>
,
<collectingCountry id="F3641D871422FF83FE8400011DFFFE92" box="[367,470,260,284]" name="Thailand" pageId="5" pageNumber="210">Thailand</collectingCountry>
,
<collectingCountry id="F3641D871422FF83FE0B00011E16FE92" box="[480,575,260,284]" name="Vietnam" pageId="5" pageNumber="210">Vietnam</collectingCountry>
and peninsular
<collectingCountry id="F3641D871422FF83FD1A00011F71FE92" box="[753,856,260,284]" name="Malaysia" pageId="5" pageNumber="210">Malaysia</collectingCountry>
to
<collectingCountry id="F3641D871422FF83FC9000011FC1FE92" box="[891,1000,260,284]" name="Indonesia" pageId="5" pageNumber="210">Indonesia</collectingCountry>
and the
<collectingCountry id="F3641D871422FF83FBA3000118E3FE92" box="[1096,1226,260,284]" name="Philippines" pageId="5" pageNumber="210">Philippines</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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