158 lines
19 KiB
XML
158 lines
19 KiB
XML
<document id="C3548170F039BC21A4ABF9487AF42A2D" ID-DOI="10.5281/zenodo.13203160" ID-ISSN="1407-8619" ID-Zenodo-Dep="13203160" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722657382176" checkinUser="felipe" docAuthor="Fanti, Fabrizio & Müller, Patrick" docDate="2022" docId="03DAD52BFF87FFC9FF0DB8DD5CE7A5B4" docLanguage="en" docName="BalticJColeopterol.22.2.331-380.pdf" docOrigin="Baltic Journal of Coleopterology 22 (2)" docStyle="DocumentStyle:9720869F1FB170B2A509257D1FA394CD.2:BalticJColeop.2022.journal_article" docStyleId="9720869F1FB170B2A509257D1FA394CD" docStyleName="BalticJColeop.2022.journal_article" docStyleVersion="2" docTitle="Sanaungulus perkovskyi Fanti & Müller, 2022, sp. nov." docType="treatment" docVersion="3" lastPageNumber="366" masterDocId="FFE3AD53FFA5FFEAFF9ABD325F46A170" masterDocTitle="Fossil Cantharidae from the Cretaceous Burmese (Kachin) amber of the Patrick Müller collection, and taxonomic information" masterLastPageNumber="380" masterPageNumber="331" pageNumber="365" updateTime="1722697113453" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="19B3F5770EB0FE8A77F21D43820C0F8D">Fossil Cantharidae from the Cretaceous Burmese (Kachin) amber of the Patrick Müller collection, and taxonomic information</mods:title>
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<mods:namePart id="4CF86ADDFD680695FEFA4533B54714B6">Fanti, Fabrizio</mods:namePart>
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<mods:namePart id="78A6260714C542C159FA5950F04E5849">Müller, Patrick</mods:namePart>
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<mods:typeOfResource id="C203E17E677B870B5700E4819FF59958">text</mods:typeOfResource>
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<mods:title id="D9F66FDECB5E2062C09ED18C9794D250">Baltic Journal of Coleopterology</mods:title>
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</mods:titleInfo>
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<mods:part id="3A7C318D0EB8D7121C3C7B48D7131B0F">
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<mods:date id="441BE9CC3AAEFBC41399F9B35CAEF415">2022</mods:date>
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<mods:detail id="843233FEA304245577E1BD840CA6003C" type="volume">
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<mods:number id="F08BAE7883D919823A28205C9D3FB8D4">22</mods:number>
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<mods:identifier id="E49076818011C7708C47F1A50308ED58" type="DOI">10.5281/zenodo.13203160</mods:identifier>
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<mods:identifier id="7C10178F650FABF2579C0F42856B012D" type="ISSN">1407-8619</mods:identifier>
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<treatment id="03DAD52BFF87FFC9FF0DB8DD5CE7A5B4" ID-DOI="http://doi.org/10.5281/zenodo.13199833" ID-Zenodo-Dep="13199833" LSID="urn:lsid:plazi:treatment:03DAD52BFF87FFC9FF0DB8DD5CE7A5B4" httpUri="http://treatment.plazi.org/id/03DAD52BFF87FFC9FF0DB8DD5CE7A5B4" lastPageId="35" lastPageNumber="366" pageId="34" pageNumber="365">
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<subSubSection id="C36937B6FF87FFC8FF0DB8DD5D47A777" box="[151,513,1519,1543]" pageId="34" pageNumber="365" type="nomenclature">
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<paragraph id="8BCC643DFF87FFC8FF0DB8DD5D47A777" blockId="34.[151,513,1519,1574]" box="[151,513,1519,1543]" pageId="34" pageNumber="365">
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<heading id="D084D351FF87FFC8FF0DB8DD5D47A777" bold="true" box="[151,513,1519,1543]" fontSize="10" level="2" pageId="34" pageNumber="365" reason="4">
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<emphasis id="B907B82FFF87FFC8FF0DB8DD5D47A777" bold="true" box="[151,513,1519,1543]" pageId="34" pageNumber="365">
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<taxonomicName id="4C731FBEFF87FFC8FF0DB8DD5EE0A777" authorityName="Fanti & Müller" authorityYear="2022" box="[151,422,1519,1543]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="365" phylum="Arthropoda" rank="species" species="perkovskyi" status="sp. nov.">
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<emphasis id="B907B82FFF87FFC8FF0DB8DD5EE0A777" bold="true" box="[151,422,1519,1543]" italics="true" pageId="34" pageNumber="365">Sanaungulus perkovskyi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2340554FF87FFC8FE36B8DD5D47A777" box="[428,513,1519,1543]" pageId="34" pageNumber="365" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C36937B6FF87FFC8FF0DBB3D5FB2A756" box="[151,244,1551,1574]" pageId="34" pageNumber="365" type="description">
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<paragraph id="8BCC643DFF87FFC8FF0DBB3D5FB2A756" blockId="34.[151,513,1519,1574]" box="[151,244,1551,1574]" pageId="34" pageNumber="365">
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(
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<figureCitation id="134878B8FF87FFC8FF05BB3D5FAAA756" box="[159,236,1551,1574]" captionStart="Fig" captionStartId="34.[151,189,1409,1432]" captionTargetBox="[151,1172,151,1376]" captionTargetId="figure-73@34.[151,1172,151,1376]" captionTargetPageId="34" captionText="Fig.19. Sanaungulus perkovskyi sp. nov. in Burmese amber.A: Holotype, dorsal view; B: Holotype, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13203217" httpUri="https://zenodo.org/record/13203217/files/figure.png" pageId="34" pageNumber="365">Fig. 19</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C36937B6FF87FFC9FF0DBB775ED6A1A3" lastPageId="35" lastPageNumber="366" pageId="34" pageNumber="365" type="materials_examined">
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<paragraph id="8BCC643DFF87FFC8FF0DBB775B13A756" blockId="34.[151,643,1605,1693]" lastBlockId="34.[680,1171,1519,1574]" pageId="34" pageNumber="365">
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<materialsCitation id="3B1B6E60FF87FFC8FF0DBB775B13A756" collectingDate="2021-12" collectionCode="BUB" county="Male" location="Catalog" municipality="Burmese" pageId="34" pageNumber="365" specimenCode="BUB3888" specimenCount="1" stateProvince="Kachin" typeStatus="holotype">
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<emphasis id="B907B82FFF87FFC8FF0DBB775E40A72D" bold="true" box="[151,262,1605,1629]" pageId="34" pageNumber="365">
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<typeStatus id="54C8DA9FFF87FFC8FF0DBB775E47A72D" box="[151,257,1605,1629]" pageId="34" pageNumber="365" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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<collectingCounty id="62AD1CB1FF87FFC8FE96BB745E03A72D" box="[268,325,1606,1629]" pageId="34" pageNumber="365">Male</collectingCounty>
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, adult specimen in a
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<collectingMunicipality id="6BA8FE47FF87FFC8FDBEBB745DC5A72D" box="[548,643,1606,1629]" pageId="34" pageNumber="365">Burmese</collectingMunicipality>
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(
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<collectingRegion id="49B7AADFFF87FFC8FF05BB545FA9A70D" box="[159,239,1638,1661]" country="Myanmar" name="Kachin" pageId="34" pageNumber="365">Kachin</collectingRegion>
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) amber piece:
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<location id="8EAC32E6FF87FFC8FE12BB545E9DA70D" LSID="urn:lsid:plazi:treatment:03DAD52BFF87FFC9FF0DB8DD5CE7A5B4:8EAC32E6FF87FFC8FE12BB545E9DA70D" box="[392,475,1638,1661]" county="Male" municipality="Burmese" name="Catalog" pageId="34" pageNumber="365" stateProvince="Kachin">Catalog</location>
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number SNSBBSPG
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<date id="FFCD42FDFF87FFC8FF79BBB45E01A7ED" box="[227,327,1670,1693]" pageId="34" pageNumber="365" value="2021-12">
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<collectingDate id="EF89BB15FF87FFC8FF79BBB45E01A7ED" box="[227,327,1670,1693]" pageId="34" pageNumber="365" value="2021-12">2021 XII</collectingDate>
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</date>
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<quantity id="4C8BC9D8FF87FFC8FECABBB45ECEA7ED" box="[336,392,1670,1693]" metricMagnitude="0" metricUnit="m" metricValue="1.7526" pageId="34" pageNumber="365" unit="in" value="69.0">69 in</quantity>
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the
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<location id="8EAC32E6FF87FFC8FE21BBB45C51A776" LSID="urn:lsid:plazi:treatment:03DAD52BFF87FFC9FF0DB8DD5CE7A5B4:8EAC32E6FF87FFC8FE21BBB45C51A776" box="[443,791,1519,1693]" county="Male" municipality="Burmese" name="Bayerische Staatssammlung" pageId="34" pageNumber="365" stateProvince="Kachin">Bayerische Staatssammlung</location>
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für
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<location id="8EAC32E6FF87FFC8FCD9B8DD5C90A776" LSID="urn:lsid:plazi:treatment:03DAD52BFF87FFC9FF0DB8DD5CE7A5B4:8EAC32E6FF87FFC8FCD9B8DD5C90A776" box="[835,982,1519,1542]" county="Male" municipality="Burmese" name="Palaontologie" pageId="34" pageNumber="365" stateProvince="Kachin">Paläontologie</location>
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und
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<location id="8EAC32E6FF87FFC8FB90B8DD5B2AA776" LSID="urn:lsid:plazi:treatment:03DAD52BFF87FFC9FF0DB8DD5CE7A5B4:8EAC32E6FF87FFC8FB90B8DD5B2AA776" box="[1034,1132,1519,1542]" county="Male" municipality="Burmese" name="Geologie" pageId="34" pageNumber="365" stateProvince="Kachin">Geologie</location>
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(ex
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<location id="8EAC32E6FF87FFC8FD32BB3D5C42A756" LSID="urn:lsid:plazi:treatment:03DAD52BFF87FFC9FF0DB8DD5CE7A5B4:8EAC32E6FF87FFC8FD32BB3D5C42A756" box="[680,772,1551,1574]" county="Male" municipality="Burmese" name="Muller's" pageId="34" pageNumber="365" stateProvince="Kachin">Müller’s</location>
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collection number:
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<specimenCode id="DBD5CC46FF87FFC8FC43BB3D5B0FA756" box="[985,1097,1551,1574]" collectionCode="BUB" pageId="34" pageNumber="365">BUB3888</specimenCode>
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).
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</materialsCitation>
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||
</paragraph>
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||
<paragraph id="8BCC643DFF87FFC8FD32BB775CBDA70D" blockId="34.[680,1172,1605,1661]" pageId="34" pageNumber="365">
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<emphasis id="B907B82FFF87FFC8FD32BB775C07A72D" bold="true" box="[680,833,1605,1629]" pageId="34" pageNumber="365">Type locality.</emphasis>
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<collectingCountry id="F36424ADFF87FFC8FCD0BB745CF0A72D" box="[842,950,1606,1629]" name="Myanmar" pageId="34" pageNumber="365">Myanmar</collectingCountry>
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:
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<collectingRegion id="49B7AADFFF87FFC8FC5BBB745B0DA72D" box="[961,1099,1606,1629]" country="Myanmar" name="Kachin" pageId="34" pageNumber="365">Kachin state</collectingRegion>
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, Myitkyina District, Hukawng Valley.
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</paragraph>
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<paragraph id="8BCC643DFF86FFC9FFEBBDA95ED6A1A3" blockId="35.[113,605,155,211]" pageId="35" pageNumber="366">
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<emphasis id="B907B82FFF86FFC9FFEBBDA95E43A1C3" bold="true" box="[113,261,155,179]" pageId="35" pageNumber="366">Type horizon.</emphasis>
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Lowermost Cenomanian (98.79 ± 0.62 Ma), midCretaceous.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C36937B6FF86FFC9FFEBBDC15D15A3FA" pageId="35" pageNumber="366" type="diagnosis">
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||
<paragraph id="8BCC643DFF86FFC9FFEBBDC15D15A3FA" blockId="35.[113,606,243,650]" pageId="35" pageNumber="366">
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<emphasis id="B907B82FFF86FFC9FFEBBDC15E26A07B" bold="true" box="[113,352,243,267]" pageId="35" pageNumber="366">Differential diagnosis.</emphasis>
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<taxonomicName id="4C731FBEFF86FFC9FEFFBDC15D1BA07A" authorityName="Fanti & Müller" authorityYear="2022" box="[357,605,243,266]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="366" phylum="Arthropoda" rank="species" species="perkovskyi" status="sp. nov.">
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<emphasis id="B907B82FFF86FFC9FEFFBDC15D1BA07A" box="[357,605,243,266]" italics="true" pageId="35" pageNumber="366">Sanaungulus perkovskyi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2340554FF86FFC9FFEBBC215F86A05A" box="[113,192,275,298]" pageId="35" pageNumber="366" rank="species">sp. nov.</taxonomicNameLabel>
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is characterized by a strongly transverse pronotum and pectinate antennae (the antennal process in antennomeres IV–VII).
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<taxonomicName id="4C731FBEFF86FFC9FE4FBC615F93A0DA" authority="Fanti, Damgaard & Ellenberger, 2018" authorityName="Fanti, Damgaard & Ellenberger" authorityYear="2018" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="366" phylum="Arthropoda" rank="species" species="ghitaenoerbyae">
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<emphasis id="B907B82FFF86FFC9FE4FBC615E51A0FA" italics="true" pageId="35" pageNumber="366">Sanaungulus ghitaenoerbyae</emphasis>
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<bibRefCitation id="EFE219CCFF86FFC9FEBABC415F93A0DA" author="Fanti F. & Damgaard A. L. & Ellenberger S." pageId="35" pageNumber="366" pagination="170 - 177" refId="ref20442" refString="Fanti F., Damgaard A. L., Ellenberger S. 2018. Two new genera of Cantharidae from Burmese amber of the Hukawng Valley (Insecta, Coleoptera). Cretaceous Research 86: 170 - 177." type="journal article" year="2018">Fanti, Damgaard & Ellenberger, 2018</bibRefCitation>
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</taxonomicName>
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is the most similar taxon and differs from the new species because of its more transverse head and its pronotum with its sides and posterior margin not straight (Fanti et al. 2018).
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<taxonomicName id="4C731FBEFF86FFC9FFEBBF215FA3A33A" authority="Fanti & Damgaard, 2019" authorityName="Fanti & Damgaard" authorityYear="2019" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="366" phylum="Arthropoda" rank="species" species="troelsikloevedali">
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<emphasis id="B907B82FFF86FFC9FFEBBF215EF0A35A" box="[113,438,531,554]" italics="true" pageId="35" pageNumber="366">Sanaungulus troelsikloevedali</emphasis>
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<bibRefCitation id="EFE219CCFF86FFC9FE5BBF215FA3A33A" author="Fanti F. & Damgaard A. L." pageId="35" pageNumber="366" pagination="101 - 125" refId="ref20361" refString="Fanti F., Damgaard A. L. 2019. New soldier beetles (Cantharidae) from Baltic, Burmese and Dominican ambers of the Anders Damgaard amber collection. Baltic Journal of Coleopterology 19 (2): 101 - 125." type="journal article" year="2019">Fanti & Damgaard, 2019</bibRefCitation>
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</taxonomicName>
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differs from the new species described here by its longer antennae with more slender last antennomeres (
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<bibRefCitation id="EFE219CCFF86FFC9FEA4BF415D01A3FA" author="Fanti F. & Damgaard A. L." box="[318,583,627,650]" pageId="35" pageNumber="366" pagination="101 - 125" refId="ref20361" refString="Fanti F., Damgaard A. L. 2019. New soldier beetles (Cantharidae) from Baltic, Burmese and Dominican ambers of the Anders Damgaard amber collection. Baltic Journal of Coleopterology 19 (2): 101 - 125." type="journal article" year="2019">Fanti & Damgaard 2019</bibRefCitation>
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).
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C36937B6FF86FFC9FFEBBF9B5CEAA2EE" pageId="35" pageNumber="366" type="description">
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||
<paragraph id="8BCC643DFF86FFC9FFEBBF9B5EC8A251" blockId="35.[113,605,681,801]" pageId="35" pageNumber="366">
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||
<emphasis id="B907B82FFF86FFC9FFEBBF9B5FB0A3B1" bold="true" box="[113,246,681,705]" pageId="35" pageNumber="366">Description.</emphasis>
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Adult, winged, rather robust. Male, defined on the basis of the last abdominal segment triangular shaped. Body length
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<quantity id="4C8BC9D8FF86FFC9FD99BFD85D1FA271" box="[515,601,746,769]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="35" pageNumber="366" unit="mm" value="3.2">3.2 mm</quantity>
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. Entirely testaceousbrown.
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||
</paragraph>
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<paragraph id="8BCC643DFF86FFC9FFEBBE725CEAA2EE" blockId="35.[113,606,832,1687]" lastBlockId="35.[643,1135,155,926]" pageId="35" pageNumber="366">Head not particularly transverse, constricted (triangularshaped) behind the eyes, pubescent with shallow punctation. Eyes prominent, large, roundish, inserted in the upper and lateral part of the head. Mandibles elongated, falciform. Maxillary palps 4 segmented with the last palpomere securiform. Labial palps 3 segmented, with the last palpomere securiform. Antennae 11 segmented, pectinate, approximately slightly surpassing the half of elytra, antennal insertion far from the eyes; scape clubshaped, robust; antennomeres II–III short with antennomere II sturdier and swollen apically; antennomeres IV– VI extremely long, each equipped with an antennal process at apex which is very long and robust and slightly pointed apically; antennomere VII shorter than previous ones, equipped with a long antennal process; antennomere VIII sturdier and shorter than previous one and without the antennal process; antennomeres IX–X very robust, flat, enlarged at sides; antennomere XI filiform, less sturdier than previous ones, with rounded apex; all the antennomeres pubescent with setae very short.Pronotum transverse, wider than head, anterior margin undulate and strongly bordered, posterior margin straight and bordered, sides straight and bordered, corners obtuse, pronotum disc pubescent with shallow punctation. Scutellum triangularshaped, very wide at base, with rounded apex. Elytra short which reveals some abdominal segments, robust, wider than pronotum, posteriorly parallelsided and not dehiscent apically, rounded apexes, surface with coarse and impressed punctation gathered in striae. Metathoracic wings transparent, longer than elytra. Sternum elongated with posterior margin almost straight, sternites transverse and pubescent, last sternite small and triangular shaped. Legs thin, long, slightly pubescent; coxae massive; trochanters small, rounded; femora almost straight, slightly flat, sturdier than tibiae; tibiae, thin, cylindrical, long, without spurs near the apex, pro and mesotibiae very slightly longer than pro and mesofemora, metatibiae longer than metafemora. Tarsal formula 5 5 5; first tarsomere long, about 1.4 times longer than second; third tarsomere short, triangularshaped; fourth strongly bilobed with lobes elongated and rounded apically; fifth tarsomere elongated, thin, cylindrical and curved; claws simple, long, pointed, without denticle at the base.</paragraph>
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||
</subSubSection>
|
||
<subSubSection id="C36937B6FF86FFC9FD19BE8C5B29A55C" pageId="35" pageNumber="366" type="etymology">
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||
<paragraph id="8BCC643DFF86FFC9FD19BE8C5B29A285" blockId="35.[643,1135,958,1013]" pageId="35" pageNumber="366">
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<emphasis id="B907B82FFF86FFC9FD19BE8C5C44A2A6" bold="true" box="[643,770,958,982]" pageId="35" pageNumber="366">Etymology.</emphasis>
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Named in honor of Evgeny E. Perkovsky (
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<collectingCountry id="F36424ADFF86FFC9FD42BEEC5C76A285" box="[728,816,990,1013]" name="Ukraine" pageId="35" pageNumber="366">Ukraine</collectingCountry>
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), a wellknown palaeontologist.
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</paragraph>
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<paragraph id="8BCC643DFF86FFC9FD19B9275B29A55C" blockId="35.[643,1135,1045,1069]" box="[643,1135,1045,1069]" pageId="35" pageNumber="366">
|
||
<emphasis id="B907B82FFF86FFC9FD19B9275C58A55D" bold="true" box="[643,798,1045,1069]" pageId="35" pageNumber="366">Syninclusions.</emphasis>
|
||
Air bubbles (very large), detritus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36937B6FF86FFC9FD19B97E5CE7A5B4" pageId="35" pageNumber="366" type="discussion">
|
||
<paragraph id="8BCC643DFF86FFC9FD19B97E5CE7A5B4" blockId="35.[643,1134,1100,1220]" pageId="35" pageNumber="366">
|
||
<emphasis id="B907B82FFF86FFC9FD19B97E5DB7A514" bold="true" box="[643,753,1100,1124]" pageId="35" pageNumber="366">Remarks.</emphasis>
|
||
Piece of very clear amber that measures 12x8x
|
||
<quantity id="4C8BC9D8FF86FFC9FC9EB95F5C00A5F4" box="[772,838,1133,1156]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="35" pageNumber="366" unit="mm" value="2.0">2 mm</quantity>
|
||
. The inclusion is complete, but many big bubbles in the amber cover the majority of its ventral part.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |