295 lines
40 KiB
XML
295 lines
40 KiB
XML
<document id="2897D2590ABF76A5ACDE5C207918C651" ID-DOI="10.11646/zootaxa.5164.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="6821026" ID-ZooBank="3BECB9C7-F4B5-4FA4-934B-1822BF3D1077" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1657611643055" checkinUser="plazi" docAuthor="Mah, Christopher L." docDate="2022" docId="03CE851E923FE96DEBF94C82FD0EFD2D" docLanguage="en" docName="zootaxa.5164.1.1.pdf" docOrigin="Zootaxa 5164 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5164.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Heligmaster kanaloa Mah 2022, n. gen." docType="treatment" docVersion="3" lastPageNumber="23" masterDocId="FFF7FD66922BE97BEB6E4C15FFC4FFDC" masterDocTitle="New Genera, Species and Occurrences of Deep-Sea Asteroidea (Valvatacea, Forcipulatacea, Echinodermata) collected from the North Pacific Ocean by the CAPSTONE Expedition" masterLastPageNumber="75" masterPageNumber="1" pageNumber="21" updateTime="1699345508031" updateUser="plazi">
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<mods:title id="7413186C227E91390B98F88D45B2697F">New Genera, Species and Occurrences of Deep-Sea Asteroidea (Valvatacea, Forcipulatacea, Echinodermata) collected from the North Pacific Ocean by the CAPSTONE Expedition</mods:title>
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<mods:namePart id="441621AB05F2796A7CEF88F2EF53B891">Mah, Christopher L.</mods:namePart>
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<mods:typeOfResource id="0DE083956BB9D96BAEC9DD8919F69B7A">text</mods:typeOfResource>
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<mods:title id="E0B76E8481834B724BB2584814EDEF7D">Zootaxa</mods:title>
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<mods:part id="C6916CBDCD3D90E888E3B184A279F08E">
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<mods:date id="4F924D46765C55636503FD39E0EC1987">2022</mods:date>
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<mods:number id="5B807C7696F267C0DCDC99378AF8CAAC">2022-07-12</mods:number>
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<mods:number id="C205779C19F500235C705627226F7202">5164</mods:number>
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<mods:url id="56CD3DDD97D1596760A6D2CC6519DC61">http://dx.doi.org/10.11646/zootaxa.5164.1.1</mods:url>
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<mods:classification id="730D37E268DB1A0D153CFFA4C786DE0A">journal article</mods:classification>
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<mods:identifier id="3D4297EEA3AF4EFC21F632996E20A1F6" type="DOI">10.11646/zootaxa.5164.1.1</mods:identifier>
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<mods:identifier id="9D707B1225009EE64CF56B4583D53C17" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03CE851E923FE96DEBF94C82FD0EFD2D" LSID="urn:lsid:plazi:treatment:03CE851E923FE96DEBF94C82FD0EFD2D" httpUri="http://treatment.plazi.org/id/03CE851E923FE96DEBF94C82FD0EFD2D" lastPageId="22" lastPageNumber="23" pageId="20" pageNumber="21">
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<subSubSection id="C37D6783923FE96FEBF94C82FDFCFF6D" box="[151,568,151,178]" pageId="20" pageNumber="21" type="nomenclature">
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<paragraph id="8BD83408923FE96FEBF94C82FDFCFF6D" blockId="20.[151,568,151,214]" box="[151,568,151,178]" pageId="20" pageNumber="21">
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<heading id="D0908364923FE96FEBF94C82FDFCFF6D" bold="true" box="[151,568,151,178]" fontSize="11" level="2" pageId="20" pageNumber="21" reason="4">
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<emphasis id="B913E81A923FE96FEBF94C82FDFCFF6D" bold="true" box="[151,568,151,178]" pageId="20" pageNumber="21">
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<taxonomicName id="4C674F8B923FE96FEBF94C82FE51FF6E" ID-CoL="9FYB2" authority="Mah, 2022" authorityName="Mah" authorityYear="2022" box="[151,405,151,178]" class="Asteroidea" family="Myxasteridae" genus="Heligmaster" kingdom="Animalia" order="Velatida" pageId="20" pageNumber="21" phylum="Echinodermata" rank="species" species="kanaloa" status="gen. nov.">
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<emphasis id="B913E81A923FE96FEBF94C82FE51FF6E" bold="true" box="[151,405,151,178]" italics="true" pageId="20" pageNumber="21">Heligmaster kanaloa</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2205561923FE96FEAF24C82FE2AFF6D" box="[412,494,151,177]" pageId="20" pageNumber="21" rank="genus">n. gen.</taxonomicNameLabel>
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n. sp.
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C37D6783923FE96FEBF94CAEFE66FF0A" box="[151,418,187,214]" pageId="20" pageNumber="21" type="description">
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<paragraph id="8BD83408923FE96FEBF94CAEFE66FF0A" blockId="20.[151,568,151,214]" box="[151,418,187,214]" pageId="20" pageNumber="21">
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<figureCitation id="135C288D923FE96FEBF94CAEFE97FF0A" box="[151,339,187,214]" captionStart="FIGURE 6" captionStartId="21.[151,250,1873,1897]" captionTargetBox="[164,1425,182,1846]" captionTargetId="figure-15@21.[163,1425,181,1849]" captionTargetPageId="21" captionText="FIGURE 6. Heligmaster kanaloa n. gen., n. sp. USNM 1457376 A. Abactinal disk surface. Scale bar=2.0 mm. B. Lateral view, whole body. Scale bar=5.0 mm. C. Cutaway of ambulacral and adambulacral ossicles. Disk is on right side. H=ambulacral head, N=ambulacral neck, B=ambulacral base, Ad= adambulacral plate. Scale bar=1.0 mm. D. Lateral view of abactinal surface, superomarginal, inferomarginal paxillae. Scale bar=2.0 mm. E. Actinal surface showing furrow spines. Scale bar= 5.0 mm. F. Close-up of furrow spination, tube foot furrow. Scale bar=1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6821042" httpUri="https://zenodo.org/record/6821042/files/figure.png" pageId="20" pageNumber="21">FIGURES 6A–F</figureCitation>
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,
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<figureCitation id="135C288D923FE96FEA304CAEFE66FF0A" box="[350,418,187,214]" captionStart="FIGURE 8" captionStartId="26.[151,250,1907,1931]" captionTargetBox="[152,1436,184,1882]" captionTargetId="figure-28@26.[151,1436,181,1883]" captionTargetPageId="26" captionText="FIGURE 8. In situ images. Heligmaster kanaloae n. gen. n. sp. A. Orange color morph. McCall Seamount, Hawaiian Islands. 2655 m. Imaged on vertical rock face. B. White color morph. South Midway Atoll region, 3917 m. Atheraster arandae C. Short armed morph. Salmon Bank, Hawaiian Islands. 2007 m. D. Long-armed morph. Salmon Bank, Hawaiian Islands, 1949 m. E. Rose Atoll, American Samoa, 2439 m. Atheraster sp. F. Bank Nine, Hawaiian Islands, 1635 m. G. Hawaiian Islands," figureDoi="http://doi.org/10.5281/zenodo.6821048" httpUri="https://zenodo.org/record/6821048/files/figure.png" pageId="20" pageNumber="21">8A–B</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C37D6783923FE96FEBF94D16FB1DFE9D" pageId="20" pageNumber="21" type="etymology">
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<paragraph id="8BD83408923FE96FEBF94D16FB1DFE9D" blockId="20.[151,1437,259,2042]" pageId="20" pageNumber="21">
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<emphasis id="B913E81A923FE96FEBF94D16FEE6FEC1" bold="true" box="[151,290,259,285]" pageId="20" pageNumber="21">Etymology:</emphasis>
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The species name honors
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<emphasis id="B913E81A923FE96FE9264D16FD6CFEC1" box="[584,680,259,285]" italics="true" pageId="20" pageNumber="21">Kanaloa</emphasis>
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, the Hawaiian god of the Ocean, whose association with squids and octopi is especially relevant to the cephalopod like arms on this species. Noun held in apposition.
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</paragraph>
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</subSubSection>
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<subSubSection id="C37D6783923FE96FEBF94D5EFDD1FD59" pageId="20" pageNumber="21" type="diagnosis">
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<paragraph id="8BD83408923FE96FEBF94D5EFC30FEBA" blockId="20.[151,1437,259,2042]" box="[151,1012,331,358]" pageId="20" pageNumber="21">
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<emphasis id="B913E81A923FE96FEBF94D5EFED6FEB9" bold="true" box="[151,274,331,357]" pageId="20" pageNumber="21">Diagnosis.</emphasis>
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Stellate (R/r=4.3 at R=7.0) body form with strongly curved arms.
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</paragraph>
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<paragraph id="8BD83408923FE96FEBA94D7AFA99FDE2" blockId="20.[151,1437,259,2042]" pageId="20" pageNumber="21">Disk with delicate, glassine, imbricate plates embedded in abactinal surface, each with one or two short spinelets. Marginal plates in single series with pointed, glassine spines, 3–6 bearing thornlets, spines on a short, rounded base. Subambulacral plates correspond 3 for every 1 marginal plate with a smooth, accessory-free dermis-covered region between them on lateral arm surface. Subambulacral plates with spines, 1–3 set on a round base. Spines similar to those on marginal plates, pointed and elongate, glassine with thornlets. Furrow spines, 3–5, webbed, dermis covered, arranged in straight to weakly curved fans, forming a near continuous series along tube foot groove.</paragraph>
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<paragraph id="8BD83408923FE96FEBA94E52FDD1FD59" blockId="20.[151,1437,259,2042]" pageId="20" pageNumber="21">Color in life: different individuals are white or orange. Orange individuals had dark orange tube feet. White individuals had dark tan tube feet.</paragraph>
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</subSubSection>
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<subSubSection id="C37D6783923FE96FEBA94E9AFA58FCE6" pageId="20" pageNumber="21" type="discussion">
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<paragraph id="8BD83408923FE96FEBA94E9AFACDFD11" blockId="20.[151,1437,259,2042]" pageId="20" pageNumber="21">
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<emphasis id="B913E81A923FE96FEBA94E9AFE8BFD75" bold="true" box="[199,335,655,681]" pageId="20" pageNumber="21">Comments.</emphasis>
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Several images of
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<taxonomicName id="4C674F8B923FE96FE94B4E9AFD6BFD75" authority="Mah, 2022" authorityName="Mah" authorityYear="2022" box="[549,687,655,681]" class="Asteroidea" family="Myxasteridae" genus="Heligmaster" kingdom="Animalia" order="Velatida" pageId="20" pageNumber="21" phylum="Echinodermata" rank="genus" status="gen. nov.">
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<emphasis id="B913E81A923FE96FE94B4E9AFD6BFD75" box="[549,687,655,681]" italics="true" pageId="20" pageNumber="21">Heligmaster</emphasis>
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</taxonomicName>
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<emphasis id="B913E81A923FE96FE9D84E9AFCC0FD75" bold="true" box="[694,772,655,681]" pageId="20" pageNumber="21">
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<taxonomicNameLabel id="A2205561923FE96FE9D84E9AFCC0FD75" box="[694,772,655,681]" pageId="20" pageNumber="21" rank="genus">n. gen.</taxonomicNameLabel>
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</emphasis>
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were made without a specimen collection. Identification of these images was based on the fewer spine counts of the superomarginal paxillae as
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<taxonomicName id="4C674F8B923FE96FEF294EA1FB07FD11" authority="Mah, 2022" authorityName="Mah" authorityYear="2022" box="[1095,1219,691,717]" class="Asteroidea" family="Myxasteridae" genus="Heligmaster" kingdom="Animalia" order="Velatida" pageId="20" pageNumber="21" phylum="Echinodermata" rank="species" species="kanaloa" status="sp. nov.">
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<emphasis id="B913E81A923FE96FEF294EA1FB07FD11" box="[1095,1219,691,717]" italics="true" pageId="20" pageNumber="21">H. kanaloa</emphasis>
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</taxonomicName>
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<emphasis id="B913E81A923FE96FEFA44EA6FACDFD11" bold="true" box="[1226,1289,691,717]" pageId="20" pageNumber="21">
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<taxonomicNameLabel id="A2205561923FE96FEFA44EA6FACDFD11" box="[1226,1289,691,717]" pageId="20" pageNumber="21" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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<paragraph id="8BD83408923FE96FEBA94EC2FBF0FCC9" blockId="20.[151,1437,259,2042]" pageId="20" pageNumber="21">Observations of differing color variations suggest two distinct color variations, one orange and one white. It was unclear, if color had any significance relative to size or if it was regional variation.</paragraph>
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<paragraph id="8BD83408923FE96FEBA94F0AFA58FCE6" blockId="20.[151,1437,259,2042]" box="[199,1436,799,826]" pageId="20" pageNumber="21">
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<emphasis id="B913E81A923FE96FEBA94F0AFE98FCE5" bold="true" box="[199,348,799,825]" pageId="20" pageNumber="21">Occurrence:</emphasis>
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Hawaiian Islands,
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<collectingCountry id="F3707498923FE96FE9434F0AFD14FCE5" box="[557,720,799,825]" name="United States Pacific Island Wildlife Refuges" pageId="20" pageNumber="21">Johnston Atoll</collectingCountry>
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, McCall Seamount, Midway and
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<collectingCountry id="F3707498923FE96FEF504F0AFB3AFCE5" box="[1086,1278,799,825]" name="American Samoa" pageId="20" pageNumber="21">American Samoa</collectingCountry>
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<quantity id="4C9F99ED923FE96FEE6D4F0AFA58FCE6" box="[1283,1436,799,826]" metricMagnitude="3" metricUnit="m" metricValue="3.0035" metricValueMax="3.916" metricValueMin="2.091" pageId="20" pageNumber="21" unit="m" value="3003.5" valueMax="3916.0" valueMin="2091.0">2091–3916 m</quantity>
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</paragraph>
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</subSubSection>
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<subSubSection id="C37D6783923FE96DEBA94F56FD0EFD2D" lastPageId="22" lastPageNumber="23" pageId="20" pageNumber="21" type="description">
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<paragraph id="8BD83408923FE96FEBA94F56FEFFFC5D" blockId="20.[151,1437,259,2042]" pageId="20" pageNumber="21">
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<emphasis id="B913E81A923FE96FEBA94F56FE9DFC81" bold="true" box="[199,345,835,861]" pageId="20" pageNumber="21">Description.</emphasis>
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Body strongly stellate (R/r=~4.3) with elongate, with tapering, curved arms, extending below the disk (
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<figureCitation id="135C288D923FE96FEBBB4F72FEEAFC5D" box="[213,302,871,897]" captionStart="FIGURE 6" captionStartId="21.[151,250,1873,1897]" captionTargetBox="[164,1425,182,1846]" captionTargetId="figure-15@21.[163,1425,181,1849]" captionTargetPageId="21" captionText="FIGURE 6. Heligmaster kanaloa n. gen., n. sp. USNM 1457376 A. Abactinal disk surface. Scale bar=2.0 mm. B. Lateral view, whole body. Scale bar=5.0 mm. C. Cutaway of ambulacral and adambulacral ossicles. Disk is on right side. H=ambulacral head, N=ambulacral neck, B=ambulacral base, Ad= adambulacral plate. Scale bar=1.0 mm. D. Lateral view of abactinal surface, superomarginal, inferomarginal paxillae. Scale bar=2.0 mm. E. Actinal surface showing furrow spines. Scale bar= 5.0 mm. F. Close-up of furrow spination, tube foot furrow. Scale bar=1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6821042" httpUri="https://zenodo.org/record/6821042/files/figure.png" pageId="20" pageNumber="21">Fig. 6B</figureCitation>
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).
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</paragraph>
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<paragraph id="8BD83408923FE96FEBA94F9EFD80FB1F" blockId="20.[151,1437,259,2042]" pageId="20" pageNumber="21">
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Abactinal surface composed of thin, delicate translucent dermis, plates weakly developed minute. Underlying features, such as ambulacrals and sediment infused stomach visible through the abactinal surface. Surface with short glassine spinelets, mostly with single pointed tip but a minority of spines with bifid tips and covered by weakly developed dermis (
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<figureCitation id="135C288D923FE96FEA054FE3FE00FBCC" box="[363,452,1014,1040]" captionStart="FIGURE 6" captionStartId="21.[151,250,1873,1897]" captionTargetBox="[164,1425,182,1846]" captionTargetId="figure-15@21.[163,1425,181,1849]" captionTargetPageId="21" captionText="FIGURE 6. Heligmaster kanaloa n. gen., n. sp. USNM 1457376 A. Abactinal disk surface. Scale bar=2.0 mm. B. Lateral view, whole body. Scale bar=5.0 mm. C. Cutaway of ambulacral and adambulacral ossicles. Disk is on right side. H=ambulacral head, N=ambulacral neck, B=ambulacral base, Ad= adambulacral plate. Scale bar=1.0 mm. D. Lateral view of abactinal surface, superomarginal, inferomarginal paxillae. Scale bar=2.0 mm. E. Actinal surface showing furrow spines. Scale bar= 5.0 mm. F. Close-up of furrow spination, tube foot furrow. Scale bar=1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6821042" httpUri="https://zenodo.org/record/6821042/files/figure.png" pageId="20" pageNumber="21">Fig. 6A</figureCitation>
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). Spinelets evenly distributed over surface, approximately 2 to 3 counted along a 1.0 mm line on disk and arms becoming more widely distributed and more weakly developed distally along arms but absent along a narrow strip interradially on disk. Papulae single, present most abundantly on proximal arm region. Absent from central disk and distal arm. Anus located centrally on disk, formed by relatively large, opening, approximately
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<quantity id="4C9F99ED923FE96FEBF94890FF2AFB43" box="[151,238,1157,1184]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="20" pageNumber="21" unit="mm" value="1.5">1.5 mm</quantity>
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in length. Madreporite large, raised above surface of body, approximately
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<quantity id="4C9F99ED923FE96FEF594890FBBCFB43" box="[1079,1144,1157,1183]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="20" pageNumber="21" unit="mm" value="3.0">3 mm</quantity>
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in width with well-developed sulci. No pedicellariae observed.
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</paragraph>
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<paragraph id="8BD83408923FE96FEBA948D9FB09F9D8" blockId="20.[151,1437,259,2042]" pageId="20" pageNumber="21">Single row of widely spaced marginal plates forming single series along upper edge of lateral side. Lateral surface and plates covered by thick dermis. Lower edge composed of adambulacral plates along lower edge of lateral surface. Approximately one marginal plate for every three adambulacral plates. Marginal plates, 44 per interradius (armtip to armtip), each with 3 to 6 sharp spines, sitting on a mound-like, pedunculate base, 1 spine demonstrably longer than the others on each plate. Spines glassine with surface bearing tiny thornlets. Spacing between marginal plates widest proximally, shortest adjacent to armtip. Lateral region between marginal plate boundary to adambulacral series, covered by distinct dermis, smooth mostly devoid of spines or other features save for one or two small spinelets proximally. Terminal plate toe-shaped, tipped with four pointed spines. Note that subambulacrals positioned adjacent to marginals on lateral surface in approximately “inferomarginal plate” position.</paragraph>
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<paragraph id="8BD83408923FE96FEBA94A1BFDCCF851" blockId="20.[151,1437,259,2042]" pageId="20" pageNumber="21">Subambulacral spines correspond roughly three plates to every one marginal plate. Large subambulacral spines emerge from adambulacral plate number 92 per interradius (armtip to armtip) corresponding roughly every 3 to every 1 marginal plate. Each with a round pedunculate base similar to marginal plates, with 1 to 3 pointed spines per plate. Furrow spines 3 to 5, forming straight to weakly curved fans, each spine short, webbed, sheathed in tissue forming a nearly continuous series adjacent to the tube foot furrow. Furrow spine arrays sit a distinct distance away from subambulacral plates. Intervening distance covered by thick dermis. Proximal-most furrow spine short, distinct, with spatulate, tissue covered tip and at an oblique angle to the other furrow spines. Adambulacral plates broadly mound-shaped, convex surface facing into furrow, each plate distinctly separated from one another by tissue. Surface covered by thick dermis. Oral plate with webbed furrow spines, 7 per plate, totaling 14 per interradius. Oral plate surface with 3 to 4 pointed spines separate from furrow spines. Fissure between oral plates well developed. No pedicellariae observed.</paragraph>
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<paragraph id="8BD83408923FE96FEBA94B82FB78F825" blockId="20.[151,1437,259,2042]" pageId="20" pageNumber="21">
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Tube feet muscular with ring-shaped tip. Podial pores remain biserial throughout, but drawing together very closely on distalmost armtip (approximately 10 plates from terminal) giving the appearance of switching from biserial to monoserial near terminus. Pyloric stomach well developed, partially extended (
|
||
<figureCitation id="135C288D923FE96FEF364BCAFB6BF825" box="[1112,1199,2015,2041]" captionStart="FIGURE 6" captionStartId="21.[151,250,1873,1897]" captionTargetBox="[164,1425,182,1846]" captionTargetId="figure-15@21.[163,1425,181,1849]" captionTargetPageId="21" captionText="FIGURE 6. Heligmaster kanaloa n. gen., n. sp. USNM 1457376 A. Abactinal disk surface. Scale bar=2.0 mm. B. Lateral view, whole body. Scale bar=5.0 mm. C. Cutaway of ambulacral and adambulacral ossicles. Disk is on right side. H=ambulacral head, N=ambulacral neck, B=ambulacral base, Ad= adambulacral plate. Scale bar=1.0 mm. D. Lateral view of abactinal surface, superomarginal, inferomarginal paxillae. Scale bar=2.0 mm. E. Actinal surface showing furrow spines. Scale bar= 5.0 mm. F. Close-up of furrow spination, tube foot furrow. Scale bar=1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6821042" httpUri="https://zenodo.org/record/6821042/files/figure.png" pageId="20" pageNumber="21">Fig. 6E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF186480923EE96EEBF94B44FCFFF825" ID-DOI="http://doi.org/10.5281/zenodo.6821042" ID-Zenodo-Dep="6821042" httpUri="https://zenodo.org/record/6821042/files/figure.png" pageId="21" pageNumber="22" startId="21.[151,250,1873,1897]" targetBox="[164,1425,182,1846]" targetPageId="21">
|
||
<paragraph id="8BD83408923EE96EEBF94B44FCFFF825" blockId="21.[151,1437,1873,2041]" pageId="21" pageNumber="22">
|
||
<emphasis id="B913E81A923EE96EEBF94B44FE3EF8B5" bold="true" box="[151,506,1873,1897]" pageId="21" pageNumber="22">
|
||
FIGURE 6.
|
||
<taxonomicName id="4C674F8B923EE96EEA704B44FE3EF8B5" authority="Mah, 2022" authorityName="Mah" authorityYear="2022" box="[286,506,1873,1897]" class="Asteroidea" family="Myxasteridae" genus="Heligmaster" kingdom="Animalia" order="Velatida" pageId="21" pageNumber="22" phylum="Echinodermata" rank="species" species="kanaloa" status="n. gen., n. sp.">
|
||
<emphasis id="B913E81A923EE96EEA704B44FE3EF8B5" bold="true" box="[286,506,1873,1897]" italics="true" pageId="21" pageNumber="22">Heligmaster kanaloa</emphasis>
|
||
</taxonomicName>
|
||
</emphasis>
|
||
<taxonomicNameLabel id="A2205561923EE96EE96D4B44FD53F8B5" box="[515,663,1873,1897]" pageId="21" pageNumber="22" rank="species">
|
||
<emphasis id="B913E81A923EE96EE96D4B44FD55F8B5" bold="true" box="[515,657,1873,1897]" pageId="21" pageNumber="22">n. gen., n. sp</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
USNM 1457376
|
||
<emphasis id="B913E81A923EE96EE83C4B44FCAFF8B5" bold="true" box="[850,875,1873,1897]" pageId="21" pageNumber="22">A.</emphasis>
|
||
Abactinal disk surface. Scale bar=2.0 mm.
|
||
<emphasis id="B913E81A923EE96EEE5B4B44FA88F8B5" bold="true" box="[1333,1356,1873,1897]" pageId="21" pageNumber="22">B.</emphasis>
|
||
Lateral view, whole body. Scale bar=5.0 mm.
|
||
<emphasis id="B913E81A923EE96EE9784B60FDEBF851" bold="true" box="[534,559,1909,1933]" pageId="21" pageNumber="22">C.</emphasis>
|
||
Cutaway of ambulacral and adambulacral ossicles. Disk is on right side. H=ambulacral head, N=ambulacral neck, B=ambulacral base, Ad= adambulacral plate. Scale bar=1.0 mm.
|
||
<emphasis id="B913E81A923EE96EEF5E4B8CFB8DF86D" bold="true" box="[1072,1097,1945,1969]" pageId="21" pageNumber="22">D.</emphasis>
|
||
Lateral view of abactinal surface, superomarginal, inferomarginal paxillae. Scale bar=2.0 mm.
|
||
<emphasis id="B913E81A923EE96EE86A4BA8FCD8F809" bold="true" box="[772,796,1981,2005]" pageId="21" pageNumber="22">E.</emphasis>
|
||
Actinal surface showing furrow spines. Scale bar= 5.0 mm.
|
||
<emphasis id="B913E81A923EE96EEEE74BA8FA58F809" bold="true" box="[1417,1436,1981,2005]" pageId="21" pageNumber="22">F.</emphasis>
|
||
Close-up of furrow spination, tube foot furrow. Scale bar=1.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD83408923DE96DEBA94C82FA4FFE9D" blockId="22.[151,1436,151,321]" pageId="22" pageNumber="23">
|
||
<materialsCitation id="3B0F3E55923DE96DEBA94C82FA4FFE9D" collectionCode="NOAA, R" collectorName="Coll. & Chris Mah & C. Kelley." country="United States Pacific Island Wildlife Refuges" county="North Pacific Ocean." elevation="2568" location="Ambulacral" municipality="Color" pageId="22" pageNumber="23" specimenCode="USNM 1457376, EX 1706" specimenCount="1" typeStatus="holotype">
|
||
<location id="8EB862D3923DE96DEBA94C82FE8FFF6D" LSID="urn:lsid:plazi:treatment:03CE851E923FE96DEBF94C82FD0EFD2D:8EB862D3923DE96DEBA94C82FE8FFF6D" box="[199,331,151,177]" country="United States Pacific Island Wildlife Refuges" county="North Pacific Ocean." municipality="Color" name="Ambulacral" pageId="22" pageNumber="23">Ambulacral</location>
|
||
ossicles viewed laterally with head projecting proximally broadly angular in outline, neck with flat surface, base quadrate n shape (
|
||
<figureCitation id="135C288D923DE96DEA944CAEFD97FF09" box="[506,595,187,213]" captionStart="FIGURE 6" captionStartId="21.[151,250,1873,1897]" captionTargetBox="[164,1425,182,1846]" captionTargetId="figure-15@21.[163,1425,181,1849]" captionTargetPageId="21" captionText="FIGURE 6. Heligmaster kanaloa n. gen., n. sp. USNM 1457376 A. Abactinal disk surface. Scale bar=2.0 mm. B. Lateral view, whole body. Scale bar=5.0 mm. C. Cutaway of ambulacral and adambulacral ossicles. Disk is on right side. H=ambulacral head, N=ambulacral neck, B=ambulacral base, Ad= adambulacral plate. Scale bar=1.0 mm. D. Lateral view of abactinal surface, superomarginal, inferomarginal paxillae. Scale bar=2.0 mm. E. Actinal surface showing furrow spines. Scale bar= 5.0 mm. F. Close-up of furrow spination, tube foot furrow. Scale bar=1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6821042" httpUri="https://zenodo.org/record/6821042/files/figure.png" pageId="22" pageNumber="23">Fig. 6C</figureCitation>
|
||
).
|
||
<collectingMunicipality id="6BBCAE72923DE96DEBA94CCAFEC3FF25" box="[199,263,223,249]" pageId="22" pageNumber="23">Color</collectingMunicipality>
|
||
of living specimens was orange and white (
|
||
<figureCitation id="135C288D923DE96DE99D4CCAFCA9FF25" box="[755,877,223,249]" captionStart="FIGURE 8" captionStartId="26.[151,250,1907,1931]" captionTargetBox="[152,1436,184,1882]" captionTargetId="figure-28@26.[151,1436,181,1883]" captionTargetPageId="26" captionText="FIGURE 8. In situ images. Heligmaster kanaloae n. gen. n. sp. A. Orange color morph. McCall Seamount, Hawaiian Islands. 2655 m. Imaged on vertical rock face. B. White color morph. South Midway Atoll region, 3917 m. Atheraster arandae C. Short armed morph. Salmon Bank, Hawaiian Islands. 2007 m. D. Long-armed morph. Salmon Bank, Hawaiian Islands, 1949 m. E. Rose Atoll, American Samoa, 2439 m. Atheraster sp. F. Bank Nine, Hawaiian Islands, 1635 m. G. Hawaiian Islands," figureDoi="http://doi.org/10.5281/zenodo.6821048" httpUri="https://zenodo.org/record/6821048/files/figure.png" pageId="22" pageNumber="23">Fig. 8A, B</figureCitation>
|
||
).
|
||
<emphasis id="B913E81A923DE96DEBA94D17FDF0FEC1" bold="true" box="[199,564,258,285]" pageId="22" pageNumber="23">
|
||
Material Examined.
|
||
<typeStatus id="54DC8AAA923DE96DEAAE4D16FDEBFEC1" box="[448,559,259,285]" pageId="22" pageNumber="23" type="holotype">Holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
<specimenCode id="DBC19C73923DE96DE9524D16FD3FFEC1" box="[572,763,258,285]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="22" pageNumber="23" type="Museum">USNM 1457376</specimenCode>
|
||
,
|
||
<collectingCountry id="F3707498923DE96DE8694D16FC69FEC1" box="[775,941,259,285]" name="United States Pacific Island Wildlife Refuges" pageId="22" pageNumber="23">Johnston Atoll</collectingCountry>
|
||
,
|
||
<collectingCounty id="62B94C84923DE96DE8D44D16FB6DFEC1" box="[954,1193,259,285]" pageId="22" pageNumber="23">North Pacific Ocean.</collectingCounty>
|
||
15.17, -171.25,
|
||
<quantity id="4C9F99ED923DE96DEE0A4D17FF6BFE9D" metricMagnitude="3" metricUnit="m" metricValue="2.568" pageId="22" pageNumber="23" unit="m" value="2568.0">
|
||
<elevation id="004AD33B923DE96DEE0A4D17FF6BFE9D" metricMagnitude="3" metricUnit="m" metricValue="2.568" pageId="22" pageNumber="23" unit="m" value="2568.0">2568 m</elevation>
|
||
</quantity>
|
||
,
|
||
<collectorName id="269251DE923DE96DEBD54D32FF37FE9D" box="[187,243,295,321]" pageId="22" pageNumber="23">Coll.</collectorName>
|
||
<collectionCode id="ED76ACCD923DE96DEB974D32FE8CFE9D" box="[249,328,295,321]" country="USA" httpUri="http://grbio.org/cool/qzhp-igg5" name="National Oceanic and Atmospeheric Administration" pageId="22" pageNumber="23">NOAA</collectionCode>
|
||
ship
|
||
<taxonomicName id="4C674F8B923DE96DEAEB4D32FD7AFE9D" authority="Explorer (NOAA)" box="[389,702,295,321]" class="Insecta" family="Pentatomidae" genus="Okeanos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B913E81A923DE96DEAEB4D32FD95FE9D" box="[389,593,295,321]" italics="true" pageId="22" pageNumber="23">Okeanos Explorer</emphasis>
|
||
(NOAA)
|
||
</taxonomicName>
|
||
,
|
||
<collectorName id="269251DE923DE96DE9A74D32FC84FE9D" box="[713,832,295,321]" pageId="22" pageNumber="23">Chris Mah</collectorName>
|
||
and
|
||
<collectorName id="269251DE923DE96DE8194D32FC2CFE9D" box="[887,1000,295,321]" pageId="22" pageNumber="23">C. Kelley.</collectorName>
|
||
<specimenCode id="DBC19C73923DE96DE8804D32FB91FE9D" box="[1006,1109,295,321]" collectionCode="EX" country="Slovenia" name="The Culture Collection of Extremophilic Fungi" pageId="22" pageNumber="23">EX 1706</specimenCode>
|
||
. 1 wet spec.
|
||
<collectionCode id="ED76ACCD923DE96DEF8B4D32FB3DFE9D" box="[1253,1273,295,321]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="22" pageNumber="23">R</collectionCode>
|
||
=~7.0, r=1.6.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BD83408923DE96DEBF94D7AFCBCFE0D" blockId="22.[151,1436,366,753]" pageId="22" pageNumber="23">
|
||
<emphasis id="B913E81A923DE96DEBF94D7AFEA9FE54" bold="true" box="[151,365,366,393]" pageId="22" pageNumber="23">Images Examined</emphasis>
|
||
<materialsCitation id="3B0F3E55923DE96DEBF94D86FCBCFE0D" collectionCode="ROVHD, ASR, AUD" elevation="2655" latitude="18.979053" location="McCall Seamount" longLatPrecision="1" longitude="-157.11024" municipality="Hawaiian Islands" pageId="22" pageNumber="23" specimenCode="EX1504, T231915" specimenCount="1">
|
||
orange forms.
|
||
<location id="8EB862D3923DE96DEA544D86FDCCFE71" LSID="urn:lsid:plazi:treatment:03CE851E923FE96DEBF94C82FD0EFD2D:8EB862D3923DE96DEA544D86FDCCFE71" box="[314,520,403,429]" latitude="18.979053" longLatPrecision="1" longitude="-157.11024" municipality="Hawaiian Islands" name="McCall Seamount" pageId="22" pageNumber="23">McCall Seamount</location>
|
||
,
|
||
<collectingMunicipality id="6BBCAE72923DE96DE97D4D86FD10FE71" box="[531,724,403,429]" pageId="22" pageNumber="23">Hawaiian Islands</collectingMunicipality>
|
||
<geoCoordinate id="EE5352CF923DE96DE9B54D87FC91FE71" box="[731,853,402,429]" degrees="18.979054" direction="north" orientation="latitude" pageId="22" pageNumber="23" precision="1" value="18.979053">18.979054</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5352CF923DE96DE80E4D87FC3BFE71" box="[864,1023,402,429]" degrees="157.1102457" direction="west" orientation="longitude" pageId="22" pageNumber="23" precision="1" value="-157.11024">-157.1102457</geoCoordinate>
|
||
,
|
||
<quantity id="4C9F99ED923DE96DEF644D87FBA6FE71" box="[1034,1122,402,429]" metricMagnitude="3" metricUnit="m" metricValue="2.655" pageId="22" pageNumber="23" unit="m" value="2655.0">
|
||
<elevation id="004AD33B923DE96DEF644D87FBA6FE71" box="[1034,1122,402,429]" metricMagnitude="3" metricUnit="m" metricValue="2.655" pageId="22" pageNumber="23" unit="m" value="2655.0">2655 m</elevation>
|
||
</quantity>
|
||
.
|
||
<specimenCode id="DBC19C73923DE96DEBF94DA3FF37FE0D" box="[151,243,438,465]" collectionCode="EX" country="Slovenia" name="The Culture Collection of Extremophilic Fungi" pageId="22" pageNumber="23">EX1504</specimenCode>
|
||
L3_IMG_20150831
|
||
<specimenCode id="DBC19C73923DE96DEABA4DA3FDFCFE0D" box="[468,568,438,465]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="22" pageNumber="23">T231915</specimenCode>
|
||
Z_
|
||
<collectionCode id="ED76ACCD923DE96DE9364DA3FD79FE0D" box="[600,701,438,465]" pageId="22" pageNumber="23">ROVHD</collectionCode>
|
||
_
|
||
<collectionCode id="ED76ACCD923DE96DE9A24DA3FCC7FE0D" box="[716,771,438,465]" pageId="22" pageNumber="23">ASR</collectionCode>
|
||
_
|
||
<collectionCode id="ED76ACCD923DE96DE87F4DA3FC8AFE0D" box="[785,846,438,465]" pageId="22" pageNumber="23">AUD</collectionCode>
|
||
.jpg
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BD83408923DE96DEBF94DCEFD0EFDC5" blockId="22.[151,1436,366,753]" pageId="22" pageNumber="23">
|
||
<materialsCitation id="3B0F3E55923DE96DEBF94DCEFD0EFDC5" collectionCode="ROVHD" country="United States Pacific Island Wildlife Refuges" elevation="2569" latitude="15.1662245" location="Wetmore Seamount" longLatPrecision="1" longitude="-171.25461" pageId="22" pageNumber="23" specimenCode="EX1706, T202554" specimenCount="1">
|
||
<location id="8EB862D3923DE96DEBF94DCEFEB3FE29" LSID="urn:lsid:plazi:treatment:03CE851E923FE96DEBF94C82FD0EFD2D:8EB862D3923DE96DEBF94DCEFEB3FE29" box="[151,375,475,501]" country="United States Pacific Island Wildlife Refuges" latitude="15.1662245" longLatPrecision="1" longitude="-171.25461" name="Wetmore Seamount" pageId="22" pageNumber="23">Wetmore Seamount</location>
|
||
,
|
||
<collectingCountry id="F3707498923DE96DEAEC4DCEFDE1FE29" box="[386,549,475,501]" name="United States Pacific Island Wildlife Refuges" pageId="22" pageNumber="23">Johnston Atoll</collectingCountry>
|
||
region,
|
||
<geoCoordinate id="EE5352CF923DE96DE9EF4DCFFCD3FE29" box="[641,791,474,501]" degrees="15.16622471" direction="north" orientation="latitude" pageId="22" pageNumber="23" precision="1" value="15.1662245">15.16622471</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5352CF923DE96DE84C4DCFFC70FE29" box="[802,948,474,501]" degrees="171.254609" direction="west" orientation="longitude" pageId="22" pageNumber="23" precision="1" value="-171.25461">-171.254609</geoCoordinate>
|
||
,
|
||
<quantity id="4C9F99ED923DE96DE8D14DCFFBD0FE29" box="[959,1044,474,501]" metricMagnitude="3" metricUnit="m" metricValue="2.569" pageId="22" pageNumber="23" unit="m" value="2569.0">
|
||
<elevation id="004AD33B923DE96DE8D14DCFFBD0FE29" box="[959,1044,474,501]" metricMagnitude="3" metricUnit="m" metricValue="2.569" pageId="22" pageNumber="23" unit="m" value="2569.0">2569 m</elevation>
|
||
</quantity>
|
||
<specimenCode id="DBC19C73923DE96DEBF94DEBFF37FDC5" box="[151,243,510,537]" collectionCode="EX" country="Slovenia" name="The Culture Collection of Extremophilic Fungi" pageId="22" pageNumber="23">EX1706</specimenCode>
|
||
_IMG_20170723
|
||
<specimenCode id="DBC19C73923DE96DEAD94DEBFDD9FDC5" box="[439,541,510,537]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="22" pageNumber="23">T202554</specimenCode>
|
||
Z_
|
||
<collectionCode id="ED76ACCD923DE96DE9524DEBFD64FDC5" box="[572,672,510,537]" pageId="22" pageNumber="23">ROVHD</collectionCode>
|
||
.jpg
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BD83408923DE96DEBF94E36FE61FDBD" blockId="22.[151,1436,366,753]" pageId="22" pageNumber="23">
|
||
<materialsCitation id="3B0F3E55923DE96DEBF94E36FE61FDBD" country="United States Pacific Island Wildlife Refuges" elevation="3916" latitude="25.359888" location="South" longLatPrecision="1" longitude="-178.42923" pageId="22" pageNumber="23" specimenCount="1">
|
||
white forms.
|
||
<location id="8EB862D3923DE96DEA404E36FEABFDE1" LSID="urn:lsid:plazi:treatment:03CE851E923FE96DEBF94C82FD0EFD2D:8EB862D3923DE96DEA404E36FEABFDE1" box="[302,367,547,573]" country="United States Pacific Island Wildlife Refuges" latitude="25.359888" longLatPrecision="1" longitude="-178.42923" name="South" pageId="22" pageNumber="23">South</location>
|
||
<collectingCountry id="F3707498923DE96DEA194E36FDD0FDE1" box="[375,532,547,573]" name="United States Pacific Island Wildlife Refuges" pageId="22" pageNumber="23">Midway Atoll</collectingCountry>
|
||
<geoCoordinate id="EE5352CF923DE96DE9724E37FD76FDE1" box="[540,690,546,573]" degrees="25.35988374" direction="north" orientation="latitude" pageId="22" pageNumber="23" precision="1" value="25.359884">25.35988374</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5352CF923DE96DE9D04E36FC99FDE1" box="[702,861,547,573]" degrees="178.4292349" direction="west" orientation="longitude" pageId="22" pageNumber="23" precision="1" value="-178.42923">-178.4292349</geoCoordinate>
|
||
,
|
||
<quantity id="4C9F99ED923DE96DE8074E36FC7BFDE1" box="[873,959,547,573]" metricMagnitude="3" metricUnit="m" metricValue="3.916" pageId="22" pageNumber="23" unit="m" value="3916.0">
|
||
<elevation id="004AD33B923DE96DE8074E36FC7BFDE1" box="[873,959,547,573]" metricMagnitude="3" metricUnit="m" metricValue="3.916" pageId="22" pageNumber="23" unit="m" value="3916.0">3916 m</elevation>
|
||
</quantity>
|
||
to
|
||
<geoCoordinate id="EE5352CF923DE96DE88A4E37FBBEFDE1" box="[996,1146,546,573]" degrees="25.35988775" direction="north" orientation="latitude" pageId="22" pageNumber="23" precision="1" value="25.359888">25.35988775</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5352CF923DE96DEFE84E37FAE2FDE1" box="[1158,1318,546,573]" degrees="178.4292251" direction="west" orientation="longitude" pageId="22" pageNumber="23" precision="1" value="-178.42923">-178.4292251</geoCoordinate>
|
||
,
|
||
<quantity id="4C9F99ED923DE96DEE5F4E36FA58FDE1" box="[1329,1436,547,573]" metricMagnitude="3" metricUnit="m" metricValue="3.9166" pageId="22" pageNumber="23" unit="m" value="3916.6">
|
||
<elevation id="004AD33B923DE96DEE5F4E36FA58FDE1" box="[1329,1436,547,573]" metricMagnitude="3" metricUnit="m" metricValue="3.9166" pageId="22" pageNumber="23" unit="m" value="3916.6">3916.6 m</elevation>
|
||
</quantity>
|
||
(for both images below)
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BD83408923DE96DEBF94E7FFC1DFD75" blockId="22.[151,1436,366,753]" pageId="22" pageNumber="23">EX1603_IMG_20160311T233015Z_ROVHD_ASR_COR_BAR_CRI.jpg EX1603_IMG_20160311T233044Z_ROVHD_ASR_COR_BAR_CRI.jpg</paragraph>
|
||
<paragraph id="8BD83408923DE96DEBF94EA6FD0EFD2D" blockId="22.[151,1436,366,753]" pageId="22" pageNumber="23">
|
||
<materialsCitation id="3B0F3E55923DE96DEBF94EA6FD0EFD2D" collectionCode="ROVHD" country="American Samoa" elevation="2091" latitude="-11.168486" location="Moki Seamount" longLatPrecision="1" longitude="-169.89217" pageId="22" pageNumber="23" specimenCode="EX1702, T222707" specimenCount="1">
|
||
<location id="8EB862D3923DE96DEBF94EA6FE8AFD11" LSID="urn:lsid:plazi:treatment:03CE851E923FE96DEBF94C82FD0EFD2D:8EB862D3923DE96DEBF94EA6FE8AFD11" box="[151,334,691,717]" country="American Samoa" latitude="-11.168486" longLatPrecision="1" longitude="-169.89217" name="Moki Seamount" pageId="22" pageNumber="23">Moki Seamount</location>
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||
,
|
||
<collectingCountry id="F3707498923DE96DEA394EA6FDDDFD11" box="[343,537,691,717]" name="American Samoa" pageId="22" pageNumber="23">American Samoa</collectingCountry>
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region
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<geoCoordinate id="EE5352CF923DE96DE9004EA7FCC9FD11" box="[622,781,690,717]" degrees="11.16848593" direction="south" orientation="latitude" pageId="22" pageNumber="23" precision="1" value="-11.168486">-11.16848593</geoCoordinate>
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,
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<geoCoordinate id="EE5352CF923DE96DE8764EA6FC73FD11" box="[792,951,691,717]" degrees="169.8921686" direction="west" orientation="longitude" pageId="22" pageNumber="23" precision="1" value="-169.89217">-169.8921686</geoCoordinate>
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,
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<quantity id="4C9F99ED923DE96DE8AC4EA6FBD3FD11" box="[962,1047,691,717]" metricMagnitude="3" metricUnit="m" metricValue="2.091" pageId="22" pageNumber="23" unit="m" value="2091.0">
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<elevation id="004AD33B923DE96DE8AC4EA6FBD3FD11" box="[962,1047,691,717]" metricMagnitude="3" metricUnit="m" metricValue="2.091" pageId="22" pageNumber="23" unit="m" value="2091.0">2091 m</elevation>
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</quantity>
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||
<specimenCode id="DBC19C73923DE96DEBF94EC2FF37FD2D" box="[151,243,727,753]" collectionCode="EX" country="Slovenia" name="The Culture Collection of Extremophilic Fungi" pageId="22" pageNumber="23">EX1702</specimenCode>
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||
_IMG_20170222
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<specimenCode id="DBC19C73923DE96DEAD64EC2FDD9FD2D" box="[440,541,727,753]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="22" pageNumber="23">T222707</specimenCode>
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||
Z_
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||
<collectionCode id="ED76ACCD923DE96DE9524EC2FD64FD2D" box="[572,672,727,753]" pageId="22" pageNumber="23">ROVHD</collectionCode>
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||
.jpg
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |