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<document id="A2497E93A88844B4790E1730BC72BDD8" ID-DOI="10.11646/zootaxa.5164.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="6821026" ID-ZooBank="3BECB9C7-F4B5-4FA4-934B-1822BF3D1077" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1657611643055" checkinUser="plazi" docAuthor="Mah, Christopher L." docDate="2022" docId="03CE851E921FE94CEBF94A22FD0EFA89" docLanguage="en" docName="zootaxa.5164.1.1.pdf" docOrigin="Zootaxa 5164 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5164.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Litonotaster gfoei Mah 2022, n. sp." docType="treatment" docVersion="3" lastPageNumber="56" masterDocId="FFF7FD66922BE97BEB6E4C15FFC4FFDC" masterDocTitle="New Genera, Species and Occurrences of Deep-Sea Asteroidea (Valvatacea, Forcipulatacea, Echinodermata) collected from the North Pacific Ocean by the CAPSTONE Expedition" masterLastPageNumber="75" masterPageNumber="1" pageNumber="53" updateTime="1699345508031" updateUser="plazi">
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<mods:title id="A114DAB38392A1CEF6ACEC5330940027">New Genera, Species and Occurrences of Deep-Sea Asteroidea (Valvatacea, Forcipulatacea, Echinodermata) collected from the North Pacific Ocean by the CAPSTONE Expedition</mods:title>
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<subSubSection id="C37D6783921FE94FEBF94A22FE73F98D" box="[151,439,1590,1617]" pageId="52" pageNumber="53" type="nomenclature">
<paragraph id="8BD83408921FE94FEBF94A22FE73F98D" blockId="52.[151,439,1590,1653]" box="[151,439,1590,1617]" pageId="52" pageNumber="53">
<heading id="D0908364921FE94FEBF94A22FE73F98D" bold="true" box="[151,439,1590,1617]" fontSize="11" level="2" pageId="52" pageNumber="53" reason="4">
<emphasis id="B913E81A921FE94FEBF94A22FE73F98D" bold="true" box="[151,439,1590,1617]" pageId="52" pageNumber="53">
<taxonomicName id="4C674F8B921FE94FEBF94A22FEA9F98C" ID-CoL="9FYBD" authority="Mah, 2022" authorityName="Mah" authorityYear="2022" box="[151,365,1590,1617]" class="Asteroidea" family="Goniasteridae" genus="Litonotaster" kingdom="Animalia" order="Valvatida" pageId="52" pageNumber="53" phylum="Echinodermata" rank="species" species="gfoei" status="sp. nov.">
<emphasis id="B913E81A921FE94FEBF94A22FEA9F98C" bold="true" box="[151,365,1590,1617]" italics="true" pageId="52" pageNumber="53">Litonotaster gfoei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2205561921FE94FEA1B4A22FE73F98D" box="[373,439,1591,1617]" pageId="52" pageNumber="53" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C37D6783921FE94FEBF94A4EFE95F9A9" box="[151,337,1627,1653]" pageId="52" pageNumber="53" type="description">
<paragraph id="8BD83408921FE94FEBF94A4EFE95F9A9" blockId="52.[151,439,1590,1653]" box="[151,337,1627,1653]" pageId="52" pageNumber="53">
<figureCitation id="135C288D921FE94FEBF94A4EFE95F9A9" box="[151,337,1627,1653]" captionStart="FIGURE 18" captionStartId="54.[151,250,1833,1857]" captionTargetBox="[205,1382,184,1809]" captionTargetId="figure-28@54.[204,1384,181,1809]" captionTargetPageId="54" captionText="FIGURE 18. Litonotaster gfoei n. sp. A. Abactinal view. Scale bar=2.0 mm. Inset showing surface detail. Largest individual plate diameter=1.0 mm. B. In situ image. Arrow showing inferomarginal spines extended, American Samoa, 2457 m. Scale bar=2.0 mm. C. Actinal view. Scale bar=2.0 mm. D. Actinal-Lateral view. Arrow showing spinelets on inferomarginal edge. Scale bar=1.0 mm. E. Actinal surface showing details. Scale bar=1.0 mm. F. Adambulacral spination, including furrow and subambulacral spines. Scale bar=0.75 mm. G. In situ image showing specimen with papular pores, Mozart Seamount, North Pacific." figureDoi="http://doi.org/10.5281/zenodo.6821080" httpUri="https://zenodo.org/record/6821080/files/figure.png" pageId="52" pageNumber="53">FIGURE 18AF</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C37D6783921FE94FEBF94AB6FD3CF93D" pageId="52" pageNumber="53" type="etymology">
<paragraph id="8BD83408921FE94FEBF94AB6FD3CF93D" blockId="52.[151,1437,1699,2013]" pageId="52" pageNumber="53">
<emphasis id="B913E81A921FE94FEBF94AB6FEE6F961" bold="true" box="[151,290,1699,1725]" pageId="52" pageNumber="53">Etymology:</emphasis>
The species epithet is named for the Global Foundation for Ocean Exploration whose efforts resulted in the collection of this very small and evasive species
</paragraph>
</subSubSection>
<subSubSection id="C37D6783921FE94EEBA94AFFFA46FF09" lastPageId="53" lastPageNumber="54" pageId="52" pageNumber="53" type="diagnosis">
<paragraph id="8BD83408921FE94EEBA94AFFFA46FF09" blockId="52.[151,1437,1699,2013]" lastBlockId="53.[151,1437,151,2013]" lastPageId="53" lastPageNumber="54" pageId="52" pageNumber="53">
<emphasis id="B913E81A921FE94FEBA94AFFFE86F8D8" bold="true" box="[199,322,1770,1796]" pageId="52" pageNumber="53">Diagnosis.</emphasis>
Body pentagonal (R/r=1.3). Abactinal plates irregular to polygonal in outline, with a bare central surface with irregularly shaped concavities surrounded by granules. Smaller plates and plates in the interradial regions are covered by granules. Papular pores, if present were not evident. Superomarginal and inferomarginal plates are identical in morphology and wider interradially. Superomarginals wide (W&gt;L) with approximately 50% of surface (abactinal facing) bare in weakly circular pattern, lower half covered by coarse granules, 2040 (mostly about 30). Inferomarginal plates with distinct cone-shaped spines present along actinolateral edge. Inferomarginal surface covered by coarse, round granules, 100150, about 130140 evenly distributed. Actinal plates covered by 320 short, pointed conical spinelets. Each plate irregular to polygonal in shape. Furrow spines four to six, tips blunt. Adambulacral plate surface with 720 short blunt spinelets similar to those on other actinal plate surfaces.
</paragraph>
</subSubSection>
<subSubSection id="C37D6783921EE94EEBA94CCBFBC8FBA1" pageId="53" pageNumber="54" type="discussion">
<paragraph id="8BD83408921EE94EEBA94CCBFDD7FE9D" blockId="53.[151,1437,151,2013]" pageId="53" pageNumber="54">
<emphasis id="B913E81A921EE94EEBA94CCBFE8BFF24" bold="true" box="[199,335,222,248]" pageId="53" pageNumber="54">Comments.</emphasis>
This species is placed within
<taxonomicName id="4C674F8B921EE94EE9C34CCAFCF2FF24" authorityName="Verrill" authorityYear="1899" box="[685,822,223,248]" class="Asteroidea" family="Goniasteridae" genus="Litonotaster" kingdom="Animalia" order="Valvatida" pageId="53" pageNumber="54" phylum="Echinodermata" rank="genus">
<emphasis id="B913E81A921EE94EE9C34CCAFCF2FF24" box="[685,822,223,248]" italics="true" pageId="53" pageNumber="54">Litonotaster</emphasis>
</taxonomicName>
primarily based on the shape and arrangement of the abactinal and marginal plates. Many of the actinal accessories, primarily the granules are pointed as they are in the Atlantic
<taxonomicName id="4C674F8B921EE94EEB964D32FDC8FE9D" baseAuthorityName="Perrier" baseAuthorityYear="1884" box="[248,524,295,321]" class="Asteroidea" family="Goniasteridae" genus="Litonotaster" kingdom="Animalia" order="Valvatida" pageId="53" pageNumber="54" phylum="Echinodermata" rank="species" species="intermedius">
<emphasis id="B913E81A921EE94EEB964D32FDC8FE9D" box="[248,524,295,321]" italics="true" pageId="53" pageNumber="54">Litonotaster intermedius</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BD83408921EE94EEBA94D5EFB6DFDE1" blockId="53.[151,1437,151,2013]" pageId="53" pageNumber="54">
This is the first species of
<taxonomicName id="4C674F8B921EE94EEA9C4D5EFDBFFEB8" authorityName="Verrill" authorityYear="1899" box="[498,635,331,356]" class="Asteroidea" family="Goniasteridae" genus="Litonotaster" kingdom="Animalia" order="Valvatida" pageId="53" pageNumber="54" phylum="Echinodermata" rank="genus">
<emphasis id="B913E81A921EE94EEA9C4D5EFDBFFEB8" box="[498,635,331,356]" italics="true" pageId="53" pageNumber="54">Litonotaster</emphasis>
</taxonomicName>
which has been observed
<taxonomicName id="4C674F8B921EE94EE8CB4D5EFC2EFEB8" box="[933,1002,331,356]" class="Asteroidea" family="Goniasteridae" genus="Iconaster" kingdom="Animalia" order="Valvatida" pageId="53" pageNumber="74" phylum="Echinodermata" rank="species" species="situ">
<emphasis id="B913E81A921EE94EE8CB4D5EFC2EFEB8" box="[933,1002,331,356]" italics="true" pageId="53" pageNumber="54">in situ</emphasis>
</taxonomicName>
and the second species to be recorded from the Pacific.
<taxonomicName id="4C674F8B921EE94EEA364D7AFDDBFE55" authority="Mah, 2022" authorityName="Mah" authorityYear="2022" box="[344,543,367,393]" class="Asteroidea" family="Goniasteridae" genus="Litonotaster" kingdom="Animalia" order="Valvatida" pageId="53" pageNumber="54" phylum="Echinodermata" rank="species" species="gfoei" status="sp. nov.">
<emphasis id="B913E81A921EE94EEA364D7AFDDBFE55" box="[344,543,367,393]" italics="true" pageId="53" pageNumber="54">Litonotaster gfoei</emphasis>
</taxonomicName>
<emphasis id="B913E81A921EE94EE94B4D7AFDA7FE55" bold="true" box="[549,611,367,393]" pageId="53" pageNumber="54">
<taxonomicNameLabel id="A2205561921EE94EE94B4D7AFDA7FE55" box="[549,611,367,393]" pageId="53" pageNumber="54" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
shows different characters that distinguish it from other
<taxonomicName id="4C674F8B921EE94EEFB64D7AFA5CFE55" box="[1240,1432,367,393]" class="Asteroidea" family="Goniasteridae" genus="Litonotaster" kingdom="Animalia" order="Valvatida" pageId="53" pageNumber="54" phylum="Echinodermata" rank="species" species="undetermined">
<emphasis id="B913E81A921EE94EEFB64D7AFAA5FE54" box="[1240,1377,367,392]" italics="true" pageId="53" pageNumber="54">Litonotaster</emphasis>
spp.
</taxonomicName>
, including its pentagonal body shape (R/r=1.3) (others are more stellate), the presence of triangular or dagger-shaped spinelets on the actinolateral edge of the inferomarginal spines (others have conical spines or granules), and the apparent absence of pedicellariae (present on all other known species).
<taxonomicName id="4C674F8B921EE94EE8F84DCEFB9AFE29" authority="Mah, 2022" authorityName="Mah" authorityYear="2022" box="[918,1118,475,501]" class="Asteroidea" family="Goniasteridae" genus="Litonotaster" kingdom="Animalia" order="Valvatida" pageId="53" pageNumber="54" phylum="Echinodermata" rank="species" species="gfoei" status="sp. nov.">
<emphasis id="B913E81A921EE94EE8F84DCEFB9AFE29" box="[918,1118,475,501]" italics="true" pageId="53" pageNumber="54">Litonotaster gfoei</emphasis>
</taxonomicName>
<emphasis id="B913E81A921EE94EEF0A4DCEFB66FE29" bold="true" box="[1124,1186,475,501]" pageId="53" pageNumber="54">
<taxonomicNameLabel id="A2205561921EE94EEF0A4DCEFB66FE29" box="[1124,1186,475,501]" pageId="53" pageNumber="54" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is most different from the Atlantic species, in that in addition to the differences, it and the East Pacific
<taxonomicName id="4C674F8B921EE94EEF704DEAFACCFDC5" authorityName="H. L. Clark" authorityYear="1920" box="[1054,1288,511,537]" class="Asteroidea" family="Goniasteridae" genus="Litonotaster" kingdom="Animalia" order="Valvatida" pageId="53" pageNumber="54" phylum="Echinodermata" rank="species" species="tumidus">
<emphasis id="B913E81A921EE94EEF704DEAFACCFDC5" box="[1054,1288,511,537]" italics="true" pageId="53" pageNumber="54">Litonotaster tumidus</emphasis>
</taxonomicName>
have significantly fewer furrow spines (four to six) versus five to eight in
<taxonomicName id="4C674F8B921EE94EE83E4E36FC0DFDE1" authorityName="Halpern" authorityYear="1969" box="[848,969,547,573]" class="Asteroidea" family="Goniasteridae" genus="Litonotaster" kingdom="Animalia" order="Valvatida" pageId="53" pageNumber="54" phylum="Echinodermata" rank="species" species="africans">
<emphasis id="B913E81A921EE94EE83E4E36FC0DFDE1" box="[848,969,547,573]" italics="true" pageId="53" pageNumber="54">L. africans</emphasis>
</taxonomicName>
and
<emphasis id="B913E81A921EE94EEF6E4E36FB6DFDE1" box="[1024,1193,547,573]" italics="true" pageId="53" pageNumber="54">
<taxonomicName id="4C674F8B921EE94EEF6E4E36FB61FDE1" baseAuthorityName="Perrier" baseAuthorityYear="1884" box="[1024,1189,547,573]" class="Asteroidea" family="Goniasteridae" genus="Litonotaster" kingdom="Animalia" order="Valvatida" pageId="53" pageNumber="54" phylum="Echinodermata" rank="species" species="intermedius">L. intermedius</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="8BD83408921EE94EEBA94E52FAA9FD2D" blockId="53.[151,1437,151,2013]" pageId="53" pageNumber="54">
Observations of
<emphasis id="B913E81A921EE94EEAED4E52FD5EFDBD" box="[387,666,583,609]" italics="true" pageId="53" pageNumber="54">
<taxonomicName id="4C674F8B921EE94EEAED4E52FE0DFDBC" box="[387,457,583,608]" class="Asteroidea" family="Goniasteridae" genus="Iconaster" kingdom="Animalia" order="Valvatida" pageId="53" pageNumber="74" phylum="Echinodermata" rank="species" species="situ">in situ</taxonomicName>
<taxonomicName id="4C674F8B921EE94EEABF4E52FD5EFDBD" authorityName="Mah" authorityYear="2022" box="[465,666,583,609]" class="Asteroidea" family="Goniasteridae" genus="Litonotaster" kingdom="Animalia" order="Valvatida" pageId="53" pageNumber="54" phylum="Echinodermata" rank="species" species="gfoei">Litonotaster gfoei</taxonomicName>
</emphasis>
suggest that some characters on the abactinal surface appear differently on living specimens than on the post-collection specimen. Most notably the distribution of granules on the abactinal surface appear more evenly distributed on the
<taxonomicName id="4C674F8B921EE94EE84E4E9AFCACFD74" box="[800,872,655,680]" class="Asteroidea" family="Goniasteridae" genus="Iconaster" kingdom="Animalia" order="Valvatida" pageId="53" pageNumber="74" phylum="Echinodermata" rank="species" species="situ">
<emphasis id="B913E81A921EE94EE84E4E9AFCACFD74" box="[800,872,655,680]" italics="true" pageId="53" pageNumber="54">in situ</emphasis>
</taxonomicName>
specimens versus the preserved specimen which shows numerous central bald areas on each plate (
<figureCitation id="135C288D921EE94EE9A54EA6FCEBFD11" box="[715,815,691,717]" captionStart="FIGURE 18" captionStartId="54.[151,250,1833,1857]" captionTargetBox="[205,1382,184,1809]" captionTargetId="figure-28@54.[204,1384,181,1809]" captionTargetPageId="54" captionText="FIGURE 18. Litonotaster gfoei n. sp. A. Abactinal view. Scale bar=2.0 mm. Inset showing surface detail. Largest individual plate diameter=1.0 mm. B. In situ image. Arrow showing inferomarginal spines extended, American Samoa, 2457 m. Scale bar=2.0 mm. C. Actinal view. Scale bar=2.0 mm. D. Actinal-Lateral view. Arrow showing spinelets on inferomarginal edge. Scale bar=1.0 mm. E. Actinal surface showing details. Scale bar=1.0 mm. F. Adambulacral spination, including furrow and subambulacral spines. Scale bar=0.75 mm. G. In situ image showing specimen with papular pores, Mozart Seamount, North Pacific." figureDoi="http://doi.org/10.5281/zenodo.6821080" httpUri="https://zenodo.org/record/6821080/files/figure.png" pageId="53" pageNumber="54">Fig. 18A</figureCitation>
vs. 18B). The presence of these irregular bald spots on granulated abactinal plates is a variable character observed in Atlantic species of
<taxonomicName id="4C674F8B921EE94EEF4D4EC2FAADFD2D" authority="(Halpern, 1969)" baseAuthorityName="Halpern" baseAuthorityYear="1969" box="[1059,1385,727,753]" class="Asteroidea" family="Goniasteridae" genus="Litonotaster" kingdom="Animalia" order="Valvatida" pageId="53" pageNumber="54" phylum="Echinodermata" rank="genus">
<emphasis id="B913E81A921EE94EEF4D4EC2FB68FD2C" box="[1059,1196,727,752]" italics="true" pageId="53" pageNumber="54">Litonotaster</emphasis>
(
<bibRefCitation id="EFF649F9921EE94EEFD54EC2FAA5FD2D" author="Halpern, J. A." box="[1211,1377,727,753]" pageId="53" pageNumber="54" pagination="129 - 142" refId="ref46522" refString="Halpern, J. A. (1969) Biological Investigations of the deep sea. 46. The genus Litonotaster (Echinodermata, Asteroidea). Proceedings of the Biological Society of Washington, 82, 129 - 142." type="journal article" year="1969">Halpern, 1969</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="8BD83408921EE94EEBA94EEEFE6AFC81" blockId="53.[151,1437,151,2013]" pageId="53" pageNumber="54">
When observed
<taxonomicName id="4C674F8B921EE94EEA114EEEFE01FCC8" box="[383,453,763,788]" class="Asteroidea" family="Goniasteridae" genus="Iconaster" kingdom="Animalia" order="Valvatida" pageId="53" pageNumber="74" phylum="Echinodermata" rank="species" species="situ">
<emphasis id="B913E81A921EE94EEA114EEEFE01FCC8" box="[383,453,763,788]" italics="true" pageId="53" pageNumber="54">in situ</emphasis>
</taxonomicName>
, the numerous small spinelets in series around the actinolateral edge on the inferomarginal plates are prominent and directed outward (
<figureCitation id="135C288D921EE94EE9A54F0AFCF4FCE5" box="[715,816,799,825]" captionStart="FIGURE 18" captionStartId="54.[151,250,1833,1857]" captionTargetBox="[205,1382,184,1809]" captionTargetId="figure-28@54.[204,1384,181,1809]" captionTargetPageId="54" captionText="FIGURE 18. Litonotaster gfoei n. sp. A. Abactinal view. Scale bar=2.0 mm. Inset showing surface detail. Largest individual plate diameter=1.0 mm. B. In situ image. Arrow showing inferomarginal spines extended, American Samoa, 2457 m. Scale bar=2.0 mm. C. Actinal view. Scale bar=2.0 mm. D. Actinal-Lateral view. Arrow showing spinelets on inferomarginal edge. Scale bar=1.0 mm. E. Actinal surface showing details. Scale bar=1.0 mm. F. Adambulacral spination, including furrow and subambulacral spines. Scale bar=0.75 mm. G. In situ image showing specimen with papular pores, Mozart Seamount, North Pacific." figureDoi="http://doi.org/10.5281/zenodo.6821080" httpUri="https://zenodo.org/record/6821080/files/figure.png" pageId="53" pageNumber="54">Fig. 18B</figureCitation>
) versus on the preserved specimen where they appear more loosely articulated.
</paragraph>
<paragraph id="8BD83408921EE94EEBA94F72FB94FC79" blockId="53.[151,1437,151,2013]" pageId="53" pageNumber="54">
<taxonomicName id="4C674F8B921EE94EEBA94F72FE4AFC5D" authority="Mah, 2022" authorityName="Mah" authorityYear="2022" box="[199,398,871,897]" class="Asteroidea" family="Goniasteridae" genus="Litonotaster" kingdom="Animalia" order="Valvatida" pageId="53" pageNumber="54" phylum="Echinodermata" rank="species" species="gfoei" status="sp. nov.">
<emphasis id="B913E81A921EE94EEBA94F72FE4AFC5D" box="[199,398,871,897]" italics="true" pageId="53" pageNumber="54">Litonotaster gfoei</emphasis>
</taxonomicName>
<emphasis id="B913E81A921EE94EEAFB4F72FE16FC5D" bold="true" box="[405,466,871,897]" pageId="53" pageNumber="54">
<taxonomicNameLabel id="A2205561921EE94EEAFB4F72FE16FC5D" box="[405,466,871,897]" pageId="53" pageNumber="54" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
was never observed in abundance but it does appear to show a widespread distribution, ranging from Johnson Atoll and Musicians Seamount to the
<collectingCountry id="F3707498921EE94EE8564F9EFC3DFC79" box="[824,1017,907,933]" name="American Samoa" pageId="53" pageNumber="54">American Samoa</collectingCountry>
region.
</paragraph>
<paragraph id="8BD83408921EE94EEBA94FBBFBF2FBE9" blockId="53.[151,1437,151,2013]" pageId="53" pageNumber="54">
<emphasis id="B913E81A921EE94EEBA94FBBFE16FC14" bold="true" box="[199,466,942,968]" pageId="53" pageNumber="54">Ecological Comments.</emphasis>
All the observations of this species (e.g.
<figureCitation id="135C288D921EE94EE8CB4FBAFBCDFC15" box="[933,1033,943,969]" captionStart="FIGURE 18" captionStartId="54.[151,250,1833,1857]" captionTargetBox="[205,1382,184,1809]" captionTargetId="figure-28@54.[204,1384,181,1809]" captionTargetPageId="54" captionText="FIGURE 18. Litonotaster gfoei n. sp. A. Abactinal view. Scale bar=2.0 mm. Inset showing surface detail. Largest individual plate diameter=1.0 mm. B. In situ image. Arrow showing inferomarginal spines extended, American Samoa, 2457 m. Scale bar=2.0 mm. C. Actinal view. Scale bar=2.0 mm. D. Actinal-Lateral view. Arrow showing spinelets on inferomarginal edge. Scale bar=1.0 mm. E. Actinal surface showing details. Scale bar=1.0 mm. F. Adambulacral spination, including furrow and subambulacral spines. Scale bar=0.75 mm. G. In situ image showing specimen with papular pores, Mozart Seamount, North Pacific." figureDoi="http://doi.org/10.5281/zenodo.6821080" httpUri="https://zenodo.org/record/6821080/files/figure.png" pageId="53" pageNumber="54">Fig. 18B</figureCitation>
) were made of individuals on manganese encrusted hard rock substrata, which were almost always topped with light colored covering, thought to be either sediment or fallen organic flocculence (i.e. marine snow). Although the oral surface was not observed, it seems possible that this light-colored flocculence/sediment is food for this species.
</paragraph>
<paragraph id="8BD83408921EE94EEBA9482BFBC8FBA1" blockId="53.[151,1437,151,2013]" pageId="53" pageNumber="54">
<emphasis id="B913E81A921EE94EEBA9482BFE98FB84" bold="true" box="[199,348,1086,1112]" pageId="53" pageNumber="54">Occurrence:</emphasis>
<materialsCitation id="3B0F3E55921EE94EEA0C482AFA90FB85" box="[354,1364,1087,1113]" country="American Samoa" location="Rose Atoll" pageId="53" pageNumber="54" specimenCount="1" stateProvince="Rose Island">
<collectingCountry id="F3707498921EE94EEA0C482AFDE2FB85" box="[354,550,1087,1113]" name="American Samoa" pageId="53" pageNumber="54">American Samoa</collectingCountry>
,
<collectingRegion id="49A3FAEA921EE94EE95E482AFD6DFB85" box="[560,681,1087,1113]" country="American Samoa" name="Rose Island" pageId="53" pageNumber="54">Rose Atoll</collectingRegion>
, Moki seamount, Utu seamount, Malulu seamount, Se Guyot
</materialsCitation>
,
<materialsCitation id="3B0F3E55921EE94EEE30482AFBC8FBA1" country="American Samoa" elevation="2859" location="Musicians Seamount region" municipality="Mozart" pageId="53" pageNumber="54" specimenCount="1" stateProvince="Johnson Atoll">
<collectingRegion id="49A3FAEA921EE94EEE30482AFF3AFBA1" inferredBySuffix="true" pageId="53" pageNumber="54">Johnson Atoll</collectingRegion>
region,
<collectingMunicipality id="6BBCAE72921EE94EEA344876FE6FFBA1" box="[346,427,1123,1149]" pageId="53" pageNumber="54">Mozart</collectingMunicipality>
seamount,
<location id="8EB862D3921EE94EE9454876FCA2FBA1" LSID="urn:lsid:plazi:treatment:03CE851E921FE94CEBF94A22FD0EFA89:8EB862D3921EE94EE9454876FCA2FBA1" box="[555,870,1123,1149]" country="American Samoa" municipality="Mozart" name="Musicians Seamount region" pageId="53" pageNumber="54" stateProvince="Johnson Atoll">Musicians Seamount region</location>
,
<quantity id="4C9F99ED921EE94EE81F4876FBC8FBA1" box="[881,1036,1123,1149]" metricMagnitude="3" metricUnit="m" metricValue="2.8589999999999995" metricValueMax="3.7079999999999997" metricValueMin="2.0100000000000002" pageId="53" pageNumber="54" unit="m" value="2859.0" valueMax="3708.0" valueMin="2010.0">
<elevation id="004AD33B921EE94EE81F4876FBC8FBA1" box="[881,1036,1123,1149]" metricMagnitude="3" metricUnit="m" metricValue="2.8589999999999995" metricValueMax="3.7079999999999997" metricValueMin="2.0100000000000002" pageId="53" pageNumber="54" unit="m" value="2859.0" valueMax="3708.0" valueMin="2010.0">20103708 m</elevation>
</quantity>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C37D6783921EE94CEBA94893FD27FDBD" lastPageId="55" lastPageNumber="56" pageId="53" pageNumber="54" type="description">
<paragraph id="8BD83408921EE94EEBA94893FE56FB19" blockId="53.[151,1437,151,2013]" pageId="53" pageNumber="54">
<emphasis id="B913E81A921EE94EEBA94893FE9DFB7C" bold="true" box="[199,345,1158,1184]" pageId="53" pageNumber="54">Description.</emphasis>
Body pentagonal (R/r=1.3), interradial arcs straight to weakly curved (
<figureCitation id="135C288D921EE94EEF044892FB15FB7D" box="[1130,1233,1159,1185]" captionStart="FIGURE 18" captionStartId="54.[151,250,1833,1857]" captionTargetBox="[205,1382,184,1809]" captionTargetId="figure-28@54.[204,1384,181,1809]" captionTargetPageId="54" captionText="FIGURE 18. Litonotaster gfoei n. sp. A. Abactinal view. Scale bar=2.0 mm. Inset showing surface detail. Largest individual plate diameter=1.0 mm. B. In situ image. Arrow showing inferomarginal spines extended, American Samoa, 2457 m. Scale bar=2.0 mm. C. Actinal view. Scale bar=2.0 mm. D. Actinal-Lateral view. Arrow showing spinelets on inferomarginal edge. Scale bar=1.0 mm. E. Actinal surface showing details. Scale bar=1.0 mm. F. Adambulacral spination, including furrow and subambulacral spines. Scale bar=0.75 mm. G. In situ image showing specimen with papular pores, Mozart Seamount, North Pacific." figureDoi="http://doi.org/10.5281/zenodo.6821080" httpUri="https://zenodo.org/record/6821080/files/figure.png" pageId="53" pageNumber="54">Fig. 18A</figureCitation>
). Body thick. Arm tips distinct, upturned.
</paragraph>
<paragraph id="8BD83408921EE94EEBA948DAFC36F9A9" blockId="53.[151,1437,151,2013]" pageId="53" pageNumber="54">
Abactinal plates flattened, trigonal, polygonal to round in outline, arranged irregularly. Proximal plates wider becoming smaller and more irregular in shape distally adjacent to superomarginals. Central surface of most plates bare with peripheral edges of each plate covered by round granules (
<figureCitation id="135C288D921EE94EE8F04902FBC6FAED" box="[926,1026,1303,1329]" captionStart="FIGURE 18" captionStartId="54.[151,250,1833,1857]" captionTargetBox="[205,1382,184,1809]" captionTargetId="figure-28@54.[204,1384,181,1809]" captionTargetPageId="54" captionText="FIGURE 18. Litonotaster gfoei n. sp. A. Abactinal view. Scale bar=2.0 mm. Inset showing surface detail. Largest individual plate diameter=1.0 mm. B. In situ image. Arrow showing inferomarginal spines extended, American Samoa, 2457 m. Scale bar=2.0 mm. C. Actinal view. Scale bar=2.0 mm. D. Actinal-Lateral view. Arrow showing spinelets on inferomarginal edge. Scale bar=1.0 mm. E. Actinal surface showing details. Scale bar=1.0 mm. F. Adambulacral spination, including furrow and subambulacral spines. Scale bar=0.75 mm. G. In situ image showing specimen with papular pores, Mozart Seamount, North Pacific." figureDoi="http://doi.org/10.5281/zenodo.6821080" httpUri="https://zenodo.org/record/6821080/files/figure.png" pageId="53" pageNumber="54">Fig. 18A</figureCitation>
and inset). Granules, 1030, mostly 20, covering plates interradially especially at contact with superomarginal plates. Observations of living animals show the granules forming a more evenly distributed, bumpy texture with no bare regions (
<figureCitation id="135C288D921EE94EEFC7494AFAF1FAA5" box="[1193,1333,1375,1401]" captionStart="FIGURE 18" captionStartId="54.[151,250,1833,1857]" captionTargetBox="[205,1382,184,1809]" captionTargetId="figure-28@54.[204,1384,181,1809]" captionTargetPageId="54" captionText="FIGURE 18. Litonotaster gfoei n. sp. A. Abactinal view. Scale bar=2.0 mm. Inset showing surface detail. Largest individual plate diameter=1.0 mm. B. In situ image. Arrow showing inferomarginal spines extended, American Samoa, 2457 m. Scale bar=2.0 mm. C. Actinal view. Scale bar=2.0 mm. D. Actinal-Lateral view. Arrow showing spinelets on inferomarginal edge. Scale bar=1.0 mm. E. Actinal surface showing details. Scale bar=1.0 mm. F. Adambulacral spination, including furrow and subambulacral spines. Scale bar=0.75 mm. G. In situ image showing specimen with papular pores, Mozart Seamount, North Pacific." figureDoi="http://doi.org/10.5281/zenodo.6821080" httpUri="https://zenodo.org/record/6821080/files/figure.png" pageId="53" pageNumber="54">Fig. 18B, G</figureCitation>
). Carinal plates largest, especially distally adjacent to the arm tip with only a single peripheral series of granules, 2040. Arm plates extend along arm tip to terminus. Most of distalmost arm plate surface bare. Plates weakly textured, not smooth. Weakly developed fasciolar groove present among distalmost plates. Living observations showed 330 papular openings per radial region (
<figureCitation id="135C288D921EE94EE94449FAFD56F9D5" box="[554,658,1519,1545]" captionStart="FIGURE 18" captionStartId="54.[151,250,1833,1857]" captionTargetBox="[205,1382,184,1809]" captionTargetId="figure-28@54.[204,1384,181,1809]" captionTargetPageId="54" captionText="FIGURE 18. Litonotaster gfoei n. sp. A. Abactinal view. Scale bar=2.0 mm. Inset showing surface detail. Largest individual plate diameter=1.0 mm. B. In situ image. Arrow showing inferomarginal spines extended, American Samoa, 2457 m. Scale bar=2.0 mm. C. Actinal view. Scale bar=2.0 mm. D. Actinal-Lateral view. Arrow showing spinelets on inferomarginal edge. Scale bar=1.0 mm. E. Actinal surface showing details. Scale bar=1.0 mm. F. Adambulacral spination, including furrow and subambulacral spines. Scale bar=0.75 mm. G. In situ image showing specimen with papular pores, Mozart Seamount, North Pacific." figureDoi="http://doi.org/10.5281/zenodo.6821080" httpUri="https://zenodo.org/record/6821080/files/figure.png" pageId="53" pageNumber="54">Fig. 18G</figureCitation>
). Images of the
<typeStatus id="54DC8AAA921EE94EE82649FAFC6EF9D5" box="[840,938,1519,1545]" pageId="53" pageNumber="54" type="holotype">holotype</typeStatus>
<taxonomicName id="4C674F8B921EE94EE8DC49FAFC33F9D4" box="[946,1015,1519,1544]" class="Asteroidea" family="Goniasteridae" genus="Iconaster" kingdom="Animalia" order="Valvatida" pageId="53" pageNumber="74" phylum="Echinodermata" rank="species" species="situ">
<emphasis id="B913E81A921EE94EE8DC49FAFC33F9D4" box="[946,1015,1519,1544]" italics="true" pageId="53" pageNumber="54">in situ</emphasis>
</taxonomicName>
showed only one or two opening per radial region with larger plates flanking the papular pores (
<figureCitation id="135C288D921EE94EE8404A06FC51F9F1" box="[814,917,1555,1581]" captionStart="FIGURE 18" captionStartId="54.[151,250,1833,1857]" captionTargetBox="[205,1382,184,1809]" captionTargetId="figure-28@54.[204,1384,181,1809]" captionTargetPageId="54" captionText="FIGURE 18. Litonotaster gfoei n. sp. A. Abactinal view. Scale bar=2.0 mm. Inset showing surface detail. Largest individual plate diameter=1.0 mm. B. In situ image. Arrow showing inferomarginal spines extended, American Samoa, 2457 m. Scale bar=2.0 mm. C. Actinal view. Scale bar=2.0 mm. D. Actinal-Lateral view. Arrow showing spinelets on inferomarginal edge. Scale bar=1.0 mm. E. Actinal surface showing details. Scale bar=1.0 mm. F. Adambulacral spination, including furrow and subambulacral spines. Scale bar=0.75 mm. G. In situ image showing specimen with papular pores, Mozart Seamount, North Pacific." figureDoi="http://doi.org/10.5281/zenodo.6821080" httpUri="https://zenodo.org/record/6821080/files/figure.png" pageId="53" pageNumber="54">Fig. 18B</figureCitation>
). The
<typeStatus id="54DC8AAA921EE94EE8B24A06FBFAF9F1" box="[988,1086,1555,1581]" pageId="53" pageNumber="54" type="holotype">holotype</typeStatus>
did not clearly display papular pores suggesting that they are cryptic or concealed, as only one or two pores were observed. Madreporite round flanked by approximately 1012 abactinal plates. No pedicellariae observed.
</paragraph>
<paragraph id="8BD83408921EE94CEBA94A6AFA59FEC1" blockId="53.[151,1437,151,2013]" lastBlockId="55.[151,1437,151,717]" lastPageId="55" lastPageNumber="56" pageId="53" pageNumber="54">
Superomarginals 12, inferomarginals 16 per interradius (arm tip to arm tip) (
<figureCitation id="135C288D921EE94EEF414A6AFB5EF945" box="[1071,1178,1663,1689]" captionStart="FIGURE 18" captionStartId="54.[151,250,1833,1857]" captionTargetBox="[205,1382,184,1809]" captionTargetId="figure-28@54.[204,1384,181,1809]" captionTargetPageId="54" captionText="FIGURE 18. Litonotaster gfoei n. sp. A. Abactinal view. Scale bar=2.0 mm. Inset showing surface detail. Largest individual plate diameter=1.0 mm. B. In situ image. Arrow showing inferomarginal spines extended, American Samoa, 2457 m. Scale bar=2.0 mm. C. Actinal view. Scale bar=2.0 mm. D. Actinal-Lateral view. Arrow showing spinelets on inferomarginal edge. Scale bar=1.0 mm. E. Actinal surface showing details. Scale bar=1.0 mm. F. Adambulacral spination, including furrow and subambulacral spines. Scale bar=0.75 mm. G. In situ image showing specimen with papular pores, Mozart Seamount, North Pacific." figureDoi="http://doi.org/10.5281/zenodo.6821080" httpUri="https://zenodo.org/record/6821080/files/figure.png" pageId="53" pageNumber="54">Fig. 18A</figureCitation>
). Superomarginals and inferomarginals offset, contact in zig zag arrangement. Lateral side distinct (
<figureCitation id="135C288D921EE94EE8904AB6FB4DF961" box="[1022,1161,1699,1725]" captionStart="FIGURE 18" captionStartId="54.[151,250,1833,1857]" captionTargetBox="[205,1382,184,1809]" captionTargetId="figure-28@54.[204,1384,181,1809]" captionTargetPageId="54" captionText="FIGURE 18. Litonotaster gfoei n. sp. A. Abactinal view. Scale bar=2.0 mm. Inset showing surface detail. Largest individual plate diameter=1.0 mm. B. In situ image. Arrow showing inferomarginal spines extended, American Samoa, 2457 m. Scale bar=2.0 mm. C. Actinal view. Scale bar=2.0 mm. D. Actinal-Lateral view. Arrow showing spinelets on inferomarginal edge. Scale bar=1.0 mm. E. Actinal surface showing details. Scale bar=1.0 mm. F. Adambulacral spination, including furrow and subambulacral spines. Scale bar=0.75 mm. G. In situ image showing specimen with papular pores, Mozart Seamount, North Pacific." figureDoi="http://doi.org/10.5281/zenodo.6821080" httpUri="https://zenodo.org/record/6821080/files/figure.png" pageId="53" pageNumber="54">Fig. 18A, B</figureCitation>
). Superomarginals wide (W&gt;L) with approximately 50% of surface (abactinal facing) bare in weakly circular pattern, bare surface becoming more narrower distalmost, widest interradially. Bare surface on superomarginals completely dominates surface on distal plates adjacent to terminal plates. Lower half of superomarginal plates covered by coarse granules, 2040 (mostly 30) evenly spaced. Superomarginal edge with convex contact on abactinal plates. Peripheral granules, approximately 1525 per side, approximately 3080 around each superomarginal plate. Inferomarginals interradially squarish in outline, more elongate than corresponding superomarginal plate becoming narrower distally. Inferomarginals covered by coarse, round granules, 100150, about 130140 evenly distributed (
<figureCitation id="135C288D921EE94EEF0A4B8AFB37F865" box="[1124,1267,1951,1977]" captionStart="FIGURE 18" captionStartId="54.[151,250,1833,1857]" captionTargetBox="[205,1382,184,1809]" captionTargetId="figure-28@54.[204,1384,181,1809]" captionTargetPageId="54" captionText="FIGURE 18. Litonotaster gfoei n. sp. A. Abactinal view. Scale bar=2.0 mm. Inset showing surface detail. Largest individual plate diameter=1.0 mm. B. In situ image. Arrow showing inferomarginal spines extended, American Samoa, 2457 m. Scale bar=2.0 mm. C. Actinal view. Scale bar=2.0 mm. D. Actinal-Lateral view. Arrow showing spinelets on inferomarginal edge. Scale bar=1.0 mm. E. Actinal surface showing details. Scale bar=1.0 mm. F. Adambulacral spination, including furrow and subambulacral spines. Scale bar=0.75 mm. G. In situ image showing specimen with papular pores, Mozart Seamount, North Pacific." figureDoi="http://doi.org/10.5281/zenodo.6821080" httpUri="https://zenodo.org/record/6821080/files/figure.png" pageId="53" pageNumber="54">Fig. 18 C, D</figureCitation>
). Central interradial inferomarginal plates each with one distinctly enlarged (approximately
<quantity id="4C9F99ED921EE94EE8904BD6FB9FF801" box="[1022,1115,1987,2013]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="53" pageNumber="54" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">23 mm</quantity>
wide) paddle-shaped pedicellariae. Actinolateral edge along inferomarginal plates distinct. Actinolateral edge on inferomarginal plates with flat, triangular spines, 715 per plate (
<figureCitation id="135C288D921CE94CE9844CAEFCB9FF09" box="[746,893,187,213]" captionStart="FIGURE 18" captionStartId="54.[151,250,1833,1857]" captionTargetBox="[205,1382,184,1809]" captionTargetId="figure-28@54.[204,1384,181,1809]" captionTargetPageId="54" captionText="FIGURE 18. Litonotaster gfoei n. sp. A. Abactinal view. Scale bar=2.0 mm. Inset showing surface detail. Largest individual plate diameter=1.0 mm. B. In situ image. Arrow showing inferomarginal spines extended, American Samoa, 2457 m. Scale bar=2.0 mm. C. Actinal view. Scale bar=2.0 mm. D. Actinal-Lateral view. Arrow showing spinelets on inferomarginal edge. Scale bar=1.0 mm. E. Actinal surface showing details. Scale bar=1.0 mm. F. Adambulacral spination, including furrow and subambulacral spines. Scale bar=0.75 mm. G. In situ image showing specimen with papular pores, Mozart Seamount, North Pacific." figureDoi="http://doi.org/10.5281/zenodo.6821080" httpUri="https://zenodo.org/record/6821080/files/figure.png" pageId="55" pageNumber="56">Fig. 18B, D</figureCitation>
). Spines absent from distalmost inferomarginal plates. Similar triangular spines, 2040, 50% of the width of those on the actinolateral edge, present on actinal surface of inferomarginal plates with large, round bald spot. Terminal plate diamond-shaped, surface bare.
</paragraph>
<caption id="DF186480921DE94DEBF94B3CFF21F829" ID-DOI="http://doi.org/10.5281/zenodo.6821080" ID-Zenodo-Dep="6821080" httpUri="https://zenodo.org/record/6821080/files/figure.png" pageId="54" pageNumber="55" startId="54.[151,250,1833,1857]" targetBox="[205,1382,184,1809]" targetPageId="54">
<paragraph id="8BD83408921DE94DEBF94B3CFF21F829" blockId="54.[151,1437,1833,2037]" pageId="54" pageNumber="55">
<emphasis id="B913E81A921DE94DEBF94B3CFD85F89D" bold="true" box="[151,577,1833,1858]" pageId="54" pageNumber="55">
FIGURE 18.
<taxonomicName id="4C674F8B921DE94DEA464B3CFE26F89D" authority="Mah, 2022" authorityName="Mah" authorityYear="2022" box="[296,482,1833,1857]" class="Asteroidea" family="Goniasteridae" genus="Litonotaster" kingdom="Animalia" order="Valvatida" pageId="54" pageNumber="55" phylum="Echinodermata" rank="species" species="gfoei" status="sp. nov.">
<emphasis id="B913E81A921DE94DEA464B3CFE26F89D" bold="true" box="[296,482,1833,1857]" italics="true" pageId="54" pageNumber="55">Litonotaster gfoei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2205561921DE94DEA874B3FFDE7F89E" box="[489,547,1834,1858]" pageId="54" pageNumber="55" rank="species">n. sp.</taxonomicNameLabel>
A.
</emphasis>
Abactinal view. Scale bar=2.0 mm. Inset showing surface detail. Largest individual plate diameter=1.0 mm.
<emphasis id="B913E81A921DE94DEAF94B5BFE6AF8BA" bold="true" box="[407,430,1870,1894]" pageId="54" pageNumber="55">B.</emphasis>
<taxonomicName id="4C674F8B921DE94DEAD94B5BFE3EF8B9" box="[439,506,1870,1893]" class="Asteroidea" family="Goniasteridae" genus="Iconaster" kingdom="Animalia" order="Valvatida" pageId="54" pageNumber="55" phylum="Echinodermata" rank="species" species="situ">
<emphasis id="B913E81A921DE94DEAD94B5BFE3EF8B9" box="[439,506,1870,1893]" italics="true" pageId="54" pageNumber="55">In situ</emphasis>
</taxonomicName>
image. Arrow showing inferomarginal spines extended, American Samoa, 2457 m. Scale bar=2.0 mm.
<emphasis id="B913E81A921DE94DEA4D4B64FEF8F855" bold="true" box="[291,316,1905,1929]" pageId="54" pageNumber="55">C.</emphasis>
Actinal view. Scale bar=2.0 mm.
<emphasis id="B913E81A921DE94DE9F14B67FD73F856" bold="true" box="[671,695,1906,1930]" pageId="54" pageNumber="55">D.</emphasis>
Actinal-Lateral view. Arrow showing spinelets on inferomarginal edge. Scale bar=1.0 mm.
<emphasis id="B913E81A921DE94DEA0D4B83FEBFF872" bold="true" box="[355,379,1942,1966]" pageId="54" pageNumber="55">E.</emphasis>
Actinal surface showing details. Scale bar=1.0 mm.
<emphasis id="B913E81A921DE94DE8CD4B83FC72F872" bold="true" box="[931,950,1942,1966]" pageId="54" pageNumber="55">F.</emphasis>
Adambulacral spination, including furrow and subambulacral spines. Scale bar=0.75 mm.
<emphasis id="B913E81A921DE94DE9364BACFDB6F80D" bold="true" box="[600,626,1977,2001]" pageId="54" pageNumber="55">G.</emphasis>
<taxonomicName id="4C674F8B921DE94DE9144BAFFD78F80D" box="[634,700,1978,2001]" class="Asteroidea" family="Goniasteridae" genus="Iconaster" kingdom="Animalia" order="Valvatida" pageId="54" pageNumber="55" phylum="Echinodermata" rank="species" species="situ">
<emphasis id="B913E81A921DE94DE9144BAFFD78F80D" box="[634,700,1978,2001]" italics="true" pageId="54" pageNumber="55">In situ</emphasis>
</taxonomicName>
image showing specimen with papular pores, Mozart Seamount, North Pacific.
</paragraph>
</caption>
<paragraph id="8BD83408921CE94CEBA94D32FCB3FDE1" blockId="55.[151,1437,151,717]" pageId="55" pageNumber="56">
Actinal surface in three complete series in chevron arrangement (
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). Actinal plates covered by 320 short, pointed conical spinelets (
<figureCitation id="135C288D921CE94CE9694D5EFDA9FEB9" box="[519,621,331,357]" captionStart="FIGURE 18" captionStartId="54.[151,250,1833,1857]" captionTargetBox="[205,1382,184,1809]" captionTargetId="figure-28@54.[204,1384,181,1809]" captionTargetPageId="54" captionText="FIGURE 18. Litonotaster gfoei n. sp. A. Abactinal view. Scale bar=2.0 mm. Inset showing surface detail. Largest individual plate diameter=1.0 mm. B. In situ image. Arrow showing inferomarginal spines extended, American Samoa, 2457 m. Scale bar=2.0 mm. C. Actinal view. Scale bar=2.0 mm. D. Actinal-Lateral view. Arrow showing spinelets on inferomarginal edge. Scale bar=1.0 mm. E. Actinal surface showing details. Scale bar=1.0 mm. F. Adambulacral spination, including furrow and subambulacral spines. Scale bar=0.75 mm. G. In situ image showing specimen with papular pores, Mozart Seamount, North Pacific." figureDoi="http://doi.org/10.5281/zenodo.6821080" httpUri="https://zenodo.org/record/6821080/files/figure.png" pageId="55" pageNumber="56">Fig. 18E</figureCitation>
). Each plate irregular to polygonal in shape, plate series adjacent to adambulacrals rectangular in outline, quadrate, plates becoming smaller adjacent to inferomarginals, 710 plates in this series from oral plates to terminal plate. Furrow spines 46 (
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), tips blunt. Adambulacral plate surface with 720 short blunt spinelets similar to those on other actinal plate surfaces. Oral plates each teardrop shaped with round convex edge facing onto actinal interradius. Each oral plate with ~2028 short spinelets facing into the tube foot furrow. Oral plate with exposed surface with approximately 2025 granules, with 1015 short spinelets on paired contact with other oral plate in each interradius (
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).
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Color in life was yellowish-white (
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).
</paragraph>
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Material Examined:
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Malulu, American Samoa-
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Mozart Seamount, Musicians Seamount Chain.
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