448 lines
58 KiB
XML
448 lines
58 KiB
XML
<document id="0A68A1199BD420ABDE0746AA47DDE14D" ID-DOI="10.11646/zootaxa.3852.4.5" ID-GBIF-Dataset="0d568f39-128d-4b01-b6ae-0e7d4f92a06a" ID-ISSN="1175-5326" ID-Zenodo-Dep="231188" ID-ZooBank="033A771C-839D-41C5-91C3-7D02A661F4C9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460885311263" checkinUser="plazi" docAuthor="Lojková, Soňa, Degma, Peter & Kodada, Ján" docDate="2014" docId="03C91474FFFC8E07FF097907FAF89BFC" docLanguage="en" docName="zt03852p488.pdf" docOrigin="Zootaxa 3852 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Parahelichus granulosus Deleve 1974" docType="treatment" docVersion="10" lastPageNumber="480" masterDocId="FFF06C0CFFFD8E02FF9E7D7DFF8A987C" masterDocTitle="Morphometry and redescription of Parahelichus granulosus (Delève, 1974) with description of P. pseudogranulosus, a new cryptic species of Long-toed Water Beetles (Coleoptera: Dryopidae) from Indochinese peninsula, and proposal of a new synonym for Praehelichus sericatus (Waterhouse, 1881) from China" masterLastPageNumber="488" masterPageNumber="475" pageNumber="476" updateTime="1698358941206" updateUser="plazi">
|
||
<mods:mods id="F9DBDA0F2939A25ECAE48BC4D5CAF16F" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="C8F438B7261560670F0AB7A28E748E85">
|
||
<mods:title id="5D6D444CA6C1F1C9E0BB4AB754C04B68">Morphometry and redescription of Parahelichus granulosus (Delève, 1974) with description of P. pseudogranulosus, a new cryptic species of Long-toed Water Beetles (Coleoptera: Dryopidae) from Indochinese peninsula, and proposal of a new synonym for Praehelichus sericatus (Waterhouse, 1881) from China</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="1AFFCDC41C4533B6775D4A67FCA26150" type="personal">
|
||
<mods:role id="EB4F864B245F72812AB6EF13FDF66974">
|
||
<mods:roleTerm id="4BE4928A7946795EC6F0D00E8175976C">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="D5ED7CFFBC99C372CABDA0C7AD9594F6">Lojková, Soňa</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="B0CC3DBA37EB16E7D633EAB72C92D200" type="personal">
|
||
<mods:role id="C596AA53B4BE69120A3C4FEA2005D61C">
|
||
<mods:roleTerm id="571C84EC3CF708ED79B4CA3519A7E2B9">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="D7148D87EC1BB35D168C0B623C3DFFCE">Degma, Peter</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="6E7C6D1AF8CC6312B6929AF65ACB5ECD" type="personal">
|
||
<mods:role id="B45C0E10C56A63FBE4AE8E4EC77BDA07">
|
||
<mods:roleTerm id="7B6AFA2A15CE709234BDC03386E038E2">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="744CA1FD7F6D6587B9543D34C781CB87">Kodada, Ján</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="9F69066EC87C667F0C123258D0D9E2DA">text</mods:typeOfResource>
|
||
<mods:relatedItem id="2E8918E631C124FC124E0BF41831A3F8" type="host">
|
||
<mods:titleInfo id="83C24B36963F589E49499AC2CCFEB881">
|
||
<mods:title id="70603452E2E55E47EE360580F9B1676A">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="B3D2E35CA7E661E9C7B105B24922B421">
|
||
<mods:date id="FFC676AE6C726CF6AF846E539BF93C3B">2014</mods:date>
|
||
<mods:detail id="73FAD0D8FCDE31B73CCC438ADCCCADF9" type="volume">
|
||
<mods:number id="9970D1C0228D721417E6BAB4ECD1DC5C">3852</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="62144E5914176E8392AE52511653A796" type="issue">
|
||
<mods:number id="96F01DA27386E02518F6D1B42D502B03">4</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="6C86F5DFDFA49F5CEE45687369F1E414" unit="page">
|
||
<mods:start id="A9F3813EF4B23E45774355FA13BFCFBA">475</mods:start>
|
||
<mods:end id="ECE5055B18088CF6FE6279CC47C6CE44">488</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="199135529170336A7CA6ED477F02F7D1">journal article</mods:classification>
|
||
<mods:identifier id="F32F4A54443B4B63709F4FF5FDD6EDAC" type="DOI">10.11646/zootaxa.3852.4.5</mods:identifier>
|
||
<mods:identifier id="2EAB4AF2D16B2AD136F4BFCB00619F3D" type="GBIF-Dataset">0d568f39-128d-4b01-b6ae-0e7d4f92a06a</mods:identifier>
|
||
<mods:identifier id="0E927D8B0B665D69A91A8ED44AEF6B77" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="BC625E956B0E22850004A832C0D801D1" type="Zenodo-Dep">231188</mods:identifier>
|
||
<mods:identifier id="20CF17ACA548A2811A013E6FC792E843" type="ZooBank">033A771C-839D-41C5-91C3-7D02A661F4C9</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03C91474FFFC8E07FF097907FAF89BFC" ID-DOI="http://doi.org/10.5281/zenodo.5630235" ID-GBIF-Taxon="119586606" ID-Zenodo-Dep="5630235" LSID="urn:lsid:plazi:treatment:03C91474FFFC8E07FF097907FAF89BFC" httpUri="http://treatment.plazi.org/id/03C91474FFFC8E07FF097907FAF89BFC" lastPageId="5" lastPageNumber="480" pageId="1" pageNumber="476">
|
||
<subSubSection id="C37AF6E9FFFC8E03FF097907FDF79CE9" box="[151,637,1146,1173]" pageId="1" pageNumber="476" type="nomenclature">
|
||
<paragraph id="8BDFA562FFFC8E03FF097907FDF79CE9" blockId="1.[151,637,1146,1173]" box="[151,637,1146,1173]" pageId="1" pageNumber="476">
|
||
<heading id="D097120EFFFC8E03FF097907FDF79CE9" bold="true" box="[151,637,1146,1173]" fontSize="11" level="1" pageId="1" pageNumber="476" reason="1">
|
||
<taxonomicName id="4C60DEE1FFFC8E03FF097907FDF79CE9" authority="Deleve, 1974" authorityName="Deleve" authorityYear="1974" box="[151,637,1146,1173]" class="Insecta" family="Dryopidae" genus="Parahelichus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="476" phylum="Arthropoda" rank="species" species="granulosus">
|
||
<emphasis id="B9147970FFFC8E03FF097907FDF79CE9" bold="true" box="[151,637,1146,1173]" pageId="1" pageNumber="476">
|
||
<emphasis id="B9147970FFFC8E03FF097907FE4C9CE8" bold="true" box="[151,454,1146,1172]" italics="true" pageId="1" pageNumber="476">Parahelichus granulosus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFF1D893FFFC8E03FE487906FDFF9CE9" author="Deleve" box="[470,629,1147,1173]" pageId="1" pageNumber="476" refString="Deleve, J. (1974) Helichus granulosus (Coleoptera, Dryopidae) une nouvelle espece du Vietnam. Annales Historico-Naturales Musei Nationalis Hungarici, 66, 171 - 172." type="journal article" year="1974">Delève, 1974</bibRefCitation>
|
||
)
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37AF6E9FFFC8E03FF0979BCFAF49E41" pageId="1" pageNumber="476" type="reference_group">
|
||
<paragraph id="8BDFA562FFFC8E03FF0979BCFDA89CAB" blockId="1.[151,546,1217,1239]" box="[151,546,1217,1239]" pageId="1" pageNumber="476">
|
||
<treatmentCitationGroup id="AB70824CFFFC8E03FF0979BCFDA89CAB" box="[151,546,1217,1239]" pageId="1" pageNumber="476">
|
||
<taxonomicName id="4C60DEE1FFFC8E03FF0979BCFDA89CAB" authority="Deleve 1974: 171" authorityName="Deleve" authorityPageNumber="171" authorityYear="1974" box="[151,546,1217,1239]" class="Insecta" family="Dryopidae" genus="Helichus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="476" phylum="Arthropoda" rank="species" species="granulosus">
|
||
<emphasis id="B9147970FFFC8E03FF0979BCFEE29CAB" box="[151,360,1217,1239]" italics="true" pageId="1" pageNumber="476">Helichus granulosus</emphasis>
|
||
<treatmentCitation id="0AC18373FFFC8E03FEF079BCFDA89CAB" author="Deleve" box="[366,546,1217,1239]" page="171" pageId="1" pageNumber="476" year="1974">
|
||
<bibRefCitation id="EFF1D893FFFC8E03FEF079BCFE789CAB" author="Deleve" box="[366,498,1217,1239]" pageId="1" pageNumber="476" refString="Deleve, J. (1974) Helichus granulosus (Coleoptera, Dryopidae) une nouvelle espece du Vietnam. Annales Historico-Naturales Musei Nationalis Hungarici, 66, 171 - 172." type="journal article" year="1974">Delève 1974</bibRefCitation>
|
||
: 171
|
||
</treatmentCitation>
|
||
</taxonomicName>
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BDFA562FFFC8E03FF097879FAF49E41" blockId="1.[151,1437,1284,2029]" pageId="1" pageNumber="476">
|
||
<emphasis id="B9147970FFFC8E03FF097879FE909D61" bold="true" box="[151,282,1284,1309]" pageId="1" pageNumber="476">Taxonomy.</emphasis>
|
||
<bibRefCitation id="EFF1D893FFFC8E03FEBC7879FE4F9D60" author="Deleve" box="[290,453,1284,1309]" pageId="1" pageNumber="476" refString="Deleve, J. (1974) Helichus granulosus (Coleoptera, Dryopidae) une nouvelle espece du Vietnam. Annales Historico-Naturales Musei Nationalis Hungarici, 66, 171 - 172." type="journal article" year="1974">Delève (1974)</bibRefCitation>
|
||
described this species in the genus
|
||
<taxonomicName id="4C60DEE1FFFC8E03FCFF7879FC4F9D60" ID-CoL="635L2" box="[865,965,1284,1308]" class="Insecta" family="Dryopidae" genus="Helichus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="476" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9147970FFFC8E03FCFF7879FC4F9D60" box="[865,965,1284,1308]" italics="true" pageId="1" pageNumber="476">Helichus</emphasis>
|
||
</taxonomicName>
|
||
, providing the following comment on its placement into the subgenus
|
||
<emphasis id="B9147970FFFC8E03FE7F7855FD0A9D3C" box="[481,640,1320,1344]" italics="true" pageId="1" pageNumber="476">
|
||
<taxonomicName id="4C60DEE1FFFC8E03FE7F7855FDF09D3C" ID-CoL="63JVH" box="[481,634,1320,1344]" class="Insecta" family="Dryopidae" genus="Parahelichus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="476" phylum="Arthropoda" rank="genus">Parahelichus</taxonomicName>
|
||
:
|
||
</emphasis>
|
||
“Par la forme du pronotum cette espèce pourrait rentrer dans le sousgenre
|
||
<emphasis id="B9147970FFFC8E03FF437831FEED9D18" box="[221,359,1356,1380]" italics="true" pageId="1" pageNumber="476">Parahelichu</emphasis>
|
||
s
|
||
<bibRefCitation id="EFF1D893FFFC8E03FEE57831FDAB9D18" author="Bollow" box="[379,545,1356,1381]" pageId="1" pageNumber="476" refString="Bollow, H. (1940) Monographie der palaearktischen Dryopidae, mit Berucksichtigung der eventuell transgredierenden Arten. (Coleoptera). Mitteilungen der Munchener Entomologischen Gesellschaft, 30 (1), 24 - 71." type="journal article" year="1940">Bollow (1940)</bibRefCitation>
|
||
mais le corps est dépourvu de la pubescence serrée masquant la coloration du tegument, caractère ayant servi à fonder le sous-genre.” The main difference between the subgenera
|
||
<taxonomicName id="4C60DEE1FFFC8E03FA96780DFA169DF4" ID-CoL="6VCJ" box="[1288,1436,1392,1416]" class="Insecta" family="Dryopidae" genus="Praehelichus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="476" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9147970FFFC8E03FA96780DFA169DF4" box="[1288,1436,1392,1416]" italics="true" pageId="1" pageNumber="476">Praehelichus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C60DEE1FFFC8E03FF5778E9FED49DD0" ID-CoL="63JVH" box="[201,350,1428,1452]" class="Insecta" family="Dryopidae" genus="Parahelichus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="476" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9147970FFFC8E03FF5778E9FED49DD0" box="[201,350,1428,1452]" italics="true" pageId="1" pageNumber="476">Parahelichus</emphasis>
|
||
</taxonomicName>
|
||
is in the form of lateral pronotal carina that is regularly bent in
|
||
<taxonomicName id="4C60DEE1FFFC8E03FBD778E9FB579DD0" ID-CoL="6VCJ" box="[1097,1245,1428,1452]" class="Insecta" family="Dryopidae" genus="Praehelichus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="476" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9147970FFFC8E03FBD778E9FB579DD0" box="[1097,1245,1428,1452]" italics="true" pageId="1" pageNumber="476">Praehelichus</emphasis>
|
||
</taxonomicName>
|
||
, while unevenly bent or lobate in
|
||
<taxonomicName id="4C60DEE1FFFC8E03FEC278C5FE7B9DAC" ID-CoL="63JVH" box="[348,497,1464,1488]" class="Insecta" family="Dryopidae" genus="Parahelichus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="476" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9147970FFFC8E03FEC278C5FE7B9DAC" box="[348,497,1464,1488]" italics="true" pageId="1" pageNumber="476">Parahelichus</emphasis>
|
||
</taxonomicName>
|
||
. Beside this key feature
|
||
<bibRefCitation id="EFF1D893FFFC8E03FC8E78CAFC3C9DAC" author="Bollow" box="[784,950,1463,1488]" pageId="1" pageNumber="476" refString="Bollow, H. (1940) Monographie der palaearktischen Dryopidae, mit Berucksichtigung der eventuell transgredierenden Arten. (Coleoptera). Mitteilungen der Munchener Entomologischen Gesellschaft, 30 (1), 24 - 71." type="journal article" year="1940">Bollow (1940)</bibRefCitation>
|
||
mentioned the 10-segmented antennae for
|
||
<taxonomicName id="4C60DEE1FFFC8E03FF0978A1FEA69D88" ID-CoL="63JVH" box="[151,300,1500,1524]" class="Insecta" family="Dryopidae" genus="Parahelichus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="476" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9147970FFFC8E03FF0978A1FEA69D88" box="[151,300,1500,1524]" italics="true" pageId="1" pageNumber="476">Parahelichus</emphasis>
|
||
</taxonomicName>
|
||
and 11-segmented for
|
||
<taxonomicName id="4C60DEE1FFFC8E03FDAD78A1FD4D9D88" ID-CoL="6VCJ" box="[563,711,1500,1524]" class="Insecta" family="Dryopidae" genus="Praehelichus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="476" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9147970FFFC8E03FDAD78A1FD4D9D88" box="[563,711,1500,1524]" italics="true" pageId="1" pageNumber="476">Praehelichus</emphasis>
|
||
</taxonomicName>
|
||
. Although the classification of these two supraspecific taxa has not yet been sufficiently resolved, we consider
|
||
<taxonomicName id="4C60DEE1FFFC8E03FD2B7B7CFCDA9E64" box="[693,848,1536,1560]" class="Insecta" family="Dryopidae" genus="Helichus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="476" phylum="Arthropoda" rank="species" species="granulosus">
|
||
<emphasis id="B9147970FFFC8E03FD2B7B7CFCDA9E64" box="[693,848,1536,1560]" italics="true" pageId="1" pageNumber="476">P. granulosus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C60DEE1FFFC8E03FC147B7CFBF99E64" box="[906,1139,1536,1560]" class="Insecta" family="Dryopidae" genus="Parahelichus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="476" phylum="Arthropoda" rank="species" species="pseudogranulosus">
|
||
<emphasis id="B9147970FFFC8E03FC147B7CFBF99E64" box="[906,1139,1536,1560]" italics="true" pageId="1" pageNumber="476">P. pseudogranulosus</emphasis>
|
||
</taxonomicName>
|
||
to form a distinct species group within the genus based on the shape of pronotal carina and tightly arranged small granules on the surface.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37AF6E9FFFC8E00FF597B3AFF6599F4" lastPageId="2" lastPageNumber="477" pageId="1" pageNumber="476" type="materials_examined">
|
||
<paragraph id="8BDFA562FFFC8E03FF597B3AFF559EF8" blockId="1.[151,1437,1284,2029]" pageId="1" pageNumber="476">
|
||
<emphasis id="B9147970FFFC8E03FF597B3AFEE39E1C" bold="true" box="[199,361,1607,1632]" pageId="1" pageNumber="476">
|
||
<typeStatus id="54DB1BC0FFFC8E03FF597B3AFE889E1C" box="[199,258,1607,1632]" pageId="1" pageNumber="476">Type</typeStatus>
|
||
locality.
|
||
</emphasis>
|
||
Hanoi,
|
||
<collectingCountry id="F377E5F2FFFC8E03FE567B3AFDA09E1C" box="[456,554,1607,1632]" name="Vietnam" pageId="1" pageNumber="476">Vietnam</collectingCountry>
|
||
, single
|
||
<typeStatus id="54DB1BC0FFFC8E03FD187B3AFD629E1C" box="[646,744,1607,1632]" pageId="1" pageNumber="476" type="holotype">holotype</typeStatus>
|
||
specimen was collected by light in hotel Kim lien (
|
||
<bibRefCitation id="EFF1D893FFFC8E03FAD57B3AFF599EF8" author="Deleve" pageId="1" pageNumber="476" refString="Deleve, J. (1974) Helichus granulosus (Coleoptera, Dryopidae) une nouvelle espece du Vietnam. Annales Historico-Naturales Musei Nationalis Hungarici, 66, 171 - 172." type="journal article" year="1974">Delève 1974</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BDFA562FFFC8E03FF597BF2FA869F91" blockId="1.[151,1437,1284,2029]" pageId="1" pageNumber="476">
|
||
<emphasis id="B9147970FFFC8E03FF597BF2FD5D9ED4" bold="true" box="[199,727,1679,1704]" pageId="1" pageNumber="476">
|
||
Material examined.
|
||
<typeStatus id="54DB1BC0FFFC8E03FE5A7BF2FDBB9ED4" box="[452,561,1679,1704]" pageId="1" pageNumber="476" type="holotype">Holotype</typeStatus>
|
||
♂ (
|
||
<collectionCode id="ED713DA7FFFC8E03FDF77BF2FD409ED4" box="[617,714,1679,1704]" pageId="1" pageNumber="476">MTMB</collectionCode>
|
||
):
|
||
</emphasis>
|
||
“Hanoi, Hotel Kim lien,
|
||
<date id="FFDE83A2FFFC8E03FB8A7BEDFB0F9ED4" box="[1044,1157,1679,1704]" pageId="1" pageNumber="476" value="1931-05-28">28. 31-V-</date>
|
||
1966 Gy. Topál, n-588, collected by lamp.”
|
||
<emphasis id="B9147970FFFC8E03FEE77BC9FDE69EB1" bold="true" box="[377,620,1716,1741]" pageId="1" pageNumber="476">Additional material.</emphasis>
|
||
442 ex. (
|
||
<collectionCode id="ED713DA7FFFC8E03FD4B7BC9FC989EB0" box="[725,786,1716,1740]" pageId="1" pageNumber="476">CKB</collectionCode>
|
||
,
|
||
<collectionCode id="ED713DA7FFFC8E03FC807BC8FCFC9EB0" box="[798,886,1717,1740]" pageId="1" pageNumber="476">MTMB</collectionCode>
|
||
,
|
||
<collectionCode id="ED713DA7FFFC8E03FC1C7BC8FC409EB0" box="[898,970,1717,1740]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="1" pageNumber="476">NMW</collectionCode>
|
||
,
|
||
<collectionCode id="ED713DA7FFFC8E03FC487BC9FBA79EB0" box="[982,1069,1716,1740]" pageId="1" pageNumber="476">NHGH</collectionCode>
|
||
): 16 ♂♂, 12 ♀♀: “
|
||
<collectingCountry id="F377E5F2FFFC8E03FA9F7BC9FAEF9EB1" box="[1281,1381,1716,1741]" name="Vietnam" pageId="1" pageNumber="476">Vietnam</collectingCountry>
|
||
, Yen Bai,
|
||
<date id="FFDE83A2FFFC8E03FF527BA5FEC29E8C" box="[204,328,1752,1776]" pageId="1" pageNumber="476" value="1990-05-15">15.V. 1990</date>
|
||
, J. Horák leg.”; 10 ♂♂, 10 ♀♀: “
|
||
<collectingCountry id="F377E5F2FFFC8E03FD217BA5FCA99E8C" box="[703,803,1752,1776]" name="Vietnam" pageId="1" pageNumber="476">Vietnam</collectingCountry>
|
||
, Yen Bai,
|
||
<date id="FFDE83A2FFFC8E03FC087BA5FBB09E8C" box="[918,1082,1752,1776]" pageId="1" pageNumber="476" value="1990-05-06" valueMax="1990-05-25" valueMin="1990-05-06">6.–25. 5. 1990</date>
|
||
, O. Šauša lgt.”;
|
||
<specimenCount id="9D666EEBFFFC8E03FB717BA5FA919E8C" box="[1263,1307,1751,1776]" pageId="1" pageNumber="476" type="generic">
|
||
1
|
||
<emphasis id="B9147970FFFC8E03FA9A7BAAFA919E8C" bold="true" box="[1284,1307,1751,1776]" pageId="1" pageNumber="476">♂</emphasis>
|
||
</specimenCount>
|
||
: “
|
||
<collectingCountry id="F377E5F2FFFC8E03FAAA7BAAFA129E8C" box="[1332,1432,1751,1776]" name="Vietnam" pageId="1" pageNumber="476">Vietnam</collectingCountry>
|
||
, Vinh phu prov., Tam dao, 27.5.–
|
||
<date id="FFDE83A2FFFC8E03FD977B81FDFF9F69" box="[521,629,1788,1813]" pageId="1" pageNumber="476" value="1986-06-02">2.6. 1986</date>
|
||
, Jan Horák lgt.”; 3 ♀♀: “
|
||
<collectingCountry id="F377E5F2FFFC8E03FC077B81FC779F69" box="[921,1021,1788,1813]" name="Vietnam" pageId="1" pageNumber="476">Vietnam</collectingCountry>
|
||
, Ha son bin prov., Hoa binh, 4.–
|
||
<date id="FFDE83A2FFFC8E03FAEC7B81FF589F44" pageId="1" pageNumber="476" value="1986-06-07">7.6. 1986</date>
|
||
, Jan Horák lgt.”; 91 ♂♂, 92 ♀♀: “N -
|
||
<collectingCountry id="F377E5F2FFFC8E03FD187A62FD629F44" box="[646,744,1823,1848]" name="Vietnam" pageId="1" pageNumber="476">Vietnam</collectingCountry>
|
||
, Sa Pa (Lao Cai), 25.5.–
|
||
<date id="FFDE83A2FFFC8E03FC657A62FBF99F44" box="[1019,1139,1823,1848]" pageId="1" pageNumber="476" value="1991-10-06">10.6. 1991</date>
|
||
,
|
||
<geoCoordinate id="EE54C3A5FFFC8E03FBE17A5DFB7A9F44" box="[1151,1264,1824,1848]" direction="north" orientation="latitude" pageId="1" pageNumber="476" precision="925" value="22.333334">22° 20`N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE54C3A5FFFC8E03FB627A5DFAF29F44" box="[1276,1400,1823,1848]" direction="east" orientation="longitude" pageId="1" pageNumber="476" precision="925" value="103.833336">103° 50`E</geoCoordinate>
|
||
, E. Jendek leg.”; 25 ♂♂, 17 ♀♀: “
|
||
<collectingCountry id="F377E5F2FFFC8E03FE6C7A39FDA19F20" box="[498,555,1860,1884]" name="Laos" pageId="1" pageNumber="476">Laos</collectingCountry>
|
||
north, Louang Namtha,
|
||
<quantity id="4C980887FFFC8E03FCA17A39FCF59F20" box="[831,895,1860,1884]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="476" unit="km" value="20.0">20km</quantity>
|
||
NW, N 21° 0 9.2, E 101° 18.7, alt. 900 ±
|
||
<quantity id="4C980887FFFC8E03FAD17A39FA1D9F27" box="[1359,1431,1860,1884]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="476" unit="m" value="100.0">100 m</quantity>
|
||
,
|
||
<date id="FFDE83A2FFFC8E03FF097A15FEB69FFC" box="[151,316,1896,1920]" pageId="1" pageNumber="476" value="1997-05-05" valueMax="1997-05-11" valueMin="1997-05-05">5.–11. V. 1997</date>
|
||
, E. Jendek & O. Šauša leg.”; 70 ♂♂, 86 ♀♀: “
|
||
<collectingCountry id="F377E5F2FFFC8E03FCD87A15FCF59FFC" box="[838,895,1896,1920]" name="Laos" pageId="1" pageNumber="476">Laos</collectingCountry>
|
||
- N, Louangphrabang pr.
|
||
<geoCoordinate id="EE54C3A5FFFC8E03FB3E7A15FA9B9F03" box="[1184,1297,1896,1920]" direction="north" orientation="latitude" pageId="1" pageNumber="476" precision="925" value="19.883333">19° 53`N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE54C3A5FFFC8E03FA827A15FA1D9F03" box="[1308,1431,1896,1920]" direction="east" orientation="longitude" pageId="1" pageNumber="476" precision="925" value="102.15">102° 09`E</geoCoordinate>
|
||
, Khan riv.,
|
||
<quantity id="4C980887FFFC8E03FE8F7AF1FED79FD8" box="[273,349,1932,1956]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="476" unit="m" value="300.0">300 m</quantity>
|
||
,
|
||
<date id="FFDE83A2FFFC8E03FEF47AF1FE789FD8" box="[362,498,1932,1956]" pageId="1" pageNumber="476" value="1999-04-21">21.IV. 1999</date>
|
||
, Vít Kubáň leg.”;
|
||
<specimenCount id="9D666EEBFFFC8E03FD597AF1FD7F9FD9" box="[711,757,1932,1957]" pageId="1" pageNumber="476" type="generic">
|
||
1
|
||
<emphasis id="B9147970FFFC8E03FD407AF1FD7F9FD9" bold="true" box="[734,757,1932,1957]" pageId="1" pageNumber="476">♂</emphasis>
|
||
</specimenCount>
|
||
: “
|
||
<collectingCountry id="F377E5F2FFFC8E03FC8F7AF1FCC69FD8" box="[785,844,1932,1956]" name="Laos" pageId="1" pageNumber="476">Laos</collectingCountry>
|
||
, Louangphrabang pr. 20° 33-4`N,
|
||
<geoCoordinate id="EE54C3A5FFFC8E03FB7F7AF1FAEB9FD8" box="[1249,1377,1932,1956]" direction="east" orientation="longitude" pageId="1" pageNumber="476" precision="925" value="102.23333">102° 14`E</geoCoordinate>
|
||
, Ban Song Cha (
|
||
<quantity id="4C980887FFFC8E03FE8B7ACDFEC49FB4" box="[277,334,1968,1992]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="476" unit="km" value="5.0">5 km</quantity>
|
||
W),
|
||
<quantity id="4C980887FFFC8E03FE197ACDFE549FB4" box="[391,478,1968,1992]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="1" pageNumber="476" unit="m" value="1200.0">1200 m</quantity>
|
||
,
|
||
<date id="FFDE83A2FFFC8E03FE757ACDFD189FB4" box="[491,658,1968,1992]" pageId="1" pageNumber="476" value="1999-05-01" valueMax="1999-05-16" valueMin="1999-05-01">1.–16. V. 1999</date>
|
||
, Vít Kubáň leg.”; 3 ♂♂, 5 ♀♀: “
|
||
<collectingCountry id="F377E5F2FFFC8E03FB9B7ACDFBCA9FB4" box="[1029,1088,1968,1992]" name="Laos" pageId="1" pageNumber="476">Laos</collectingCountry>
|
||
, centr., Khammouan prov.,
|
||
<geoCoordinate id="EE54C3A5FFFC8E03FAEB7ACDFF509F90" direction="north" orientation="latitude" pageId="1" pageNumber="476" precision="925" value="18.116667">18° 07`N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE54C3A5FFFC8E03FF7B7AA9FEE29F90" box="[229,360,2004,2028]" direction="east" orientation="longitude" pageId="1" pageNumber="476" precision="925" value="104.48333">104° 29`E</geoCoordinate>
|
||
, alt.
|
||
<quantity id="4C980887FFFC8E03FE007AA9FE6D9F90" box="[414,487,2004,2029]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="1" pageNumber="476" unit="m" value="250.0">250 m</quantity>
|
||
, Ban Khoun Ngeun env.,
|
||
<date id="FFDE83A2FFFC8E03FC967AA9FC439F90" box="[776,969,2004,2028]" pageId="1" pageNumber="476" value="2001-06-10" valueMax="2001-06-13" valueMin="2001-06-10">10.–13. VI. 2001</date>
|
||
, E. Jendek & O. Šauša leg.”.
|
||
</paragraph>
|
||
<paragraph id="8BDFA562FFFF8E00FF597DEAFF6599F4" blockId="2.[151,1437,151,536]" pageId="2" pageNumber="477">
|
||
<emphasis id="B9147970FFFF8E00FF597DEAFE4798CC" bold="true" box="[199,461,151,176]" pageId="2" pageNumber="477">Differential diagnosis.</emphasis>
|
||
<taxonomicName id="4C60DEE1FFFF8E00FE4D7DE4FDE198CC" box="[467,619,152,176]" class="Insecta" family="Dryopidae" genus="Helichus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="477" phylum="Arthropoda" rank="species" species="granulosus">
|
||
<emphasis id="B9147970FFFF8E00FE4D7DE4FDE198CC" box="[467,619,152,176]" italics="true" pageId="2" pageNumber="477">P. granulosus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C60DEE1FFFF8E00FD3E7DE4FC0D98CC" box="[672,903,152,176]" class="Insecta" family="Dryopidae" genus="Parahelichus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="477" phylum="Arthropoda" rank="species" species="pseudogranulosus" status="sp. nov.">
|
||
<emphasis id="B9147970FFFF8E00FD3E7DE4FC0D98CC" box="[672,903,152,176]" italics="true" pageId="2" pageNumber="477">P. pseudogranulosus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A227C40BFFFF8E00FC137DE5FC6998D3" box="[909,995,152,175]" pageId="2" pageNumber="477" rank="species">sp. nov.</taxonomicNameLabel>
|
||
are large species distinguished from all described Asian
|
||
<taxonomicName id="4C60DEE1FFFF8E00FECA7DC1FE6298A8" box="[340,488,188,212]" class="Insecta" family="Dryopidae" genus="Praehelichus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="477" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9147970FFFF8E00FECA7DC1FE6298A8" box="[340,488,188,212]" italics="true" pageId="2" pageNumber="477">Praehelichus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C60DEE1FFFF8E00FE667DC1FD0798A8" box="[504,653,188,212]" class="Insecta" family="Dryopidae" genus="Parahelichus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="477" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9147970FFFF8E00FE667DC1FD0798A8" box="[504,653,188,212]" italics="true" pageId="2" pageNumber="477">Parahelichus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C60DEE1FFFF8E00FD597DC1FCA198A8" box="[711,811,188,212]" class="Insecta" family="Dryopidae" genus="Helichus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="477" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9147970FFFF8E00FD597DC1FCA198A8" box="[711,811,188,212]" italics="true" pageId="2" pageNumber="477">Helichus</emphasis>
|
||
</taxonomicName>
|
||
species by the very tightly arranged small granules on the dorsal and ventral surface (causing its reticulate appearance at smaller magnifications). Further, both species differ from other relatives by the lack of a conspicuous area on the dorsal surface covered by plastron setae. Thus, plastron setae are confined only to a small area on ventrite 1 posteriad of metacoxae in
|
||
<taxonomicName id="4C60DEE1FFFF8E00FB327C54FAC3993C" box="[1196,1353,296,320]" class="Insecta" family="Dryopidae" genus="Helichus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="477" phylum="Arthropoda" rank="species" species="granulosus">
|
||
<emphasis id="B9147970FFFF8E00FB327C54FAC3993C" box="[1196,1353,296,320]" italics="true" pageId="2" pageNumber="477">P. granulosus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C60DEE1FFFF8E00FA197C54FEE99918" class="Insecta" family="Dryopidae" genus="Parahelichus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="477" phylum="Arthropoda" rank="species" species="pseudogranulosus">
|
||
<emphasis id="B9147970FFFF8E00FA197C54FEE99918" italics="true" pageId="2" pageNumber="477">P. pseudogranulosus</emphasis>
|
||
</taxonomicName>
|
||
, while the remaining congeners bear more or less densely arranged plastron setae on the dorsal surface.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37AF6E9FFFF8E07FF597CE9FEB69AB1" lastPageId="5" lastPageNumber="480" pageId="2" pageNumber="477" type="description">
|
||
<paragraph id="8BDFA562FFFF8E00FF597CE9FAA49A64" blockId="2.[151,1437,151,536]" pageId="2" pageNumber="477">
|
||
<emphasis id="B9147970FFFF8E00FF597CE9FEFE99D1" bold="true" box="[199,372,404,429]" pageId="2" pageNumber="477">Redescription.</emphasis>
|
||
Habitus of male as shown in
|
||
<figureCitation id="135BB9E7FFFF8E00FD6E7CE9FCB399D0" box="[752,825,404,429]" captionStart="FIGURE 1" captionStartId="2.[151,250,1836,1858]" captionText="FIGURE 1. Habitus of Parahelichus granulosus, male, TL = 4.5 mm." pageId="2" pageNumber="477">Fig. 1</figureCitation>
|
||
. Body elongated, heavily sclerotized and compact, moderately convex dorsally. Length (BL) in ♂♂:
|
||
<quantity id="4C980887FFFF8E00FD567CCAFCE699B3" box="[712,876,439,464]" metricMagnitude="-3" metricUnit="m" metricValue="4.984999999999999" metricValueMax="6.39" metricValueMin="3.58" pageId="2" pageNumber="477" unit="mm" value="4.984999999999999" valueMax="6.39" valueMin="3.58">3.58–6.39 mm</quantity>
|
||
(5.36 ±
|
||
<quantity id="4C980887FFFF8E00FC527CC5FBBF99AC" box="[972,1077,440,464]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" pageId="2" pageNumber="477" unit="mm" value="0.48">0.48 mm</quantity>
|
||
), in ♀♀:
|
||
<quantity id="4C980887FFFF8E00FB3E7CCAFACE99B3" box="[1184,1348,439,464]" metricMagnitude="-3" metricUnit="m" metricValue="5.46" metricValueMax="6.95" metricValueMin="3.97" pageId="2" pageNumber="477" unit="mm" value="5.46" valueMax="6.95" valueMin="3.97">3.97–6.95 mm</quantity>
|
||
(5.84 ± 0.46); maximum width (EW) in ♂♂:
|
||
<quantity id="4C980887FFFF8E00FDDD7CA1FD6D998F" box="[579,743,476,500]" metricMagnitude="-3" metricUnit="m" metricValue="2.065" metricValueMax="2.63" metricValueMin="1.5" pageId="2" pageNumber="477" unit="mm" value="2.065" valueMax="2.63" valueMin="1.5">1.50–2.63 mm</quantity>
|
||
(2.21 ±
|
||
<quantity id="4C980887FFFF8E00FCD47CA1FC399988" box="[842,947,476,500]" metricMagnitude="-4" metricUnit="m" metricValue="1.9" pageId="2" pageNumber="477" unit="mm" value="0.19">0.19 mm</quantity>
|
||
), in ♀♀:
|
||
<quantity id="4C980887FFFF8E00FBBE7CA1FB4E998F" box="[1056,1220,476,500]" metricMagnitude="-3" metricUnit="m" metricValue="2.28" metricValueMax="2.95" metricValueMin="1.61" pageId="2" pageNumber="477" unit="mm" value="2.28" valueMax="2.95" valueMin="1.61">1.61–2.95 mm</quantity>
|
||
(2.46 ±
|
||
<quantity id="4C980887FFFF8E00FAB97CA1FA1A9988" box="[1319,1424,476,500]" metricMagnitude="-4" metricUnit="m" metricValue="1.9" pageId="2" pageNumber="477" unit="mm" value="0.19">0.19 mm</quantity>
|
||
). Colour nearly black, antennal clubs, mouthparts and tarsi reddish-brown; immature specimens brownish.
|
||
</paragraph>
|
||
<caption id="DF1FF5EAFFFF8E00FF097A51FCEC9F3E" box="[151,870,1836,1858]" pageId="2" pageNumber="477">
|
||
<paragraph id="8BDFA562FFFF8E00FF097A51FCEC9F3E" blockId="2.[151,870,1836,1858]" box="[151,870,1836,1858]" pageId="2" pageNumber="477">
|
||
<emphasis id="B9147970FFFF8E00FF097A51FE999F3D" bold="true" box="[151,275,1836,1858]" pageId="2" pageNumber="477">FIGURE 1.</emphasis>
|
||
Habitus of
|
||
<taxonomicName id="4C60DEE1FFFF8E00FE177A51FD0D9F3E" box="[393,647,1836,1858]" class="Insecta" family="Dryopidae" genus="Parahelichus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="477" phylum="Arthropoda" rank="species" species="granulosus">
|
||
<emphasis id="B9147970FFFF8E00FE177A51FD0D9F3E" box="[393,647,1836,1858]" italics="true" pageId="2" pageNumber="477">Parahelichus granulosus</emphasis>
|
||
</taxonomicName>
|
||
, male, TL = 4.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BDFA562FFFF8E00FF597A14FCD49F92" blockId="2.[151,1436,1897,2030]" pageId="2" pageNumber="477">Surface of pronotum, elytra and venter opaque, covered with tightly arranged round small granules and larger granules. Small granules about one third of a diameter of a facet. Larger granules shiny, nearly rounded, subequal with a diameter of a facet, associated with moderately long yellowish setae and more or less regularly intermixed within small granules. Elytra lacking distinct striae or intervals.</paragraph>
|
||
<paragraph id="8BDFA562FFFE8E01FF597DEAFC4699F4" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="478">
|
||
Head wider than long; surface shiny with sparse flat granules; granules moderately smaller than a facet diameter, with yellowish recumbent setae, more densely arranged on anterolateral portion than mesally. Frons with narrow, small shiny longitudinal area; eyes large and protuberant, strongly facetted; interfacetal setae fine, short, scattered and present on central portion of eye. ID in ♂♂:
|
||
<quantity id="4C980887FFFE8E01FCB87C79FC439960" box="[806,969,260,285]" metricMagnitude="-4" metricUnit="m" metricValue="5.1" metricValueMax="6.8" metricValueMin="3.4" pageId="3" pageNumber="478" unit="mm" value="0.51" valueMax="0.68" valueMin="0.34">0.34–0.68 mm</quantity>
|
||
(0.58 ±
|
||
<quantity id="4C980887FFFE8E01FBB67C79FB1A9960" box="[1064,1168,260,285]" metricMagnitude="-5" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="478" unit="mm" value="0.06">0.06 mm</quantity>
|
||
), in ♀♀:
|
||
<quantity id="4C980887FFFE8E01FB647C79FA179960" box="[1274,1437,260,284]" metricMagnitude="-4" metricUnit="m" metricValue="6.2" metricValueMax="8.2" metricValueMin="4.2" pageId="3" pageNumber="478" unit="mm" value="0.62" valueMax="0.82" valueMin="0.42">0.42–0.82 mm</quantity>
|
||
(0.66 ±
|
||
<quantity id="4C980887FFFE8E01FF737C55FEDE993C" box="[237,340,296,320]" metricMagnitude="-5" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="478" unit="mm" value="0.06">0.06 mm</quantity>
|
||
); HW in ♂♂:
|
||
<quantity id="4C980887FFFE8E01FE6F7C55FD199943" box="[497,659,296,320]" metricMagnitude="-3" metricUnit="m" metricValue="1.01" metricValueMax="1.26" metricValueMin="0.76" pageId="3" pageNumber="478" unit="mm" value="1.01" valueMax="1.26" valueMin="0.76">0.76–1.26 mm</quantity>
|
||
(1.10 ±
|
||
<quantity id="4C980887FFFE8E01FD6E7C55FCDD993C" box="[752,855,296,320]" metricMagnitude="-5" metricUnit="m" metricValue="9.0" pageId="3" pageNumber="478" unit="mm" value="0.09">0.09 mm</quantity>
|
||
), in ♀♀:
|
||
<quantity id="4C980887FFFE8E01FC237C55FBEA9943" box="[957,1120,296,320]" metricMagnitude="-3" metricUnit="m" metricValue="1.145" metricValueMax="1.42" metricValueMin="0.87" pageId="3" pageNumber="478" unit="mm" value="1.145" valueMax="1.42" valueMin="0.87">0.87–1.42 mm</quantity>
|
||
(1.21 ±
|
||
<quantity id="4C980887FFFE8E01FB227C55FAAE993C" box="[1212,1316,296,320]" metricMagnitude="-5" metricUnit="m" metricValue="9.0" pageId="3" pageNumber="478" unit="mm" value="0.09">0.09 mm</quantity>
|
||
). Antennal insertion exposed from above, antennae widely separated; subantennal groove shallow, reaching about middle of eye; antenna pectinate, short, club with eight antennomeres (
|
||
<figureCitation id="135BB9E7FFFE8E01FCA37C12FC3599F4" box="[829,959,367,392]" captionStart="FIGURES 17 – 20" captionStartId="9.[151,264,1103,1126]" captionTargetBox="[238,1350,196,1075]" captionTargetId="figure@9.[231,1355,193,1082]" captionTargetPageId="9" captionText="FIGURES 17 – 20. Parahelichus granulosus; (17 ‒ 18) female antennal club: (17) ventral view, (18) dorsal view, scale 0.1 mm; (19) abdomen of a male, ventral view; (20) abdomen of a female, ventral view; scale 1 mm." httpUri="https://zenodo.org/record/231191/files/figure.png" pageId="3" pageNumber="478">Figs 17, 18</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BDFA562FFFE8E01FF597CE9FD429EC1" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="478">
|
||
Thorax. Pronotum wider than long; disc convex dorsally, without gibbosities, carinae or furrows; granules more distinct than those on head, separated usually by distance of one granule diameter or less. Sides arcuate; lateral pronotal carina distinct, moderately raised, not crenulate; posterior angles almost rectangular, moderately acute; anterior angles acute, protruding; posterior margin trisinuate, smooth; prescutellar foveae absent. MPL in ♂♂:
|
||
<quantity id="4C980887FFFE8E01FF4E7F59FEF89A47" box="[208,370,548,572]" metricMagnitude="-3" metricUnit="m" metricValue="1.22" metricValueMax="1.55" metricValueMin="0.89" pageId="3" pageNumber="478" unit="mm" value="1.22" valueMax="1.55" valueMin="0.89">0.89–1.55 mm</quantity>
|
||
(1.22 ±
|
||
<quantity id="4C980887FFFE8E01FE4F7F59FDB39A40" box="[465,569,548,572]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="478" unit="mm" value="0.1">0.10 mm</quantity>
|
||
), in ♀♀:
|
||
<quantity id="4C980887FFFE8E01FD3F7F59FCCE9A47" box="[673,836,548,572]" metricMagnitude="-3" metricUnit="m" metricValue="1.235" metricValueMax="1.55" metricValueMin="0.92" pageId="3" pageNumber="478" unit="mm" value="1.235" valueMax="1.55" valueMin="0.92">0.92–1.55 mm</quantity>
|
||
(1.34 ±
|
||
<quantity id="4C980887FFFE8E01FC3C7F59FB809A40" box="[930,1034,548,572]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="478" unit="mm" value="0.1">0.10 mm</quantity>
|
||
); APW in ♂♂:
|
||
<quantity id="4C980887FFFE8E01FB267F59FAD19A47" box="[1208,1371,548,572]" metricMagnitude="-3" metricUnit="m" metricValue="1.145" metricValueMax="1.42" metricValueMin="0.87" pageId="3" pageNumber="478" unit="mm" value="1.145" valueMax="1.42" valueMin="0.87">0.87–1.42 mm</quantity>
|
||
(1.22 ±
|
||
<quantity id="4C980887FFFE8E01FF2E7F35FE919A1C" box="[176,283,584,608]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="478" unit="mm" value="0.1">0.10 mm</quantity>
|
||
), in ♀♀:
|
||
<quantity id="4C980887FFFE8E01FE137F35FDB99A23" box="[397,563,584,608]" metricMagnitude="-3" metricUnit="m" metricValue="1.22" metricValueMax="1.55" metricValueMin="0.89" pageId="3" pageNumber="478" unit="mm" value="1.22" valueMax="1.55" valueMin="0.89">0.89–1.55 mm</quantity>
|
||
(1.33 ±
|
||
<quantity id="4C980887FFFE8E01FD047F35FC8F9A1C" box="[666,773,584,608]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="478" unit="mm" value="0.1">0.10 mm</quantity>
|
||
); PPW in ♂♂: 1.16–2.00 mm (1.69 ±
|
||
<quantity id="4C980887FFFE8E01FB577F3AFABE9A1C" box="[1225,1332,583,608]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="478" unit="mm" value="0.15">0.15 mm</quantity>
|
||
), in ♀♀:
|
||
<quantity id="4C980887FFFE8E01FF097F11FEB79AF8" box="[151,317,620,644]" metricMagnitude="-3" metricUnit="m" metricValue="1.765" metricValueMax="2.24" metricValueMin="1.29" pageId="3" pageNumber="478" unit="mm" value="1.765" valueMax="2.24" valueMin="1.29">1.29–2.24 mm</quantity>
|
||
(1.84 ±
|
||
<quantity id="4C980887FFFE8E01FE3B7F11FD9B9AF8" box="[421,529,620,645]" metricMagnitude="-4" metricUnit="m" metricValue="1.6" pageId="3" pageNumber="478" unit="mm" value="0.16">0.16 mm</quantity>
|
||
). Hypomeron broadest near middle, distinctly narrowed anteriorly and slightly posteriorly; separated from prosternum by a gap along anterior fourth; postcoxal process very short, triangular, not reach prosternal process. Prosternum in front of coxae approximately as long as prosternal process, anterolateral portion rounded. Setae in middle of prosternum and on prosternal process longer than those on dorsal surface; longest setae in disc of metaventrite. Prosternal process wide, strongly overlapping mesoventrite, fitting into deep cavity of mesoventrite, apex long and subacute, sides raised. Mesoventrite very short, with conspicuous ventral projection on each side of mesoventral cavity; exposed portion of mesanepisternum large, about as long as width of hypomeron; mesepimeron fourth as long as metanepisternum, both divided by fine line and both reaching mesocoxal cavity. Mesocoxal cavities subcircular, transversally oriented; mesocoxae subglobular, with short projection near attachment of trochanter, less widely separated than procoxae; exposed portion of mesotrochantin small. Scutellum (scutellar shield) large, broadly rounded, in same level with elytra, surface granulated. Mesoventral and metaventral processes solidly fused. Metaventrite distinctly longer than prosternum in front of coxae, without grooves or depressions for reception of femur and tibia; discrimen long and fine; transverse suture fine but distinct; anterior edge of metaventrite without transverse carina between mesocoxal cavities; metanepisternum wide and long. Metacoxal cavities large, transverse, oblique and not reaching elytra. Metacoxae separated by suture from metaventrite, posteriorly deeply excavate for reception of femora; moderately widely separated mesally, posterior projection distinct. Elytra convex dorsally, lateral margin not explanate, gradually extended from shoulders up to 0.75 of their length, elytral apices subacute; granulation on elytra similar to that on pronotum but large granules less dense and nearly regularly spaced on disc, denser laterally; epipleura widest basally, inflected at level of metacoxa and near apex, partly concealed by lateral margin of ventrites. EW in ♂♂:
|
||
<quantity id="4C980887FFFE8E01FF097841FEB09D28" box="[151,314,1340,1365]" metricMagnitude="-3" metricUnit="m" metricValue="2.065" metricValueMax="2.63" metricValueMin="1.5" pageId="3" pageNumber="478" unit="mm" value="2.065" valueMax="2.63" valueMin="1.5">1.50–2.63 mm</quantity>
|
||
(2.21 ±
|
||
<quantity id="4C980887FFFE8E01FE077841FD8B9D28" box="[409,513,1340,1364]" metricMagnitude="-4" metricUnit="m" metricValue="1.9" pageId="3" pageNumber="478" unit="mm" value="0.19">0.19 mm</quantity>
|
||
), in ♀♀:
|
||
<quantity id="4C980887FFFE8E01FDF57841FC849D2F" box="[619,782,1340,1364]" metricMagnitude="-3" metricUnit="m" metricValue="2.28" metricValueMax="2.95" metricValueMin="1.61" pageId="3" pageNumber="478" unit="mm" value="2.28" valueMax="2.95" valueMin="1.61">1.61–2.95 mm</quantity>
|
||
(2.46 ±
|
||
<quantity id="4C980887FFFE8E01FCF37841FC5F9D28" box="[877,981,1340,1364]" metricMagnitude="-4" metricUnit="m" metricValue="1.9" pageId="3" pageNumber="478" unit="mm" value="0.19">0.19 mm</quantity>
|
||
); EL in ♂♂:
|
||
<quantity id="4C980887FFFE8E01FBF77841FA869D2F" box="[1129,1292,1340,1364]" metricMagnitude="-3" metricUnit="m" metricValue="3.785" metricValueMax="4.89" metricValueMin="2.68" pageId="3" pageNumber="478" unit="mm" value="3.785" valueMax="4.89" valueMin="2.68">2.68–4.89 mm</quantity>
|
||
(4.14 ±
|
||
<quantity id="4C980887FFFE8E01FAF57841FF4D9D04" metricMagnitude="-4" metricUnit="m" metricValue="3.9" pageId="3" pageNumber="478" unit="mm" value="0.39">0.39 mm</quantity>
|
||
), in ♀♀:
|
||
<quantity id="4C980887FFFE8E01FEDC7822FE619D04" box="[322,491,1375,1400]" metricMagnitude="-3" metricUnit="m" metricValue="4.26" metricValueMax="5.47" metricValueMin="3.05" pageId="3" pageNumber="478" unit="mm" value="4.26" valueMax="5.47" valueMin="3.05">3.05–5.47 mm</quantity>
|
||
(4.50 ±
|
||
<quantity id="4C980887FFFE8E01FDC2781DFD419D04" box="[604,715,1376,1400]" metricMagnitude="-4" metricUnit="m" metricValue="3.8" pageId="3" pageNumber="478" unit="mm" value="0.38">0.38 mm</quantity>
|
||
). Legs slightly longer than elytra; trochanters large, femoral attachment oblique. Femora excavate for reception of tibiae, strongly widened, widest near middle, with scattered fine punctures; mesofemora without rows of conspicuous strong setae on posterior face. Tibiae slightly longer than femora; mesotibia as long as protibia, apically moderately widened; metatibia longest, apically strongly widened; each tibia weakly curved, narrow, with various specialized setae along anterior (protibia) or posterior face (meso- and metatibia) and with numerous short setae on remaining parts. Tarsi approximately as long as protibia and mesotibia, respectively, and slightly shorter than metatibia, with few short strong setae ventrally and longer setae mainly dorsally and laterally; terminal tarsomere on metatibia slightly shorter than combined lengths of tarsomeres 1–4, those on protibia and mesotibia approximately as long as combined lengths of tarsomeres 1–4. Claws robust and simple; empodium squamiform.
|
||
</paragraph>
|
||
<paragraph id="8BDFA562FFFE8E01FF597BBAFDB59FA1" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="478">
|
||
Abdomen with five ventrites, first two connate; ventrite 1 without lateral depression for reception of femora and tibiae; abdominal intercoxal process acute; ventrite 1 longest, ventrite 5 posteriorly slightly excised in males and rounded in females. Male segment VIII: sternite gently rounded posteriorly, not emarginated, distinctly shorter than ventrite 5, with short median process anteriorly (
|
||
<figureCitation id="135BB9E7FFFE8E01FC867A49FCE89F31" box="[792,866,1844,1869]" captionStart="FIGURES 3 – 13" captionStartId="7.[151,264,1950,1973]" captionTargetBox="[217,1367,201,1913]" captionTargetId="figure@7.[206,1381,193,1929]" captionTargetPageId="7" captionText="FIGURES 3 – 13. Parahelichus granulosus, male; (3 – 5, 8 – 13) aedeagus, penis and parameres in lateral, ventral and dorsal view; figures show differences in proportions between parameres and phallobase; (6) male sternite VIII, ventral view; (7) male sternite IX, ventral view. Scale 0.1 mm." httpUri="https://zenodo.org/record/231189/files/figure.png" pageId="3" pageNumber="478">Fig. 6</figureCitation>
|
||
); tergite more strongly sclerotized laterally than mesally, rounded posteriorly. Female segment VIII: sternite posteriorly sub-triangular, sclerotized laterally, with anterior median process approximately half as long as abdomen (
|
||
<figureCitation id="135BB9E7FFFE8E01FC0E7A01FC6B9FE9" box="[912,993,1916,1941]" captionStart="FIGURES 14 – 16" captionStartId="8.[151,264,1859,1882]" captionTargetBox="[196,1391,199,1829]" captionTargetId="figure@8.[184,1403,193,1838]" captionTargetPageId="8" captionText="FIGURES 14 – 16. Parahelichus granulosus, female; (14 ‒ 15) ovipositor, lateral view: (14) ovipositor of a specimen from the locality Louang Namtha, Laos, (15) a specimen from the locality Sa Pa (Lao Cai), Vietnam; (16) female sternite VIII, ventral view. Scale 0.1 mm." httpUri="https://zenodo.org/record/231190/files/figure.png" pageId="3" pageNumber="478">Fig 16</figureCitation>
|
||
); tergite similar to that of male. Male sternite IX (
|
||
<figureCitation id="135BB9E7FFFE8E01FEBA7ADDFEE19FC4" box="[292,363,1952,1977]" captionStart="FIGURES 3 – 13" captionStartId="7.[151,264,1950,1973]" captionTargetBox="[217,1367,201,1913]" captionTargetId="figure@7.[206,1381,193,1929]" captionTargetPageId="7" captionText="FIGURES 3 – 13. Parahelichus granulosus, male; (3 – 5, 8 – 13) aedeagus, penis and parameres in lateral, ventral and dorsal view; figures show differences in proportions between parameres and phallobase; (6) male sternite VIII, ventral view; (7) male sternite IX, ventral view. Scale 0.1 mm." httpUri="https://zenodo.org/record/231189/files/figure.png" pageId="3" pageNumber="478">Fig. 7</figureCitation>
|
||
) with anterior median process narrow and nearly as long as ventrites 4–5 combined; female sternite IX forming part of ovipositor.
|
||
</paragraph>
|
||
<caption id="DF1FF5EAFFF98E06FF097A6BFB889F50" box="[151,1026,1814,1836]" pageId="4" pageNumber="479">
|
||
<paragraph id="8BDFA562FFF98E06FF097A6BFB889F50" blockId="4.[151,1026,1814,1836]" box="[151,1026,1814,1836]" pageId="4" pageNumber="479">
|
||
<emphasis id="B9147970FFF98E06FF097A6BFE989F57" bold="true" box="[151,274,1814,1836]" pageId="4" pageNumber="479">FIGURE 2.</emphasis>
|
||
Habitus of
|
||
<taxonomicName id="4C60DEE1FFF98E06FE177A6BFD459F50" box="[393,719,1814,1836]" class="Insecta" family="Dryopidae" genus="Parahelichus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="479" phylum="Arthropoda" rank="species" species="pseudogranulosus" status="sp. nov.">
|
||
<emphasis id="B9147970FFF98E06FE177A6BFD459F50" box="[393,719,1814,1836]" italics="true" pageId="4" pageNumber="479">Parahelichus pseudogranulosus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A227C40BFFF98E06FD4B7A6AFCAC9F50" box="[725,806,1815,1836]" pageId="4" pageNumber="479" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, male, TL = 5.4 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BDFA562FFF98E07FF597A2EFF549884" blockId="4.[151,1436,1875,2008]" lastBlockId="5.[151,1437,151,896]" lastPageId="5" lastPageNumber="480" pageId="4" pageNumber="479">
|
||
Aedeagus (
|
||
<figureCitation id="135BB9E7FFF98E06FEDA7A2EFE679F10" box="[324,493,1875,1900]" captionStart="FIGURES 3 – 13" captionStartId="7.[151,264,1950,1973]" captionTargetBox="[217,1367,201,1913]" captionTargetId="figure@7.[206,1381,193,1929]" captionTargetPageId="7" captionText="FIGURES 3 – 13. Parahelichus granulosus, male; (3 – 5, 8 – 13) aedeagus, penis and parameres in lateral, ventral and dorsal view; figures show differences in proportions between parameres and phallobase; (6) male sternite VIII, ventral view; (7) male sternite IX, ventral view. Scale 0.1 mm." httpUri="https://zenodo.org/record/231189/files/figure.png" pageId="4" pageNumber="479">Figs 3–5, 8–13</figureCitation>
|
||
) with phallobase robust, long, subcylindrical, moderately curved, 1.14–1.77 (1.40 ± 0.12) times as long as parameres, basally wider than apically (lateral view), with basal orifice asymmetrical, large; parameres individually articulated with phallobase, widest basally, moderately long, gradually curved ventrad and gradually narrowed toward apices; apices rounded in lateral and ventral view. Penis narrow, symmetrical, moderately shorter than parameres, widest basally, gradually narrowed apically; apex narrowly rounded (ventral view); membranous ventral sac with fine longitudinal furrows; fibula narrow, reaching middle of penis (ventral view).
|
||
</paragraph>
|
||
<paragraph id="8BDFA562FFF88E07FF597C79FCF099F4" blockId="5.[151,1437,151,896]" pageId="5" pageNumber="480">
|
||
Ovipositor (
|
||
<figureCitation id="135BB9E7FFF88E07FECC7C79FE519961" box="[338,475,260,285]" captionStart="FIGURES 14 – 16" captionStartId="8.[151,264,1859,1882]" captionTargetBox="[196,1391,199,1829]" captionTargetId="figure@8.[184,1403,193,1838]" captionTargetPageId="8" captionText="FIGURES 14 – 16. Parahelichus granulosus, female; (14 ‒ 15) ovipositor, lateral view: (14) ovipositor of a specimen from the locality Louang Namtha, Laos, (15) a specimen from the locality Sa Pa (Lao Cai), Vietnam; (16) female sternite VIII, ventral view. Scale 0.1 mm." httpUri="https://zenodo.org/record/231190/files/figure.png" pageId="5" pageNumber="480">Figs 14, 15</figureCitation>
|
||
) about as long as ventrites 3–5 combined, strongly sclerotized; composed of two proximal valvifers and two distal coxites; valvifers wide, long and flattened, medially fused; coxites flattened, plate-like, vertical; left and right coxite slightly asymmetrical, shorter than valvifers, without styli, narrowed posteriorly. Bursa copulatrix tubular, without sclerotized spinules.
|
||
</paragraph>
|
||
<paragraph id="8BDFA562FFF88E07FF597CE9FE1F9A6B" blockId="5.[151,1437,151,896]" pageId="5" pageNumber="480">
|
||
<emphasis id="B9147970FFF88E07FF597CE9FE3299D1" bold="true" box="[199,440,404,429]" pageId="5" pageNumber="480">Sexual dimorphism.</emphasis>
|
||
Females are generally larger than males and both sexes differ in the shape and excision of the apex of ventrite 5. The apex of ventrite 5 is slightly excised in males (
|
||
<figureCitation id="135BB9E7FFF88E07FB907CCAFBE999AC" box="[1038,1123,439,464]" captionStart="FIGURES 17 – 20" captionStartId="9.[151,264,1103,1126]" captionTargetBox="[238,1350,196,1075]" captionTargetId="figure@9.[231,1355,193,1082]" captionTargetPageId="9" captionText="FIGURES 17 – 20. Parahelichus granulosus; (17 ‒ 18) female antennal club: (17) ventral view, (18) dorsal view, scale 0.1 mm; (19) abdomen of a male, ventral view; (20) abdomen of a female, ventral view; scale 1 mm." httpUri="https://zenodo.org/record/231191/files/figure.png" pageId="5" pageNumber="480">Fig. 19</figureCitation>
|
||
), while rounded in females (
|
||
<figureCitation id="135BB9E7FFF88E07FF017CA1FF799988" box="[159,243,476,501]" captionStart="FIGURES 17 – 20" captionStartId="9.[151,264,1103,1126]" captionTargetBox="[238,1350,196,1075]" captionTargetId="figure@9.[231,1355,193,1082]" captionTargetPageId="9" captionText="FIGURES 17 – 20. Parahelichus granulosus; (17 ‒ 18) female antennal club: (17) ventral view, (18) dorsal view, scale 0.1 mm; (19) abdomen of a male, ventral view; (20) abdomen of a female, ventral view; scale 1 mm." httpUri="https://zenodo.org/record/231191/files/figure.png" pageId="5" pageNumber="480">Fig. 20</figureCitation>
|
||
). Since ranges of morphometric characters overlap between males and females, they cannot be used for sex recognition of its own.
|
||
</paragraph>
|
||
<paragraph id="8BDFA562FFF88E07FF597F59FEB69AB1" blockId="5.[151,1437,151,896]" pageId="5" pageNumber="480">
|
||
<emphasis id="B9147970FFF88E07FF597F59FEC79A41" bold="true" box="[199,333,548,573]" pageId="5" pageNumber="480">Variability.</emphasis>
|
||
The PCA resulted from 12-selected metric characters of males demonstrated that all geographical populations are mutually overlapped with each other in the graph (Fig. 33b). Since the metric data overlaped mutually (
|
||
<tableCitation id="C6E290D9FFF88E07FE967F11FED99AF8" box="[264,339,620,645]" captionStart="TABLE 1" captionStartId="6.[119,197,594,614]" captionTargetBox="[127,1893,759,1047]" captionTargetPageId="6" captionText="TABLE 1. Variability of selected metric characters in males of Parahelichus granulosus. HW – head width including eyes, ID – interocular distance, APW – anterior pronotal width, PPW – posterior pronotal width, PW – pronotal width, maximum width of pronotum, EW – elytral width, maximum width combined, MPL – pronotal length along the body midline, EL – elytral length along suture, BL – body length, AEP – aedeagus proportion, ratio of length of parameres to length of phallobase. First rows show ranges of measured values for specimens for different localities, in second rows mean and standard deviation for the respective measurements, n – number of measured specimens. Numbers of missing values of aedeagus proportion in some localities – see Material" httpUri="http://table.plazi.org/id/DF1FF5EAFFFB8E04FFE97F2FF8029AC8" pageId="5" pageNumber="480" tableUuid="DF1FF5EAFFFB8E04FFE97F2FF8029AC8">Tabs 1</tableCitation>
|
||
and
|
||
<tableCitation id="C6E290D9FFF88E07FE177F11FE139AF8" box="[393,409,620,644]" captionStart="TABLE 2" captionStartId="6.[119,197,1098,1118]" captionTargetBox="[127,1762,1238,1582]" captionTargetPageId="6" captionText="TABLE 2. Variability of selected metric characters in females of Parahelichus granulosus. HW – head width including eyes, ID – interocular distance, APW – anterior pronotal width, PPW – posterior pronotal width, PW – pronotal width, maximum width of pronotum, EW – elytral width, maximum width combined, MPL – pronotal length along the body midline, EL – elytral length along suture, BL – body length. First rows show ranges of measured values for specimens for different localities, in second rows mean and standard deviation for the respective measurements, n – number of measured specimens. HW and ID were not measured in one female from Sa Pa." httpUri="http://table.plazi.org/id/DF1FF5EAFFFB8E04FFE97937FC389CD1" pageId="5" pageNumber="480" tableUuid="DF1FF5EAFFFB8E04FFE97937FC389CD1">2</tableCitation>
|
||
), we consider specimens from different localities to fit within the species variability. Except for variability of metric characters, specimens moderately vary in the curvature of pronotal sides and in the density of granulation.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37AF6E9FFF88E07FF597FAAFAF89BFC" pageId="5" pageNumber="480" type="distribution">
|
||
<paragraph id="8BDFA562FFF88E07FF597FAAFAB89B68" blockId="5.[151,1437,151,896]" pageId="5" pageNumber="480">
|
||
<emphasis id="B9147970FFF88E07FF597FAAFEEA9A8C" bold="true" box="[199,352,727,752]" pageId="5" pageNumber="480">Distribution.</emphasis>
|
||
So far known from
|
||
<collectingCountry id="F377E5F2FFF88E07FDD87FAAFD2F9A8C" box="[582,677,727,752]" name="Vietnam" pageId="5" pageNumber="480">Vietnam</collectingCountry>
|
||
and
|
||
<collectingCountry id="F377E5F2FFF88E07FD427FA5FC989A93" box="[732,786,728,751]" name="Laos" pageId="5" pageNumber="480">Laos</collectingCountry>
|
||
(
|
||
<figureCitation id="135BB9E7FFF88E07FCBD7FAAFCF29A8C" box="[803,888,727,752]" captionStart="FIGURE 32" captionStartId="11.[151,250,1596,1618]" captionTargetBox="[172,1412,180,1572]" captionTargetId="figure@11.[166,1420,177,1579]" captionTargetPageId="11" captionText="FIGURE 32. Geographical distribution of Parahelichus granulosu s (black squares) and P. pseudogranulosus sp. nov. (black circles)." httpUri="https://zenodo.org/record/231193/files/figure.png" pageId="5" pageNumber="480">Fig. 32</figureCitation>
|
||
). All specimens were collected using light traps. This species probably represents one of the most commonly light collected dryopid in
|
||
<collectingCountry id="F377E5F2FFF88E07FBC17F81FB349B69" box="[1119,1214,764,789]" name="Vietnam" pageId="5" pageNumber="480">Vietnam</collectingCountry>
|
||
and
|
||
<collectingCountry id="F377E5F2FFF88E07FB6A7F80FAA49B68" box="[1268,1326,765,788]" name="Laos" pageId="5" pageNumber="480">Laos</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BDFA562FFF88E07FF597E62FAF89BFC" blockId="5.[151,1437,151,896]" pageId="5" pageNumber="480">
|
||
<emphasis id="B9147970FFF88E07FF597E62FE8C9B44" bold="true" box="[199,262,799,824]" pageId="5" pageNumber="480">Note.</emphasis>
|
||
The collection of A. Grouvelle (MNHP) contains a single specimen labelled as: “
|
||
<taxonomicName id="4C60DEE1FFF88E07FB5C7E5DFF549B20" authority="Vaulogeri Grouv" authorityName="Vaulogeri Grouv" class="Insecta" family="Dryopidae" genus="Helichus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="480" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9147970FFF88E07FB5C7E5DFA169B44" box="[1218,1436,800,824]" italics="true" pageId="5" pageNumber="480">Helichus Vaulogeri</emphasis>
|
||
Grouv
|
||
</taxonomicName>
|
||
/
|
||
<typeStatus id="54DB1BC0FFF88E07FF737E39FEAE9B20" box="[237,292,836,860]" pageId="5" pageNumber="480">Type</typeStatus>
|
||
/ Hanoi, Vaulogue”. We consider this specimen to be conspecific with
|
||
<taxonomicName id="4C60DEE1FFF88E07FBA07E38FB5C9B20" box="[1086,1238,836,860]" class="Insecta" family="Dryopidae" genus="Parahelichus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="480" phylum="Arthropoda" rank="species" species="granulosus">
|
||
<emphasis id="B9147970FFF88E07FBA07E38FB5C9B20" box="[1086,1238,836,860]" italics="true" pageId="5" pageNumber="480">P. granulosus</emphasis>
|
||
</taxonomicName>
|
||
. To date we have not been able to find any published description of
|
||
<taxonomicName id="4C60DEE1FFF88E07FD577E14FCDC9BFC" box="[713,854,872,896]" class="Insecta" family="Dryopidae" genus="Helichus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="480" phylum="Arthropoda" rank="species" species="vaulogeri">
|
||
<emphasis id="B9147970FFF88E07FD577E14FCDC9BFC" box="[713,854,872,896]" italics="true" pageId="5" pageNumber="480">H. vaulogeri</emphasis>
|
||
</taxonomicName>
|
||
, and thus we consider this name as unavailable.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |