739 lines
83 KiB
XML
739 lines
83 KiB
XML
<document id="3155633857938B7C4F25EEBA313D30C3" ID-DOI="10.11646/zootaxa.3852.4.5" ID-GBIF-Dataset="0d568f39-128d-4b01-b6ae-0e7d4f92a06a" ID-ISSN="1175-5326" ID-Zenodo-Dep="231188" ID-ZooBank="033A771C-839D-41C5-91C3-7D02A661F4C9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460885311263" checkinUser="plazi" docAuthor="Lojková, Soňa, Degma, Peter & Kodada, Ján" docDate="2014" docId="03C91474FFF88E0EFF097EBBFF7B9F92" docLanguage="en" docName="zt03852p488.pdf" docOrigin="Zootaxa 3852 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Parahelichus pseudogranulosus Lojková, Degma & Kodada, 2014, sp. nov." docType="treatment" docVersion="10" lastPageNumber="487" masterDocId="FFF06C0CFFFD8E02FF9E7D7DFF8A987C" masterDocTitle="Morphometry and redescription of Parahelichus granulosus (Delève, 1974) with description of P. pseudogranulosus, a new cryptic species of Long-toed Water Beetles (Coleoptera: Dryopidae) from Indochinese peninsula, and proposal of a new synonym for Praehelichus sericatus (Waterhouse, 1881) from China" masterLastPageNumber="488" masterPageNumber="475" pageNumber="480" updateTime="1698358941206" updateUser="plazi">
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<mods:mods id="E7EF954B4364DABC91880733468C2A5A" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="5D916FD1E2606BB4692B78EA2862645A">
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<mods:title id="4DD1488D06AFA3A0B3CACB37E7918FA5">Morphometry and redescription of Parahelichus granulosus (Delève, 1974) with description of P. pseudogranulosus, a new cryptic species of Long-toed Water Beetles (Coleoptera: Dryopidae) from Indochinese peninsula, and proposal of a new synonym for Praehelichus sericatus (Waterhouse, 1881) from China</mods:title>
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</mods:titleInfo>
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<mods:name id="676FC758D7F51A62B23736405B8DA391" type="personal">
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<mods:role id="B4E265694869FD4109A7A9C6108C63A2">
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<mods:roleTerm id="D1FD71BD2DCBF2F75C3B421C8E285B67">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="EF3B6AADF4323F60C6D8C841031770F3">Lojková, Soňa</mods:namePart>
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</mods:name>
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<mods:name id="7862C8D8D80AAB3C7BB9AA8A8DEC187F" type="personal">
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<mods:role id="7180EBC733C702D8823E7361AE10262B">
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<mods:roleTerm id="4270B518B514D4845DE823EBE34C34F5">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="B668830C3C90763FFF8E00AB25A19C41">Degma, Peter</mods:namePart>
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</mods:name>
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<mods:name id="6A4FA7ADC2533FF5ED294E72AA4FF7A7" type="personal">
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<mods:role id="C1CA2E0DD8E15629FDD3362A5E2F56DC">
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<mods:roleTerm id="C03EBB3A4F2F570460D0865A98DE45D6">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="D0499CEA01ECBA0BE7C05167CF46FF94">Kodada, Ján</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="995FA40A233BB905B54777EA778B482E">text</mods:typeOfResource>
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<mods:titleInfo id="56A258CCCC9BAEEEA97DF03191C4A437">
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<mods:title id="031A0C82C20397D7FA7D11955A2106CD">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="B0BC9CCD8B9A46343EAFA391E2832B21">
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<mods:date id="1DA77267ED407027E39A2FB9004A977B">2014</mods:date>
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<mods:detail id="3637D4BD907023CCF783E30A02671EFE" type="volume">
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<mods:number id="0F21BA39466F8380166F9ED765445D47">3852</mods:number>
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</mods:detail>
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<mods:detail id="A890CEF28045605B537579461C4F6F59" type="issue">
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<mods:number id="A132E64EBD0DA9AD7AF5771870624AC7">4</mods:number>
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<mods:classification id="D3C8B22B623D9FA95E50337CDF23483D">journal article</mods:classification>
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<mods:identifier id="B61C9F7E85D3C06792B50FD13D07FC52" type="DOI">10.11646/zootaxa.3852.4.5</mods:identifier>
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<mods:identifier id="9D59208FDAD079DC77A57897FDE43BA0" type="GBIF-Dataset">0d568f39-128d-4b01-b6ae-0e7d4f92a06a</mods:identifier>
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<mods:identifier id="B20B0D97DDF725E24D4A0CF3D5D3E247" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="0BBC6B5579B0F5588469B3435B0F0E22" type="Zenodo-Dep">231188</mods:identifier>
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<mods:identifier id="BFCA1C09F1D523DB5E211DC37A5528C7" type="ZooBank">033A771C-839D-41C5-91C3-7D02A661F4C9</mods:identifier>
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</mods:mods>
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<treatment id="03C91474FFF88E0EFF097EBBFF7B9F92" ID-DOI="http://doi.org/10.5281/zenodo.5630237" ID-GBIF-Taxon="119586607" ID-Zenodo-Dep="5630237" LSID="urn:lsid:plazi:treatment:03C91474FFF88E0EFF097EBBFF7B9F92" httpUri="http://treatment.plazi.org/id/03C91474FFF88E0EFF097EBBFF7B9F92" lastPageId="12" lastPageNumber="487" pageId="5" pageNumber="480">
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<subSubSection id="C37AF6E9FFF88E07FF097EBBFD0A9B9C" box="[151,640,966,992]" pageId="5" pageNumber="480" type="nomenclature">
|
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<paragraph id="8BDFA562FFF88E07FF097EBBFD0A9B9C" blockId="5.[151,640,966,992]" box="[151,640,966,992]" pageId="5" pageNumber="480">
|
||
<heading id="D097120EFFF88E07FF097EBBFD0A9B9C" bold="true" box="[151,640,966,992]" fontSize="11" level="1" pageId="5" pageNumber="480" reason="1">
|
||
<emphasis id="B9147970FFF88E07FF097EBBFD0A9B9C" bold="true" box="[151,640,966,992]" pageId="5" pageNumber="480">
|
||
<taxonomicName id="4C60DEE1FFF88E07FF097EBBFD919B9C" box="[151,539,966,992]" class="Insecta" family="Dryopidae" genus="Parahelichus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="480" phylum="Arthropoda" rank="species" species="pseudogranulosus" status="sp. nov.">
|
||
<emphasis id="B9147970FFF88E07FF097EBBFD919B9C" bold="true" box="[151,539,966,992]" italics="true" pageId="5" pageNumber="480">Parahelichus pseudogranulosus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A227C40BFFF88E07FDBC7EBBFD0A9B9C" box="[546,640,966,992]" pageId="5" pageNumber="480" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37AF6E9FFF88E07FF097970FCE79C59" box="[151,877,1037,1062]" pageId="5" pageNumber="480" type="materials_examined">
|
||
<paragraph id="8BDFA562FFF88E07FF097970FCE79C59" blockId="5.[151,1437,1037,1998]" box="[151,877,1037,1062]" pageId="5" pageNumber="480">
|
||
<emphasis id="B9147970FFF88E07FF097970FEBF9C5A" bold="true" box="[151,309,1037,1062]" pageId="5" pageNumber="480">
|
||
<typeStatus id="54DB1BC0FFF88E07FF097970FF589C5A" box="[151,210,1037,1062]" pageId="5" pageNumber="480">Type</typeStatus>
|
||
locality.
|
||
</emphasis>
|
||
Pakka Ding, Bolikhamxai province, Central
|
||
<collectingCountry id="F377E5F2FFF88E07FCB17973FCE39C59" box="[815,873,1038,1061]" name="Laos" pageId="5" pageNumber="480">Laos</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37AF6E9FFF88E07FF59794CFD279C11" pageId="5" pageNumber="480" type="etymology">
|
||
<paragraph id="8BDFA562FFF88E07FF59794CFD279C11" blockId="5.[151,1437,1037,1998]" pageId="5" pageNumber="480">
|
||
<emphasis id="B9147970FFF88E07FF59794CFEC49C36" bold="true" box="[199,334,1073,1098]" pageId="5" pageNumber="480">Etymology.</emphasis>
|
||
The name is a combination of the prefix
|
||
<emphasis id="B9147970FFF88E07FCBA794CFCF99C35" box="[804,883,1073,1097]" italics="true" pageId="5" pageNumber="480">pseudo</emphasis>
|
||
(apparently similar) and
|
||
<taxonomicName id="4C60DEE1FFF88E07FB0C794CFA849C35" box="[1170,1294,1073,1097]" class="Insecta" family="Dryopidae" genus="Parahelichus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="480" phylum="Arthropoda" rank="species" species="granulosus">
|
||
<emphasis id="B9147970FFF88E07FB0C794CFA849C35" box="[1170,1294,1073,1097]" italics="true" pageId="5" pageNumber="480">granulosus</emphasis>
|
||
</taxonomicName>
|
||
in reference to the very similar appearance as
|
||
<emphasis id="B9147970FFF88E07FD93792BFD279C11" box="[525,685,1109,1133]" italics="true" pageId="5" pageNumber="480">
|
||
<taxonomicName id="4C60DEE1FFF88E07FD93792BFD239C11" box="[525,681,1109,1133]" class="Insecta" family="Dryopidae" genus="Parahelichus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="480" phylum="Arthropoda" rank="species" species="granulosus">P. granulosus</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37AF6E9FFF88E07FF597904FBB19EAE" pageId="5" pageNumber="480" type="materials_examined">
|
||
<paragraph id="8BDFA562FFF88E07FF597904FC2D9D16" blockId="5.[151,1437,1037,1998]" pageId="5" pageNumber="480">
|
||
<emphasis id="B9147970FFF88E07FF597904FDA39CEE" bold="true" box="[199,553,1145,1170]" pageId="5" pageNumber="480">
|
||
Material examined.
|
||
<typeStatus id="54DB1BC0FFF88E07FE227904FDA39CEE" box="[444,553,1145,1170]" pageId="5" pageNumber="480" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
♂
|
||
<emphasis id="B9147970FFF88E07FDCE7904FD3C9CEE" bold="true" box="[592,694,1145,1170]" pageId="5" pageNumber="480">
|
||
(
|
||
<collectionCode id="ED713DA7FFF88E07FDC67904FD239CEE" box="[600,681,1145,1170]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="5" pageNumber="480">NMW</collectionCode>
|
||
):
|
||
</emphasis>
|
||
“LAOS-C,
|
||
<date id="FFDE83A2FFF88E07FCDF7904FC699CEE" box="[833,995,1145,1170]" pageId="5" pageNumber="480" value="2003-05-26" valueMax="2003-05-27" valueMin="2003-05-26">26.-27.V.2003</date>
|
||
, Bolikhamxai prov.,
|
||
<collectionCode id="ED713DA7FFF88E07FB4D7907FA0A9CEE" box="[1235,1408,1146,1170]" pageId="5" pageNumber="480">PAKKA DING</collectionCode>
|
||
, ~
|
||
<quantity id="4C980887FFF88E07FF0979E3FF6A9CC9" box="[151,224,1182,1206]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="480" unit="m" value="300.0">300 m</quantity>
|
||
,
|
||
<geoCoordinate id="EE54C3A5FFF88E07FF7579E3FEC19CCA" box="[235,331,1182,1206]" direction="north" orientation="latitude" pageId="5" pageNumber="480" precision="925" value="18.333334">18°20´N</geoCoordinate>
|
||
<geoCoordinate id="EE54C3A5FFF88E07FECD79E3FE4B9CCA" box="[339,449,1182,1206]" direction="east" orientation="longitude" pageId="5" pageNumber="480" precision="925" value="104.0">104°00´E</geoCoordinate>
|
||
, Vít Kubáň leg.“
|
||
<emphasis id="B9147970FFF88E07FD1B79E0FC8E9CCA" bold="true" box="[645,772,1181,1206]" pageId="5" pageNumber="480">
|
||
<typeStatus id="54DB1BC0FFF88E07FD1B79E0FD759CCA" box="[645,767,1181,1206]" pageId="5" pageNumber="480" type="paratype">Paratypes</typeStatus>
|
||
.
|
||
</emphasis>
|
||
<specimenCount id="9D666EEBFFF88E07FC9479E3FCA79CCA" box="[778,813,1182,1206]" pageId="5" pageNumber="480" type="generic">1♂</specimenCount>
|
||
(
|
||
<collectionCode id="ED713DA7FFF88E07FCA379E3FCF79CCA" box="[829,893,1182,1206]" pageId="5" pageNumber="480">CKB</collectionCode>
|
||
): “NW
|
||
<collectingCountry id="F377E5F2FFF88E07FC4F79E3FB9E9CCA" box="[977,1044,1182,1206]" name="Thailand" pageId="5" pageNumber="480">THAI</collectingCountry>
|
||
<date id="FFDE83A2FFF88E07FB8579E3FB089CCA" box="[1051,1154,1182,1206]" pageId="5" pageNumber="480">23.-31.V.</date>
|
||
MAE
|
||
<collectionCode id="ED713DA7FFF88E07FB5079E3FA959CCA" box="[1230,1311,1182,1206]" pageId="5" pageNumber="480">HONG</collectionCode>
|
||
SON
|
||
<quantity id="4C980887FFF88E07FAFA79E3FF259CA5" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="5" pageNumber="480" unit="m" value="1200.0">1200 m</quantity>
|
||
, Ban Si Lang 1991, J. Horák leg.“;
|
||
<specimenCount id="9D666EEBFFF88E07FDDE79BFFDE09CA5" box="[576,618,1218,1242]" pageId="5" pageNumber="480" type="generic">1 ♂</specimenCount>
|
||
(
|
||
<collectionCode id="ED713DA7FFF88E07FDE779BFFD429CA6" box="[633,712,1218,1242]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="5" pageNumber="480">NMW</collectionCode>
|
||
): “
|
||
<collectingCountry id="F377E5F2FFF88E07FD7A79BFFCA59CA6" box="[740,815,1218,1242]" name="Laos" pageId="5" pageNumber="480">LAOS</collectingCountry>
|
||
centr., Khammouan prov.,
|
||
<date id="FFDE83A2FFF88E07FBFF79BFFA859CA6" box="[1121,1295,1218,1242]" pageId="5" pageNumber="480" value="2001-06-10" valueMax="2001-06-13" valueMin="2001-06-10">10.-13.VI.2001</date>
|
||
, Ban Khoun Ngeun env.
|
||
<geoCoordinate id="EE54C3A5FFF88E07FEB3799BFE059C82" box="[301,399,1254,1278]" direction="north" orientation="latitude" pageId="5" pageNumber="480" precision="925" value="18.116667">18°07´N</geoCoordinate>
|
||
<geoCoordinate id="EE54C3A5FFF88E07FE03799BFD869C82" box="[413,524,1254,1278]" direction="east" orientation="longitude" pageId="5" pageNumber="480" precision="925" value="104.48333">104°29´E</geoCoordinate>
|
||
, alt.
|
||
<quantity id="4C980887FFF88E07FDDE7998FD199C81" box="[576,659,1253,1278]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="5" pageNumber="480" unit="m" value="250.0">250 m</quantity>
|
||
, E. Jendek & O.Šauša leg.“.
|
||
<emphasis id="B9147970FFF88E07FB967998FE029D5E" bold="true" pageId="5" pageNumber="480">
|
||
Non
|
||
<typeStatus id="54DB1BC0FFF88E07FBD77998FBF69C82" box="[1097,1148,1253,1278]" pageId="5" pageNumber="480">type</typeStatus>
|
||
material of presumed conspecific females.
|
||
</emphasis>
|
||
2 ♀♀ (
|
||
<collectionCode id="ED713DA7FFF88E07FE717877FDA49D5E" box="[495,558,1290,1314]" pageId="5" pageNumber="480">CKB</collectionCode>
|
||
): “LAOS-C,
|
||
<date id="FFDE83A2FFF88E07FD537874FCE59D5E" box="[717,879,1289,1314]" pageId="5" pageNumber="480" value="2003-05-26" valueMax="2003-05-27" valueMin="2003-05-26">26.-27.V.2003</date>
|
||
, Bolikhamxai prov.,
|
||
<collectionCode id="ED713DA7FFF88E07FBF57877FA919D5E" box="[1131,1307,1290,1314]" pageId="5" pageNumber="480">PAKKA DING</collectionCode>
|
||
, ~
|
||
<quantity id="4C980887FFF88E07FAD67877FA1D9D5D" box="[1352,1431,1290,1314]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="480" unit="m" value="300.0">300 m</quantity>
|
||
,
|
||
<geoCoordinate id="EE54C3A5FFF88E07FF097850FF7D9D39" box="[151,247,1325,1349]" direction="north" orientation="latitude" pageId="5" pageNumber="480" precision="925" value="18.333334">18°20´N</geoCoordinate>
|
||
<geoCoordinate id="EE54C3A5FFF88E07FF617850FEE79D39" box="[255,365,1325,1349]" direction="east" orientation="longitude" pageId="5" pageNumber="480" precision="925" value="104.0">104°00´E</geoCoordinate>
|
||
, Vít Kubáň leg.“; 2 ♀♀ (
|
||
<collectionCode id="ED713DA7FFF88E07FD167850FD429D39" box="[648,712,1325,1349]" pageId="5" pageNumber="480">CKB</collectionCode>
|
||
): “
|
||
<collectingCountry id="F377E5F2FFF88E07FD7B7850FCBA9D39" box="[741,816,1325,1349]" name="Laos" pageId="5" pageNumber="480">LAOS</collectingCountry>
|
||
centr., Khammouan prov.,
|
||
<date id="FFDE83A2FFF88E07FBFC7850FA9A9D39" box="[1122,1296,1325,1349]" pageId="5" pageNumber="480" value="2001-06-10" valueMax="2001-06-13" valueMin="2001-06-10">10.-13.VI.2001</date>
|
||
, Ban Khoun Ngeun env.
|
||
<geoCoordinate id="EE54C3A5FFF88E07FE83782FFEF49D16" box="[285,382,1362,1386]" direction="north" orientation="latitude" pageId="5" pageNumber="480" precision="925" value="18.116667">18°07´N</geoCoordinate>
|
||
<geoCoordinate id="EE54C3A5FFF88E07FE1B782FFE799D16" box="[389,499,1362,1386]" direction="east" orientation="longitude" pageId="5" pageNumber="480" precision="925" value="104.48333">104°29´E</geoCoordinate>
|
||
, alt.
|
||
<quantity id="4C980887FFF88E07FD81782CFDE09D15" box="[543,618,1361,1386]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="5" pageNumber="480" unit="m" value="250.0">250 m</quantity>
|
||
, E. Jendek & O.Šauša leg.“.
|
||
</paragraph>
|
||
<paragraph id="8BDFA562FFF88E07FF597808FE919E19" blockId="5.[151,1437,1037,1998]" pageId="5" pageNumber="480">
|
||
<emphasis id="B9147970FFF88E07FF597808FE449DF2" bold="true" box="[199,462,1397,1422]" pageId="5" pageNumber="480">Differential diagnosis.</emphasis>
|
||
<taxonomicName id="4C60DEE1FFF88E07FE4B780BFD379DF1" box="[469,701,1397,1421]" class="Insecta" family="Dryopidae" genus="Parahelichus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="480" phylum="Arthropoda" rank="species" species="pseudogranulosus">
|
||
<emphasis id="B9147970FFF88E07FE4B780BFD379DF1" box="[469,701,1397,1421]" italics="true" pageId="5" pageNumber="480">P. pseudogranulosus</emphasis>
|
||
</taxonomicName>
|
||
is externally very similar to
|
||
<emphasis id="B9147970FFF88E07FB9D780BFB299DF1" box="[1027,1187,1397,1421]" italics="true" pageId="5" pageNumber="480">
|
||
<taxonomicName id="4C60DEE1FFF88E07FB9D780BFB159DF1" box="[1027,1183,1397,1421]" class="Insecta" family="Dryopidae" genus="Parahelichus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="480" phylum="Arthropoda" rank="species" species="granulosus">P. granulosus</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
Males and females of both species cannot be recognized from each other according to metric characters as their ranges overlap. The only exceptions are values of aedeagus proportions: phallobase in
|
||
<taxonomicName id="4C60DEE1FFF88E07FCC778C3FBC99DA9" box="[857,1091,1469,1493]" class="Insecta" family="Dryopidae" genus="Parahelichus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="480" phylum="Arthropoda" rank="species" species="pseudogranulosus">
|
||
<emphasis id="B9147970FFF88E07FCC778C3FBC99DA9" box="[857,1091,1469,1493]" italics="true" pageId="5" pageNumber="480">P. pseudogranulosus</emphasis>
|
||
</taxonomicName>
|
||
is 2.00–2.22 times as long as parameres, while the ratio is
|
||
<quantity id="4C980887FFF88E07FE75789FFDF69D86" box="[491,636,1505,1530]" metricMagnitude="-2" metricUnit="m" metricValue="3.6957" metricValueMax="4.4958" metricValueMin="2.8956" pageId="5" pageNumber="480" unit="in" value="1.455" valueMax="1.77" valueMin="1.14">1.14–1.77 in</quantity>
|
||
<taxonomicName id="4C60DEE1FFF88E07FD1B789FFCAA9D85" box="[645,800,1505,1529]" class="Insecta" family="Dryopidae" genus="Parahelichus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="480" phylum="Arthropoda" rank="species" species="granulosus">
|
||
<emphasis id="B9147970FFF88E07FD1B789FFCAA9D85" box="[645,800,1505,1529]" italics="true" pageId="5" pageNumber="480">P. granulosus</emphasis>
|
||
</taxonomicName>
|
||
. These higher values for this character were shown to clearly separate males of
|
||
<taxonomicName id="4C60DEE1FFF88E07FE227B7BFD2F9E61" box="[444,677,1541,1565]" class="Insecta" family="Dryopidae" genus="Parahelichus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="480" phylum="Arthropoda" rank="species" species="pseudogranulosus">
|
||
<emphasis id="B9147970FFF88E07FE227B7BFD2F9E61" box="[444,677,1541,1565]" italics="true" pageId="5" pageNumber="480">P. pseudogranulosus</emphasis>
|
||
</taxonomicName>
|
||
from those of
|
||
<taxonomicName id="4C60DEE1FFF88E07FCCC7B7BFC669E61" box="[850,1004,1541,1565]" class="Insecta" family="Dryopidae" genus="Parahelichus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="480" phylum="Arthropoda" rank="species" species="granulosus">
|
||
<emphasis id="B9147970FFF88E07FCCC7B7BFC669E61" box="[850,1004,1541,1565]" italics="true" pageId="5" pageNumber="480">P. granulosus</emphasis>
|
||
</taxonomicName>
|
||
in the PCA (Fig. 33). Except for this difference, the flat larger granules on pronotum and elytra of
|
||
<taxonomicName id="4C60DEE1FFF88E07FCDC7B57FBA09E3D" box="[834,1066,1577,1601]" class="Insecta" family="Dryopidae" genus="Parahelichus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="480" phylum="Arthropoda" rank="species" species="pseudogranulosus">
|
||
<emphasis id="B9147970FFF88E07FCDC7B57FBA09E3D" box="[834,1066,1577,1601]" italics="true" pageId="5" pageNumber="480">P. pseudogranulosus</emphasis>
|
||
</taxonomicName>
|
||
look a little bit smaller than in
|
||
<emphasis id="B9147970FFF88E07FA197B57FE919E19" italics="true" pageId="5" pageNumber="480">
|
||
<taxonomicName id="4C60DEE1FFF88E07FA197B57FE9D9E19" class="Insecta" family="Dryopidae" genus="Parahelichus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="480" phylum="Arthropoda" rank="species" species="granulosus">P. granulosus</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BDFA562FFF88E07FF597B0CFBB19EAE" blockId="5.[151,1437,1037,1998]" pageId="5" pageNumber="480">
|
||
Since it is not possible to unambiguously assign females to
|
||
<taxonomicName id="4C60DEE1FFF88E07FC387B0FFBCF9EF5" box="[934,1093,1649,1673]" class="Insecta" family="Dryopidae" genus="Parahelichus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="480" phylum="Arthropoda" rank="species" species="granulosus">
|
||
<emphasis id="B9147970FFF88E07FC387B0FFBCF9EF5" box="[934,1093,1649,1673]" italics="true" pageId="5" pageNumber="480">P. granulosus</emphasis>
|
||
</taxonomicName>
|
||
or
|
||
<taxonomicName id="4C60DEE1FFF88E07FBE67B0FFAED9EF5" box="[1144,1383,1649,1673]" class="Insecta" family="Dryopidae" genus="Parahelichus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="480" phylum="Arthropoda" rank="species" species="pseudogranulosus">
|
||
<emphasis id="B9147970FFF88E07FBE67B0FFAED9EF5" box="[1144,1383,1649,1673]" italics="true" pageId="5" pageNumber="480">P. pseudogranulosus</emphasis>
|
||
</taxonomicName>
|
||
, we arbitrarily ascribed four females to
|
||
<taxonomicName id="4C60DEE1FFF88E07FDCA7BEBFCCC9ED1" box="[596,838,1685,1709]" class="Insecta" family="Dryopidae" genus="Parahelichus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="480" phylum="Arthropoda" rank="species" species="pseudogranulosus">
|
||
<emphasis id="B9147970FFF88E07FDCA7BEBFCCC9ED1" box="[596,838,1685,1709]" italics="true" pageId="5" pageNumber="480">P. pseudogranulosus</emphasis>
|
||
</taxonomicName>
|
||
of the basis of co-occurence with males of
|
||
<taxonomicName id="4C60DEE1FFF88E07FA197BEBFEE99EAD" class="Insecta" family="Dryopidae" genus="Parahelichus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="480" phylum="Arthropoda" rank="species" species="pseudogranulosus">
|
||
<emphasis id="B9147970FFF88E07FA197BEBFEE99EAD" italics="true" pageId="5" pageNumber="480">P. pseudogranulosus</emphasis>
|
||
</taxonomicName>
|
||
. Therefore, they presumed females were not included into
|
||
<typeStatus id="54DB1BC0FFF88E07FC667BC7FBBC9EAE" box="[1016,1078,1722,1746]" pageId="5" pageNumber="480">types</typeStatus>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37AF6E9FFF88E0EFF597BA0FA179B44" lastPageId="12" lastPageNumber="487" pageId="5" pageNumber="480" type="description">
|
||
<paragraph id="8BDFA562FFF88E07FF597BA0FEC89F1E" blockId="5.[151,1437,1037,1998]" pageId="5" pageNumber="480">
|
||
<emphasis id="B9147970FFF88E07FF597BA0FED39E8A" bold="true" box="[199,345,1757,1782]" pageId="5" pageNumber="480">Description.</emphasis>
|
||
Habitus of male as shown in
|
||
<figureCitation id="135BB9E7FFF88E07FD3D7BA0FD6D9E8A" box="[675,743,1757,1782]" captionStart="FIGURE 2" captionStartId="4.[151,250,1814,1836]" captionText="FIGURE 2. Habitus of Parahelichus pseudogranulosus sp. nov., male, TL = 5.4 mm." pageId="5" pageNumber="480">Fig. 2</figureCitation>
|
||
. Body elongated, moderately convex dorsally. Length (BL) in ♂♂:
|
||
<quantity id="4C980887FFF88E07FF4F7A7CFEFF9F65" box="[209,373,1793,1817]" metricMagnitude="-3" metricUnit="m" metricValue="5.92" metricValueMax="6.63" metricValueMin="5.21" pageId="5" pageNumber="480" unit="mm" value="5.92" valueMax="6.63" valueMin="5.21">5.21–6.63 mm</quantity>
|
||
(5.71 ± 0.80), in ♀♀:
|
||
<quantity id="4C980887FFF88E07FDE47A7CFC949F65" box="[634,798,1793,1817]" metricMagnitude="-3" metricUnit="m" metricValue="6.015" metricValueMax="6.71" metricValueMin="5.32" pageId="5" pageNumber="480" unit="mm" value="6.015" valueMax="6.71" valueMin="5.32">5.32–6.71 mm</quantity>
|
||
(5.89 ± 0.65); maximum width (EW) in ♂♂:
|
||
<quantity id="4C980887FFF88E07FAB27A7CFF499F41" metricMagnitude="-3" metricUnit="m" metricValue="2.45" metricValueMax="2.66" metricValueMin="2.24" pageId="5" pageNumber="480" unit="mm" value="2.45" valueMax="2.66" valueMin="2.24">2.24–2.66 mm</quantity>
|
||
(2.39 ± 0.24), in ♀♀:
|
||
<quantity id="4C980887FFF88E07FE587A5BFDE09F41" box="[454,618,1830,1854]" metricMagnitude="-3" metricUnit="m" metricValue="2.565" metricValueMax="2.89" metricValueMin="2.24" pageId="5" pageNumber="480" unit="mm" value="2.565" valueMax="2.89" valueMin="2.24">2.24–2.89 mm</quantity>
|
||
(2.53 ± 0.35). Colour nearly black, antennal clubs, mouthparts and tarsi reddish-brown.
|
||
</paragraph>
|
||
<paragraph id="8BDFA562FFF88E07FF597A10FB339FB2" blockId="5.[151,1437,1037,1998]" pageId="5" pageNumber="480">Surface opaque; densely covered with small rounded granules and larger shiny flat ones. Small granules about one third of a diameter of a facet. Larger granules moderately smaller than facet, nearly rounded and more or less regularly arranged, associated with yellowish setae. Elytra lacking distinct striae and intervals.</paragraph>
|
||
<caption id="DF1FF5EAFFFB8E04FFE97F2FF8029AC8" ID-Table-UUID="DF1FF5EAFFFB8E04FFE97F2FF8029AC8" httpUri="http://table.plazi.org/id/DF1FF5EAFFFB8E04FFE97F2FF8029AC8" pageId="6" pageNumber="481" targetBox="[127,1893,759,1047]" targetIsTable="true" targetPageId="6">
|
||
<paragraph id="8BDFA562FFFB8E04FFE97F2FF8029AC8" blockId="6.[119,1929,593,719]" pageId="6" pageNumber="481">
|
||
<emphasis id="B9147970FFFB8E04FFE97F2FFF549A1A" bold="true" box="[119,222,594,614]" pageId="6" pageNumber="481">TABLE 1.</emphasis>
|
||
Variability of selected metric characters in males of
|
||
<emphasis id="B9147970FFFB8E04FD497F2CFC4F9A1A" box="[727,965,593,614]" italics="true" pageId="6" pageNumber="481">
|
||
<taxonomicName id="4C60DEE1FFFB8E04FD497F2CFC4A9A1A" box="[727,960,593,614]" class="Insecta" family="Dryopidae" genus="Parahelichus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="481" phylum="Arthropoda" rank="species" species="granulosus">Parahelichus granulosus</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
<emphasis id="B9147970FFFB8E04FC527F2FFC7E9A1A" bold="true" box="[972,1012,594,614]" pageId="6" pageNumber="481">HW</emphasis>
|
||
– head width including eyes,
|
||
<emphasis id="B9147970FFFB8E04FA8A7F2FFAA79A1A" bold="true" box="[1300,1325,594,614]" pageId="6" pageNumber="481">ID</emphasis>
|
||
– interocular distance,
|
||
<emphasis id="B9147970FFFB8E04F9937F2FF9C89A1A" bold="true" box="[1549,1602,594,614]" pageId="6" pageNumber="481">APW</emphasis>
|
||
– anterior pronotal width,
|
||
<emphasis id="B9147970FFFB8E04F8DD7F2FF8FF9A1A" bold="true" box="[1859,1909,594,614]" pageId="6" pageNumber="481">PPW</emphasis>
|
||
– posterior pronotal width,
|
||
<emphasis id="B9147970FFFB8E04FEFD7F11FE0D9AFC" bold="true" box="[355,391,620,640]" pageId="6" pageNumber="481">PW</emphasis>
|
||
– pronotal width, maximum width of pronotum,
|
||
<emphasis id="B9147970FFFB8E04FCCF7F11FCFD9AFC" bold="true" box="[849,887,620,640]" pageId="6" pageNumber="481">EW</emphasis>
|
||
– elytral width, maximum width combined,
|
||
<emphasis id="B9147970FFFB8E04FA887F11FAC39AFC" bold="true" box="[1302,1353,620,640]" pageId="6" pageNumber="481">MPL</emphasis>
|
||
– pronotal length along the body midline,
|
||
<emphasis id="B9147970FFFB8E04F9467F11F97C9AFC" bold="true" box="[1752,1782,620,640]" pageId="6" pageNumber="481">EL</emphasis>
|
||
– elytral length along suture,
|
||
<emphasis id="B9147970FFFB8E04FF6D7FFBFE989AE6" bold="true" box="[243,274,646,666]" pageId="6" pageNumber="481">BL</emphasis>
|
||
– body length,
|
||
<emphasis id="B9147970FFFB8E04FE3F7FFBFE459AE6" bold="true" box="[417,463,646,666]" pageId="6" pageNumber="481">AEP</emphasis>
|
||
– aedeagus proportion, ratio of length of parameres to length of phallobase. First rows show ranges of measured values for specimens for different localities, in second rows mean and standard deviation for the respective measurements,
|
||
<emphasis id="B9147970FFFB8E04FCAC7FDDFCB49AC8" bold="true" box="[818,830,672,692]" pageId="6" pageNumber="481">n</emphasis>
|
||
– number of measured specimens. Numbers of missing values of aedeagus proportion in some localities – see Material
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BDFA562FFFB8E04FF007FC7FF789AB3" blockId="6.[119,1929,593,719]" box="[158,242,698,719]" pageId="6" pageNumber="481">methods.</paragraph>
|
||
<paragraph id="8BDFA562FFFB8E04FFE17F8AF8EF9C6B" pageId="6" pageNumber="481">
|
||
<table id="F96057C2FFFB71FDFFE17F8AF8EF9C6B" box="[127,1893,759,1047]" gridcols="13" gridrows="5" pageId="6" pageNumber="481">
|
||
<tr id="3550A720FFFB71FDFFE17F8AF8EF9B4E" box="[127,1893,759,818]" gridrow="0" pageId="6" pageNumber="481">
|
||
<th id="7681CE5CFFFB71FDFFE17F8AFEBF9B4E" box="[127,309,759,818]" gridcol="0" gridrow="0" pageId="6" pageNumber="481">
|
||
<emphasis id="B9147970FFFB8E04FFE17F8AFF379B7B" bold="true" box="[127,189,759,775]" pageId="6" pageNumber="481">Locality</emphasis>
|
||
Yen Bai, Vietnam n = 26
|
||
</th>
|
||
<th id="7681CE5CFFFB71FDFE1F7F8AFE6E9B4E" box="[385,484,759,818]" gridcol="1" gridrow="0" pageId="6" pageNumber="481">
|
||
<emphasis id="B9147970FFFB8E04FE1F7F8AFE459B7B" bold="true" box="[385,463,759,775]" pageId="6" pageNumber="481">HW [mm]</emphasis>
|
||
0.92‒1.24 1.053 ± 0.089
|
||
</th>
|
||
<th id="7681CE5CFFFB71FDFD9F7F8AFDEE9B4E" box="[513,612,759,818]" gridcol="2" gridrow="0" pageId="6" pageNumber="481">
|
||
<emphasis id="B9147970FFFB8E04FD9F7F8AFDC99B7B" bold="true" box="[513,579,759,775]" pageId="6" pageNumber="481">ID [mm]</emphasis>
|
||
0.42‒0.66 0.566 ± 0.059
|
||
</th>
|
||
<th id="7681CE5CFFFB71FDFD1F7F8AFD6E9B4E" box="[641,740,759,818]" gridcol="3" gridrow="0" pageId="6" pageNumber="481">
|
||
<emphasis id="B9147970FFFB8E04FD1F7F8AFD539B7B" bold="true" box="[641,729,759,775]" pageId="6" pageNumber="481">APW [mm]</emphasis>
|
||
0.97‒1.42 1.179 ± 0.113
|
||
</th>
|
||
<th id="7681CE5CFFFB71FDFC9F7F8AFCEE9B4E" box="[769,868,759,818]" gridcol="4" gridrow="0" pageId="6" pageNumber="481">
|
||
<emphasis id="B9147970FFFB8E04FC9F7F8AFCDD9B7B" bold="true" box="[769,855,759,775]" pageId="6" pageNumber="481">PPW [mm]</emphasis>
|
||
1.32‒2.00 1.620 ± 0.155
|
||
</th>
|
||
<th id="7681CE5CFFFB71FDFC1F7F8AFC6E9B4E" box="[897,996,759,818]" gridcol="5" gridrow="0" pageId="6" pageNumber="481">
|
||
<emphasis id="B9147970FFFB8E04FC1F7F8AFC469B7B" bold="true" box="[897,972,759,775]" pageId="6" pageNumber="481">PW [mm]</emphasis>
|
||
1.39‒2.00 1.660 ± 0.148
|
||
</th>
|
||
<th id="7681CE5CFFFB71FDFB9E7F8AFBEE9B4E" box="[1024,1124,759,818]" gridcol="6" gridrow="0" pageId="6" pageNumber="481">
|
||
<emphasis id="B9147970FFFB8E04FB9E7F8AFBC69B7B" bold="true" box="[1024,1100,759,775]" pageId="6" pageNumber="481">EW [mm]</emphasis>
|
||
1.76‒2.63 2.142 ± 0.199
|
||
</th>
|
||
<th id="7681CE5CFFFB71FDFB1F7F8AFB6E9B4E" box="[1153,1252,759,818]" gridcol="7" gridrow="0" pageId="6" pageNumber="481">
|
||
<emphasis id="B9147970FFFB8E04FB1F7F8AFB5C9B7B" bold="true" box="[1153,1238,759,775]" pageId="6" pageNumber="481">MPL [mm]</emphasis>
|
||
0.95‒1.42 1,176 ± 0.112
|
||
</th>
|
||
<th id="7681CE5CFFFB71FDFA9E7F8AFAE99B4E" box="[1280,1379,759,818]" gridcol="8" gridrow="0" pageId="6" pageNumber="481">
|
||
<emphasis id="B9147970FFFB8E04FA9E7F8AFACC9B7B" bold="true" box="[1280,1350,759,775]" pageId="6" pageNumber="481">EL [mm]</emphasis>
|
||
3.21‒4.84 3.957 ± 0.402
|
||
</th>
|
||
<th id="7681CE5CFFFB71FDFA1E7F8AFA699B4E" box="[1408,1507,759,818]" gridcol="9" gridrow="0" pageId="6" pageNumber="481">
|
||
<emphasis id="B9147970FFFB8E04FA1E7F8AFA4C9B7B" bold="true" box="[1408,1478,759,775]" pageId="6" pageNumber="481">BL [mm]</emphasis>
|
||
4.16‒6.26 5.134 ± 0.500
|
||
</th>
|
||
<th id="7681CE5CFFFB71FDF99E7F8AF9E89B4E" box="[1536,1634,759,818]" gridcol="10" gridrow="0" pageId="6" pageNumber="481">
|
||
<emphasis id="B9147970FFFB8E04F99E7F8AF9B09B7B" bold="true" box="[1536,1594,759,775]" pageId="6" pageNumber="481">BL/EW</emphasis>
|
||
2.20‒2.51 2.397 ± 0.070
|
||
</th>
|
||
<th id="7681CE5CFFFB71FDF91E7F8AF9699B4E" box="[1664,1763,759,818]" gridcol="11" gridrow="0" pageId="6" pageNumber="481">
|
||
<emphasis id="B9147970FFFB8E04F91E7F8AF9439B7B" bold="true" box="[1664,1737,759,775]" pageId="6" pageNumber="481">MPL/PW</emphasis>
|
||
0.65‒0.77 0.709 ± 0.034
|
||
</th>
|
||
<th id="7681CE5CFFFB71FDF89C7F8AF8EF9B4E" box="[1794,1893,759,818]" gridcol="12" gridrow="0" pageId="6" pageNumber="481">
|
||
<emphasis id="B9147970FFFB8E04F89D7F8AF8AD9B7B" bold="true" box="[1795,1831,759,775]" pageId="6" pageNumber="481">AEP</emphasis>
|
||
1.20‒1.64 1.418 ± 0.111
|
||
</th>
|
||
</tr>
|
||
<tr id="3550A720FFFB71FDFFE17E3AF8EF9B17" box="[127,1893,839,875]" gridrow="1" pageId="6" pageNumber="481">
|
||
<th id="7681CE5CFFFB71FDFFE17E3AFEBF9B17" box="[127,309,839,875]" gridcol="0" gridrow="1" pageId="6" pageNumber="481">Sa Pa (Lao Cai), Vietnam n = 91</th>
|
||
<td id="7681CE5CFFFB71FDFE1F7E3AFE6E9B17" box="[385,484,839,875]" gridcol="1" gridrow="1" pageId="6" pageNumber="481">1.00‒1.26 1.121 ± 0.064</td>
|
||
<td id="7681CE5CFFFB71FDFD9F7E3AFDEE9B17" box="[513,612,839,875]" gridcol="2" gridrow="1" pageId="6" pageNumber="481">0.50‒0.68 0.598 ± 0.046</td>
|
||
<td id="7681CE5CFFFB71FDFD1F7E3AFD6E9B17" box="[641,740,839,875]" gridcol="3" gridrow="1" pageId="6" pageNumber="481">1.00‒1.39 1.230 ± 0.075</td>
|
||
<td id="7681CE5CFFFB71FDFC9F7E3AFCEE9B17" box="[769,868,839,875]" gridcol="4" gridrow="1" pageId="6" pageNumber="481">1.21‒1.97 1.727 ± 0.123</td>
|
||
<td id="7681CE5CFFFB71FDFC1F7E3AFC6E9B17" box="[897,996,839,875]" gridcol="5" gridrow="1" pageId="6" pageNumber="481">1.24‒1.97 1.758 ± 0.123</td>
|
||
<td id="7681CE5CFFFB71FDFB9E7E3AFBEE9B17" box="[1024,1124,839,875]" gridcol="6" gridrow="1" pageId="6" pageNumber="481">1.97‒2.61 2.248 ± 0.142</td>
|
||
<td id="7681CE5CFFFB71FDFB1F7E3AFB6E9B17" box="[1153,1252,839,875]" gridcol="7" gridrow="1" pageId="6" pageNumber="481">1.05‒1.55 1.263 ± 0.093</td>
|
||
<td id="7681CE5CFFFB71FDFA9E7E3AFAE99B17" box="[1280,1379,839,875]" gridcol="8" gridrow="1" pageId="6" pageNumber="481">3.68‒4.89 4.239 ± 0.280</td>
|
||
<td id="7681CE5CFFFB71FDFA1E7E3AFA699B17" box="[1408,1507,839,875]" gridcol="9" gridrow="1" pageId="6" pageNumber="481">4.84‒6.39 5.501 ± 0.360</td>
|
||
<td id="7681CE5CFFFB71FDF99E7E3AF9E89B17" box="[1536,1634,839,875]" gridcol="10" gridrow="1" pageId="6" pageNumber="481">2.34‒2.58 2.447 ± 0.051</td>
|
||
<td id="7681CE5CFFFB71FDF91E7E3AF9699B17" box="[1664,1763,839,875]" gridcol="11" gridrow="1" pageId="6" pageNumber="481">0.63‒0.87 0.719 ± 0.037</td>
|
||
<td id="7681CE5CFFFB71FDF89C7E3AF8EF9B17" box="[1794,1893,839,875]" gridcol="12" gridrow="1" pageId="6" pageNumber="481">1.14‒1.77 1.323 ± 0.097</td>
|
||
</tr>
|
||
<tr id="3550A720FFFB71FDFFE17EFDF8EF9BD8" box="[127,1893,896,932]" gridrow="2" pageId="6" pageNumber="481">
|
||
<th id="7681CE5CFFFB71FDFFE17EFDFEBF9BD8" box="[127,309,896,932]" gridcol="0" gridrow="2" pageId="6" pageNumber="481">Louang Namtha, Laos n = 25</th>
|
||
<td id="7681CE5CFFFB71FDFE1F7EFDFE6E9BD8" box="[385,484,896,932]" gridcol="1" gridrow="2" pageId="6" pageNumber="481">0.76‒1.18 0.996 ± 0.140</td>
|
||
<td id="7681CE5CFFFB71FDFD9F7EFDFDEE9BD8" box="[513,612,896,932]" gridcol="2" gridrow="2" pageId="6" pageNumber="481">0.34‒0.63 0.509 ± 0.097</td>
|
||
<td id="7681CE5CFFFB71FDFD1F7EFDFD6E9BD8" box="[641,740,896,932]" gridcol="3" gridrow="2" pageId="6" pageNumber="481">0.87‒1.32 1.104 ± 0.140</td>
|
||
<td id="7681CE5CFFFB71FDFC9F7EFDFCEE9BD8" box="[769,868,896,932]" gridcol="4" gridrow="2" pageId="6" pageNumber="481">1.16‒1.79 1.511 ± 0.207</td>
|
||
<td id="7681CE5CFFFB71FDFC1F7EFDFC6E9BD8" box="[897,996,896,932]" gridcol="5" gridrow="2" pageId="6" pageNumber="481">1.18‒1.84 1.545 ± 0.214</td>
|
||
<td id="7681CE5CFFFB71FDFB9E7EFDFBEE9BD8" box="[1024,1124,896,932]" gridcol="6" gridrow="2" pageId="6" pageNumber="481">1.50‒2.32 1.964 ± 0.269</td>
|
||
<td id="7681CE5CFFFB71FDFB1F7EFDFB6E9BD8" box="[1153,1252,896,932]" gridcol="7" gridrow="2" pageId="6" pageNumber="481">0.89‒1.32 1.097 ± 0.117</td>
|
||
<td id="7681CE5CFFFB71FDFA9E7EFDFAE99BD8" box="[1280,1379,896,932]" gridcol="8" gridrow="2" pageId="6" pageNumber="481">2.68‒4.32 3.619 ± 0.557</td>
|
||
<td id="7681CE5CFFFB71FDFA1E7EFDFA699BD8" box="[1408,1507,896,932]" gridcol="9" gridrow="2" pageId="6" pageNumber="481">3.58‒5.63 4.716 ± 0.669</td>
|
||
<td id="7681CE5CFFFB71FDF99E7EFDF9E89BD8" box="[1536,1634,896,932]" gridcol="10" gridrow="2" pageId="6" pageNumber="481">2.28‒2.49 2.400 ± 0.058</td>
|
||
<td id="7681CE5CFFFB71FDF91E7EFDF9699BD8" box="[1664,1763,896,932]" gridcol="11" gridrow="2" pageId="6" pageNumber="481">0.64‒0.96 0.715 ± 0.064</td>
|
||
<td id="7681CE5CFFFB71FDF89C7EFDF8EF9BD8" box="[1794,1893,896,932]" gridcol="12" gridrow="2" pageId="6" pageNumber="481">1.33‒1.75 1.534 ± 0.132</td>
|
||
</tr>
|
||
<tr id="3550A720FFFB71FDFFE17EC4F8EF9BA2" box="[127,1893,953,990]" gridrow="3" pageId="6" pageNumber="481">
|
||
<th id="7681CE5CFFFB71FDFFE17EC4FEBF9BA2" box="[127,309,953,990]" gridcol="0" gridrow="3" pageId="6" pageNumber="481">Khan riv., Laos n = 70</th>
|
||
<td id="7681CE5CFFFB71FDFE1F7EC4FE6E9BA2" box="[385,484,953,990]" gridcol="1" gridrow="3" pageId="6" pageNumber="481">0.92‒1.26 1.138 ± 0.063</td>
|
||
<td id="7681CE5CFFFB71FDFD9F7EC4FDEE9BA2" box="[513,612,953,990]" gridcol="2" gridrow="3" pageId="6" pageNumber="481">0.50‒0.68 0.586 ± 0.045</td>
|
||
<td id="7681CE5CFFFB71FDFD1F7EC4FD6E9BA2" box="[641,740,953,990]" gridcol="3" gridrow="3" pageId="6" pageNumber="481">1.05‒1.39 1.252 ± 0.070</td>
|
||
<td id="7681CE5CFFFB71FDFC9F7EC4FCEE9BA2" box="[769,868,953,990]" gridcol="4" gridrow="3" pageId="6" pageNumber="481">1.50‒1.97 1.746 ± 0.101</td>
|
||
<td id="7681CE5CFFFB71FDFC1F7EC4FC6E9BA2" box="[897,996,953,990]" gridcol="5" gridrow="3" pageId="6" pageNumber="481">1.50‒1.97 1.782 ± 0.100</td>
|
||
<td id="7681CE5CFFFB71FDFB9E7EC4FBEE9BA2" box="[1024,1124,953,990]" gridcol="6" gridrow="3" pageId="6" pageNumber="481">1.87‒2.50 2.270 ± 0.124</td>
|
||
<td id="7681CE5CFFFB71FDFB1F7EC4FB6E9BA2" box="[1153,1252,953,990]" gridcol="7" gridrow="3" pageId="6" pageNumber="481">1.03‒1.39 1.224 ± 0.070</td>
|
||
<td id="7681CE5CFFFB71FDFA9E7EC4FAE99BA2" box="[1280,1379,953,990]" gridcol="8" gridrow="3" pageId="6" pageNumber="481">3.68‒4.74 4.279 ± 0.223</td>
|
||
<td id="7681CE5CFFFB71FDFA1E7EC4FA699BA2" box="[1408,1507,953,990]" gridcol="9" gridrow="3" pageId="6" pageNumber="481">4.71‒6.11 5.503 ± 0.280</td>
|
||
<td id="7681CE5CFFFB71FDF99E7EC4F9E89BA2" box="[1536,1634,953,990]" gridcol="10" gridrow="3" pageId="6" pageNumber="481">2.30‒2.54 2.425 ± 0.056</td>
|
||
<td id="7681CE5CFFFB71FDF91E7EC4F9699BA2" box="[1664,1763,953,990]" gridcol="11" gridrow="3" pageId="6" pageNumber="481">0.63‒0.74 0.687 ± 0.026</td>
|
||
<td id="7681CE5CFFFB71FDF89C7EC4F8EF9BA2" box="[1794,1893,953,990]" gridcol="12" gridrow="3" pageId="6" pageNumber="481">1.14‒1.65 1.463 ± 0.077</td>
|
||
</tr>
|
||
<tr id="3550A720FFFB71FDFFE17E8FF8EF9C6B" box="[127,1893,1010,1047]" gridrow="4" pageId="6" pageNumber="481">
|
||
<th id="7681CE5CFFFB71FDFFE17E8FFEBF9C6B" box="[127,309,1010,1047]" gridcol="0" gridrow="4" pageId="6" pageNumber="481">Ban Khoun Ngeun, Laos n = 3</th>
|
||
<td id="7681CE5CFFFB71FDFE1F7E8FFE6E9C6B" box="[385,484,1010,1047]" gridcol="1" gridrow="4" pageId="6" pageNumber="481">1.03‒1.13 1.079 ± 0.053</td>
|
||
<td id="7681CE5CFFFB71FDFD9F7E8FFDEE9C6B" box="[513,612,1010,1047]" gridcol="2" gridrow="4" pageId="6" pageNumber="481">0.50‒0.61 0.544 ± 0.055</td>
|
||
<td id="7681CE5CFFFB71FDFD1F7E8FFD6E9C6B" box="[641,740,1010,1047]" gridcol="3" gridrow="4" pageId="6" pageNumber="481">1.16‒1.24 1.193 ± 0.040</td>
|
||
<td id="7681CE5CFFFB71FDFC9F7E8FFCEE9C6B" box="[769,868,1010,1047]" gridcol="4" gridrow="4" pageId="6" pageNumber="481">1.58‒1.71 1.649 ± 0.066</td>
|
||
<td id="7681CE5CFFFB71FDFC1F7E8FFC6E9C6B" box="[897,996,1010,1047]" gridcol="5" gridrow="4" pageId="6" pageNumber="481">1.61‒1.74 1.684 ± 0.070</td>
|
||
<td id="7681CE5CFFFB71FDFB9E7E8FFBEE9C6B" box="[1024,1124,1010,1047]" gridcol="6" gridrow="4" pageId="6" pageNumber="481">2.11‒2.29 2.211 ± 0.095</td>
|
||
<td id="7681CE5CFFFB71FDFB1F7E8FFB6E9C6B" box="[1153,1252,1010,1047]" gridcol="7" gridrow="4" pageId="6" pageNumber="481">1.18‒1.24 1.211 ± 0.026</td>
|
||
<td id="7681CE5CFFFB71FDFA9E7E8FFAE99C6B" box="[1280,1379,1010,1047]" gridcol="8" gridrow="4" pageId="6" pageNumber="481">3.68‒4.42 4.053 ± 0.368</td>
|
||
<td id="7681CE5CFFFB71FDFA1E7E8FFA699C6B" box="[1408,1507,1010,1047]" gridcol="9" gridrow="4" pageId="6" pageNumber="481">4.87‒5.66 5.263 ± 0.395</td>
|
||
<td id="7681CE5CFFFB71FDF99E7E8FF9E89C6B" box="[1536,1634,1010,1047]" gridcol="10" gridrow="4" pageId="6" pageNumber="481">2.31‒2.47 2.379 ± 0.083</td>
|
||
<td id="7681CE5CFFFB71FDF91E7E8FF9699C6B" box="[1664,1763,1010,1047]" gridcol="11" gridrow="4" pageId="6" pageNumber="481">0.71‒0.74 0.719 ± 0.016</td>
|
||
<td id="7681CE5CFFFB71FDF89C7E8FF8EF9C6B" box="[1794,1893,1010,1047]" gridcol="12" gridrow="4" pageId="6" pageNumber="481">1.32‒1.35 1.331 ± 0.020</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<caption id="DF1FF5EAFFFB8E04FFE97937FC389CD1" ID-Table-UUID="DF1FF5EAFFFB8E04FFE97937FC389CD1" httpUri="http://table.plazi.org/id/DF1FF5EAFFFB8E04FFE97937FC389CD1" pageId="6" pageNumber="481" targetBox="[127,1762,1238,1582]" targetIsTable="true" targetPageId="6">
|
||
<paragraph id="8BDFA562FFFB8E04FFE97937FC389CD1" blockId="6.[119,1929,1098,1197]" pageId="6" pageNumber="481">
|
||
<emphasis id="B9147970FFFB8E04FFE97937FF549C22" bold="true" box="[119,222,1098,1118]" pageId="6" pageNumber="481">TABLE 2.</emphasis>
|
||
Variability of selected metric characters in females of
|
||
<emphasis id="B9147970FFFB8E04FD7C7937FC5A9C23" box="[738,976,1098,1119]" italics="true" pageId="6" pageNumber="481">
|
||
<taxonomicName id="4C60DEE1FFFB8E04FD7C7937FC419C23" box="[738,971,1098,1119]" class="Insecta" family="Dryopidae" genus="Parahelichus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="481" phylum="Arthropoda" rank="species" species="granulosus">Parahelichus granulosus</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
<emphasis id="B9147970FFFB8E04FC487937FC749C22" bold="true" box="[982,1022,1098,1118]" pageId="6" pageNumber="481">HW</emphasis>
|
||
– head width including eyes,
|
||
<emphasis id="B9147970FFFB8E04FA847937FAB99C22" bold="true" box="[1306,1331,1098,1118]" pageId="6" pageNumber="481">ID</emphasis>
|
||
– interocular distance,
|
||
<emphasis id="B9147970FFFB8E04F98F7937F9CC9C22" bold="true" box="[1553,1606,1098,1118]" pageId="6" pageNumber="481">APW</emphasis>
|
||
– anterior pronotal width,
|
||
<emphasis id="B9147970FFFB8E04F8DD7937F8FF9C22" bold="true" box="[1859,1909,1098,1118]" pageId="6" pageNumber="481">PPW</emphasis>
|
||
– posterior pronotal width,
|
||
<emphasis id="B9147970FFFB8E04FEFD7919FE0D9C04" bold="true" box="[355,391,1124,1144]" pageId="6" pageNumber="481">PW</emphasis>
|
||
– pronotal width, maximum width of pronotum,
|
||
<emphasis id="B9147970FFFB8E04FCCF7919FCFD9C04" bold="true" box="[849,887,1124,1144]" pageId="6" pageNumber="481">EW</emphasis>
|
||
– elytral width, maximum width combined,
|
||
<emphasis id="B9147970FFFB8E04FA887919FAC39C04" bold="true" box="[1302,1353,1124,1144]" pageId="6" pageNumber="481">MPL</emphasis>
|
||
– pronotal length along the body midline,
|
||
<emphasis id="B9147970FFFB8E04F9467919F97C9C04" bold="true" box="[1752,1782,1124,1144]" pageId="6" pageNumber="481">EL</emphasis>
|
||
– elytral length along suture,
|
||
<emphasis id="B9147970FFFB8E04FF6A7903FE999CEE" bold="true" box="[244,275,1150,1170]" pageId="6" pageNumber="481">BL</emphasis>
|
||
– body length. First rows show ranges of measured values for specimens for different localities, in second rows mean and standard deviation for the respective measurements,
|
||
<emphasis id="B9147970FFFB8E04F8F47903F8FC9CEE" bold="true" box="[1898,1910,1150,1170]" pageId="6" pageNumber="481">n</emphasis>
|
||
– number of measured specimens. HW and ID were not measured in one female from Sa Pa.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BDFA562FFFB8E04FFE179ABF9689E52" pageId="6" pageNumber="481">
|
||
<table id="F96057C2FFFB71FDFFE179ABF9689E52" box="[127,1762,1238,1582]" gridcols="12" gridrows="6" pageId="6" pageNumber="481">
|
||
<tr id="3550A720FFFB71FDFFE179ABF9689D6D" box="[127,1762,1238,1297]" gridrow="0" pageId="6" pageNumber="481">
|
||
<th id="7681CE5CFFFB71FDFFE179ABFEBF9D6D" box="[127,309,1238,1297]" gridcol="0" gridrow="0" pageId="6" pageNumber="481">
|
||
<emphasis id="B9147970FFFB8E04FFE179ABFF379C9A" bold="true" box="[127,189,1238,1254]" pageId="6" pageNumber="481">Locality</emphasis>
|
||
Yen Bai, Vietnam n = 22
|
||
</th>
|
||
<th id="7681CE5CFFFB71FDFE1F79ABFE6E9D6D" box="[385,484,1238,1297]" gridcol="1" gridrow="0" pageId="6" pageNumber="481">
|
||
<emphasis id="B9147970FFFB8E04FE1F79ABFE459C9A" bold="true" box="[385,463,1238,1254]" pageId="6" pageNumber="481">HW [mm]</emphasis>
|
||
0.87‒1.39 1.158 ± 0.098
|
||
</th>
|
||
<th id="7681CE5CFFFB71FDFD9F79ABFDEE9D6D" box="[513,612,1238,1297]" gridcol="2" gridrow="0" pageId="6" pageNumber="481">
|
||
<emphasis id="B9147970FFFB8E04FD9F79ABFDC99C9A" bold="true" box="[513,579,1238,1254]" pageId="6" pageNumber="481">ID [mm]</emphasis>
|
||
0.42‒0.79 0.644 ± 0.071
|
||
</th>
|
||
<th id="7681CE5CFFFB71FDFD1E79ABFD699D6D" box="[640,739,1238,1297]" gridcol="3" gridrow="0" pageId="6" pageNumber="481">
|
||
<emphasis id="B9147970FFFB8E04FD1E79ABFD529C9A" bold="true" box="[640,728,1238,1254]" pageId="6" pageNumber="481">APW [mm]</emphasis>
|
||
0.89‒1.53 1.293 ± 0.126
|
||
</th>
|
||
<th id="7681CE5CFFFB71FDFC9E79ABFCE99D6D" box="[768,867,1238,1297]" gridcol="4" gridrow="0" pageId="6" pageNumber="481">
|
||
<emphasis id="B9147970FFFB8E04FC9E79ABFCDC9C9A" bold="true" box="[768,854,1238,1254]" pageId="6" pageNumber="481">PPW [mm]</emphasis>
|
||
1.29‒2.13 1.762 ± 0.163
|
||
</th>
|
||
<th id="7681CE5CFFFB71FDFC1E79ABFC699D6D" box="[896,995,1238,1297]" gridcol="5" gridrow="0" pageId="6" pageNumber="481">
|
||
<emphasis id="B9147970FFFB8E04FC1E79ABFC419C9A" bold="true" box="[896,971,1238,1254]" pageId="6" pageNumber="481">PW [mm]</emphasis>
|
||
1.29‒2.21 1.822 ± 0.177
|
||
</th>
|
||
<th id="7681CE5CFFFB71FDFB9E79ABFBE99D6D" box="[1024,1123,1238,1297]" gridcol="6" gridrow="0" pageId="6" pageNumber="481">
|
||
<emphasis id="B9147970FFFB8E04FB9E79ABFBC69C9A" bold="true" box="[1024,1100,1238,1254]" pageId="6" pageNumber="481">EW [mm]</emphasis>
|
||
1.61‒2.82 2.313 ± 0.237
|
||
</th>
|
||
<th id="7681CE5CFFFB71FDFB1E79ABFB699D6D" box="[1152,1251,1238,1297]" gridcol="7" gridrow="0" pageId="6" pageNumber="481">
|
||
<emphasis id="B9147970FFFB8E04FB1E79ABFB5F9C9A" bold="true" box="[1152,1237,1238,1254]" pageId="6" pageNumber="481">MPL [mm]</emphasis>
|
||
0.92‒1.47 1.287 ± 0.121
|
||
</th>
|
||
<th id="7681CE5CFFFB71FDFA9E79ABFAE99D6D" box="[1280,1379,1238,1297]" gridcol="8" gridrow="0" pageId="6" pageNumber="481">
|
||
<emphasis id="B9147970FFFB8E04FA9E79ABFACF9C9A" bold="true" box="[1280,1349,1238,1254]" pageId="6" pageNumber="481">EL [mm]</emphasis>
|
||
3.05‒5.47 4.335 ± 0.455
|
||
</th>
|
||
<th id="7681CE5CFFFB71FDFAE179ABFA689D6D" box="[1407,1506,1238,1297]" gridcol="9" gridrow="0" pageId="6" pageNumber="481">
|
||
<emphasis id="B9147970FFFB8E04FAE179ABFA4F9C9A" bold="true" box="[1407,1477,1238,1254]" pageId="6" pageNumber="481">BL [mm]</emphasis>
|
||
3.97‒6.95 5.622 ± 0.568
|
||
</th>
|
||
<th id="7681CE5CFFFB71FDFA6179ABF9E89D6D" box="[1535,1634,1238,1297]" gridcol="10" gridrow="0" pageId="6" pageNumber="481">
|
||
<emphasis id="B9147970FFFB8E04FA6179ABF9B39C9A" bold="true" box="[1535,1593,1238,1254]" pageId="6" pageNumber="481">BL/EW</emphasis>
|
||
2.29‒2.56 2.432 ± 0.068
|
||
</th>
|
||
<th id="7681CE5CFFFB71FDF9E179ABF9689D6D" box="[1663,1762,1238,1297]" gridcol="11" gridrow="0" pageId="6" pageNumber="481">
|
||
<emphasis id="B9147970FFFB8E04F9E179ABF9429C9A" bold="true" box="[1663,1736,1238,1254]" pageId="6" pageNumber="481">MPL/PW</emphasis>
|
||
0.64‒0.77 0.707 ± 0.032
|
||
</th>
|
||
</tr>
|
||
<tr id="3550A720FFFB71FDFFE17858F9689D36" box="[127,1762,1317,1354]" gridrow="1" pageId="6" pageNumber="481">
|
||
<th id="7681CE5CFFFB71FDFFE17858FEBF9D36" box="[127,309,1317,1354]" gridcol="0" gridrow="1" pageId="6" pageNumber="481">Hoa binh, Vietnam n = 3</th>
|
||
<td id="7681CE5CFFFB71FDFE1F7858FE6E9D36" box="[385,484,1317,1354]" gridcol="1" gridrow="1" pageId="6" pageNumber="481">1.16‒1.26 1.202 ± 0.055</td>
|
||
<td id="7681CE5CFFFB71FDFD9F7858FDEE9D36" box="[513,612,1317,1354]" gridcol="2" gridrow="1" pageId="6" pageNumber="481">0.66‒0.66 0.658 ± 0.000</td>
|
||
<td id="7681CE5CFFFB71FDFD1E7858FD699D36" box="[640,739,1317,1354]" gridcol="3" gridrow="1" pageId="6" pageNumber="481">1.26‒1.34 1.298 ± 0.040</td>
|
||
<td id="7681CE5CFFFB71FDFC9E7858FCE99D36" box="[768,867,1317,1354]" gridcol="4" gridrow="1" pageId="6" pageNumber="481">1.68‒1.87 1.789 ± 0.095</td>
|
||
<td id="7681CE5CFFFB71FDFC1E7858FC699D36" box="[896,995,1317,1354]" gridcol="5" gridrow="1" pageId="6" pageNumber="481">1.82‒1.95 1.886 ± 0.066</td>
|
||
<td id="7681CE5CFFFB71FDFB9E7858FBE99D36" box="[1024,1123,1317,1354]" gridcol="6" gridrow="1" pageId="6" pageNumber="481">2.42‒2.47 2.439 ± 0.030</td>
|
||
<td id="7681CE5CFFFB71FDFB1E7858FB699D36" box="[1152,1251,1317,1354]" gridcol="7" gridrow="1" pageId="6" pageNumber="481">1.32‒1.42 1.368 ± 0.053</td>
|
||
<td id="7681CE5CFFFB71FDFA9E7858FAE99D36" box="[1280,1379,1317,1354]" gridcol="8" gridrow="1" pageId="6" pageNumber="481">4.37‒4.58 4.456 ± 0.110</td>
|
||
<td id="7681CE5CFFFB71FDFAE17858FA689D36" box="[1407,1506,1317,1354]" gridcol="9" gridrow="1" pageId="6" pageNumber="481">5.74‒5.89 5.825 ± 0.080</td>
|
||
<td id="7681CE5CFFFB71FDFA617858F9E89D36" box="[1535,1634,1317,1354]" gridcol="10" gridrow="1" pageId="6" pageNumber="481">2.36‒2.43 2.389 ± 0.040</td>
|
||
<td id="7681CE5CFFFB71FDF9E17858F9689D36" box="[1663,1762,1317,1354]" gridcol="11" gridrow="1" pageId="6" pageNumber="481">0.68‒0.75 0.726 ± 0.044</td>
|
||
</tr>
|
||
<tr id="3550A720FFFB71FDFFE17822F9689DFF" box="[127,1762,1375,1411]" gridrow="2" pageId="6" pageNumber="481">
|
||
<th id="7681CE5CFFFB71FDFFE17822FEBF9DFF" box="[127,309,1375,1411]" gridcol="0" gridrow="2" pageId="6" pageNumber="481">Sa Pa (Lao Cai), Vietnam n = 92</th>
|
||
<td id="7681CE5CFFFB71FDFE1F7822FE6E9DFF" box="[385,484,1375,1411]" gridcol="1" gridrow="2" pageId="6" pageNumber="481">1.05‒1.39 1.214 ± 0.076</td>
|
||
<td id="7681CE5CFFFB71FDFD9F7822FDEE9DFF" box="[513,612,1375,1411]" gridcol="2" gridrow="2" pageId="6" pageNumber="481">0.55‒0.82 0.678 ± 0.049</td>
|
||
<td id="7681CE5CFFFB71FDFD1E7822FD699DFF" box="[640,739,1375,1411]" gridcol="3" gridrow="2" pageId="6" pageNumber="481">1.16‒1.55 1.330 ± 0.086</td>
|
||
<td id="7681CE5CFFFB71FDFC9E7822FCE99DFF" box="[768,867,1375,1411]" gridcol="4" gridrow="2" pageId="6" pageNumber="481">1.37‒2.24 1.844 ± 0.145</td>
|
||
<td id="7681CE5CFFFB71FDFC1E7822FC699DFF" box="[896,995,1375,1411]" gridcol="5" gridrow="2" pageId="6" pageNumber="481">1.61‒2.24 1.900 ± 0.128</td>
|
||
<td id="7681CE5CFFFB71FDFB9E7822FBE99DFF" box="[1024,1123,1375,1411]" gridcol="6" gridrow="2" pageId="6" pageNumber="481">2.11‒2.95 2.503 ± 0.164</td>
|
||
<td id="7681CE5CFFFB71FDFB1E7822FB699DFF" box="[1152,1251,1375,1411]" gridcol="7" gridrow="2" pageId="6" pageNumber="481">1.18‒1.55 1.377 ± 0.093</td>
|
||
<td id="7681CE5CFFFB71FDFA9E7822FAE99DFF" box="[1280,1379,1375,1411]" gridcol="8" gridrow="2" pageId="6" pageNumber="481">3.95‒5.21 4.533 ± 0.300</td>
|
||
<td id="7681CE5CFFFB71FDFAE17822FA689DFF" box="[1407,1506,1375,1411]" gridcol="9" gridrow="2" pageId="6" pageNumber="481">5.13‒6.76 5.910 ± 0.379</td>
|
||
<td id="7681CE5CFFFB71FDFA617822F9E89DFF" box="[1535,1634,1375,1411]" gridcol="10" gridrow="2" pageId="6" pageNumber="481">2.23‒2.51 2.362 ± 0.065</td>
|
||
<td id="7681CE5CFFFB71FDF9E17822F9689DFF" box="[1663,1762,1375,1411]" gridcol="11" gridrow="2" pageId="6" pageNumber="481">0.66‒0.80 0.725 ± 0.030</td>
|
||
</tr>
|
||
<tr id="3550A720FFFB71FDFFE178E5F9689DC0" box="[127,1762,1432,1468]" gridrow="3" pageId="6" pageNumber="481">
|
||
<th id="7681CE5CFFFB71FDFFE178E5FEBF9DC0" box="[127,309,1432,1468]" gridcol="0" gridrow="3" pageId="6" pageNumber="481">Louang Namtha, Laos n = 17</th>
|
||
<td id="7681CE5CFFFB71FDFE1F78E5FE6E9DC0" box="[385,484,1432,1468]" gridcol="1" gridrow="3" pageId="6" pageNumber="481">0.87‒1.32 1.107 ± 0.145</td>
|
||
<td id="7681CE5CFFFB71FDFD9F78E5FDEE9DC0" box="[513,612,1432,1468]" gridcol="2" gridrow="3" pageId="6" pageNumber="481">0.42‒0.76 0.598 ± 0.117</td>
|
||
<td id="7681CE5CFFFB71FDFD1E78E5FD699DC0" box="[640,739,1432,1468]" gridcol="3" gridrow="3" pageId="6" pageNumber="481">0.97‒1.47 1.231 ± 0.157</td>
|
||
<td id="7681CE5CFFFB71FDFC9E78E5FCE99DC0" box="[768,867,1432,1468]" gridcol="4" gridrow="3" pageId="6" pageNumber="481">1.32‒2.11 1.684 ± 0.224</td>
|
||
<td id="7681CE5CFFFB71FDFC1E78E5FC699DC0" box="[896,995,1432,1468]" gridcol="5" gridrow="3" pageId="6" pageNumber="481">1.37‒2.13 1.752 ± 0.231</td>
|
||
<td id="7681CE5CFFFB71FDFB9E78E5FBE99DC0" box="[1024,1123,1432,1468]" gridcol="6" gridrow="3" pageId="6" pageNumber="481">1.71‒2.66 2.259 ± 0.296</td>
|
||
<td id="7681CE5CFFFB71FDFB1E78E5FB699DC0" box="[1152,1251,1432,1468]" gridcol="7" gridrow="3" pageId="6" pageNumber="481">1.00‒1.50 1.224 ± 0.134</td>
|
||
<td id="7681CE5CFFFB71FDFA9E78E5FAE99DC0" box="[1280,1379,1432,1468]" gridcol="8" gridrow="3" pageId="6" pageNumber="481">3.05‒4.84 3.978 ± 0.617</td>
|
||
<td id="7681CE5CFFFB71FDFAE178E5FA689DC0" box="[1407,1506,1432,1468]" gridcol="9" gridrow="3" pageId="6" pageNumber="481">4.05‒6.24 5.203 ± 0.741</td>
|
||
<td id="7681CE5CFFFB71FDFA6178E5F9E89DC0" box="[1535,1634,1432,1468]" gridcol="10" gridrow="3" pageId="6" pageNumber="481">2.21‒2.44 2.301 ± 0.061</td>
|
||
<td id="7681CE5CFFFB71FDF9E178E5F9689DC0" box="[1663,1762,1432,1468]" gridcol="11" gridrow="3" pageId="6" pageNumber="481">0.63‒0.74 0.701 ± 0.033</td>
|
||
</tr>
|
||
<tr id="3550A720FFFB71FDFFE178ACF9689D89" box="[127,1762,1489,1525]" gridrow="4" pageId="6" pageNumber="481">
|
||
<th id="7681CE5CFFFB71FDFFE178ACFEBF9D89" box="[127,309,1489,1525]" gridcol="0" gridrow="4" pageId="6" pageNumber="481">Khan riv., Laos n = 86</th>
|
||
<td id="7681CE5CFFFB71FDFE1F78ACFE6E9D89" box="[385,484,1489,1525]" gridcol="1" gridrow="4" pageId="6" pageNumber="481">1.05‒1.42 1.237 ± 0.072</td>
|
||
<td id="7681CE5CFFFB71FDFD9F78ACFDEE9D89" box="[513,612,1489,1525]" gridcol="2" gridrow="4" pageId="6" pageNumber="481">0.53‒0.82 0.670 ± 0.053</td>
|
||
<td id="7681CE5CFFFB71FDFD1E78ACFD699D89" box="[640,739,1489,1525]" gridcol="3" gridrow="4" pageId="6" pageNumber="481">1.18‒1.55 1.356 ± 0.087</td>
|
||
<td id="7681CE5CFFFB71FDFC9E78ACFCE99D89" box="[768,867,1489,1525]" gridcol="4" gridrow="4" pageId="6" pageNumber="481">1.58‒2.21 1.887 ± 0.134</td>
|
||
<td id="7681CE5CFFFB71FDFC1E78ACFC699D89" box="[896,995,1489,1525]" gridcol="5" gridrow="4" pageId="6" pageNumber="481">1.66‒2.24 1.946 ± 0.133</td>
|
||
<td id="7681CE5CFFFB71FDFB9E78ACFBE99D89" box="[1024,1123,1489,1525]" gridcol="6" gridrow="4" pageId="6" pageNumber="481">2.16‒2.79 2.510 ± 0.138</td>
|
||
<td id="7681CE5CFFFB71FDFB1E78ACFB699D89" box="[1152,1251,1489,1525]" gridcol="7" gridrow="4" pageId="6" pageNumber="481">1.16‒1.53 1.338 ± 0.081</td>
|
||
<td id="7681CE5CFFFB71FDFA9E78ACFAE99D89" box="[1280,1379,1489,1525]" gridcol="8" gridrow="4" pageId="6" pageNumber="481">4.00‒5.21 4.618 ± 0.264</td>
|
||
<td id="7681CE5CFFFB71FDFAE178ACFA689D89" box="[1407,1506,1489,1525]" gridcol="9" gridrow="4" pageId="6" pageNumber="481">5.24‒6.74 5.956 ± 0.330</td>
|
||
<td id="7681CE5CFFFB71FDFA6178ACF9E89D89" box="[1535,1634,1489,1525]" gridcol="10" gridrow="4" pageId="6" pageNumber="481">2.17‒2.50 2.373 ± 0.062</td>
|
||
<td id="7681CE5CFFFB71FDF9E178ACF9689D89" box="[1663,1762,1489,1525]" gridcol="11" gridrow="4" pageId="6" pageNumber="481">0.62‒0.76 0.688 ± 0.031</td>
|
||
</tr>
|
||
<tr id="3550A720FFFB71FDFFE17B77F9689E52" box="[127,1762,1546,1582]" gridrow="5" pageId="6" pageNumber="481">
|
||
<th id="7681CE5CFFFB71FDFFE17B77FEBF9E52" box="[127,309,1546,1582]" gridcol="0" gridrow="5" pageId="6" pageNumber="481">Ban Khoun Ngeun, Laos n = 5</th>
|
||
<td id="7681CE5CFFFB71FDFE1F7B77FE6E9E52" box="[385,484,1546,1582]" gridcol="1" gridrow="5" pageId="6" pageNumber="481">1.08‒1.21 1.137 ± 0.051</td>
|
||
<td id="7681CE5CFFFB71FDFD9F7B77FDEE9E52" box="[513,612,1546,1582]" gridcol="2" gridrow="5" pageId="6" pageNumber="481">0.58‒0.66 0.632 ± 0.032</td>
|
||
<td id="7681CE5CFFFB71FDFD1E7B77FD699E52" box="[640,739,1546,1582]" gridcol="3" gridrow="5" pageId="6" pageNumber="481">1.18‒1.34 1.284 ± 0.063</td>
|
||
<td id="7681CE5CFFFB71FDFC9E7B77FCE99E52" box="[768,867,1546,1582]" gridcol="4" gridrow="5" pageId="6" pageNumber="481">1.58‒1.89 1.711 ± 0.121</td>
|
||
<td id="7681CE5CFFFB71FDFC1E7B77FC699E52" box="[896,995,1546,1582]" gridcol="5" gridrow="5" pageId="6" pageNumber="481">1.63‒2.00 1.774 ± 0.147</td>
|
||
<td id="7681CE5CFFFB71FDFB9E7B77FBE99E52" box="[1024,1123,1546,1582]" gridcol="6" gridrow="5" pageId="6" pageNumber="481">2.16‒2.68 2.363 ± 0.212</td>
|
||
<td id="7681CE5CFFFB71FDFB1E7B77FB699E52" box="[1152,1251,1546,1582]" gridcol="7" gridrow="5" pageId="6" pageNumber="481">1.21‒1.45 1.332 ± 0.088</td>
|
||
<td id="7681CE5CFFFB71FDFA9E7B77FAE99E52" box="[1280,1379,1546,1582]" gridcol="8" gridrow="5" pageId="6" pageNumber="481">4.16‒4.74 4.358 ± 0.245</td>
|
||
<td id="7681CE5CFFFB71FDFAE17B77FA689E52" box="[1407,1506,1546,1582]" gridcol="9" gridrow="5" pageId="6" pageNumber="481">5.37‒6.18 5.689 ± 0.321</td>
|
||
<td id="7681CE5CFFFB71FDFA617B77F9E89E52" box="[1535,1634,1546,1582]" gridcol="10" gridrow="5" pageId="6" pageNumber="481">2.30‒2.52 2.414 ± 0.089</td>
|
||
<td id="7681CE5CFFFB71FDF9E17B77F9689E52" box="[1663,1762,1546,1582]" gridcol="11" gridrow="5" pageId="6" pageNumber="481">0.72‒0.79 0.752 ± 0.027</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<caption id="DF1FF5EAFFFA8E05FF097AE3FDA39F8E" httpUri="https://zenodo.org/record/231189/files/figure.png" pageId="7" pageNumber="482" targetBox="[217,1367,201,1913]" targetPageId="7">
|
||
<paragraph id="8BDFA562FFFA8E05FF097AE3FDA39F8E" blockId="7.[151,1436,1950,2034]" pageId="7" pageNumber="482">
|
||
<emphasis id="B9147970FFFA8E05FF097AE3FEC09FC8" bold="true" box="[151,330,1950,1973]" pageId="7" pageNumber="482">FIGURES 3–13.</emphasis>
|
||
<taxonomicName id="4C60DEE1FFFA8E05FECD7AE3FDDE9FC8" box="[339,596,1950,1972]" class="Insecta" family="Dryopidae" genus="Parahelichus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="482" phylum="Arthropoda" rank="species" species="granulosus">
|
||
<emphasis id="B9147970FFFA8E05FECD7AE3FDDE9FC8" box="[339,596,1950,1972]" italics="true" pageId="7" pageNumber="482">Parahelichus granulosus</emphasis>
|
||
</taxonomicName>
|
||
, male; (3–5, 8–13) aedeagus, penis and parameres in lateral, ventral and dorsal view; figures show differences in proportions between parameres and phallobase; (6) male sternite VIII, ventral view; (7) male sternite IX, ventral view. Scale 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF1FF5EAFFF58E0AFF097A3EFEEF9FEA" httpUri="https://zenodo.org/record/231190/files/figure.png" pageId="8" pageNumber="483" targetBox="[196,1391,199,1829]" targetPageId="8">
|
||
<paragraph id="8BDFA562FFF58E0AFF097A3EFEEF9FEA" blockId="8.[151,1436,1859,1942]" pageId="8" pageNumber="483">
|
||
<emphasis id="B9147970FFF58E0AFF097A3EFEDF9F24" bold="true" box="[151,341,1859,1882]" pageId="8" pageNumber="483">FIGURES 14–16.</emphasis>
|
||
<taxonomicName id="4C60DEE1FFF58E0AFEC27A3EFDD19F25" box="[348,603,1859,1881]" class="Insecta" family="Dryopidae" genus="Parahelichus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="483" phylum="Arthropoda" rank="species" species="granulosus">
|
||
<emphasis id="B9147970FFF58E0AFEC27A3EFDD19F25" box="[348,603,1859,1881]" italics="true" pageId="8" pageNumber="483">Parahelichus granulosus</emphasis>
|
||
</taxonomicName>
|
||
, female; (14‒15) ovipositor, lateral view: (14) ovipositor of a specimen from the locality Louang Namtha, Laos, (15) a specimen from the locality Sa Pa (Lao Cai), Vietnam; (16) female sternite VIII, ventral view. Scale 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF1FF5EAFFF48E0BFF097932FBB29CF8" httpUri="https://zenodo.org/record/231191/files/figure.png" pageId="9" pageNumber="484" targetBox="[238,1350,196,1075]" targetPageId="9">
|
||
<paragraph id="8BDFA562FFF48E0BFF097932FBB29CF8" blockId="9.[151,1436,1103,1156]" pageId="9" pageNumber="484">
|
||
<emphasis id="B9147970FFF48E0BFF097932FEDE9C18" bold="true" box="[151,340,1103,1126]" pageId="9" pageNumber="484">FIGURES 17–20.</emphasis>
|
||
<taxonomicName id="4C60DEE1FFF48E0BFEC57932FDD39C19" box="[347,601,1103,1125]" class="Insecta" family="Dryopidae" genus="Parahelichus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="484" phylum="Arthropoda" rank="species" species="granulosus">
|
||
<emphasis id="B9147970FFF48E0BFEC57932FDD39C19" box="[347,601,1103,1125]" italics="true" pageId="9" pageNumber="484">Parahelichus granulosus</emphasis>
|
||
</taxonomicName>
|
||
; (17‒18) female antennal club: (17) ventral view, (18) dorsal view, scale 0.1 mm; (19) abdomen of a male, ventral view; (20) abdomen of a female, ventral view; scale 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BDFA562FFF48E0BFF5979D6FF529D04" blockId="9.[151,1436,1195,2012]" pageId="9" pageNumber="484">
|
||
Head wider than long; surface shiny with sparse granules and fine punctures; granules distinctly smaller than a facet diameter, associated with yellowish recumbent setae; granules and punctures irregularly spaced, more densely arranged on lateral portion of clypeus. Frontal region with a small shiny longitudinal area. Eyes large, protuberant, strongly facetted; interfacetal setae fine, scattered, confined to central portion. ID in ♂♂:
|
||
<quantity id="4C980887FFF48E0BFB02786AFACB9D53" box="[1180,1345,1303,1327]" metricMagnitude="-4" metricUnit="m" metricValue="6.55" metricValueMax="7.6" metricValueMin="5.5" pageId="9" pageNumber="484" unit="mm" value="0.655" valueMax="0.76" valueMin="0.55">0.55–0.76 mm</quantity>
|
||
(0.62 ± 0.12), in ♀♀:
|
||
<quantity id="4C980887FFF48E0BFEAA7846FE5C9D2F" box="[308,470,1339,1363]" metricMagnitude="-4" metricUnit="m" metricValue="6.85" metricValueMax="7.9" metricValueMin="5.8" pageId="9" pageNumber="484" unit="mm" value="0.685" valueMax="0.79" valueMin="0.58">0.58–0.79 mm</quantity>
|
||
(0.66 ± 0.10); HW in ♂♂:
|
||
<quantity id="4C980887FFF48E0BFC997846FC209D2F" box="[775,938,1339,1363]" metricMagnitude="-3" metricUnit="m" metricValue="1.185" metricValueMax="1.32" metricValueMin="1.05" pageId="9" pageNumber="484" unit="mm" value="1.185" valueMax="1.32" valueMin="1.05">1.05–1.32 mm</quantity>
|
||
(1.14 ± 0.15), in ♀♀:
|
||
<quantity id="4C980887FFF48E0BFB3A7841FACC9D2F" box="[1188,1350,1340,1364]" metricMagnitude="-3" metricUnit="m" metricValue="1.24" metricValueMax="1.37" metricValueMin="1.11" pageId="9" pageNumber="484" unit="mm" value="1.24" valueMax="1.37" valueMin="1.11">1.11–1.37 mm</quantity>
|
||
(1.23 ± 0.12).
|
||
</paragraph>
|
||
<paragraph id="8BDFA562FFF48E09FF5978FEFC7F9FF4" blockId="9.[151,1436,1195,2012]" lastBlockId="11.[151,1437,1724,2037]" lastPageId="11" lastPageNumber="486" pageId="9" pageNumber="484">
|
||
Thorax. Pronotum wider than long; disc convex dorsally; larger granules more distinct than those on head, separated usually by distance of more or less one granule diameter. Lateral pronotal carina distinct; sides moderately arcuate to sinuate; posterior angles nearly rectangular, moderately acute; anterior angles acute, deflected and strongly protruding; posterior margin trisinuate, smooth; prescutellar foveae absent. MPL in ♂♂:
|
||
<quantity id="4C980887FFF48E0BFF097B69FEB19E57" box="[151,315,1556,1580]" metricMagnitude="-3" metricUnit="m" metricValue="1.2999999999999998" metricValueMax="1.47" metricValueMin="1.13" pageId="9" pageNumber="484" unit="mm" value="1.2999999999999998" valueMax="1.47" valueMin="1.13">1.13–1.47 mm</quantity>
|
||
(1.25 ± 0.19), in ♀♀:
|
||
<quantity id="4C980887FFF48E0BFDDC7B6EFD6C9E57" box="[578,742,1555,1579]" metricMagnitude="-3" metricUnit="m" metricValue="1.33" metricValueMax="1.45" metricValueMin="1.21" pageId="9" pageNumber="484" unit="mm" value="1.33" valueMax="1.45" valueMin="1.21">1.21–1.45 mm</quantity>
|
||
(1.32 ± 0.10); APW in ♂♂:
|
||
<quantity id="4C980887FFF48E0BFBAA7B6EFB539E57" box="[1076,1241,1555,1579]" metricMagnitude="-3" metricUnit="m" metricValue="1.315" metricValueMax="1.45" metricValueMin="1.18" pageId="9" pageNumber="484" unit="mm" value="1.315" valueMax="1.45" valueMin="1.18">1.18–1.45 mm</quantity>
|
||
(1.28 ± 0.14), in ♀♀:
|
||
<quantity id="4C980887FFF48E0BFF4E7B4AFEFE9E33" box="[208,372,1591,1615]" metricMagnitude="-3" metricUnit="m" metricValue="1.37" metricValueMax="1.53" metricValueMin="1.21" pageId="9" pageNumber="484" unit="mm" value="1.37" valueMax="1.53" valueMin="1.21">1.21–1.53 mm</quantity>
|
||
(1.37 ± 0.14); PPW in ♂♂:
|
||
<quantity id="4C980887FFF48E0BFD287B4AFCD29E33" box="[694,856,1591,1615]" metricMagnitude="-3" metricUnit="m" metricValue="1.845" metricValueMax="2.11" metricValueMin="1.58" pageId="9" pageNumber="484" unit="mm" value="1.845" valueMax="2.11" valueMin="1.58">1.58–2.11 mm</quantity>
|
||
(1.82 ± 0.27), in ♀♀:
|
||
<quantity id="4C980887FFF48E0BFBC67B4AFB709E33" box="[1112,1274,1591,1616]" metricMagnitude="-3" metricUnit="m" metricValue="1.845" metricValueMax="2.11" metricValueMin="1.58" pageId="9" pageNumber="484" unit="mm" value="1.845" valueMax="2.11" valueMin="1.58">1.58–2.11 mm</quantity>
|
||
(1.86 ± 0.28). Hypomeron widest near middle, distinctly narrowed anteriorly and slightly posteriorly; separated from prosternum by a gap along anterior third. Prosternum in front of coxae approximately as long as prosternal process, anterolateral portion rounded. Setae in middle of prosternum and on prosternal process longer than those on dorsal surface, longest setae in middle of metaventrite. Prosternal process wide, strongly overlapping mesoventrite, apex subacute; sides raised. Scutellum (scutellar shield) large, broadly rounded, in same level with elytra, surface granulated. Metaventrite distinctly longer than prosternum in front of coxae; discrimen distinct along entire length, in depression; transverse suture fine. Elytra convex dorsally; lateral margin not explanate; gradually extended posteriad from shoulders up to 0.75 of their length; shoulders rounded, prominent; elytral apices nearly acute; granulation on elytra similar to that on pronotum but less dense, granules more or less regularly spaced; epipleura widest basally, inflected at level of metacoxa and near apex, partly concealed by lateral margin of ventrites. EW in ♂♂:
|
||
<quantity id="4C980887FFF48E0BFF517AB9FEF89FA7" box="[207,370,1988,2012]" metricMagnitude="-3" metricUnit="m" metricValue="2.45" metricValueMax="2.66" metricValueMin="2.24" pageId="9" pageNumber="484" unit="mm" value="2.45" valueMax="2.66" valueMin="2.24">2.24–2.66 mm</quantity>
|
||
(2.39 ± 0.24), in ♀♀:
|
||
<quantity id="4C980887FFF48E0BFDEB7AB9FC929FA7" box="[629,792,1988,2012]" metricMagnitude="-3" metricUnit="m" metricValue="2.565" metricValueMax="2.89" metricValueMin="2.24" pageId="9" pageNumber="484" unit="mm" value="2.565" valueMax="2.89" valueMin="2.24">2.24–2.89 mm</quantity>
|
||
(2.53 ± 0.35); EL in ♂♂:
|
||
<quantity id="4C980887FFF48E0BFBA37ABEFB6A9FA7" box="[1085,1248,1987,2012]" metricMagnitude="-3" metricUnit="m" metricValue="4.605" metricValueMax="5.16" metricValueMin="4.05" pageId="9" pageNumber="484" unit="mm" value="4.605" valueMax="5.16" valueMin="4.05">4.05–5.16 mm</quantity>
|
||
(4.46 ± 0.61), in ♀♀:
|
||
<quantity id="4C980887FFF68E09FF4F7BC1FEFE9EA8" box="[209,372,1724,1749]" metricMagnitude="-3" metricUnit="m" metricValue="4.685" metricValueMax="5.26" metricValueMin="4.11" pageId="11" pageNumber="486" unit="mm" value="4.685" valueMax="5.26" valueMin="4.11">4.11–5.26 mm</quantity>
|
||
(4.57 ± 0.57). Legs slightly longer than elytra; femora strongly widened, widest near middle, with scattered fine punctures. Tibiae slightly longer than femora, apically widened: especially apex of pro- and metatibia; each tibia weakly curved, narrow. Tarsi approximately as long as protibia and mesotibia, respectively; slightly shorter than metatibia, with few short strong setae ventrally and longer setae mainly dorsally and laterally; terminal tarsomere of metatibia slightly shorter than combined lengths of tarsomeres 1–4, those on pro- and mesotibia approximately as long as combined lengths of tarsomeres 1–4.
|
||
</paragraph>
|
||
<caption id="DF1FF5EAFFF78E08FF097A0BFC2C9F95" httpUri="https://zenodo.org/record/231192/files/figure.png" pageId="10" pageNumber="485" targetBox="[249,1346,182,1891]" targetPageId="10">
|
||
<paragraph id="8BDFA562FFF78E08FF097A0BFC2C9F95" blockId="10.[151,1436,1910,2025]" pageId="10" pageNumber="485">
|
||
<emphasis id="B9147970FFF78E08FF097A0BFEDC9FF0" bold="true" box="[151,342,1910,1933]" pageId="10" pageNumber="485">FIGURES 21–31.</emphasis>
|
||
<taxonomicName id="4C60DEE1FFF78E08FEC07A0BFD2F9FF0" box="[350,677,1910,1932]" class="Insecta" family="Dryopidae" genus="Parahelichus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="485" phylum="Arthropoda" rank="species" species="pseudogranulosus" status="sp. nov.">
|
||
<emphasis id="B9147970FFF78E08FEC07A0BFD2F9FF0" box="[350,677,1910,1932]" italics="true" pageId="10" pageNumber="485">Parahelichus pseudogranulosus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A227C40BFFF78E08FD337A0AFC8A9FF0" box="[685,768,1911,1932]" pageId="10" pageNumber="485" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, male; (21‒23, 26–31) aedeagus, penis and parameres in lateral, ventral and dorsal view; (21, 26, 27) a male from the locality Mae Hong Son, Ban Si Lang, Thailand; (22, 28, 29) a male from the Bolikhamxai province, Laos; (23, 30, 31) a male from the Khammouan province, Ban Khoun Ngeun env., Laos; (24) male sternite VIII, ventral view; (25) male sternite IX, ventral view. Scale 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF1FF5EAFFF68E09FF097B41FF639E0D" httpUri="https://zenodo.org/record/231193/files/figure.png" pageId="11" pageNumber="486" targetBox="[172,1412,180,1572]" targetPageId="11">
|
||
<paragraph id="8BDFA562FFF68E09FF097B41FF639E0D" blockId="11.[151,1436,1596,1649]" pageId="11" pageNumber="486">
|
||
<emphasis id="B9147970FFF68E09FF097B41FEAB9E2D" bold="true" box="[151,289,1596,1618]" pageId="11" pageNumber="486">FIGURE 32.</emphasis>
|
||
Geographical distribution of
|
||
<taxonomicName id="4C60DEE1FFF68E09FDCD7B41FCC39E2E" box="[595,841,1596,1618]" class="Insecta" family="Dryopidae" genus="Parahelichus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="486" phylum="Arthropoda" rank="species" species="granulosu">
|
||
<emphasis id="B9147970FFF68E09FDCD7B41FCC39E2E" box="[595,841,1596,1618]" italics="true" pageId="11" pageNumber="486">Parahelichus granulosu</emphasis>
|
||
</taxonomicName>
|
||
s (black squares) and
|
||
<taxonomicName id="4C60DEE1FFF68E09FBB57B40FB779E2E" box="[1067,1277,1596,1618]" class="Insecta" family="Dryopidae" genus="Parahelichus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="486" phylum="Arthropoda" rank="species" species="pseudogranulosus" status="sp. nov.">
|
||
<emphasis id="B9147970FFF68E09FBB57B40FB779E2E" box="[1067,1277,1596,1618]" italics="true" pageId="11" pageNumber="486">P. pseudogranulosus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A227C40BFFF68E09FA9B7B40FADC9E2E" box="[1285,1366,1597,1618]" pageId="11" pageNumber="486" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(black circles).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BDFA562FFF68E0EFF597AE9FC4D9961" blockId="11.[151,1437,1724,2037]" lastBlockId="12.[151,1437,151,1040]" lastPageId="12" lastPageNumber="487" pageId="11" pageNumber="486">Abdomen with ventrite 1 longest, ventrite 5 posteriorly slightly excised in males and rounded in females. Segments VIII and IX sexually dimorphic. Male segment VIII: sternite gently rounded posteriorly, not emarginated, distinctly shorter than ventrite 5, with short median symmetrical process anteriorly; tergite more strongly sclerotized laterally than mesally, rounded posteriorly. Female segment VIII: sternite posteriorly subtriangular, sclerotized laterally, with anterior median process approximately half as long as abdomen; tergite similar to that of male. Male sternite IX with anterior median process nearly as long as ventrites 4–5 combined, process narrow and long; female sternite IX forming a part of ovipositor.</paragraph>
|
||
<paragraph id="8BDFA562FFF18E0EFF597C5AFEB69A64" blockId="12.[151,1437,151,1040]" pageId="12" pageNumber="487">
|
||
Aedeagus (
|
||
<figureCitation id="135BB9E7FFF18E0EFED37C5AFDBE993C" box="[333,564,295,320]" captionStart="FIGURES 21 – 31" captionStartId="10.[151,264,1910,1933]" captionTargetBox="[249,1346,182,1891]" captionTargetId="figure@10.[241,1346,167,1903]" captionTargetPageId="10" captionText="FIGURES 21 – 31. Parahelichus pseudogranulosus sp. nov., male; (21 ‒ 23, 26 – 31) aedeagus, penis and parameres in lateral, ventral and dorsal view; (21, 26, 27) a male from the locality Mae Hong Son, Ban Si Lang, Thailand; (22, 28, 29) a male from the Bolikhamxai province, Laos; (23, 30, 31) a male from the Khammouan province, Ban Khoun Ngeun env., Laos; (24) male sternite VIII, ventral view; (25) male sternite IX, ventral view. Scale 0.1 mm." httpUri="https://zenodo.org/record/231192/files/figure.png" pageId="12" pageNumber="487">Figs 21–23, 26–31</figureCitation>
|
||
), with phallobase narrow, very long, subcylindrical, moderately curved, 2.00–2.22 (2.15 ± 0.13) times as long as parameres, almost as wide basally as apically (lateral view), with basal orifice asymmetrical, large; parameres individually articulated with phallobase, widest basally, short, slightly curved ventrad and gradually narrowed toward apices; apices rounded in lateral and ventral view. Penis narrow, symmetrical, moderately shorter than parameres, widest basally, gradually narrowed apically; apex subacute (ventral view); membranous ventral sac with fine longitudinal furrows; fibula narrow, reaching middle of penis (ventral view).
|
||
</paragraph>
|
||
<paragraph id="8BDFA562FFF18E0EFF597F59FEF19A1C" blockId="12.[151,1437,151,1040]" pageId="12" pageNumber="487">Ovipositor about as long as ventrites 3–5 combined, strongly sclerotized. Bursa copulatrix tubular, without sclerotized spinules.</paragraph>
|
||
<paragraph id="8BDFA562FFF18E0EFF597F11FA139AD4" blockId="12.[151,1437,151,1040]" pageId="12" pageNumber="487">
|
||
<emphasis id="B9147970FFF18E0EFF597F11FE369AF9" bold="true" box="[199,444,620,645]" pageId="12" pageNumber="487">Sexual dimorphism.</emphasis>
|
||
Intersexual differences in metric characters were not found, as their values overlap between males and females. Females are recognized only on the basis of the shape of the apex of the fifth ventrite.
|
||
</paragraph>
|
||
<paragraph id="8BDFA562FFF18E0EFF597FC9FA179B44" blockId="12.[151,1437,151,1040]" pageId="12" pageNumber="487">
|
||
<emphasis id="B9147970FFF18E0EFF597FC9FEC79AB1" bold="true" box="[199,333,692,717]" pageId="12" pageNumber="487">Variability.</emphasis>
|
||
Specimens vary in size and arrangement of larger flat granules on surface. In the
|
||
<typeStatus id="54DB1BC0FFF18E0EFA997FC9FAE69AB1" box="[1287,1388,692,717]" pageId="12" pageNumber="487" type="holotype">holotype</typeStatus>
|
||
, the granules on pronotum and elytra are distinctly smaller than facet and are separated by a distance larger than one facet diameter. On the other hand, the granules of
|
||
<typeStatus id="54DB1BC0FFF18E0EFD4B7F80FCCA9B69" box="[725,832,765,789]" pageId="12" pageNumber="487" type="paratype">paratypes</typeStatus>
|
||
are subequal to the diameter of a facet and are more densely arranged. The pronotal form varies moderately in curvature of sides from evenly arcuate to slightly sinuate.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37AF6E9FFF18E0EFF597E39FF7B9F92" pageId="12" pageNumber="487" type="distribution">
|
||
<paragraph id="8BDFA562FFF18E0EFF597E39FDA09BFC" blockId="12.[151,1437,151,1040]" pageId="12" pageNumber="487">
|
||
<emphasis id="B9147970FFF18E0EFF597E39FEEA9B21" bold="true" box="[199,352,836,861]" pageId="12" pageNumber="487">Distribution.</emphasis>
|
||
As yet known from
|
||
<collectingCountry id="F377E5F2FFF18E0EFDD07E38FD0F9B20" box="[590,645,837,860]" name="Laos" pageId="12" pageNumber="487">Laos</collectingCountry>
|
||
and
|
||
<collectingCountry id="F377E5F2FFF18E0EFD207E39FCA89B21" box="[702,802,836,861]" name="Thailand" pageId="12" pageNumber="487">Thailand</collectingCountry>
|
||
(
|
||
<figureCitation id="135BB9E7FFF18E0EFCAD7E39FC029B20" box="[819,904,836,861]" captionStart="FIGURE 32" captionStartId="11.[151,250,1596,1618]" captionTargetBox="[172,1412,180,1572]" captionTargetId="figure@11.[166,1420,177,1579]" captionTargetPageId="11" captionText="FIGURE 32. Geographical distribution of Parahelichus granulosu s (black squares) and P. pseudogranulosus sp. nov. (black circles)." httpUri="https://zenodo.org/record/231193/files/figure.png" pageId="12" pageNumber="487">Fig. 32</figureCitation>
|
||
). However, the distribution pattern is probably different from that of
|
||
<emphasis id="B9147970FFF18E0EFE147E14FDA09BFC" box="[394,554,872,896]" italics="true" pageId="12" pageNumber="487">
|
||
<taxonomicName id="4C60DEE1FFF18E0EFE147E14FDAF9BFC" box="[394,549,872,896]" class="Insecta" family="Dryopidae" genus="Parahelichus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="487" phylum="Arthropoda" rank="species" species="granulosus">P. granulosus</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BDFA562FFF18E0EFF597EF1FEDC9C6C" blockId="12.[151,1437,151,1040]" pageId="12" pageNumber="487">
|
||
<emphasis id="B9147970FFF18E0EFF597EF1FE8C9BD9" bold="true" box="[199,262,908,933]" pageId="12" pageNumber="487">Note.</emphasis>
|
||
Except for two species treated above, we have examined two other similar taxa from this small species complex. They came from
|
||
<collectingCountry id="F377E5F2FFF18E0EFE547ED2FD8E9BB4" box="[458,516,943,968]" name="India" pageId="12" pageNumber="487">India</collectingCountry>
|
||
and differ from
|
||
<taxonomicName id="4C60DEE1FFF18E0EFD5C7ECCFCD09BB4" box="[706,858,944,968]" class="Insecta" family="Dryopidae" genus="Parahelichus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="487" phylum="Arthropoda" rank="species" species="granulosus">
|
||
<emphasis id="B9147970FFF18E0EFD5C7ECCFCD09BB4" box="[706,858,944,968]" italics="true" pageId="12" pageNumber="487">P. granulosus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C60DEE1FFF18E0EFC0A7ECCFBF79BB4" box="[916,1149,944,968]" class="Insecta" family="Dryopidae" genus="Parahelichus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="487" phylum="Arthropoda" rank="species" species="pseudogranulosus">
|
||
<emphasis id="B9147970FFF18E0EFC0A7ECCFBF79BB4" box="[916,1149,944,968]" italics="true" pageId="12" pageNumber="487">P. pseudogranulosus</emphasis>
|
||
</taxonomicName>
|
||
in the form of elytra and pronotum. However, all Indian specimens investigated are females and, therefore, we keep from describing new species for them.
|
||
</paragraph>
|
||
<paragraph id="8BDFA562FFF18E0EFF097BC5FB229F1B" blockId="12.[151,1437,1720,2030]" pageId="12" pageNumber="487">
|
||
<emphasis id="B9147970FFF18E0EFF097BC5FE959EB1" bold="true" box="[151,287,1720,1742]" pageId="12" pageNumber="487">FIGURE 33.</emphasis>
|
||
Morphometric analyses of males of
|
||
<taxonomicName id="4C60DEE1FFF18E0EFD0C7BC5FC1A9EB2" box="[658,912,1720,1742]" class="Insecta" family="Dryopidae" genus="Parahelichus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="487" phylum="Arthropoda" rank="species" species="granulosus">
|
||
<emphasis id="B9147970FFF18E0EFD0C7BC5FC1A9EB2" box="[658,912,1720,1742]" italics="true" pageId="12" pageNumber="487">Parahelichus granulosus</emphasis>
|
||
</taxonomicName>
|
||
(n = 202) and
|
||
<taxonomicName id="4C60DEE1FFF18E0EFBBB7BC4FB7C9EB2" box="[1061,1270,1720,1742]" class="Insecta" family="Dryopidae" genus="Parahelichus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="487" phylum="Arthropoda" rank="species" species="pseudogranulosus" status="sp. nov.">
|
||
<emphasis id="B9147970FFF18E0EFBBB7BC4FB7C9EB2" box="[1061,1270,1720,1742]" italics="true" pageId="12" pageNumber="487">P. pseudogranulosus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A227C40BFFF18E0EFB627BC4FAC19EB2" box="[1276,1355,1721,1742]" pageId="12" pageNumber="487" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(n = 3), documenting that both taxa represent distinct species. (A) Box-plot graph to show the variation of aedeagus proportions (ratio of length of parameres to length of phallobase) of
|
||
<taxonomicName id="4C60DEE1FFF18E0EFD117B8BFC929F77" box="[655,792,1781,1803]" class="Insecta" family="Dryopidae" genus="Parahelichus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="487" phylum="Arthropoda" rank="species" species="granulosus">
|
||
<emphasis id="B9147970FFF18E0EFD117B8BFC929F77" box="[655,792,1781,1803]" italics="true" pageId="12" pageNumber="487">P. granulosus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C60DEE1FFF18E0EFCD77B8BFB939F77" box="[841,1049,1781,1803]" class="Insecta" family="Dryopidae" genus="Parahelichus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="487" phylum="Arthropoda" rank="species" species="pseudogranulosus" status="sp. nov.">
|
||
<emphasis id="B9147970FFF18E0EFCD77B8BFB939F77" box="[841,1049,1781,1803]" italics="true" pageId="12" pageNumber="487">P. pseudogranulosus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A227C40BFFF18E0EFB817B8BFBE79F77" box="[1055,1133,1782,1803]" pageId="12" pageNumber="487" rank="species">sp. nov.</taxonomicNameLabel>
|
||
Rectangles represent the 25th and 75th percentiles, horizontal lines show the median, whiskers are from the 10th to the 90th percentiles, and asterisks denote the extreme values. (B) PCA of males of
|
||
<taxonomicName id="4C60DEE1FFF18E0EFDC67A4EFD6F9F34" box="[600,741,1842,1864]" class="Insecta" family="Dryopidae" genus="Parahelichus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="487" phylum="Arthropoda" rank="species" species="granulosus">
|
||
<emphasis id="B9147970FFF18E0EFDC67A4EFD6F9F34" box="[600,741,1842,1864]" italics="true" pageId="12" pageNumber="487">P. granulosus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C60DEE1FFF18E0EFC807A4EFC799F34" box="[798,1011,1842,1864]" class="Insecta" family="Dryopidae" genus="Parahelichus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="487" phylum="Arthropoda" rank="species" species="pseudogranulosus" status="sp. nov.">
|
||
<emphasis id="B9147970FFF18E0EFC807A4EFC799F34" box="[798,1011,1842,1864]" italics="true" pageId="12" pageNumber="487">P. pseudogranulosus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A227C40BFFF18E0EFC637A4EFBDA9F34" box="[1021,1104,1843,1864]" pageId="12" pageNumber="487" rank="species">sp. nov.</taxonomicNameLabel>
|
||
based on 12 metric characters. Eigenvalues of two first axes are λ1 = 0.9985 and λ2 = 0.0010, explaining 99.95% of the total variation.
|
||
</paragraph>
|
||
<paragraph id="8BDFA562FFF18E0EFF097A0FFF7B9F92" blockId="12.[151,1437,1720,2030]" pageId="12" pageNumber="487">
|
||
Explanations and colour code of
|
||
<taxonomicName id="4C60DEE1FFF18E0EFD857A0EFD259FF4" box="[539,687,1906,1928]" class="Insecta" family="Dryopidae" genus="Parahelichus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="487" phylum="Arthropoda" rank="species" species="granulosus">
|
||
<emphasis id="B9147970FFF18E0EFD857A0EFD259FF4" box="[539,687,1906,1928]" italics="true" pageId="12" pageNumber="487">P. granulosus</emphasis>
|
||
</taxonomicName>
|
||
populations: YB (blue)—specimens from Yen Bai [
|
||
<collectingCountry id="F377E5F2FFF18E0EFA967A0FFAEF9FF4" box="[1288,1381,1906,1928]" name="Vietnam" pageId="12" pageNumber="487">Vietnam</collectingCountry>
|
||
], SP (orange)—specimens from Sa Pa [
|
||
<collectingCountry id="F377E5F2FFF18E0EFD8E7AE9FDE79FD6" box="[528,621,1940,1962]" name="Vietnam" pageId="12" pageNumber="487">Vietnam</collectingCountry>
|
||
], LN (red)—Louang Namtha [
|
||
<collectingCountry id="F377E5F2FFF18E0EFC237AE9FC7F9FD6" box="[957,1013,1940,1962]" name="Laos" pageId="12" pageNumber="487">Laos</collectingCountry>
|
||
], KR (green)—Khan river [
|
||
<collectingCountry id="F377E5F2FFF18E0EFAB77AE9FAEB9FD6" box="[1321,1377,1940,1962]" name="Laos" pageId="12" pageNumber="487">Laos</collectingCountry>
|
||
], BK (yellow)—Ban Khoun Ngeun env. [
|
||
<collectingCountry id="F377E5F2FFF18E0EFD987AC8FDB49FB7" box="[518,574,1973,1995]" name="Laos" pageId="12" pageNumber="487">Laos</collectingCountry>
|
||
]. Specimens of
|
||
<taxonomicName id="4C60DEE1FFF18E0EFD7C7ACBFC3E9FB7" box="[738,948,1973,1995]" class="Insecta" family="Dryopidae" genus="Parahelichus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="487" phylum="Arthropoda" rank="species" species="pseudogranulosus" status="sp. nov.">
|
||
<emphasis id="B9147970FFF18E0EFD7C7ACBFC3E9FB7" box="[738,948,1973,1995]" italics="true" pageId="12" pageNumber="487">P. pseudogranulosus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A227C40BFFF18E0EFC227ACBFB869FB7" box="[956,1036,1974,1995]" pageId="12" pageNumber="487" rank="species">sp. nov.</taxonomicNameLabel>
|
||
are shown as black circles in the PCA diagram.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |