treatments-xml/data/03/BF/DC/03BFDC60AB68695CFF47FC26FBE5FF1C.xml

<document id="05CC56B41BBA1659134B05C3756BDDB1" ID-CLB-Dataset="36832" ID-DOI="10.11646/zootaxa.3820.1.1" ID-GBIF-Dataset="e3d7d016-1d73-438f-95e4-1b028b2b9627" ID-ISSN="1175-5326" ID-Zenodo-Dep="286303" ID-ZooBank="BBE4177B-5A2C-4911-987C-454BB8FA767C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460439774061" checkinUser="plazi" docAuthor="Pešić, Vladimir &amp; Smit, Harry" docDate="2014" docId="03BFDC60AB68695CFF47FC26FBE5FF1C" docLanguage="en" docName="zt03820p080.pdf" docOrigin="Zootaxa 3820 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Monatractides urania Pešić &amp; Smit, 2014, n. sp." docType="treatment" docVersion="7" lastPageNumber="70" masterDocId="FF86A418AB5D6919FFD0FFCBFF8CFFE4" masterDocTitle="Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Ghana" masterLastPageNumber="80" masterPageNumber="1" pageNumber="54" updateTime="1698332736548" updateUser="ExternalLinkService">
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<mods:title id="8E66CBEA7E2EFECAAFB69D1857666D35">Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Ghana</mods:title>
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<mods:namePart id="8009955109A69298B62BC11CD2322A4F">Pešić, Vladimir</mods:namePart>
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<mods:namePart id="E4277CC22F576BE522DF5B958DBEF783">Smit, Harry</mods:namePart>
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<treatment id="03BFDC60AB68695CFF47FC26FBE5FF1C" ID-DOI="http://doi.org/10.5281/zenodo.6143481" ID-GBIF-Taxon="127668411" ID-Zenodo-Dep="6143481" LSID="urn:lsid:plazi:treatment:03BFDC60AB68695CFF47FC26FBE5FF1C" httpUri="http://treatment.plazi.org/id/03BFDC60AB68695CFF47FC26FBE5FF1C" lastPageId="69" lastPageNumber="70" pageId="53" pageNumber="54">
<subSubSection id="C30C3EFDAB68692CFF47FC26FE2CFBCC" pageId="53" pageNumber="54" type="nomenclature">
<paragraph id="8BA96D76AB68692CFF47FC26FE6BFBE3" blockId="53.[151,487,1005,1064]" box="[151,487,1005,1031]" pageId="53" pageNumber="54">
<heading id="D0E1DA1AAB68692CFF47FC26FE6BFBE3" bold="true" box="[151,487,1005,1031]" fontSize="11" level="1" pageId="53" pageNumber="54" reason="1">
<emphasis id="B962B164AB68692CFF47FC26FE6BFBE3" bold="true" box="[151,487,1005,1031]" pageId="53" pageNumber="54">
<taxonomicName id="4C1616F5AB68692CFF47FC26FE12FBE3" box="[151,414,1005,1031]" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="urania" status="sp. nov.">
<emphasis id="B962B164AB68692CFF47FC26FE12FBE3" bold="true" box="[151,414,1005,1031]" italics="true" pageId="53" pageNumber="54">Monatractides urania</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2510C1FAB68692CFE75FC26FE6BFBE3" box="[421,487,1005,1031]" pageId="53" pageNumber="54" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BA96D76AB68692CFF47FBC4FE2CFBCC" blockId="53.[151,487,1005,1064]" box="[151,416,1039,1064]" pageId="53" pageNumber="54">
(
<figureCitation id="132D71F3AB68692CFF4FFBC4FEB8FBCC" box="[159,308,1039,1064]" captionStart="FIGURE 36 A – H" captionStartId="54.[151,250,1940,1962]" captionTargetBox="[226,1361,205,1915]" captionTargetId="figure@54.[202,1385,193,1932]" captionTargetPageId="54" captionText="FIGURE 36 A – H. Monatractides urania n. sp., male, Kintampo Falls: A = frontal margin of idiosoma, dorsal view; B = dorsal shield; C = idiosoma, ventral view; D = genital field; E = palp, lateral view; F = palp, medial view; G = gnathosoma; H = I-L- 5 and - 6. Scale bars = 100 Μm." httpUri="https://zenodo.org/record/286339/files/figure.png" pageId="53" pageNumber="54">Figs. 36A–H</figureCitation>
, 
<figureCitation id="132D71F3AB68692CFEEFFBDBFEEBFBCC" box="[319,359,1040,1064]" captionStart="FIGURE 37 A – I" captionStartId="55.[151,250,1851,1873]" captionTargetBox="[166,1436,195,1811]" captionTargetId="figure@55.[166,1451,195,1813]" captionTargetPageId="55" captionText="FIGURE 37 A – I. Photographs of dorsal shield. A – B Monatractides melpomene n. sp., Amedzofe Falls: A = male holotype, B = female paratype. C – D M. euterpe n. sp., creek upstream Sagyimase: C = male holotype, D = female paratype. E – F M. polyhymnia n. sp., Kue River: E = male holotype, F = female paratype. G M. thalia n. sp., male holotype, Agumatsa River. H M. terpsichore n. sp., female holotype, Plunge pool. I M. urania n. sp., male paratype, Kintampo Falls." httpUri="https://zenodo.org/record/286340/files/figure.png" pageId="53" pageNumber="54">
<date id="FFA84BB6AB68692CFEEFFBDBFEEBFBCC" box="[319,359,1040,1064]" pageId="53" pageNumber="54" value="1937-01">37I</date>
</figureCitation>
, 
<figureCitation id="132D71F3AB68692CFEA2FBDBFE15FBCC" box="[370,409,1040,1064]" captionStart="FIGURE 38 A – I" captionStartId="56.[151,250,1900,1922]" captionTargetBox="[164,1407,195,1862]" captionTargetId="figure@56.[164,1408,195,1863]" captionTargetPageId="56" captionText="FIGURE 38 A – I. Photographs of ventral shield. A – B Monatractides melpomene n. sp., Amedzofe Falls: A = male holotype, B = female paratype. C – D M. euterpe n. sp., creek upstream Sagyimase: C = male holotype, D = female paratype. E – F M. polyhymnia n. sp., Kue River: E = male holotype, F = female paratype. G M. thalia n. sp., male holotype, Agumatsa River. H M. terpsichore n. sp., female holotype, Plunge pool. I M. urania n. sp., male paratype, Kintampo Falls." httpUri="https://zenodo.org/record/286341/files/figure.png" pageId="53" pageNumber="54">
<date id="FFA84BB6AB68692CFEA2FBDBFE15FBCC" box="[370,409,1040,1064]" pageId="53" pageNumber="54" value="1938-01">38I</date>
</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30C3EFDAB68692CFF47FB9CFB1EFB71" pageId="53" pageNumber="54" type="materials_examined">
<paragraph id="8BA96D76AB68692CFF47FB9CFB1EFB71" blockId="53.[151,1437,1111,2037]" pageId="53" pageNumber="54">
<emphasis id="B962B164AB68692CFF47FB9CFEADFB94" bold="true" box="[151,289,1111,1136]" pageId="53" pageNumber="54">
<typeStatus id="54ADD3D4AB68692CFF47FB9CFF5EFB94" box="[151,210,1111,1136]" pageId="53" pageNumber="54">Type</typeStatus>
series
</emphasis>
. 
<typeStatus id="54ADD3D4AB68692CFEE4FB9CFE10FB94" box="[308,412,1111,1136]" pageId="53" pageNumber="54" type="holotype">Holotype</typeStatus>
male, dissected and slide mounted, 
<collectingCountry id="F3012DE6AB68692CFC82FB9CFC12FB94" box="[850,926,1111,1136]" name="Ghana" pageId="53" pageNumber="54">Ghana</collectingCountry>
, Kintampo Falls, 8º 
<geoCoordinate id="EE220BB1AB68692CFB5AFB9CFB70FB8B" box="[1162,1276,1111,1136]" direction="north" orientation="latitude" pageId="53" pageNumber="54" precision="55" value="5.413">05.413 N</geoCoordinate>
, 1º 
<geoCoordinate id="EE220BB1AB68692CFACFFB93FA14FB8B" box="[1311,1432,1112,1136]" direction="west" orientation="longitude" pageId="53" pageNumber="54" precision="55" value="-41.881">41.881 W</geoCoordinate>
, 
<date id="FFA84BB6AB68692CFF47FBB7FE8FFB71" box="[151,259,1148,1173]" pageId="53" pageNumber="54" value="2011-03-03">3.iii.2011</date>
, Smit. 
<typeStatus id="54ADD3D4AB68692CFE82FBB6FE35FB71" box="[338,441,1149,1173]" pageId="53" pageNumber="54" type="paratype">Paratype</typeStatus>
: 3/0/0, same data as 
<typeStatus id="54ADD3D4AB68692CFD4FFBB7FC88FB71" box="[671,772,1148,1173]" pageId="53" pageNumber="54" type="holotype">holotype</typeStatus>
, 2/0/0 dissected and slide mounted.
</paragraph>
</subSubSection>
<subSubSection id="C30C3EFDAB68692CFF17FB54FDB9FAC1" pageId="53" pageNumber="54" type="diagnosis">
<paragraph id="8BA96D76AB68692CFF17FB54FDB9FAC1" blockId="53.[151,1437,1111,2037]" pageId="53" pageNumber="54">
<emphasis id="B962B164AB68692CFF17FB54FEB6FB5C" bold="true" box="[199,314,1183,1208]" pageId="53" pageNumber="54">Diagnosis</emphasis>
(Female unknown). Shoulder platelets shorter than frontal platelets (shoulder/frontal platelets L ratio 0.93–0.96); medial margin with convex protrusions medial to the eyes; medial margin of Cx-II/III very short (21µm); genital field large (L 140–160 µm) and strongly trapezoid in shape, anteriorly enlarged and pointed; I-L-6 L/H ratio 2.0–2.1, distally thickened.
</paragraph>
</subSubSection>
<subSubSection id="C30C3EFDAB686920FF17FAE4FE15FF54" lastPageId="57" lastPageNumber="58" pageId="53" pageNumber="54" type="description">
<paragraph id="8BA96D76AB68692CFF17FAE4FC3CF98C" blockId="53.[151,1437,1111,2037]" pageId="53" pageNumber="54">
<emphasis id="B962B164AB68692CFF17FAE4FED5FAAC" bold="true" box="[199,345,1327,1352]" pageId="53" pageNumber="54">Description.</emphasis>
Male. 
<emphasis id="B962B164AB68692CFE78FAE4FDE8FAAC" box="[424,612,1327,1352]" italics="true" pageId="53" pageNumber="54">General features</emphasis>
—Idiosoma elongated-oval; shoulder platelets short, slightly shorter than frontal platelets (shoulder/frontal platelets L ratio 0.93–0.96); frontal margin medially convexly rounded between anterolaterally orientated apodemes (
<figureCitation id="132D71F3AB68692CFDE9FABCFD2EFA74" box="[569,674,1399,1424]" captionStart="FIGURE 36 A – H" captionStartId="54.[151,250,1940,1962]" captionTargetBox="[226,1361,205,1915]" captionTargetId="figure@54.[202,1385,193,1932]" captionTargetPageId="54" captionText="FIGURE 36 A – H. Monatractides urania n. sp., male, Kintampo Falls: A = frontal margin of idiosoma, dorsal view; B = dorsal shield; C = idiosoma, ventral view; D = genital field; E = palp, lateral view; F = palp, medial view; G = gnathosoma; H = I-L- 5 and - 6. Scale bars = 100 Μm." httpUri="https://zenodo.org/record/286339/files/figure.png" pageId="53" pageNumber="54">Fig. 36A</figureCitation>
); Cxgl–4 located far anteriorly, near tips of Cx-I; gnathosomal bay, U-shaped, proximally rectangular box-shaped; medial margin of Cx-II/III relatively very short; genital field large and strongly trapezoid in shape, anteriorly enlarged and pointed, laterally convex, tapering towards posterior (
<figureCitation id="132D71F3AB68692CFAB4FA74FF60FA19" captionStart="FIGURE 36 A – H" captionStartId="54.[151,250,1940,1962]" captionTargetBox="[226,1361,205,1915]" captionTargetId="figure@54.[202,1385,193,1932]" captionTargetPageId="54" captionText="FIGURE 36 A – H. Monatractides urania n. sp., male, Kintampo Falls: A = frontal margin of idiosoma, dorsal view; B = dorsal shield; C = idiosoma, ventral view; D = genital field; E = palp, lateral view; F = palp, medial view; G = gnathosoma; H = I-L- 5 and - 6. Scale bars = 100 Μm." httpUri="https://zenodo.org/record/286339/files/figure.png" pageId="53" pageNumber="54">Figs. 36C–D</figureCitation>
); suture line of Cx-IV extending back beyond posterior margin of genital field; excretory pore slightly away from the line of primary sclerotization, excretory pore anterior to Vgl–2; gnathosoma nearly triangular in lateral view (
<figureCitation id="132D71F3AB68692CFEFAF9E7FE1FF9A1" box="[298,403,1580,1605]" captionStart="FIGURE 36 A – H" captionStartId="54.[151,250,1940,1962]" captionTargetBox="[226,1361,205,1915]" captionTargetId="figure@54.[202,1385,193,1932]" captionTargetPageId="54" captionText="FIGURE 36 A – H. Monatractides urania n. sp., male, Kintampo Falls: A = frontal margin of idiosoma, dorsal view; B = dorsal shield; C = idiosoma, ventral view; D = genital field; E = palp, lateral view; F = palp, medial view; G = gnathosoma; H = I-L- 5 and - 6. Scale bars = 100 Μm." httpUri="https://zenodo.org/record/286339/files/figure.png" pageId="53" pageNumber="54">Fig. 36G</figureCitation>
); P-2 and P-3 distal margins without pointed tips, ventral seta on P-4 short (
<figureCitation id="132D71F3AB68692CFB37F9E7FAFAF9A1" box="[1255,1398,1580,1605]" captionStart="FIGURE 36 A – H" captionStartId="54.[151,250,1940,1962]" captionTargetBox="[226,1361,205,1915]" captionTargetId="figure@54.[202,1385,193,1932]" captionTargetPageId="54" captionText="FIGURE 36 A – H. Monatractides urania n. sp., male, Kintampo Falls: A = frontal margin of idiosoma, dorsal view; B = dorsal shield; C = idiosoma, ventral view; D = genital field; E = palp, lateral view; F = palp, medial view; G = gnathosoma; H = I-L- 5 and - 6. Scale bars = 100 Μm." httpUri="https://zenodo.org/record/286339/files/figure.png" pageId="53" pageNumber="54">Figs. 36E–F</figureCitation>
); I- L-6 L/H ratio 2.0–2.1, basally narrowed, distally thickened (
<figureCitation id="132D71F3AB68692CFCEAF984FC2FF98C" box="[826,931,1615,1640]" captionStart="FIGURE 36 A – H" captionStartId="54.[151,250,1940,1962]" captionTargetBox="[226,1361,205,1915]" captionTargetId="figure@54.[202,1385,193,1932]" captionTargetPageId="54" captionText="FIGURE 36 A – H. Monatractides urania n. sp., male, Kintampo Falls: A = frontal margin of idiosoma, dorsal view; B = dorsal shield; C = idiosoma, ventral view; D = genital field; E = palp, lateral view; F = palp, medial view; G = gnathosoma; H = I-L- 5 and - 6. Scale bars = 100 Μm." httpUri="https://zenodo.org/record/286339/files/figure.png" pageId="53" pageNumber="54">Fig. 36H</figureCitation>
).
</paragraph>
<paragraph id="8BA96D76AB68692CFF17F9BEFBBDF810" blockId="53.[151,1437,1111,2037]" pageId="53" pageNumber="54">
<emphasis id="B962B164AB68692CFF17F9BEFE3FF969" box="[199,435,1652,1677]" italics="true" pageId="53" pageNumber="54">Measurements. Male</emphasis>
(
<typeStatus id="54ADD3D4AB68692CFE14F9BFFDA6F969" box="[452,554,1652,1677]" pageId="53" pageNumber="54" type="holotype">holotype</typeStatus>
, in parentheses measurements of 
<typeStatus id="54ADD3D4AB68692CFC7DF9BEFB97F969" box="[941,1051,1653,1677]" pageId="53" pageNumber="54" type="paratype">paratypes</typeStatus>
, n = 1)—Idiosoma (ventral view: 
<figureCitation id="132D71F3AB68692CFF47F95CFE8BF954" box="[151,263,1687,1712]" captionStart="FIGURE 36 A – H" captionStartId="54.[151,250,1940,1962]" captionTargetBox="[226,1361,205,1915]" captionTargetId="figure@54.[202,1385,193,1932]" captionTargetPageId="54" captionText="FIGURE 36 A – H. Monatractides urania n. sp., male, Kintampo Falls: A = frontal margin of idiosoma, dorsal view; B = dorsal shield; C = idiosoma, ventral view; D = genital field; E = palp, lateral view; F = palp, medial view; G = gnathosoma; H = I-L- 5 and - 6. Scale bars = 100 Μm." httpUri="https://zenodo.org/record/286339/files/figure.png" pageId="53" pageNumber="54">Figs. 36C</figureCitation>
, 
<figureCitation id="132D71F3AB68692CFEC2F953FEBBF954" box="[274,311,1688,1712]" captionStart="FIGURE 38 A – I" captionStartId="56.[151,250,1900,1922]" captionTargetBox="[164,1407,195,1862]" captionTargetId="figure@56.[164,1408,195,1863]" captionTargetPageId="56" captionText="FIGURE 38 A – I. Photographs of ventral shield. A – B Monatractides melpomene n. sp., Amedzofe Falls: A = male holotype, B = female paratype. C – D M. euterpe n. sp., creek upstream Sagyimase: C = male holotype, D = female paratype. E – F M. polyhymnia n. sp., Kue River: E = male holotype, F = female paratype. G M. thalia n. sp., male holotype, Agumatsa River. H M. terpsichore n. sp., female holotype, Plunge pool. I M. urania n. sp., male paratype, Kintampo Falls." httpUri="https://zenodo.org/record/286341/files/figure.png" pageId="53" pageNumber="54">
<date id="FFA84BB6AB68692CFEC2F953FEBBF954" box="[274,311,1688,1712]" pageId="53" pageNumber="54" value="1938-01">38I</date>
</figureCitation>
) L 619 (642), W 425 (434); dorsal shield (
<figureCitation id="132D71F3AB68692CFCC5F95CFC0AF954" box="[789,902,1687,1712]" captionStart="FIGURE 36 A – H" captionStartId="54.[151,250,1940,1962]" captionTargetBox="[226,1361,205,1915]" captionTargetId="figure@54.[202,1385,193,1932]" captionTargetPageId="54" captionText="FIGURE 36 A – H. Monatractides urania n. sp., male, Kintampo Falls: A = frontal margin of idiosoma, dorsal view; B = dorsal shield; C = idiosoma, ventral view; D = genital field; E = palp, lateral view; F = palp, medial view; G = gnathosoma; H = I-L- 5 and - 6. Scale bars = 100 Μm." httpUri="https://zenodo.org/record/286339/files/figure.png" pageId="53" pageNumber="54">Figs. 36B</figureCitation>
, 
<figureCitation id="132D71F3AB68692CFC41F953FC34F954" box="[913,952,1688,1712]" captionStart="FIGURE 37 A – I" captionStartId="55.[151,250,1851,1873]" captionTargetBox="[166,1436,195,1811]" captionTargetId="figure@55.[166,1451,195,1813]" captionTargetPageId="55" captionText="FIGURE 37 A – I. Photographs of dorsal shield. A – B Monatractides melpomene n. sp., Amedzofe Falls: A = male holotype, B = female paratype. C – D M. euterpe n. sp., creek upstream Sagyimase: C = male holotype, D = female paratype. E – F M. polyhymnia n. sp., Kue River: E = male holotype, F = female paratype. G M. thalia n. sp., male holotype, Agumatsa River. H M. terpsichore n. sp., female holotype, Plunge pool. I M. urania n. sp., male paratype, Kintampo Falls." httpUri="https://zenodo.org/record/286340/files/figure.png" pageId="53" pageNumber="54">
<date id="FFA84BB6AB68692CFC41F953FC34F954" box="[913,952,1688,1712]" pageId="53" pageNumber="54" value="1937-01">37I</date>
</figureCitation>
) L 525 (550), W 366 (384), L/W ratio 1.43 (1.43); dorsal plate L 491 (509); shoulder platelets L 116–119 (127–131), W 50–52 (55–58), L/W ratio 2.2–2.4 (2.3); frontal platelets L 123–125 (127–131), W 48 (50–52), L/W ratio 2.5–2.6 (2.5–2.6); shoulder/frontal platelets L ratio 0.93–0.96 (0.96). Gnathosomal bay L 150 (153), Cx-I total L 275 (306), Cx-I mL 125 (122), Cx-II+III mL 21 (21); ratio Cx-I L/Cx-II+III mL 12.9 (14.6); Cx-I mL/Cx-II+III mL 5.9 (5.8). Genital field L/W 141 (155)/127 (130), ratio 1.11 (1.19); distance genital field-excretory pore 133 (131), genital field-caudal idiosoma margin 181 (189). Gnathosoma vL 157 (155); chelicera total L 189 (198); palp total L 188–190 (188–191), dL/H, dL/H ratio: P-1, 26/20, 1.3 (26/21, 1.26); P-2, 57/38–39, 1.48 (54–55/37, 1.48); P-3, 38–39/32, 1.2 (38–39/31, 1.25); P-4, 49/ 22, 2.2 (52/21–22, 2.4); P-5, 18–19/11–12, 1.6 (18–19/12, 1.5); P-2/P-4 ratio 1.16 (1.04); dL of I-L-2–6: 63 (65), 76 (80), 97 (99), 92 (97), 100 (106); I-L-6 H 48 (52), dL/H I-L-6 ratio 2.08 (2.03).
</paragraph>
<caption id="DF693DFEAB6B692FFF47F85FFE37F808" httpUri="https://zenodo.org/record/286339/files/figure.png" pageId="54" pageNumber="55" targetBox="[226,1361,205,1915]" targetPageId="54">
<paragraph id="8BA96D76AB6B692FFF47F85FFE37F808" blockId="54.[151,1436,1940,2028]" pageId="54" pageNumber="55">
<emphasis id="B962B164AB6B692FFF47F85FFEDCF84E" bold="true" box="[151,336,1940,1962]" pageId="54" pageNumber="55">FIGURE 36A–H.</emphasis>
<taxonomicName id="4C1616F5AB6B692FFE86F85FFDB8F84E" box="[342,564,1940,1962]" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="urania" status="sp. nov.">
<emphasis id="B962B164AB6B692FFE86F85FFDB8F84E" box="[342,564,1940,1962]" italics="true" pageId="54" pageNumber="55">Monatractides urania</emphasis>
</taxonomicName>
<emphasis id="B962B164AB6B692FFDEAF85FFDFFF84E" bold="true" box="[570,627,1940,1962]" pageId="54" pageNumber="55">
<taxonomicNameLabel id="A2510C1FAB6B692FFDEAF85FFDFFF84E" box="[570,627,1940,1962]" pageId="54" pageNumber="55" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, male, Kintampo Falls: A = frontal margin of idiosoma, dorsal view; B = dorsal shield; C = idiosoma, ventral view; D = genital field; E = palp, lateral view; F = palp, medial view; G = gnathosoma; H = I-L-5 and -6. Scale bars = 100 Μm.
</paragraph>
</caption>
<caption id="DF693DFEAB6A692EFF47F8F0FB22F851" httpUri="https://zenodo.org/record/286340/files/figure.png" pageId="55" pageNumber="56" targetBox="[166,1436,195,1811]" targetPageId="55">
<paragraph id="8BA96D76AB6A692EFF47F8F0FB22F851" blockId="55.[151,1437,1851,1973]" pageId="55" pageNumber="56">
<emphasis id="B962B164AB6A692EFF47F8F0FECBF8B5" bold="true" box="[151,327,1851,1873]" pageId="55" pageNumber="56">FIGURE 37A–I.</emphasis>
Photographs of dorsal shield. 
<emphasis id="B962B164AB6A692EFDAEF8F0FD21F8B5" bold="true" box="[638,685,1851,1873]" pageId="55" pageNumber="56">A–B</emphasis>
<taxonomicName id="4C1616F5AB6A692EFD64F8F0FC4DF8B5" box="[692,961,1851,1873]" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="melpomene" status="sp. nov.">
<emphasis id="B962B164AB6A692EFD64F8F0FC4DF8B5" box="[692,961,1851,1873]" italics="true" pageId="55" pageNumber="56">Monatractides melpomene</emphasis>
</taxonomicName>
<emphasis id="B962B164AB6A692EFC18F8F0FB8EF8B5" bold="true" box="[968,1026,1851,1873]" pageId="55" pageNumber="56">
<taxonomicNameLabel id="A2510C1FAB6A692EFC18F8F0FB8EF8B5" box="[968,1026,1851,1873]" pageId="55" pageNumber="56" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, Amedzofe Falls: A = male holotype, B = female paratype. 
<emphasis id="B962B164AB6A692EFEB4F897FE18F896" bold="true" box="[356,404,1884,1906]" pageId="55" pageNumber="56">C–D</emphasis>
<taxonomicName id="4C1616F5AB6A692EFE4EF896FD83F896" box="[414,527,1885,1906]" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="euterpe" status="sp. nov.">
<emphasis id="B962B164AB6A692EFE4EF896FD83F896" box="[414,527,1885,1906]" italics="true" pageId="55" pageNumber="56">M. euterpe</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2510C1FAB6A692EFDC9F897FDDAF896" box="[537,598,1884,1906]" pageId="55" pageNumber="56" rank="species">
<emphasis id="B962B164AB6A692EFDC9F897FDDAF896" bold="true" box="[537,598,1884,1906]" pageId="55" pageNumber="56">n. sp.</emphasis>
</taxonomicNameLabel>
, creek upstream Sagyimase: C = male holotype, D = female paratype. 
<emphasis id="B962B164AB6A692EFA9BF897FAFAF896" bold="true" box="[1355,1398,1884,1906]" pageId="55" pageNumber="56">E–F</emphasis>
<taxonomicName id="4C1616F5AB6A692EFA50F896FE81F877" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="polyhymnia" status="sp. nov.">
<emphasis id="B962B164AB6A692EFA50F896FE81F877" italics="true" pageId="55" pageNumber="56">M. polyhymnia</emphasis>
</taxonomicName>
<emphasis id="B962B164AB6A692EFEC4F8B6FEC1F877" bold="true" box="[276,333,1917,1939]" pageId="55" pageNumber="56">
<taxonomicNameLabel id="A2510C1FAB6A692EFEC4F8B6FEC1F877" box="[276,333,1917,1939]" pageId="55" pageNumber="56" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, Kue River: E = male holotype, F = female paratype. 
<emphasis id="B962B164AB6A692EFCA4F8B6FC04F877" bold="true" box="[884,904,1917,1939]" pageId="55" pageNumber="56">G</emphasis>
<taxonomicName id="4C1616F5AB6A692EFC5EF8B4FC60F877" box="[910,1004,1917,1940]" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="thalia" status="sp. nov.">
<emphasis id="B962B164AB6A692EFC5EF8B4FC60F877" box="[910,1004,1917,1940]" italics="true" pageId="55" pageNumber="56">M. thalia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2510C1FAB6A692EFC23F8B6FBA0F877" box="[1011,1068,1917,1939]" pageId="55" pageNumber="56" rank="species">
<emphasis id="B962B164AB6A692EFC23F8B6FBA0F877" bold="true" box="[1011,1068,1917,1939]" pageId="55" pageNumber="56">n. sp.</emphasis>
</taxonomicNameLabel>
, male holotype, Agumatsa River. 
<emphasis id="B962B164AB6A692EFA58F8B6FA10F877" bold="true" box="[1416,1436,1917,1939]" pageId="55" pageNumber="56">H</emphasis>
<taxonomicName id="4C1616F5AB6A692EFF47F86BFEA7F851" box="[151,299,1951,1973]" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="terpsichore" status="sp. nov.">
<emphasis id="B962B164AB6A692EFF47F86BFEA7F851" box="[151,299,1951,1973]" italics="true" pageId="55" pageNumber="56">M. terpsichore</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2510C1FAB6A692EFEE1F854FEE6F851" box="[305,362,1951,1973]" pageId="55" pageNumber="56" rank="species">
<emphasis id="B962B164AB6A692EFEE1F854FEE6F851" bold="true" box="[305,362,1951,1973]" pageId="55" pageNumber="56">n. sp.</emphasis>
</taxonomicNameLabel>
, female holotype, Plunge pool. 
<emphasis id="B962B164AB6A692EFD7DF854FD3BF851" bold="true" box="[685,695,1951,1973]" pageId="55" pageNumber="56">I</emphasis>
<taxonomicName id="4C1616F5AB6A692EFD6EF86BFCAFF851" box="[702,803,1952,1973]" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="urania" status="sp. nov.">
<emphasis id="B962B164AB6A692EFD6EF86BFCAFF851" box="[702,803,1952,1973]" italics="true" pageId="55" pageNumber="56">M. urania</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2510C1FAB6A692EFCF9F854FCEEF851" box="[809,866,1951,1973]" pageId="55" pageNumber="56" rank="species">
<emphasis id="B962B164AB6A692EFCF9F854FCEEF851" bold="true" box="[809,866,1951,1973]" pageId="55" pageNumber="56">n. sp.</emphasis>
</taxonomicNameLabel>
, male paratype, Kintampo Falls.
</paragraph>
</caption>
<caption id="DF693DFEAB656921FF47F8A7FB22F802" httpUri="https://zenodo.org/record/286341/files/figure.png" pageId="56" pageNumber="57" targetBox="[164,1407,195,1862]" targetPageId="56">
<paragraph id="8BA96D76AB656921FF47F8A7FB22F802" blockId="56.[151,1437,1900,2023]" pageId="56" pageNumber="57">
<emphasis id="B962B164AB656921FF47F8A7FECAF866" bold="true" box="[151,326,1900,1922]" pageId="56" pageNumber="57">FIGURE 38A–I.</emphasis>
Photographs of ventral shield. 
<emphasis id="B962B164AB656921FD54F8A7FD3EF866" bold="true" box="[644,690,1900,1922]" pageId="56" pageNumber="57">A–B</emphasis>
<taxonomicName id="4C1616F5AB656921FD68F8A7FC49F866" box="[696,965,1900,1922]" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="melpomene" status="sp. nov.">
<emphasis id="B962B164AB656921FD68F8A7FC49F866" box="[696,965,1900,1922]" italics="true" pageId="56" pageNumber="57">Monatractides melpomene</emphasis>
</taxonomicName>
<emphasis id="B962B164AB656921FC1CF8A7FB89F866" bold="true" box="[972,1029,1900,1922]" pageId="56" pageNumber="57">
<taxonomicNameLabel id="A2510C1FAB656921FC1CF8A7FB89F866" box="[972,1029,1900,1922]" pageId="56" pageNumber="57" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, Amedzofe Falls: A = male holotype, B = female paratype. 
<emphasis id="B962B164AB656921FEB4F846FE18F847" bold="true" box="[356,404,1933,1955]" pageId="56" pageNumber="57">C–D</emphasis>
<taxonomicName id="4C1616F5AB656921FE4EF844FD83F840" box="[414,527,1935,1956]" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="euterpe" status="sp. nov.">
<emphasis id="B962B164AB656921FE4EF844FD83F840" box="[414,527,1935,1956]" italics="true" pageId="56" pageNumber="57">M. euterpe</emphasis>
</taxonomicName>
<emphasis id="B962B164AB656921FDC9F846FDDAF847" bold="true" box="[537,598,1933,1955]" pageId="56" pageNumber="57">
<taxonomicNameLabel id="A2510C1FAB656921FDC9F846FDDAF847" box="[537,598,1933,1955]" pageId="56" pageNumber="57" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, creek upstream Sagyimase: C = male holotype, D = female paratype. 
<emphasis id="B962B164AB656921FA9BF846FAFAF847" bold="true" box="[1355,1398,1933,1955]" pageId="56" pageNumber="57">E–F</emphasis>
<taxonomicName id="4C1616F5AB656921FA50F844FE81F821" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="polyhymnia" status="sp. nov.">
<emphasis id="B962B164AB656921FA50F844FE81F821" italics="true" pageId="56" pageNumber="57">M. polyhymnia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2510C1FAB656921FEC4F864FEC1F821" box="[276,333,1967,1989]" pageId="56" pageNumber="57" rank="species">
<emphasis id="B962B164AB656921FEC4F864FEC1F821" bold="true" box="[276,333,1967,1989]" pageId="56" pageNumber="57">n. sp.</emphasis>
</taxonomicNameLabel>
, Kue River: E = male holotype, F = female paratype. 
<emphasis id="B962B164AB656921FCA4F864FC04F821" bold="true" box="[884,904,1967,1989]" pageId="56" pageNumber="57">G</emphasis>
<taxonomicName id="4C1616F5AB656921FC5EF87BFC60F821" box="[910,1004,1967,1989]" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="thalia" status="sp. nov.">
<emphasis id="B962B164AB656921FC5EF87BFC60F821" box="[910,1004,1967,1989]" italics="true" pageId="56" pageNumber="57">M. thalia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2510C1FAB656921FC23F864FBA0F821" box="[1011,1068,1967,1989]" pageId="56" pageNumber="57" rank="species">
<emphasis id="B962B164AB656921FC23F864FBA0F821" bold="true" box="[1011,1068,1967,1989]" pageId="56" pageNumber="57">n. sp.</emphasis>
</taxonomicNameLabel>
, male holotype, Agumatsa River. 
<emphasis id="B962B164AB656921FA58F864FA10F821" bold="true" box="[1416,1436,1967,1989]" pageId="56" pageNumber="57">H</emphasis>
<taxonomicName id="4C1616F5AB656921FF47F819FEA7F802" box="[151,299,2000,2023]" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="terpsichore" status="sp. nov.">
<emphasis id="B962B164AB656921FF47F819FEA7F802" box="[151,299,2000,2023]" italics="true" pageId="56" pageNumber="57">M. terpsichore</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2510C1FAB656921FEE1F81BFEE6F802" box="[305,362,2000,2022]" pageId="56" pageNumber="57" rank="species">
<emphasis id="B962B164AB656921FEE1F81BFEE6F802" bold="true" box="[305,362,2000,2022]" pageId="56" pageNumber="57">n. sp.</emphasis>
</taxonomicNameLabel>
, female holotype, Plunge pool. 
<emphasis id="B962B164AB656921FD7DF81BFD3BF802" bold="true" box="[685,695,2000,2022]" pageId="56" pageNumber="57">I</emphasis>
<taxonomicName id="4C1616F5AB656921FD6EF819FCAFF802" box="[702,803,2001,2023]" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="urania" status="sp. nov.">
<emphasis id="B962B164AB656921FD6EF819FCAFF802" box="[702,803,2001,2023]" italics="true" pageId="56" pageNumber="57">M. urania</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2510C1FAB656921FCF9F81BFCEEF802" box="[809,866,2000,2022]" pageId="56" pageNumber="57" rank="species">
<emphasis id="B962B164AB656921FCF9F81BFCEEF802" bold="true" box="[809,866,2000,2022]" pageId="56" pageNumber="57">n. sp.</emphasis>
</taxonomicNameLabel>
, male paratype, Kintampo Falls.
</paragraph>
</caption>
<paragraph id="8BA96D76AB646920FF17FF5CFE15FF54" blockId="57.[151,1437,151,645]" box="[199,409,151,176]" pageId="57" pageNumber="58">
<emphasis id="B962B164AB646920FF17FF5CFE97FF54" box="[199,283,151,176]" italics="true" pageId="57" pageNumber="58">Female</emphasis>
: unknown.
</paragraph>
</subSubSection>
<subSubSection id="C30C3EFDAB646920FF17FF77FAC3FF1C" pageId="57" pageNumber="58" type="etymology">
<paragraph id="8BA96D76AB646920FF17FF77FAC3FF1C" blockId="57.[151,1437,151,645]" pageId="57" pageNumber="58">
<emphasis id="B962B164AB646920FF17FF77FECAFF31" bold="true" box="[199,326,188,213]" pageId="57" pageNumber="58">Etymology</emphasis>
. The species is named after 
<taxonomicName id="4C1616F5AB646920FD4BFF77FD6BFF31" box="[667,743,188,213]" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="urania" status="sp. nov.">Urania</taxonomicName>
(Ancient Greek: Οὐρανία), one of nine Muses from Greek mythology, who was a patron of tragedy. The species name is a noun in apposition (in the nominative case).
</paragraph>
</subSubSection>
<subSubSection id="C30C3EFDAB646920FF17FECFFC39FD84" pageId="57" pageNumber="58" type="discussion">
<paragraph id="8BA96D76AB646920FF17FECFFE65FE34" blockId="57.[151,1437,151,645]" pageId="57" pageNumber="58">
<emphasis id="B962B164AB646920FF17FECFFEC0FEF9" bold="true" box="[199,332,260,285]" pageId="57" pageNumber="58">Discussion.</emphasis>
The combination of very short medial margin of Cx-II/III, the box-shaped gnathosomal bay, the large and strongly trapezoid genital field, makes the new species close to 
<taxonomicName id="4C1616F5AB646920FC01FEECFF54FE80" authority="Lundblad, 1927" authorityName="Lundblad" authorityYear="1927" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="jucundus" subGenus="Monatractides">
<emphasis id="B962B164AB646920FC01FEECFBF9FEA4" box="[977,1141,295,320]" italics="true" pageId="57" pageNumber="58">Monatractides</emphasis>
(
<emphasis id="B962B164AB646920FB55FEE2FB28FEA4" box="[1157,1188,297,320]" italics="true" pageId="57" pageNumber="58">M.</emphasis>
) 
<emphasis id="B962B164AB646920FB64FEECFA95FEA4" box="[1204,1305,295,320]" italics="true" pageId="57" pageNumber="58">jucundus</emphasis>
(Lundblad, 1927)
</taxonomicName>
(syn. 
<taxonomicName id="4C1616F5AB646920FECDFE86FDCBFE80" authority="Lundblad, 1927" authorityName="Lundblad" authorityYear="1927" box="[285,583,332,357]" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="afer">
<emphasis id="B962B164AB646920FECDFE86FEFCFE80" box="[285,368,332,356]" italics="true" pageId="57" pageNumber="58">M. afer</emphasis>
(Lundblad, 1927))
</taxonomicName>
, known from East Africa (
<bibRefCitation id="EF871087AB646920FCA1FE87FBACFE81" author="Lundblad" box="[881,1056,332,357]" pageId="57" pageNumber="58" refString="Lundblad, O. (1952) Hydracarinen von den ostafrikanischen Gebirgen. Arkiv for Zoology, Serie 2, 3 (31), 391 - 525." type="journal article" year="1952">Lundblad 1952</bibRefCitation>
, 
<bibRefCitation id="EF871087AB646920FBFCFE87FAACFE81" author="Walter" box="[1068,1312,332,357]" pageId="57" pageNumber="58" refString="Walter, C. &amp; Bader, C. (1952) Hydracarina. In: Mission scientifique de l`Omo. Memoires du Museum National Histoire naturelle, Paris, (n. s.), Series A (Zoologie), 4 (3), 87 - 236." type="journal article" year="1952">Walter &amp; Bader 1952</bibRefCitation>
). The male of latter species differs from 
<taxonomicName id="4C1616F5AB646920FE39FEBAFDD1FE6C" box="[489,605,369,392]" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="urania" status="sp. nov.">
<emphasis id="B962B164AB646920FE39FEBAFDD1FE6C" box="[489,605,369,392]" italics="true" pageId="57" pageNumber="58">M. urania</emphasis>
</taxonomicName>
<emphasis id="B962B164AB646920FDB6FEA4FD24FE6C" bold="true" box="[614,680,367,392]" pageId="57" pageNumber="58">
<taxonomicNameLabel id="A2510C1FAB646920FDB6FEA4FD24FE6C" box="[614,680,367,392]" pageId="57" pageNumber="58" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, in its larger dimensions of idiosoma and palps, a frontal margin medially straight, a shoulder platelets longer than medial ones and a genital field anteriorly less pointed (see 
<bibRefCitation id="EF871087AB646920FF47FE7CFEC8FE34" author="Lundblad" box="[151,324,439,464]" pageId="57" pageNumber="58" refString="Lundblad, O. (1952) Hydracarinen von den ostafrikanischen Gebirgen. Arkiv for Zoology, Serie 2, 3 (31), 391 - 525." type="journal article" year="1952">Lundblad 1952</bibRefCitation>
, figs. 38A–B).
</paragraph>
<paragraph id="8BA96D76AB646920FF17FE17FF63FDD9" blockId="57.[151,1437,151,645]" pageId="57" pageNumber="58">
<emphasis id="B962B164AB646920FF17FE17FEB7FE11" bold="true" box="[199,315,476,501]" pageId="57" pageNumber="58">Remarks.</emphasis>
The ejaculatory complex of the male was not found, but sclerotized framework immediately above the genital field can be visible. For a discussion on the ejaculatory complex in 
<taxonomicName id="4C1616F5AB646920FBEBFDCAFB45FDFC" box="[1083,1225,511,536]" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="jucundus">
<emphasis id="B962B164AB646920FBEBFDCAFB45FDFC" box="[1083,1225,511,536]" italics="true" pageId="57" pageNumber="58">M. jucundus</emphasis>
</taxonomicName>
see the following species.
</paragraph>
<paragraph id="8BA96D76AB646920FF17FD8CFC39FD84" blockId="57.[151,1437,151,645]" box="[199,949,583,608]" pageId="57" pageNumber="58">
<emphasis id="B962B164AB646920FF17FD8CFEA6FD84" bold="true" box="[199,298,583,608]" pageId="57" pageNumber="58">Habitat.</emphasis>
A sandy/bouldary streams, shaded by riparian vegetation.
</paragraph>
</subSubSection>
<subSubSection id="C30C3EFDAB646920FF17FDA7FCCEFD61" box="[199,834,620,645]" pageId="57" pageNumber="58" type="distribution">
<paragraph id="8BA96D76AB646920FF17FDA7FCCEFD61" blockId="57.[151,1437,151,645]" box="[199,834,620,645]" pageId="57" pageNumber="58">
<emphasis id="B962B164AB646920FF17FDA7FED5FD61" bold="true" box="[199,345,620,645]" pageId="57" pageNumber="58">Distribution</emphasis>
. 
<collectingCountry id="F3012DE6AB646920FEB7FDA7FE3FFD61" box="[359,435,620,645]" name="Ghana" pageId="57" pageNumber="58">Ghana</collectingCountry>
, known only from the 
<typeStatus id="54ADD3D4AB646920FD61FDA6FD6DFD61" box="[689,737,621,645]" pageId="57" pageNumber="58">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C30C3EFDAB64695CFF47FD13FBE5FF1C" lastPageId="69" lastPageNumber="70" pageId="57" pageNumber="58" type="reference_group">
<paragraph id="8BA96D76AB646920FF47FD13FC09FD16" blockId="57.[151,990,728,787]" box="[151,901,728,754]" pageId="57" pageNumber="58">
<emphasis id="B962B164AB646920FF47FD13FC09FD16" bold="true" box="[151,901,728,754]" pageId="57" pageNumber="58">
<taxonomicName id="4C1616F5AB646920FF47FD13FD2AFD16" authority="Koenike, 1898" authorityName="Koenike" authorityYear="1898" box="[151,678,728,754]" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="microstoma">
<emphasis id="B962B164AB646920FF47FD13FE55FD16" bold="true" box="[151,473,728,754]" italics="true" pageId="57" pageNumber="58">Monatractides microstoma</emphasis>
(
<bibRefCitation id="EF871087AB646920FE39FD13FD12FD16" author="Koenike" box="[489,670,728,754]" pageId="57" pageNumber="58" refString="Koenike, F. (1898) Hydrachniden-Fauna von Madagaskar und Nossi-Be. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft in Frankfurt am Main, 21 (2), 295 - 435." type="journal article" year="1898">Koenike, 1898</bibRefCitation>
)
</taxonomicName>
—species complex
</emphasis>
</paragraph>
<paragraph id="8BA96D76AB646920FF47FD31FC52FCF7" blockId="57.[151,990,728,787]" box="[151,990,762,787]" pageId="57" pageNumber="58">
(
<figureCitation id="132D71F3AB646920FF4FFD31FEBCFCF7" box="[159,304,762,787]" captionStart="FIGURE 39 A – G" captionStartId="58.[151,250,1945,1967]" captionTargetBox="[210,1376,197,1927]" captionTargetId="figure@58.[202,1385,195,1935]" captionTargetPageId="58" captionText="FIGURE 39 A – G. Monatractides microstoma — species complex (Form '' A' '), male, Tagbo River: A = frontal margin of idiosoma, dorsal view; B = dorsal shield; C = idiosoma, ventral view; D = palp, medial view; E = palp, lateral view; F = gnathosoma; G = I-L- 5 and - 6. Scale bars = 100 Μm." httpUri="https://zenodo.org/record/286342/files/figure.png" pageId="57" pageNumber="58">Figs. 39A–G</figureCitation>
, 
<figureCitation id="132D71F3AB646920FEECFD30FE02FCF7" box="[316,398,763,787]" captionStart="FIGURE 40 A – G" captionStartId="59.[151,250,1946,1968]" captionTargetBox="[156,1421,200,1921]" captionTargetId="figure@59.[151,1436,195,1922]" captionTargetPageId="59" captionText="FIGURE 40 A – G. Monatractides microstoma - species complex (Form '' B' '), male, Apkonu stream: A = frontal margin of idiosoma, dorsal view; B = dorsal shield; C = idiosoma, ventral view; D = palp, medial view; E = palp, lateral view; F = gnathosoma; G = I-L- 5 and - 6. Scale bars = 100 Μm." httpUri="https://zenodo.org/record/286343/files/figure.png" pageId="57" pageNumber="58">40A–G</figureCitation>
, 
<figureCitation id="132D71F3AB646920FE48FD30FE62FCF7" box="[408,494,763,787]" captionStart="FIGURE 41 A – D" captionStartId="60.[151,250,1957,1979]" captionTargetBox="[215,1372,202,1919]" captionTargetId="figure@60.[198,1388,195,1933]" captionTargetPageId="60" captionText="FIGURE 41 A – D. Monatractides microstoma - species complex, female (A – B—Form '' A' ', Plunge pool Agumatsa Falls; C – D—Form '' B' ', Apkonu stream): A, C = dorsal shield; B, D = idiosoma, ventral view. Scale bar = 100 Μm." httpUri="https://zenodo.org/record/286344/files/figure.png" pageId="57" pageNumber="58">41A–D</figureCitation>
, 
<figureCitation id="132D71F3AB646920FE29FD30FDC3FCF7" box="[505,591,763,787]" captionStart="FIGURE 42 A – K. A – H" captionStartId="61.[151,250,1673,1695]" captionTargetBox="[160,1430,212,1641]" captionTargetId="figure@61.[151,1436,195,1648]" captionTargetPageId="61" captionText="FIGURE 42 A – K. A – H Monatractides microstoma - species complex, male (A—Form '' A' ', B – H—Form '' B' '; A, G – H = specimens from Tagbo River, B – E = specimens from Apkonu stream, F = specimen from Kintampo Falls): A – D = genital field; E – H = ejaculatory complex. I – K Monatractides jucundus (Lundblad, 1927) (redrawn from Lundblad 1952): I = ejaculatory complex, K = genital flap with apical part of ejaculatory complex opened, lateral view. Scale bars = 100 Μm." httpUri="https://zenodo.org/record/286345/files/figure.png" pageId="57" pageNumber="58">42A–H</figureCitation>
, 
<figureCitation id="132D71F3AB646920FD89FD30FD2FFCF7" box="[601,675,763,787]" captionStart="FIGURE 43 A – I" captionStartId="62.[151,250,1931,1953]" captionTargetBox="[165,1436,195,1911]" captionTargetId="figure@62.[165,1450,195,1912]" captionTargetPageId="62" captionText="FIGURE 43 A – I. Photographs of dorsal (A – C) and ventral shields (D – I) of Monatractides microstoma - species complex (A, C D – G—Form '' A' ', B, H – I—Form '' B' '): A, D = male from Tagbo River, E = male from Wanjakli stream, C, F = aberrant male from Kintampo Falls, G = female from Plunge pool, B – H = male from Apkonu stream, I = female from Apkonu stream." httpUri="https://zenodo.org/record/286346/files/figure.png" pageId="57" pageNumber="58">43A–I</figureCitation>
, 
<figureCitation id="132D71F3AB646920FD7FFD30FD72FCF7" box="[687,766,763,787]" captionStart="FIGURE 44 A – L" captionStartId="63.[151,250,1687,1709]" captionTargetBox="[184,1389,195,1651]" captionTargetId="figure@63.[184,1389,195,1651]" captionTargetPageId="63" captionText="FIGURE 44 A – L. Photographs of ejaculatory complex: A Monatractides convexiscutata (K. Viets, 1958), male, Asukuma River. B M. melpomene n. sp., Amedzofe Falls. C M. euterpe n. sp., creek upstream Sagyimase. D M. cf. ventriosus (K. Viets, 1916), male, Laboun River. E – L Monatractides microstoma - species complex (E – G—Form '' A' ', H – L—Form '' B' '; E – H—ejaculatory complex reduced to sclerotized framework above the genital field, I – L ejaculatory complex slide-mounted): E, L = specimens from Tagbo River, F, H – K = specimens from Apkonu stream, G = specimen from Plunge pool Agumatsa Falls." httpUri="https://zenodo.org/record/286347/files/figure.png" pageId="57" pageNumber="58">44E–L</figureCitation>
, 
<figureCitation id="132D71F3AB646920FCD9FD31FCD0FCF7" box="[777,860,762,787]" captionStart="FIGURE 46 A – C" captionStartId="68.[151,250,1283,1305]" captionTargetBox="[169,1426,483,1258]" captionTargetId="figure@68.[163,1437,473,1266]" captionTargetPageId="68" captionText="FIGURE 46 A – C. Monatractides microstoma - species complex, male, Tagbo River: A = dorsal shield; B = idiosoma, ventral view; C = slide-mounted ejaculatory complex reduced to sclerotized framework. Scale bars = 100 Μm." httpUri="https://zenodo.org/record/286349/files/figure.png" pageId="57" pageNumber="58">46A–C</figureCitation>
, 
<tableCitation id="C69458CDAB646920FCB8FD31FC5AFCF7" box="[872,982,762,787]" captionStart-0="TABLE 4" captionStart-1="TABLE 5" captionStartId-0="64.[151,239,152,175]" captionStartId-1="65.[151,239,664,687]" captionTargetBox-0="[157,1429,234,1960]" captionTargetBox-1="[151,1435,896,1662]" captionTargetPageId-0="64" captionTargetPageId-1="65" captionText-0="TABLE 4. Measurements of Monatractides microstoma (Koenike, 1898) — species complex (Form “ A “) from Ghana, and female of M. microstoma from Cameroon (from K. Viets 1958, prep. no. 1390; * calculated from figure)." captionText-1="TABLE 5. Measurements of Monatractides microstoma (Koenike, 1898) — species complex (Form “ B “) from Ghana, and M. procursa (K. O. Viets, 1974) from Congo (from K. O. Viets &amp; Böttger 1974)." httpUri-0="http://table.plazi.org/id/DF693DFEAB1D6959FF47FF53FAA8FF2B" httpUri-1="http://table.plazi.org/id/DF693DFEAB1C6958FF47FD53FB98FD2B" pageId="57" pageNumber="58" tableUuid-0="DF693DFEAB1D6959FF47FF53FAA8FF2B" tableUuid-1="DF693DFEAB1C6958FF47FD53FB98FD2B">Tabs. 4–5</tableCitation>
)
</paragraph>
<paragraph id="8BA96D76AB646920FF47FC89FE36F9CF" blockId="57.[151,1437,834,1867]" pageId="57" pageNumber="58">
<emphasis id="B962B164AB646920FF47FC89FEA3FCBF" bold="true" box="[151,303,834,859]" pageId="57" pageNumber="58">New records</emphasis>
. 
<collectingCountry id="F3012DE6AB646920FEEFFC89FE01FCBF" box="[319,397,834,859]" name="Ghana" pageId="57" pageNumber="58">Ghana</collectingCountry>
: Apkonu stream, downstream of falls, 6º 
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, 0º 
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, alt. 
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asl., 
<date id="FFA84BB6AB646920FAF3FC89FA14FCBF" box="[1315,1432,834,859]" pageId="57" pageNumber="58" value="2013-02-21">21.ii.2013</date>
, Smit, 10/14/0 (2/2/0 Form ''A'' [2/1/0 mounted] + 8/13/0 Form ''B'' [6/2/0 mounted]); Small rainforest stream upstream Sagyimase, Atewa Hills, 6º 
<geoCoordinate id="EE220BB1AB646920FD93FC41FD3EFC46" box="[579,690,906,931]" direction="north" orientation="latitude" pageId="57" pageNumber="58" precision="55" value="13.964">13.964 N</geoCoordinate>
, 0º 
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, alt. 
<quantity id="4CEEC093AB646920FC53FC41FC42FC46" box="[899,974,906,931]" metricMagnitude="2" metricUnit="m" metricValue="6.54" pageId="57" pageNumber="58" unit="m" value="654.0">654 m</quantity>
asl., 
<date id="FFA84BB6AB646920FBDFFC41FBF7FC47" box="[1039,1147,906,931]" pageId="57" pageNumber="58" value="2011-03-08">8.iii.2011</date>
, Smit, 0/1/1 (Form ''B''); Kuluga River, N of Biakpa, 6º 
<geoCoordinate id="EE220BB1AB646920FE39FC65FDD9FC22" box="[489,597,942,967]" direction="north" orientation="latitude" pageId="57" pageNumber="58" precision="55" value="51.365">51.365 N</geoCoordinate>
, 0º 
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, alt. 
<quantity id="4CEEC093AB646920FCC5FC64FCD0FC22" box="[789,860,943,967]" metricMagnitude="2" metricUnit="m" metricValue="3.88" pageId="57" pageNumber="58" unit="m" value="388.0">388 m</quantity>
asl., 
<date id="FFA84BB6AB646920FC48FC65FB87FC23" box="[920,1035,942,967]" pageId="57" pageNumber="58" value="2013-02-19">19.ii.2013</date>
, Smit, 0/1/0 (mounted) (Form ''B''); Kuluga River, upstream, hyporheic, 6º 
<geoCoordinate id="EE220BB1AB646920FD97FC19FD38FC0E" box="[583,692,978,1003]" direction="north" orientation="latitude" pageId="57" pageNumber="58" precision="55" value="51.223">51.223 N</geoCoordinate>
, 0º 
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, alt. 
<quantity id="4CEEC093AB646920FCA6FC18FC31FC0E" box="[886,957,979,1003]" metricMagnitude="2" metricUnit="m" metricValue="4.1" pageId="57" pageNumber="58" unit="m" value="410.0">410 m</quantity>
asl., 
<date id="FFA84BB6AB646920FC2AFC19FBE2FC0F" box="[1018,1134,978,1003]" pageId="57" pageNumber="58" value="2013-02-20">20.ii.2013</date>
, Smit, 4/3 (one juvenile)/0 (3/0/0 mounted) (1/0/0 Form ''A'' [mounted] + 2/3/0 Form ''B'' [2/0/0 mounted]); Agumatsa River at first bridge, Agumatsa Wildlife Sanctuary, 7º 
<geoCoordinate id="EE220BB1AB646920FDD7FBD1FDF8FBD6" box="[519,628,1050,1075]" direction="north" orientation="latitude" pageId="57" pageNumber="58" precision="55" value="6.83">06.830 N</geoCoordinate>
, 0º 
<geoCoordinate id="EE220BB1AB646920FD46FBD1FD73FBD6" box="[662,767,1050,1075]" direction="east" orientation="longitude" pageId="57" pageNumber="58" precision="55" value="35.76">35.760 E</geoCoordinate>
, alt. 
<quantity id="4CEEC093AB646920FCE7FBD1FCF3FBD6" box="[823,895,1050,1075]" metricMagnitude="2" metricUnit="m" metricValue="2.53" pageId="57" pageNumber="58" unit="m" value="253.0">253 m</quantity>
asl., 
<date id="FFA84BB6AB646920FC6CFBD1FBBDFBD7" box="[956,1073,1050,1075]" pageId="57" pageNumber="58" value="2013-02-22">22.ii.2013</date>
, Smit, 5/9/0 (Form ''A''); stream Boti Falls, 6º 
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, 0º 
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, alt. 
<quantity id="4CEEC093AB646920FD89FBF4FD2CFBB2" box="[601,672,1087,1111]" metricMagnitude="2" metricUnit="m" metricValue="2.73" pageId="57" pageNumber="58" unit="m" value="273.0">273 m</quantity>
asl, 
<date id="FFA84BB6AB646920FD03FBF5FCCCFBB3" box="[723,832,1086,1111]" pageId="57" pageNumber="58" value="2011-03-09">9.iii.2011</date>
, Smit, 0/1/0 (Form ''B''); Kintampo Falls, 8º 
<geoCoordinate id="EE220BB1AB646920FAFBFBF5FA14FBB2" box="[1323,1432,1086,1111]" direction="north" orientation="latitude" pageId="57" pageNumber="58" precision="55" value="5.413">05.413 N</geoCoordinate>
, 1º 
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, alt. 
<quantity id="4CEEC093AB646920FE82FBA9FE15FB9E" box="[338,409,1122,1147]" metricMagnitude="2" metricUnit="m" metricValue="2.35" pageId="57" pageNumber="58" unit="m" value="235.0">235 m</quantity>
asl, 
<date id="FFA84BB6AB646920FE1DFBA9FDB6FB9F" box="[461,570,1122,1147]" pageId="57" pageNumber="58" value="2011-03-03">3.iii.2011</date>
, Smit, 8/7 (one juvenile)/0 (3/1/0 mounted) (4/5/0 Form ''A'' [1/1/0 mounted] + 4/2/0 Form ''B'' [2/0/0 mounted]); Afiaso stream, Kakum NP, 5º 
<geoCoordinate id="EE220BB1AB646920FCA2FB4DFC53FB7A" box="[882,991,1158,1183]" direction="north" orientation="latitude" pageId="57" pageNumber="58" precision="55" value="30.087">30.087 N</geoCoordinate>
, 1º 
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, alt. 
<quantity id="4CEEC093AB646920FB73FB4CFB65FB7A" box="[1187,1257,1159,1183]" metricMagnitude="2" metricUnit="m" metricValue="1.14" pageId="57" pageNumber="58" unit="m" value="114.0">114 m</quantity>
asl., 
<date id="FFA84BB6AB646920FAF3FB4DFA14FB7B" box="[1315,1432,1158,1183]" pageId="57" pageNumber="58" value="2013-02-12">12.ii.2013</date>
, Smit, 0/1/0 (Form A); Fuller Falls, 8º 
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, 1º 
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, alt. 
<quantity id="4CEEC093AB646920FC5AFB60FC58FB26" box="[906,980,1195,1219]" metricMagnitude="2" metricUnit="m" metricValue="1.89" pageId="57" pageNumber="58" unit="m" value="189.0">189 m</quantity>
asl, 
<date id="FFA84BB6AB646920FBDEFB61FBF7FB27" box="[1038,1147,1194,1219]" pageId="57" pageNumber="58" value="2011-03-06">6.iii.2011</date>
, Smit, 0/1/0 (Form ''A''); Supon stream near Asiakwa, Atewa Hills, 6º 
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, 0º 
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, alt. 
<quantity id="4CEEC093AB646920FC03FB05FB90FB02" box="[979,1052,1230,1255]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="57" pageNumber="58" unit="m" value="250.0">250 m</quantity>
asl, 
<date id="FFA84BB6AB646920FB84FB05FB4CFB03" box="[1108,1216,1230,1255]" pageId="57" pageNumber="58" value="2011-03-07">7.iii.2011</date>
, Smit, 1/0/0 (Form ''A''); Tagbo River, downstream of falls, 7º 
<geoCoordinate id="EE220BB1AB646920FD5DFB38FD72FAEE" box="[653,766,1267,1291]" direction="north" orientation="latitude" pageId="57" pageNumber="58" precision="55" value="0.708">00.708 N</geoCoordinate>
, 0º 
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, alt. 
<quantity id="4CEEC093AB646920FC00FB38FB97FAEE" box="[976,1051,1267,1291]" metricMagnitude="2" metricUnit="m" metricValue="3.94" pageId="57" pageNumber="58" unit="m" value="394.0">394 m</quantity>
asl., 
<date id="FFA84BB6AB646920FBB0FB39FB5FFAEF" box="[1120,1235,1266,1291]" pageId="57" pageNumber="58" value="2013-02-23">23.ii.2013</date>
, Smit, 19/11(one juvenile)/0 (6/2/0 mounted) (10/5/0 Form ''A'' [4/1/0 mounted] +8/6/0 Form ''B'' [2/1/0 mounted]); Akaa Falls, 6º 
<geoCoordinate id="EE220BB1AB646920FF7DFAF1FE92FAB6" box="[173,286,1338,1363]" direction="north" orientation="latitude" pageId="57" pageNumber="58" precision="55" value="10.516">10.516 N</geoCoordinate>
, 0º 
<geoCoordinate id="EE220BB1AB646920FE96FAF0FE3FFAB6" box="[326,435,1339,1363]" direction="east" orientation="longitude" pageId="57" pageNumber="58" precision="55" value="11.723">11.723 E</geoCoordinate>
, alt. 
<quantity id="4CEEC093AB646920FE25FAF0FDCEFAB6" box="[501,578,1339,1363]" metricMagnitude="2" metricUnit="m" metricValue="1.8" pageId="57" pageNumber="58" unit="m" value="180.0">180 m</quantity>
asl., 
<date id="FFA84BB6AB646920FD59FAF1FD79FAB7" box="[649,757,1338,1363]" pageId="57" pageNumber="58" value="2011-03-09">9.iii.2011</date>
, Smit, 
<date id="FFA84BB6AB646920FC80FAF0FC1BFAB7" box="[848,919,1339,1363]" pageId="57" pageNumber="58" value="2012-08-01">8/12/1</date>
(1/0/0 mounted) (Form ''B''); Nubui River, Agumatsa Wildlife Sanctuary, 7º 
<geoCoordinate id="EE220BB1AB646920FDD2FA95FDE2FA92" box="[514,622,1374,1399]" direction="north" orientation="latitude" pageId="57" pageNumber="58" precision="55" value="6.986">06.986 N</geoCoordinate>
, 0º 
<geoCoordinate id="EE220BB1AB646920FD5FFA95FD7BFA92" box="[655,759,1374,1399]" direction="east" orientation="longitude" pageId="57" pageNumber="58" precision="55" value="35.548">35.548 E</geoCoordinate>
, alt. 
<quantity id="4CEEC093AB646920FCFCFA95FCFEFA92" box="[812,882,1374,1399]" metricMagnitude="2" metricUnit="m" metricValue="2.54" pageId="57" pageNumber="58" unit="m" value="254.0">254 m</quantity>
asl., 
<date id="FFA84BB6AB646920FC7CFA95FBACFA93" box="[940,1056,1374,1399]" pageId="57" pageNumber="58" value="2013-02-22">22.ii.2013</date>
, Smit, 10/14(
<specimenCount id="9D10A6FFAB646920FB67FA94FABEFA93" box="[1207,1330,1374,1399]" pageId="57" pageNumber="58" type="juvenile">2 juveniles</specimenCount>
)/0 (Form ''A''); Laboun River, downstream of falls, Kyabobo NP, 8º 
<geoCoordinate id="EE220BB1AB646920FCF9FA49FC19FA7E" box="[809,917,1410,1435]" direction="north" orientation="latitude" pageId="57" pageNumber="58" precision="55" value="19.836">19.836 N</geoCoordinate>
, 0º 
<geoCoordinate id="EE220BB1AB646920FC6FFA49FBAAFA7E" box="[959,1062,1410,1435]" direction="east" orientation="longitude" pageId="57" pageNumber="58" precision="55" value="35.487">35.487 E</geoCoordinate>
, alt. 
<quantity id="4CEEC093AB646920FB8DFA48FB28FA7E" box="[1117,1188,1411,1435]" metricMagnitude="2" metricUnit="m" metricValue="3.42" pageId="57" pageNumber="58" unit="m" value="342.0">342 m</quantity>
asl., 
<date id="FFA84BB6AB646920FB30FA49FAD9FA7F" box="[1248,1365,1410,1435]" pageId="57" pageNumber="58" value="2013-02-24">24.ii.2013</date>
, Smit, 2/0/0 (1/0/0 Form ''A'' + 1/0/0 Form ''B''); stream downstream of falls, Wanjakli River, Likpe Todome, 7º 
<geoCoordinate id="EE220BB1AB646920FAFBFA6DFA14FA5A" box="[1323,1432,1446,1471]" direction="north" orientation="latitude" pageId="57" pageNumber="58" precision="55" value="9.976">09.976 N</geoCoordinate>
, 0º 
<geoCoordinate id="EE220BB1AB646920FF7DFA01FE94FA06" box="[173,280,1482,1507]" direction="east" orientation="longitude" pageId="57" pageNumber="58" precision="55" value="36.456">36.456 E</geoCoordinate>
, alt. 
<quantity id="4CEEC093AB646920FE86FA00FE2DFA06" box="[342,417,1483,1507]" metricMagnitude="2" metricUnit="m" metricValue="4.779999999999999" pageId="57" pageNumber="58" unit="m" value="478.0">478 m</quantity>
asl., 
<date id="FFA84BB6AB646920FE34FA01FDD4FA07" box="[484,600,1482,1507]" pageId="57" pageNumber="58" value="2013-02-23">23.ii.2013</date>
, Smit, 3/9/2 (1/1/0 Form ''A'' + 0/1/0 Form ''B'' mounted); Plunge pool Agumatsa Falls, Agumatsa Wildlife Sanctuary, 7º 
<geoCoordinate id="EE220BB1AB646920FD10FA25FCA1F9E2" box="[704,813,1518,1543]" direction="north" orientation="latitude" pageId="57" pageNumber="58" precision="55" value="6.35">06.350 N</geoCoordinate>
, 0º 
<geoCoordinate id="EE220BB1AB646920FC9EFA25FC39F9E2" box="[846,949,1518,1543]" direction="east" orientation="longitude" pageId="57" pageNumber="58" precision="55" value="36.476">36.476 E</geoCoordinate>
, alt. 
<quantity id="4CEEC093AB646920FC3BFA25FBBEF9E2" box="[1003,1074,1518,1543]" metricMagnitude="2" metricUnit="m" metricValue="2.58" pageId="57" pageNumber="58" unit="m" value="258.0">258 m</quantity>
asl., 
<date id="FFA84BB6AB646920FBBDFA25FB6DF9E3" box="[1133,1249,1518,1543]" pageId="57" pageNumber="58" value="2013-02-22">22.ii.2013</date>
, Smit, 12/8/0 (1/ 2/0 mounted) (Form ''A'').
</paragraph>
<paragraph id="8BA96D76AB646920FF17F9FDFE7BF8AF" blockId="57.[151,1437,834,1867]" pageId="57" pageNumber="58">
<emphasis id="B962B164AB646920FF17F9FDFEEDF9AB" bold="true" box="[199,353,1590,1615]" pageId="57" pageNumber="58">Morphology.</emphasis>
<emphasis id="B962B164AB646920FEB7F9FDFDA8F9AB" box="[359,548,1590,1615]" italics="true" pageId="57" pageNumber="58">General features</emphasis>
—Idiosoma elongated oval; frontal platelets only slightly shorter than shoulder platelets; frontal margin medially straight or slightly convex, between flat, anterolaterally pointed protrusions (
<figureCitation id="132D71F3AB646920FF4FF9B5FE9FF973" box="[159,275,1662,1687]" captionStart="FIGURE 39 A – G" captionStartId="58.[151,250,1945,1967]" captionTargetBox="[210,1376,197,1927]" captionTargetId="figure@58.[202,1385,195,1935]" captionTargetPageId="58" captionText="FIGURE 39 A – G. Monatractides microstoma — species complex (Form '' A' '), male, Tagbo River: A = frontal margin of idiosoma, dorsal view; B = dorsal shield; C = idiosoma, ventral view; D = palp, medial view; E = palp, lateral view; F = gnathosoma; G = I-L- 5 and - 6. Scale bars = 100 Μm." httpUri="https://zenodo.org/record/286342/files/figure.png" pageId="57" pageNumber="58">Figs. 39A</figureCitation>
, 
<figureCitation id="132D71F3AB646920FEF1F9B4FEDAF973" box="[289,342,1663,1687]" captionStart="FIGURE 40 A – G" captionStartId="59.[151,250,1946,1968]" captionTargetBox="[156,1421,200,1921]" captionTargetId="figure@59.[151,1436,195,1922]" captionTargetPageId="59" captionText="FIGURE 40 A – G. Monatractides microstoma - species complex (Form '' B' '), male, Apkonu stream: A = frontal margin of idiosoma, dorsal view; B = dorsal shield; C = idiosoma, ventral view; D = palp, medial view; E = palp, lateral view; F = gnathosoma; G = I-L- 5 and - 6. Scale bars = 100 Μm." httpUri="https://zenodo.org/record/286343/files/figure.png" pageId="57" pageNumber="58">40A</figureCitation>
); Cxgl–4 located far anteriorly, near tips of Cx-I; gnathosomal bay moderately deep, U-shaped, proximally rectangular box-shaped; suture line of Cx-IV strongly extending posteriorly beyond posterior margin of genital field, laterally curved (
<figureCitation id="132D71F3AB646920FDD8F90DFD0CF93B" box="[520,640,1734,1759]" captionStart="FIGURE 39 A – G" captionStartId="58.[151,250,1945,1967]" captionTargetBox="[210,1376,197,1927]" captionTargetId="figure@58.[202,1385,195,1935]" captionTargetPageId="58" captionText="FIGURE 39 A – G. Monatractides microstoma — species complex (Form '' A' '), male, Tagbo River: A = frontal margin of idiosoma, dorsal view; B = dorsal shield; C = idiosoma, ventral view; D = palp, medial view; E = palp, lateral view; F = gnathosoma; G = I-L- 5 and - 6. Scale bars = 100 Μm." httpUri="https://zenodo.org/record/286342/files/figure.png" pageId="57" pageNumber="58">Figs. 39C</figureCitation>
, 
<figureCitation id="132D71F3AB646920FD43F90CFD4AF93B" box="[659,710,1735,1759]" captionStart="FIGURE 40 A – G" captionStartId="59.[151,250,1946,1968]" captionTargetBox="[156,1421,200,1921]" captionTargetId="figure@59.[151,1436,195,1922]" captionTargetPageId="59" captionText="FIGURE 40 A – G. Monatractides microstoma - species complex (Form '' B' '), male, Apkonu stream: A = frontal margin of idiosoma, dorsal view; B = dorsal shield; C = idiosoma, ventral view; D = palp, medial view; E = palp, lateral view; F = gnathosoma; G = I-L- 5 and - 6. Scale bars = 100 Μm." httpUri="https://zenodo.org/record/286343/files/figure.png" pageId="57" pageNumber="58">40C</figureCitation>
); excretory pore and Vgl–2 away from the line of primary sclerotization, excretory pore anterior to Vgl–2; ventral seta on P-4 short (
<figureCitation id="132D71F3AB646920FC27F921FB00F8E7" box="[1015,1164,1770,1795]" captionStart="FIGURE 39 A – G" captionStartId="58.[151,250,1945,1967]" captionTargetBox="[210,1376,197,1927]" captionTargetId="figure@58.[202,1385,195,1935]" captionTargetPageId="58" captionText="FIGURE 39 A – G. Monatractides microstoma — species complex (Form '' A' '), male, Tagbo River: A = frontal margin of idiosoma, dorsal view; B = dorsal shield; C = idiosoma, ventral view; D = palp, medial view; E = palp, lateral view; F = gnathosoma; G = I-L- 5 and - 6. Scale bars = 100 Μm." httpUri="https://zenodo.org/record/286342/files/figure.png" pageId="57" pageNumber="58">Figs. 39D–E</figureCitation>
, 
<figureCitation id="132D71F3AB646920FB4AF920FB62F8E7" box="[1178,1262,1771,1795]" captionStart="FIGURE 40 A – G" captionStartId="59.[151,250,1946,1968]" captionTargetBox="[156,1421,200,1921]" captionTargetId="figure@59.[151,1436,195,1922]" captionTargetPageId="59" captionText="FIGURE 40 A – G. Monatractides microstoma - species complex (Form '' B' '), male, Apkonu stream: A = frontal margin of idiosoma, dorsal view; B = dorsal shield; C = idiosoma, ventral view; D = palp, medial view; E = palp, lateral view; F = gnathosoma; G = I-L- 5 and - 6. Scale bars = 100 Μm." httpUri="https://zenodo.org/record/286343/files/figure.png" pageId="57" pageNumber="58">40D–E</figureCitation>
). Male: medial margin of Cx-II/III relatively short; genital field elongated, anteriorly enlarged. Female: genital field elongated pentagonal, anteriorly rounded.
</paragraph>
<caption id="DF693DFEAB676923FF47F852FD20F816" httpUri="https://zenodo.org/record/286342/files/figure.png" pageId="58" pageNumber="59" targetBox="[210,1376,197,1927]" targetPageId="58">
<paragraph id="8BA96D76AB676923FF47F852FD20F816" blockId="58.[151,1436,1945,2034]" pageId="58" pageNumber="59">
<emphasis id="B962B164AB676923FF47F852FEDCF84B" bold="true" box="[151,336,1945,1967]" pageId="58" pageNumber="59">FIGURE 39A–G.</emphasis>
<taxonomicName id="4C1616F5AB676923FE8FF852FDFAF84B" box="[351,630,1945,1967]" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="microstoma">
<emphasis id="B962B164AB676923FE8FF852FDFAF84B" box="[351,630,1945,1967]" italics="true" pageId="58" pageNumber="59">Monatractides microstoma</emphasis>
</taxonomicName>
—species complex (Form ''A''), male, Tagbo River: A = frontal margin of idiosoma, dorsal view; B = dorsal shield; C = idiosoma, ventral view; D = palp, medial view; E = palp, lateral view; F = gnathosoma; G = I-L-5 and -6. Scale bars = 100 Μm.
</paragraph>
</caption>
<caption id="DF693DFEAB666922FF47F851FD20F817" httpUri="https://zenodo.org/record/286343/files/figure.png" pageId="59" pageNumber="60" targetBox="[156,1421,200,1921]" targetPageId="59">
<paragraph id="8BA96D76AB666922FF47F851FD20F817" blockId="59.[151,1436,1946,2035]" pageId="59" pageNumber="60">
<emphasis id="B962B164AB666922FF47F851FEC2F854" bold="true" box="[151,334,1946,1968]" pageId="59" pageNumber="60">FIGURE 40A–G.</emphasis>
<taxonomicName id="4C1616F5AB666922FE8BF851FDE3F854" box="[347,623,1946,1968]" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="microstoma">
<emphasis id="B962B164AB666922FE8BF851FDE3F854" box="[347,623,1946,1968]" italics="true" pageId="59" pageNumber="60">Monatractides microstoma</emphasis>
</taxonomicName>
- species complex (Form ''B''), male, Apkonu stream: A = frontal margin of idiosoma, dorsal view; B = dorsal shield; C = idiosoma, ventral view; D = palp, medial view; E = palp, lateral view; F = gnathosoma; G = I-L-5 and -6. Scale bars = 100 Μm.
</paragraph>
</caption>
<caption id="DF693DFEAB616925FF47F86EFB6CF838" httpUri="https://zenodo.org/record/286344/files/figure.png" pageId="60" pageNumber="61" targetBox="[215,1372,202,1919]" targetPageId="60">
<paragraph id="8BA96D76AB616925FF47F86EFB6CF838" blockId="60.[151,1436,1957,2012]" pageId="60" pageNumber="61">
<emphasis id="B962B164AB616925FF47F86EFED8F85F" bold="true" box="[151,340,1957,1979]" pageId="60" pageNumber="61">FIGURE 41A–D.</emphasis>
<taxonomicName id="4C1616F5AB616925FE8FF86EFDF9F85F" box="[351,629,1957,1979]" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="microstoma">
<emphasis id="B962B164AB616925FE8FF86EFDF9F85F" box="[351,629,1957,1979]" italics="true" pageId="60" pageNumber="61">Monatractides microstoma</emphasis>
</taxonomicName>
- species complex, female (A–B—Form ''A'', Plunge pool Agumatsa Falls; C–D—Form ''B'', Apkonu stream): A, C = dorsal shield; B, D = idiosoma, ventral view. Scale bar = 100 Μm.
</paragraph>
</caption>
<caption id="DF693DFEAB606924FF47F942FB51F8E7" httpUri="https://zenodo.org/record/286345/files/figure.png" pageId="61" pageNumber="62" targetBox="[160,1430,212,1641]" targetPageId="61">
<paragraph id="8BA96D76AB606924FF47F942FB51F8E7" blockId="61.[151,1436,1673,1795]" pageId="61" pageNumber="62">
<emphasis id="B962B164AB606924FF47F942FE1DF97B" bold="true" box="[151,401,1673,1695]" pageId="61" pageNumber="62">FIGURE 42A–K. A–H</emphasis>
<taxonomicName id="4C1616F5AB606924FE4CF942FD3DF97B" box="[412,689,1673,1695]" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="microstoma">
<emphasis id="B962B164AB606924FE4CF942FD3DF97B" box="[412,689,1673,1695]" italics="true" pageId="61" pageNumber="62">Monatractides microstoma</emphasis>
</taxonomicName>
- species complex, male (A—Form ''A'', B–H—Form ''B''; A, G–H = specimens from Tagbo River, B–E = specimens from Apkonu stream, F = specimen from Kintampo Falls): A–D = genital field; E–H = ejaculatory complex. 
<emphasis id="B962B164AB606924FE13F907FE60F906" bold="true" box="[451,492,1740,1762]" pageId="61" pageNumber="62">I–K</emphasis>
<taxonomicName id="4C1616F5AB606924FE24F907FC24F906" authority="Lundblad, 1927" authorityName="Lundblad" authorityYear="1927" box="[500,936,1740,1762]" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="jucundus">
<emphasis id="B962B164AB606924FE24F907FD04F906" box="[500,648,1740,1762]" italics="true" pageId="61" pageNumber="62">Monatractides</emphasis>
<emphasis id="B962B164AB606924FD41F907FD61F906" box="[657,749,1740,1762]" italics="true" pageId="61" pageNumber="62">jucundus</emphasis>
(Lundblad, 1927)
</taxonomicName>
(redrawn from Lundblad 1952): I = ejaculatory complex, K = genital flap with apical part of ejaculatory complex opened, lateral view. Scale bars =100 Μm.
</paragraph>
</caption>
<caption id="DF693DFEAB636927FF47F840FADBF800" httpUri="https://zenodo.org/record/286346/files/figure.png" pageId="62" pageNumber="63" targetBox="[165,1436,195,1911]" targetPageId="62">
<paragraph id="8BA96D76AB636927FF47F840FADBF800" blockId="62.[151,1436,1931,2020]" pageId="62" pageNumber="63">
<emphasis id="B962B164AB636927FF47F840FECCF845" bold="true" box="[151,320,1931,1953]" pageId="62" pageNumber="63">FIGURE 43A–I</emphasis>
. Photographs of dorsal (A–C) and ventral shields (D–I) of 
<taxonomicName id="4C1616F5AB636927FC42F840FB2FF845" box="[914,1187,1931,1953]" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="microstoma">
<emphasis id="B962B164AB636927FC42F840FB2FF845" box="[914,1187,1931,1953]" italics="true" pageId="62" pageNumber="63">Monatractides microstoma</emphasis>
</taxonomicName>
- species complex (A, C D–G—Form ''A'', B, H–I—Form ''B''): A, D = male from Tagbo River, E = male from Wanjakli stream, C, F = aberrant male from Kintampo Falls, G = female from Plunge pool, B–H = male from Apkonu stream, I = female from Apkonu stream.
</paragraph>
</caption>
<caption id="DF693DFEAB626926FF47F95CFF43F8A3" httpUri="https://zenodo.org/record/286347/files/figure.png" pageId="63" pageNumber="64" targetBox="[184,1389,195,1651]" targetPageId="63">
<paragraph id="8BA96D76AB626926FF47F95CFF43F8A3" blockId="63.[151,1437,1687,1863]" pageId="63" pageNumber="64">
<emphasis id="B962B164AB626926FF47F95CFEC7F949" bold="true" box="[151,331,1687,1709]" pageId="63" pageNumber="64">FIGURE 44A–L</emphasis>
. Photographs of ejaculatory complex: A 
<taxonomicName id="4C1616F5AB626926FCD1F95CFB60F949" authority="K. Viets, 1958" authorityName="K. Viets" authorityYear="1958" box="[769,1260,1687,1710]" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="convexiscutata">
<emphasis id="B962B164AB626926FCD1F95CFC19F949" box="[769,917,1687,1709]" italics="true" pageId="63" pageNumber="64">Monatractides</emphasis>
<emphasis id="B962B164AB626926FC4FF952FBB9F94A" box="[927,1077,1689,1710]" italics="true" pageId="63" pageNumber="64">convexiscutata</emphasis>
(K. Viets, 1958)
</taxonomicName>
, male, Asukuma River. B 
<taxonomicName id="4C1616F5AB626926FF22F97CFE0BF928" box="[242,391,1718,1740]" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="melpomene" status="sp. nov.">
<emphasis id="B962B164AB626926FF22F97CFE0BF928" box="[242,391,1718,1740]" italics="true" pageId="63" pageNumber="64">M. melpomene</emphasis>
</taxonomicName>
<emphasis id="B962B164AB626926FE5EF97DFE4BF928" bold="true" box="[398,455,1718,1740]" pageId="63" pageNumber="64">
<taxonomicNameLabel id="A2510C1FAB626926FE5EF97DFE4BF928" box="[398,455,1718,1740]" pageId="63" pageNumber="64" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, Amedzofe Falls. C 
<taxonomicName id="4C1616F5AB626926FD4AF97CFC84F928" box="[666,776,1719,1740]" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="euterpe" status="sp. nov.">
<emphasis id="B962B164AB626926FD4AF97CFC84F928" box="[666,776,1719,1740]" italics="true" pageId="63" pageNumber="64">M. euterpe</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2510C1FAB626926FCDEF97DFCC4F928" box="[782,840,1718,1740]" pageId="63" pageNumber="64" rank="species">
<emphasis id="B962B164AB626926FCDEF97DFCC4F928" bold="true" box="[782,840,1718,1740]" pageId="63" pageNumber="64">n. sp.</emphasis>
</taxonomicNameLabel>
, creek upstream Sagyimase. D 
<taxonomicName id="4C1616F5AB626926FB59F97CFF58F90F" authority="K. Viets, 1916" authorityName="K. Viets" authorityYear="1916" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="ventriosus">
<emphasis id="B962B164AB626926FB59F97CFB29F928" box="[1161,1189,1719,1740]" italics="true" pageId="63" pageNumber="64">M.</emphasis>
cf. 
<emphasis id="B962B164AB626926FB1BF97CFABFF928" box="[1227,1331,1719,1740]" italics="true" pageId="63" pageNumber="64">ventriosus</emphasis>
(K. Viets, 1916)
</taxonomicName>
, male, Laboun River. E–L 
<taxonomicName id="4C1616F5AB626926FDC3F91EFCA1F90F" box="[531,813,1749,1771]" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="microstoma">
<emphasis id="B962B164AB626926FDC3F91EFCA1F90F" box="[531,813,1749,1771]" italics="true" pageId="63" pageNumber="64">Monatractides microstoma</emphasis>
</taxonomicName>
- species complex (E–G—Form ''A'', H–L—Form ''B''; E–H—ejaculatory complex reduced to sclerotized framework above the genital field, I–L ejaculatory complex slide-mounted): E, L = specimens from Tagbo River, F, H–K = specimens from Apkonu stream, G = specimen from Plunge pool Agumatsa Falls.
</paragraph>
</caption>
<caption id="DF693DFEAB1D6959FF47FF53FAA8FF2B" ID-Table-UUID="DF693DFEAB1D6959FF47FF53FAA8FF2B" httpUri="http://table.plazi.org/id/DF693DFEAB1D6959FF47FF53FAA8FF2B" pageId="64" pageNumber="65" targetBox="[157,1429,234,1960]" targetIsTable="true" targetPageId="64">
<paragraph id="8BA96D76AB1D6959FF47FF53FAA8FF2B" blockId="64.[151,1436,152,207]" pageId="64" pageNumber="65">
<emphasis id="B962B164AB1D6959FF47FF53FE80FF4B" bold="true" box="[151,268,152,175]" pageId="64" pageNumber="65">TABLE 4.</emphasis>
Measurements of 
<taxonomicName id="4C1616F5AB1D6959FE0AFF53FC39FF4B" authority="Koenike, 1898" authorityName="Koenike" authorityYear="1898" box="[474,949,152,175]" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="microstoma">
<emphasis id="B962B164AB1D6959FE0AFF53FDF9FF4B" box="[474,629,152,175]" italics="true" pageId="64" pageNumber="65">Monatractides</emphasis>
<emphasis id="B962B164AB1D6959FDAEFF53FD77FF4B" box="[638,763,152,175]" italics="true" pageId="64" pageNumber="65">microstoma</emphasis>
(Koenike, 1898)
</taxonomicName>
—species complex (Form “A“) from Ghana, and female of 
<taxonomicName id="4C1616F5AB1D6959FEE0FF73FE5DFF2B" box="[304,465,184,207]" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="microstoma">
<emphasis id="B962B164AB1D6959FEE0FF73FE5DFF2B" box="[304,465,184,207]" italics="true" pageId="64" pageNumber="65">M. microstoma</emphasis>
</taxonomicName>
from Cameroon (from K. Viets 1958, prep. no. 1390; * calculated from figure).
</paragraph>
</caption>
<paragraph id="8BA96D76AB1D6959FE3DFF21FB6BF84C" pageId="64" pageNumber="65">
<table id="F9169FD6AB1D96E6FF4DFF21FA19F84C" box="[157,1429,234,1960]" gridcols="4" gridrows="15" pageId="64" pageNumber="65">
<tr id="35266F34AB1D96E6FF4DFF21FA19FEB8" box="[157,1429,234,348]" gridrow="0" pageId="64" pageNumber="65" rowspan-0="1">
<th id="76F70648AB1D96E6FE3DFF21FC42FEB8" box="[493,974,234,348]" gridcol="1" gridrow="0" pageId="64" pageNumber="65">Tagbo River ♂, Plunge pool, ♂ Plunge pool, n = 2 2♀</th>
<th id="76F70648AB1D96E6FC26FF21FB03FEB8" box="[1014,1167,234,348]" gridcol="2" gridrow="0" pageId="64" pageNumber="65">
<taxonomicName id="4C1616F5AB1D6959FC26FF20FBA7FEDA" authority="Cook, 1966" authorityName="Cook" authorityYear="1966" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="microstoma">
<emphasis id="B962B164AB1D6959FC26FF20FB03FEE4" box="[1014,1167,235,256]" italics="true" pageId="64" pageNumber="65">M. microstoma</emphasis>
sensu Cook, 1966
</taxonomicName>
, Tagbo River, ♂
</th>
<th id="76F70648AB1D96E6FB6BFF21FA19FEB8" box="[1211,1429,234,348]" gridcol="3" gridrow="0" pageId="64" pageNumber="65">
K. Viets, 1958 Cameroon, 
<taxonomicName id="4C1616F5AB1D6959FAE1FEC1FABDFEDA" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="microstoma">
<emphasis id="B962B164AB1D6959FAE1FEC1FABDFEDA" italics="true" pageId="64" pageNumber="65">M. microstoma</emphasis>
</taxonomicName>
, prep. no. 1390, ♀
</th>
</tr>
<tr id="35266F34AB1D96E6FF4DFEBBFA19FE3C" box="[157,1429,368,472]" gridrow="1" pageId="64" pageNumber="65">
<th id="76F70648AB1D96E6FF4DFEBBFE35FE3C" box="[157,441,368,472]" gridcol="0" gridrow="1" pageId="64" pageNumber="65">Idiosoma L Idiosoma W Idiosoma L/W ratio</th>
<td id="76F70648AB1D96E6FE3DFEBBFC42FE3C" box="[493,974,368,472]" gridcol="1" gridrow="1" pageId="64" pageNumber="65">573–647 641 675–703 398–447 459 466–506 1.44–1.45 1.4 1.39–1.45</td>
<td id="76F70648AB1D96E6FC26FEBBFB03FE3C" box="[1014,1167,368,472]" gridcol="2" gridrow="1" pageId="64" pageNumber="65">733 476 1.54</td>
<td id="76F70648AB1D96E6FB6BFEBBFA19FE3C" box="[1211,1429,368,472]" gridcol="3" gridrow="1" pageId="64" pageNumber="65">640 535 1.2</td>
</tr>
<tr id="35266F34AB1D96E6FF4DFE27FA19FDB0" box="[157,1429,492,596]" gridrow="2" pageId="64" pageNumber="65">
<th id="76F70648AB1D96E6FF4DFE27FE35FDB0" box="[157,441,492,596]" gridcol="0" gridrow="2" pageId="64" pageNumber="65">Ds L Ds W Ds L/W ratio</th>
<td id="76F70648AB1D96E6FE3DFE27FC42FDB0" box="[493,974,492,596]" gridcol="1" gridrow="2" pageId="64" pageNumber="65">468–533 534 548–586 340–381 381 389–405 1.38–1.4 1.4 1.41–1.45</td>
<td id="76F70648AB1D96E6FC26FE27FB03FDB0" box="[1014,1167,492,596]" gridcol="2" gridrow="2" pageId="64" pageNumber="65">600 406 1.48</td>
<td id="76F70648AB1D96E6FB6BFE27FA19FDB0" box="[1211,1429,492,596]" gridcol="3" gridrow="2" pageId="64" pageNumber="65">590 420 1.4</td>
</tr>
<tr id="35266F34AB1D96E6FF4DFDA3FA19FD34" box="[157,1429,616,720]" gridrow="3" pageId="64" pageNumber="65">
<th id="76F70648AB1D96E6FF4DFDA3FE35FD34" box="[157,441,616,720]" gridcol="0" gridrow="3" pageId="64" pageNumber="65">Dp L Sh plate L Sh plate W</th>
<td id="76F70648AB1D96E6FE3DFDA3FC42FD34" box="[493,974,616,720]" gridcol="1" gridrow="3" pageId="64" pageNumber="65">436–491 497 513–545 109–131 122–123 125–127 42–69 50 45–55</td>
<td id="76F70648AB1D96E6FC26FDA3FB03FD34" box="[1014,1167,616,720]" gridcol="2" gridrow="3" pageId="64" pageNumber="65">550 169–172 68–73</td>
<td id="76F70648AB1D96E6FB6BFDA3FA19FD34" box="[1211,1429,616,720]" gridcol="3" gridrow="3" pageId="64" pageNumber="65">525 145 -</td>
</tr>
<tr id="35266F34AB1D96E6FF4DFD2FFA19FCA8" box="[157,1429,740,844]" gridrow="4" pageId="64" pageNumber="65">
<th id="76F70648AB1D96E6FF4DFD2FFE35FCA8" box="[157,441,740,844]" gridcol="0" gridrow="4" pageId="64" pageNumber="65">Sh plate L/W F plate L F plate W</th>
<td id="76F70648AB1D96E6FE3DFD2FFC42FCA8" box="[493,974,740,844]" gridcol="1" gridrow="4" pageId="64" pageNumber="65">1.9–2.6 2.4–2.5 2.3–2.8 108–122 125–127 116–125 45–56 53 50–55</td>
<td id="76F70648AB1D96E6FC26FD2FFB03FCA8" box="[1014,1167,740,844]" gridcol="2" gridrow="4" pageId="64" pageNumber="65">2.3–2.5 142–144 72</td>
<td id="76F70648AB1D96E6FB6BFD2FFA19FCA8" box="[1211,1429,740,844]" gridcol="3" gridrow="4" pageId="64" pageNumber="65">- 115 65</td>
</tr>
<tr id="35266F34AB1D96E6FF4DFCABFA19FC2C" box="[157,1429,864,968]" gridrow="5" pageId="64" pageNumber="65">
<th id="76F70648AB1D96E6FF4DFCABFE35FC2C" box="[157,441,864,968]" gridcol="0" gridrow="5" pageId="64" pageNumber="65">F plate L/W ratio Sh pl L/f pl L ratio Gnathosomal bay L</th>
<td id="76F70648AB1D96E6FE3DFCABFC42FC2C" box="[493,974,864,968]" gridcol="1" gridrow="5" pageId="64" pageNumber="65">2.2–2.4 2.3–2.4 2.2–2.4 0.97–1.07 0.97–1.0 1.04–1.1 123–143 145 141–158</td>
<td id="76F70648AB1D96E6FC26FCABFB03FC2C" box="[1014,1167,864,968]" gridcol="2" gridrow="5" pageId="64" pageNumber="65">2.0 1.19–2.0 153</td>
<td id="76F70648AB1D96E6FB6BFCABFA19FC2C" box="[1211,1429,864,968]" gridcol="3" gridrow="5" pageId="64" pageNumber="65">1.8 1.26 154</td>
</tr>
<tr id="35266F34AB1D96E6FF4DFC17FA19FBA0" box="[157,1429,988,1092]" gridrow="6" pageId="64" pageNumber="65">
<th id="76F70648AB1D96E6FF4DFC17FE35FBA0" box="[157,441,988,1092]" gridcol="0" gridrow="6" pageId="64" pageNumber="65">Cx-I L Cx-I mL Cx-II+III mL</th>
<td id="76F70648AB1D96E6FE3DFC17FC42FBA0" box="[493,974,988,1092]" gridcol="1" gridrow="6" pageId="64" pageNumber="65">209–241 247 236–253 85–97 102 95–96 25–31 23 29–34</td>
<td id="76F70648AB1D96E6FC26FC17FB03FBA0" box="[1014,1167,988,1092]" gridcol="2" gridrow="6" pageId="64" pageNumber="65">313 159 83</td>
<td id="76F70648AB1D96E6FB6BFC17FA19FBA0" box="[1211,1429,988,1092]" gridcol="3" gridrow="6" pageId="64" pageNumber="65">260 - 40</td>
</tr>
<tr id="35266F34AB1D96E6FF4DFB93FA19FB24" box="[157,1429,1112,1216]" gridrow="7" pageId="64" pageNumber="65">
<th id="76F70648AB1D96E6FF4DFB93FE35FB24" box="[157,441,1112,1216]" gridcol="0" gridrow="7" pageId="64" pageNumber="65">Cx-1 L/Cx-II+III mL Cx-I mL/Cx-II+III mL ratio Genital field L</th>
<td id="76F70648AB1D96E6FE3DFB93FC42FB24" box="[493,974,1112,1216]" gridcol="1" gridrow="7" pageId="64" pageNumber="65">8.4–7.9 10.7 7.4–8.1 3.2–3.4 4.4 2.8–3.3 120–133 137 134–144</td>
<td id="76F70648AB1D96E6FC26FB93FB03FB24" box="[1014,1167,1112,1216]" gridcol="2" gridrow="7" pageId="64" pageNumber="65">3.8 1.9 137</td>
<td id="76F70648AB1D96E6FB6BFB93FA19FB24" box="[1211,1429,1112,1216]" gridcol="3" gridrow="7" pageId="64" pageNumber="65">6.5 - 149</td>
</tr>
<tr id="35266F34AB1D96E6FF4DFB1FFA19FAD8" box="[157,1429,1236,1340]" gridrow="8" pageId="64" pageNumber="65">
<th id="76F70648AB1D96E6FF4DFB1FFE35FAD8" box="[157,441,1236,1340]" gridcol="0" gridrow="8" pageId="64" pageNumber="65">Genital field W Gf L/W ratio Dist gf—expo</th>
<td id="76F70648AB1D96E6FE3DFB1FFC42FAD8" box="[493,974,1236,1340]" gridcol="1" gridrow="8" pageId="64" pageNumber="65">101–111 111 120–127 1.19 1.23 1.12–1.14 156–181 186 175–194</td>
<td id="76F70648AB1D96E6FC26FB1FFB03FAD8" box="[1014,1167,1236,1340]" gridcol="2" gridrow="8" pageId="64" pageNumber="65">116 1.18 144</td>
<td id="76F70648AB1D96E6FB6BFB1FFA19FAD8" box="[1211,1429,1236,1340]" gridcol="3" gridrow="8" pageId="64" pageNumber="65">133 1.12 -</td>
</tr>
<tr id="35266F34AB1D96E6FF4DFA9BFA19FA82" box="[157,1429,1360,1382]" gridrow="9" pageId="64" pageNumber="65">
<th id="76F70648AB1D96E6FF4DFA9BFE35FA82" box="[157,441,1360,1382]" gridcol="0" gridrow="9" pageId="64" pageNumber="65">Dist gf—cauda</th>
<td id="76F70648AB1D96E6FE3DFA9BFC42FA82" box="[493,974,1360,1382]" gridcol="1" gridrow="9" pageId="64" pageNumber="65">219–241 233 269–270</td>
<td id="76F70648AB1D96E6FC26FA9BFB03FA82" box="[1014,1167,1360,1382]" gridcol="2" gridrow="9" pageId="64" pageNumber="65">198</td>
<td id="76F70648AB1D96E6FB6BFA9BFA19FA82" box="[1211,1429,1360,1382]" gridcol="3" gridrow="9" pageId="64" pageNumber="65">-</td>
</tr>
<tr id="35266F34AB1D96E6FF4DFAB1FA19FA5C" box="[157,1429,1402,1464]" gridrow="10" pageId="64" pageNumber="65">
<th id="76F70648AB1D96E6FF4DFAB1FE35FA5C" box="[157,441,1402,1464]" gridcol="0" gridrow="10" pageId="64" pageNumber="65">Ec L Egg (maximum diameter)</th>
<td id="76F70648AB1D96E6FE3DFAB1FC42FA5C" box="[493,974,1402,1464]" gridcol="1" gridrow="10" pageId="64" pageNumber="65">- - - - - 169–178</td>
<td id="76F70648AB1D96E6FC26FAB1FB03FA5C" box="[1014,1167,1402,1464]" gridcol="2" gridrow="10" pageId="64" pageNumber="65">- -</td>
<td id="76F70648AB1D96E6FB6BFAB1FA19FA5C" box="[1211,1429,1402,1464]" gridcol="3" gridrow="10" pageId="64" pageNumber="65">- -</td>
</tr>
<tr id="35266F34AB1D96E6FF4DFA07FA19F9D0" box="[157,1429,1484,1588]" gridrow="11" pageId="64" pageNumber="65">
<th id="76F70648AB1D96E6FF4DFA07FE35F9D0" box="[157,441,1484,1588]" gridcol="0" gridrow="11" pageId="64" pageNumber="65">Gnathosoma vL Chelicera L Palp, total L</th>
<td id="76F70648AB1D96E6FE3DFA07FC42F9D0" box="[493,974,1484,1588]" gridcol="1" gridrow="11" pageId="64" pageNumber="65">148–160 152 160–164 165–189 - 192 148–163 172 163–176</td>
<td id="76F70648AB1D96E6FC26FA07FB03F9D0" box="[1014,1167,1484,1588]" gridcol="2" gridrow="11" pageId="64" pageNumber="65">192 211 182</td>
<td id="76F70648AB1D96E6FB6BFA07FA19F9D0" box="[1211,1429,1484,1588]" gridcol="3" gridrow="11" pageId="64" pageNumber="65">- 205 -</td>
</tr>
<tr id="35266F34AB1D96E6FF4DF983FA19F954" box="[157,1429,1608,1712]" gridrow="12" pageId="64" pageNumber="65">
<th id="76F70648AB1D96E6FF4DF983FE35F954" box="[157,441,1608,1712]" gridcol="0" gridrow="12" pageId="64" pageNumber="65">P-1 dL/H P-2 dL/H P-3 dL/H</th>
<td id="76F70648AB1D96E6FE3DF983FC42F954" box="[493,974,1608,1712]" gridcol="1" gridrow="12" pageId="64" pageNumber="65">19–22/17–20 23/20 22–24/21–22 43–46/31–33 48/33 46–51/35 31–34/25–27 34/28 34–35/28</td>
<td id="76F70648AB1D96E6FC26F983FB03F954" box="[1014,1167,1608,1712]" gridcol="2" gridrow="12" pageId="64" pageNumber="65">25/22 51/37 39/31</td>
<td id="76F70648AB1D96E6FB6BF983FA19F954" box="[1211,1429,1608,1712]" gridcol="3" gridrow="12" pageId="64" pageNumber="65">- 53 33/26</td>
</tr>
<tr id="35266F34AB1D96E6FF4DF90FFA19F8C8" box="[157,1429,1732,1836]" gridrow="13" pageId="64" pageNumber="65">
<th id="76F70648AB1D96E6FF4DF90FFE35F8C8" box="[157,441,1732,1836]" gridcol="0" gridrow="13" pageId="64" pageNumber="65">P-4 dL/H P-5 dL/H P-1 dL/H ratio</th>
<td id="76F70648AB1D96E6FE3DF90FFC42F8C8" box="[493,974,1732,1836]" gridcol="1" gridrow="13" pageId="64" pageNumber="65">38–43/18–19 46/18–19 43–46/18–19 17–19/9–11 21/10 18–20/10–12 1.08–1.1 1.16 1.04–1.12</td>
<td id="76F70648AB1D96E6FC26F90FFB03F8C8" box="[1014,1167,1732,1836]" gridcol="2" gridrow="13" pageId="64" pageNumber="65">46/24 21/12 1.14</td>
<td id="76F70648AB1D96E6FB6BF90FFA19F8C8" box="[1211,1429,1732,1836]" gridcol="3" gridrow="13" pageId="64" pageNumber="65">45 13 -</td>
</tr>
<tr id="35266F34AB1D96E6FF4DF88BFA19F84C" box="[157,1429,1856,1960]" gridrow="14" pageId="64" pageNumber="65">
<th id="76F70648AB1D96E6FF4DF88BFE35F84C" box="[157,441,1856,1960]" gridcol="0" gridrow="14" pageId="64" pageNumber="65">P-2 dL/H ratio P-3 dL/H ratio P-4 dL/H ratio</th>
<td id="76F70648AB1D96E6FE3DF88BFC42F84C" box="[493,974,1856,1960]" gridcol="1" gridrow="14" pageId="64" pageNumber="65">1.4–1.41 1.46 1.34–1.46 1.21–1.25 1.22 1.25 2.1–2.27 2.5 2.37–2.5</td>
<td id="76F70648AB1D96E6FC26F88BFB03F84C" box="[1014,1167,1856,1960]" gridcol="2" gridrow="14" pageId="64" pageNumber="65">1.37 1.26 1.93</td>
<td id="76F70648AB1D96E6FB6BF88BFA19F84C" box="[1211,1429,1856,1960]" gridcol="3" gridrow="14" pageId="64" pageNumber="65">1.45* 1.27 2.3*</td>
</tr>
</table>
</paragraph>
<paragraph id="8BA96D76AB1D6958FB8EF877FB77FCC4" blockId="64.[1118,1429,1980,2002]" lastBlockId="65.[449,1367,778,877]" lastPageId="65" lastPageNumber="66" pageId="64" pageNumber="65">
<emphasis id="B962B164AB1D6959FB8EF877FA19F836" box="[1118,1429,1980,2002]" italics="true" pageId="64" pageNumber="65">......continued on the next page</emphasis>
Apkonu Apkonu 
<taxonomicName id="4C1616F5AB1C6958FCC1FCC0FB21FCC4" authority="K.O. Viets &amp; Bottger 1974" authorityName="K.O. Viets &amp; Bottger" authorityYear="1974" box="[785,1197,778,800]" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="procursa">
<emphasis id="B962B164AB1C6958FCC1FCC0FC01FCC4" box="[785,909,779,800]" italics="true" pageId="65" pageNumber="66">M. procursa</emphasis>
, 
<bibRefCitation id="EF871087AB1C6958FC4AFCC1FB21FCC4" author="Viets" box="[922,1197,778,800]" pageId="65" pageNumber="66" refString="Viets, K. O. &amp; Bottger, K. (1974) Zur Systematik und Okologie rheophiler Hydrachnellae (Acari) Zentralafrikas. Teil I. Acarologia, 16 (1), 106 - 159." type="journal article" year="1974">K.O. Viets &amp; Böttger 1974</bibRefCitation>
</taxonomicName>
, 
<collectingCountry id="F3012DE6AB1C6958FB67FCC1FB77FCC4" box="[1207,1275,778,800]" name="Republic of the Congo" pageId="65" pageNumber="66">Congo</collectingCountry>
</paragraph>
<caption id="DF693DFEAB1C6958FF4DFF6BFE07FF52" ID-Table-UUID="DF693DFEAB1C6958FF4DFF6BFE07FF52" box="[157,395,160,182]" httpUri="http://table.plazi.org/id/DF693DFEAB1C6958FF4DFF6BFE07FF52" pageId="65" pageNumber="66" targetBox="[157,1394,201,604]" targetIsTable="true" targetPageId="65">
<paragraph id="8BA96D76AB1C6958FF4DFF6BFE07FF52" blockId="65.[157,395,160,182]" box="[157,395,160,182]" pageId="65" pageNumber="66">
<emphasis id="B962B164AB1C6958FF4DFF6BFE87FF52" bold="true" box="[157,267,160,182]" pageId="65" pageNumber="66">TABLE 4.</emphasis>
(Continued)
</paragraph>
</caption>
<paragraph id="8BA96D76AB1C6958FF4DFE85FB4FFDB8" pageId="65" pageNumber="66">
<table id="F9169FD6AB1C96E6FF4DFF02FAFEFDB8" box="[157,1394,201,604]" gridcols="4" gridrows="3" pageId="65" pageNumber="66">
<tr id="35266F34AB1C96E6FF4DFF02FAFEFE80" box="[157,1394,201,356]" gridrow="0" pageId="65" pageNumber="66">
<th id="76F70648AB1C96E6FF4DFF02FECEFE80" box="[157,322,201,356]" gridcol="0" gridrow="0" pageId="65" pageNumber="66">P-5 dL/H ratio</th>
<th id="76F70648AB1C96E6FE3DFF02FC45FE80" box="[493,969,201,356]" gridcol="1" gridrow="0" pageId="65" pageNumber="66">Tagbo River ♂, Plunge pool, ♂ Plunge pool, n = 2 2♀ 1.7–1.8 2.1 1.7–1.9</th>
<th id="76F70648AB1C96E6FC26FF02FB03FE80" box="[1014,1167,201,356]" gridcol="2" gridrow="0" pageId="65" pageNumber="66">
<taxonomicName id="4C1616F5AB1C6958FC26FF01FBA7FEF8" authority="Cook, 1966" authorityName="Cook" authorityYear="1966" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="microstoma">
<emphasis id="B962B164AB1C6958FC26FF01FB03FF3B" box="[1014,1167,202,223]" italics="true" pageId="65" pageNumber="66">M. microstoma</emphasis>
sensu Cook, 1966
</taxonomicName>
, Tagbo River, ♂ 1.7
</th>
<th id="76F70648AB1C96E6FB6BFF02FAFEFE80" box="[1211,1394,201,356]" gridcol="3" gridrow="0" pageId="65" pageNumber="66">
K. Viets, 1958 Cameroon, 
<taxonomicName id="4C1616F5AB1C6958FAE1FF22FABDFEF9" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="microstoma">
<emphasis id="B962B164AB1C6958FAE1FF22FABDFEF9" italics="true" pageId="65" pageNumber="66">M. microstoma</emphasis>
</taxonomicName>
, prep. no. 1390, ♀ -
</th>
</tr>
<tr id="35266F34AB1C96E6FF4DFEB3FAFEFE04" box="[157,1394,376,480]" gridrow="1" pageId="65" pageNumber="66">
<th id="76F70648AB1C96E6FF4DFEB3FECEFE04" box="[157,322,376,480]" gridcol="0" gridrow="1" pageId="65" pageNumber="66">dL P-2/P-4 ratio I-L-4 L I-L-5 L</th>
<td id="76F70648AB1C96E6FE3DFEB3FC45FE04" box="[493,969,376,480]" gridcol="1" gridrow="1" pageId="65" pageNumber="66">1.07–1.12 1.03 1.07–1.11 72–82 84 78–88 77–88 88 83–92</td>
<td id="76F70648AB1C96E6FC26FEB3FB03FE04" box="[1014,1167,376,480]" gridcol="2" gridrow="1" pageId="65" pageNumber="66">1.1 102 108</td>
<td id="76F70648AB1C96E6FB6BFEB3FAFEFE04" box="[1211,1394,376,480]" gridcol="3" gridrow="1" pageId="65" pageNumber="66">1.17 - -</td>
</tr>
<tr id="35266F34AB1C96E6FF4DFE3FFAFEFDB8" box="[157,1394,500,604]" gridrow="2" pageId="65" pageNumber="66">
<th id="76F70648AB1C96E6FF4DFE3FFECEFDB8" box="[157,322,500,604]" gridcol="0" gridrow="2" pageId="65" pageNumber="66">I-L-6 L I-L-6 H I-L-6 L/H ratio</th>
<td id="76F70648AB1C96E6FE3DFE3FFC45FDB8" box="[493,969,500,604]" gridcol="1" gridrow="2" pageId="65" pageNumber="66">84–94 93 86–92 38–41 41–42 38–42 2.23–2.28 2.24 2.17–2.23</td>
<td id="76F70648AB1C96E6FC26FE3FFB03FDB8" box="[1014,1167,500,604]" gridcol="2" gridrow="2" pageId="65" pageNumber="66">116 53 2.2</td>
<td id="76F70648AB1C96E6FB6BFE3FFAFEFDB8" box="[1211,1394,500,604]" gridcol="3" gridrow="2" pageId="65" pageNumber="66">- - -</td>
</tr>
</table>
</paragraph>
<caption id="DF693DFEAB1C6958FF47FD53FB98FD2B" ID-Table-UUID="DF693DFEAB1C6958FF47FD53FB98FD2B" httpUri="http://table.plazi.org/id/DF693DFEAB1C6958FF47FD53FB98FD2B" pageId="65" pageNumber="66" targetBox="[151,1435,896,1662]" targetIsTable="true" targetPageId="65">
<paragraph id="8BA96D76AB1C6958FF47FD53FB98FD2B" blockId="65.[151,1436,664,719]" pageId="65" pageNumber="66">
<emphasis id="B962B164AB1C6958FF47FD53FE81FD4B" bold="true" box="[151,269,664,687]" pageId="65" pageNumber="66">TABLE 5.</emphasis>
Measurements of 
<taxonomicName id="4C1616F5AB1C6958FE0AFD53FC3AFD4B" authority="Koenike, 1898" authorityName="Koenike" authorityYear="1898" box="[474,950,664,687]" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="microstoma">
<emphasis id="B962B164AB1C6958FE0AFD53FDF9FD4B" box="[474,629,664,687]" italics="true" pageId="65" pageNumber="66">Monatractides</emphasis>
<emphasis id="B962B164AB1C6958FDAFFD53FD70FD4B" box="[639,764,664,687]" italics="true" pageId="65" pageNumber="66">microstoma</emphasis>
(Koenike, 1898)
</taxonomicName>
—species complex (Form “B“) from Ghana, and 
<taxonomicName id="4C1616F5AB1C6958FF14FD73FD9FFD2B" authority="K.O. Viets, 1974" authorityName="K.O. Viets" authorityYear="1974" box="[196,531,696,719]" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="procursa">
<emphasis id="B962B164AB1C6958FF14FD73FF6EFD2B" box="[196,226,696,719]" italics="true" pageId="65" pageNumber="66">M.</emphasis>
<emphasis id="B962B164AB1C6958FF37FD73FECAFD2B" box="[231,326,696,719]" italics="true" pageId="65" pageNumber="66">procursa</emphasis>
(K.O. Viets, 1974)
</taxonomicName>
from Congo (from K.O. Viets &amp; Böttger 1974).
</paragraph>
</caption>
<paragraph id="8BA96D76AB1C6958FE11FCE1FADBFCAA" blockId="65.[449,1367,778,877]" box="[449,1367,810,846]" pageId="65" pageNumber="66">
stream, ♂, stream, ♀, 
<typeStatus id="54ADD3D4AB1C6958FCC1FCF3FCE6FCAA" box="[785,874,824,846]" pageId="65" pageNumber="66" type="holotype">holotype</typeStatus>
♀ 
<typeStatus id="54ADD3D4AB1C6958FC68FCF2FB83FCAA" box="[952,1039,825,846]" pageId="65" pageNumber="66" type="paratype">paratype</typeStatus>
♂, ♂, n = 8 ♀, n = 9
</paragraph>
<paragraph id="8BA96D76AB1C6958FE11FC82FBCBFC89" blockId="65.[449,1367,778,877]" box="[449,1095,840,877]" pageId="65" pageNumber="66">n = 5 n = 2 slide no. 4768 slide no. 4797</paragraph>
<paragraph id="8BA96D76AB1C6958FF47FC4BFA80F99A" pageId="65" pageNumber="66">
<table id="F9169FD6AB1C96E6FF47FC4BFA17F99A" box="[151,1435,896,1662]" gridcols="7" gridrows="7" pageId="65" pageNumber="66">
<tr id="35266F34AB1C96E6FF47FC4BFA17FC72" box="[151,1435,896,918]" gridrow="0" pageId="65" pageNumber="66">
<th id="76F70648AB1C96E6FF47FC4BFEE2FC72" box="[151,366,896,918]" gridcol="0" gridrow="0" pageId="65" pageNumber="66">Idiosoma L</th>
<th id="76F70648AB1C96E6FE11FC4BFDABFC72" box="[449,551,896,918]" gridcol="1" gridrow="0" pageId="65" pageNumber="66">650–736</th>
<th id="76F70648AB1C96E6FDB9FC4BFD43FC72" box="[617,719,896,918]" gridcol="2" gridrow="0" pageId="65" pageNumber="66">677–818</th>
<th id="76F70648AB1C96E6FCC1FC4BFCFAFC72" box="[785,886,896,918]" gridcol="3" gridrow="0" pageId="65" pageNumber="66">701</th>
<th id="76F70648AB1C96E6FC69FC4BFB92FC72" box="[953,1054,896,918]" gridcol="4" gridrow="0" pageId="65" pageNumber="66">681</th>
<th id="76F70648AB1C96E6FB8CFC4BFB3AFC72" box="[1116,1206,896,918]" gridcol="5" gridrow="0" pageId="65" pageNumber="66">665–697</th>
<th id="76F70648AB1C96E6FAD4FC4BFA17FC72" box="[1284,1435,896,918]" gridcol="6" gridrow="0" pageId="65" pageNumber="66">677–750</th>
</tr>
<tr id="35266F34AB1C96E6FF47FC61FA17FBF6" box="[151,1435,938,1042]" gridrow="1" pageId="65" pageNumber="66">
<th id="76F70648AB1C96E6FF47FC61FEE2FBF6" box="[151,366,938,1042]" gridcol="0" gridrow="1" pageId="65" pageNumber="66">Idiosoma W Idiosoma L/W ratio Ds L</th>
<td id="76F70648AB1C96E6FE11FC61FDABFBF6" box="[449,551,938,1042]" gridcol="1" gridrow="1" pageId="65" pageNumber="66">469–489 1.36–1.5 522–591</td>
<td id="76F70648AB1C96E6FDB9FC61FD43FBF6" box="[617,719,938,1042]" gridcol="2" gridrow="1" pageId="65" pageNumber="66">481–534 1.4–1.5 544–666</td>
<td id="76F70648AB1C96E6FCC1FC61FCFAFBF6" box="[785,886,938,1042]" gridcol="3" gridrow="1" pageId="65" pageNumber="66">515 1.36 585</td>
<td id="76F70648AB1C96E6FC69FC61FB92FBF6" box="[953,1054,938,1042]" gridcol="4" gridrow="1" pageId="65" pageNumber="66">487 1.4 564</td>
<td id="76F70648AB1C96E6FB8CFC61FB3AFBF6" box="[1116,1206,938,1042]" gridcol="5" gridrow="1" pageId="65" pageNumber="66">478–508 - 545–573</td>
<td id="76F70648AB1C96E6FAD4FC61FA17FBF6" box="[1284,1435,938,1042]" gridcol="6" gridrow="1" pageId="65" pageNumber="66">474–528 - 548–606</td>
</tr>
<tr id="35266F34AB1C96E6FF47FBEDFA17FB6A" box="[151,1435,1062,1166]" gridrow="2" pageId="65" pageNumber="66">
<th id="76F70648AB1C96E6FF47FBEDFEE2FB6A" box="[151,366,1062,1166]" gridcol="0" gridrow="2" pageId="65" pageNumber="66">Ds W Ds L/W ratio Dp L</th>
<td id="76F70648AB1C96E6FE11FBEDFDABFB6A" box="[449,551,1062,1166]" gridcol="1" gridrow="2" pageId="65" pageNumber="66">409–428 1.24–1.39 474–548</td>
<td id="76F70648AB1C96E6FDB9FBEDFD43FB6A" box="[617,719,1062,1166]" gridcol="2" gridrow="2" pageId="65" pageNumber="66">425–453 1.28–1.47 498–619</td>
<td id="76F70648AB1C96E6FCC1FBEDFCFAFB6A" box="[785,886,1062,1166]" gridcol="3" gridrow="2" pageId="65" pageNumber="66">454 1.29 540</td>
<td id="76F70648AB1C96E6FC69FBEDFB92FB6A" box="[953,1054,1062,1166]" gridcol="4" gridrow="2" pageId="65" pageNumber="66">429 1.32 523</td>
<td id="76F70648AB1C96E6FB8CFBEDFB3AFB6A" box="[1116,1206,1062,1166]" gridcol="5" gridrow="2" pageId="65" pageNumber="66">424–437 - 503–535</td>
<td id="76F70648AB1C96E6FAD4FBEDFA17FB6A" box="[1284,1435,1062,1166]" gridcol="6" gridrow="2" pageId="65" pageNumber="66">404–462 - 498–564</td>
</tr>
<tr id="35266F34AB1C96E6FF47FB69FA17FAEE" box="[151,1435,1186,1290]" gridrow="3" pageId="65" pageNumber="66">
<th id="76F70648AB1C96E6FF47FB69FEE2FAEE" box="[151,366,1186,1290]" gridcol="0" gridrow="3" pageId="65" pageNumber="66">Sh plate L Sh plate W Sh plate L/W</th>
<td id="76F70648AB1C96E6FE11FB69FDABFAEE" box="[449,551,1186,1290]" gridcol="1" gridrow="3" pageId="65" pageNumber="66">131–139 55–66 2.05–2.46</td>
<td id="76F70648AB1C96E6FDB9FB69FD43FAEE" box="[617,719,1186,1290]" gridcol="2" gridrow="3" pageId="65" pageNumber="66">138–144 61–67 2.1–2.4</td>
<td id="76F70648AB1C96E6FCC1FB69FCFAFAEE" box="[785,886,1186,1290]" gridcol="3" gridrow="3" pageId="65" pageNumber="66">133–136 - -</td>
<td id="76F70648AB1C96E6FC69FB69FB92FAEE" box="[953,1054,1186,1290]" gridcol="4" gridrow="3" pageId="65" pageNumber="66">135–137 - -</td>
<td id="76F70648AB1C96E6FB8CFB69FB3AFAEE" box="[1116,1206,1186,1290]" gridcol="5" gridrow="3" pageId="65" pageNumber="66">128–137 - -</td>
<td id="76F70648AB1C96E6FAD4FB69FA17FAEE" box="[1284,1435,1186,1290]" gridcol="6" gridrow="3" pageId="65" pageNumber="66">130–139 - -</td>
</tr>
<tr id="35266F34AB1C96E6FF47FAD5FA17FA62" box="[151,1435,1310,1414]" gridrow="4" pageId="65" pageNumber="66">
<th id="76F70648AB1C96E6FF47FAD5FEE2FA62" box="[151,366,1310,1414]" gridcol="0" gridrow="4" pageId="65" pageNumber="66">F plate L F plate W F plate L/W ratio</th>
<td id="76F70648AB1C96E6FE11FAD5FDABFA62" box="[449,551,1310,1414]" gridcol="1" gridrow="4" pageId="65" pageNumber="66">119–141 55–67 1.86–2.57</td>
<td id="76F70648AB1C96E6FDB9FAD5FD43FA62" box="[617,719,1310,1414]" gridcol="2" gridrow="4" pageId="65" pageNumber="66">119–150 59–63 1.9–2.5</td>
<td id="76F70648AB1C96E6FCC1FAD5FCFAFA62" box="[785,886,1310,1414]" gridcol="3" gridrow="4" pageId="65" pageNumber="66">132–133 - -</td>
<td id="76F70648AB1C96E6FC69FAD5FB92FA62" box="[953,1054,1310,1414]" gridcol="4" gridrow="4" pageId="65" pageNumber="66">122–126 - -</td>
<td id="76F70648AB1C96E6FB8CFAD5FB3AFA62" box="[1116,1206,1310,1414]" gridcol="5" gridrow="4" pageId="65" pageNumber="66">114–129 - -</td>
<td id="76F70648AB1C96E6FAD4FAD5FA17FA62" box="[1284,1435,1310,1414]" gridcol="6" gridrow="4" pageId="65" pageNumber="66">114–133 - -</td>
</tr>
<tr id="35266F34AB1C96E6FF47FA51FA17F9E6" box="[151,1435,1434,1538]" gridrow="5" pageId="65" pageNumber="66">
<th id="76F70648AB1C96E6FF47FA51FEE2F9E6" box="[151,366,1434,1538]" gridcol="0" gridrow="5" pageId="65" pageNumber="66">Sh pl L/f pl L ratio Gnathosomal bay L Cx-I L</th>
<td id="76F70648AB1C96E6FE11FA51FDABF9E6" box="[449,551,1434,1538]" gridcol="1" gridrow="5" pageId="65" pageNumber="66">0.96–1.13 147–178 256–287</td>
<td id="76F70648AB1C96E6FDB9FA51FD43F9E6" box="[617,719,1434,1538]" gridcol="2" gridrow="5" pageId="65" pageNumber="66">0.96–1.18 163–184 266–313</td>
<td id="76F70648AB1C96E6FCC1FA51FCFAF9E6" box="[785,886,1434,1538]" gridcol="3" gridrow="5" pageId="65" pageNumber="66">0.98–1.01 150 ca.250</td>
<td id="76F70648AB1C96E6FC69FA51FB92F9E6" box="[953,1054,1434,1538]" gridcol="4" gridrow="5" pageId="65" pageNumber="66">1.07–1.12 143 ca. 249</td>
<td id="76F70648AB1C96E6FB8CFA51FB3AF9E6" box="[1116,1206,1434,1538]" gridcol="5" gridrow="5" pageId="65" pageNumber="66">- 130–161 -</td>
<td id="76F70648AB1C96E6FAD4FA51FA17F9E6" box="[1284,1435,1434,1538]" gridcol="6" gridrow="5" pageId="65" pageNumber="66">- 134–157 -</td>
</tr>
<tr id="35266F34AB1C96E6FF47F9DDFA17F99A" box="[151,1435,1558,1662]" gridrow="6" pageId="65" pageNumber="66">
<th id="76F70648AB1C96E6FF47F9DDFEE2F99A" box="[151,366,1558,1662]" gridcol="0" gridrow="6" pageId="65" pageNumber="66">Cx-I mL Cx-II+III mL Cx-1 L/Cx-II+III mL</th>
<td id="76F70648AB1C96E6FE11F9DDFDABF99A" box="[449,551,1558,1662]" gridcol="1" gridrow="6" pageId="65" pageNumber="66">103–115 24–50 5.7–11.2</td>
<td id="76F70648AB1C96E6FDB9F9DDFD43F99A" box="[617,719,1558,1662]" gridcol="2" gridrow="6" pageId="65" pageNumber="66">103–128 18–50 6.3–14.7</td>
<td id="76F70648AB1C96E6FCC1F9DDFCFAF99A" box="[785,886,1558,1662]" gridcol="3" gridrow="6" pageId="65" pageNumber="66">100 20 12.5</td>
<td id="76F70648AB1C96E6FC69F9DDFB92F99A" box="[953,1054,1558,1662]" gridcol="4" gridrow="6" pageId="65" pageNumber="66">106 27 9.2</td>
<td id="76F70648AB1C96E6FB8CF9DDFB3AF99A" box="[1116,1206,1558,1662]" gridcol="5" gridrow="6" pageId="65" pageNumber="66">94–106 20–32 -</td>
<td id="76F70648AB1C96E6FAD4F9DDFA17F99A" box="[1284,1435,1558,1662]" gridcol="6" gridrow="6" pageId="65" pageNumber="66">93–106 17–29 -</td>
</tr>
</table>
</paragraph>
<paragraph id="8BA96D76AB1C6958FF47F959FA80F844" pageId="65" pageNumber="66">
<table id="F9169FD6AB1C96E6FF47F959FA17F844" box="[151,1435,1682,1952]" gridcols="6" gridrows="3" pageId="65" pageNumber="66">
<tr id="35266F34AB1C96E6FF47F959FA17F91E" box="[151,1435,1682,1786]" gridrow="0" pageId="65" pageNumber="66">
<th id="76F70648AB1C96E6FF47F959FDABF91E" box="[151,551,1682,1786]" gridcol="0" gridrow="0" pageId="65" pageNumber="66">Cx-I mL/Cx-II+III mL ratio 2.2–4.8 Genital field L 138–150 Genital field W 105–122</th>
<th id="76F70648AB1C96E6FDB9F959FD43F91E" box="[617,719,1682,1786]" gridcol="1" gridrow="0" pageId="65" pageNumber="66">2.6–5.7 152–161 128–139</th>
<th id="76F70648AB1C96E6FCC1F959FCB1F91E" box="[785,829,1682,1786]" gridcol="2" gridrow="0" pageId="65" pageNumber="66">5.0 156 141</th>
<th id="76F70648AB1C96E6FC69F959FC69F91E" box="[953,997,1682,1786]" gridcol="3" gridrow="0" pageId="65" pageNumber="66">3.9 147 114</th>
<th id="76F70648AB1C96E6FB8CF959FB39F91E" box="[1116,1205,1682,1786]" gridcol="4" gridrow="0" pageId="65" pageNumber="66">- 142–149 110–117</th>
<th id="76F70648AB1C96E6FAD4F959FA17F91E" box="[1284,1435,1682,1786]" gridcol="5" gridrow="0" pageId="65" pageNumber="66">- 155–160 135–146</th>
</tr>
<tr id="35266F34AB1C96E6FF47F8C5FA17F892" box="[151,1435,1806,1910]" gridrow="1" pageId="65" pageNumber="66">
<th id="76F70648AB1C96E6FF47F8C5FDABF892" box="[151,551,1806,1910]" gridcol="0" gridrow="1" pageId="65" pageNumber="66">Gf L/W ratio 1.23–1.33 Dist gf—expo 153–169 Dist gf—cauda 211–248</th>
<td id="76F70648AB1C96E6FDB9F8C5FD43F892" box="[617,719,1806,1910]" gridcol="1" gridrow="1" pageId="65" pageNumber="66">1.16–1.18 156–200 239–294</td>
<td id="76F70648AB1C96E6FCC1F8C5FCB1F892" box="[785,829,1806,1910]" gridcol="2" gridrow="1" pageId="65" pageNumber="66">1.11 - -</td>
<td id="76F70648AB1C96E6FC69F8C5FC69F892" box="[953,997,1806,1910]" gridcol="3" gridrow="1" pageId="65" pageNumber="66">1.29 - -</td>
<td id="76F70648AB1C96E6FB8CF8C5FB39F892" box="[1116,1205,1806,1910]" gridcol="4" gridrow="1" pageId="65" pageNumber="66">- - -</td>
<td id="76F70648AB1C96E6FAD4F8C5FA17F892" box="[1284,1435,1806,1910]" gridcol="5" gridrow="1" pageId="65" pageNumber="66">- - -</td>
</tr>
<tr id="35266F34AB1C96E6FF47F841FA17F844" box="[151,1435,1930,1952]" gridrow="2" pageId="65" pageNumber="66">
<th id="76F70648AB1C96E6FF47F841FDABF844" box="[151,551,1930,1952]" gridcol="0" gridrow="2" pageId="65" pageNumber="66">Ec L 119–177</th>
<td id="76F70648AB1C96E6FDB9F841FD43F844" box="[617,719,1930,1952]" gridcol="1" gridrow="2" pageId="65" pageNumber="66">-</td>
<td id="76F70648AB1C96E6FCC1F841FCB1F844" box="[785,829,1930,1952]" gridcol="2" gridrow="2" pageId="65" pageNumber="66">-</td>
<td id="76F70648AB1C96E6FC69F841FC69F844" box="[953,997,1930,1952]" gridcol="3" gridrow="2" pageId="65" pageNumber="66">-</td>
<td id="76F70648AB1C96E6FB8CF841FB39F844" box="[1116,1205,1930,1952]" gridcol="4" gridrow="2" pageId="65" pageNumber="66">-</td>
<td id="76F70648AB1C96E6FAD4F841FA17F844" box="[1284,1435,1930,1952]" gridcol="5" gridrow="2" pageId="65" pageNumber="66">-</td>
</tr>
</table>
</paragraph>
<paragraph id="8BA96D76AB1C695BFBB5F878FC85F9AA" blockId="65.[1125,1435,1971,1993]" lastBlockId="66.[151,1436,1202,2037]" lastPageId="66" lastPageNumber="67" pageId="65" pageNumber="66">
<emphasis id="B962B164AB1C6958FBB5F878FA17F82D" box="[1125,1435,1971,1993]" italics="true" pageId="65" pageNumber="66">......continued on the next page</emphasis>
<emphasis id="B962B164AB1F695BFF17FB79FEB8FB2F" bold="true" box="[199,308,1202,1227]" pageId="66" pageNumber="67">Remarks</emphasis>
. There appear to be two forms of this species (or two sibling species within 
<taxonomicName id="4C1616F5AB1F695BFB29FB79FE96FB0A" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="microstoma">
<emphasis id="B962B164AB1F695BFB29FB79FE96FB0A" italics="true" pageId="66" pageNumber="67">Monatractides microstoma</emphasis>
</taxonomicName>
species - complex, see above for a discussion). At one extreme end (Form ''A''), specimens are generally of smaller dimensions, the dorsal plate with the area of primary sclerotization bears four dorsoglandularia (
<figureCitation id="132D71F3AB1F695BFF4FFAD7FE98FAD1" box="[159,276,1308,1333]" captionStart="FIGURE 39 A – G" captionStartId="58.[151,250,1945,1967]" captionTargetBox="[210,1376,197,1927]" captionTargetId="figure@58.[202,1385,195,1935]" captionTargetPageId="58" captionText="FIGURE 39 A – G. Monatractides microstoma — species complex (Form '' A' '), male, Tagbo River: A = frontal margin of idiosoma, dorsal view; B = dorsal shield; C = idiosoma, ventral view; D = palp, medial view; E = palp, lateral view; F = gnathosoma; G = I-L- 5 and - 6. Scale bars = 100 Μm." httpUri="https://zenodo.org/record/286342/files/figure.png" pageId="66" pageNumber="67">Figs. 39B</figureCitation>
, 
<figureCitation id="132D71F3AB1F695BFEF5FAD6FEDBFAD1" box="[293,343,1309,1333]" captionStart="FIGURE 41 A – D" captionStartId="60.[151,250,1957,1979]" captionTargetBox="[215,1372,202,1919]" captionTargetId="figure@60.[198,1388,195,1933]" captionTargetPageId="60" captionText="FIGURE 41 A – D. Monatractides microstoma - species complex, female (A – B—Form '' A' ', Plunge pool Agumatsa Falls; C – D—Form '' B' ', Apkonu stream): A, C = dorsal shield; B, D = idiosoma, ventral view. Scale bar = 100 Μm." httpUri="https://zenodo.org/record/286344/files/figure.png" pageId="66" pageNumber="67">41A</figureCitation>
, 
<figureCitation id="132D71F3AB1F695BFEB8FAD6FE10FAD1" box="[360,412,1309,1333]" captionStart="FIGURE 43 A – I" captionStartId="62.[151,250,1931,1953]" captionTargetBox="[165,1436,195,1911]" captionTargetId="figure@62.[165,1450,195,1912]" captionTargetPageId="62" captionText="FIGURE 43 A – I. Photographs of dorsal (A – C) and ventral shields (D – I) of Monatractides microstoma - species complex (A, C D – G—Form '' A' ', B, H – I—Form '' B' '): A, D = male from Tagbo River, E = male from Wanjakli stream, C, F = aberrant male from Kintampo Falls, G = female from Plunge pool, B – H = male from Apkonu stream, I = female from Apkonu stream." httpUri="https://zenodo.org/record/286346/files/figure.png" pageId="66" pageNumber="67">43A</figureCitation>
), the gnathosoma is more elongated (
<figureCitation id="132D71F3AB1F695BFC8BFAD7FC4DFAD1" box="[859,961,1308,1333]" captionStart="FIGURE 39 A – G" captionStartId="58.[151,250,1945,1967]" captionTargetBox="[210,1376,197,1927]" captionTargetId="figure@58.[202,1385,195,1935]" captionTargetPageId="58" captionText="FIGURE 39 A – G. Monatractides microstoma — species complex (Form '' A' '), male, Tagbo River: A = frontal margin of idiosoma, dorsal view; B = dorsal shield; C = idiosoma, ventral view; D = palp, medial view; E = palp, lateral view; F = gnathosoma; G = I-L- 5 and - 6. Scale bars = 100 Μm." httpUri="https://zenodo.org/record/286342/files/figure.png" pageId="66" pageNumber="67">Fig. 39F</figureCitation>
) and the male genital field is anteriorly projecting and strongly pointed (
<figureCitation id="132D71F3AB1F695BFDD6FAF4FDFBFABC" box="[518,631,1343,1368]" captionStart="FIGURE 39 A – G" captionStartId="58.[151,250,1945,1967]" captionTargetBox="[210,1376,197,1927]" captionTargetId="figure@58.[202,1385,195,1935]" captionTargetPageId="58" captionText="FIGURE 39 A – G. Monatractides microstoma — species complex (Form '' A' '), male, Tagbo River: A = frontal margin of idiosoma, dorsal view; B = dorsal shield; C = idiosoma, ventral view; D = palp, medial view; E = palp, lateral view; F = gnathosoma; G = I-L- 5 and - 6. Scale bars = 100 Μm." httpUri="https://zenodo.org/record/286342/files/figure.png" pageId="66" pageNumber="67">Figs. 39C</figureCitation>
, 
<figureCitation id="132D71F3AB1F695BFD53FA8BFD34FABC" box="[643,696,1344,1368]" captionStart="FIGURE 42 A – K. A – H" captionStartId="61.[151,250,1673,1695]" captionTargetBox="[160,1430,212,1641]" captionTargetId="figure@61.[151,1436,195,1648]" captionTargetPageId="61" captionText="FIGURE 42 A – K. A – H Monatractides microstoma - species complex, male (A—Form '' A' ', B – H—Form '' B' '; A, G – H = specimens from Tagbo River, B – E = specimens from Apkonu stream, F = specimen from Kintampo Falls): A – D = genital field; E – H = ejaculatory complex. I – K Monatractides jucundus (Lundblad, 1927) (redrawn from Lundblad 1952): I = ejaculatory complex, K = genital flap with apical part of ejaculatory complex opened, lateral view. Scale bars = 100 Μm." httpUri="https://zenodo.org/record/286345/files/figure.png" pageId="66" pageNumber="67">42A</figureCitation>
). At the other extreme end (form ''B''), specimens are generally of larger dimensions, the dorsal plate with the area of primary sclerotization bears two dorsoglandularia (
<figureCitation id="132D71F3AB1F695BFAF4FAA9FA14FA9F" box="[1316,1432,1378,1403]" captionStart="FIGURE 40 A – G" captionStartId="59.[151,250,1946,1968]" captionTargetBox="[156,1421,200,1921]" captionTargetId="figure@59.[151,1436,195,1922]" captionTargetPageId="59" captionText="FIGURE 40 A – G. Monatractides microstoma - species complex (Form '' B' '), male, Apkonu stream: A = frontal margin of idiosoma, dorsal view; B = dorsal shield; C = idiosoma, ventral view; D = palp, medial view; E = palp, lateral view; F = gnathosoma; G = I-L- 5 and - 6. Scale bars = 100 Μm." httpUri="https://zenodo.org/record/286343/files/figure.png" pageId="66" pageNumber="67">Figs. 40A</figureCitation>
, 
<figureCitation id="132D71F3AB1F695BFF47FA4DFF44FA7A" box="[151,200,1414,1438]" captionStart="FIGURE 41 A – D" captionStartId="60.[151,250,1957,1979]" captionTargetBox="[215,1372,202,1919]" captionTargetId="figure@60.[198,1388,195,1933]" captionTargetPageId="60" captionText="FIGURE 41 A – D. Monatractides microstoma - species complex, female (A – B—Form '' A' ', Plunge pool Agumatsa Falls; C – D—Form '' B' ', Apkonu stream): A, C = dorsal shield; B, D = idiosoma, ventral view. Scale bar = 100 Μm." httpUri="https://zenodo.org/record/286344/files/figure.png" pageId="66" pageNumber="67">41C</figureCitation>
, 
<figureCitation id="132D71F3AB1F695BFF04FA4DFE89FA7A" box="[212,261,1414,1438]" captionStart="FIGURE 43 A – I" captionStartId="62.[151,250,1931,1953]" captionTargetBox="[165,1436,195,1911]" captionTargetId="figure@62.[165,1450,195,1912]" captionTargetPageId="62" captionText="FIGURE 43 A – I. Photographs of dorsal (A – C) and ventral shields (D – I) of Monatractides microstoma - species complex (A, C D – G—Form '' A' ', B, H – I—Form '' B' '): A, D = male from Tagbo River, E = male from Wanjakli stream, C, F = aberrant male from Kintampo Falls, G = female from Plunge pool, B – H = male from Apkonu stream, I = female from Apkonu stream." httpUri="https://zenodo.org/record/286346/files/figure.png" pageId="66" pageNumber="67">43B</figureCitation>
), the gnathosoma is more triangular in shape (
<figureCitation id="132D71F3AB1F695BFCDFFA4EFCF8FA7A" box="[783,884,1413,1438]" captionStart="FIGURE 40 A – G" captionStartId="59.[151,250,1946,1968]" captionTargetBox="[156,1421,200,1921]" captionTargetId="figure@59.[151,1436,195,1922]" captionTargetPageId="59" captionText="FIGURE 40 A – G. Monatractides microstoma - species complex (Form '' B' '), male, Apkonu stream: A = frontal margin of idiosoma, dorsal view; B = dorsal shield; C = idiosoma, ventral view; D = palp, medial view; E = palp, lateral view; F = gnathosoma; G = I-L- 5 and - 6. Scale bars = 100 Μm." httpUri="https://zenodo.org/record/286343/files/figure.png" pageId="66" pageNumber="67">Fig. 40F</figureCitation>
), P-4 has a small but clearly visible denticle near insertion of ventral setae (
<figureCitation id="132D71F3AB1F695BFE6DFA62FDC1FA25" box="[445,589,1449,1474]" captionStart="FIGURE 40 A – G" captionStartId="59.[151,250,1946,1968]" captionTargetBox="[156,1421,200,1921]" captionTargetId="figure@59.[151,1436,195,1922]" captionTargetPageId="59" captionText="FIGURE 40 A – G. Monatractides microstoma - species complex (Form '' B' '), male, Apkonu stream: A = frontal margin of idiosoma, dorsal view; B = dorsal shield; C = idiosoma, ventral view; D = palp, medial view; E = palp, lateral view; F = gnathosoma; G = I-L- 5 and - 6. Scale bars = 100 Μm." httpUri="https://zenodo.org/record/286343/files/figure.png" pageId="66" pageNumber="67">Figs. 40D–E</figureCitation>
) and the male genital field is anteriorly less projecting and slightly pointed (
<figureCitation id="132D71F3AB1F695BFF4FFA07FE9CFA01" box="[159,272,1484,1509]" captionStart="FIGURE 40 A – G" captionStartId="59.[151,250,1946,1968]" captionTargetBox="[156,1421,200,1921]" captionTargetId="figure@59.[151,1436,195,1922]" captionTargetPageId="59" captionText="FIGURE 40 A – G. Monatractides microstoma - species complex (Form '' B' '), male, Apkonu stream: A = frontal margin of idiosoma, dorsal view; B = dorsal shield; C = idiosoma, ventral view; D = palp, medial view; E = palp, lateral view; F = gnathosoma; G = I-L- 5 and - 6. Scale bars = 100 Μm." httpUri="https://zenodo.org/record/286343/files/figure.png" pageId="66" pageNumber="67">Figs. 40C</figureCitation>
, 
<figureCitation id="132D71F3AB1F695BFECBFA06FEFDFA01" box="[283,369,1485,1509]" captionStart="FIGURE 42 A – K. A – H" captionStartId="61.[151,250,1673,1695]" captionTargetBox="[160,1430,212,1641]" captionTargetId="figure@61.[151,1436,195,1648]" captionTargetPageId="61" captionText="FIGURE 42 A – K. A – H Monatractides microstoma - species complex, male (A—Form '' A' ', B – H—Form '' B' '; A, G – H = specimens from Tagbo River, B – E = specimens from Apkonu stream, F = specimen from Kintampo Falls): A – D = genital field; E – H = ejaculatory complex. I – K Monatractides jucundus (Lundblad, 1927) (redrawn from Lundblad 1952): I = ejaculatory complex, K = genital flap with apical part of ejaculatory complex opened, lateral view. Scale bars = 100 Μm." httpUri="https://zenodo.org/record/286345/files/figure.png" pageId="66" pageNumber="67">42C–D</figureCitation>
). The figures 39 A–G and 40 A–G illustrate the two extremes. However it is necessary to stress that in many cases we observed intermediate characters (in shape of male genital field, see 
<figureCitation id="132D71F3AB1F695BFB4DFA24FABCF9EC" box="[1181,1328,1519,1544]" captionStart="FIGURE 42 A – K. A – H" captionStartId="61.[151,250,1673,1695]" captionTargetBox="[160,1430,212,1641]" captionTargetId="figure@61.[151,1436,195,1648]" captionTargetPageId="61" captionText="FIGURE 42 A – K. A – H Monatractides microstoma - species complex, male (A—Form '' A' ', B – H—Form '' B' '; A, G – H = specimens from Tagbo River, B – E = specimens from Apkonu stream, F = specimen from Kintampo Falls): A – D = genital field; E – H = ejaculatory complex. I – K Monatractides jucundus (Lundblad, 1927) (redrawn from Lundblad 1952): I = ejaculatory complex, K = genital flap with apical part of ejaculatory complex opened, lateral view. Scale bars = 100 Μm." httpUri="https://zenodo.org/record/286345/files/figure.png" pageId="66" pageNumber="67">Figs. 42A–D</figureCitation>
) between these two extremes, even though at most sampling sites where these two forms are living together it is easily possible to identify the two groups (see ''New records'').
</paragraph>
<caption id="DF693DFEAB1F695BFF47FF6BFE09FF52" ID-Table-UUID="DF693DFEAB1F695BFF47FF6BFE09FF52" box="[151,389,160,182]" httpUri="http://table.plazi.org/id/DF693DFEAB1F695BFF47FF6BFE09FF52" pageId="66" pageNumber="67" targetBox="[151,1374,218,482]" targetIsTable="true" targetPageId="66">
<paragraph id="8BA96D76AB1F695BFF47FF6BFE09FF52" blockId="66.[151,389,160,182]" box="[151,389,160,182]" pageId="66" pageNumber="67">
<emphasis id="B962B164AB1F695BFF47FF6BFE88FF52" bold="true" box="[151,260,160,182]" pageId="66" pageNumber="67">TABLE 5.</emphasis>
(Continued)
</paragraph>
</caption>
<paragraph id="8BA96D76AB1F695BFE11FF11FA81FE06" pageId="66" pageNumber="67">
<table id="F9169FD6AB1F96E6FF47FF11FAD2FE06" box="[151,1374,218,482]" gridcols="3" gridrows="3" pageId="66" pageNumber="67">
<tr id="35266F34AB1F96E6FF47FF11FAD2FED8" box="[151,1374,218,316]" gridrow="0" pageId="66" pageNumber="67">
<th id="76F70648AB1F96E6FF47FF11FDA7FED8" box="[151,555,218,316]" gridcol="0" gridrow="0" pageId="66" pageNumber="67">Apkonu stream, ♂, n = 5</th>
<th id="76F70648AB1F96E6FDB9FF11FD5EFED8" box="[617,722,218,316]" gridcol="1" gridrow="0" pageId="66" pageNumber="67">Apkonu stream, ♀, n = 2</th>
<th id="76F70648AB1F96E6FCC1FF11FAD2FED8" box="[785,1374,218,316]" gridcol="2" gridrow="0" pageId="66" pageNumber="67">
<taxonomicName id="4C1616F5AB1F695BFCC1FF10FB21FF14" authority="K.O. Viets &amp; Bottger 1974" authorityName="K.O. Viets &amp; Bottger" authorityYear="1974" box="[785,1197,218,240]" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="procursa">
<emphasis id="B962B164AB1F695BFCC1FF10FC01FF14" box="[785,909,219,240]" italics="true" pageId="66" pageNumber="67">M. procursa</emphasis>
, K.O. Viets &amp; Böttger 1974
</taxonomicName>
, Congo holotype ♀ paratype ♂, ♂, n = 8 ♀, n = 9 slide no. 4768 slide no. 4797
</th>
</tr>
<tr id="35266F34AB1F96E6FF47FE9BFAD2FE6B" box="[151,1374,336,399]" gridrow="1" pageId="66" pageNumber="67">
<th id="76F70648AB1F96E6FF47FE9BFDA7FE6B" box="[151,555,336,399]" gridcol="0" gridrow="1" pageId="66" pageNumber="67">Egg (maximum diameter) - Gnathosoma vL 138–180</th>
<td id="76F70648AB1F96E6FDB9FE9BFD5EFE6B" box="[617,722,336,399]" gridcol="1" gridrow="1" pageId="66" pageNumber="67">- 163–178</td>
<td id="76F70648AB1F96E6FCC1FE9BFAD2FE6B" box="[785,1374,336,399]" gridcol="2" gridrow="1" pageId="66" pageNumber="67">172 - - - 162 162 158–170 162–178</td>
</tr>
<tr id="35266F34AB1F96E6FF47FE68FAD2FE06" box="[151,1374,419,482]" gridrow="2" pageId="66" pageNumber="67">
<th id="76F70648AB1F96E6FF47FE68FDA7FE06" box="[151,555,419,482]" gridcol="0" gridrow="2" pageId="66" pageNumber="67">Chelicera L 155–215 Palp, total L 169–175</th>
<td id="76F70648AB1F96E6FDB9FE68FD5EFE06" box="[617,722,419,482]" gridcol="1" gridrow="2" pageId="66" pageNumber="67">180–222 192–194</td>
<td id="76F70648AB1F96E6FCC1FE68FAD2FE06" box="[785,1374,419,482]" gridcol="2" gridrow="2" pageId="66" pageNumber="67">180–185 174 168–185 168–189 163–166 154 - -</td>
</tr>
</table>
</paragraph>
<paragraph id="8BA96D76AB1F695BFF47FE3EFA80FB93" pageId="66" pageNumber="67">
<table id="F9169FD6AB1F96E6FF47FE3EFAD2FB93" box="[151,1374,501,1143]" gridcols="5" gridrows="6" pageId="66" pageNumber="67">
<tr id="35266F34AB1F96E6FF47FE3EFAD2FDD0" box="[151,1374,501,564]" gridrow="0" pageId="66" pageNumber="67">
<th id="76F70648AB1F96E6FF47FE3EFEB0FDD0" box="[151,316,501,564]" gridcol="0" gridrow="0" pageId="66" pageNumber="67">P-1 dL/H P-2 dL/H</th>
<th id="76F70648AB1F96E6FE11FE3EFCFBFDD0" box="[449,887,501,564]" gridcol="1" gridrow="0" pageId="66" pageNumber="67">21–23/19–22 23–29/21–24 22–23 47–50/33–35 51–58/36–38 49–50</th>
<th id="76F70648AB1F96E6FC69FE3EFB93FDD0" box="[953,1055,501,564]" gridcol="2" gridrow="0" pageId="66" pageNumber="67">20 45–48</th>
<th id="76F70648AB1F96E6FB8CFE3EFB10FDD0" box="[1116,1180,501,564]" gridcol="3" gridrow="0" pageId="66" pageNumber="67">20–23 45–50</th>
<th id="76F70648AB1F96E6FAD4FE3EFAD2FDD0" box="[1284,1374,501,564]" gridcol="4" gridrow="0" pageId="66" pageNumber="67">22–26 46–51</th>
</tr>
<tr id="35266F34AB1F96E6FF47FD83FAD2FD63" box="[151,1374,584,647]" gridrow="1" pageId="66" pageNumber="67">
<th id="76F70648AB1F96E6FF47FD83FEB0FD63" box="[151,316,584,647]" gridcol="0" gridrow="1" pageId="66" pageNumber="67">P-3 dL/H P-4 dL/H</th>
<td id="76F70648AB1F96E6FE11FD83FCFBFD63" box="[449,887,584,647]" gridcol="1" gridrow="1" pageId="66" pageNumber="67">34–36/27–28 35–39/28–31 32–33 45–46/18–20 45–48/20–22 42–43</td>
<td id="76F70648AB1F96E6FC69FD83FB93FD63" box="[953,1055,584,647]" gridcol="2" gridrow="1" pageId="66" pageNumber="67">32 40–41</td>
<td id="76F70648AB1F96E6FB8CFD83FB10FD63" box="[1116,1180,584,647]" gridcol="3" gridrow="1" pageId="66" pageNumber="67">30–34 40–45</td>
<td id="76F70648AB1F96E6FAD4FD83FAD2FD63" box="[1284,1374,584,647]" gridcol="4" gridrow="1" pageId="66" pageNumber="67">32–35 40–45</td>
</tr>
<tr id="35266F34AB1F96E6FF47FD50FAD2FCC8" box="[151,1374,667,812]" gridrow="2" pageId="66" pageNumber="67">
<th id="76F70648AB1F96E6FF47FD50FEB0FCC8" box="[151,316,667,812]" gridcol="0" gridrow="2" pageId="66" pageNumber="67">P-5 dL/H P-1 dL/H ratio P-2 dL/H ratio P-3 dL/H ratio</th>
<td id="76F70648AB1F96E6FE11FD50FCFBFCC8" box="[449,887,667,812]" gridcol="1" gridrow="2" pageId="66" pageNumber="67">18–20/10–12 19–20/11–12 17–18 1.0–1.2 1.1–1.2 - 1.35–1.52 1.4–1.54 - 1.23–1.31 1.24–1.25 -</td>
<td id="76F70648AB1F96E6FC69FD50FB93FCC8" box="[953,1055,667,812]" gridcol="2" gridrow="2" pageId="66" pageNumber="67">17 - - -</td>
<td id="76F70648AB1F96E6FB8CFD50FB10FCC8" box="[1116,1180,667,812]" gridcol="3" gridrow="2" pageId="66" pageNumber="67">15–19 - - -</td>
<td id="76F70648AB1F96E6FAD4FD50FAD2FCC8" box="[1284,1374,667,812]" gridcol="4" gridrow="2" pageId="66" pageNumber="67">16–18 - - -</td>
</tr>
<tr id="35266F34AB1F96E6FF47FC8BFAD2FC9B" box="[151,1374,832,895]" gridrow="3" pageId="66" pageNumber="67">
<th id="76F70648AB1F96E6FF47FC8BFEB0FC9B" box="[151,316,832,895]" gridcol="0" gridrow="3" pageId="66" pageNumber="67">P-4 dL/H ratio P-5 dL/H ratio</th>
<td id="76F70648AB1F96E6FE11FC8BFCFBFC9B" box="[449,887,832,895]" gridcol="1" gridrow="3" pageId="66" pageNumber="67">2.3–2.5 2.22–2.27 - 1.74–2.0 1.7 -</td>
<td id="76F70648AB1F96E6FC69FC8BFB93FC9B" box="[953,1055,832,895]" gridcol="2" gridrow="3" pageId="66" pageNumber="67">- -</td>
<td id="76F70648AB1F96E6FB8CFC8BFB10FC9B" box="[1116,1180,832,895]" gridcol="3" gridrow="3" pageId="66" pageNumber="67">- -</td>
<td id="76F70648AB1F96E6FAD4FC8BFAD2FC9B" box="[1284,1374,832,895]" gridcol="4" gridrow="3" pageId="66" pageNumber="67">- -</td>
</tr>
<tr id="35266F34AB1F96E6FF47FC58FAD2FBC0" box="[151,1374,915,1060]" gridrow="4" pageId="66" pageNumber="67">
<th id="76F70648AB1F96E6FF47FC58FEB0FBC0" box="[151,316,915,1060]" gridcol="0" gridrow="4" pageId="66" pageNumber="67">dL P-2/P-4 ratio I-L-4 L I-L-5 L I-L-6 L</th>
<td id="76F70648AB1F96E6FE11FC58FCFBFBC0" box="[449,887,915,1060]" gridcol="1" gridrow="4" pageId="66" pageNumber="67">1.03–1.12 1.1–1.2 1.14–1.19 82–101 97 - 86–102 100 - 88–98 91–92 -</td>
<td id="76F70648AB1F96E6FC69FC58FB93FBC0" box="[953,1055,915,1060]" gridcol="2" gridrow="4" pageId="66" pageNumber="67">1.12–1.17 - - -</td>
<td id="76F70648AB1F96E6FB8CFC58FB10FBC0" box="[1116,1180,915,1060]" gridcol="3" gridrow="4" pageId="66" pageNumber="67">- - - -</td>
<td id="76F70648AB1F96E6FAD4FC58FAD2FBC0" box="[1284,1374,915,1060]" gridcol="4" gridrow="4" pageId="66" pageNumber="67">- - - -</td>
</tr>
<tr id="35266F34AB1F96E6FF47FBF3FAD2FB93" box="[151,1374,1080,1143]" gridrow="5" pageId="66" pageNumber="67">
<th id="76F70648AB1F96E6FF47FBF3FEB0FB93" box="[151,316,1080,1143]" gridcol="0" gridrow="5" pageId="66" pageNumber="67">I-L-6 H I-L-6 L/H ratio</th>
<td id="76F70648AB1F96E6FE11FBF3FCFBFB93" box="[449,887,1080,1143]" gridcol="1" gridrow="5" pageId="66" pageNumber="67">35–37 38–39 - 2.38–2.6 2.38 -</td>
<td id="76F70648AB1F96E6FC69FBF3FB93FB93" box="[953,1055,1080,1143]" gridcol="2" gridrow="5" pageId="66" pageNumber="67">- -</td>
<td id="76F70648AB1F96E6FB8CFBF3FB10FB93" box="[1116,1180,1080,1143]" gridcol="3" gridrow="5" pageId="66" pageNumber="67">- -</td>
<td id="76F70648AB1F96E6FAD4FBF3FAD2FB93" box="[1284,1374,1080,1143]" gridcol="4" gridrow="5" pageId="66" pageNumber="67">- -</td>
</tr>
</table>
</paragraph>
<paragraph id="8BA96D76AB1F695BFF17F992FB09F91A" blockId="66.[151,1436,1202,2037]" pageId="66" pageNumber="67">
There is a large group of African species sharing the more or less elongated idiosoma and the box-shaped gnathosomal bay. This group includes several similar but still weakly defined species, i.e., 
<taxonomicName id="4C1616F5AB1F695BFB29F9B7FE6DF95C" authority="Koenike, 1898" authorityName="Koenike" authorityYear="1898" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="microstoma">
<emphasis id="B962B164AB1F695BFB29F9B7FE96F953" italics="true" pageId="66" pageNumber="67">Monatractides microstoma</emphasis>
(
<bibRefCitation id="EF871087AB1F695BFEFCF954FE54F95C" author="Koenike" box="[300,472,1695,1720]" pageId="66" pageNumber="67" refString="Koenike, F. (1898) Hydrachniden-Fauna von Madagaskar und Nossi-Be. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft in Frankfurt am Main, 21 (2), 295 - 435." type="journal article" year="1898">Koenike, 1898</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="4C1616F5AB1F695BFE20F96BFCFCF95C" authority="Lundblad, 1951" authorityName="Lundblad" authorityYear="1951" box="[496,880,1695,1720]" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="amplipalpis">
<emphasis id="B962B164AB1F695BFE20F96BFD11F95C" box="[496,669,1695,1720]" italics="true" pageId="66" pageNumber="67">M. amplipalpis</emphasis>
(Lundblad, 1951)
</taxonomicName>
, 
<taxonomicName id="4C1616F5AB1F695BFCAFF96BFB59F95C" authority="Cook, 1966" authorityName="Cook" authorityYear="1966" box="[895,1237,1695,1720]" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="longicoxalis">
<emphasis id="B962B164AB1F695BFCAFF96BFBBDF95C" box="[895,1073,1695,1720]" italics="true" pageId="66" pageNumber="67">M. longicoxalis</emphasis>
(
<bibRefCitation id="EF871087AB1F695BFB92F954FB42F95C" author="Cook" box="[1090,1230,1695,1720]" pageId="66" pageNumber="67" refString="Cook, D. R. (1966) The water mites of Liberia. Memoirs of the American Entomological Institute, 6, 1 - 418." type="journal article" year="1966">Cook, 1966</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="4C1616F5AB1F695BFB34F96BFEA2F93F" authority="Cook, 1966" authorityName="Cook" authorityYear="1966" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="dolichosoma">
<emphasis id="B962B164AB1F695BFB34F96BFA10F95C" box="[1252,1436,1695,1720]" italics="true" pageId="66" pageNumber="67">M. dolichosoma</emphasis>
(
<bibRefCitation id="EF871087AB1F695BFF4FF909FEABF93F" author="Cook" box="[159,295,1730,1755]" pageId="66" pageNumber="67" refString="Cook, D. R. (1966) The water mites of Liberia. Memoirs of the American Entomological Institute, 6, 1 - 418." type="journal article" year="1966">Cook, 1966</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="4C1616F5AB1F695BFEEAF90FFD3FF93F" authority="K.O. Viets, 1974" authorityName="K.O. Viets" authorityYear="1974" box="[314,691,1730,1755]" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="congoensis">
<emphasis id="B962B164AB1F695BFEEAF90FFE57F93F" box="[314,475,1732,1755]" italics="true" pageId="66" pageNumber="67">M. congoensis</emphasis>
(
<bibRefCitation id="EF871087AB1F695BFE3BF908FD26F93F" author="Viets" box="[491,682,1730,1755]" pageId="66" pageNumber="67" refString="Viets, K. O. &amp; Bottger, K. (1974) Zur Systematik und Okologie rheophiler Hydrachnellae (Acari) Zentralafrikas. Teil I. Acarologia, 16 (1), 106 - 159." type="journal article" year="1974">K.O. Viets, 1974</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="4C1616F5AB1F695BFD6EF90FFB8FF93F" authority="K.O. Viets, 1974" authorityName="K.O. Viets" authorityYear="1974" box="[702,1027,1730,1755]" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="rivalis">
<emphasis id="B962B164AB1F695BFD6EF90FFCA7F93F" box="[702,811,1730,1755]" italics="true" pageId="66" pageNumber="67">M. rivalis</emphasis>
(
<bibRefCitation id="EF871087AB1F695BFCEBF908FC77F93F" author="Viets" box="[827,1019,1730,1755]" pageId="66" pageNumber="67" refString="Viets, K. O. &amp; Bottger, K. (1974) Zur Systematik und Okologie rheophiler Hydrachnellae (Acari) Zentralafrikas. Teil I. Acarologia, 16 (1), 106 - 159." type="journal article" year="1974">K.O. Viets, 1974</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="4C1616F5AB1F695BFBDEF90FFAE1F93F" authority="K.O. Viets, 1974" authorityName="K.O. Viets" authorityYear="1974" box="[1038,1389,1730,1755]" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="procursa">
<emphasis id="B962B164AB1F695BFBDEF90FFB1BF93F" box="[1038,1175,1732,1755]" italics="true" pageId="66" pageNumber="67">M. procursa</emphasis>
(
<bibRefCitation id="EF871087AB1F695BFB77F908FAE9F93F" author="Viets" box="[1191,1381,1730,1755]" pageId="66" pageNumber="67" refString="Viets, K. O. &amp; Bottger, K. (1974) Zur Systematik und Okologie rheophiler Hydrachnellae (Acari) Zentralafrikas. Teil I. Acarologia, 16 (1), 106 - 159." type="journal article" year="1974">K.O. Viets, 1974</bibRefCitation>
)
</taxonomicName>
and 
<taxonomicName id="4C1616F5AB1F695BFF47F92CFD18F91A" authority="Goldschmidt &amp; Smit 2009" authorityName="Goldschmidt &amp; Smit" authorityYear="2009" box="[151,660,1765,1790]" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="nigromontanus">
<emphasis id="B962B164AB1F695BFF47F92CFEE8F91A" box="[151,356,1767,1790]" italics="true" pageId="66" pageNumber="67">M. nigromontanus</emphasis>
<bibRefCitation id="EF871087AB1F695BFEBCF92EFD18F91A" author="Goldschmidt" box="[364,660,1765,1790]" pageId="66" pageNumber="67" refString="Goldschmidt, T. &amp; Smit. H. (2009) Studies on torrenticolid water mites mainly from South Africa - Torrenticola Piersig, 1896 and Monatractides K. Viets, 1926 (Acari: Hydrachnidia: Torrenticolidae). International Journal of Acarology, 35 (3), 179 - 234. http: // dx. doi. org / 10.1080 / 01647950902980286" type="journal article" year="2009">Goldschmidt &amp; Smit 2009</bibRefCitation>
</taxonomicName>
(see 
<figureCitation id="132D71F3AB1F695BFD1FF92EFC92F91A" box="[719,798,1765,1790]" captionStart="FIGURE 45" captionStartId="67.[151,250,1276,1298]" captionTargetBox="[151,1436,195,1251]" captionTargetId="figure@67.[151,1436,195,1251]" captionTargetPageId="67" captionText="FIGURE 45. Distribution map of some selected Monatractides species (species number corresponds with numbers in Map): 1 — M. microstoma Koenike, 1898, 2 — M. amplipalpis Lundblad, 1951, 3 — M. longicoxalis Cook, 1966, 4 — M. dolichosoma Cook, 1966, 5 — M. congoensis K. O. Viets, 1974, 6 — M. rivalis K. O. Viets, 1974, 7 — M. procursa K. O. Viets, 1974, 8 — M. nigromontanus Goldschmidt &amp; Smit, 2009; populations of Monatractides microstoma species-complex from Ghana are marked by X." httpUri="https://zenodo.org/record/286348/files/figure.png" pageId="66" pageNumber="67">Fig. 45</figureCitation>
for their distribution in Africa).
</paragraph>
<paragraph id="8BA96D76AB1F695BFF17F8C1FD79F811" blockId="66.[151,1436,1202,2037]" pageId="66" pageNumber="67">
Due to the similar shape of the idiosoma and palp the specimens from our study most closely resemble 
<taxonomicName id="4C1616F5AB1F695BFF47F8E7FD09F8A1" authority="Koenike, 1898" authorityName="Koenike" authorityYear="1898" box="[151,645,1836,1861]" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="microstoma">
<emphasis id="B962B164AB1F695BFF47F8E7FE49F8A0" box="[151,453,1836,1861]" italics="true" pageId="66" pageNumber="67">Monatractides microstoma</emphasis>
(
<bibRefCitation id="EF871087AB1F695BFE05F8E7FDF1F8A1" author="Koenike" box="[469,637,1836,1861]" pageId="66" pageNumber="67" refString="Koenike, F. (1898) Hydrachniden-Fauna von Madagaskar und Nossi-Be. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft in Frankfurt am Main, 21 (2), 295 - 435." type="journal article" year="1898">Koenike, 1898</bibRefCitation>
)
</taxonomicName>
and 
<taxonomicName id="4C1616F5AB1F695BFD6DF8E6FBA8F8A1" authority="K.O. Viets, 1974" authorityName="K.O. Viets" authorityYear="1974" box="[701,1060,1836,1861]" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="procursa">
<emphasis id="B962B164AB1F695BFD6DF8E6FCC4F8A0" box="[701,840,1837,1860]" italics="true" pageId="66" pageNumber="67">M. procursa</emphasis>
(
<bibRefCitation id="EF871087AB1F695BFC89F8E6FB97F8A1" author="Viets" box="[857,1051,1836,1861]" pageId="66" pageNumber="67" refString="Viets, K. O. &amp; Bottger, K. (1974) Zur Systematik und Okologie rheophiler Hydrachnellae (Acari) Zentralafrikas. Teil I. Acarologia, 16 (1), 106 - 159." type="journal article" year="1974">K.O. Viets, 1974</bibRefCitation>
)
</taxonomicName>
. The latter species was described by K.O. Viets (in 
<bibRefCitation id="EF871087AB1F695BFE8DF89BFD17F88C" author="Viets" box="[349,667,1871,1896]" pageId="66" pageNumber="67" refString="Viets, K. O. &amp; Bottger, K. (1974) Zur Systematik und Okologie rheophiler Hydrachnellae (Acari) Zentralafrikas. Teil I. Acarologia, 16 (1), 106 - 159." type="journal article" year="1974">K.O. Viets &amp; Böttger (1974)</bibRefCitation>
from 
<collectingCountry id="F3012DE6AB1F695BFD0FF89BFCA6F88C" box="[735,810,1872,1896]" name="Republic of the Congo" pageId="66" pageNumber="67">Congo</collectingCountry>
(
<collectingCountry id="F3012DE6AB1F695BFCE8F884FCF5F88C" box="[824,889,1871,1896]" name="Democratic Republic of the Congo" pageId="66" pageNumber="67">Zaire</collectingCountry>
). Due to the dorsal plate with the area of primary sclerotization bearing two dorsoglandularia and similar shape of the genital field, 
<taxonomicName id="4C1616F5AB1F695BFBE0F8BFFB36F86F" box="[1072,1210,1908,1931]" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="procursa">
<emphasis id="B962B164AB1F695BFBE0F8BFFB36F86F" box="[1072,1210,1908,1931]" italics="true" pageId="66" pageNumber="67">M. procursa</emphasis>
</taxonomicName>
resembles more the specimens of Form ''B'' in our study. Differences with figures and measurements in the original description are found in the excretory pore shifted to the line of primary sclerotization in the female (see Fig. 
<quantity id="4CEEC093AB1F695BFB15F872FB73F836" box="[1221,1279,1977,2002]" metricMagnitude="0" metricUnit="m" metricValue="1.905" pageId="66" pageNumber="67" unit="in" value="75.0">75 in</quantity>
<bibRefCitation id="EF871087AB1F695BFAD7F871FEA0F811" author="Viets" pageId="66" pageNumber="67" refString="Viets, K. O. &amp; Bottger, K. (1974) Zur Systematik und Okologie rheophiler Hydrachnellae (Acari) Zentralafrikas. Teil I. Acarologia, 16 (1), 106 - 159." type="journal article" year="1974">K.O. Viets &amp; Böttger 1974</bibRefCitation>
, postgenital area in male not illustrated).
</paragraph>
<caption id="DF693DFEAB1E695AFF47FB37FF5CFA68" httpUri="https://zenodo.org/record/286348/files/figure.png" pageId="67" pageNumber="68" targetBox="[151,1436,195,1251]" targetPageId="67">
<paragraph id="8BA96D76AB1E695AFF47FB37FF5CFA68" blockId="67.[151,1437,1276,1420]" pageId="67" pageNumber="68">
<emphasis id="B962B164AB1E695AFF47FB37FEADFAF6" bold="true" box="[151,289,1276,1298]" pageId="67" pageNumber="68">FIGURE 45.</emphasis>
Distribution map of some selected 
<taxonomicName id="4C1616F5AB1E695AFD42FB37FCAAFAF6" box="[658,806,1276,1298]" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="67" pageNumber="68" phylum="Arthropoda" rank="genus">
<emphasis id="B962B164AB1E695AFD42FB37FCAAFAF6" box="[658,806,1276,1298]" italics="true" pageId="67" pageNumber="68">Monatractides</emphasis>
</taxonomicName>
species (species number corresponds with numbers in Map): 1— 
<taxonomicName id="4C1616F5AB1E695AFF6DFAD0FE74FAD4" authority="Koenike, 1898" authorityName="Koenike" authorityYear="1898" box="[189,504,1306,1328]" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="microstoma">
<emphasis id="B962B164AB1E695AFF6DFAD0FEDBFAD4" box="[189,343,1307,1328]" italics="true" pageId="67" pageNumber="68">M. microstoma</emphasis>
Koenike, 1898
</taxonomicName>
, 2— 
<taxonomicName id="4C1616F5AB1E695AFDFAFAD0FCFDFAD4" authority="Lundblad, 1951" authorityName="Lundblad" authorityYear="1951" box="[554,881,1306,1328]" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="amplipalpis">
<emphasis id="B962B164AB1E695AFDFAFAD0FDCAFAD4" box="[554,582,1307,1328]" italics="true" pageId="67" pageNumber="68">M.</emphasis>
<emphasis id="B962B164AB1E695AFD9DFAD1FD49FAD4" box="[589,709,1306,1328]" italics="true" pageId="67" pageNumber="68">amplipalpis</emphasis>
Lundblad, 1951
</taxonomicName>
, 3— 
<taxonomicName id="4C1616F5AB1E695AFC74FAD0FB49FAD4" authority="Cook, 1966" authorityName="Cook" authorityYear="1966" box="[932,1221,1306,1328]" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="longicoxalis">
<emphasis id="B962B164AB1E695AFC74FAD0FC4CFAD4" box="[932,960,1307,1328]" italics="true" pageId="67" pageNumber="68">M.</emphasis>
<emphasis id="B962B164AB1E695AFC17FAD1FBCFFAD4" box="[967,1091,1306,1328]" italics="true" pageId="67" pageNumber="68">longicoxalis</emphasis>
Cook, 1966
</taxonomicName>
, 4— 
<taxonomicName id="4C1616F5AB1E695AFB27FAD0FE9FFAAB" authority="Cook, 1966" authorityName="Cook" authorityYear="1966" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="dolichosoma">
<emphasis id="B962B164AB1E695AFB27FAD0FA98FAD4" box="[1271,1300,1307,1328]" italics="true" pageId="67" pageNumber="68">M.</emphasis>
<emphasis id="B962B164AB1E695AFACBFAD1FA11FAD4" box="[1307,1437,1306,1328]" italics="true" pageId="67" pageNumber="68">dolichosoma</emphasis>
Cook, 1966
</taxonomicName>
, 5— 
<taxonomicName id="4C1616F5AB1E695AFE97FAF1FD1FFAAB" authority="K.O.Viets, 1974" authorityName="K.O.Viets" authorityYear="1974" box="[327,659,1337,1359]" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="congoensis">
<emphasis id="B962B164AB1E695AFE97FAF1FEE8FAAB" box="[327,356,1338,1359]" italics="true" pageId="67" pageNumber="68">M.</emphasis>
<emphasis id="B962B164AB1E695AFEBDFAF1FE51FAAB" box="[365,477,1338,1359]" italics="true" pageId="67" pageNumber="68">congoensis</emphasis>
K.O.Viets, 1974
</taxonomicName>
, 6— 
<taxonomicName id="4C1616F5AB1E695AFD17FAF1FC6FFAAB" authority="K.O.Viets, 1974" authorityName="K.O.Viets" authorityYear="1974" box="[711,995,1337,1359]" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="rivalis">
<emphasis id="B962B164AB1E695AFD17FAF1FD68FAAB" box="[711,740,1338,1359]" italics="true" pageId="67" pageNumber="68">M.</emphasis>
<emphasis id="B962B164AB1E695AFD3DFAF2FCA2FAAB" box="[749,814,1337,1359]" italics="true" pageId="67" pageNumber="68">rivalis</emphasis>
K.O.Viets, 1974
</taxonomicName>
, 7— 
<taxonomicName id="4C1616F5AB1E695AFBC7FAF1FAC0FAAB" authority="K.O.Viets, 1974" authorityName="K.O.Viets" authorityYear="1974" box="[1047,1356,1337,1359]" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="procursa">
<emphasis id="B962B164AB1E695AFBC7FAF1FBBFFAAB" box="[1047,1075,1338,1359]" italics="true" pageId="67" pageNumber="68">M.</emphasis>
<emphasis id="B962B164AB1E695AFBECFAF1FB1AFAAB" box="[1084,1174,1338,1359]" italics="true" pageId="67" pageNumber="68">procursa</emphasis>
K.O.Viets, 1974
</taxonomicName>
, 8— 
<taxonomicName id="4C1616F5AB1E695AFA50FAF1FDC4FA8A" authority="Goldschmidt &amp; Smit, 2009" authorityName="Goldschmidt &amp; Smit" authorityYear="2009" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="nigromontanus">
<emphasis id="B962B164AB1E695AFA50FAF1FEBCFA8A" italics="true" pageId="67" pageNumber="68">M. nigromontanus</emphasis>
Goldschmidt &amp; Smit, 2009
</taxonomicName>
; populations of 
<taxonomicName id="4C1616F5AB1E695AFD39FA93FC75FA8A" box="[745,1017,1368,1390]" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="microstoma">
<emphasis id="B962B164AB1E695AFD39FA93FC75FA8A" box="[745,1017,1368,1390]" italics="true" pageId="67" pageNumber="68">Monatractides microstoma</emphasis>
</taxonomicName>
species-complex from Ghana are marked by X.
</paragraph>
</caption>
<paragraph id="8BA96D76AB1E695AFF17FA7FFB74F8BC" blockId="67.[151,1437,1460,2025]" pageId="67" pageNumber="68">
<taxonomicName id="4C1616F5AB1E695AFF17FA7FFE7BFA28" box="[199,503,1460,1485]" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="microstoma">
<emphasis id="B962B164AB1E695AFF17FA7FFE7BFA28" box="[199,503,1460,1485]" italics="true" pageId="67" pageNumber="68">Monatractides microstoma</emphasis>
</taxonomicName>
was decribed by 
<bibRefCitation id="EF871087AB1E695AFD10FA7FFCFEFA28" author="Koenike" box="[704,882,1460,1485]" pageId="67" pageNumber="68" refString="Koenike, F. (1898) Hydrachniden-Fauna von Madagaskar und Nossi-Be. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft in Frankfurt am Main, 21 (2), 295 - 435." type="journal article" year="1898">Koenike (1898)</bibRefCitation>
from 
<collectingCountry id="F3012DE6AB1E695AFC6AFA7FFBA3FA28" box="[954,1071,1460,1484]" name="Cameroon" pageId="67" pageNumber="68">Cameroon</collectingCountry>
based on a single female. Later on a detailed description of the female (from Manoka near Duala, prep. no. 1390) was given by 
<bibRefCitation id="EF871087AB1E695AFB35FA13FA15FA14" author="Viets" box="[1253,1433,1495,1520]" pageId="67" pageNumber="68" refString="Viets, K. (1958) Bemerkungen zu zwei afrikanischen Torrenticola-Arten (Hydrachnellae, Acari). Zoologischer Anzeiger, 161 (3 - 4), 90 - 104." type="journal article" year="1958">K. Viets (1958)</bibRefCitation>
. 
<bibRefCitation id="EF871087AB1E695AFF47FA37FEA4F9F1" author="Cook" box="[151,296,1532,1557]" pageId="67" pageNumber="68" refString="Cook, D. R. (1966) The water mites of Liberia. Memoirs of the American Entomological Institute, 6, 1 - 418." type="journal article" year="1966">Cook (1966)</bibRefCitation>
reported 
<taxonomicName id="4C1616F5AB1E695AFE49FA36FDC8F9F0" box="[409,580,1533,1556]" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="microstoma">
<emphasis id="B962B164AB1E695AFE49FA36FDC8F9F0" box="[409,580,1533,1556]" italics="true" pageId="67" pageNumber="68">M. microstoma</emphasis>
</taxonomicName>
as the most frequently collected and most widely distributed torrenticolid species in 
<collectingCountry id="F3012DE6AB1E695AFEC6F9D4FEEBF9DC" box="[278,359,1567,1592]" name="Liberia" pageId="67" pageNumber="68">Liberia</collectingCountry>
and provided illustrations and descriptions of the male. 
<bibRefCitation id="EF871087AB1E695AFBDDF9EBFB6CF9DC" author="Viets" box="[1037,1248,1567,1592]" pageId="67" pageNumber="68" refString="Viets, K. O. (1971) Wassermilben aus Sudafrika (Hydrachnellae, Acari). Acarologia, 12 (4), 752 - 779." type="journal article" year="1971">K.O. Viets (1971)</bibRefCitation>
published from 
<collectingCountry id="F3012DE6AB1E695AFF47F98FFE83F9B9" box="[151,271,1604,1629]" name="Zimbabwe" pageId="67" pageNumber="68">Zimbabwe</collectingCountry>
a female slightly larger than those from 
<collectingCountry id="F3012DE6AB1E695AFD02F98FFCAFF9B9" box="[722,803,1604,1629]" name="Liberia" pageId="67" pageNumber="68">Liberia</collectingCountry>
also under the name 
<taxonomicName id="4C1616F5AB1E695AFBDFF98EFB1EF9B8" box="[1039,1170,1605,1628]" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="microstoma">
<emphasis id="B962B164AB1E695AFBDFF98EFB1EF9B8" box="[1039,1170,1605,1628]" italics="true" pageId="67" pageNumber="68">microstoma</emphasis>
</taxonomicName>
. Due to the dorsal plate with the area of primary sclerotization bearing four dorsoglandularia, 
<taxonomicName id="4C1616F5AB1E695AFC0BF9A2FB06F964" box="[987,1162,1641,1664]" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="microstoma">
<emphasis id="B962B164AB1E695AFC0BF9A2FB06F964" box="[987,1162,1641,1664]" italics="true" pageId="67" pageNumber="68">M. microstoma</emphasis>
</taxonomicName>
is more similar to the specimens of Form ''A'' of our study. Males from 
<collectingCountry id="F3012DE6AB1E695AFD1CF947FC91F941" box="[716,797,1676,1701]" name="Liberia" pageId="67" pageNumber="68">Liberia</collectingCountry>
differ from those in our study in a medial suture of Cx- II+III of moderate length, the genital field anteriorly less projecting and slightly pointed and a less extended postgenital area (see 
<bibRefCitation id="EF871087AB1E695AFE54F91FFD8EF909" author="Cook" box="[388,514,1748,1773]" pageId="67" pageNumber="68" refString="Cook, D. R. (1966) The water mites of Liberia. Memoirs of the American Entomological Institute, 6, 1 - 418." type="journal article" year="1966">Cook 1966</bibRefCitation>
, fig. 262). Among the specimens from 
<collectingCountry id="F3012DE6AB1E695AFC6CF91FFB89F909" box="[956,1029,1748,1773]" name="Ghana" pageId="67" pageNumber="68">Ghana</collectingCountry>
we found a single male from Tagbo stream which fits well Cook’s (1966) description of 
<taxonomicName id="4C1616F5AB1E695AFD3BF932FCE2F8F4" box="[747,878,1785,1808]" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="microstoma">
<emphasis id="B962B164AB1E695AFD3BF932FCE2F8F4" box="[747,878,1785,1808]" italics="true" pageId="67" pageNumber="68">microstoma</emphasis>
</taxonomicName>
(see 
<figureCitation id="132D71F3AB1E695AFC7CF93CFBB3F8F4" box="[940,1087,1783,1808]" captionStart="FIGURE 46 A – C" captionStartId="68.[151,250,1283,1305]" captionTargetBox="[169,1426,483,1258]" captionTargetId="figure@68.[163,1437,473,1266]" captionTargetPageId="68" captionText="FIGURE 46 A – C. Monatractides microstoma - species complex, male, Tagbo River: A = dorsal shield; B = idiosoma, ventral view; C = slide-mounted ejaculatory complex reduced to sclerotized framework. Scale bars = 100 Μm." httpUri="https://zenodo.org/record/286349/files/figure.png" pageId="67" pageNumber="68">Figs. 46A–C</figureCitation>
, and measurements in 
<tableCitation id="C69458CDAB1E695AFA94F93CFA02F8F4" box="[1348,1422,1783,1808]" captionStart="TABLE 4" captionStartId="64.[151,239,152,175]" captionTargetBox="[157,1429,234,1960]" captionTargetPageId="64" captionText="TABLE 4. Measurements of Monatractides microstoma (Koenike, 1898) — species complex (Form “ A “) from Ghana, and female of M. microstoma from Cameroon (from K. Viets 1958, prep. no. 1390; * calculated from figure)." httpUri="http://table.plazi.org/id/DF693DFEAB1D6959FF47FF53FAA8FF2B" pageId="67" pageNumber="68" tableUuid="DF693DFEAB1D6959FF47FF53FAA8FF2B">Tab. 4</tableCitation>
). This specimen also agrees in the shape of the dorsal shield, gnathosoma elongated in lateral view and P-4 without lacking denticle at insertion of ventral setae, but it is notably larger than our specimens from 
<collectingCountry id="F3012DE6AB1E695AFB78F8F4FB79F8BC" box="[1192,1269,1855,1880]" name="Ghana" pageId="67" pageNumber="68">Ghana</collectingCountry>
.
</paragraph>
<paragraph id="8BA96D76AB1E695AFF17F8AFFB58F844" blockId="67.[151,1437,1460,2025]" pageId="67" pageNumber="68">
As the 
<typeStatus id="54ADD3D4AB1E695AFECDF8AFFEF3F899" box="[285,383,1892,1917]" pageId="67" pageNumber="68" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="4C1616F5AB1E695AFE7BF8AEFDDAF898" box="[427,598,1893,1916]" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="microstoma">
<emphasis id="B962B164AB1E695AFE7BF8AEFDDAF898" box="[427,598,1893,1916]" italics="true" pageId="67" pageNumber="68">M. microstoma</emphasis>
</taxonomicName>
is lost (see 
<bibRefCitation id="EF871087AB1E695AFD38F8AEFC00F899" author="Viets" box="[744,908,1892,1917]" pageId="67" pageNumber="68" refString="Viets, K. (1958) Bemerkungen zu zwei afrikanischen Torrenticola-Arten (Hydrachnellae, Acari). Zoologischer Anzeiger, 161 (3 - 4), 90 - 104." type="journal article" year="1958">K. Viets 1958</bibRefCitation>
), additional material, including the male, and selection of a 
<typeStatus id="54ADD3D4AB1E695AFEE3F842FE00F845" box="[307,396,1929,1953]" pageId="67" pageNumber="68" type="neotype">neotype</typeStatus>
from the locus typicus is necessary to clarify the taxonomy of this species.
</paragraph>
<paragraph id="8BA96D76AB1E695DFF17F867FCDEFEF8" blockId="67.[151,1437,1460,2025]" lastBlockId="68.[151,1436,151,429]" lastPageId="68" lastPageNumber="69" pageId="67" pageNumber="68">
<taxonomicName id="4C1616F5AB1E695AFF17F867FD90F820" authority="(Congo)" baseAuthorityName="Congo" box="[199,540,1964,1989]" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="rivalis">
<emphasis id="B962B164AB1E695AFF17F867FE34F821" box="[199,440,1964,1989]" italics="true" pageId="67" pageNumber="68">Monatractides rivalis</emphasis>
(
<collectingCountry id="F3012DE6AB1E695AFE17F867FD99F820" box="[455,533,1964,1988]" name="Republic of the Congo" pageId="67" pageNumber="68">Congo</collectingCountry>
)
</taxonomicName>
and 
<taxonomicName id="4C1616F5AB1E695AFD81F866FC93F820" box="[593,799,1965,1988]" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="nigromontanus">
<emphasis id="B962B164AB1E695AFD81F866FC93F820" box="[593,799,1965,1988]" italics="true" pageId="67" pageNumber="68">M. nigromontanus</emphasis>
</taxonomicName>
(
<collectingCountry id="F3012DE6AB1E695AFCFDF867FC33F821" box="[813,959,1964,1989]" name="South Africa" pageId="67" pageNumber="68">South Africa</collectingCountry>
) are two other species similar in having an elongated genital field, a box-shaped gnathosomal bay and a short medial margin of Cx-II+III. The latter species differs in a more elongated idiosoma in both sexes, a more trapezoid shape of the genital field, anteriorly less enlarged and slightly pointed, and a sligtly longer medial margin of Cx-II+III in the male (see 
<bibRefCitation id="EF871087AB19695DFB63FF77FF58FF1C" author="Goldschmidt" pageId="68" pageNumber="69" refString="Goldschmidt, T. &amp; Smit. H. (2009) Studies on torrenticolid water mites mainly from South Africa - Torrenticola Piersig, 1896 and Monatractides K. Viets, 1926 (Acari: Hydrachnidia: Torrenticolidae). International Journal of Acarology, 35 (3), 179 - 234. http: // dx. doi. org / 10.1080 / 01647950902980286" type="journal article" year="2009">Goldschmidt &amp; Smit 2009</bibRefCitation>
). 
<taxonomicName id="4C1616F5AB19695DFF38FF14FE57FF1C" box="[232,475,223,248]" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="rivalis">
<emphasis id="B962B164AB19695DFF38FF14FE57FF1C" box="[232,475,223,248]" italics="true" pageId="68" pageNumber="69">Monatractides rivalis</emphasis>
</taxonomicName>
can be distinguished by the larger dimensions in both sexes and a characteristically rounded lateral margin Cx-II (see 
<bibRefCitation id="EF871087AB19695DFDC4FECFFCCAFEF8" author="Viets" box="[532,838,260,285]" pageId="68" pageNumber="69" refString="Viets, K. O. &amp; Bottger, K. (1974) Zur Systematik und Okologie rheophiler Hydrachnellae (Acari) Zentralafrikas. Teil I. Acarologia, 16 (1), 106 - 159." type="journal article" year="1974">K.O. Viets &amp; Böttger 1974</bibRefCitation>
).
</paragraph>
<paragraph id="8BA96D76AB19695DFF17FEE3FE20FE49" blockId="68.[151,1436,151,429]" pageId="68" pageNumber="69">
For the time being we keep the populations from 
<collectingCountry id="F3012DE6AB19695DFD24FEECFCB1FEA4" box="[756,829,295,320]" name="Ghana" pageId="68" pageNumber="69">Ghana</collectingCountry>
not assigned to either 
<taxonomicName id="4C1616F5AB19695DFBECFEE2FB69FEA4" box="[1084,1253,297,320]" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="microstoma">
<emphasis id="B962B164AB19695DFBECFEE2FB69FEA4" box="[1084,1253,297,320]" italics="true" pageId="68" pageNumber="69">M. microstoma</emphasis>
</taxonomicName>
or 
<taxonomicName id="4C1616F5AB19695DFADBFEE2FA19FEA4" box="[1291,1429,297,320]" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="procursa">
<emphasis id="B962B164AB19695DFADBFEE2FA19FEA4" box="[1291,1429,297,320]" italics="true" pageId="68" pageNumber="69">M. procursa</emphasis>
</taxonomicName>
, mainly because the unclear taxonomy of 
<taxonomicName id="4C1616F5AB19695DFD40FE86FCCCFE80" box="[656,832,333,356]" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="microstoma">
<emphasis id="B962B164AB19695DFD40FE86FCCCFE80" box="[656,832,333,356]" italics="true" pageId="68" pageNumber="69">M. microstoma</emphasis>
</taxonomicName>
. Understanding the taxonomic position of the 
<taxonomicName id="4C1616F5AB19695DFAADFE86FE96FE6C" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="microstoma">
<emphasis id="B962B164AB19695DFAADFE86FE96FE6C" italics="true" pageId="68" pageNumber="69">M. microstoma</emphasis>
</taxonomicName>
- complex is not possible without additional material from a wide area and/or without the application of molecular techniques.
</paragraph>
<caption id="DF693DFEAB19695DFF47FAC8FB28FADE" httpUri="https://zenodo.org/record/286349/files/figure.png" pageId="68" pageNumber="69" targetBox="[169,1426,483,1258]" targetPageId="68">
<paragraph id="8BA96D76AB19695DFF47FAC8FB28FADE" blockId="68.[151,1435,1283,1338]" pageId="68" pageNumber="69">
<emphasis id="B962B164AB19695DFF47FAC8FEDCFAFD" bold="true" box="[151,336,1283,1305]" pageId="68" pageNumber="69">FIGURE 46A–C.</emphasis>
<taxonomicName id="4C1616F5AB19695DFE87FAC8FDE5FAFD" box="[343,617,1283,1305]" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="microstoma">
<emphasis id="B962B164AB19695DFE87FAC8FDE5FAFD" box="[343,617,1283,1305]" italics="true" pageId="68" pageNumber="69">Monatractides microstoma</emphasis>
</taxonomicName>
- species complex, male, Tagbo River: A = dorsal shield; B = idiosoma, ventral view; C = slide-mounted ejaculatory complex reduced to sclerotized framework. Scale bars = 100 Μm.
</paragraph>
</caption>
<paragraph id="8BA96D76AB19695DFF17FAA3FACBF990" blockId="68.[151,1437,1384,2034]" pageId="68" pageNumber="69">
<emphasis id="B962B164AB19695DFF17FAA3FE49FA65" bold="true" box="[199,453,1384,1409]" pageId="68" pageNumber="69">Ejaculatory complex.</emphasis>
In male specimens of form ''A'' the ejaculatory complex was not found but a sclerotized framework immediately above the genital field can be hardly visible (see 
<figureCitation id="132D71F3AB19695DFC0CFA40FBE3FA40" box="[988,1135,1419,1444]" captionStart="FIGURE 44 A – L" captionStartId="63.[151,250,1687,1709]" captionTargetBox="[184,1389,195,1651]" captionTargetId="figure@63.[184,1389,195,1651]" captionTargetPageId="63" captionText="FIGURE 44 A – L. Photographs of ejaculatory complex: A Monatractides convexiscutata (K. Viets, 1958), male, Asukuma River. B M. melpomene n. sp., Amedzofe Falls. C M. euterpe n. sp., creek upstream Sagyimase. D M. cf. ventriosus (K. Viets, 1916), male, Laboun River. E – L Monatractides microstoma - species complex (E – G—Form '' A' ', H – L—Form '' B' '; E – H—ejaculatory complex reduced to sclerotized framework above the genital field, I – L ejaculatory complex slide-mounted): E, L = specimens from Tagbo River, F, H – K = specimens from Apkonu stream, G = specimen from Plunge pool Agumatsa Falls." httpUri="https://zenodo.org/record/286347/files/figure.png" pageId="68" pageNumber="69">Figs. 44E–G</figureCitation>
). In some examined males of form “B” the ejaculatory complex was reduced to a sclerotized framework (
<figureCitation id="132D71F3AB19695DFBC6FA65FBF2FA23" box="[1046,1150,1454,1479]" captionStart="FIGURE 44 A – L" captionStartId="63.[151,250,1687,1709]" captionTargetBox="[184,1389,195,1651]" captionTargetId="figure@63.[184,1389,195,1651]" captionTargetPageId="63" captionText="FIGURE 44 A – L. Photographs of ejaculatory complex: A Monatractides convexiscutata (K. Viets, 1958), male, Asukuma River. B M. melpomene n. sp., Amedzofe Falls. C M. euterpe n. sp., creek upstream Sagyimase. D M. cf. ventriosus (K. Viets, 1916), male, Laboun River. E – L Monatractides microstoma - species complex (E – G—Form '' A' ', H – L—Form '' B' '; E – H—ejaculatory complex reduced to sclerotized framework above the genital field, I – L ejaculatory complex slide-mounted): E, L = specimens from Tagbo River, F, H – K = specimens from Apkonu stream, G = specimen from Plunge pool Agumatsa Falls." httpUri="https://zenodo.org/record/286347/files/figure.png" pageId="68" pageNumber="69">Fig. 44H</figureCitation>
) as in the previous form. However, in other specimens of form ''B'' a genital skeleton with more or less developed series of membranous chambers attached to a sclerotized framework can be observed (see 
<figureCitation id="132D71F3AB19695DFC44FA38FB97F9E8" box="[916,1051,1523,1548]" captionStart="FIGURE 44 A – L" captionStartId="63.[151,250,1687,1709]" captionTargetBox="[184,1389,195,1651]" captionTargetId="figure@63.[184,1389,195,1651]" captionTargetPageId="63" captionText="FIGURE 44 A – L. Photographs of ejaculatory complex: A Monatractides convexiscutata (K. Viets, 1958), male, Asukuma River. B M. melpomene n. sp., Amedzofe Falls. C M. euterpe n. sp., creek upstream Sagyimase. D M. cf. ventriosus (K. Viets, 1916), male, Laboun River. E – L Monatractides microstoma - species complex (E – G—Form '' A' ', H – L—Form '' B' '; E – H—ejaculatory complex reduced to sclerotized framework above the genital field, I – L ejaculatory complex slide-mounted): E, L = specimens from Tagbo River, F, H – K = specimens from Apkonu stream, G = specimen from Plunge pool Agumatsa Falls." httpUri="https://zenodo.org/record/286347/files/figure.png" pageId="68" pageNumber="69">
Figs. 
<date id="FFA84BB6AB19695DFC03FA3FFC75F9E8" box="[979,1017,1524,1548]" pageId="68" pageNumber="69" value="1944-01">44I</date>
–L
</figureCitation>
, 
<figureCitation id="132D71F3AB19695DFBF6FA38FBD5F9E8" box="[1062,1113,1523,1548]" captionStart="FIGURE 46 A – C" captionStartId="68.[151,250,1283,1305]" captionTargetBox="[169,1426,483,1258]" captionTargetId="figure@68.[163,1437,473,1266]" captionTargetPageId="68" captionText="FIGURE 46 A – C. Monatractides microstoma - species complex, male, Tagbo River: A = dorsal shield; B = idiosoma, ventral view; C = slide-mounted ejaculatory complex reduced to sclerotized framework. Scale bars = 100 Μm." httpUri="https://zenodo.org/record/286349/files/figure.png" pageId="68" pageNumber="69">46C</figureCitation>
). This ejaculatory organ was shortened apically, the proximal chamber is large, the proximal and distal arms are small, the 
<emphasis id="B962B164AB19695DFB7CF9DCFAD4F9CA" box="[1196,1368,1559,1582]" italics="true" pageId="68" pageNumber="69">carina anterior</emphasis>
is not developed and the apical setae is very long (
<figureCitation id="132D71F3AB19695DFD55F9F3FC94F9B5" box="[645,792,1592,1617]" captionStart="FIGURE 42 A – K. A – H" captionStartId="61.[151,250,1673,1695]" captionTargetBox="[160,1430,212,1641]" captionTargetId="figure@61.[151,1436,195,1648]" captionTargetPageId="61" captionText="FIGURE 42 A – K. A – H Monatractides microstoma - species complex, male (A—Form '' A' ', B – H—Form '' B' '; A, G – H = specimens from Tagbo River, B – E = specimens from Apkonu stream, F = specimen from Kintampo Falls): A – D = genital field; E – H = ejaculatory complex. I – K Monatractides jucundus (Lundblad, 1927) (redrawn from Lundblad 1952): I = ejaculatory complex, K = genital flap with apical part of ejaculatory complex opened, lateral view. Scale bars = 100 Μm." httpUri="https://zenodo.org/record/286345/files/figure.png" pageId="68" pageNumber="69">Figs. 42E–G</figureCitation>
). 
<figureCitation id="132D71F3AB19695DFCFAF9F3FC54F9B5" box="[810,984,1592,1617]" captionStart="FIGURE 42 A – K. A – H" captionStartId="61.[151,250,1673,1695]" captionTargetBox="[160,1430,212,1641]" captionTargetId="figure@61.[151,1436,195,1648]" captionTargetPageId="61" captionText="FIGURE 42 A – K. A – H Monatractides microstoma - species complex, male (A—Form '' A' ', B – H—Form '' B' '; A, G – H = specimens from Tagbo River, B – E = specimens from Apkonu stream, F = specimen from Kintampo Falls): A – D = genital field; E – H = ejaculatory complex. I – K Monatractides jucundus (Lundblad, 1927) (redrawn from Lundblad 1952): I = ejaculatory complex, K = genital flap with apical part of ejaculatory complex opened, lateral view. Scale bars = 100 Μm." httpUri="https://zenodo.org/record/286345/files/figure.png" pageId="68" pageNumber="69">Figures 42G–H</figureCitation>
illustrate the ejaculatory complex found in the largest specimens of form ''B'', possibly the last stage in the development of the ejaculatory complex.
</paragraph>
<paragraph id="8BA96D76AB19695DFF17F9B4FBA9F8C5" blockId="68.[151,1437,1384,2034]" pageId="68" pageNumber="69">We considered several possibilities: 1) it is possible that populations (or species?) with developed or reduced ejaculatory complex exist? 2), it is possible that populations (or species?) exist in which the coalescence of membranous chambers to make up a normally developed ejaculatory complex is more or less progressed and 3), the ejaculatory complex may be more or less developed in animals of different ages. At the time being we think that differences in development of ejaculatory complex might be the result of growth.</paragraph>
<paragraph id="8BA96D76AB19695DFF17F8E7FE1FF815" blockId="68.[151,1437,1384,2034]" pageId="68" pageNumber="69">
No ejaculatory complex has been described for 
<taxonomicName id="4C1616F5AB19695DFCD3F8E0FBB9F8A0" box="[771,1077,1835,1860]" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="microstoma">
<emphasis id="B962B164AB19695DFCD3F8E0FBB9F8A0" box="[771,1077,1835,1860]" italics="true" pageId="68" pageNumber="69">Monatractides microstoma</emphasis>
</taxonomicName>
, nor for 
<taxonomicName id="4C1616F5AB19695DFB77F8E6FABAF8A0" box="[1191,1334,1837,1860]" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="procursa">
<emphasis id="B962B164AB19695DFB77F8E6FABAF8A0" box="[1191,1334,1837,1860]" italics="true" pageId="68" pageNumber="69">M. procursa</emphasis>
</taxonomicName>
. A very unusually shaped ejaculatory complex was found in 
<taxonomicName id="4C1616F5AB19695DFCD4F885FB94F883" box="[772,1048,1870,1895]" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="jucundus">
<emphasis id="B962B164AB19695DFCD4F885FB94F883" box="[772,1048,1870,1895]" italics="true" pageId="68" pageNumber="69">Monatractides jucundus</emphasis>
</taxonomicName>
(see 
<figureCitation id="132D71F3AB19695DFB8DF885FB31F883" box="[1117,1213,1870,1895]" captionStart="FIGURE 42 A – K. A – H" captionStartId="61.[151,250,1673,1695]" captionTargetBox="[160,1430,212,1641]" captionTargetId="figure@61.[151,1436,195,1648]" captionTargetPageId="61" captionText="FIGURE 42 A – K. A – H Monatractides microstoma - species complex, male (A—Form '' A' ', B – H—Form '' B' '; A, G – H = specimens from Tagbo River, B – E = specimens from Apkonu stream, F = specimen from Kintampo Falls): A – D = genital field; E – H = ejaculatory complex. I – K Monatractides jucundus (Lundblad, 1927) (redrawn from Lundblad 1952): I = ejaculatory complex, K = genital flap with apical part of ejaculatory complex opened, lateral view. Scale bars = 100 Μm." httpUri="https://zenodo.org/record/286345/files/figure.png" pageId="68" pageNumber="69">
Fig. 
<date id="FFA84BB6AB19695DFB46F884FB31F883" box="[1174,1213,1871,1895]" pageId="68" pageNumber="69" value="1942-01">42I</date>
</figureCitation>
, figure taken from 
<bibRefCitation id="EF871087AB19695DFF47F8BBFEC6F86D" author="Lundblad" box="[151,330,1904,1929]" pageId="68" pageNumber="69" refString="Lundblad, O. (1952) Hydracarinen von den ostafrikanischen Gebirgen. Arkiv for Zoology, Serie 2, 3 (31), 391 - 525." type="journal article" year="1952">Lundblad 1952</bibRefCitation>
). 
<figureCitation id="132D71F3AB19695DFEB3F8BBFE64F86D" box="[355,488,1904,1929]" captionStart="FIGURE 42 A – K. A – H" captionStartId="61.[151,250,1673,1695]" captionTargetBox="[160,1430,212,1641]" captionTargetId="figure@61.[151,1436,195,1648]" captionTargetPageId="61" captionText="FIGURE 42 A – K. A – H Monatractides microstoma - species complex, male (A—Form '' A' ', B – H—Form '' B' '; A, G – H = specimens from Tagbo River, B – E = specimens from Apkonu stream, F = specimen from Kintampo Falls): A – D = genital field; E – H = ejaculatory complex. I – K Monatractides jucundus (Lundblad, 1927) (redrawn from Lundblad 1952): I = ejaculatory complex, K = genital flap with apical part of ejaculatory complex opened, lateral view. Scale bars = 100 Μm." httpUri="https://zenodo.org/record/286345/files/figure.png" pageId="68" pageNumber="69">Figure 42K</figureCitation>
(taken from 
<bibRefCitation id="EF871087AB19695DFD59F8BBFCB5F86D" author="Lundblad" box="[649,825,1904,1929]" pageId="68" pageNumber="69" refString="Lundblad, O. (1952) Hydracarinen von den ostafrikanischen Gebirgen. Arkiv for Zoology, Serie 2, 3 (31), 391 - 525." type="journal article" year="1952">Lundblad 1952</bibRefCitation>
) shows the genital flap in one of Lundblad’s fixed specimen, with the apical part of the opened ejaculatory complex ending in two apical setae and a two-parted, apically pointed membranous structure, similar to condition found in ejaculatory complex of our specimens illustrated in 
<figureCitation id="132D71F3AB19695DFEFBF812FE02F815" box="[299,398,2009,2034]" captionStart="FIGURE 42 A – K. A – H" captionStartId="61.[151,250,1673,1695]" captionTargetBox="[160,1430,212,1641]" captionTargetId="figure@61.[151,1436,195,1648]" captionTargetPageId="61" captionText="FIGURE 42 A – K. A – H Monatractides microstoma - species complex, male (A—Form '' A' ', B – H—Form '' B' '; A, G – H = specimens from Tagbo River, B – E = specimens from Apkonu stream, F = specimen from Kintampo Falls): A – D = genital field; E – H = ejaculatory complex. I – K Monatractides jucundus (Lundblad, 1927) (redrawn from Lundblad 1952): I = ejaculatory complex, K = genital flap with apical part of ejaculatory complex opened, lateral view. Scale bars = 100 Μm." httpUri="https://zenodo.org/record/286345/files/figure.png" pageId="68" pageNumber="69">Fig. 42G</figureCitation>
.
</paragraph>
<paragraph id="8BA96D76AB18695CFF17FF5CFDA7FF30" blockId="69.[151,1436,151,248]" pageId="69" pageNumber="70">
<emphasis id="B962B164AB18695CFF17FF5CFEFAFF54" bold="true" box="[199,374,151,176]" pageId="69" pageNumber="70">Malformation.</emphasis>
In one male (form ''A'') from Kintampo Falls the anterolateral platelets of dorsal shield were asymetrically developed (
<figureCitation id="132D71F3AB18695CFE67FF77FD92FF30" box="[439,542,188,213]" captionStart="FIGURE 43 A – I" captionStartId="62.[151,250,1931,1953]" captionTargetBox="[165,1436,195,1911]" captionTargetId="figure@62.[165,1450,195,1912]" captionTargetPageId="62" captionText="FIGURE 43 A – I. Photographs of dorsal (A – C) and ventral shields (D – I) of Monatractides microstoma - species complex (A, C D – G—Form '' A' ', B, H – I—Form '' B' '): A, D = male from Tagbo River, E = male from Wanjakli stream, C, F = aberrant male from Kintampo Falls, G = female from Plunge pool, B – H = male from Apkonu stream, I = female from Apkonu stream." httpUri="https://zenodo.org/record/286346/files/figure.png" pageId="69" pageNumber="70">Fig. 43C</figureCitation>
).
</paragraph>
<paragraph id="8BA96D76AB18695CFF17FF14FBE5FF1C" blockId="69.[151,1436,151,248]" box="[199,1129,223,248]" pageId="69" pageNumber="70">
<emphasis id="B962B164AB18695CFF17FF14FEA6FF1C" bold="true" box="[199,298,223,248]" pageId="69" pageNumber="70">Habitat.</emphasis>
Sandy/bouldary streams, shaded by riparian vegetation (
<figureCitation id="132D71F3AB18695CFC76FF14FBD0FF1C" box="[934,1116,223,248]" captionStart="FIGURE 53 A – E" captionStartId="77.[151,250,1655,1677]" captionTargetBox="[150,1430,237,1585]" captionTargetId="figure@77.[146,1431,237,1585]" captionTargetPageId="77" captionText="FIGURE 53 A – E. Photographs of selected sampling sites: A—Laboun River (sampling site of Torrenticola motasi, T. harrisoni, Monatractides acutiscutatus, M. cf. ventriosa, M. microstoma - species complex), B—Apkonu stream (sampling site of Torrenticola anomallela, T. harrisoni, Monatractides stigeophora, M. microstoma - species complex). C—Agumatsa River at first bridge (type locality of Monatractides thalia n. sp. and sampling site of Torrenticola harrisoni, Monatractides cf. ventriosus). D—Tagbo River, downstream of falls (sampling site of Torrenticola harrisoni, T. motasi, T. bomiensis, Monatractides microstoma - species complex). E—Tagbo River, downstream (sampling site of Torrenticola harrisoni). Photos. H. Smit." httpUri="https://zenodo.org/record/286356/files/figure.png" pageId="69" pageNumber="70">Figs. 53A–B, D</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>