312 lines
46 KiB
XML
312 lines
46 KiB
XML
<document id="5CDD7E17A9EE8D263F0C87AA1BD0532D" ID-DOI="10.5281/zenodo.193333" ID-GBIF-Dataset="a2d3a699-7d95-4ead-8e8e-00de1c6b970b" ID-ISSN="1175-5326" ID-Zenodo-Dep="193333" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460133148736" checkinUser="plazi" docAuthor="Hershler, Robert & Liu, Hsiu-Ping" docDate="2010" docId="03BF655E3E068A0AFF3D6527EA71FDB6" docLanguage="en" docName="zt02343p017.pdf" docOrigin="Zootaxa 2343" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Pyrgulopsis milleri Hershler & Liu, 2010, sp. nov." docType="treatment" docVersion="8" lastPageNumber="15" masterDocId="FF861D263E0F8A04FFAA6200E934FFC4" masterDocTitle="Two new, possibly threatened species of Pyrgulopsis (Gastropoda: Hydrobiidae) from southwestern California" masterLastPageNumber="17" masterPageNumber="1" pageNumber="10" updateTime="1698235187927" updateUser="plazi">
|
||
<mods:mods id="7BD848C1DC254D1EACA1C9E95AE13000" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="B073E26A36A06AA7518A9A02F21A3FDA">
|
||
<mods:title id="595FFF7E400C8B2CC8CCF974195CBD8F">Two new, possibly threatened species of Pyrgulopsis (Gastropoda: Hydrobiidae) from southwestern California</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="4EBF3FD04A743FE08B93CA3567367251" type="personal">
|
||
<mods:role id="E328432B75DBB9B38AB8A46246F9D95A">
|
||
<mods:roleTerm id="3A44451F37331427E1AC5D924CA877BA">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="F86758424A772B9849608AC967226A10">Hershler, Robert</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="6455E851F9E2D3795FFC87854F1164EC" type="personal">
|
||
<mods:role id="25EC02B73769737BBFC5A88E18637AE5">
|
||
<mods:roleTerm id="C5788546C0DE4CC12C95A586FB65999A">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="F0DDBB436A6CEA72D4A3F06041B9D52C">Liu, Hsiu-Ping</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="54E3210AB64F8B67BE9CA638C2BD47E6">text</mods:typeOfResource>
|
||
<mods:relatedItem id="8A45FEE513F8A4B10440013A82248220" type="host">
|
||
<mods:titleInfo id="2F5D4BE07C05F74665FFB78CD2D84499">
|
||
<mods:title id="C87FD1B4A1B6150D19507F6BD75E85CA">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="6A6BDB139DCA9C0670C004EC585590FB">
|
||
<mods:date id="8367CDEEC9EEE06921E9FA1BB0D1640D">2010</mods:date>
|
||
<mods:detail id="50D4F3857F2938D77AD78E60FFA04F51" type="volume">
|
||
<mods:number id="B06D584D8CECA207CD8C865EE3BF40D6">2343</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="05741E0889AFF992B5A361C4C9911302" unit="page">
|
||
<mods:start id="DB73D29C9150A1B7A14578BF3599AC71">1</mods:start>
|
||
<mods:end id="C96B5E77A090FA4233F3BEF888C2BD5A">17</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="D983E599805DBCF1AEDAE65D718D6D00">journal article</mods:classification>
|
||
<mods:identifier id="629B86961ED6F60687A5F0199C9657F4" type="DOI">10.5281/zenodo.193333</mods:identifier>
|
||
<mods:identifier id="D2563B14473103D0BC15D67E3E4ED619" type="GBIF-Dataset">a2d3a699-7d95-4ead-8e8e-00de1c6b970b</mods:identifier>
|
||
<mods:identifier id="5F783DB2E4EEC2673DCFDEBDBEF39DCA" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="FDB2414F61FBCAEE07194DB229017F83" type="Zenodo-Dep">193333</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03BF655E3E068A0AFF3D6527EA71FDB6" ID-DOI="http://doi.org/10.5281/zenodo.5678764" ID-GBIF-Taxon="124955708" ID-Zenodo-Dep="5678764" LSID="urn:lsid:plazi:treatment:03BF655E3E068A0AFF3D6527EA71FDB6" httpUri="http://treatment.plazi.org/id/03BF655E3E068A0AFF3D6527EA71FDB6" lastPageId="14" lastPageNumber="15" pageId="9" pageNumber="10">
|
||
<subSubSection id="C30C87C33E068A0DFF3D6527E825F8A0" pageId="9" pageNumber="10" type="nomenclature">
|
||
<paragraph id="8BA9D4483E068A0DFF3D6527E8D4F885" blockId="9.[151,480,1831,1892]" box="[151,480,1831,1857]" pageId="9" pageNumber="10">
|
||
<heading id="D0E163243E068A0DFF3D6527E8D4F885" bold="true" box="[151,480,1831,1857]" fontSize="11" level="1" pageId="9" pageNumber="10" reason="1">
|
||
<emphasis id="B962085A3E068A0DFF3D6527E8D4F885" bold="true" box="[151,480,1831,1857]" pageId="9" pageNumber="10">
|
||
<taxonomicName id="4C16AFCB3E068A0DFF3D6527E848F885" ID-CoL="7SYW9" box="[151,380,1831,1857]" class="Gastropoda" family="Hydrobiidae" genus="Pyrgulopsis" kingdom="Animalia" order="Littorinimorpha" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="milleri" status="sp. nov.">
|
||
<emphasis id="B962085A3E068A0DFF3D6527E848F885" bold="true" box="[151,380,1831,1857]" italics="true" pageId="9" pageNumber="10">Pyrgulopsis milleri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A251B5213E068A0DFE296527E8D4F885" box="[387,480,1831,1857]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BA9D4483E068A0DFF3D654AE825F8A0" blockId="9.[151,480,1831,1892]" box="[151,273,1866,1892]" pageId="9" pageNumber="10">
|
||
(
|
||
<figureCitation id="132DC8CD3E068A0DFF35654AE833F8A0" box="[159,263,1866,1892]" captionStart-0="FIGURE 6. P" captionStart-1="FIGURE 7. P" captionStart-2="FIGURE 8. P" captionStartId-0="10.[151,256,1914,1938]" captionStartId-1="11.[151,255,1779,1803]" captionStartId-2="12.[151,256,1704,1728]" captionTargetBox-0="[151,1436,260,1898]" captionTargetBox-1="[287,1300,194,1758]" captionTargetBox-2="[245,1339,350,1703]" captionTargetId-0="figure@10.[151,1436,260,1898]" captionTargetId-1="figure@11.[287,1301,194,1758]" captionTargetId-2="figure@12.[228,1360,320,1705]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionTargetPageId-2="12" captionText-0="FIGURE 6. P. milleri sp. nov. A – D. Shells. Scale = 1.0 mm. A. Holotype, SBMNH 83651. B – C. Paratypes, SBMNH 74688. D. SBMNH 74687. E. Shell apex, SBMNH 74688. Scale = 100 µm. F. Close-up of protoconch sculpture, SBMNH 74688. Scale = 100 µm. G – I. Opercula, USNM 90527. Scale = 500 µm. G. Outer side. H – I. Inner side, showing variation in thickening of attachment scar margin." captionText-1="FIGURE 7. P. milleri sp. nov., USNM 90527, scales = 10 µm. A. Portion of radula ribbon. B. Central radular teeth. C. Lateral tooth. D. Inner marginal teeth. E – F. Outer marginal teeth. Wg = wing." captionText-2="FIGURE 8. P. milleri sp. nov., USNM 90527. Scale = 0.5 mm. A – C. Penis. A. Dorsal surface. B – C. Ventral surface, showing ventral and terminal glands. D. Female glandular oviduct and associated structures (viewed from the left side). E. Bursa copulatrix and its duct. F. Seminal receptacle and its duct. Ag = albumen gland, Bu = bursa copulatrix, Cd = common duct of seminal receptacle and coiled oviduct, Cg = capsule gland, Co = coiled oviduct, Dbu = bursal duct, Ga = genital aperture, Pd = penial duct, Pf = penial filament, Pl = penial lobe, Pw = posterior wall of pallial cavity, Sr = seminal receptacle, Tg = terminal gland, Vc = ventral channel, Vg = ventral gland." httpUri-0="https://zenodo.org/record/193339/files/figure.png" httpUri-1="https://zenodo.org/record/193340/files/figure.png" httpUri-2="https://zenodo.org/record/193341/files/figure.png" pageId="9" pageNumber="10">Figs 6–8</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30C87C33E068A0EFF3D6595E882FF13" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10" type="description">
|
||
<paragraph id="8BA9D4483E068A0EFF3D6595E882FF13" blockId="9.[151,1437,1941,2004]" lastBlockId="10.[151,1436,152,215]" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
|
||
<typeStatus id="54AD6AEA3E068A0DFF3D6595E9EAF86B" box="[151,222,1941,1967]" pageId="9" pageNumber="10">
|
||
<emphasis id="B962085A3E068A0DFF3D6595E9EAF86B" bold="true" box="[151,222,1941,1967]" pageId="9" pageNumber="10">Types</emphasis>
|
||
</typeStatus>
|
||
.
|
||
<typeStatus id="54AD6AEA3E068A0DFF446595E86FF86B" box="[238,347,1941,1967]" pageId="9" pageNumber="10" type="holotype">Holotype</typeStatus>
|
||
(
|
||
<figureCitation id="132DC8CD3E068A0DFEC16595E885F86B" box="[363,433,1941,1967]" captionStart="FIGURE 6. P" captionStartId="10.[151,256,1914,1938]" captionTargetBox="[151,1436,260,1898]" captionTargetId="figure@10.[151,1436,260,1898]" captionTargetPageId="10" captionText="FIGURE 6. P. milleri sp. nov. A – D. Shells. Scale = 1.0 mm. A. Holotype, SBMNH 83651. B – C. Paratypes, SBMNH 74688. D. SBMNH 74687. E. Shell apex, SBMNH 74688. Scale = 100 µm. F. Close-up of protoconch sculpture, SBMNH 74688. Scale = 100 µm. G – I. Opercula, USNM 90527. Scale = 500 µm. G. Outer side. H – I. Inner side, showing variation in thickening of attachment scar margin." httpUri="https://zenodo.org/record/193339/files/figure.png" pageId="9" pageNumber="10">Fig. 6</figureCitation>
|
||
A), SBMNH 83651, creek
|
||
<quantity id="4CEE79AD3E068A0DFD476595EA79F86B" box="[749,845,1941,1967]" metricMagnitude="3" metricUnit="m" metricValue="1.1265408" pageId="9" pageNumber="10" unit="mi" value="0.7">0.7 mile</quantity>
|
||
(
|
||
<quantity id="4CEE79AD3E068A0DFCF76595EAF4F86B" box="[861,960,1941,1967]" metricMagnitude="3" metricUnit="m" metricValue="1.13" pageId="9" pageNumber="10" unit="km" value="1.13">1.13 km</quantity>
|
||
) east of Pierpoint Spring on (California) Hwy 190, approximately
|
||
<quantity id="4CEE79AD3E068A0DFE7F65BAEB63F810" box="[469,599,1978,2004]" metricMagnitude="4" metricUnit="m" metricValue="2.51057664" pageId="9" pageNumber="10" unit="mi" value="15.6">15.6 miles</quantity>
|
||
(
|
||
<quantity id="4CEE79AD3E068A0DFDC365BAEBE6F810" box="[617,722,1978,2004]" metricMagnitude="4" metricUnit="m" metricValue="2.51" pageId="9" pageNumber="10" unit="km" value="25.1">25.1 km</quantity>
|
||
) east of Springville, Tule River drainage, Tulare County, California, Walter B. Miller,
|
||
<date id="FFA8F2883E058A0EFE436298EB59FF76" box="[489,621,152,178]" pageId="10" pageNumber="11" value="1964-08-01">1/viii/1964</date>
|
||
.
|
||
<typeStatus id="54AD6AEA3E058A0EFDD36298EBD9FF76" box="[633,749,152,178]" pageId="10" pageNumber="11" type="paratype">Paratypes</typeStatus>
|
||
, SBMNH 74688, USNM 1132568 (from same lot, 211 + 6 dry shells, respectively).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF6984C03E058A0EFF3D657AEB98F835" httpUri="https://zenodo.org/record/193339/files/figure.png" pageId="10" pageNumber="11" targetBox="[151,1436,260,1898]" targetPageId="10">
|
||
<paragraph id="8BA9D4483E058A0EFF3D657AEB98F835" blockId="10.[151,1436,1914,2033]" pageId="10" pageNumber="11">
|
||
<emphasis id="B962085A3E058A0EFF3D657AE820F856" bold="true" box="[151,276,1914,1938]" pageId="10" pageNumber="11">FIGURE 6</emphasis>
|
||
.
|
||
<taxonomicName id="4C16AFCB3E058A0EFE89657AE8B2F855" box="[291,390,1914,1937]" class="Gastropoda" family="Hydrobiidae" genus="Pyrgulopsis" kingdom="Animalia" order="Littorinimorpha" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="milleri" status="sp. nov.">
|
||
<emphasis id="B962085A3E058A0EFE89657AE8B2F855" box="[291,390,1914,1937]" italics="true" pageId="10" pageNumber="11">P. milleri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A251B5213E058A0EFE27657AE8D1F856" box="[397,485,1914,1938]" pageId="10" pageNumber="11" rank="species">
|
||
<emphasis id="B962085A3E058A0EFE27657AE8D1F856" bold="true" box="[397,485,1914,1938]" pageId="10" pageNumber="11">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
A–D. Shells. Scale = 1.0 mm. A. Holotype, SBMNH 83651. B–C. Paratypes, SBMNH 74688. D. SBMNH 74687. E. Shell apex, SBMNH 74688. Scale = 100 µm. F. Close-up of protoconch sculpture, SBMNH 74688. Scale = 100 µm. G–I. Opercula, USNM 90527. Scale = 500 µm. G. Outer side. H–I. Inner side, showing variation in thickening of attachment scar margin.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF6984C03E048A0FFF3D64F3EAE7F8EF" httpUri="https://zenodo.org/record/193340/files/figure.png" pageId="11" pageNumber="12" targetBox="[287,1300,194,1758]" targetPageId="11">
|
||
<paragraph id="8BA9D4483E048A0FFF3D64F3EAE7F8EF" blockId="11.[151,1436,1779,1835]" pageId="11" pageNumber="12">
|
||
<emphasis id="B962085A3E048A0FFF3D64F3E827F8CF" bold="true" box="[151,275,1779,1803]" pageId="11" pageNumber="12">FIGURE 7</emphasis>
|
||
.
|
||
<taxonomicName id="4C16AFCB3E048A0FFE8B64F4E8B7F8CF" box="[289,387,1780,1803]" class="Gastropoda" family="Hydrobiidae" genus="Pyrgulopsis" kingdom="Animalia" order="Littorinimorpha" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="milleri" status="sp. nov.">
|
||
<emphasis id="B962085A3E048A0FFE8B64F4E8B7F8CF" box="[289,387,1780,1803]" italics="true" pageId="11" pageNumber="12">P. milleri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A251B5213E048A0FFE2164F3E8D5F8CF" box="[395,481,1779,1803]" pageId="11" pageNumber="12" rank="species">
|
||
<emphasis id="B962085A3E048A0FFE2164F3E8D5F8CF" bold="true" box="[395,481,1779,1803]" pageId="11" pageNumber="12">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, USNM 90527, scales = 10 µm. A. Portion of radula ribbon. B. Central radular teeth. C. Lateral tooth. D. Inner marginal teeth. E–F. Outer marginal teeth. Wg = wing.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C30C87C33E048A0FFF6E6556EC29F852" pageId="11" pageNumber="12" type="etymology">
|
||
<paragraph id="8BA9D4483E048A0FFF6E6556EC29F852" blockId="11.[151,1436,1878,2017]" pageId="11" pageNumber="12">
|
||
<emphasis id="B962085A3E048A0FFF6E6556E863F8B4" bold="true" box="[196,343,1878,1904]" pageId="11" pageNumber="12">Etymology.</emphasis>
|
||
In honor of deceased zoologist Walter B. Miller, an intrepid collector of western North American nonmarine mollusks (
|
||
<bibRefCitation id="EF87A9B93E048A0FFDA5657CEA12F852" author="Hochberg" box="[527,806,1916,1942]" pageId="11" pageNumber="16" refString="Hochberg, F. G. & Roth, B. (2001) Walter B. Miller (1918 - 2000): molluscan taxa and bibliography. The Veliger, 44: 95 - 98." type="journal article" year="2001">Hochberg & Roth 2001</bibRefCitation>
|
||
) who first discovered this novelty in 1964.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30C87C33E048A08FF6E65A1EAD5FF13" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12" type="materials_examined">
|
||
<paragraph id="8BA9D4483E048A08FF6E65A1EAD5FF13" blockId="11.[151,1436,1878,2017]" lastBlockId="12.[151,1436,152,290]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
|
||
<emphasis id="B962085A3E048A0FFF6E65A1E884F87F" bold="true" box="[196,432,1953,1979]" pageId="11" pageNumber="12">Referred material.</emphasis>
|
||
CALIFORNIA. Tulare County:
|
||
<collectionCode id="ED074C8D3E048A0FFC9365A1EA96F87F" LSID="urn:lsid:biocol.org:col:34834" box="[825,930,1953,1979]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34834" name="Santa Barbara Museum of Natural History" pageId="11" pageNumber="12">SBMNH</collectionCode>
|
||
74687, Pierpoint Spring,
|
||
<quantity id="4CEE79AD3E048A0FFB7365A1EC7CF87F" box="[1241,1352,1953,1979]" metricMagnitude="4" metricUnit="m" metricValue="1.0460736" pageId="11" pageNumber="12" unit="mi" value="6.5">6.5 miles</quantity>
|
||
(
|
||
<quantity id="4CEE79AD3E048A0FFAFD65A1E98AF825" metricMagnitude="4" metricUnit="m" metricValue="1.046" pageId="11" pageNumber="12" unit="km" value="10.46">10.46 km</quantity>
|
||
) east of Tule River power house, 353028 E, 4000829 N (approximate coordinates), Walter B. Miller,
|
||
<date id="FFA8F2883E048A08FADC65C7E9C8FF76" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12" value="1964-07-31">31/ vii/1964</date>
|
||
.
|
||
<collectionCode id="ED074C8D3E038A08FEA26298E869FF76" LSID="urn:lsid:biocol.org:col:34871" box="[264,349,152,178]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="12" pageNumber="13">USNM</collectionCode>
|
||
905257,
|
||
<emphasis id="B962085A3E038A08FE676298E8CFFF75" box="[461,507,152,177]" italics="true" pageId="12" pageNumber="13">ibid</emphasis>
|
||
., Robert Hershler,
|
||
<date id="FFA8F2883E038A08FD776298EA54FF76" box="[733,864,152,178]" pageId="12" pageNumber="13" value="2000-06-26">26/vi/2000</date>
|
||
.
|
||
<collectionCode id="ED074C8D3E038A08FCC66298EAE1FF76" LSID="urn:lsid:biocol.org:col:34834" box="[876,981,152,178]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34834" name="Santa Barbara Museum of Natural History" pageId="12" pageNumber="13">SBMNH</collectionCode>
|
||
74540, spring along Hwy 190
|
||
<quantity id="4CEE79AD3E038A08FAEC6298ECA8FF76" box="[1350,1436,152,178]" metricMagnitude="4" metricUnit="m" metricValue="1.1265408" pageId="12" pageNumber="13" unit="mi" value="7.0">7 miles</quantity>
|
||
(
|
||
<quantity id="4CEE79AD3E038A08FF3562BDE824FF13" box="[159,272,189,215]" metricMagnitude="4" metricUnit="m" metricValue="1.123" pageId="12" pageNumber="13" unit="km" value="11.23">11.23 km</quantity>
|
||
) east of Tule hydroelectric plant, Walter B. Miller,
|
||
<date id="FFA8F2883E038A08FCC362BDEAE8FF13" box="[873,988,189,215]" pageId="12" pageNumber="13" value="1963-01-25">25/i/1963</date>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30C87C33E038A08FF6E62E2ECB6FEE6" pageId="12" pageNumber="13" type="diagnosis">
|
||
<paragraph id="8BA9D4483E038A08FF6E62E2ECB6FEE6" blockId="12.[151,1436,152,290]" pageId="12" pageNumber="13">
|
||
<emphasis id="B962085A3E038A08FF6E62E2E87DFF38" bold="true" box="[196,329,226,252]" pageId="12" pageNumber="13">Diagnosis.</emphasis>
|
||
A medium-sized species having ovate- to narrow-conic shell with weakly convex whorls. Penis usually alobate; filament medium length; penial ornament usually consisting of a small ventral gland.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF6984C03E038A08FF3D64A8ED3CF8A4" httpUri="https://zenodo.org/record/193341/files/figure.png" pageId="12" pageNumber="13" targetBox="[245,1339,350,1703]" targetPageId="12">
|
||
<paragraph id="8BA9D4483E038A08FF3D64A8ED3CF8A4" blockId="12.[151,1437,1704,1888]" pageId="12" pageNumber="13">
|
||
<emphasis id="B962085A3E038A08FF3D64A8E820F904" bold="true" box="[151,276,1704,1728]" pageId="12" pageNumber="13">FIGURE 8</emphasis>
|
||
.
|
||
<taxonomicName id="4C16AFCB3E038A08FE8964A9E8BCF904" box="[291,392,1705,1728]" class="Gastropoda" family="Hydrobiidae" genus="Pyrgulopsis" kingdom="Animalia" order="Littorinimorpha" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="milleri" status="sp. nov.">
|
||
<emphasis id="B962085A3E038A08FE8964A9E8BCF904" box="[291,392,1705,1728]" italics="true" pageId="12" pageNumber="13">P. milleri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A251B5213E038A08FE2564A8E8D3F904" box="[399,487,1704,1728]" pageId="12" pageNumber="13" rank="species">
|
||
<emphasis id="B962085A3E038A08FE2564A8E8D3F904" bold="true" box="[399,487,1704,1728]" pageId="12" pageNumber="13">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, USNM 90527. Scale = 0.5 mm. A–C. Penis. A. Dorsal surface. B–C. Ventral surface, showing ventral and terminal glands. D. Female glandular oviduct and associated structures (viewed from the left side). E. Bursa copulatrix and its duct. F. Seminal receptacle and its duct. Ag = albumen gland, Bu = bursa copulatrix, Cd = common duct of seminal receptacle and coiled oviduct, Cg = capsule gland, Co = coiled oviduct, Dbu = bursal duct, Ga =genital aperture, Pd = penial duct, Pf = penial filament, Pl = penial lobe, Pw = posterior wall of pallial cavity, Sr = seminal receptacle, Tg = terminal gland, Vc = ventral channel, Vg = ventral gland.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C30C87C33E038A09FF6E658EEAF3F9A6" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13" type="description">
|
||
<paragraph id="8BA9D4483E038A09FF6E658EEC20FF38" blockId="12.[151,1437,1934,2035]" lastBlockId="13.[151,1437,152,2007]" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
|
||
<emphasis id="B962085A3E038A08FF6E658EE856F86C" bold="true" box="[196,354,1934,1960]" pageId="12" pageNumber="13">Description.</emphasis>
|
||
Shell (
|
||
<figureCitation id="132DC8CD3E038A08FE11658EEB37F86C" box="[443,515,1934,1960]" captionStart="FIGURE 6. P" captionStartId="10.[151,256,1914,1938]" captionTargetBox="[151,1436,260,1898]" captionTargetId="figure@10.[151,1436,260,1898]" captionTargetPageId="10" captionText="FIGURE 6. P. milleri sp. nov. A – D. Shells. Scale = 1.0 mm. A. Holotype, SBMNH 83651. B – C. Paratypes, SBMNH 74688. D. SBMNH 74687. E. Shell apex, SBMNH 74688. Scale = 100 µm. F. Close-up of protoconch sculpture, SBMNH 74688. Scale = 100 µm. G – I. Opercula, USNM 90527. Scale = 500 µm. G. Outer side. H – I. Inner side, showing variation in thickening of attachment scar margin." httpUri="https://zenodo.org/record/193339/files/figure.png" pageId="12" pageNumber="13">Fig. 6</figureCitation>
|
||
A–D) ovate- or narrow-conic, height about 2.5–4.0 mm; whorls 4.25–5.25. Periostracum tan, thin, often covered with dark deposits. Protoconch (
|
||
<figureCitation id="132DC8CD3E038A08FC5465B3ED7CF809" box="[1022,1096,1971,1997]" captionStart="FIGURE 6. P" captionStartId="10.[151,256,1914,1938]" captionTargetBox="[151,1436,260,1898]" captionTargetId="figure@10.[151,1436,260,1898]" captionTargetPageId="10" captionText="FIGURE 6. P. milleri sp. nov. A – D. Shells. Scale = 1.0 mm. A. Holotype, SBMNH 83651. B – C. Paratypes, SBMNH 74688. D. SBMNH 74687. E. Shell apex, SBMNH 74688. Scale = 100 µm. F. Close-up of protoconch sculpture, SBMNH 74688. Scale = 100 µm. G – I. Opercula, USNM 90527. Scale = 500 µm. G. Outer side. H – I. Inner side, showing variation in thickening of attachment scar margin." httpUri="https://zenodo.org/record/193339/files/figure.png" pageId="12" pageNumber="13">Fig. 6</figureCitation>
|
||
E) near planispiral, 1.3–1.5 whorls, diameter about 380 µm, surface weakly wrinkled near apex (
|
||
<figureCitation id="132DC8CD3E038A08FC6F65D9ED38F837" box="[965,1036,2009,2035]" captionStart="FIGURE 6. P" captionStartId="10.[151,256,1914,1938]" captionTargetBox="[151,1436,260,1898]" captionTargetId="figure@10.[151,1436,260,1898]" captionTargetPageId="10" captionText="FIGURE 6. P. milleri sp. nov. A – D. Shells. Scale = 1.0 mm. A. Holotype, SBMNH 83651. B – C. Paratypes, SBMNH 74688. D. SBMNH 74687. E. Shell apex, SBMNH 74688. Scale = 100 µm. F. Close-up of protoconch sculpture, SBMNH 74688. Scale = 100 µm. G – I. Opercula, USNM 90527. Scale = 500 µm. G. Outer side. H – I. Inner side, showing variation in thickening of attachment scar margin." httpUri="https://zenodo.org/record/193339/files/figure.png" pageId="12" pageNumber="13">Fig. 6</figureCitation>
|
||
F), otherwise smooth. Teleoconch whorls weakly convex, sometimes narrowly shouldered, last 0.25 whorl rarely slightly loosened; sculpture of collabral growth lines. Aperture ovate, angled adapically. Inner lip complete, usually adnate, slightly thickened internally; columellar shelf absent; outer lip thin, weakly prosocline. Umbilicus narrow.
|
||
</paragraph>
|
||
<paragraph id="8BA9D4483E028A09FF6E6308E8A7FD88" blockId="13.[151,1437,152,2007]" pageId="13" pageNumber="14">
|
||
Operculum (
|
||
<figureCitation id="132DC8CD3E028A09FEF06308E895FEE6" box="[346,417,264,290]" captionStart="FIGURE 6. P" captionStartId="10.[151,256,1914,1938]" captionTargetBox="[151,1436,260,1898]" captionTargetId="figure@10.[151,1436,260,1898]" captionTargetPageId="10" captionText="FIGURE 6. P. milleri sp. nov. A – D. Shells. Scale = 1.0 mm. A. Holotype, SBMNH 83651. B – C. Paratypes, SBMNH 74688. D. SBMNH 74687. E. Shell apex, SBMNH 74688. Scale = 100 µm. F. Close-up of protoconch sculpture, SBMNH 74688. Scale = 100 µm. G – I. Opercula, USNM 90527. Scale = 500 µm. G. Outer side. H – I. Inner side, showing variation in thickening of attachment scar margin." httpUri="https://zenodo.org/record/193339/files/figure.png" pageId="13" pageNumber="14">Fig. 6</figureCitation>
|
||
G) thin, flat, light amber, multispiral with eccentric nucleus; last quarter whorl frilled on outer side; inner side (
|
||
<figureCitation id="132DC8CD3E028A09FE78632DEB28FE83" box="[466,540,301,327]" captionStart="FIGURE 6. P" captionStartId="10.[151,256,1914,1938]" captionTargetBox="[151,1436,260,1898]" captionTargetId="figure@10.[151,1436,260,1898]" captionTargetPageId="10" captionText="FIGURE 6. P. milleri sp. nov. A – D. Shells. Scale = 1.0 mm. A. Holotype, SBMNH 83651. B – C. Paratypes, SBMNH 74688. D. SBMNH 74687. E. Shell apex, SBMNH 74688. Scale = 100 µm. F. Close-up of protoconch sculpture, SBMNH 74688. Scale = 100 µm. G – I. Opercula, USNM 90527. Scale = 500 µm. G. Outer side. H – I. Inner side, showing variation in thickening of attachment scar margin." httpUri="https://zenodo.org/record/193339/files/figure.png" pageId="13" pageNumber="14">Fig. 6</figureCitation>
|
||
H–I) having attachment scar border slightly thickened almost all around. Radula taenioglossate (
|
||
<figureCitation id="132DC8CD3E028A09FE026352E8DAFEA8" box="[424,494,338,364]" captionStart="FIGURE 7. P" captionStartId="11.[151,255,1779,1803]" captionTargetBox="[287,1300,194,1758]" captionTargetId="figure@11.[287,1301,194,1758]" captionTargetPageId="11" captionText="FIGURE 7. P. milleri sp. nov., USNM 90527, scales = 10 µm. A. Portion of radula ribbon. B. Central radular teeth. C. Lateral tooth. D. Inner marginal teeth. E – F. Outer marginal teeth. Wg = wing." httpUri="https://zenodo.org/record/193340/files/figure.png" pageId="13" pageNumber="14">Fig. 7</figureCitation>
|
||
A), having about 55 well-formed rows of teeth. Central teeth (
|
||
<figureCitation id="132DC8CD3E028A09FB616352EC25FEA8" box="[1227,1297,338,364]" captionStart="FIGURE 7. P" captionStartId="11.[151,255,1779,1803]" captionTargetBox="[287,1300,194,1758]" captionTargetId="figure@11.[287,1301,194,1758]" captionTargetPageId="11" captionText="FIGURE 7. P. milleri sp. nov., USNM 90527, scales = 10 µm. A. Portion of radula ribbon. B. Central radular teeth. C. Lateral tooth. D. Inner marginal teeth. E – F. Outer marginal teeth. Wg = wing." httpUri="https://zenodo.org/record/193340/files/figure.png" pageId="13" pageNumber="14">Fig. 7</figureCitation>
|
||
B) about 28 µm wide, cutting edge concave; lateral cusps 3–6; central cusp large, U-shaped; basal cusp 1, small; basal tongue V-shaped, about as long as lateral margins. Lateral tooth (
|
||
<figureCitation id="132DC8CD3E028A09FC01639DEAC7FE73" box="[939,1011,413,439]" captionStart="FIGURE 7. P" captionStartId="11.[151,255,1779,1803]" captionTargetBox="[287,1300,194,1758]" captionTargetId="figure@11.[287,1301,194,1758]" captionTargetPageId="11" captionText="FIGURE 7. P. milleri sp. nov., USNM 90527, scales = 10 µm. A. Portion of radula ribbon. B. Central radular teeth. C. Lateral tooth. D. Inner marginal teeth. E – F. Outer marginal teeth. Wg = wing." httpUri="https://zenodo.org/record/193340/files/figure.png" pageId="13" pageNumber="14">Fig. 7</figureCitation>
|
||
C) face rectangular, angled; central cusp large, U-shaped; lateral cusps 2–4 (inner), 3–5 (outer); outer wing medium width, straight, about 200% length of cutting edge; basal tongue well developed. Inner marginal teeth (
|
||
<figureCitation id="132DC8CD3E028A09FC5363E8ED0AFDC6" box="[1017,1086,488,514]" captionStart="FIGURE 7. P" captionStartId="11.[151,255,1779,1803]" captionTargetBox="[287,1300,194,1758]" captionTargetId="figure@11.[287,1301,194,1758]" captionTargetPageId="11" captionText="FIGURE 7. P. milleri sp. nov., USNM 90527, scales = 10 µm. A. Portion of radula ribbon. B. Central radular teeth. C. Lateral tooth. D. Inner marginal teeth. E – F. Outer marginal teeth. Wg = wing." httpUri="https://zenodo.org/record/193340/files/figure.png" pageId="13" pageNumber="14">Fig. 7</figureCitation>
|
||
D) having 17–21 cusps. Outer marginal teeth (
|
||
<figureCitation id="132DC8CD3E028A09FEFB600DE8A3FDE3" box="[337,407,525,551]" captionStart="FIGURE 7. P" captionStartId="11.[151,255,1779,1803]" captionTargetBox="[287,1300,194,1758]" captionTargetId="figure@11.[287,1301,194,1758]" captionTargetPageId="11" captionText="FIGURE 7. P. milleri sp. nov., USNM 90527, scales = 10 µm. A. Portion of radula ribbon. B. Central radular teeth. C. Lateral tooth. D. Inner marginal teeth. E – F. Outer marginal teeth. Wg = wing." httpUri="https://zenodo.org/record/193340/files/figure.png" pageId="13" pageNumber="14">Fig. 7</figureCitation>
|
||
E–F) having 19–28 cusps; inner edge having long, rectangular wing. Radula data were from USNM 905257.
|
||
</paragraph>
|
||
<paragraph id="8BA9D4483E028A09FF6E6058EB52FB66" blockId="13.[151,1437,152,2007]" pageId="13" pageNumber="14">
|
||
Head-foot generally pale. Snout usually pale, but sometimes light to dark brown. Pallial roof, visceral coil dark brown or black. Ctenidium well developed, positioned a little in front of pericardium; ctenidial filaments about 20, broadly triangular. Osphradium narrow, positioned slightly posterior to middle of ctenidium. Prostate gland large, bean-shaped, with about 40% of length in pallial roof. Anterior vas deferens opening from ventral edge of prostate gland a little in front of posterior pallial wall, section of duct on columellar muscle straight. Penis (
|
||
<figureCitation id="132DC8CD3E028A09FE1F6112E8C8FCE8" box="[437,508,786,812]" captionStart="FIGURE 8. P" captionStartId="12.[151,256,1704,1728]" captionTargetBox="[245,1339,350,1703]" captionTargetId="figure@12.[228,1360,320,1705]" captionTargetPageId="12" captionText="FIGURE 8. P. milleri sp. nov., USNM 90527. Scale = 0.5 mm. A – C. Penis. A. Dorsal surface. B – C. Ventral surface, showing ventral and terminal glands. D. Female glandular oviduct and associated structures (viewed from the left side). E. Bursa copulatrix and its duct. F. Seminal receptacle and its duct. Ag = albumen gland, Bu = bursa copulatrix, Cd = common duct of seminal receptacle and coiled oviduct, Cg = capsule gland, Co = coiled oviduct, Dbu = bursal duct, Ga = genital aperture, Pd = penial duct, Pf = penial filament, Pl = penial lobe, Pw = posterior wall of pallial cavity, Sr = seminal receptacle, Tg = terminal gland, Vc = ventral channel, Vg = ventral gland." httpUri="https://zenodo.org/record/193341/files/figure.png" pageId="13" pageNumber="14">Fig. 8</figureCitation>
|
||
A–C) medium-sized, base rectangular, inner edge smooth; filament short or medium length, tapering, slightly oblique; lobe usually absent, nub-like when present (
|
||
<figureCitation id="132DC8CD3E028A09FB136138EC35FC96" box="[1209,1281,824,850]" captionStart="FIGURE 8. P" captionStartId="12.[151,256,1704,1728]" captionTargetBox="[245,1339,350,1703]" captionTargetId="figure@12.[228,1360,320,1705]" captionTargetPageId="12" captionText="FIGURE 8. P. milleri sp. nov., USNM 90527. Scale = 0.5 mm. A – C. Penis. A. Dorsal surface. B – C. Ventral surface, showing ventral and terminal glands. D. Female glandular oviduct and associated structures (viewed from the left side). E. Bursa copulatrix and its duct. F. Seminal receptacle and its duct. Ag = albumen gland, Bu = bursa copulatrix, Cd = common duct of seminal receptacle and coiled oviduct, Cg = capsule gland, Co = coiled oviduct, Dbu = bursal duct, Ga = genital aperture, Pd = penial duct, Pf = penial filament, Pl = penial lobe, Pw = posterior wall of pallial cavity, Sr = seminal receptacle, Tg = terminal gland, Vc = ventral channel, Vg = ventral gland." httpUri="https://zenodo.org/record/193341/files/figure.png" pageId="13" pageNumber="14">Fig. 8</figureCitation>
|
||
C). Terminal gland (observed in one specimen) small, ovate, positioned on ventral surface of lobe. Ventral gland small, ovate, centrally positioned, borne on raised swelling. Penial duct narrow, nearly straight. Penis entirely pale or with filament variably pigmented by black granules. Female glandular oviduct and associated structures shown in
|
||
<figureCitation id="132DC8CD3E028A09FEA661CDE85BFC23" box="[268,367,973,999]" captionStart="FIGURE 8. P" captionStartId="12.[151,256,1704,1728]" captionTargetBox="[245,1339,350,1703]" captionTargetId="figure@12.[228,1360,320,1705]" captionTargetPageId="12" captionText="FIGURE 8. P. milleri sp. nov., USNM 90527. Scale = 0.5 mm. A – C. Penis. A. Dorsal surface. B – C. Ventral surface, showing ventral and terminal glands. D. Female glandular oviduct and associated structures (viewed from the left side). E. Bursa copulatrix and its duct. F. Seminal receptacle and its duct. Ag = albumen gland, Bu = bursa copulatrix, Cd = common duct of seminal receptacle and coiled oviduct, Cg = capsule gland, Co = coiled oviduct, Dbu = bursal duct, Ga = genital aperture, Pd = penial duct, Pf = penial filament, Pl = penial lobe, Pw = posterior wall of pallial cavity, Sr = seminal receptacle, Tg = terminal gland, Vc = ventral channel, Vg = ventral gland." httpUri="https://zenodo.org/record/193341/files/figure.png" pageId="13" pageNumber="14">Figure 8</figureCitation>
|
||
D–F. Coiled oviduct a small, proximally kinked, posterior-oblique loop. Bursa copulatrix small, narrowly ovate, horizontal, about 50% overlapped by albumen gland. Bursal duct about as long as bursa, medium width, opening from distal edge, partly embedded in albumen gland. Seminal receptacle small, pouch-shaped, positioned along ventral edge of proximal bursal duct; duct short. Albumen gland a little shorter than capsule gland, having very short pallial section. Capsule gland composed of a single tissue section. Genital aperture a terminal slit.
|
||
</paragraph>
|
||
<paragraph id="8BA9D4483E028A09FF6E66ADEA56FA36" blockId="13.[151,1437,152,2007]" pageId="13" pageNumber="14">
|
||
<emphasis id="B962085A3E028A09FF6E66ADE8FCFB03" bold="true" box="[196,456,1197,1223]" pageId="13" pageNumber="14">Shell measurements</emphasis>
|
||
(mean + standard deviation in parentheses): height 3.00–4.00 mm (3.42+0.35), width 2.00–
|
||
<quantity id="4CEE79AD3E028A09FE8966D2E8BFFB28" box="[291,395,1234,1260]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="13" pageNumber="14" unit="mm" value="2.5">2.50 mm</quantity>
|
||
(2.17+0.12), body whorl height
|
||
<quantity id="4CEE79AD3E028A09FCAC66D2EA84FB28" box="[774,944,1234,1260]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="2.75" metricValueMin="2.25" pageId="13" pageNumber="14" unit="mm" value="2.5" valueMax="2.75" valueMin="2.25">2.25–2.75 mm</quantity>
|
||
(2.45+0.15), body whorl width
|
||
<quantity id="4CEE79AD3E028A09FA8C66D2E9F1FAD6" metricMagnitude="-3" metricUnit="m" metricValue="1.9449999999999998" metricValueMax="2.19" metricValueMin="1.7" pageId="13" pageNumber="14" unit="mm" value="1.9449999999999998" valueMax="2.19" valueMin="1.7">1.70–2.19 mm</quantity>
|
||
(1.87+0.13), aperture height
|
||
<quantity id="4CEE79AD3E028A09FDB766F8EBF3FAD6" box="[541,711,1272,1298]" metricMagnitude="-3" metricUnit="m" metricValue="1.485" metricValueMax="1.61" metricValueMin="1.36" pageId="13" pageNumber="14" unit="mm" value="1.485" valueMax="1.61" valueMin="1.36">1.36–1.61 mm</quantity>
|
||
(1.46+0.07), aperture width
|
||
<quantity id="4CEE79AD3E028A09FBB366F8EDF0FAD6" box="[1049,1220,1272,1298]" metricMagnitude="-3" metricUnit="m" metricValue="1.2799999999999998" metricValueMax="1.41" metricValueMin="1.15" pageId="13" pageNumber="14" unit="mm" value="1.2799999999999998" valueMax="1.41" valueMin="1.15">1.15–1.41 mm</quantity>
|
||
(1.25+0.06), shell width/height 0.58–0.69 (0.64+0.03), body whorl height/shell height 0.67–0.76 (0.72+0.02), aperture height/ shell height 0.39–0.47 (0.43+0.02) (
|
||
<typeStatus id="54AD6AEA3E028A09FDEF6742EB88FA98" box="[581,700,1346,1372]" pageId="13" pageNumber="14" type="paratype">paratypes</typeStatus>
|
||
, SBMNH 74688,
|
||
<emphasis id="B962085A3E028A09FC396743EA96FA98" box="[915,930,1347,1372]" italics="true" pageId="13" pageNumber="14">n</emphasis>
|
||
= 30). Height
|
||
<quantity id="4CEE79AD3E028A09FBF96742EC34FA98" box="[1107,1280,1346,1372]" metricMagnitude="-3" metricUnit="m" metricValue="3.185" metricValueMax="3.46" metricValueMin="2.91" pageId="13" pageNumber="14" unit="mm" value="3.185" valueMax="3.46" valueMin="2.91">2.91–3.46 mm</quantity>
|
||
(3.12+0.14), width
|
||
<quantity id="4CEE79AD3E028A09FF4A6768E8BFFA46" box="[224,395,1384,1410]" metricMagnitude="-3" metricUnit="m" metricValue="1.965" metricValueMax="2.16" metricValueMin="1.77" pageId="13" pageNumber="14" unit="mm" value="1.965" valueMax="2.16" valueMin="1.77">1.77–2.16 mm</quantity>
|
||
(1.97+0.08), body whorl height
|
||
<quantity id="4CEE79AD3E028A09FCAC6768EA84FA46" box="[774,944,1384,1410]" metricMagnitude="-3" metricUnit="m" metricValue="2.335" metricValueMax="2.57" metricValueMin="2.1" pageId="13" pageNumber="14" unit="mm" value="2.335" valueMax="2.57" valueMin="2.1">2.10–2.57 mm</quantity>
|
||
(2.30+0.10), body whorl width
|
||
<quantity id="4CEE79AD3E028A09FA8C6768E9F1FA63" metricMagnitude="-3" metricUnit="m" metricValue="1.66" metricValueMax="1.8" metricValueMin="1.52" pageId="13" pageNumber="14" unit="mm" value="1.66" valueMax="1.8" valueMin="1.52">1.52–1.80 mm</quantity>
|
||
(1.65+0.07), aperture height
|
||
<quantity id="4CEE79AD3E028A09FDB7678DEBF3FA63" box="[541,711,1421,1447]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" metricValueMax="1.57" metricValueMin="1.23" pageId="13" pageNumber="14" unit="mm" value="1.4" valueMax="1.57" valueMin="1.23">1.23–1.57 mm</quantity>
|
||
(1.39+0.07), aperture width
|
||
<quantity id="4CEE79AD3E028A09FBB3678DEDF0FA63" box="[1049,1220,1421,1447]" metricMagnitude="-3" metricUnit="m" metricValue="1.195" metricValueMax="1.31" metricValueMin="1.08" pageId="13" pageNumber="14" unit="mm" value="1.195" valueMax="1.31" valueMin="1.08">1.08–1.31 mm</quantity>
|
||
(1.16+0.06), shell width/height 0.57–0.66 (0.63+0.02), body whorl height/shell height 0.68–0.77 (0.74+0.02), aperture height/ shell height 0.39–0.48 (0.45+0.02) (SBMNH 74687,
|
||
<emphasis id="B962085A3E028A09FCAF67D8EA20FA35" box="[773,788,1496,1521]" italics="true" pageId="13" pageNumber="14">n</emphasis>
|
||
= 30).
|
||
</paragraph>
|
||
<paragraph id="8BA9D4483E028A09FF6E67FDEAF3F9A6" blockId="13.[151,1437,152,2007]" pageId="13" pageNumber="14">
|
||
<emphasis id="B962085A3E028A09FF6E67FDEB2FF9D3" bold="true" box="[196,539,1533,1559]" pageId="13" pageNumber="14">
|
||
Measurements of
|
||
<typeStatus id="54AD6AEA3E028A09FE0F67FDEB22F9D3" box="[421,534,1533,1559]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
height,
|
||
<quantity id="4CEE79AD3E028A09FDD667FDEBDDF9D3" box="[636,745,1533,1559]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="13" pageNumber="14" unit="mm" value="3.7">3.70 mm</quantity>
|
||
, width
|
||
<quantity id="4CEE79AD3E028A09FCEA67FDEA99F9D3" box="[832,941,1533,1559]" metricMagnitude="-3" metricUnit="m" metricValue="2.36" pageId="13" pageNumber="14" unit="mm" value="2.36">2.36 mm</quantity>
|
||
, body whorl height
|
||
<quantity id="4CEE79AD3E028A09FB3167FDEC3CF9D3" box="[1179,1288,1533,1559]" metricMagnitude="-3" metricUnit="m" metricValue="2.67" pageId="13" pageNumber="14" unit="mm" value="2.67">2.67 mm</quantity>
|
||
, body whorl width
|
||
<quantity id="4CEE79AD3E028A09FF4F6422E862F9F8" box="[229,342,1570,1596]" metricMagnitude="-3" metricUnit="m" metricValue="2.01" pageId="13" pageNumber="14" unit="mm" value="2.01">2.01 mm</quantity>
|
||
, aperture height
|
||
<quantity id="4CEE79AD3E028A09FD8D6422EBACF9F8" box="[551,664,1570,1596]" metricMagnitude="-3" metricUnit="m" metricValue="1.59" pageId="13" pageNumber="14" unit="mm" value="1.59">1.59 mm</quantity>
|
||
, aperture width
|
||
<quantity id="4CEE79AD3E028A09FCC86422EAE7F9F8" box="[866,979,1570,1596]" metricMagnitude="-3" metricUnit="m" metricValue="1.39" pageId="13" pageNumber="14" unit="mm" value="1.39">1.39 mm</quantity>
|
||
, shell width/height 0.64, body whorl height/shell height 0.72, aperture height/shell height 0.43, 5.0 whorls.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30C87C33E028A0AFF6E646DED18FF13" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14" type="distribution">
|
||
<paragraph id="8BA9D4483E028A09FF6E646DED9CF8A3" blockId="13.[151,1437,152,2007]" pageId="13" pageNumber="14">
|
||
<emphasis id="B962085A3E028A09FF6E646DEA34F943" bold="true" box="[196,768,1645,1671]" pageId="13" pageNumber="14">Distribution, habitat and conservation status.</emphasis>
|
||
<taxonomicName id="4C16AFCB3E028A09FCA2646DEADFF942" box="[776,1003,1645,1670]" class="Gastropoda" family="Hydrobiidae" genus="Pyrgulopsis" kingdom="Animalia" order="Littorinimorpha" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="milleri">
|
||
<emphasis id="B962085A3E028A09FCA2646DEADFF942" box="[776,1003,1645,1670]" italics="true" pageId="13" pageNumber="14">Pyrgulopsis milleri</emphasis>
|
||
</taxonomicName>
|
||
is known only from its
|
||
<typeStatus id="54AD6AEA3E028A09FAA2646DEC0FF943" box="[1288,1339,1645,1671]" pageId="13" pageNumber="14">type</typeStatus>
|
||
locality area, which consists of spring-fed waters along a very short (ca. 1.0 km) reach of the South Fork of the Middle Fork Tule River drainage. Pierpoint Spring spills down a rock face into a ditch right alongside Hwy 190; part of its flow issues from a pipe (see
|
||
<bibRefCitation id="EF87A9B93E028A09FD9564DDEA52F933" author="Jenkins" box="[575,870,1757,1783]" pageId="13" pageNumber="16" refString="Jenkins, J. C. & Jenkins, R. J. (1995) Exploring the Southern Sierra: west side. Wilderness Press, Berkeley, California. 256 pp." type="book" year="1995">Jenkins & Jenkins 1995</bibRefCitation>
|
||
:153 for a photograph of this spring). The first author collected
|
||
<taxonomicName id="4C16AFCB3E028A09FEF76503E8FEF8D8" box="[349,458,1795,1820]" class="Gastropoda" family="Hydrobiidae" genus="Pyrgulopsis" kingdom="Animalia" order="Littorinimorpha" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="milleri">
|
||
<emphasis id="B962085A3E028A09FEF76503E8FEF8D8" box="[349,458,1795,1820]" italics="true" pageId="13" pageNumber="14">P. milleri</emphasis>
|
||
</taxonomicName>
|
||
from aquatic vegetation in this ditch; density was low. There is no information on the habitat and current status of the other two populations which Miller sampled during the early 1960’s. (The first author did not attempt to access these sites during his brief visit to the area in 2000.)
|
||
</paragraph>
|
||
<paragraph id="8BA9D4483E028A0AFF6E6573ED18FF13" blockId="13.[151,1437,152,2007]" lastBlockId="14.[151,1437,152,626]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
|
||
<taxonomicName id="4C16AFCB3E028A09FF6E6573E891F848" box="[196,421,1907,1932]" class="Gastropoda" family="Hydrobiidae" genus="Pyrgulopsis" kingdom="Animalia" order="Littorinimorpha" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="milleri">
|
||
<emphasis id="B962085A3E028A09FF6E6573E891F848" box="[196,421,1907,1932]" italics="true" pageId="13" pageNumber="14">Pyrgulopsis milleri</emphasis>
|
||
</taxonomicName>
|
||
is distributed within a parcel of patented (private) land that is nested within the Giant Sequoia National Monument (Sequoia National Forest). This species currently has no protection and may be threatened by the diversion of Pierpoint Spring, physical disturbance (e.g., trampling, pollution) associated with the frequent use of this spring as a drinking water supply by travelers on CA Highway 190 (
|
||
<bibRefCitation id="EF87A9B93E018A0AFA886298E85CFF13" author="Jenkins" pageId="14" pageNumber="16" refString="Jenkins, J. C. & Jenkins, R. J. (1995) Exploring the Southern Sierra: west side. Wilderness Press, Berkeley, California. 256 pp." type="book" year="1995">Jenkins & Jenkins 1995:153</bibRefCitation>
|
||
), and the planned local widening of this road (CDOT 2009).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30C87C33E018A0AFF6E62E2EA71FDB6" pageId="14" pageNumber="15" type="discussion">
|
||
<paragraph id="8BA9D4483E018A0AFF6E62E2EBE6FDC6" blockId="14.[151,1437,152,626]" pageId="14" pageNumber="15">
|
||
<emphasis id="B962085A3E018A0AFF6E62E2E874FF38" bold="true" box="[196,320,226,252]" pageId="14" pageNumber="15">Remarks.</emphasis>
|
||
<taxonomicName id="4C16AFCB3E018A0AFEE262E2EB05FF3F" box="[328,561,226,251]" class="Gastropoda" family="Hydrobiidae" genus="Pyrgulopsis" kingdom="Animalia" order="Littorinimorpha" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="milleri">
|
||
<emphasis id="B962085A3E018A0AFEE262E2EB05FF3F" box="[328,561,226,251]" italics="true" pageId="14" pageNumber="15">Pyrgulopsis milleri</emphasis>
|
||
</taxonomicName>
|
||
differs from closely similar
|
||
<taxonomicName id="4C16AFCB3E018A0AFC2462E2ED0DFF3F" box="[910,1081,226,251]" class="Gastropoda" family="Hydrobiidae" genus="Pyrgulopsis" kingdom="Animalia" order="Littorinimorpha" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="stearnsiana">
|
||
<emphasis id="B962085A3E018A0AFC2462E2ED0DFF3F" box="[910,1081,226,251]" italics="true" pageId="14" pageNumber="15">P. stearnsiana</emphasis>
|
||
</taxonomicName>
|
||
, which ranges into the lower portion of the Tule River basin (Hershler unpublished), in its broader central cusps on the central radular teeth, shorter pallial section of the albumen gland, greater overlap of the bursa copulatrix by the albumen gland, absence of a bend or loop in the anterior vas deferens, presence of a ventral gland on the penis and absence (except in one specimen) of a terminal gland.
|
||
<emphasis id="B962085A3E018A0AFD1E6378EAA0FE55" box="[692,916,376,401]" italics="true" pageId="14" pageNumber="15">
|
||
Pyrgulospis
|
||
<taxonomicName id="4C16AFCB3E018A0AFCEC6378EAA0FE55" box="[838,916,376,401]" class="Gastropoda" family="Hydrobiidae" genus="Pyrgulopsis" kingdom="Animalia" order="Littorinimorpha" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="milleri">milleri</taxonomicName>
|
||
</emphasis>
|
||
was most similar to
|
||
<taxonomicName id="4C16AFCB3E018A0AFB206378EC1BFE55" box="[1162,1327,376,401]" class="Gastropoda" family="Hydrobiidae" genus="Pyrgulopsis" kingdom="Animalia" order="Littorinimorpha" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="stearnsiana">
|
||
<emphasis id="B962085A3E018A0AFB206378EC1BFE55" box="[1162,1327,376,401]" italics="true" pageId="14" pageNumber="15">P. stearnsiana</emphasis>
|
||
</taxonomicName>
|
||
from the Santa Clara River basin (21AA) in its mtCOI sequences (2.8% divergence) and differed from other specimens of this species by 3.3–8.4%. Thus, as with
|
||
<taxonomicName id="4C16AFCB3E018A0AFD3863C2EA09FE1F" box="[658,829,450,475]" class="Gastropoda" family="Hydrobiidae" genus="Pyrgulopsis" kingdom="Animalia" order="Littorinimorpha" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="castaicensis">
|
||
<emphasis id="B962085A3E018A0AFD3863C2EA09FE1F" box="[658,829,450,475]" italics="true" pageId="14" pageNumber="15">P. castaicensis</emphasis>
|
||
</taxonomicName>
|
||
(described above), both morphological and genetic evidence indicate that this snail is a new species.
|
||
</paragraph>
|
||
<paragraph id="8BA9D4483E018A0AFF6E600DEA71FDB6" blockId="14.[151,1437,152,626]" pageId="14" pageNumber="15">
|
||
Based on the road distances provided by Miller we infer that Pierpoint Spring is the well known (unnamed) spring that is shown on USGS maps along the north side of Hwy 190 about
|
||
<quantity id="4CEE79AD3E018A0AFB6B6032EC22FD88" box="[1217,1302,562,588]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="14" pageNumber="15" unit="km" value="0.6">0.6 km</quantity>
|
||
east of the Pierpoint Springs Resort and
|
||
<quantity id="4CEE79AD3E018A0AFE5A6058EB76FDB6" box="[496,578,600,626]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="14" pageNumber="15" unit="km" value="1.6">1.6 km</quantity>
|
||
east of Camp Nelson.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |