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<document id="586C744DE01B4051A63FA35306BB6FE3" ID-DOI="10.26107/RBZ-2019-0041" ID-ISSN="2345-7600" ID-Zenodo-Dep="4576444" ID-ZooBank="AAA866A1-FBAF-486D-AB84-3AB989A269C1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1614778122966" checkinUser="carolina" docAuthor="Páll-Gergely, Barna, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Hunyadi, András &amp; Jochum, Adrienne" docDate="2019" docId="03BF447DFF97FFA1FF1EFAC9FE97F88D" docLanguage="en" docName="RafflesBZool.67.517-535.pdf" docOrigin="Raffles Bulletin of Zoology 67" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Tonkinospira depressa" docType="treatment" docVersion="4" lastPageNumber="527" masterDocId="FF863C05FF9DFFABFFEEFA75FFE6FFC4" masterDocTitle="New Tonkinospira Jochum, Slapnik &amp; Páll-Gergely, 2014 species from Laos and Vietnam (Gastropoda: Pulmonata: Hypselostomatidae)" masterLastPageNumber="535" masterPageNumber="517" pageNumber="527" updateTime="1698878161501" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="3094FBAACC88780149259036FD1796B7">New Tonkinospira Jochum, Slapnik &amp; Páll-Gergely, 2014 species from Laos and Vietnam (Gastropoda: Pulmonata: Hypselostomatidae)</mods:title>
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<mods:namePart id="5D78CAEABE9C9ABEC202CE67B062B568">Páll-Gergely, Barna</mods:namePart>
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<mods:namePart id="53BEDC4BDCB180B285769BCB147CC64B">Grego, Jozef</mods:namePart>
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<mods:namePart id="F1845F60BA9F40758B2EB283C626703B">Vermeulen, Jaap J.</mods:namePart>
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<mods:namePart id="451673ABE5D12E813D52922A7D10942B">Reischütz, Alexander</mods:namePart>
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<mods:namePart id="F5F5DDDF5E57C94728280734BC8FA856">Hunyadi, András</mods:namePart>
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<mods:namePart id="FAE38622A5051640C6C40B5569579476">Jochum, Adrienne</mods:namePart>
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<mods:title id="1CEA3F4EDE8DEFFCECAD7EDEC43A8EEE">Raffles Bulletin of Zoology</mods:title>
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<mods:date id="A5F9749FE559DB5AD265B5366631E089">2019</mods:date>
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<mods:number id="C76957368B98C5394C8A3B2349DA7B06">2019-11-08</mods:number>
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<mods:number id="07E955495083601AA44A12F50889E411">67</mods:number>
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<treatment id="03BF447DFF97FFA1FF1EFAC9FE97F88D" ID-DOI="http://doi.org/10.5281/zenodo.4576555" ID-Zenodo-Dep="4576555" LSID="urn:lsid:plazi:treatment:03BF447DFF97FFA1FF1EFAC9FE97F88D" httpUri="http://treatment.plazi.org/id/03BF447DFF97FFA1FF1EFAC9FE97F88D" lastPageNumber="527" pageId="10" pageNumber="527">
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<paragraph id="8BA9F56BFF97FFA1FF1EFAC9FD79FF11" blockId="10.[240,671,188,245]" box="[240,671,188,213]" pageId="10" pageNumber="527">
<heading id="D0E14207FF97FFA1FF1EFAC9FD79FF11" bold="true" box="[240,671,188,213]" centered="true" fontSize="10" level="2" pageId="10" pageNumber="527" reason="2">
<taxonomicName id="4C168EE8FF97FFA1FF1EFAC9FD79FF11" ID-CoL="7CGCX" authority="(Jaeckel, 1950)" baseAuthorityName="Jaeckel" baseAuthorityYear="1950" box="[240,671,188,213]" class="Gastropoda" family="Gastrocoptidae" genus="Tonkinospira" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="527" phylum="Mollusca" rank="species" species="depressa">
<emphasis id="B9622979FF97FFA1FF1EFAC9FD79FF11" bold="true" box="[240,671,188,213]" pageId="10" pageNumber="527">
<emphasis id="B9622979FF97FFA1FF1EFAC9FE0AFF10" bold="true" box="[240,492,188,212]" italics="true" pageId="10" pageNumber="527">Tonkinospira depressa</emphasis>
(
<bibRefCitation id="EF87889AFF97FFA1FE15FAC8FD7EFF11" author="Jaeckel SH" box="[507,664,189,213]" pageId="10" pageNumber="527" pagination="15 - 20" refId="ref7243" refString="Jaeckel SH (1950) Die Mollusken eines tropischen Flussgenistes aus Tonkin. Archiv fur Molluskenkunde, 79: 15 - 20." type="journal article" year="1950">Jaeckel, 1950</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C30CA6E0FF97FFA1FE70FAA8FE16FF31" box="[414,496,221,245]" pageId="10" pageNumber="527" type="description">
<paragraph id="8BA9F56BFF97FFA1FE70FAA8FE16FF31" blockId="10.[240,671,188,245]" box="[414,496,221,245]" pageId="10" pageNumber="527">
(
<figureCitation id="132DE9EEFF97FFA1FE48FAA8FE0EFF31" box="[422,488,221,245]" captionStart="Fig" captionStartId="8.[136,171,1342,1364]" captionTargetBox="[136,1474,189,1327]" captionTargetId="figure-304@8.[136,1474,189,1327]" captionTargetPageId="8" captionText="Fig. 7. Tonkinospira depressa (Jaeckel, 1950), SMF 202927 (paratype). Scale bar represents 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4576468" httpUri="https://zenodo.org/record/4576468/files/figure.png" pageId="10" pageNumber="527">Fig. 7</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30CA6E0FF97FFA1FF66FB69FEDDFD8C" pageId="10" pageNumber="527" type="reference_group">
<paragraph id="8BA9F56BFF97FFA1FF66FB69FDFAFE8B" blockId="10.[136,775,283,392]" pageId="10" pageNumber="527">
<treatmentCitationGroup id="AB06D245FF97FFA1FF66FB69FDFAFE8B" pageId="10" pageNumber="527">
<treatmentCitation id="0AB7D37AFF97FFA1FF66FB69FDA1FEF5" author="Jaeckel SH" box="[136,583,283,305]" page="16" pageId="10" pageNumber="527" year="1950">
<taxonomicName id="4C168EE8FF97FFA1FF66FB69FDA1FEF5" ID-CoL="544QW" authority="Jaeckel, 1950: 15 - 16" authorityName="Jaeckel" authorityPageNumber="15 - 16" authorityYear="1950" box="[136,583,283,305]" class="Gastropoda" family="Gastrocoptidae" genus="Systenostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="527" phylum="Mollusca" rank="species" species="depressa">
<emphasis id="B9622979FF97FFA1FF66FB69FE8CFEF4" box="[136,362,283,305]" italics="true" pageId="10" pageNumber="527">Systenostoma depressa</emphasis>
<materialsCitation id="3B7EFF36FF97FFA1FE9DFB6EFDA1FEF5" box="[371,583,283,305]" country="Vietnam" location="Vietnam" pageId="10" pageNumber="527" specimenCount="1">
<bibRefCitation id="EF87889AFF97FFA1FE9DFB6EFDA1FEF5" author="Jaeckel SH" box="[371,583,283,305]" pageId="10" pageNumber="527" pagination="15 - 20" refId="ref7243" refString="Jaeckel SH (1950) Die Mollusken eines tropischen Flussgenistes aus Tonkin. Archiv fur Molluskenkunde, 79: 15 - 20." type="journal article" year="1950">Jaeckel, 1950: 1516</bibRefCitation>
</materialsCitation>
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</treatmentCitation>
, pl. 1, fig. 1. (
<typeStatus id="54AD4BC9FF97FFA1FD32FB69FCE1FEF5" box="[732,775,284,305]" pageId="10" pageNumber="527">type</typeStatus>
locality: Tonkin [northern
<collectingCountry id="F301B5FBFF97FFA1FE5DFB4DFDEAFE8B" box="[435,524,312,335]" name="Vietnam" pageId="10" pageNumber="527">Vietnam</collectingCountry>
]).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA9F56BFF97FFA1FF66FB20FD76FEAE" blockId="10.[136,775,283,392]" box="[136,656,341,363]" pageId="10" pageNumber="527">
<treatmentCitationGroup id="AB06D245FF97FFA1FF66FB20FD76FEAE" box="[136,656,341,363]" pageId="10" pageNumber="527">
<taxonomicName id="4C168EE8FF97FFA1FF66FB20FEBCFEAE" ID-CoL="7Q8WL" baseAuthorityName="Jaeckel" baseAuthorityYear="1950" box="[136,346,341,362]" class="Gastropoda" family="Gastrocoptidae" genus="Aulacospira" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="527" phylum="Mollusca" rank="species" species="depressa">
<emphasis id="B9622979FF97FFA1FF66FB20FEBCFEAE" box="[136,346,341,362]" italics="true" pageId="10" pageNumber="527">Aulacospira depressa</emphasis>
</taxonomicName>
<treatmentCitation id="0AB7D37AFF97FFA1FE6CFB20FD6AFEAE" author="Vermeulen JJ &amp; Phung LC &amp; Truong QT" box="[386,652,341,363]" page="91" pageId="10" pageNumber="527" year="2007">
<bibRefCitation id="EF87889AFF97FFA1FE6CFB20FD6AFEAE" author="Vermeulen JJ &amp; Phung LC &amp; Truong QT" box="[386,652,341,363]" pageId="10" pageNumber="527" pagination="81 - 92" refId="ref7890" refString="Vermeulen JJ, Phung LC &amp; Truong QT (2007) New species of terrestrial molluscs (Caenogastropoda, Pupinidae &amp; Pulmonata, Vertiginidae) of the Hon Chong - Ha Tien limestone hills, Southern Vietnam. Basteria, 71: 81 - 92." type="journal article" year="2007">Vermeulen et al., 2007: 91</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA9F56BFF97FFA1FF66FB07FD9DFE43" blockId="10.[136,775,283,392]" box="[136,635,370,392]" pageId="10" pageNumber="527">
<treatmentCitationGroup id="AB06D245FF97FFA1FF66FB07FD9DFE43" box="[136,635,370,392]" pageId="10" pageNumber="527">
<taxonomicName id="4C168EE8FF97FFA1FF66FB07FE83FE43" ID-CoL="7CGCX" baseAuthorityName="Jaeckel" baseAuthorityYear="1950" box="[136,357,370,391]" class="Gastropoda" family="Gastrocoptidae" genus="Tonkinospira" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="527" phylum="Mollusca" rank="species" species="depressa">
<emphasis id="B9622979FF97FFA1FF66FB07FE83FE43" box="[136,357,370,391]" italics="true" pageId="10" pageNumber="527">Tonkinospira depressa</emphasis>
</taxonomicName>
<treatmentCitation id="0AB7D37AFF97FFA1FE63FB07FD91FE43" author="Jochum A &amp; Slapnik R &amp; Kampschulte M &amp; Martels G &amp; Heneka M &amp; Pall-Gergely B" box="[397,631,370,392]" page="32" pageId="10" pageNumber="527" year="2014">
<bibRefCitation id="EF87889AFF97FFA1FE63FB07FD91FE43" author="Jochum A &amp; Slapnik R &amp; Kampschulte M &amp; Martels G &amp; Heneka M &amp; Pall-Gergely B" box="[397,631,370,392]" pageId="10" pageNumber="527" pagination="23 - 40" refId="ref7266" refString="Jochum A, Slapnik R, Kampschulte M, Martels G, Heneka M &amp; Pall-Gergely B (2014) A review of the microgastropod genus Systenostoma Bavay &amp; Dautzenberg, 1908 and a new subterranean species from China (Gastropoda, Pulmonata, Hypselostomatidae). ZooKeys, 410: 23 - 40." type="journal article" year="2014">Jochum et al., 2014: 32</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA9F56BFF97FFA1FF66FBC5FEDDFD8C" blockId="10.[136,776,432,584]" pageId="10" pageNumber="527">
<emphasis id="B9622979FF97FFA1FF66FBC5FEACFE0C" bold="true" box="[136,330,432,456]" pageId="10" pageNumber="527">Types examined.</emphasis>
<materialsCitation id="3B7EFF36FF97FFA1FEBDFBC5FE73FDCC" collectionCode="SMF" collectorName="S. H. Jaeckel &amp; H. Rolle" country="Vietnam" location="Flussgenist" pageId="10" pageNumber="527" specimenCode="SMF 202927" specimenCount="33" typeStatus="paratype">
<specimenCount id="9D103EE2FF97FFA1FEBDFBC5FE3BFE0D" box="[339,477,432,457]" pageId="10" pageNumber="527" type="generic" typeStatus="paratypes">33 paratypes</specimenCount>
(H = 1.6, D = 2.3,
<figureCitation id="132DE9EEFF97FFA1FD53FBC5FD18FE0C" box="[701,766,432,456]" captionStart="Fig" captionStartId="8.[136,171,1342,1364]" captionTargetBox="[136,1474,189,1327]" captionTargetId="figure-304@8.[136,1474,189,1327]" captionTargetPageId="8" captionText="Fig. 7. Tonkinospira depressa (Jaeckel, 1950), SMF 202927 (paratype). Scale bar represents 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4576468" httpUri="https://zenodo.org/record/4576468/files/figure.png" pageId="10" pageNumber="527">Fig. 7</figureCitation>
) (
<specimenCode id="DBB05D10FF97FFA1FF7FFBA5FEF9FE2C" box="[145,287,464,488]" collectionCode="SMF" country="Germany" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="10" pageNumber="527">SMF 202927</specimenCode>
),
<collectingCountry id="F301B5FBFF97FFA1FEDAFBA5FE63FE2C" box="[308,389,464,488]" name="Vietnam" pageId="10" pageNumber="527">Tonkin</collectingCountry>
:
<location id="8EC9A3B0FF97FFA1FE7EFBA5FDE7FE2C" LSID="urn:lsid:plazi:treatment:03BF447DFF97FFA1FF1EFAC9FE97F88D:8EC9A3B0FF97FFA1FE7EFBA5FDE7FE2C" box="[400,513,464,488]" country="Vietnam" name="Flussgenist" pageId="10" pageNumber="527">Flussgenist</location>
(river debris), coll.
<collectorName id="26E390BDFF97FFA1FD39FBA5FF30FDCC" pageId="10" pageNumber="527">S.H. Jaeckel</collectorName>
ex coll.
<collectorName id="26E390BDFF97FFA1FED8FB84FE73FDCC" box="[310,405,496,520]" pageId="10" pageNumber="527">H. Rolle</collectorName>
</materialsCitation>
;
<materialsCitation id="3B7EFF36FF97FFA1FE4FFB85FED1FD8C" accessionNumber="SMF 63875" collectingDate="1949-12-08" collectorName="Genist &amp; Zilch &amp; H. Rolle &amp; Jaeckel" country="Vietnam" location="Tonkin" pageId="10" pageNumber="527" specimenCount="8" typeStatus="paratype">
<specimenCount id="9D103EE2FF97FFA1FE4FFB85FDFAFDCD" box="[417,540,496,521]" pageId="10" pageNumber="527" type="generic" typeStatus="paratypes">8 paratypes</specimenCount>
SMF 63875,
<collectingCountry id="F301B5FBFF97FFA1FD5AFB85FCE2FDCC" box="[692,772,496,520]" name="Vietnam" pageId="10" pageNumber="527">Tonkin</collectingCountry>
,
<collectorName id="26E390BDFF97FFA1FF66F865FF34FDEC" box="[136,210,528,552]" pageId="10" pageNumber="527">Genist</collectorName>
, coll.
<collectorName id="26E390BDFF97FFA1FEF2F865FEBFFDEC" box="[284,345,528,552]" pageId="10" pageNumber="527">Zilch</collectorName>
, ex coll.
<collectorName id="26E390BDFF97FFA1FE28F864FDCFFDEC" box="[454,553,528,552]" pageId="10" pageNumber="527">H. Rolle</collectorName>
, ex coll.
<collectorName id="26E390BDFF97FFA1FD79F865FD0DFDEC" box="[663,747,528,552]" pageId="10" pageNumber="527">Jaeckel</collectorName>
,
<date id="FFA8D3ABFF97FFA1FD17F865FED1FD8C" pageId="10" pageNumber="527" value="1949-12-08">
<collectingDate id="EFEC2A43FF97FFA1FD17F865FED1FD8C" pageId="10" pageNumber="527" value="1949-12-08">8 December 1949</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30CA6E0FF97FFA1FF66F805FEB2FD2C" pageId="10" pageNumber="527" type="diagnosis">
<paragraph id="8BA9F56BFF97FFA1FF66F805FEB2FD2C" blockId="10.[136,775,624,744]" pageId="10" pageNumber="527">
<emphasis id="B9622979FF97FFA1FF66F805FF19FD4C" bold="true" box="[136,255,624,648]" pageId="10" pageNumber="527">Diagnosis.</emphasis>
A large
<taxonomicName id="4C168EE8FF97FFA1FE86F805FE1FFD4C" authorityName="Jochum, Slapnik &amp; Pall-Gergely" authorityYear="2014" box="[360,505,624,648]" class="Gastropoda" family="Gastrocoptidae" genus="Tonkinospira" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="527" phylum="Mollusca" rank="genus">
<emphasis id="B9622979FF97FFA1FE86F805FE1FFD4C" box="[360,505,624,648]" italics="true" pageId="10" pageNumber="527">Tonkinospira</emphasis>
</taxonomicName>
species with a strongly depressed shell, strongly angled body whorl, dense spiral striation, and a peristome that is more or less adnate to penultimate whorl.
</paragraph>
</subSubSection>
<subSubSection id="C30CA6E0FF97FFA1FF66F965FD83FB4C" pageId="10" pageNumber="527" type="description">
<paragraph id="8BA9F56BFF97FFA1FF66F965FD83FB4C" blockId="10.[135,776,784,1161]" pageId="10" pageNumber="527">
<emphasis id="B9622979FF97FFA1FF66F965FEFFFCEC" bold="true" box="[136,281,784,808]" pageId="10" pageNumber="527">Description.</emphasis>
Shell large-sized for the genus; depressed conical with rapidly increasing whorls; body whorl strongly angled, keel not symmetrical, situated above mid line of body whorl, flattened above; keel not very sharp; whorls 3.754; suture moderately deep; protoconch consists of 1.25 whorls, roughly rugose and spirally striated; teleoconch with rather regular spiral striation and irregular, rough radial lines; aperture oblique to shell axis from lateral view; aperture ovate-oblong; peristome expanded especially in direction of the columella/umbilicus; aperture more or less adnate to penultimate whorl, but peristome continuous; umbilicus narrow, very slightly covered by peristome.
</paragraph>
</subSubSection>
<subSubSection id="C30CA6E0FF97FFA1FF66FEC5FE8AFACC" pageId="10" pageNumber="527" type="diagnosis">
<paragraph id="8BA9F56BFF97FFA1FF66FEC5FE8AFACC" blockId="10.[136,776,1200,1288]" pageId="10" pageNumber="527">
<emphasis id="B9622979FF97FFA1FF66FEC5FE71FB0C" bold="true" box="[136,407,1200,1224]" pageId="10" pageNumber="527">Differential diagnosis.</emphasis>
The most similar species is
<taxonomicName id="4C168EE8FF97FFA1FD1FFEC4FF0DFB2C" baseAuthorityName="Bavay &amp; Dautzenberg" baseAuthorityYear="1909" class="Gastropoda" family="Gastrocoptidae" genus="Tonkinospira" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="527" phylum="Mollusca" rank="species" species="pulverea">
<emphasis id="B9622979FF97FFA1FD1FFEC4FF0DFB2C" italics="true" pageId="10" pageNumber="527">T. pulverea</emphasis>
</taxonomicName>
, which has a narrower umbilicus and a more rounded body whorl.
</paragraph>
</subSubSection>
<subSubSection id="C30CA6E0FF97FFA1FF66FF45FE97F88D" pageId="10" pageNumber="527" type="discussion">
<paragraph id="8BA9F56BFF97FFA1FF66FF45FE97F88D" blockId="10.[135,776,1328,1865]" pageId="10" pageNumber="527">
<emphasis id="B9622979FF97FFA1FF66FF45FF1CFA8C" bold="true" box="[136,250,1328,1352]" pageId="10" pageNumber="527">Remarks.</emphasis>
<taxonomicName id="4C168EE8FF97FFA1FEEBFF45FDE3FA8C" baseAuthorityName="Jaeckel" baseAuthorityYear="1950" box="[261,517,1328,1352]" class="Gastropoda" family="Gastrocoptidae" genus="Tonkinospira" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="527" phylum="Mollusca" rank="species" species="depressa">
<emphasis id="B9622979FF97FFA1FEEBFF45FDE3FA8C" box="[261,517,1328,1352]" italics="true" pageId="10" pageNumber="527">Tonkinospira depressa</emphasis>
</taxonomicName>
is most similar to the Philippine members of the genus
<taxonomicName id="4C168EE8FF97FFA1FE15FF25FD9DFAAC" authorityName="Moellendorff" authorityYear="1890" box="[507,635,1360,1384]" class="Gastropoda" family="Gastrocoptidae" genus="Aulacospira" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="527" phylum="Mollusca" rank="genus">
<emphasis id="B9622979FF97FFA1FE15FF25FD9DFAAC" box="[507,635,1360,1384]" italics="true" pageId="10" pageNumber="527">Aulacospira</emphasis>
</taxonomicName>
, and reflects this classification by
<bibRefCitation id="EF87889AFF97FFA1FE95FF05FD69FA4C" author="Vermeulen JJ &amp; Phung LC &amp; Truong QT" box="[379,655,1392,1416]" pageId="10" pageNumber="527" pagination="81 - 92" refId="ref7890" refString="Vermeulen JJ, Phung LC &amp; Truong QT (2007) New species of terrestrial molluscs (Caenogastropoda, Pupinidae &amp; Pulmonata, Vertiginidae) of the Hon Chong - Ha Tien limestone hills, Southern Vietnam. Basteria, 71: 81 - 92." type="journal article" year="2007">Vermeulen et al. (2007)</bibRefCitation>
. The most similar species is
<emphasis id="B9622979FF97FFA1FED3FFE5FDC1FA6C" box="[317,551,1424,1448]" italics="true" pageId="10" pageNumber="527">
<taxonomicName id="4C168EE8FF97FFA1FED3FFE5FE58FA6C" authorityName="Moellendorff" authorityYear="1890" box="[317,446,1424,1448]" class="Gastropoda" family="Gastrocoptidae" genus="Aulacospira" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="527" phylum="Mollusca" rank="genus">Aulacospira</taxonomicName>
hololoma
</emphasis>
(
<bibRefCitation id="EF87889AFF97FFA1FDDAFFE5FD1DFA6C" author="Mollendorff OF von" box="[564,763,1424,1448]" pageId="10" pageNumber="527" pagination="235 - 283" refId="ref7394" refString="Mollendorff OF von (1887) Von den Philippinen. II. Die Insel Cebu. Jahrbucher der Deutschen Malakozoologischen Gesellschaft, 14: 235 - 283. https: // biodiversitylibrary. org / page / 15886780." type="journal article" year="1887">Möllendorff, 1887</bibRefCitation>
), which has a regularly growing protoconch (tightly spired and projected above the teleoconch in
<taxonomicName id="4C168EE8FF97FFA1FDA6FFA4FD2DFA2C" baseAuthorityName="Jaeckel" baseAuthorityYear="1950" box="[584,715,1488,1512]" class="Gastropoda" family="Gastrocoptidae" genus="Tonkinospira" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="527" phylum="Mollusca" rank="species" species="depressa">
<emphasis id="B9622979FF97FFA1FDA6FFA4FD2DFA2C" box="[584,715,1488,1512]" italics="true" pageId="10" pageNumber="527">T. depressa</emphasis>
</taxonomicName>
) and bears a columellar tooth (lacks in
<taxonomicName id="4C168EE8FF97FFA1FDEFFF84FD98F9CC" baseAuthorityName="Jaeckel" baseAuthorityYear="1950" box="[513,638,1520,1544]" class="Gastropoda" family="Gastrocoptidae" genus="Tonkinospira" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="527" phylum="Mollusca" rank="species" species="depressa">
<emphasis id="B9622979FF97FFA1FDEFFF84FD98F9CC" box="[513,638,1520,1544]" italics="true" pageId="10" pageNumber="527">T. depressa</emphasis>
</taxonomicName>
). Moreover, the Philippine species typically have a prominent subsutural furrow on the body whorl, with the keel situated above the middle line of the body whorl (i.e., the keel is not symmetrical) (see
<bibRefCitation id="EF87889AFF97FFA1FEBCFC05FD82F94C" author="Pall-Gergely B &amp; Schilthuizen M &amp; Orstan A &amp; Auffenberg K" box="[338,612,1648,1672]" pageId="10" pageNumber="527" pagination="67 - 83" refId="ref7584" refString="Pall-Gergely B, Schilthuizen M, Orstan A &amp; Auffenberg K (2019) A review of Aulacospira Mollendorff, 1890 and Pseudostreptaxis Mollendorff, 1890 in the Philippines (Gastropoda, Pupilloidea, Hypselostomatidae). ZooKeys, 842: 67 - 83. https: // doi. org / 10.3897 / zookeys. 842.33052." type="journal article" year="2019">Páll-Gergely et al., 2019</bibRefCitation>
). Although the similarity between
<taxonomicName id="4C168EE8FF97FFA1FEBEFCE5FE38F96C" authorityName="Jochum, Slapnik &amp; Pall-Gergely" authorityYear="2014" box="[336,478,1680,1704]" class="Gastropoda" family="Gastrocoptidae" genus="Tonkinospira" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="527" phylum="Mollusca" rank="genus">
<emphasis id="B9622979FF97FFA1FEBEFCE5FE38F96C" box="[336,478,1680,1704]" italics="true" pageId="10" pageNumber="527">Tonkinospira</emphasis>
</taxonomicName>
and
<taxonomicName id="4C168EE8FF97FFA1FDE1FCE5FD77F96C" authorityName="Moellendorff" authorityYear="1890" box="[527,657,1680,1704]" class="Gastropoda" family="Gastrocoptidae" genus="Aulacospira" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="527" phylum="Mollusca" rank="genus">
<emphasis id="B9622979FF97FFA1FDE1FCE5FD77F96C" box="[527,657,1680,1704]" italics="true" pageId="10" pageNumber="527">Aulacospira</emphasis>
</taxonomicName>
is striking, the true phylogenetic relationships cannot be resolved based on conchological characters only.
<taxonomicName id="4C168EE8FF97FFA1FD80FCA5FF01F8CC" baseAuthorityName="Jaeckel" baseAuthorityYear="1950" class="Gastropoda" family="Gastrocoptidae" genus="Tonkinospira" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="527" phylum="Mollusca" rank="species" species="depressa">
<emphasis id="B9622979FF97FFA1FD80FCA5FF01F8CC" italics="true" pageId="10" pageNumber="527">Tonkinospira depressa</emphasis>
</taxonomicName>
further differs from other
<taxonomicName id="4C168EE8FF97FFA1FDE6FC85FD70F8CC" authorityName="Jochum, Slapnik &amp; Pall-Gergely" authorityYear="2014" box="[520,662,1776,1800]" class="Gastropoda" family="Gastrocoptidae" genus="Tonkinospira" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="527" phylum="Mollusca" rank="genus">
<emphasis id="B9622979FF97FFA1FDE6FC85FD70F8CC" box="[520,662,1776,1800]" italics="true" pageId="10" pageNumber="527">Tonkinospira</emphasis>
</taxonomicName>
species in the brownish shell colour, which is colourless in the other species of this genus.
</paragraph>
</subSubSection>
</treatment>
</document>