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<document id="B54D3517B61A704CCCC80994A5F1F9BB" ID="10.11646/zootaxa.4157.1.1" ID-DOI="10.11646/zootaxa.4157.1.1" ID-GBIF-Dataset="2389f078-1812-4b4e-826a-561cd7e46654" ID-ISSN="1175-5326" ID-Zenodo-Dep="262413" ID-ZooBank="6BF4514A-892F-499F-BC1E-B7920C7A00B0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1472198526050" checkinUser="plazi" docAuthor="Jałoszyński, Paweł &amp; Perkovsky, Evgeny" docDate="2016" docId="03A187A92927FFEEFF03C5BA9941FE91" docLanguage="en" docName="zootaxa.4157.1.1.pdf" docOrigin="Zootaxa 4157 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Cephennomicrus" docType="treatment" docVersion="9" lastPageNumber="32" masterDocId="FF98FFD12939FFF1FF94C52D9D54FFD1" masterDocTitle="Diversity of Scydmaeninae (Coleoptera: Staphylinidae) in Upper Eocene Rovno amber" masterLastPageNumber="85" masterPageNumber="1" pageNumber="31" updateTime="1698382676775" updateUser="plazi">
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<mods:title id="3677CBFC2BBC5344275783403834CA80">Diversity of Scydmaeninae (Coleoptera: Staphylinidae) in Upper Eocene Rovno amber</mods:title>
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<mods:namePart id="3EAC7B109A53B56DEE1862F7CCE877A7">Jałoszyński, Paweł</mods:namePart>
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<mods:namePart id="4CD5326A36485D4F6E09B6181344892A">Perkovsky, Evgeny</mods:namePart>
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<treatment id="03A187A92927FFEEFF03C5BA9941FE91" ID-DOI="http://doi.org/10.5281/zenodo.5681751" ID-GBIF-Taxon="124407840" ID-Zenodo-Dep="5681751" LSID="urn:lsid:plazi:treatment:03A187A92927FFEEFF03C5BA9941FE91" httpUri="http://treatment.plazi.org/id/03A187A92927FFEEFF03C5BA9941FE91" lastPageId="31" lastPageNumber="32" pageId="30" pageNumber="31">
<subSubSection id="C31265342927FFEFFF03C5BA9C17FF02" pageId="30" pageNumber="31" type="nomenclature">
<paragraph id="8BB736BF2927FFEFFF03C5BA9CF2FF63" blockId="30.[151,422,151,211]" box="[151,422,151,178]" pageId="30" pageNumber="31">
<heading id="D0FF81D32927FFEFFF03C5BA9CF2FF63" bold="true" box="[151,422,151,178]" fontSize="11" level="1" pageId="30" pageNumber="31" reason="1">
<emphasis id="B97CEAAD2927FFEFFF03C5BA9CF2FF63" bold="true" box="[151,422,151,178]" pageId="30" pageNumber="31">
<taxonomicName id="4C084D3C2927FFEFFF03C5BA9C32FF60" ID-CoL="8XKBK" box="[151,358,151,177]" class="Insecta" family="Staphylinidae" genus="Cephennomicrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="B97CEAAD2927FFEFFF03C5BA9C32FF60" bold="true" box="[151,358,151,177]" italics="true" pageId="30" pageNumber="31">Cephennomicrus</emphasis>
</taxonomicName>
sp. 4
</emphasis>
</heading>
</paragraph>
<paragraph id="8BB736BF2927FFEFFF03C5979C17FF02" blockId="30.[151,422,151,211]" box="[151,323,186,211]" pageId="30" pageNumber="31">
(
<figureCitation id="13332A3A2927FFEFFF0BC5979DB3FF02" box="[159,231,186,211]" captionStart="FIGURES 1 20" captionStartId="7.[151,264,1794,1817]" captionTargetBox="[157,1423,512,1763]" captionTargetId="figure@7.[151,1436,503,1772]" captionTargetPageId="7" captionText="FIGURES 1 20. Specimens of Rovno amber with inclusions of Scydmaeninae. K- 27418, Rovnoleptochromus ableptonoides sp. n. (1); K- 243, Vertheia quadrisetosa sp. n. (2); UA- 442, Vertheia sp. 1 (3); K- 800, Cephennodes sp. 1 (4); K- 6011, Cephennomicrus giganteus sp. n. (5); K- 8289, Cephennomicrus sp. 1 (6); K- 7010, Cephennomicrus sp. 2 (7); K- 27565, Cephennomicrus sp. 3 (8); K- 8912, Cephennomicrus sp. 4 (9); DU- 32, Cephennomicrus sp. 5 (10); K- 5770, Glaesoconnus unicus sp. n. (11); K- 9579, Rovnoscydmus frontalis sp. n. (12); K- 347, Rovnoscydmus microscopicus sp. n. (13); K- 25702, Rovnoscydmus sp. specimen 1 (14); K- 26348, Rovnoscydmus sp. specimen 2 (15); UA- 1556, Rovnoscydmus sp. specimen 3 (16); K- 8970, Rovnoscydmus sp. specimen 4 (17); K- 8968, Rovnoscydmus sp. specimen 5 (18); K- 7350, Rovnoscydmus sp. specimen 6 (19); K- 3037, Rovnoscydmus sp. specimen 7 (20)." httpUri="https://zenodo.org/record/262414/files/figure.png" pageId="30" pageNumber="31">Figs 9</figureCitation>
,
<figureCitation id="13332A3A2927FFEFFF67C5969C6EFF02" box="[243,314,187,211]" captionStart="FIGURES 93 98" captionStartId="30.[151,264,1984,2007]" captionTargetBox="[151,1436,687,1952]" captionTargetId="figure@30.[151,1436,687,1963]" captionTargetPageId="30" captionText="FIGURES 93 98. Cephennomicrus sp. 4 (K- 8912). Anterodorsal (93, 94, 96, 97) and ventrolateral (95, 98) view. Abbreviations: bst, basisternal part of prosternum; psp, prosternal process; mtvp, metaventral intercoxal process." httpUri="https://zenodo.org/record/262429/files/figure.png" pageId="30" pageNumber="31">9398</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31265342927FFEFFF03C42F9909FEEE" pageId="30" pageNumber="31" type="materials_examined">
<paragraph id="8BB736BF2927FFEFFF03C42F9909FEEE" blockId="30.[151,1437,258,643]" pageId="30" pageNumber="31">
<materialsCitation id="3B603CE22927FFEFFF03C42F9909FEEE" ID-GBIF-Occurrence="1300093502" collectionCode="SIZK" county="Material" location="Europe" municipality="Late Eocene" pageId="30" pageNumber="31" specimenCount="1" stateProvince="Rovno">
<emphasis id="B97CEAAD2927FFEFFF03C42F9E34FECA" bold="true" box="[151,864,258,283]" pageId="30" pageNumber="31">
<collectingCounty id="62D64E332927FFEFFF03C42F9DABFECA" box="[151,255,258,283]" pageId="30" pageNumber="31">Material</collectingCounty>
studied.
<collectingMunicipality id="6BD3ACC52927FFEFFEE3C42F9F45FECA" box="[375,529,258,283]" pageId="30" pageNumber="31">Late Eocene</collectingMunicipality>
of
<location id="8ED760642927FFEFFDD6C42F9FC8FECA" LSID="urn:lsid:plazi:treatment:03A187A92927FFEEFF03C5BA9941FE91:8ED760642927FFEFFDD6C42F9FC8FECA" box="[578,668,258,283]" county="Material" municipality="Late Eocene" name="Europe" pageId="30" pageNumber="31" stateProvince="Rovno">Europe</location>
,
<collectingRegion id="49CCF85D2927FFEFFD3AC42F9FA8FECA" box="[686,764,258,283]" country="Ukraine" name="Rivne" pageId="30" pageNumber="31">Rovno</collectingRegion>
amber:
</emphasis>
sex unknown; inclusion in elongate, irregularly subtriangular piece of amber
<quantity id="4CF09B5A2927FFEFFE4BC40B9F61FEEF" box="[479,565,294,319]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="30" pageNumber="31" unit="mm" value="9.5">9.5 mm</quantity>
long (
<figureCitation id="13332A3A2927FFEFFDE9C40B9F95FEEE" box="[637,705,294,319]" captionStart="FIGURES 1 20" captionStartId="7.[151,264,1794,1817]" captionTargetBox="[157,1423,512,1763]" captionTargetId="figure@7.[151,1436,503,1772]" captionTargetPageId="7" captionText="FIGURES 1 20. Specimens of Rovno amber with inclusions of Scydmaeninae. K- 27418, Rovnoleptochromus ableptonoides sp. n. (1); K- 243, Vertheia quadrisetosa sp. n. (2); UA- 442, Vertheia sp. 1 (3); K- 800, Cephennodes sp. 1 (4); K- 6011, Cephennomicrus giganteus sp. n. (5); K- 8289, Cephennomicrus sp. 1 (6); K- 7010, Cephennomicrus sp. 2 (7); K- 27565, Cephennomicrus sp. 3 (8); K- 8912, Cephennomicrus sp. 4 (9); DU- 32, Cephennomicrus sp. 5 (10); K- 5770, Glaesoconnus unicus sp. n. (11); K- 9579, Rovnoscydmus frontalis sp. n. (12); K- 347, Rovnoscydmus microscopicus sp. n. (13); K- 25702, Rovnoscydmus sp. specimen 1 (14); K- 26348, Rovnoscydmus sp. specimen 2 (15); UA- 1556, Rovnoscydmus sp. specimen 3 (16); K- 8970, Rovnoscydmus sp. specimen 4 (17); K- 8968, Rovnoscydmus sp. specimen 5 (18); K- 7350, Rovnoscydmus sp. specimen 6 (19); K- 3037, Rovnoscydmus sp. specimen 7 (20)." httpUri="https://zenodo.org/record/262414/files/figure.png" pageId="30" pageNumber="31">Fig. 9</figureCitation>
), collection number K-8912 (
<collectionCode id="ED19AE7A2927FFEFFB99C40A9905FEEE" box="[1037,1105,295,319]" country="Ukraine" httpUri="http://grbio.org/cool/c442-9tdv" name="Schmaulhausen Institute of Zoology" pageId="30" pageNumber="31">SIZK</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C31265342927FFEEFF53C4679EEDFF29" lastPageId="31" lastPageNumber="32" pageId="30" pageNumber="31" type="description">
<paragraph id="8BB736BF2927FFEFFF53C46799EBFEB3" blockId="30.[151,1437,258,643]" box="[199,1215,330,355]" pageId="30" pageNumber="31">
<emphasis id="B97CEAAD2927FFEFFF53C4679C0DFEB2" bold="true" box="[199,345,330,355]" pageId="30" pageNumber="31">Description.</emphasis>
Body (
<figureCitation id="13332A3A2927FFEFFE3EC4679F7EFEB2" box="[426,554,330,355]" captionStart="FIGURES 93 98" captionStartId="30.[151,264,1984,2007]" captionTargetBox="[151,1436,687,1952]" captionTargetId="figure@30.[151,1436,687,1963]" captionTargetPageId="30" captionText="FIGURES 93 98. Cephennomicrus sp. 4 (K- 8912). Anterodorsal (93, 94, 96, 97) and ventrolateral (95, 98) view. Abbreviations: bst, basisternal part of prosternum; psp, prosternal process; mtvp, metaventral intercoxal process." httpUri="https://zenodo.org/record/262429/files/figure.png" pageId="30" pageNumber="31">Figs 9398</figureCitation>
) elongate and strongly convex, nearly black; BL 1.00 mm.
</paragraph>
<paragraph id="8BB736BF2927FFEFFF53C4439ED1FE22" blockId="30.[151,1437,258,643]" pageId="30" pageNumber="31">
Head (
<figureCitation id="13332A3A2927FFEFFE85C4439CB2FE56" box="[273,486,366,391]" captionStart="FIGURES 93 98" captionStartId="30.[151,264,1984,2007]" captionTargetBox="[151,1436,687,1952]" captionTargetId="figure@30.[151,1436,687,1963]" captionTargetPageId="30" captionText="FIGURES 93 98. Cephennomicrus sp. 4 (K- 8912). Anterodorsal (93, 94, 96, 97) and ventrolateral (95, 98) view. Abbreviations: bst, basisternal part of prosternum; psp, prosternal process; mtvp, metaventral intercoxal process." httpUri="https://zenodo.org/record/262429/files/figure.png" pageId="30" pageNumber="31">Figs 9394, 9697</figureCitation>
) broadest at eyes, HL
<quantity id="4CF09B5A2927FFEFFD76C4429E1DFE57" box="[738,841,367,391]" metricMagnitude="-4" metricUnit="m" metricValue="1.3" pageId="30" pageNumber="31" unit="mm" value="0.13">0.13 mm</quantity>
, HW
<quantity id="4CF09B5A2927FFEFFC1EC4429EA5FE57" box="[906,1009,367,391]" metricMagnitude="-4" metricUnit="m" metricValue="2.8" pageId="30" pageNumber="31" unit="mm" value="0.28">0.28 mm</quantity>
; head structures poorly visible, vertex and frons confluent, weakly convex and together strongly transverse; supraantennal tubercles weakly marked; eyes large and strongly convex, moderately coarsely faceted. Antennae (
<figureCitation id="13332A3A2927FFEFFC36C49B9971FE1E" box="[930,1061,438,463]" captionStart="FIGURES 93 98" captionStartId="30.[151,264,1984,2007]" captionTargetBox="[151,1436,687,1952]" captionTargetId="figure@30.[151,1436,687,1963]" captionTargetPageId="30" captionText="FIGURES 93 98. Cephennomicrus sp. 4 (K- 8912). Anterodorsal (93, 94, 96, 97) and ventrolateral (95, 98) view. Abbreviations: bst, basisternal part of prosternum; psp, prosternal process; mtvp, metaventral intercoxal process." httpUri="https://zenodo.org/record/262429/files/figure.png" pageId="30" pageNumber="31">Figs 9398</figureCitation>
) about as long as half BL, AnL about
<quantity id="4CF09B5A2927FFEFFF48C4F79C16FE23" box="[220,322,474,499]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="30" pageNumber="31" unit="mm" value="0.5">0.50 mm</quantity>
, shapes of antennomeres obscured by turbid amber.
</paragraph>
<paragraph id="8BB736BF2927FFEFFF53C4D29C40FD52" blockId="30.[151,1437,258,643]" pageId="30" pageNumber="31">
Pronotum (
<figureCitation id="13332A3A2927FFEFFED2C4D39F4AFDC6" box="[326,542,510,535]" captionStart="FIGURES 93 98" captionStartId="30.[151,264,1984,2007]" captionTargetBox="[151,1436,687,1952]" captionTargetId="figure@30.[151,1436,687,1963]" captionTargetPageId="30" captionText="FIGURES 93 98. Cephennomicrus sp. 4 (K- 8912). Anterodorsal (93, 94, 96, 97) and ventrolateral (95, 98) view. Abbreviations: bst, basisternal part of prosternum; psp, prosternal process; mtvp, metaventral intercoxal process." httpUri="https://zenodo.org/record/262429/files/figure.png" pageId="30" pageNumber="31">Figs 9394, 9697</figureCitation>
) subrectangular, but exact shape difficult to asses in the studied specimen; PL
<quantity id="4CF09B5A2927FFEFFF03C70E9DABFDEB" box="[151,255,547,571]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="30" pageNumber="31" unit="mm" value="0.3">0.30 mm</quantity>
, PW
<quantity id="4CF09B5A2927FFEFFED4C70E9CFDFDEB" box="[320,425,547,571]" metricMagnitude="-4" metricUnit="m" metricValue="3.8" pageId="30" pageNumber="31" unit="mm" value="0.38">0.38 mm</quantity>
; pronotal disc broadest most likely near anterior third, sides rounded; posterior pronotal corners obtuse-angled; posterior margin arcuate; antebasal structures (i.e., pits and grooves), punctures and setae not visible.
</paragraph>
<caption id="DF7766372927FFEFFF03C2ED9858F824" httpUri="https://zenodo.org/record/262429/files/figure.png" pageId="30" pageNumber="31" targetBox="[151,1436,687,1952]" targetPageId="30">
<paragraph id="8BB736BF2927FFEFFF03C2ED9858F824" blockId="30.[151,1437,1984,2037]" pageId="30" pageNumber="31">
<emphasis id="B97CEAAD2927FFEFFF03C2ED9C09F807" bold="true" box="[151,349,1984,2007]" pageId="30" pageNumber="31">FIGURES 9398.</emphasis>
<taxonomicName id="4C084D3C2927FFEFFEFFC2EC9F43F806" box="[363,535,1985,2007]" class="Insecta" family="Staphylinidae" genus="Cephennomicrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="B97CEAAD2927FFEFFEFFC2EC9F43F806" box="[363,535,1985,2007]" italics="true" pageId="30" pageNumber="31">Cephennomicrus</emphasis>
</taxonomicName>
sp. 4 (K-8912). Anterodorsal (93, 94, 96, 97) and ventrolateral (95, 98) view. Abbreviations: bst, basisternal part of prosternum; psp, prosternal process; mtvp, metaventral intercoxal process.
</paragraph>
</caption>
<paragraph id="8BB736BF2926FFEEFF53C5BA9CA2FF04" blockId="31.[151,1437,151,320]" pageId="31" pageNumber="32">
Elytra (
<figureCitation id="13332A3A2926FFEEFE8FC5BA9CA6FF61" box="[283,498,151,176]" captionStart="FIGURES 93 98" captionStartId="30.[151,264,1984,2007]" captionTargetBox="[151,1436,687,1952]" captionTargetId="figure@30.[151,1436,687,1963]" captionTargetPageId="30" captionText="FIGURES 93 98. Cephennomicrus sp. 4 (K- 8912). Anterodorsal (93, 94, 96, 97) and ventrolateral (95, 98) view. Abbreviations: bst, basisternal part of prosternum; psp, prosternal process; mtvp, metaventral intercoxal process." httpUri="https://zenodo.org/record/262429/files/figure.png" pageId="31" pageNumber="32">Figs 9394, 9697</figureCitation>
) elongate oval, strongly convex, EL about
<quantity id="4CF09B5A2926FFEEFC4AC5BA9911FF7E" box="[990,1093,151,176]" metricMagnitude="-4" metricUnit="m" metricValue="5.8" pageId="31" pageNumber="32" unit="mm" value="0.58">0.58 mm</quantity>
, EW
<quantity id="4CF09B5A2926FFEEFB12C5BA99B9FF7E" box="[1158,1261,151,176]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="31" pageNumber="32" unit="mm" value="0.5">0.50 mm</quantity>
, EI about 1.15; punctures and setae not visible.
</paragraph>
<paragraph id="8BB736BF2926FFEEFF53C5CD9EEDFF29" blockId="31.[151,1437,151,320]" box="[199,953,223,248]" pageId="31" pageNumber="32">
Legs (
<figureCitation id="13332A3A2926FFEEFE98C5F29CD8FF29" box="[268,396,223,248]" captionStart="FIGURES 93 98" captionStartId="30.[151,264,1984,2007]" captionTargetBox="[151,1436,687,1952]" captionTargetId="figure@30.[151,1436,687,1963]" captionTargetPageId="30" captionText="FIGURES 93 98. Cephennomicrus sp. 4 (K- 8912). Anterodorsal (93, 94, 96, 97) and ventrolateral (95, 98) view. Abbreviations: bst, basisternal part of prosternum; psp, prosternal process; mtvp, metaventral intercoxal process." httpUri="https://zenodo.org/record/262429/files/figure.png" pageId="31" pageNumber="32">Figs 95, 98</figureCitation>
) only partly visible, moderately long and slender.
</paragraph>
</subSubSection>
<subSubSection id="C31265342926FFEEFF53C4299941FE91" pageId="31" pageNumber="32" type="discussion">
<paragraph id="8BB736BF2926FFEEFF53C4299941FE91" blockId="31.[151,1437,151,320]" pageId="31" pageNumber="32">
<emphasis id="B97CEAAD2926FFEEFF53C4299C6FFECC" bold="true" box="[199,315,260,285]" pageId="31" pageNumber="32">Remarks.</emphasis>
This specimen is poorly preserved, but the general body form and some ventral characters (
<figureCitation id="13332A3A2926FFEEFAD6C42998C0FECD" box="[1346,1428,260,285]" captionStart="FIGURES 93 98" captionStartId="30.[151,264,1984,2007]" captionTargetBox="[151,1436,687,1952]" captionTargetId="figure@30.[151,1436,687,1963]" captionTargetPageId="30" captionText="FIGURES 93 98. Cephennomicrus sp. 4 (K- 8912). Anterodorsal (93, 94, 96, 97) and ventrolateral (95, 98) view. Abbreviations: bst, basisternal part of prosternum; psp, prosternal process; mtvp, metaventral intercoxal process." httpUri="https://zenodo.org/record/262429/files/figure.png" pageId="31" pageNumber="32">Fig. 98</figureCitation>
) allow for placing it in
<taxonomicName id="4C084D3C2926FFEEFE04C40A9F1AFE91" box="[400,590,295,320]" class="Insecta" family="Staphylinidae" genus="Cephennomicrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="B97CEAAD2926FFEEFE04C40A9F1AFE91" box="[400,590,295,320]" italics="true" pageId="31" pageNumber="32">Cephennomicrus</emphasis>
</taxonomicName>
. See remarks for
<taxonomicName id="4C084D3C2926FFEEFC80C40A9E86FE91" box="[788,978,295,320]" class="Insecta" family="Staphylinidae" genus="Cephennomicrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="B97CEAAD2926FFEEFC80C40A9E86FE91" box="[788,978,295,320]" italics="true" pageId="31" pageNumber="32">Cephennomicrus</emphasis>
</taxonomicName>
sp. 2.
</paragraph>
</subSubSection>
</treatment>
</document>