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<document id="28EF0DBB33DB8214E7A5337D6998726F" ID="10.11646/zootaxa.4157.1.1" ID-DOI="10.11646/zootaxa.4157.1.1" ID-GBIF-Dataset="2389f078-1812-4b4e-826a-561cd7e46654" ID-ISSN="1175-5326" ID-Zenodo-Dep="262413" ID-ZooBank="6BF4514A-892F-499F-BC1E-B7920C7A00B0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1472198526050" checkinUser="plazi" docAuthor="Jałoszyński, Paweł &amp; Perkovsky, Evgeny" docDate="2016" docId="03A187A92907FFB0FF03C0B49ED7F933" docLanguage="en" docName="zootaxa.4157.1.1.pdf" docOrigin="Zootaxa 4157 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Stenichnus Jałoszyński &amp; Perkovsky, 2016, sp. n." docType="treatment" docVersion="9" lastPageNumber="66" masterDocId="FF98FFD12939FFF1FF94C52D9D54FFD1" masterDocTitle="Diversity of Scydmaeninae (Coleoptera: Staphylinidae) in Upper Eocene Rovno amber" masterLastPageNumber="85" masterPageNumber="1" pageNumber="63" updateTime="1698382676775" updateUser="plazi">
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<mods:title id="040DF223AB0F1155E99AE789789CB77F">Diversity of Scydmaeninae (Coleoptera: Staphylinidae) in Upper Eocene Rovno amber</mods:title>
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<mods:namePart id="D9D6E0E43E8B2B73252C067E9AB011E4">Jałoszyński, Paweł</mods:namePart>
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<mods:namePart id="AF5753F61CFA4DA7B5C7A92FDA65D3A2">Perkovsky, Evgeny</mods:namePart>
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<treatment id="03A187A92907FFB0FF03C0B49ED7F933" ID-DOI="http://doi.org/10.5281/zenodo.5681835" ID-GBIF-Taxon="124407861" ID-Zenodo-Dep="5681835" LSID="urn:lsid:plazi:treatment:03A187A92907FFB0FF03C0B49ED7F933" httpUri="http://treatment.plazi.org/id/03A187A92907FFB0FF03C0B49ED7F933" lastPageId="65" lastPageNumber="66" pageId="62" pageNumber="63">
<subSubSection id="C31265342907FFCFFF03C0B49C38FA05" pageId="62" pageNumber="63" type="nomenclature">
<paragraph id="8BB736BF2907FFCFFF03C0B49F6CFA62" blockId="62.[151,568,1433,1493]" box="[151,568,1433,1459]" pageId="62" pageNumber="63">
<heading id="D0FF81D32907FFCFFF03C0B49F6CFA62" bold="true" box="[151,568,1433,1459]" fontSize="11" level="1" pageId="62" pageNumber="63" reason="1">
<emphasis id="B97CEAAD2907FFCFFF03C0B49F6CFA62" bold="true" box="[151,568,1433,1459]" pageId="62" pageNumber="63">
<taxonomicName id="4C084D3C2907FFCFFF31C0B49C7FFA62" ID-CoL="8VS44" box="[165,299,1433,1459]" class="Insecta" family="Staphylinidae" genus="Stenichnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="genus">
<emphasis id="B97CEAAD2907FFCFFF31C0B49C7FFA62" bold="true" box="[165,299,1433,1459]" italics="true" pageId="62" pageNumber="63">Stenichnus</emphasis>
</taxonomicName>
(s. str.)
<taxonomicName id="4C084D3C2907FFCFFE1AC0B49CBAFA62" box="[398,494,1433,1459]" class="Insecta" family="Staphylinidae" genus="Stenichnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="proavus" status="sp. nov.">
<emphasis id="B97CEAAD2907FFCFFE1AC0B49CBAFA62" bold="true" box="[398,494,1433,1459]" italics="true" pageId="62" pageNumber="63">proavus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24F57D62907FFCFFE62C0B49F6CFA62" box="[502,568,1433,1459]" pageId="62" pageNumber="63" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BB736BF2907FFCFFF03C0919C38FA05" blockId="62.[151,568,1433,1493]" box="[151,364,1468,1493]" pageId="62" pageNumber="63">
(
<figureCitation id="13332A3A2907FFCFFF0BC0919DA1FA05" box="[159,245,1468,1493]" captionStart="FIGURES 21 34" captionStartId="8.[151,264,1436,1459]" captionTargetBox="[163,1421,202,1404]" captionTargetId="figure@8.[151,1436,193,1415]" captionTargetPageId="8" captionText="FIGURES 21 34. Specimens of Rovno amber with inclusions of Scydmaeninae. K- 8972, Rovnoscydmus sp. specimen 8 (21); K- 3322, Rovnoscydmus sp. specimen 9 (22); K- 15548,? Rovnoscydmus sp. specimen 10 (23); K- 8971,? Rovnoscydmus sp. specimen 11 (24); K- 6045, Euconnus palaeogenus sp. n. (25); K- 9126, Euconnus sp. 1 (26); K- 24859,? Euconnus sp. 2 (27); K- 24302, Stenichnus proavus sp. n. (28); UA- 2216, Microscydmus sp. 1 (29); K- 6943,? Microscydmus sp. 2 (30); K- 3657 a,? Microscydmus sp. 3 (31); K- 6764,? Scydmoraphes sp. 1 (32); K- 6274, Glandulariini gen. &amp; sp. indet. 1 (33); K- 7089, Glandulariini gen. &amp; sp. indet. 2 (34)." httpUri="https://zenodo.org/record/262415/files/figure.png" pageId="62" pageNumber="63">Figs 28</figureCitation>
,
<figureCitation id="13332A3A2907FFCFFE95C0919C31FA05" box="[257,357,1468,1492]" captionStart-0="FIGURES 193 196" captionStart-1="FIGURES 197 198" captionStartId-0="64.[151,264,1956,1979]" captionStartId-1="65.[151,264,1286,1309]" captionTargetBox-0="[171,1404,193,1925]" captionTargetBox-1="[160,1425,202,1258]" captionTargetId-0="figure@64.[171,1405,193,1927]" captionTargetId-1="figure@65.[151,1435,193,1265]" captionTargetPageId-0="64" captionTargetPageId-1="65" captionText-0="FIGURES 193 196. Holotype of Stenichnus proavus sp. n. (K- 24302). Dorsolateral (193), right lateral (194), left lateroventral (195) and nearly anterior (196) views." captionText-1="FIGURES 197 198. Holotype of Stenichnus proavus sp. n. (K- 24302). Dorsolateral (197) and ventrolateral (198) views. Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral fovea; lbr, labrum; mtvp, metaventral intercoxal process; mxp 2 4, maxillary palpomere II IV; pg, pygidium; v 3, metaventrite." httpUri-0="https://zenodo.org/record/262456/files/figure.png" httpUri-1="https://zenodo.org/record/262457/files/figure.png" pageId="62" pageNumber="63">193198</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31265342907FFCFFF03C3299831F991" pageId="62" pageNumber="63" type="materials_examined">
<paragraph id="8BB736BF2907FFCFFF03C3299831F991" blockId="62.[151,1437,1540,2033]" pageId="62" pageNumber="63">
<materialsCitation id="3B603CE22907FFCFFF03C3299831F991" ID-GBIF-Occurrence="1300093493" collectionCode="SIZK" location="Europe" municipality="Late Eocene" pageId="62" pageNumber="63" specimenCount="2" stateProvince="Rovno" typeStatus="holotype">
<emphasis id="B97CEAAD2907FFCFFF03C3299E83F9CC" bold="true" box="[151,983,1540,1565]" pageId="62" pageNumber="63">
<typeStatus id="54B3881D2907FFCFFF03C3299D86F9CC" box="[151,210,1540,1565]" pageId="62" pageNumber="63">Type</typeStatus>
material.
<typeStatus id="54B3881D2907FFCFFEC8C3299C99F9CC" box="[348,461,1540,1565]" pageId="62" pageNumber="63" type="holotype">Holotype</typeStatus>
:
<collectingMunicipality id="6BD3ACC52907FFCFFE76C3299F29F9CC" box="[482,637,1540,1565]" pageId="62" pageNumber="63">Late Eocene</collectingMunicipality>
of
<location id="8ED760642907FFCFFD20C3299E5AF9CC" LSID="urn:lsid:plazi:treatment:03A187A92907FFB0FF03C0B49ED7F933:8ED760642907FFCFFD20C3299E5AF9CC" box="[692,782,1540,1565]" municipality="Late Eocene" name="Europe" pageId="62" pageNumber="63" stateProvince="Rovno">Europe</location>
,
<collectingRegion id="49CCF85D2907FFCFFCB7C3299E25F9CC" box="[803,881,1540,1565]" country="Ukraine" name="Rivne" pageId="62" pageNumber="63">Rovno</collectingRegion>
amber:
</emphasis>
sex unknown; inclusion in elongate, subtriangular piece of amber
<quantity id="4CF09B5A2907FFCFFE4BC3059F74F991" box="[479,544,1576,1600]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="62" pageNumber="63" unit="mm" value="9.0">9 mm</quantity>
long with truncated corners (
<figureCitation id="13332A3A2907FFCFFCFDC3059EE8F991" box="[873,956,1576,1601]" captionStart="FIGURES 21 34" captionStartId="8.[151,264,1436,1459]" captionTargetBox="[163,1421,202,1404]" captionTargetId="figure@8.[151,1436,193,1415]" captionTargetPageId="8" captionText="FIGURES 21 34. Specimens of Rovno amber with inclusions of Scydmaeninae. K- 8972, Rovnoscydmus sp. specimen 8 (21); K- 3322, Rovnoscydmus sp. specimen 9 (22); K- 15548,? Rovnoscydmus sp. specimen 10 (23); K- 8971,? Rovnoscydmus sp. specimen 11 (24); K- 6045, Euconnus palaeogenus sp. n. (25); K- 9126, Euconnus sp. 1 (26); K- 24859,? Euconnus sp. 2 (27); K- 24302, Stenichnus proavus sp. n. (28); UA- 2216, Microscydmus sp. 1 (29); K- 6943,? Microscydmus sp. 2 (30); K- 3657 a,? Microscydmus sp. 3 (31); K- 6764,? Scydmoraphes sp. 1 (32); K- 6274, Glandulariini gen. &amp; sp. indet. 1 (33); K- 7089, Glandulariini gen. &amp; sp. indet. 2 (34)." httpUri="https://zenodo.org/record/262415/files/figure.png" pageId="62" pageNumber="63">Fig. 28</figureCitation>
), collection number K-24302 (
<collectionCode id="ED19AE7A2907FFCFFA81C305980DF991" box="[1301,1369,1576,1600]" country="Ukraine" httpUri="http://grbio.org/cool/c442-9tdv" name="Schmaulhausen Institute of Zoology" pageId="62" pageNumber="63">SIZK</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C31265342907FFCFFF53C3619EBBF959" pageId="62" pageNumber="63" type="diagnosis">
<paragraph id="8BB736BF2907FFCFFF53C3619EBBF959" blockId="62.[151,1437,1540,2033]" pageId="62" pageNumber="63">
<emphasis id="B97CEAAD2907FFCFFF53C3619C15F9B4" bold="true" box="[199,321,1612,1637]" pageId="62" pageNumber="63">Diagnosis.</emphasis>
<taxonomicName id="4C084D3C2907FFCFFED8C3619C90F9B4" box="[332,452,1612,1637]" class="Insecta" family="Staphylinidae" genus="Stenichnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B97CEAAD2907FFCFFED8C3619C90F9B4" box="[332,452,1612,1637]" italics="true" pageId="62" pageNumber="63">Stenichnus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24F57D62907FFCFFE5BC3609F5AF9B5" box="[463,526,1613,1636]" pageId="62" pageNumber="63" sensu="stricto">s. str.</taxonomicNameLabel>
with pronotal base bearing indistinct median transverse groove or impression flanked at each side by three or four small pits; BL about
<quantity id="4CF09B5A2907FFCFFC88C35D9ED7F959" box="[796,899,1648,1673]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" pageId="62" pageNumber="63" unit="mm" value="1.45">1.45 mm</quantity>
; EI 1.48..
</paragraph>
</subSubSection>
<subSubSection id="C31265342907FFCEFF53C3B99E37FE59" lastPageId="63" lastPageNumber="64" pageId="62" pageNumber="63" type="description">
<paragraph id="8BB736BF2907FFCFFF53C3B99D9EF901" blockId="62.[151,1437,1540,2033]" pageId="62" pageNumber="63">
<emphasis id="B97CEAAD2907FFCFFF53C3B99C0DF97C" bold="true" box="[199,345,1684,1709]" pageId="62" pageNumber="63">Description.</emphasis>
Body (
<figureCitation id="13332A3A2907FFCFFE3EC3B99F13F97D" box="[426,583,1684,1709]" captionStart-0="FIGURES 193 196" captionStart-1="FIGURES 197 198" captionStartId-0="64.[151,264,1956,1979]" captionStartId-1="65.[151,264,1286,1309]" captionTargetBox-0="[171,1404,193,1925]" captionTargetBox-1="[160,1425,202,1258]" captionTargetId-0="figure@64.[171,1405,193,1927]" captionTargetId-1="figure@65.[151,1435,193,1265]" captionTargetPageId-0="64" captionTargetPageId-1="65" captionText-0="FIGURES 193 196. Holotype of Stenichnus proavus sp. n. (K- 24302). Dorsolateral (193), right lateral (194), left lateroventral (195) and nearly anterior (196) views." captionText-1="FIGURES 197 198. Holotype of Stenichnus proavus sp. n. (K- 24302). Dorsolateral (197) and ventrolateral (198) views. Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral fovea; lbr, labrum; mtvp, metaventral intercoxal process; mxp 2 4, maxillary palpomere II IV; pg, pygidium; v 3, metaventrite." httpUri-0="https://zenodo.org/record/262456/files/figure.png" httpUri-1="https://zenodo.org/record/262457/files/figure.png" pageId="62" pageNumber="63">Figs 193198</figureCitation>
) elongate, relatively stout and strongly convex, nearly black; BL about
<quantity id="4CF09B5A2907FFCFFAFFC3B99D92F901" metricMagnitude="-3" metricUnit="m" metricValue="1.45" pageId="62" pageNumber="63" unit="mm" value="1.45">1.45 mm</quantity>
.
</paragraph>
<paragraph id="8BB736BF2907FFCFFF53C3F19C62F879" blockId="62.[151,1437,1540,2033]" pageId="62" pageNumber="63">
Head (
<figureCitation id="13332A3A2907FFCFFE85C3F19CF9F925" box="[273,429,1756,1781]" captionStart-0="FIGURES 193 196" captionStart-1="FIGURES 197 198" captionStartId-0="64.[151,264,1956,1979]" captionStartId-1="65.[151,264,1286,1309]" captionTargetBox-0="[171,1404,193,1925]" captionTargetBox-1="[160,1425,202,1258]" captionTargetId-0="figure@64.[171,1405,193,1927]" captionTargetId-1="figure@65.[151,1435,193,1265]" captionTargetPageId-0="64" captionTargetPageId-1="65" captionText-0="FIGURES 193 196. Holotype of Stenichnus proavus sp. n. (K- 24302). Dorsolateral (193), right lateral (194), left lateroventral (195) and nearly anterior (196) views." captionText-1="FIGURES 197 198. Holotype of Stenichnus proavus sp. n. (K- 24302). Dorsolateral (197) and ventrolateral (198) views. Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral fovea; lbr, labrum; mtvp, metaventral intercoxal process; mxp 2 4, maxillary palpomere II IV; pg, pygidium; v 3, metaventrite." httpUri-0="https://zenodo.org/record/262456/files/figure.png" httpUri-1="https://zenodo.org/record/262457/files/figure.png" pageId="62" pageNumber="63">Figs 193198</figureCitation>
) broadest at eyes, HL about
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, HW about
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; vertex and frons confluent but poorly visible; compound eyes large, strongly convex, oval; tempora seem very short. Punctures and setae on frons and vertex poorly visible. Antennae (
<figureCitation id="13332A3A2907FFCFFDD4C2099F8AF8ED" box="[576,734,1828,1853]" captionStart-0="FIGURES 193 196" captionStart-1="FIGURES 197 198" captionStartId-0="64.[151,264,1956,1979]" captionStartId-1="65.[151,264,1286,1309]" captionTargetBox-0="[171,1404,193,1925]" captionTargetBox-1="[160,1425,202,1258]" captionTargetId-0="figure@64.[171,1405,193,1927]" captionTargetId-1="figure@65.[151,1435,193,1265]" captionTargetPageId-0="64" captionTargetPageId-1="65" captionText-0="FIGURES 193 196. Holotype of Stenichnus proavus sp. n. (K- 24302). Dorsolateral (193), right lateral (194), left lateroventral (195) and nearly anterior (196) views." captionText-1="FIGURES 197 198. Holotype of Stenichnus proavus sp. n. (K- 24302). Dorsolateral (197) and ventrolateral (198) views. Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral fovea; lbr, labrum; mtvp, metaventral intercoxal process; mxp 2 4, maxillary palpomere II IV; pg, pygidium; v 3, metaventrite." httpUri-0="https://zenodo.org/record/262456/files/figure.png" httpUri-1="https://zenodo.org/record/262457/files/figure.png" pageId="62" pageNumber="63">Figs 193198</figureCitation>
) nearly as short as 1/3 BL, gradually thickened distally, AnL about
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, five proximal antennomeres elongate, distal flagellomeres poorly visible, but most likely not elongate, antennomere XI about as long as IXX together or slightly shorter, about twice as long as broad, with rounded apex.
</paragraph>
<paragraph id="8BB736BF2907FFCEFF53C2989E0EFF29" blockId="62.[151,1437,1540,2033]" lastBlockId="63.[151,1437,151,717]" lastPageId="63" lastPageNumber="64" pageId="62" pageNumber="63">
Pronotum (
<figureCitation id="13332A3A2907FFCFFED3C2999CB3F81D" box="[327,487,1972,1997]" captionStart="FIGURES 193 196" captionStartId="64.[151,264,1956,1979]" captionTargetBox="[171,1404,193,1925]" captionTargetId="figure@64.[171,1405,193,1927]" captionTargetPageId="64" captionText="FIGURES 193 196. Holotype of Stenichnus proavus sp. n. (K- 24302). Dorsolateral (193), right lateral (194), left lateroventral (195) and nearly anterior (196) views." httpUri="https://zenodo.org/record/262456/files/figure.png" pageId="62" pageNumber="63">Figs 193194</figureCitation>
,
<figureCitation id="13332A3A2907FFCFFE61C2999F74F81D" box="[501,544,1972,1996]" captionStart="FIGURES 197 198" captionStartId="65.[151,264,1286,1309]" captionTargetBox="[160,1425,202,1258]" captionTargetId="figure@65.[151,1435,193,1265]" captionTargetPageId="65" captionText="FIGURES 197 198. Holotype of Stenichnus proavus sp. n. (K- 24302). Dorsolateral (197) and ventrolateral (198) views. Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral fovea; lbr, labrum; mtvp, metaventral intercoxal process; mxp 2 4, maxillary palpomere II IV; pg, pygidium; v 3, metaventrite." httpUri="https://zenodo.org/record/262457/files/figure.png" pageId="62" pageNumber="63">197</figureCitation>
) broadest in anterior half; PL
<quantity id="4CF09B5A2907FFCFFC17C2999EB8F81D" box="[899,1004,1972,1997]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="62" pageNumber="63" unit="mm" value="0.35">0.35 mm</quantity>
, PW
<quantity id="4CF09B5A2907FFCFFBBAC29999C3F81D" box="[1070,1175,1972,1996]" metricMagnitude="-4" metricUnit="m" metricValue="3.8" pageId="62" pageNumber="63" unit="mm" value="0.38">0.38 mm</quantity>
; anterior and posterior margins arcuate, lateral margins sinuate, anterior and posterior pronotal corners blunt; base with shallow and diffused transverse antebasal groove (
<figureCitation id="13332A3A2906FFCEFDDEC5BA9FFAFF61" box="[586,686,151,176]" captionStart="FIGURES 197 198" captionStartId="65.[151,264,1286,1309]" captionTargetBox="[160,1425,202,1258]" captionTargetId="figure@65.[151,1435,193,1265]" captionTargetPageId="65" captionText="FIGURES 197 198. Holotype of Stenichnus proavus sp. n. (K- 24302). Dorsolateral (197) and ventrolateral (198) views. Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral fovea; lbr, labrum; mtvp, metaventral intercoxal process; mxp 2 4, maxillary palpomere II IV; pg, pygidium; v 3, metaventrite." httpUri="https://zenodo.org/record/262457/files/figure.png" pageId="63" pageNumber="64">Fig. 197</figureCitation>
;
<emphasis id="B97CEAAD2906FFCEFD28C5BA9FB2FF61" box="[700,742,151,176]" italics="true" pageId="63" pageNumber="64">abg</emphasis>
) and lateral groups of 34 small and shallow antebasal pits (
<figureCitation id="13332A3A2906FFCEFF0BC5919C57FF05" box="[159,259,188,213]" captionStart="FIGURES 197 198" captionStartId="65.[151,264,1286,1309]" captionTargetBox="[160,1425,202,1258]" captionTargetId="figure@65.[151,1435,193,1265]" captionTargetPageId="65" captionText="FIGURES 197 198. Holotype of Stenichnus proavus sp. n. (K- 24302). Dorsolateral (197) and ventrolateral (198) views. Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral fovea; lbr, labrum; mtvp, metaventral intercoxal process; mxp 2 4, maxillary palpomere II IV; pg, pygidium; v 3, metaventrite." httpUri="https://zenodo.org/record/262457/files/figure.png" pageId="63" pageNumber="64">Fig. 197</figureCitation>
;
<emphasis id="B97CEAAD2906FFCEFE86C5919C68FF04" box="[274,316,188,213]" italics="true" pageId="63" pageNumber="64">abp</emphasis>
). Punctures on pronotal disc small but distinct (
<figureCitation id="13332A3A2906FFCEFCF9C5919E85FF05" box="[877,977,188,213]" captionStart="FIGURES 197 198" captionStartId="65.[151,264,1286,1309]" captionTargetBox="[160,1425,202,1258]" captionTargetId="figure@65.[151,1435,193,1265]" captionTargetPageId="65" captionText="FIGURES 197 198. Holotype of Stenichnus proavus sp. n. (K- 24302). Dorsolateral (197) and ventrolateral (198) views. Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral fovea; lbr, labrum; mtvp, metaventral intercoxal process; mxp 2 4, maxillary palpomere II IV; pg, pygidium; v 3, metaventrite." httpUri="https://zenodo.org/record/262457/files/figure.png" pageId="63" pageNumber="64">Fig. 197</figureCitation>
), separated by spaces 23 × as wide as diameters of punctures; setae (
<figureCitation id="13332A3A2906FFCEFE78C5F29F1FFF29" box="[492,587,223,248]" captionStart="FIGURES 197 198" captionStartId="65.[151,264,1286,1309]" captionTargetBox="[160,1425,202,1258]" captionTargetId="figure@65.[151,1435,193,1265]" captionTargetPageId="65" captionText="FIGURES 197 198. Holotype of Stenichnus proavus sp. n. (K- 24302). Dorsolateral (197) and ventrolateral (198) views. Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral fovea; lbr, labrum; mtvp, metaventral intercoxal process; mxp 2 4, maxillary palpomere II IV; pg, pygidium; v 3, metaventrite." httpUri="https://zenodo.org/record/262457/files/figure.png" pageId="63" pageNumber="64">Fig. 197</figureCitation>
) short, sparse, suberect.
</paragraph>
<paragraph id="8BB736BF2906FFCEFF53C429983CFEB4" blockId="63.[151,1437,151,717]" pageId="63" pageNumber="64">
Elytra (
<figureCitation id="13332A3A2906FFCEFE88C4299CEFFECD" box="[284,443,260,285]" captionStart="FIGURES 193 196" captionStartId="64.[151,264,1956,1979]" captionTargetBox="[171,1404,193,1925]" captionTargetId="figure@64.[171,1405,193,1927]" captionTargetPageId="64" captionText="FIGURES 193 196. Holotype of Stenichnus proavus sp. n. (K- 24302). Dorsolateral (193), right lateral (194), left lateroventral (195) and nearly anterior (196) views." httpUri="https://zenodo.org/record/262456/files/figure.png" pageId="63" pageNumber="64">Figs 193194</figureCitation>
,
<figureCitation id="13332A3A2906FFCEFE5DC4299CA0FECD" box="[457,500,260,284]" captionStart="FIGURES 197 198" captionStartId="65.[151,264,1286,1309]" captionTargetBox="[160,1425,202,1258]" captionTargetId="figure@65.[151,1435,193,1265]" captionTargetPageId="65" captionText="FIGURES 197 198. Holotype of Stenichnus proavus sp. n. (K- 24302). Dorsolateral (197) and ventrolateral (198) views. Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral fovea; lbr, labrum; mtvp, metaventral intercoxal process; mxp 2 4, maxillary palpomere II IV; pg, pygidium; v 3, metaventrite." httpUri="https://zenodo.org/record/262457/files/figure.png" pageId="63" pageNumber="64">197</figureCitation>
) elongate oval, strongly convex, broadest most likely in front of middle, EL
<quantity id="4CF09B5A2906FFCEFAFFC4299D91FEEE" metricMagnitude="-4" metricUnit="m" metricValue="8.5" pageId="63" pageNumber="64" unit="mm" value="0.85">0.85 mm</quantity>
, EW
<quantity id="4CF09B5A2906FFCEFE91C40A9C3FFEEE" box="[261,363,295,320]" metricMagnitude="-4" metricUnit="m" metricValue="5.8" pageId="63" pageNumber="64" unit="mm" value="0.58">0.58 mm</quantity>
, EI 1.48; punctures (
<figureCitation id="13332A3A2906FFCEFDCCC40A9FECFE91" box="[600,696,295,320]" captionStart="FIGURES 197 198" captionStartId="65.[151,264,1286,1309]" captionTargetBox="[160,1425,202,1258]" captionTargetId="figure@65.[151,1435,193,1265]" captionTargetPageId="65" captionText="FIGURES 197 198. Holotype of Stenichnus proavus sp. n. (K- 24302). Dorsolateral (197) and ventrolateral (198) views. Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral fovea; lbr, labrum; mtvp, metaventral intercoxal process; mxp 2 4, maxillary palpomere II IV; pg, pygidium; v 3, metaventrite." httpUri="https://zenodo.org/record/262457/files/figure.png" pageId="63" pageNumber="64">Fig. 197</figureCitation>
) slightly larger but otherwise similar to those on pronotum; setae (
<figureCitation id="13332A3A2906FFCEFF0BC4619DABFEB5" box="[159,255,332,357]" captionStart="FIGURES 197 198" captionStartId="65.[151,264,1286,1309]" captionTargetBox="[160,1425,202,1258]" captionTargetId="figure@65.[151,1435,193,1265]" captionTargetPageId="65" captionText="FIGURES 197 198. Holotype of Stenichnus proavus sp. n. (K- 24302). Dorsolateral (197) and ventrolateral (198) views. Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral fovea; lbr, labrum; mtvp, metaventral intercoxal process; mxp 2 4, maxillary palpomere II IV; pg, pygidium; v 3, metaventrite." httpUri="https://zenodo.org/record/262457/files/figure.png" pageId="63" pageNumber="64">Fig. 197</figureCitation>
) similar to those on pronotum. Scutellum large, subtriangular and well visible between elytral bases.
</paragraph>
<paragraph id="8BB736BF2906FFCEFF53C45D9E37FE59" blockId="63.[151,1437,151,717]" box="[199,867,367,392]" pageId="63" pageNumber="64">
Legs (
<figureCitation id="13332A3A2906FFCEFE98C4429CFCFE59" box="[268,424,367,392]" captionStart-0="FIGURES 193 196" captionStart-1="FIGURES 197 198" captionStartId-0="64.[151,264,1956,1979]" captionStartId-1="65.[151,264,1286,1309]" captionTargetBox-0="[171,1404,193,1925]" captionTargetBox-1="[160,1425,202,1258]" captionTargetId-0="figure@64.[171,1405,193,1927]" captionTargetId-1="figure@65.[151,1435,193,1265]" captionTargetPageId-0="64" captionTargetPageId-1="65" captionText-0="FIGURES 193 196. Holotype of Stenichnus proavus sp. n. (K- 24302). Dorsolateral (193), right lateral (194), left lateroventral (195) and nearly anterior (196) views." captionText-1="FIGURES 197 198. Holotype of Stenichnus proavus sp. n. (K- 24302). Dorsolateral (197) and ventrolateral (198) views. Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral fovea; lbr, labrum; mtvp, metaventral intercoxal process; mxp 2 4, maxillary palpomere II IV; pg, pygidium; v 3, metaventrite." httpUri-0="https://zenodo.org/record/262456/files/figure.png" httpUri-1="https://zenodo.org/record/262457/files/figure.png" pageId="63" pageNumber="64">Figs 193198</figureCitation>
) moderately long, slender, unmodified.
</paragraph>
</subSubSection>
<subSubSection id="C31265342906FFCEFF53C4B998CEFE7C" box="[199,1434,404,429]" pageId="63" pageNumber="64" type="etymology">
<paragraph id="8BB736BF2906FFCEFF53C4B998CEFE7C" blockId="63.[151,1437,151,717]" box="[199,1434,404,429]" pageId="63" pageNumber="64">
<emphasis id="B97CEAAD2906FFCEFF53C4B99C1AFE7C" bold="true" box="[199,334,404,429]" pageId="63" pageNumber="64">Etymology.</emphasis>
The name
<taxonomicName id="4C084D3C2906FFCEFE5FC4B89F70FE7D" box="[459,548,405,428]" class="Insecta" family="Staphylinidae" genus="Stenichnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="proavus">
<emphasis id="B97CEAAD2906FFCEFE5FC4B89F70FE7D" box="[459,548,405,428]" italics="true" pageId="63" pageNumber="64">proavus</emphasis>
</taxonomicName>
(a noun in apposition) means &quot;great grandfather, forefather, ancestor&quot; in Latin.
</paragraph>
</subSubSection>
<subSubSection id="C31265342906FFB0FF53C49A9ED7F933" lastPageId="65" lastPageNumber="66" pageId="63" pageNumber="64" type="discussion">
<paragraph id="8BB736BF2906FFB0FF53C49A997EFA36" blockId="63.[151,1437,151,717]" lastBlockId="65.[151,1436,1414,1763]" lastPageId="65" lastPageNumber="66" pageId="63" pageNumber="64">
<emphasis id="B97CEAAD2906FFCEFF53C49A9C6FFE01" bold="true" box="[199,315,439,464]" pageId="63" pageNumber="64">Remarks.</emphasis>
Placement of this species in the extant genus
<taxonomicName id="4C084D3C2906FFCEFCECC49A9EA4FE01" box="[888,1008,439,464]" class="Insecta" family="Staphylinidae" genus="Stenichnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="genus">
<emphasis id="B97CEAAD2906FFCEFCECC49A9EA4FE01" box="[888,1008,439,464]" italics="true" pageId="63" pageNumber="64">Stenichnus</emphasis>
</taxonomicName>
is not strongly supported, as some morphological characters in the studied specimen are obscured by turbid amber or hidden under declined head. However, the general body form, the shape and antebasal structures of the pronotum, absent lateral pronotal carinae and sublateral pronotal carinae, the gradually thickened antennae, the 'wooly' setae forming a fringe along posterior margins of the mesocoxal cavities and the large scutellum exposed between elytral bases in combination with the metacoxae touching the lateral margins of metaventrite and the narrow metaventral intercoxal process indicate that this is either a
<taxonomicName id="4C084D3C2906FFCEFED6C7A29CEEFD79" box="[322,442,655,680]" class="Insecta" family="Staphylinidae" genus="Stenichnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="genus">
<emphasis id="B97CEAAD2906FFCEFED6C7A29CEEFD79" box="[322,442,655,680]" italics="true" pageId="63" pageNumber="64">Stenichnus</emphasis>
</taxonomicName>
or a genus very close to
<taxonomicName id="4C084D3C2906FFCEFD4AC7A29E02FD79" box="[734,854,655,680]" class="Insecta" family="Staphylinidae" genus="Stenichnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="genus">
<emphasis id="B97CEAAD2906FFCEFD4AC7A29E02FD79" box="[734,854,655,680]" italics="true" pageId="63" pageNumber="64">Stenichnus</emphasis>
</taxonomicName>
. The arrangement of the antebasal groove and pits make this species easily identifiable, and its generic placement will be possible to verify if new specimens become available in future. If the placement in
<taxonomicName id="4C084D3C2978FFB0FDDEC0AB9F96FA4E" box="[586,706,1414,1439]" class="Insecta" family="Staphylinidae" genus="Stenichnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis id="B97CEAAD2978FFB0FDDEC0AB9F96FA4E" box="[586,706,1414,1439]" italics="true" pageId="65" pageNumber="66">Stenichnus</emphasis>
</taxonomicName>
is accepted, then the presence of one well-visible and large basal elytral fovea adjacent to the scutellum indicates that this is the nominotypical subgenus;
<taxonomicName id="4C084D3C2978FFB0FAB0C08798C8FA12" box="[1316,1436,1450,1475]" class="Insecta" family="Staphylinidae" genus="Stenichnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis id="B97CEAAD2978FFB0FAB0C08798C8FA12" box="[1316,1436,1450,1475]" italics="true" pageId="65" pageNumber="66">Stenichnus</emphasis>
</taxonomicName>
(
<taxonomicName id="4C084D3C2978FFB0FF34C0E39CA0FA36" authority="Reitter, 1905" authorityName="Reitter" authorityYear="1905" box="[160,500,1486,1511]" class="Insecta" family="Staphylinidae" genus="Scydmaenichnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis id="B97CEAAD2978FFB0FF34C0E39C0FFA36" box="[160,347,1486,1511]" italics="true" pageId="65" pageNumber="66">Scydmaenichnus</emphasis>
<bibRefCitation id="EF994B4E2978FFB0FEF6C0E39CA0FA36" author="Reitter" box="[354,500,1486,1511]" pageId="65" pageNumber="66" refString="Reitter, E. (1905) Uber die Stenichnus - Arten von der Insel Cypern (Cyrtoscydmus Motsch., Coleoptera.). Wiener Entomologische Zeitung, 24, 98 - 99." type="journal article" year="1905">Reitter, 1905</bibRefCitation>
</taxonomicName>
) has two rudimentary foveae (
<bibRefCitation id="EF994B4E2978FFB0FCDEC0E3994AFA36" author="Jaloszynski" box="[842,1054,1486,1511]" pageId="65" pageNumber="66" refString="Jaloszynski, P. (2013 b) Revision of subgenera of Stenichnus Thomson, with review of Australo-Pacific species (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa, 3630 (1), 39 - 79. http: // dx. doi. org / 10.11646 / zootaxa. 3630.1.2" type="journal article" year="2013" yearSuffix="b">Jałoszyński 2013b</bibRefCitation>
).
</paragraph>
<caption id="DF7766372906FFCEFF03C2AB982CF80B" httpUri="https://zenodo.org/record/262455/files/figure.png" pageId="63" pageNumber="64" targetBox="[169,1426,779,1894]" targetPageId="63">
<paragraph id="8BB736BF2906FFCEFF03C2AB982CF80B" blockId="63.[151,1437,1926,2010]" pageId="63" pageNumber="64">
<emphasis id="B97CEAAD2906FFCEFF03C2AB9C3BF84D" bold="true" box="[151,367,1926,1949]" pageId="63" pageNumber="64">FIGURES 191192.</emphasis>
?
<taxonomicName id="4C084D3C2906FFCEFE16C2A59CB2F84C" box="[386,486,1928,1949]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="genus">
<emphasis id="B97CEAAD2906FFCEFE16C2A59CB2F84C" box="[386,486,1928,1949]" italics="true" pageId="63" pageNumber="64">Euconnus</emphasis>
</taxonomicName>
sp. 2 (K-24859). Right ventrolateral (191) and left dorsolateral (192) views. Abbreviations: abp, antebasal pit; bef, basal elytral fovea; bst, basisternal part of prosternum; cl, clypeus; fr, frons; hyr, hypomeral ridge; lbr, labrum; mcp, mesocoxal projection; md, mandible; mxp34, maxillary palpomere IIIIV; slc, sublateral carina; tm, temple.
</paragraph>
</caption>
<caption id="DF7766372979FFB1FF03C2899F4DF808" httpUri="https://zenodo.org/record/262456/files/figure.png" pageId="64" pageNumber="65" targetBox="[171,1404,193,1925]" targetPageId="64">
<paragraph id="8BB736BF2979FFB1FF03C2899F4DF808" blockId="64.[151,1436,1956,2009]" pageId="64" pageNumber="65">
<emphasis id="B97CEAAD2979FFB1FF03C2899C39F868" bold="true" box="[151,365,1956,1979]" pageId="64" pageNumber="65">FIGURES 193196.</emphasis>
Holotype of
<taxonomicName id="4C084D3C2979FFB1FE65C2899FE1F86B" box="[497,693,1956,1978]" class="Insecta" family="Staphylinidae" genus="Stenichnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="proavus" status="sp. nov.">
<emphasis id="B97CEAAD2979FFB1FE65C2899FE1F86B" box="[497,693,1956,1978]" italics="true" pageId="64" pageNumber="65">Stenichnus proavus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24F57D62979FFB1FD2FC2889FA7F86B" box="[699,755,1957,1978]" pageId="64" pageNumber="65" rank="species">
<emphasis id="B97CEAAD2979FFB1FD2FC2889FA7F86B" bold="true" box="[699,755,1957,1978]" pageId="64" pageNumber="65">sp. n.</emphasis>
</taxonomicNameLabel>
(K-24302). Dorsolateral (193), right lateral (194), left lateroventral (195) and nearly anterior (196) views.
</paragraph>
</caption>
<caption id="DF7766372978FFB0FF03C02B9EF8FA8B" httpUri="https://zenodo.org/record/262457/files/figure.png" pageId="65" pageNumber="66" targetBox="[160,1425,202,1258]" targetPageId="65">
<paragraph id="8BB736BF2978FFB0FF03C02B9EF8FA8B" blockId="65.[151,1437,1286,1370]" pageId="65" pageNumber="66">
<emphasis id="B97CEAAD2978FFB0FF03C02B9C25FACD" bold="true" box="[151,369,1286,1309]" pageId="65" pageNumber="66">FIGURES 197198.</emphasis>
Holotype of
<taxonomicName id="4C084D3C2978FFB0FD97C02B9F9FFACC" box="[515,715,1286,1309]" class="Insecta" family="Staphylinidae" genus="Stenichnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="proavus" status="sp. nov.">
<emphasis id="B97CEAAD2978FFB0FD97C02B9F9FFACC" box="[515,715,1286,1309]" italics="true" pageId="65" pageNumber="66">Stenichnus proavus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24F57D62978FFB0FD41C02A9E46FACD" box="[725,786,1287,1308]" pageId="65" pageNumber="66" rank="species">
<emphasis id="B97CEAAD2978FFB0FD41C02A9E46FACD" bold="true" box="[725,786,1287,1308]" pageId="65" pageNumber="66">sp. n.</emphasis>
</taxonomicNameLabel>
(K-24302). Dorsolateral (197) and ventrolateral (198) views. Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral fovea; lbr, labrum; mtvp, metaventral intercoxal process; mxp24, maxillary palpomere IIIV; pg, pygidium; v3, metaventrite.
</paragraph>
</caption>
<paragraph id="8BB736BF2978FFB0FF53C0DE9ED7F933" blockId="65.[151,1436,1414,1763]" pageId="65" pageNumber="66">
Most extant species of
<taxonomicName id="4C084D3C2978FFB0FE53C0DF9F14F9DA" box="[455,576,1522,1547]" class="Insecta" family="Staphylinidae" genus="Stenichnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis id="B97CEAAD2978FFB0FE53C0DF9F14F9DA" box="[455,576,1522,1547]" italics="true" pageId="65" pageNumber="66">Stenichnus</emphasis>
</taxonomicName>
have the pronotal base with a transverse row of four or six pits, also a row of large, numerous and irregularly distributed punctures (larger than those on disc) can be found in some representatives of this genus. A transverse impression in addition to pits is unique for
<taxonomicName id="4C084D3C2978FFB0FB08C3169849F982" box="[1180,1309,1595,1619]" class="Insecta" family="Staphylinidae" genus="Stenichnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="proavus">
<emphasis id="B97CEAAD2978FFB0FB08C3169849F982" box="[1180,1309,1595,1619]" italics="true" pageId="65" pageNumber="66">St. proavus</emphasis>
</taxonomicName>
. However, recently a species with the pronotal base with one lateral pair of shallow pits and a pair of short longitudinal grooves near middle composed of diffused and connected punctures was redescribed (
<bibRefCitation id="EF994B4E2978FFB0FB16C3AF9800F94A" author="Jaloszynski" box="[1154,1364,1666,1691]" pageId="65" pageNumber="66" refString="Jaloszynski, P. (2015 i) Scydmoraphes rothangensis Franz, a misplaced Himalayan species of Cyrtoscydmini (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa, 4013 (3), 443 - 448. http: // dx. doi. org / 10.11646 / zootaxa. 4013.3.9" type="journal article" year="2015" yearSuffix="i">
Jałoszyński
<date id="FFB6107F2978FFB0FA84C3AF9800F94A" box="[1296,1364,1666,1691]" pageId="65" pageNumber="66" value="2015-01">2015i</date>
</bibRefCitation>
). This demonstrated that the spectrum of antebasal structures in
<taxonomicName id="4C084D3C2978FFB0FCA0C38B9EF8F96E" box="[820,940,1702,1727]" class="Insecta" family="Staphylinidae" genus="Stenichnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis id="B97CEAAD2978FFB0FCA0C38B9EF8F96E" box="[820,940,1702,1727]" italics="true" pageId="65" pageNumber="66">Stenichnus</emphasis>
</taxonomicName>
is greater than previously thought, and
<taxonomicName id="4C084D3C2978FFB0FA14C38A9DA4F932" class="Insecta" family="Staphylinidae" genus="Stenichnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="proavus">
<emphasis id="B97CEAAD2978FFB0FA14C38A9DA4F932" italics="true" pageId="65" pageNumber="66">St. proavus</emphasis>
</taxonomicName>
is not as unusual with its antebasal groove as it may seem.
</paragraph>
</subSubSection>
</treatment>
</document>