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<mods:title id="9CFF6FA0A4AF21D096E40484DFD194A6">Review of the Malagasy lecithocerid species described by Pierre Viette and deposited in MNHN (Paris), with new generic combinations and descriptions of a new subfamily and genus of Momphidae (Lepidoptera: Gelechioidea)</mods:title>
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<heading id="D0CC7F10FFBAFFD15CE19A01FDEAFA73" bold="true" box="[151,586,1387,1414]" fontSize="11" level="1" pageId="13" pageNumber="164" reason="1">
<taxonomicName id="4C3BB3FFFFBAFFD15CE19A01FDEAFA73" ID-CoL="9FPWQ" authority="(Viette, 1986)" baseAuthorityName="Viette" baseAuthorityYear="1986" box="[151,586,1387,1414]" class="Insecta" family="Lecithoceridae" genus="Torodora" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="kambanella">
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<emphasis id="B94F146EFFBAFFD15CE19A01FE3CFA70" bold="true" box="[151,412,1387,1414]" italics="true" pageId="13" pageNumber="164">Torodora kambanella</emphasis>
(
<bibRefCitation id="EFAAB58DFFBAFFD15DDD9A01FDE1FA73" author="Viette, P. E. L." box="[427,577,1387,1413]" pageId="13" pageNumber="164" pagination="109 - 118" refId="ref24658" refString="Viette, P. E. L. (1986) Nouveaux Microlepidopteres de Madagascar (Oecophoridae et Gelechiidae) (New species of Malagasy Microlepidoptera-Oecophoridae and Gelechiidae). Bulletin mensuel de la Societe linneenne de Lyon, 55 (4), 109 - 118. https: // doi. org / 10.3406 / linly. 1986.10753" type="journal article" year="1986">Viette, 1986</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<paragraph id="8B84C87CFFBAFFD15CE19AFAFE88FA5C" blockId="13.[151,586,1387,1450]" box="[151,296,1424,1450]" pageId="13" pageNumber="164">
(
<figureCitation id="1300D4F9FFBAFFD15CE99AFAFF4CFA5C" box="[159,236,1424,1450]" captionStart="FIGURE 9" captionStartId="14.[151,250,1645,1669]" captionTargetBox="[161,1419,178,1591]" captionTargetId="figure-25@14.[151,1436,166,1622]" captionTargetPageId="14" captionText="FIGURE 9. Torodora kambanella (Viette, 1986):A, holotype; B, labels; C, hind wing (ventral surface); D, male genitalia, prep. P. Viette no. 3810; E, aedeagus; F, abdomen. Scale bar for genitalia and aedeagus: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4406478" httpUri="https://zenodo.org/record/4406478/files/figure.png" pageId="13" pageNumber="164">Figs. 9</figureCitation>
AF)
</paragraph>
</subSubSection>
<subSubSection id="C3219BF7FFBAFFD15CE19ABEFD8EF9FD" pageId="13" pageNumber="164" type="reference_group">
<paragraph id="8B84C87CFFBAFFD15CE19ABEFCDDFA1A" blockId="13.[151,893,1492,1547]" box="[151,893,1492,1516]" pageId="13" pageNumber="164">
<treatmentCitationGroup id="AB2BEF52FFBAFFD15CE19ABEFCDDFA1A" box="[151,893,1492,1516]" pageId="13" pageNumber="164">
<treatmentCitation id="0A9AEE6DFFBAFFD15CE19ABEFDE4FA1A" author="Viette, P. E. L." box="[151,580,1492,1516]" page="114" pageId="13" pageNumber="164" year="1986">
<taxonomicName id="4C3BB3FFFFBAFFD15CE19ABEFDE4FA1A" ID-CoL="9F8BQ" authority="Viette, 1986: 114" authorityName="Viette" authorityPageNumber="114" authorityYear="1986" box="[151,580,1492,1516]" class="Insecta" family="Lecithoceridae" genus="Lecithocera" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="kambanella">
<emphasis id="B94F146EFFBAFFD15CE19ABEFE2DFA1A" box="[151,397,1492,1516]" italics="true" pageId="13" pageNumber="164">Lecithocera kambanella</emphasis>
<bibRefCitation id="EFAAB58DFFBAFFD15DE59ABEFDE4FA1A" author="Viette, P. E. L." box="[403,580,1492,1516]" pageId="13" pageNumber="164" pagination="109 - 118" refId="ref24658" refString="Viette, P. E. L. (1986) Nouveaux Microlepidopteres de Madagascar (Oecophoridae et Gelechiidae) (New species of Malagasy Microlepidoptera-Oecophoridae and Gelechiidae). Bulletin mensuel de la Societe linneenne de Lyon, 55 (4), 109 - 118. https: // doi. org / 10.3406 / linly. 1986.10753" type="journal article" year="1986">Viette, 1986: 114</bibRefCitation>
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. TL: Zahamena National Park.
</treatmentCitationGroup>
</paragraph>
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<treatmentCitationGroup id="AB2BEF52FFBAFFD15CE19A99FD8EF9FD" box="[151,558,1523,1547]" pageId="13" pageNumber="164">
<taxonomicName id="4C3BB3FFFFBAFFD15CE19A99FED1F9FD" ID-CoL="9FPWQ" baseAuthorityName="Viette" baseAuthorityYear="1986" box="[151,369,1523,1547]" class="Insecta" family="Lecithoceridae" genus="Torodora" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="kambanella">
<emphasis id="B94F146EFFBAFFD15CE19A99FED1F9FD" box="[151,369,1523,1547]" italics="true" pageId="13" pageNumber="164">Torodora kambanella</emphasis>
</taxonomicName>
;
<treatmentCitation id="0A9AEE6DFFBAFFD15D099A99FD8AF9FD" author="Park, K. - T." box="[383,554,1523,1547]" page="1085" pageId="13" pageNumber="164" year="2018">
<bibRefCitation id="EFAAB58DFFBAFFD15D099A99FD8AF9FD" author="Park, K. - T." box="[383,554,1523,1547]" pageId="13" pageNumber="164" pagination="1085 - 1093" refId="ref23829" refString="Park, K. - T. (2018) A new genus Thubdora Park, gen. nov. and seven new species of the subfamily Torodorinae (Lepidoptera, Lecithoceridae) from Cameroon and Rep. of Congo, Africa. Journal of Asia-Pacific Entomology, 21, 1085 - 1093. https: // doi. org / 10.1016 / j. aspen. 2018.07.018" type="journal article" year="2018">Park 2018: 1085</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3219BF7FFBAFFD15CE19953FCB2F861" pageId="13" pageNumber="164" type="materials_examined">
<paragraph id="8B84C87CFFBAFFD15CE19953FAD4F981" blockId="13.[151,1437,1593,2016]" pageId="13" pageNumber="164">
<emphasis id="B94F146EFFBAFFD15CE19953FF79F9A5" bold="true" box="[151,217,1593,1619]" pageId="13" pageNumber="164">Type.</emphasis>
<materialsCitation id="3B53C221FFBAFFD15C939953FAD1F981" ID-GBIF-Occurrence="3027957302" collectingDate="1954-11-16" collectorName="P. Viette" country="Madagascar" county="Nosivola" elevation="1000" location="East" municipality="Manakambahiny East" pageId="13" pageNumber="164" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<specimenCount id="9D3D03F5FFBAFFD15C939953FEA8F9A5" box="[229,264,1593,1619]" pageId="13" pageNumber="164" type="male">1♂</specimenCount>
(
<typeStatus id="548076DEFFBAFFD15D6A9953FE24F9A2" box="[284,388,1593,1620]" pageId="13" pageNumber="164" type="holotype">holotype</typeStatus>
),
<location id="8EE49EA7FFBAFFD15DEA9950FE6CF9A2" LSID="urn:lsid:plazi:treatment:0392796AFFBAFFD15CE19A01FD42F816:8EE49EA7FFBAFFD15DEA9950FE6CF9A2" box="[412,460,1594,1620]" country="Madagascar" county="Nosivola" municipality="Manakambahiny East" name="East" pageId="13" pageNumber="164">East</location>
<collectingCountry id="F32C88ECFFBAFFD15DAF9953FDC2F9A5" box="[473,610,1593,1619]" name="Madagascar" pageId="13" pageNumber="164">Madagascar</collectingCountry>
, near 3
<superScript id="7C4E6534FFBAFFD15ECB9953FD6BF9B1" attach="left" box="[701,715,1593,1607]" fontSize="6" pageId="13" pageNumber="164">rd</superScript>
<location id="8EE49EA7FFBAFFD15EA19953FC55F9A2" LSID="urn:lsid:plazi:treatment:0392796AFFBAFFD15CE19A01FD42F816:8EE49EA7FFBAFFD15EA19953FC55F9A2" box="[727,1013,1593,1620]" country="Madagascar" county="Nosivola" municipality="Manakambahiny East" name="Integral Natural Reserve" pageId="13" pageNumber="164">Integral Natural Reserve</location>
[Zahamena National Park], road to
<collectingMunicipality id="6BE05206FFBAFFD15CE19937FE2DF98E" box="[151,397,1629,1656]" pageId="13" pageNumber="164">Manakambahiny East</collectingMunicipality>
, near
<collectingCounty id="62E5B0F0FFBAFFD15DA69937FD98F981" box="[464,568,1629,1655]" pageId="13" pageNumber="164">Nosivola</collectingCounty>
, alt.
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<elevation id="00162F4FFFBAFFD15E1B9937FD6BF98E" box="[621,715,1629,1656]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="13" pageNumber="164" unit="m" value="1000.0">1,000 m</elevation>
</quantity>
,
<date id="FF85EEBCFFBAFFD15EA19937FCF1F98E" box="[727,849,1629,1656]" pageId="13" pageNumber="164" value="1954-11-16">
<collectingDate id="EFC11754FFBAFFD15EA19937FCF1F98E" box="[727,849,1629,1656]" pageId="13" pageNumber="164" value="1954-11-16">16 xi 1954</collectingDate>
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,
<collectorName id="26CEADAAFFBAFFD15F2A9934FC19F981" box="[860,953,1629,1656]" pageId="13" pageNumber="164">P. Viette</collectorName>
leg.; genitalia: prep.
<collectorName id="26CEADAAFFBAFFD158D19934FAA4F981" box="[1191,1284,1629,1656]" pageId="13" pageNumber="164">P. Viette</collectorName>
no. 3810
</materialsCitation>
.
</paragraph>
<paragraph id="8B84C87CFFBAFFD15CB199EBFE05F8F1" blockId="13.[151,1437,1593,2016]" pageId="13" pageNumber="164">
<emphasis id="B94F146EFFBAFFD15CB199EBFEABF96D" bold="true" box="[199,267,1665,1691]" pageId="13" pageNumber="164">Adult</emphasis>
(
<figureCitation id="1300D4F9FFBAFFD15D6B99EBFE04F96A" box="[285,420,1665,1692]" captionStart="FIGURE 9" captionStartId="14.[151,250,1645,1669]" captionTargetBox="[161,1419,178,1591]" captionTargetId="figure-25@14.[151,1436,166,1622]" captionTargetPageId="14" captionText="FIGURE 9. Torodora kambanella (Viette, 1986):A, holotype; B, labels; C, hind wing (ventral surface); D, male genitalia, prep. P. Viette no. 3810; E, aedeagus; F, abdomen. Scale bar for genitalia and aedeagus: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4406478" httpUri="https://zenodo.org/record/4406478/files/figure.png" pageId="13" pageNumber="164">Figs. 9A, B</figureCitation>
). Wingspan,
<quantity id="4CC36599FFBAFFD15E4D99EBFD2EF96A" box="[571,654,1665,1692]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="13" pageNumber="164" unit="mm" value="18.0">18 mm</quantity>
. The species is characterized by the trapezoidal blackish antemedian band discontinued to inner margin, and triangular blackish costal patch developed in 2/3 of costa on the forewing. The hind wing venation has M
<subScript id="17BFCA39FFBAFFD15D9B99B6FE55F91C" attach="left" box="[493,501,1756,1770]" fontSize="6" pageId="13" pageNumber="164">3</subScript>
stalked with CuA
<subScript id="17BFCA39FFBAFFD15EB699B6FD68F91C" attach="left" box="[704,712,1756,1770]" fontSize="6" pageId="13" pageNumber="164">1</subScript>
at distal 1/5 (
<figureCitation id="1300D4F9FFBAFFD15F1799A3FC19F915" box="[865,953,1737,1763]" captionStart="FIGURE 9" captionStartId="14.[151,250,1645,1669]" captionTargetBox="[161,1419,178,1591]" captionTargetId="figure-25@14.[151,1436,166,1622]" captionTargetPageId="14" captionText="FIGURE 9. Torodora kambanella (Viette, 1986):A, holotype; B, labels; C, hind wing (ventral surface); D, male genitalia, prep. P. Viette no. 3810; E, aedeagus; F, abdomen. Scale bar for genitalia and aedeagus: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4406478" httpUri="https://zenodo.org/record/4406478/files/figure.png" pageId="13" pageNumber="164">Fig. 9C</figureCitation>
). The abdomen has broad spinose zones on dorsal surface (
<figureCitation id="1300D4F9FFBAFFD15D359987FE38F8F1" box="[323,408,1773,1799]" captionStart="FIGURE 9" captionStartId="14.[151,250,1645,1669]" captionTargetBox="[161,1419,178,1591]" captionTargetId="figure-25@14.[151,1436,166,1622]" captionTargetPageId="14" captionText="FIGURE 9. Torodora kambanella (Viette, 1986):A, holotype; B, labels; C, hind wing (ventral surface); D, male genitalia, prep. P. Viette no. 3810; E, aedeagus; F, abdomen. Scale bar for genitalia and aedeagus: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4406478" httpUri="https://zenodo.org/record/4406478/files/figure.png" pageId="13" pageNumber="164">Fig. 9F</figureCitation>
).
</paragraph>
<paragraph id="8B84C87CFFBAFFD15CB1987BFCB2F861" blockId="13.[151,1437,1593,2016]" pageId="13" pageNumber="164">
<emphasis id="B94F146EFFBAFFD15CB1987BFECCF8DD" box="[199,364,1809,1835]" italics="true" pageId="13" pageNumber="164">Male genitalia</emphasis>
(
<figureCitation id="1300D4F9FFBAFFD15D0A987BFDA2F8DA" box="[380,514,1809,1836]" captionStart="FIGURE 9" captionStartId="14.[151,250,1645,1669]" captionTargetBox="[161,1419,178,1591]" captionTargetId="figure-25@14.[151,1436,166,1622]" captionTargetPageId="14" captionText="FIGURE 9. Torodora kambanella (Viette, 1986):A, holotype; B, labels; C, hind wing (ventral surface); D, male genitalia, prep. P. Viette no. 3810; E, aedeagus; F, abdomen. Scale bar for genitalia and aedeagus: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4406478" httpUri="https://zenodo.org/record/4406478/files/figure.png" pageId="13" pageNumber="164">Figs. 9D, E</figureCitation>
): See the line drawing given by
<bibRefCitation id="EFAAB58DFFBAFFD15F06987BFC5EF8DD" author="Viette, P. E. L." box="[880,1022,1809,1835]" pageId="13" pageNumber="164" pagination="109 - 118" refId="ref24658" refString="Viette, P. E. L. (1986) Nouveaux Microlepidopteres de Madagascar (Oecophoridae et Gelechiidae) (New species of Malagasy Microlepidoptera-Oecophoridae and Gelechiidae). Bulletin mensuel de la Societe linneenne de Lyon, 55 (4), 109 - 118. https: // doi. org / 10.3406 / linly. 1986.10753" type="journal article" year="1986">Viette (1986</bibRefCitation>
,
<figureCitation id="1300D4F9FFBAFFD1587C987BFBFDF8DD" box="[1034,1117,1809,1835]" captionStart="FIGURE 10" captionStartId="15.[151,250,1833,1857]" captionTargetBox="[163,1419,538,1791]" captionTargetId="figure-164@15.[151,1436,525,1810]" captionTargetPageId="15" captionText="FIGURE 10. Torodora lecithocerella (Viette, 1956), comb. nov.: A, holotype; B, labels; C, female genitalia, prep. J. Minet no. 1663; D, close-up of abdominal segment VIII and ostium bursae; E, close-up of signum; F, abdomen. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4406480" httpUri="https://zenodo.org/record/4406480/files/figure.png" pageId="13" pageNumber="164">Fig. 10</figureCitation>
). Uncus long, elongate, narrowed apically. Basal plate of gnathos elongated with round apex. Valva elongated with spatulate cucullus; cucullus broadened distally with round apex. Aedeagus as long as valva, globular at base; cornuti consist of a long sac with numerous spinules and heavily sclerotized, shorter plate.
</paragraph>
</subSubSection>
<subSubSection id="C3219BF7FFBAFFD15CB198CBFAA8F84D" box="[199,1288,1953,1980]" pageId="13" pageNumber="164" type="discussion">
<paragraph id="8B84C87CFFBAFFD15CB198CBFAA8F84D" blockId="13.[151,1437,1593,2016]" box="[199,1288,1953,1980]" pageId="13" pageNumber="164">
<emphasis id="B94F146EFFBAFFD15CB198CBFE9BF84D" bold="true" box="[199,315,1953,1979]" pageId="13" pageNumber="164">Remarks.</emphasis>
This species was previously transferred from
<taxonomicName id="4C3BB3FFFFBAFFD15F3698CBFC66F84D" authorityName="Herrich-Schaffer" authorityYear="1853" box="[832,966,1953,1979]" class="Insecta" family="Lecithoceridae" genus="Lecithocera" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="164" phylum="Arthropoda" rank="genus">
<emphasis id="B94F146EFFBAFFD15F3698CBFC66F84D" box="[832,966,1953,1979]" italics="true" pageId="13" pageNumber="164">Lecithocera</emphasis>
</taxonomicName>
to
<taxonomicName id="4C3BB3FFFFBAFFD15F9C98CBFBF2F84D" box="[1002,1106,1953,1979]" class="Insecta" family="Lecithoceridae" genus="Torodora" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="164" phylum="Arthropoda" rank="genus">
<emphasis id="B94F146EFFBAFFD15F9C98CBFBF2F84D" box="[1002,1106,1953,1979]" italics="true" pageId="13" pageNumber="164">Torodora</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFAAB58DFFBAFFD1580D98CBFAA4F84D" author="Park, K. - T." box="[1147,1284,1953,1979]" pageId="13" pageNumber="164" pagination="1085 - 1093" refId="ref23829" refString="Park, K. - T. (2018) A new genus Thubdora Park, gen. nov. and seven new species of the subfamily Torodorinae (Lepidoptera, Lecithoceridae) from Cameroon and Rep. of Congo, Africa. Journal of Asia-Pacific Entomology, 21, 1085 - 1093. https: // doi. org / 10.1016 / j. aspen. 2018.07.018" type="journal article" year="2018">Park (2018)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3219BF7FFBAFFD15CB198AFFD42F816" box="[199,738,1989,2016]" pageId="13" pageNumber="164" type="distribution">
<paragraph id="8B84C87CFFBAFFD15CB198AFFD42F816" blockId="13.[151,1437,1593,2016]" box="[199,738,1989,2016]" pageId="13" pageNumber="164">
<emphasis id="B94F146EFFBAFFD15CB198AFFEF9F829" bold="true" box="[199,345,1989,2015]" pageId="13" pageNumber="164">Distribution</emphasis>
.
<collectingCountry id="F32C88ECFFBAFFD15D1198AFFE4EF829" box="[359,494,1989,2015]" name="Madagascar" pageId="13" pageNumber="164">Madagascar</collectingCountry>
(East) (
<bibRefCitation id="EFAAB58DFFBAFFD15E3198AFFD76F816" author="Viette, P. E. L." box="[583,726,1989,2016]" pageId="13" pageNumber="164" pagination="109 - 118" refId="ref24658" refString="Viette, P. E. L. (1986) Nouveaux Microlepidopteres de Madagascar (Oecophoridae et Gelechiidae) (New species of Malagasy Microlepidoptera-Oecophoridae and Gelechiidae). Bulletin mensuel de la Societe linneenne de Lyon, 55 (4), 109 - 118. https: // doi. org / 10.3406 / linly. 1986.10753" type="journal article" year="1986">Viette, 1986</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>