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<mods:title id="EE05DEDB87F2D4B0D9B7DCC24472A072">A never-ending story: updated 3 D cyber-taxonomic revision of the ant genus Zasphinctus Wheeler (Hymenoptera, Formicidae, Dorylinae) for the Afrotropical region</mods:title>
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<mods:affiliation id="382CE49D89780376CBAA125DD7318E63">Garraf, Barcelona, Spain</mods:affiliation>
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<taxonomicName id="0D7F1668CACAB51BBD8360921747DB9D" ID-CoL="CN4FR" authority="Hita Garcia, 2017" authorityName="Hita Garcia" authorityYear="2017" class="Insecta" family="Formicidae" genus="Zasphinctus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="wilsoni">
<emphasis id="FEDA7D7AC808167ADB0451F25F0624CE" italics="true">Zasphinctus wilsoni</emphasis>
Hita Garcia, 2017
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<figureCitation id="AD47A9325036F97506431007566DE4FC" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Maps showing the known distribution ranges of Afrotropical Zasphinctus species." figureDoi="10.3897/zookeys.1223.131238.figure3" httpUri="https://binary.pensoft.net/fig/1213417">Figs 3 H</figureCitation>
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<figureCitation id="69B0B3BBAA7EBDFDE206B502982CDE0A" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Diagnostic plate showing still images from surface volume renderings of the head in full-face view (the remainder of the body virtually removed) (torular-posttorular complex in semi-transparent red; anterior projections of parafrontal ridges in semi-transparent yellow) A Z. aprilia sp. nov. holotype (CASENT 0764763) B Z. kouakoui sp. nov. paratype (CASENT 0764653) C Z. lolae sp. nov. holotype (KGCOL 02270) D Z. lumumbai sp. nov. holotype (MRACFOR 0010007) E Z. ndouri sp. nov. holotype (KGCOL 01883) F Z. obamai Hita Garcia, 2017 holotype (CASENT 0764125) G Z. sarowiwai Hita Garcia, 2017 holotype (CASENT 0764654) H Z. wilsoni Hita Garcia, 2017 holotype (MCZ-ENT 00512764)." figureDoi="10.3897/zookeys.1223.131238.figure4" httpUri="https://binary.pensoft.net/fig/1213418">4 H</figureCitation>
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<figureCitation id="A725969C0692182F0F052F08F261BD7E" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Diagnostic plate showing still images from surface volume renderings of the head in profile view (the remainder of the body virtually removed) (torular-posttorular complex in semi-transparent red) A Z. aprilia sp. nov. holotype (CASENT 0764763) B Z. kouakoui sp. nov. paratype (CASENT 0764653) C Z. lolae sp. nov. paratype (CASENT 0764651) D Z. lumumbai sp. nov. holotype (KGCOL 02270) E Z. ndouri sp. nov. holotype (KGCOL 01883) F Z. obamai Hita Garcia, 2017 holotype (CASENT 0764125) G Z. sarowiwai Hita Garcia, 2017 holotype (CASENT 0764654) H Z. wilsoni Hita Garcia, 2017 holotype (MCZ-ENT 00512764)." figureDoi="10.3897/zookeys.1223.131238.figure5" httpUri="https://binary.pensoft.net/fig/1213419">5 H</figureCitation>
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<figureCitation id="CECB8EBD75650FA7B0E9A8E69EABA413" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Diagnostic plate showing still images from surface volume renderings of the head in posterodorsal view (the remainder of the body virtually removed) (occipital margin in semi-transparent red; vertexal margin in semi-transparent yellow) A Z. aprilia sp. nov. holotype (CASENT 0764763) B Z. kouakoui sp. nov. paratype (CASENT 0764653) C Z. lolae sp. nov. holotype (KGCOL 02270) D Z. lumumbai sp. nov. holotype (MRACFOR 0010007) E Z. ndouri sp. nov. holotype (KGCOL 01883) F Z. obamai Hita Garcia, 2017 holotype (CASENT 0764125) G Z. sarowiwai Hita Garcia, 2017 holotype (CASENT 0764654) H Z. wilsoni Hita Garcia, 2017 holotype (MCZ-ENT 00512764)." figureDoi="10.3897/zookeys.1223.131238.figure6" httpUri="https://binary.pensoft.net/fig/1213420">6 H</figureCitation>
,
<figureCitation id="B585FDF612B715DDAF326C152E757AAC" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Diagnostic plate showing still images from surface volume renderings of the head in posterior view (the remainder of the body virtually removed) (occipital margin in semi-transparent red; vertexal margin in semi-transparent yellow) A Z. aprilia sp. nov. holotype (CASENT 0764763) B Z. kouakoui sp. nov. paratype (CASENT 0764653) C Z. lolae sp. nov. holotype (KGCOL 02270) D Z. lumumbai sp. nov. holotype (MRACFOR 0010007) E Z. ndouri sp. nov. holotype (KGCOL 01883) F Z. obamai Hita Garcia, 2017 holotype (CASENT 0764125) G Z. sarowiwai Hita Garcia, 2017 holotype (CASENT 0764654) H Z. wilsoni Hita Garcia, 2017 holotype (MCZ-ENT 00512764)." figureDoi="10.3897/zookeys.1223.131238.figure7" httpUri="https://binary.pensoft.net/fig/1213421">7 H</figureCitation>
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<figureCitation id="4E5EC958148463F2FAD5C94DF9D86A7E" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Diagnostic plate showing still images from surface volume renderings of the head in ventral view (the remainder of the body virtually removed) (hypostoma in semi-transparent red; postgenal sulcus in semi-transparent yellow; postoccipital carina in semi-transparent green) A Z. aprilia sp. nov. holotype (CASENT 0764763) B Z. kouakoui sp. nov. paratype (CASENT 0764653) C Z. lolae sp. nov. paratype (CASENT 0764651) D Z. lumumbai sp. nov. holotype (MRACFOR 0010007) E Z. ndouri sp. nov. holotype (KGCOL 01883) F Z. obamai Hita Garcia, 2017 holotype (CASENT 0764125) G Z. sarowiwai Hita Garcia, 2017 paratype (CASENT 0764650) H Z. wilsoni Hita Garcia, 2017 holotype (MCZ-ENT 00512764)." figureDoi="10.3897/zookeys.1223.131238.figure8" httpUri="https://binary.pensoft.net/fig/1213422">8 H</figureCitation>
,
<figureCitation id="145C9CAEE9C047B28A078F56FF2E5306" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Diagnostic plate showing still images from surface volume renderings of the mesosoma in profile view (the remainder of the body virtually removed) (pleural endophragmal pit concavity in semi-transparent yellow) A Z. aprilia sp. nov. holotype (CASENT 0764763) B Z. kouakoui sp. nov. paratype (CASENT 0764653) C Z. lolae sp. nov. holotype (KGCOL 02270) D Z. lumumbai sp. nov. holotype (MRACFOR 0010007) E Z. ndouri sp. nov. holotype (KGCOL 01883) F Z. obamai Hita Garcia, 2017 holotype (CASENT 0764125) G Z. sarowiwai Hita Garcia, 2017 paratype (CASENT 0764650) H Z. wilsoni Hita Garcia, 2017 holotype (MCZ-ENT 00512764)." figureDoi="10.3897/zookeys.1223.131238.figure10" httpUri="https://binary.pensoft.net/fig/1213424">10 H</figureCitation>
,
<figureCitation id="DB9463080C114DF0D667111B046CA3EB" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Diagnostic plate showing still images from surface volume renderings of the mesosoma in dorsal view (the remainder of the body virtually removed) A Z. aprilia sp. nov. holotype (CASENT 0764763) B Z. kouakoui sp. nov. paratype (CASENT 0764653) C Z. lolae sp. nov. holotype (KGCOL 02270) D Z. lumumbai sp. nov. holotype (MRACFOR 0010007) E Z. ndouri sp. nov. holotype (KGCOL 01883) F Z. obamai Hita Garcia, 2017 holotype (CASENT 0764125) G Z. sarowiwai Hita Garcia, 2017 paratype (CASENT 0764650) H Z. wilsoni Hita Garcia, 2017 holotype (MCZ-ENT 00512764)." figureDoi="10.3897/zookeys.1223.131238.figure11" httpUri="https://binary.pensoft.net/fig/1213425">11 H</figureCitation>
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<figureCitation id="90261F7E8A6C2D9936141C385AFB7735" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Diagnostic plate showing still images from surface volume renderings of the mesosoma in posterior view focusing on propodeal declivity (outline of declivity in semi-transparent yellow) A Z. aprilia sp. nov. holotype (CASENT 0764763) B Z. kouakoui sp. nov. paratype (CASENT 0764653) C Z. lolae sp. nov. paratype (CASENT 0764651) D Z. lumumbai sp. nov. holotype (MRACFOR 0010007) E Z. ndouri sp. nov. holotype (KGCOL 01883) F Z. obamai Hita Garcia, 2017 holotype (CASENT 0764125) G Z. sarowiwai Hita Garcia, 2017 paratype (CASENT 0764650) H Z. wilsoni Hita Garcia, 2017 holotype (MCZ-ENT 00512764)." figureDoi="10.3897/zookeys.1223.131238.figure12" httpUri="https://binary.pensoft.net/fig/1213426">12 H</figureCitation>
,
<figureCitation id="52E2BBE2D4AD7A741D3252A7D8087966" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Diagnostic plate showing still images from surface volume renderings of the tergum of AS II (petiole) in profile view (the remainder of the body virtually removed) (tergum in semi-transparent red, laterotergite in semi-transparent yellow) A Z. aprilia sp. nov. holotype (CASENT 0764763) B Z. kouakoui sp. nov. paratype (CASENT 0764653) C Z. lolae sp. nov. holotype (KGCOL 02270) D Z. lumumbai sp. nov. holotype (MRACFOR 0010007) E Z. ndouri sp. nov. holotype (KGCOL 01883) F Z. obamai Hita Garcia, 2017 holotype (CASENT 0764125) G Z. sarowiwai Hita Garcia, 2017 paratype (CASENT 0764650) H Z. wilsoni Hita Garcia, 2017 holotype (MCZ-ENT 00512764)." figureDoi="10.3897/zookeys.1223.131238.figure13" httpUri="https://binary.pensoft.net/fig/1213427">13 H</figureCitation>
,
<figureCitation id="5C28E8DA371A32DF1F34CC246D75335E" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Diagnostic plate showing still images from surface volume renderings of the tergum of AS II (petiole) in dorsal view (the remainder of the body virtually removed) A Z. aprilia sp. nov. holotype (CASENT 0764763) B Z. kouakoui sp. nov. holotype (KGCOL 00589) C Z. lolae sp. nov. holotype (KGCOL 02270) D Z. lumumbai sp. nov. holotype (MRACFOR 0010007) E Z. ndouri sp. nov. holotype (KGCOL 01883) F Z. obamai Hita Garcia, 2017 holotype (CASENT 0764125) G Z. sarowiwai Hita Garcia, 2017 paratype (CASENT 0764650) H Z. wilsoni Hita Garcia, 2017 holotype (MCZ-ENT 00512764)." figureDoi="10.3897/zookeys.1223.131238.figure14" httpUri="https://binary.pensoft.net/fig/1213428">14 H</figureCitation>
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<figureCitation id="08943490927FD4B370240225D6E268B5" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Diagnostic plate showing still images from surface volume renderings of the tergum of AS III in dorsal view (the remainder of the body virtually removed) A Z. aprilia sp. nov. holotype (CASENT 0764763) B Z. kouakoui sp. nov. paratype (CASENT 0764653) C Z. lolae sp. nov. paratype (CASENT 0764651) D Z. lumumbai sp. nov. holotype (MRACFOR 0010007) E Z. ndouri sp. nov. holotype (KGCOL 01883) F Z. obamai Hita Garcia, 2017 holotype (CASENT 0764125) G Z. sarowiwai Hita Garcia, 2017 paratype (CASENT 0764650) H Z. wilsoni Hita Garcia, 2017 holotype (MCZ-ENT 00512764)." figureDoi="10.3897/zookeys.1223.131238.figure15" httpUri="https://binary.pensoft.net/fig/1213429">15 H</figureCitation>
,
<figureCitation id="C9571719AFC622DEA3B1D1E20A4CAC50" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Diagnostic plate showing still images from surface volume renderings of the sternum of AS III in ventral view (the remainder of the body virtually removed) (Bum in semi-transparent yellow, prora in semi-transparent red) A Z. aprilia sp. nov. holotype (CASENT 0764763) B Z. kouakoui sp. nov. paratype (CASENT 0764653) C Z. lolae sp. nov. paratype (CASENT 0764651) D Z. lumumbai sp. nov. holotype (MRACFOR 0010007) E Z. ndouri sp. nov. holotype (KGCOL 01883) F Z. obamai Hita Garcia, 2017 holotype (CASENT 0764125) G Z. sarowiwai Hita Garcia, 2017 paratype (CASENT 0764650) H Z. wilsoni Hita Garcia, 2017 holotype (MCZ-ENT 00512764)." figureDoi="10.3897/zookeys.1223.131238.figure16" httpUri="https://binary.pensoft.net/fig/1213430">16 H</figureCitation>
,
<figureCitation id="A3078D499D5BFB8D08B9BC5D3295F829" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Diagnostic plate showing still images from surface volume renderings of the tergum of AS VI in dorsal view (the remainder of the body virtually removed) (outline of post-sclerite tergum in semi-transparent red) A Z. aprilia sp. nov. holotype (CASENT 0764763) B Z. kouakoui sp. nov. paratype (CASENT 0764653) C Zasphinctus lolae sp. nov. paratype (CASENT 0764651) D Z. lumumbai sp. nov. holotype (MRACFOR 0010007) E Z. ndouri sp. nov. holotype (KGCOL 01883) F Z. obamai Hita Garcia, 2017 holotype (CASENT 0764125) G Z. sarowiwai Hita Garcia, 2017 paratype (CASENT 0764650) H Z. wilsoni Hita Garcia, 2017 holotype (MCZ-ENT 00512764)." figureDoi="10.3897/zookeys.1223.131238.figure17" httpUri="https://binary.pensoft.net/fig/1213431">17 H</figureCitation>
,
<figureCitation id="57048054B2E28A9EAB303C6F06547763" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Diagnostic plate showing still images from surface volume renderings of the sternites of AS III-VII in ventral view (the remainder of the body virtually removed) (girdling constrictions with cross-ribbed sculpture are in semi-transparent red; unsculptured girdling constrictions are in semi-transparent yellow) A Z. aprilia sp. nov. holotype (CASENT 0764763) B Z. kouakoui sp. nov. paratype (CASENT 0764653) C Z. lolae sp. nov. paratype (CASENT 0764651) D Z. lumumbai sp. nov. holotype (MRACFOR 0010007) E Z. ndouri sp. nov. holotype (KGCOL 01883) F Z. obamai Hita Garcia, 2017 holotype (CASENT 0764125) G Z. sarowiwai Hita Garcia, 2017 paratype (CASENT 0764650) H Z. wilsoni Hita Garcia, 2017 holotype (MCZ-ENT 00512764)." figureDoi="10.3897/zookeys.1223.131238.figure18" httpUri="https://binary.pensoft.net/fig/1213432">18 H</figureCitation>
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<figureCitation id="449417AB1E38B6836BAF5100F4FBA9FC" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Diagnostic plate showing head in full-face view of all species treated herein (stacked colour images) A Z. aprilia sp. nov. holotype (CASENT 0764763) B Z. kouakoui sp. nov. holotype (KGCOL 00589) C Z. lolae sp. nov. holotype (KGCOL 02270) D Z. lumumbai sp. nov. holotype (MRACFOR 0010007) E Z. ndouri sp. nov. holotype (KGCOL 01883) F Z. obamai Hita Garcia, 2017 holotype (CASENT 0764125) G Z. sarowiwai Hita Garcia, 2017 paratype (CASENT 0764650) H Z. wilsoni Hita Garcia, 2017 holotype (MCZ-ENT 00512764)." figureDoi="10.3897/zookeys.1223.131238.figure19" httpUri="https://binary.pensoft.net/fig/1213433">19 H</figureCitation>
,
<figureCitation id="F816DAEB35682B5B093C2BECBB13CA22" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Diagnostic plate showing full body in profile view of all species treated herein (stacked colour images) A Z. aprilia sp. nov. holotype (CASENT 0764763) B Z. kouakoui sp. nov. holotype (KGCOL 00589) C Z. lolae sp. nov. holotype (KGCOL 02270) D Z. lumumbai sp. nov. holotype (MRACFOR 0010007) E Z. ndouri sp. nov. holotype (KGCOL 01883) F Z. obamai Hita Garcia, 2017 holotype (CASENT 0764125) G Z. sarowiwai Hita Garcia, 2017 holotype (CASENT 0764654) H Z. wilsoni Hita Garcia, 2017 holotype (MCZ-ENT 00512764)." figureDoi="10.3897/zookeys.1223.131238.figure20" httpUri="https://binary.pensoft.net/fig/1213434">20 H</figureCitation>
,
<figureCitation id="AB551DE1F5B5B7FD32DC96FA65A857BF" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Diagnostic plate showing full body in dorsal view of all species treated herein (stacked colour images) A Z. aprilia sp. nov. holotype (CASENT 0764763) B Z. kouakoui sp. nov. holotype (KGCOL 00589) C Z. lolae sp. nov. holotype (KGCOL 02270) D Z. lumumbai sp. nov. holotype (MRACFOR 0010007) E Z. ndouri sp. nov. holotype (KGCOL 01883) F Z. obamai Hita Garcia, 2017 holotype (CASENT 0764125) G Z. sarowiwai Hita Garcia, 2017 holotype (CASENT 0764654) H Z. wilsoni Hita Garcia 2017 holotype (MCZ-ENT 00512764)." figureDoi="10.3897/zookeys.1223.131238.figure21" httpUri="https://binary.pensoft.net/fig/1213435">21 H</figureCitation>
,
<figureCitation id="2387FF9A0D6C4018739FA2E8ED3DD56A" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Shaded surface display volume renderings of 3 D models of Z. wilsoni Hita Garcia, 2017 holotype (MCZ-ENT 00512764) A full body in profile B full body in dorsal view C head in full-face view (with antennae) D head in full-face view (without antennae) E head in ventral view F abdominal segment II (petiole) in profile G abdominal segment II (petiole) in dorsal view H tergum of AS III in dorsal view I sternum of AS III in ventral view." figureDoi="10.3897/zookeys.1223.131238.figure24" httpUri="https://binary.pensoft.net/fig/1213438">24</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EUTAK" type="type material">
<paragraph id="DD8D54AD551D25A3D86A1A15ED8A6100">
<heading id="3BE3B1A66F7E3C5725084CBBB6CA74F9" reason="title">Type material examined.</heading>
</paragraph>
<paragraph id="D30CE21143B51312341E9178A313B595">
<materialsCitation id="F9C01F1AC03B62929E1C30A6A4A7EABE" collectedFrom="secondary forest, leaf litter" collectingDate="2012-05-30" collectionCode="MCZC" collectorName="G. D. Alpert" country="Mozambique" elevation="40" latitude="-18.99472" location="Gorongosa National Park" longLatPrecision="1" longitude="34.35769" municipality="Pinned" specimenCode="MCZ-ENT 00512764" specimenCount="1" stateProvince="Sofala" typeStatus="holotype">
<emphasis id="2A6D7D42993540EB170821CF5E06996A" bold="true" italics="true">
<typeStatus id="FCB7E239EE6AD945FCAEE52A4914B487" type="holotype">Holotype</typeStatus>
</emphasis>
• Pinned worker,
<collectingCountry id="2945EF928006DE337E95E3A27C7B186A" name="Mozambique">Mozambique</collectingCountry>
,
<collectingRegion id="AB664FEAD45A0FCE098CF4757A6A8EAA" country="Mozambique" name="Sofala">Sofala</collectingRegion>
,
<location id="28C8E0C17CC00365C5E577EE86149D45" LSID="urn:lsid:plazi:treatment:F252AF06EC075B3E804FDAF1FB296058:28C8E0C17CC00365C5E577EE86149D45" country="Mozambique" latitude="-18.99472" longLatPrecision="1" longitude="34.35769" municipality="Pinned" name="Gorongosa National Park" stateProvince="Sofala">Gorongosa National Park</location>
,
<locationDeviation id="C6D18CCD48E70AA06AB63D3C2FE9550B" location="Chitengo">
<quantity id="8DC2CC31B0AF1140BD76C67E81499CF5" metricMagnitude="3" metricUnit="m" metricValue="2.0" unit="km" value="2.0">2 km</quantity>
S Chitengo
</locationDeviation>
,
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ECUAK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[34.357690,-18.994720]}">
<geoCoordinate id="09110C9C2E629014F6DB6B0D533F244A" degrees="18.99472" direction="south" orientation="latitude" precision="1" value="-18.99472">- 18.99472</geoCoordinate>
,
<geoCoordinate id="17EFCE98DB30B26D01A338FB394AE8E3" degrees="34.35769" direction="east" orientation="longitude" precision="1" value="34.35769">34.35769</geoCoordinate>
</named-content>
,
<quantity id="FF162BAEF4E91CCC3A1E958C7D8F12A1" metricMagnitude="1" metricUnit="m" metricValue="4.0" unit="m" value="40.0">
<elevation id="089EE8CA76CD7CEECBEA8924085801D2" metricMagnitude="1" metricUnit="m" metricValue="4.0" unit="m" value="40.0">40 m</elevation>
</quantity>
,
<collectedFrom id="3BD20FC640AE145D94C468551F126732">secondary forest, leaf litter</collectedFrom>
, collection code ANTC 37418,
<date id="A501407A72512669DCF5C540CFBD9827" value="2012-05-30">
<collectingDate id="8521B8935C3BCD1E8B6F4E2D6821169E" value="2012-05-30">30. V. 2012</collectingDate>
</date>
(
<emphasis id="2FBC1C6383889DDF2020CA057DC112CF" italics="true">
<collectorName id="1B94079A5BE93878F46EAADE9F1D23D2">G. D. Alpert</collectorName>
</emphasis>
) (
<named-content id="BC9BE59C16F62CC9074CCA12C03C7633" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museum-comparative-zoology" xlink_title="Museum of Comparative Zoology, Harvard University, Cambridge, USA">
<collectionCode id="8D21FE013E44F984C521303039C8585A">MCZC</collectionCode>
</named-content>
:
<ext-link id="39FEB2834595F1AA2C1B94C6B7777C65" ext-link-type="uri" xlink_href="https://www.antweb.org/specimen.do?code=mcz-ent00512764" xlink_type="simple">
<specimenCode id="0ACAC38F656CBB9CBB4E649F56EEAA2A">MCZ-ENT 00512764</specimenCode>
</ext-link>
).
</materialsCitation>
</paragraph>
<paragraph id="C938534687C5F8E5B1DCC090F94E14B0">
<emphasis id="9EC2945F33F00DFAB5C10EE76F2946F1" bold="true" italics="true">Cybertype</emphasis>
• Dataset was published in
<bibRefCitation id="43C15BF54BCA17970ED9AF42B4678400" DOI="10.3897/zookeys.693.13012" author="Hita Garcia F &amp; Fischer G &amp; Liu C &amp; Audisio TL &amp; Economo EP" journalOrPublisher="ZooKeys" pagination="33-93" refId="B32" refString="Hita Garcia F, Fischer G, Liu C, Audisio TL, Economo EP (2017 a) Next-generation morphological character discovery and evaluation: an X-ray micro-CT enhanced revision of the ant genus Zasphinctus Wheeler (Hymenoptera, Formicidae, Dorylinae) in the Afrotropics. ZooKeys 693: 3393. https://doi.org/10.3897/zookeys.693.13012" title="Next-generation morphological character discovery and evaluation: an X-ray micro-CT enhanced revision of the ant genus Zasphinctus Wheeler (Hymenoptera, Formicidae, Dorylinae) in the Afrotropics." volume="693">Hita Garcia et al. (2017 a</bibRefCitation>
) and consists of the volumetric raw data (in DICOM format), as well as 3 D PDFs and 3 D rotation videos of scans of the head, mesosoma, metasoma, and the full body of the physical
<typeStatus id="90B9D554DCDA7B1A1670140EFE81DC55" type="holotype">holotype</typeStatus>
(
<named-content id="4D75D18786EE0C200300B1146E405971" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museum-comparative-zoology" xlink_title="Museum of Comparative Zoology, Harvard University, Cambridge, USA">
<collectionCode id="0A1781A94F8A345FC50B7E8D45D52A01">MCZC</collectionCode>
</named-content>
:
<ext-link id="58584303ACA2CF20D8CB238F5E25FF41" ext-link-type="uri" xlink_href="https://www.antweb.org/specimen.do?code=mcz-ent00512764" xlink_type="simple">MCZ-ENT 00512764</ext-link>
) in addition to montage photos illustrating head in full-face view, profile, and dorsal views of the body. The data was deposited at Dryad and can be freely accessed as virtual representation of the
<typeStatus id="FD754979808777384276531545894F05" type="holotype">holotype</typeStatus>
(
<bibRefCitation id="ECCCBE5D7D480FD0285CF14B1DF37AEA" DOI="10.5061/dryad.4s3v1" author="Hita Garcia F &amp; Fischer G &amp; Liu C &amp; Audisio TL &amp; Economo EP" refId="B34" refString="Hita Garcia F, Fischer G, Liu C, Audisio TL, Economo EP (2017 c) Data from: Next-generation morphological character discovery and evaluation: an X-ray micro-CT enhanced revision of the ant genus Zasphinctus Wheeler (Hymenoptera, Formicidae, Dorylinae) in the Afrotropics [Dataset]. Dryad Digital Repository. https://doi.org/10.5061/dryad.4s3v1">Hita Garcia et al. 2017 c</bibRefCitation>
,
<ext-link id="2727CB86E2CF927E2A23EE5DB225A55B" ext-link-type="doi" xlink_href="10.5061/dryad.4s3v1" xlink_type="simple">http://dx.doi.org/10.5061/dryad.4s3v1</ext-link>
). In addition to the cybertype data at Dryad, we also provided a freely accessible 3 D surface model of the
<typeStatus id="FECFEB67EB97A59DB5706E36EAFA89F0" type="holotype">holotype</typeStatus>
at Sketchfab (
<ext-link id="191BBBD82638A7E1AE3A3EF8A40F8662" ext-link-type="uri" xlink_href="https://skfb.ly/6sPwN" xlink_type="simple">https://skfb.ly/6sPwN</ext-link>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0E2VAK" type="type material">
<paragraph id="AA491D044BAD9B10EA2B95372797B14A">
<heading id="B22E99591B994721B8E527412757E4DA" reason="title">Non-type material examined.</heading>
</paragraph>
<paragraph id="C29187EFF560F9525FD265034822C44E">
<materialsCitation id="8171CED71E24A3E2E15CB1C85F2C1433" collectedFrom="miombo woodland, ex soil" collectingDate="2016-02-25" collectionCode="CASC" collectorName="B. L. Fisher &amp; Arthropod Team" country="Mozambique" elevation="180" latitude="-12.84778" location="Mareja Reserve" longLatPrecision="1" longitude="40.16542" municipality="Parque Nacional Quirimbas" specimenCode="CASENT 0779283, CASENT 0779285" specimenCount="5" specimenCount-worker="5" stateProvince="Cabo Delgado">
<specimenCount id="8A74B031D6BB92430E114117AD8B0A96" count="5" type="worker">Five workers</specimenCount>
from:
<collectingCountry id="43F338F3526602118E95E026E1072945" name="Mozambique">Mozambique</collectingCountry>
:
<collectingRegion id="88018C2BF485825A8C094EA1954D407F" country="Mozambique" name="Cabo Delgado">Cabo Delgado</collectingRegion>
,
<collectingMunicipality id="E8ECE1F6D2717943EC72C80882D4AA3E">Parque Nacional Quirimbas</collectingMunicipality>
,
<location id="48BF083EB27DE9647E77CA2CA03B8E2F" LSID="urn:lsid:plazi:treatment:F252AF06EC075B3E804FDAF1FB296058:48BF083EB27DE9647E77CA2CA03B8E2F" country="Mozambique" latitude="-12.84778" longLatPrecision="1" longitude="40.16542" municipality="Parque Nacional Quirimbas" name="Mareja Reserve" stateProvince="Cabo Delgado">Mareja Reserve</location>
,
<collectedFrom id="499B69340EE8824B9A30BE06DD5D374D">miombo woodland, ex soil</collectedFrom>
,
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EEWAK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[40.165420,-12.847780]}">
<geoCoordinate id="44F18129D7FF102C55AC195CCAB8E728" degrees="12.84778" direction="south" orientation="latitude" precision="1" value="-12.84778">- 12.84778</geoCoordinate>
,
<geoCoordinate id="E39765CD1A26495D0DC92576CF10AEA4" degrees="40.16542" direction="east" orientation="longitude" precision="1" value="40.16542">40.16542</geoCoordinate>
</named-content>
,
<quantity id="D1119A8D720E1D9BA0E2B4D21E230839" metricMagnitude="2" metricUnit="m" metricValue="1.8" unit="m" value="180.0">
<elevation id="D47C01F188812D95DA2A14B0CF255CF9" metricMagnitude="2" metricUnit="m" metricValue="1.8" unit="m" value="180.0">180 m</elevation>
</quantity>
, collection code BLF 38248,
<date id="ABD9A9F0AC5078E3EA42DD10A09A1985" value="2016-02-25">
<collectingDate id="44C6F2EDC283BDE37D44A860EBB0C00E" value="2016-02-25">25. II. 2016</collectingDate>
</date>
(
<emphasis id="8A53E5C0264B967CE1F404D4719C7DAE" italics="true">
<collectorName id="902B06127D9879E3FB82DF096D8A3D4C">B. L. Fisher</collectorName>
;
<collectorName id="F55851887D2EFBCFFA175B830E16936A">Arthropod Team</collectorName>
</emphasis>
) (
<named-content id="622B78F1A2002EF467CFB51BCB79FBCD" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/california-academy-sciences" xlink_title="California Academy of Sciences, San Francisco, USA">
<collectionCode id="B8B8941F69FC0CBC21A4A3A573DAEE2F">CASC</collectionCode>
</named-content>
:
<ext-link id="F3865BD8CE3ABF4BD81023F70D37CBB7" ext-link-type="uri" xlink_href="http://www.antweb.org/specimen/CASENT0779283" xlink_type="simple">
<specimenCode id="CF0221904299444E9EA9FDA9668B5F5D" collectionCode="CASC">CASENT 0779283</specimenCode>
</ext-link>
,
<ext-link id="C73CE091F9C823CEB3323C7FD782A7E7" ext-link-type="uri" xlink_href="http://www.antweb.org/specimen/CASENT0779285" xlink_type="simple">
<specimenCode id="5FF16F650989AA877DB2336E166C7161" collectionCode="CASC">CASENT 0779285</specimenCode>
</ext-link>
)
</materialsCitation>
<materialsCitation id="D6DF22FF01CEA21E4598A68AA4096AE2" collectedFrom="rainforest, ex soil" collectingDate="2016-03-12" collectionCode="CASC" collectorName="B. L. Fisher &amp; Arthropod Team" country="Mozambique" elevation="375" latitude="-16.34888" location="Mount Mabu" longLatPrecision="5" longitude="36.4081" specimenCode="CASENT 0779844" specimenCount="1" stateProvince="Zambezia">
<collectingRegion id="E9D75958FBA486CD569F6D508C9028A3" country="Mozambique" name="Zambezia">Zambezia</collectingRegion>
,
<location id="62D7906368EE93B7D3146BFDE6F7CC63" LSID="urn:lsid:plazi:treatment:F252AF06EC075B3E804FDAF1FB296058:62D7906368EE93B7D3146BFDE6F7CC63" country="Mozambique" latitude="-16.34888" longLatPrecision="5" longitude="36.4081" name="Mount Mabu" stateProvince="Zambezia">Mount Mabu</location>
,
<collectedFrom id="12B0AA2EF34B045F0BF7073CAD48E17A">rainforest, ex soil</collectedFrom>
,
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E4WAK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[36.408100,-16.348880]}">
<geoCoordinate id="3B79BEB9A45ABD741BDEF1413C85351A" degrees="16.34888" direction="south" orientation="latitude" precision="1" value="-16.34888">- 16.34888</geoCoordinate>
,
<geoCoordinate id="55D05033C753291C9B2A41595A40C9C8" degrees="36.4081" direction="east" orientation="longitude" precision="5" value="36.4081">36.4081</geoCoordinate>
</named-content>
,
<quantity id="82011BF9521E303BD5782118FB3BBA28" metricMagnitude="2" metricUnit="m" metricValue="3.75" unit="m" value="375.0">
<elevation id="C3EE051D67850E0DAEDDEB726C14B19B" metricMagnitude="2" metricUnit="m" metricValue="3.75" unit="m" value="375.0">375 m</elevation>
</quantity>
, collection code BLF 38954,
<date id="CB5865EBB7B7C3C27F2B8E8BC9C44DBA" value="2016-03-12">
<collectingDate id="602000CC198CAFF0EF9944014DEEC3A0" value="2016-03-12">12. III. 2016</collectingDate>
</date>
(
<emphasis id="2C68E9DBF903F83923ED78564309A1F5" italics="true">
<collectorName id="87AB06BA1DF281EB0436F727C29E8B67">B. L. Fisher</collectorName>
;
<collectorName id="30BFAF924104A7B457F590924F2D247A">Arthropod Team</collectorName>
</emphasis>
) (
<named-content id="9C3BBF8A42647CEB21F7184ABD18213F" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/california-academy-sciences" xlink_title="California Academy of Sciences, San Francisco, USA">
<collectionCode id="BED3162C4C6A3833FF3D7774A94A7833">CASC</collectionCode>
</named-content>
:
<ext-link id="02EABCA968863C148E8118E7D4D9C69F" ext-link-type="uri" xlink_href="http://www.antweb.org/specimen/CASENT0779844" xlink_type="simple">
<specimenCode id="C7C9D8E034F850F7A8CC95865D185343" collectionCode="CASC">CASENT 0779844</specimenCode>
</ext-link>
)
</materialsCitation>
<materialsCitation id="F5B89DA9E979849C8E6BD1037A4C616F" collectedFrom="rainforest, ex soil" collectingDate="2016-03-24" collectionCode="CASC" collectorName="B. L. Fisher &amp; Arthropod Team" country="Mozambique" elevation="375" latitude="-16.34888" location="Mount Mabu" longLatPrecision="5" longitude="36.4081" specimenCode="CASENT 0781222" specimenCount="1" stateProvince="Zambezia">
<collectingRegion id="5BF2212A2FBFCD6D9C74151AEE2E03C0" country="Mozambique" name="Zambezia">Zambezia</collectingRegion>
,
<location id="97A5DB6874559EB5C737522CE95597DC" LSID="urn:lsid:plazi:treatment:F252AF06EC075B3E804FDAF1FB296058:97A5DB6874559EB5C737522CE95597DC" country="Mozambique" latitude="-16.34888" longLatPrecision="5" longitude="36.4081" name="Mount Mabu" stateProvince="Zambezia">Mount Mabu</location>
,
<collectedFrom id="CB9EBF11ACA22D1CBF393637B7195919">rainforest, ex soil</collectedFrom>
,
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ERXAK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[36.408100,-16.348880]}">
<geoCoordinate id="B199C01D166225D9DEA0F2F268CDD480" degrees="16.34888" direction="south" orientation="latitude" precision="1" value="-16.34888">- 16.34888</geoCoordinate>
,
<geoCoordinate id="971BC512EFF3B8253E15C9EAFC853E8B" degrees="36.4081" direction="east" orientation="longitude" precision="5" value="36.4081">36.4081</geoCoordinate>
</named-content>
,
<quantity id="709C8E2F4B5CFDAC0E43FA83286EA1D7" metricMagnitude="2" metricUnit="m" metricValue="3.75" unit="m" value="375.0">
<elevation id="463B15B6AFA2FEBFD23C1A11DC3C6C08" metricMagnitude="2" metricUnit="m" metricValue="3.75" unit="m" value="375.0">375 m</elevation>
</quantity>
, collection code BLF 39161,
<date id="8D749405A9258F42535ADFB4720797D6" value="2016-03-24">
<collectingDate id="4557BDC15C906F77047854888FEB9C9E" value="2016-03-24">24. III. 2016</collectingDate>
</date>
(
<emphasis id="F046BC17A01361311BB330F5E7C1DEC7" italics="true">
<collectorName id="8101AEF1EC276FEF39301D818712E95E">B. L. Fisher</collectorName>
;
<collectorName id="0B0373276FD651950D15449F729643AF">Arthropod Team</collectorName>
</emphasis>
) (
<named-content id="14C2610B24BC10C1CE6694152F500813" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/california-academy-sciences" xlink_title="California Academy of Sciences, San Francisco, USA">
<collectionCode id="8CA947E9082310910174F23EE6A4A426">CASC</collectionCode>
</named-content>
:
<ext-link id="F76578831ABA1D122423D8C5CF569EC3" ext-link-type="uri" xlink_href="http://www.antweb.org/specimen/CASENT0781222" xlink_type="simple">
<specimenCode id="876525672B4CC082AA3215CE7B7C36EC" collectionCode="CASC">CASENT 0781222</specimenCode>
</ext-link>
)
</materialsCitation>
<materialsCitation id="DA6D7AE86164DBADE4619766E6F7E8A8" collectedFrom="rainforest, ex soil" collectingDate="2016-03-25" collectionCode="CASC" collectorName="B. L. Fisher &amp; Arthropod Team" country="Mozambique" elevation="375" latitude="-16.34888" location="Mount Mabu" longLatPrecision="5" longitude="36.4081" specimenCode="CASENT 0781285" specimenCount="1" stateProvince="Zambezia">
<collectingRegion id="F77697905375EC09C5EDC43841ECC248" country="Mozambique" name="Zambezia">Zambezia</collectingRegion>
,
<location id="43CAD073E4CA8238D10D4293DE5ABE92" LSID="urn:lsid:plazi:treatment:F252AF06EC075B3E804FDAF1FB296058:43CAD073E4CA8238D10D4293DE5ABE92" country="Mozambique" latitude="-16.34888" longLatPrecision="5" longitude="36.4081" name="Mount Mabu" stateProvince="Zambezia">Mount Mabu</location>
,
<collectedFrom id="77F2A5D90602A39A25D17721C91C27CA">rainforest, ex soil</collectedFrom>
,
<geoCoordinate id="723145E846EAE9ACA4FBA5123E2E5FD3" degrees="16.34888" direction="south" orientation="latitude" precision="1" value="-16.34888">- 16.34888</geoCoordinate>
,
<geoCoordinate id="DF953EB366AB6B001D93C5AE5DD88B98" degrees="36.4081" direction="east" orientation="longitude" precision="5" value="36.4081">36.4081</geoCoordinate>
,
<quantity id="E6E5E6F1DA2EFA80D5E0BBAD3761B1C7" metricMagnitude="2" metricUnit="m" metricValue="3.75" unit="m" value="375.0">
<elevation id="06800EED4BB974EBD0361CC54BD2D175" metricMagnitude="2" metricUnit="m" metricValue="3.75" unit="m" value="375.0">375 m</elevation>
</quantity>
, collection code BLF 39209,
<date id="48C041C61DFBCE316D99EE6856CBB64D" value="2016-03-25">
<collectingDate id="71322C050AC14EB18BF71F65405DD08B" value="2016-03-25">25. III. 2016</collectingDate>
</date>
(
<emphasis id="2453E6E04F0DAF966F3E1DA5690B6D9A" italics="true">
<collectorName id="720ECB294BD9117998736B8D0E16C648">B. L. Fisher</collectorName>
;
<collectorName id="6A6824581E00CCF9986174A3DA863B2E">Arthropod Team</collectorName>
</emphasis>
) (
<named-content id="7B9139FAD85A16B4A9C6730EBC713D12" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/california-academy-sciences" xlink_title="California Academy of Sciences, San Francisco, USA">
<collectionCode id="3FAA44274C85F094C4B78927276220E3">CASC</collectionCode>
</named-content>
:
<ext-link id="7DC5690DBC1B862B38658A0BCC20DA6D" ext-link-type="uri" xlink_href="http://www.antweb.org/specimen/CASENT0781285" xlink_type="simple">
<specimenCode id="B32E406B72897812B7EFCB5F6A0F68B4" collectionCode="CASC">CASENT 0781285</specimenCode>
</ext-link>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EOYAK" type="Differential worker diagnosis">
<paragraph id="C7291664F53584F3A9ED28E10D7E157B">
<heading id="8300DFABBDDE1117F77415C95C2124B0" reason="title">Differential worker diagnosis.</heading>
</paragraph>
<paragraph id="66142DA58B7804F77E04AB6004D8E691">
With characters of the
<taxonomicName id="D6A6A13B09785E9BA5B8D882249153D9" authorityName="Hita Garcia" authorityYear="2017" class="Insecta" family="Formicidae" genus="Zasphinctus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="obamai">
<emphasis id="B459C2492DAD66EE00F3AC612B48ACA4" italics="true">Z. obamai</emphasis>
</taxonomicName>
group plus the following: body size significantly much smaller (
<abbrev id="ABBRID0E6YAK" xlink_title="Head Length">HL</abbrev>
0.61;
<abbrev id="ABBRID0EDZAK" xlink_title="Webers Length of Mesosoma">WL</abbrev>
0.87); lateral arms of hypostomal carina strongly diverging anteriorly, relatively thick, and strongly angulate at widest points (Fig.
<figureCitation id="D045C2499458EC5EBAA1FDF926E339C6" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Diagnostic plate showing still images from surface volume renderings of the head in ventral view (the remainder of the body virtually removed) (hypostoma in semi-transparent red; postgenal sulcus in semi-transparent yellow; postoccipital carina in semi-transparent green) A Z. aprilia sp. nov. holotype (CASENT 0764763) B Z. kouakoui sp. nov. paratype (CASENT 0764653) C Z. lolae sp. nov. paratype (CASENT 0764651) D Z. lumumbai sp. nov. holotype (MRACFOR 0010007) E Z. ndouri sp. nov. holotype (KGCOL 01883) F Z. obamai Hita Garcia, 2017 holotype (CASENT 0764125) G Z. sarowiwai Hita Garcia, 2017 paratype (CASENT 0764650) H Z. wilsoni Hita Garcia, 2017 holotype (MCZ-ENT 00512764)." figureDoi="10.3897/zookeys.1223.131238.figure8" httpUri="https://binary.pensoft.net/fig/1213422">8 H</figureCitation>
); postgenal sulcus weakly impressed and running halfway to occipital margin (Fig.
<figureCitation id="4AE08DE19A139D9C725021E6FD41ABD2" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Diagnostic plate showing still images from surface volume renderings of the head in ventral view (the remainder of the body virtually removed) (hypostoma in semi-transparent red; postgenal sulcus in semi-transparent yellow; postoccipital carina in semi-transparent green) A Z. aprilia sp. nov. holotype (CASENT 0764763) B Z. kouakoui sp. nov. paratype (CASENT 0764653) C Z. lolae sp. nov. paratype (CASENT 0764651) D Z. lumumbai sp. nov. holotype (MRACFOR 0010007) E Z. ndouri sp. nov. holotype (KGCOL 01883) F Z. obamai Hita Garcia, 2017 holotype (CASENT 0764125) G Z. sarowiwai Hita Garcia, 2017 paratype (CASENT 0764650) H Z. wilsoni Hita Garcia, 2017 holotype (MCZ-ENT 00512764)." figureDoi="10.3897/zookeys.1223.131238.figure8" httpUri="https://binary.pensoft.net/fig/1213422">8 F</figureCitation>
); postoccipital margin in ventral view with anterior outline moderately or weakly and irregularly defined; anterolateral projections angulate (Fig.
<figureCitation id="D6B7F3E2F751E8C78E32307EB87281D5" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Diagnostic plate showing still images from surface volume renderings of the head in ventral view (the remainder of the body virtually removed) (hypostoma in semi-transparent red; postgenal sulcus in semi-transparent yellow; postoccipital carina in semi-transparent green) A Z. aprilia sp. nov. holotype (CASENT 0764763) B Z. kouakoui sp. nov. paratype (CASENT 0764653) C Z. lolae sp. nov. paratype (CASENT 0764651) D Z. lumumbai sp. nov. holotype (MRACFOR 0010007) E Z. ndouri sp. nov. holotype (KGCOL 01883) F Z. obamai Hita Garcia, 2017 holotype (CASENT 0764125) G Z. sarowiwai Hita Garcia, 2017 paratype (CASENT 0764650) H Z. wilsoni Hita Garcia, 2017 holotype (MCZ-ENT 00512764)." figureDoi="10.3897/zookeys.1223.131238.figure8" httpUri="https://binary.pensoft.net/fig/1213422">8 H</figureCitation>
); pleural endophragmal pit very weakly developed and inconspicuous (Fig.
<figureCitation id="03FF174BAC490C28D033A2BCA6868068" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Diagnostic plate showing still images from surface volume renderings of the mesosoma in profile view (the remainder of the body virtually removed) (pleural endophragmal pit concavity in semi-transparent yellow) A Z. aprilia sp. nov. holotype (CASENT 0764763) B Z. kouakoui sp. nov. paratype (CASENT 0764653) C Z. lolae sp. nov. holotype (KGCOL 02270) D Z. lumumbai sp. nov. holotype (MRACFOR 0010007) E Z. ndouri sp. nov. holotype (KGCOL 01883) F Z. obamai Hita Garcia, 2017 holotype (CASENT 0764125) G Z. sarowiwai Hita Garcia, 2017 paratype (CASENT 0764650) H Z. wilsoni Hita Garcia, 2017 holotype (MCZ-ENT 00512764)." figureDoi="10.3897/zookeys.1223.131238.figure10" httpUri="https://binary.pensoft.net/fig/1213424">10 H</figureCitation>
); subpetiolar process of petiole (
<collectingCountry id="9411724BF1B76A9AC84AACF2294D1675" name="American Samoa">AS</collectingCountry>
II) in profile with thickened anterior and ventral margins and weak concavity without differentiated fenestra (Fig.
<figureCitation id="10A787EE2BA8C891F7FA1736CB893D6F" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Diagnostic plate showing still images from surface volume renderings of the tergum of AS II (petiole) in profile view (the remainder of the body virtually removed) (tergum in semi-transparent red, laterotergite in semi-transparent yellow) A Z. aprilia sp. nov. holotype (CASENT 0764763) B Z. kouakoui sp. nov. paratype (CASENT 0764653) C Z. lolae sp. nov. holotype (KGCOL 02270) D Z. lumumbai sp. nov. holotype (MRACFOR 0010007) E Z. ndouri sp. nov. holotype (KGCOL 01883) F Z. obamai Hita Garcia, 2017 holotype (CASENT 0764125) G Z. sarowiwai Hita Garcia, 2017 paratype (CASENT 0764650) H Z. wilsoni Hita Garcia, 2017 holotype (MCZ-ENT 00512764)." figureDoi="10.3897/zookeys.1223.131238.figure13" httpUri="https://binary.pensoft.net/fig/1213427">13 H</figureCitation>
); posterior end of abdominal segment III in ventral view with transverse groove weak to absent, instead with irregular groves and rugosity (Fig.
<figureCitation id="14876FEF04DD0E94735483942FAFD9D2" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Diagnostic plate showing still images from surface volume renderings of the sternum of AS III in ventral view (the remainder of the body virtually removed) (Bum in semi-transparent yellow, prora in semi-transparent red) A Z. aprilia sp. nov. holotype (CASENT 0764763) B Z. kouakoui sp. nov. paratype (CASENT 0764653) C Z. lolae sp. nov. paratype (CASENT 0764651) D Z. lumumbai sp. nov. holotype (MRACFOR 0010007) E Z. ndouri sp. nov. holotype (KGCOL 01883) F Z. obamai Hita Garcia, 2017 holotype (CASENT 0764125) G Z. sarowiwai Hita Garcia, 2017 paratype (CASENT 0764650) H Z. wilsoni Hita Garcia, 2017 holotype (MCZ-ENT 00512764)." figureDoi="10.3897/zookeys.1223.131238.figure16" httpUri="https://binary.pensoft.net/fig/1213430">16 H</figureCitation>
); prora in anteroventral view very weakly developed with almost absent lateroventral margins (Fig.
<figureCitation id="108C4625C820FBAFC15DA0E2081D84FE" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Diagnostic plate showing still images from surface volume renderings of the sternum of AS III in ventral view (the remainder of the body virtually removed) (Bum in semi-transparent yellow, prora in semi-transparent red) A Z. aprilia sp. nov. holotype (CASENT 0764763) B Z. kouakoui sp. nov. paratype (CASENT 0764653) C Z. lolae sp. nov. paratype (CASENT 0764651) D Z. lumumbai sp. nov. holotype (MRACFOR 0010007) E Z. ndouri sp. nov. holotype (KGCOL 01883) F Z. obamai Hita Garcia, 2017 holotype (CASENT 0764125) G Z. sarowiwai Hita Garcia, 2017 paratype (CASENT 0764650) H Z. wilsoni Hita Garcia, 2017 holotype (MCZ-ENT 00512764)." figureDoi="10.3897/zookeys.1223.131238.figure16" httpUri="https://binary.pensoft.net/fig/1213430">16 H</figureCitation>
); surface sculpture on genae mostly smooth and shiny, on cephalic dorsum mostly reticulate-rugose (Figs
<figureCitation id="0B05114F13884AB2721A7DECB6D10C45" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Diagnostic plate showing still images from surface volume renderings of the head in full-face view (the remainder of the body virtually removed) (torular-posttorular complex in semi-transparent red; anterior projections of parafrontal ridges in semi-transparent yellow) A Z. aprilia sp. nov. holotype (CASENT 0764763) B Z. kouakoui sp. nov. paratype (CASENT 0764653) C Z. lolae sp. nov. holotype (KGCOL 02270) D Z. lumumbai sp. nov. holotype (MRACFOR 0010007) E Z. ndouri sp. nov. holotype (KGCOL 01883) F Z. obamai Hita Garcia, 2017 holotype (CASENT 0764125) G Z. sarowiwai Hita Garcia, 2017 holotype (CASENT 0764654) H Z. wilsoni Hita Garcia, 2017 holotype (MCZ-ENT 00512764)." figureDoi="10.3897/zookeys.1223.131238.figure4" httpUri="https://binary.pensoft.net/fig/1213418">4 H</figureCitation>
,
<figureCitation id="06A40BC66BF493793ED512738C2E7840" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Diagnostic plate showing still images from surface volume renderings of the head in profile view (the remainder of the body virtually removed) (torular-posttorular complex in semi-transparent red) A Z. aprilia sp. nov. holotype (CASENT 0764763) B Z. kouakoui sp. nov. paratype (CASENT 0764653) C Z. lolae sp. nov. paratype (CASENT 0764651) D Z. lumumbai sp. nov. holotype (KGCOL 02270) E Z. ndouri sp. nov. holotype (KGCOL 01883) F Z. obamai Hita Garcia, 2017 holotype (CASENT 0764125) G Z. sarowiwai Hita Garcia, 2017 holotype (CASENT 0764654) H Z. wilsoni Hita Garcia, 2017 holotype (MCZ-ENT 00512764)." figureDoi="10.3897/zookeys.1223.131238.figure5" httpUri="https://binary.pensoft.net/fig/1213419">5 H</figureCitation>
,
<figureCitation id="1C9728CAAF354A5104F94C1EACCE27DE" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Diagnostic plate showing head in full-face view of all species treated herein (stacked colour images) A Z. aprilia sp. nov. holotype (CASENT 0764763) B Z. kouakoui sp. nov. holotype (KGCOL 00589) C Z. lolae sp. nov. holotype (KGCOL 02270) D Z. lumumbai sp. nov. holotype (MRACFOR 0010007) E Z. ndouri sp. nov. holotype (KGCOL 01883) F Z. obamai Hita Garcia, 2017 holotype (CASENT 0764125) G Z. sarowiwai Hita Garcia, 2017 paratype (CASENT 0764650) H Z. wilsoni Hita Garcia, 2017 holotype (MCZ-ENT 00512764)." figureDoi="10.3897/zookeys.1223.131238.figure19" httpUri="https://binary.pensoft.net/fig/1213433">19 H</figureCitation>
,
<figureCitation id="633FBD95E706DEE042A3231ABE084FC0" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Diagnostic plate showing full body in profile view of all species treated herein (stacked colour images) A Z. aprilia sp. nov. holotype (CASENT 0764763) B Z. kouakoui sp. nov. holotype (KGCOL 00589) C Z. lolae sp. nov. holotype (KGCOL 02270) D Z. lumumbai sp. nov. holotype (MRACFOR 0010007) E Z. ndouri sp. nov. holotype (KGCOL 01883) F Z. obamai Hita Garcia, 2017 holotype (CASENT 0764125) G Z. sarowiwai Hita Garcia, 2017 holotype (CASENT 0764654) H Z. wilsoni Hita Garcia, 2017 holotype (MCZ-ENT 00512764)." figureDoi="10.3897/zookeys.1223.131238.figure20" httpUri="https://binary.pensoft.net/fig/1213434">20 H</figureCitation>
); general surface sculpture on mesosoma and metasoma mostly smooth and shiny with abundant piliferous foveae, mesosoma and petiole laterally mostly reticulate-punctate, hypopygidium reticulate-rugose (Figs
<figureCitation id="6C63ACC5D4C8ABABB32D47E920CC1BC2" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Diagnostic plate showing full body in profile view of all species treated herein (stacked colour images) A Z. aprilia sp. nov. holotype (CASENT 0764763) B Z. kouakoui sp. nov. holotype (KGCOL 00589) C Z. lolae sp. nov. holotype (KGCOL 02270) D Z. lumumbai sp. nov. holotype (MRACFOR 0010007) E Z. ndouri sp. nov. holotype (KGCOL 01883) F Z. obamai Hita Garcia, 2017 holotype (CASENT 0764125) G Z. sarowiwai Hita Garcia, 2017 holotype (CASENT 0764654) H Z. wilsoni Hita Garcia, 2017 holotype (MCZ-ENT 00512764)." figureDoi="10.3897/zookeys.1223.131238.figure20" httpUri="https://binary.pensoft.net/fig/1213434">20 H</figureCitation>
,
<figureCitation id="C01EC0DE4E52642515AECB2687B60F50" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Diagnostic plate showing full body in dorsal view of all species treated herein (stacked colour images) A Z. aprilia sp. nov. holotype (CASENT 0764763) B Z. kouakoui sp. nov. holotype (KGCOL 00589) C Z. lolae sp. nov. holotype (KGCOL 02270) D Z. lumumbai sp. nov. holotype (MRACFOR 0010007) E Z. ndouri sp. nov. holotype (KGCOL 01883) F Z. obamai Hita Garcia, 2017 holotype (CASENT 0764125) G Z. sarowiwai Hita Garcia, 2017 holotype (CASENT 0764654) H Z. wilsoni Hita Garcia 2017 holotype (MCZ-ENT 00512764)." figureDoi="10.3897/zookeys.1223.131238.figure21" httpUri="https://binary.pensoft.net/fig/1213435">21 H</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0E21AK" type="description">
<paragraph id="2F33B069E9EF15FD27633D1F08E822D5">
<heading id="3FBF5F1C7F3605183BADABE68D45D5EF" reason="title">Measurements and indices.</heading>
</paragraph>
<paragraph id="6D721FB131C1B4FC4C486BBDDF6F51F1">
Morphometric data is based on the singleton
<typeStatus id="937707C206F7F077F637F9585EB585CA" type="holotype">holotype</typeStatus>
from
<collectingCountry id="3847C87EB0A45984A5229B2EE2E3630F" name="Mozambique">Mozambique</collectingCountry>
and can be seen in Table
<tableCitation id="844E819CE22AB213A3F5EB5193329587" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Comparative data of measurements and indices used for the eight species of Afrotropical Zasphinctus (raw data is available in Suppl. material 3)." httpUri="http://table.plazi.org/id/8548FBEC1F3F4349758B2AB18390E1E9" tableUuid="8548FBEC1F3F4349758B2AB18390E1E9">2</tableCitation>
, Suppl. material
<xref id="165030F316A57CA0F992558B491F2E31" ref-type="supplementary-material" rid="S3">3</xref>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EJ2AK" type="distribution">
<paragraph id="6383B855CC91AF7A7F2F271FCB5736F0">
<heading id="59847D8A24FF0D9FD28382856005D570" reason="title">Distribution and biology.</heading>
</paragraph>
<paragraph id="1265B191811A000EEB3A48749DA6082F">
Fortunately, our knowledge of the distribution of
<taxonomicName id="6C542FF6C6708D2B5A20EFC8B8899E67" authorityName="Hita Garcia" authorityYear="2017" class="Insecta" family="Formicidae" genus="Zasphinctus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="wilsoni">
<emphasis id="EB077DAB9390713C4018D0E8341CF61D" italics="true">Z. wilsoni</emphasis>
</taxonomicName>
has increased since its original description. Whereas in
<bibRefCitation id="C5C27018B30911F71F7B5F80550A4F4C" DOI="10.3897/zookeys.693.13012" author="Hita Garcia F &amp; Fischer G &amp; Liu C &amp; Audisio TL &amp; Economo EP" journalOrPublisher="ZooKeys" pagination="33-93" refId="B32" refString="Hita Garcia F, Fischer G, Liu C, Audisio TL, Economo EP (2017 a) Next-generation morphological character discovery and evaluation: an X-ray micro-CT enhanced revision of the ant genus Zasphinctus Wheeler (Hymenoptera, Formicidae, Dorylinae) in the Afrotropics. ZooKeys 693: 3393. https://doi.org/10.3897/zookeys.693.13012" title="Next-generation morphological character discovery and evaluation: an X-ray micro-CT enhanced revision of the ant genus Zasphinctus Wheeler (Hymenoptera, Formicidae, Dorylinae) in the Afrotropics." volume="693">Hita Garcia et al. (2017 a</bibRefCitation>
) it was only known from its type locality, the Gorongosa National Park, now it also known from two additional localities considerably further northeast, namely Quirimbas in
<collectingRegion id="ADC5EDFF7992842EF80C83F5E6386058" country="Mozambique" name="Cabo Delgado">Cabo Delgado</collectingRegion>
and Mount Mabu in
<collectingRegion id="B9F08E0F1F2BF7A4EAE118A51F319AFC" country="Mozambique" name="Zambezia">Zambezia</collectingRegion>
. The species was collected in litter in Gorongosa and in soil in the other two localities. To our surprise it seems that
<taxonomicName id="0953B233E073E61B998738F889FA9392" authorityName="Hita Garcia" authorityYear="2017" class="Insecta" family="Formicidae" genus="Zasphinctus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="wilsoni">
<emphasis id="2C7E0D5EBCF18D6C5F92D538AC2451D6" italics="true">Z. wilsoni</emphasis>
</taxonomicName>
is relatively flexible in its habitat requirements since it was collected in secondary dry forest, miombo woodland and rainforest.
</paragraph>
<caption id="8ABBAC6B09C7AC14CCDBADF7E671729A" ID-DOI="10.3897/zookeys.1223.131238.figure24" ID-arpha="14A3445A-C1AD-5CD6-97A4-38A8EE372693" httpUri="https://binary.pensoft.net/fig/1213438" startId="F24">
<paragraph id="B15BC7B73CC0B330FF47B5377112811F">
<label id="DA2220F82F9324A5CD4C62D4101E5EF6">Figure 24.</label>
</paragraph>
<paragraph id="95D8420EE3F9747DCB8008249C68A065">
Shaded surface display volume renderings of 3 D models of
<taxonomicName id="63ED35303E286A624F75800418E181CB" authority="Hita Garcia, 2017" authorityName="Hita Garcia" authorityYear="2017" class="Insecta" family="Formicidae" genus="Zasphinctus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="wilsoni">
<emphasis id="8525AFB098E083571EA7B0BB626C0C28" italics="true">Z. wilsoni</emphasis>
Hita Garcia, 2017
</taxonomicName>
holotype (
<ext-link id="628E7BC9B58A393DDAF7C7F0B2404910" ext-link-type="uri" xlink_href="https://www.antweb.org/specimen.do?code=mcz-ent00512764" xlink_type="simple">MCZ-ENT 00512764</ext-link>
)
<emphasis id="C0EE316C94DE84FA75EB659E2C6288A1" bold="true">A</emphasis>
full body in profile
<emphasis id="68189764B58C3DA90EEC14B4132EBAF7" bold="true">B</emphasis>
full body in dorsal view
<emphasis id="B070624CAA4229395119B1A436CAAC8C" bold="true">C</emphasis>
head in full-face view (with antennae)
<emphasis id="C0F246FD1E906846404AB1AC7261BD9E" bold="true">D</emphasis>
head in full-face view (without antennae)
<emphasis id="6EB2C6A711A363CE4A3A0000DC34CF03" bold="true">E</emphasis>
head in ventral view
<emphasis id="939CB41488020982AC473D20A342AA80" bold="true">F</emphasis>
abdominal segment II (petiole) in profile
<emphasis id="45F5ABDD2CDAB72A37923A43DDF790DF" bold="true">G</emphasis>
abdominal segment II (petiole) in dorsal view
<emphasis id="18B774C32AC81E79AC5870D845EAD066" bold="true">H</emphasis>
tergum of AS III in dorsal view
<emphasis id="37FDE0D3E578199F4A12556CC133F956" bold="true">I</emphasis>
sternum of AS III in ventral view.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>