treatments-xml/data/E5/2A/87/E52A87BCFFAAFFB1FF79F89B89146D38.xml
2024-06-21 12:55:36 +02:00

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<document id="90B4349F61B1C96D5232F4533F3DDCE0" ID-CLB-Dataset="7784" ID-DOI="10.11646/zootaxa.4933.4.1" ID-GBIF-Dataset="2ffdac24-b2ae-448c-b88e-b4f0dd1a9547" ID-ISSN="1175-5326" ID-Zenodo-Dep="4556546" ID-ZooBank="1FC0D2E0-888E-4F79-ABFE-BC7E91ADEECE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1614066053014" checkinUser="plazi" docAuthor="Wood, Thomas James" docDate="2021" docId="E52A87BCFFAAFFB1FF79F89B89146D38" docLanguage="en" docName="zootaxa.4933.4.1.pdf" docOrigin="Zootaxa 4933 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Andrena (Truncandrena) syriensis Wood 2021, spec. nov." docType="treatment" docUuid="E653EFDF-CD62-4021-A7AD-FCE0D9AEE618" docUuidSource="ZooBank" docVersion="7" lastPageNumber="491" masterDocId="1913FFC4FF89FF99FFEEFFE18A626C79" masterDocTitle="Fifteen new Andrena species from little-visited arid, Mediterranean, and mountainous parts of the Old World (Hymenoptera: Andrenidae)" masterLastPageNumber="492" masterPageNumber="451" pageNumber="486" updateTime="1698930763140" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="97425EA3C4E3D89D524363F85309E06E">Fifteen new Andrena species from little-visited arid, Mediterranean, and mountainous parts of the Old World (Hymenoptera: Andrenidae)</mods:title>
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<mods:namePart id="F325E0F4A22EB3E2BCE1112BA5D4EFAD">Wood, Thomas James</mods:namePart>
<mods:affiliation id="D8C40C2497E7367F96D0F6463C9A9D77">Laboratoire de Zoologie, Université de Mons, Mons, 7000, Belgium</mods:affiliation>
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<mods:date id="1226ADFE35A2D99B344794E575B265CE">2021</mods:date>
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<treatment id="E52A87BCFFAAFFB1FF79F89B89146D38" ID-DOI="http://doi.org/10.5281/zenodo.4599240" ID-GBIF-Taxon="180326862" ID-Zenodo-Dep="4599240" LSID="urn:lsid:zoobank.org:act:E653EFDF-CD62-4021-A7AD-FCE0D9AEE618" httpUri="http://treatment.plazi.org/id/E52A87BCFFAAFFB1FF79F89B89146D38" lastPageId="40" lastPageNumber="491" pageId="35" pageNumber="486">
<subSubSection id="25996521FFAAFFBAFF79F89B89046BC8" pageId="35" pageNumber="486" type="nomenclature">
<paragraph id="6D3C36AAFFAAFFBAFF79F89B88D86BEC" blockId="35.[151,870,1914,1969]" box="[151,698,1914,1941]" pageId="35" pageNumber="486">
<heading id="367481C6FFAAFFBAFF79F89B88D86BEC" bold="true" box="[151,698,1914,1941]" fontSize="11" level="1" pageId="35" pageNumber="486" reason="1">
<emphasis id="5FF7EAB8FFAAFFBAFF79F89B88D86BEC" bold="true" box="[151,698,1914,1941]" pageId="35" pageNumber="486">
<taxonomicName id="AA834D29FFAAFFBAFF79F89B88586BEC" authority="Wood, 2021" authorityName="Wood" authorityYear="2021" box="[151,570,1914,1941]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="486" phylum="Arthropoda" rank="species" species="syriensis" status="sp. nov." subGenus="Truncandrena">
<emphasis id="5FF7EAB8FFAAFFBAFF79F89B8B636BEC" bold="true" box="[151,257,1914,1941]" italics="true" pageId="35" pageNumber="486">Andrena</emphasis>
(
<emphasis id="5FF7EAB8FFAAFFBAFEFCF89B8BA36BEC" bold="true" box="[274,449,1914,1941]" italics="true" pageId="35" pageNumber="486">Truncandrena</emphasis>
)
<emphasis id="5FF7EAB8FFAAFFBAFE3CF89A88586BEC" bold="true" box="[466,570,1915,1941]" italics="true" pageId="35" pageNumber="486">syriensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="44C457C3FFAAFFBAFDACF89A88D86BEC" box="[578,698,1915,1941]" pageId="35" pageNumber="486" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="6D3C36AAFFAAFFBAFF79F87A89046BC8" blockId="35.[151,870,1914,1969]" box="[151,870,1947,1969]" pageId="35" pageNumber="486">
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urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
</subSubSection>
<subSubSection id="25996521FFAAFFBDFF79F8018ED36CCA" lastPageId="36" lastPageNumber="487" pageId="35" pageNumber="486" type="materials_examined">
<paragraph id="6D3C36AAFFAAFFBAFF79F8018F776B85" blockId="35.[151,1301,2016,2044]" box="[151,1301,2016,2044]" pageId="35" pageNumber="486">
<materialsCitation id="DDEB3CF7FFAAFFBAFF79F8018F776B85" ID-GBIF-Occurrence="3050682369" box="[151,1301,2016,2044]" collectingDate="1988-03-24" collectionCode="OOLM" collectorName="L. Blank" country="Syria" elevation="250" location="Homs" pageId="35" pageNumber="486" specimenCount="1" stateProvince="Hims" typeStatus="holotype">
<emphasis id="5FF7EAB8FFAAFFBAFF79F8018BC56B83" bold="true" box="[151,423,2016,2042]" pageId="35" pageNumber="486">
<typeStatus id="B2388808FFAAFFBAFF79F8018B586B83" box="[151,314,2016,2042]" pageId="35" pageNumber="486" type="holotype">HOLOTYPE</typeStatus>
:
<collectingCountry id="1594763AFFAAFFBAFEA8F8018BC36B83" box="[326,417,2016,2042]" name="Syria" pageId="35" pageNumber="486">SYRIA</collectingCountry>
:
</emphasis>
<collectingRegion id="AF47F848FFAAFFBAFE40F8008B966B82" box="[430,500,2017,2043]" country="Syria" name="Hims" pageId="35" pageNumber="486">Homs</collectingRegion>
,
<quantity id="AA7B9B4FFFAAFFBAFE11F801882A6B82" box="[511,584,2016,2043]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="35" pageNumber="486" unit="m" value="250.0">
<elevation id="E6AED199FFAAFFBAFE11F801882A6B82" box="[511,584,2016,2043]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="35" pageNumber="486" unit="m" value="250.0">250 m</elevation>
</quantity>
, As-Shuknah,
<quantity id="AA7B9B4FFFAAFFBAFD04F80189536B83" box="[746,817,2016,2042]" metricMagnitude="4" metricUnit="m" metricValue="2.2" pageId="35" pageNumber="486" unit="km" value="22.0">22 km</quantity>
E,
<date id="193D106AFFAAFFBAFCB9F80189B16B83" box="[855,979,2016,2042]" pageId="35" pageNumber="486" value="1988-03-24">
<collectingDate id="0979E982FFAAFFBAFCB9F80189B16B83" box="[855,979,2016,2042]" pageId="35" pageNumber="486" value="1988-03-24">24.iii.1988</collectingDate>
</date>
, 1³, leg.
<collectorName id="C076537CFFAAFFBAFBAEF8008EC06B83" box="[1088,1186,2016,2043]" pageId="35" pageNumber="486">L. Blank</collectorName>
(
<collectionCode id="0B92AE6FFFAAFFBAFB5FF8008F686B85" box="[1201,1290,2017,2044]" pageId="35" pageNumber="486">OÖLM</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="6D3C36AAFFADFFBDFF29FF778ED36CCA" blockId="36.[199,1201,150,212]" box="[199,1201,150,179]" pageId="36" pageNumber="487">
<materialsCitation id="DDEB3CF7FFADFFBDFF29FF778ECF6CCA" ID-GBIF-Occurrence="3050682334" box="[199,1197,150,179]" collectingDate="1988-04-01" collectionCode="OOLM" collectorName="S. M. Blank" country="Syria" elevation="400" location="Homs" pageId="36" pageNumber="487" specimenCount="1" stateProvince="Hims" typeStatus="paratype">
<emphasis id="5FF7EAB8FFADFFBDFF29FF778BAE6CC9" bold="true" box="[199,460,150,176]" pageId="36" pageNumber="487">
<typeStatus id="B2388808FFADFFBDFF29FF778B026CC9" box="[199,352,150,176]" pageId="36" pageNumber="487" type="paratype">PARATYPE</typeStatus>
:
<collectingCountry id="1594763AFFADFFBDFE82FF778BA56CC9" box="[364,455,150,176]" name="Syria" pageId="36" pageNumber="487">SYRIA</collectingCountry>
:
</emphasis>
<collectingRegion id="AF47F848FFADFFBDFE3DFF76887B6CC8" box="[467,537,151,177]" country="Syria" name="Hims" pageId="36" pageNumber="487">Homs</collectingRegion>
, E
<quantity id="AA7B9B4FFFADFFBDFDD2FF7788E46CC9" box="[572,646,150,176]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="36" pageNumber="487" unit="km" value="20.0">20 km</quantity>
,
<quantity id="AA7B9B4FFFADFFBDFD7FFF7788B86CC8" box="[657,730,150,177]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="36" pageNumber="487" unit="m" value="400.0">
<elevation id="E6AED199FFADFFBDFD7FFF7788B86CC8" box="[657,730,150,177]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="36" pageNumber="487" unit="m" value="400.0">400 m</elevation>
</quantity>
,
<date id="193D106AFFADFFBDFD08FF77892D6CC9" box="[742,847,150,176]" pageId="36" pageNumber="487" value="1988-04-01">
<collectingDate id="0979E982FFADFFBDFD08FF77892D6CC9" box="[742,847,150,176]" pageId="36" pageNumber="487" value="1988-04-01">1.iv.1988</collectingDate>
</date>
, 1³, leg.
<collectorName id="C076537CFFADFFBDFC53FF778E5F6CC9" box="[957,1085,150,176]" pageId="36" pageNumber="487">S.M. Blank</collectorName>
(
<collectionCode id="0B92AE6FFFADFFBDFBA2FF798EC76CCA" box="[1100,1189,152,179]" pageId="36" pageNumber="487">OÖLM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="25996521FFADFFBCFF29FF5B8E036E18" lastPageId="37" lastPageNumber="488" pageId="36" pageNumber="487" type="description">
<paragraph id="6D3C36AAFFADFFBDFF29FF5B885D6CAD" blockId="36.[199,1201,150,212]" box="[199,575,186,212]" pageId="36" pageNumber="487">
<emphasis id="5FF7EAB8FFADFFBDFF29FF5B8BA06CAD" bold="true" box="[199,450,186,212]" pageId="36" pageNumber="487">Description: Female:</emphasis>
Unknown.
</paragraph>
<caption id="39FC6622FFADFFBDFF79F94D88586A91" ID-DOI="http://doi.org/10.5281/zenodo.4556592" ID-Zenodo-Dep="4556592" httpUri="https://zenodo.org/record/4556592/files/figure.png" pageId="36" pageNumber="487" startId="36.[151,264,1708,1732]" targetBox="[151,1436,236,1685]" targetPageId="36">
<paragraph id="6D3C36AAFFADFFBDFF79F94D88586A91" blockId="36.[151,1437,1708,1769]" pageId="36" pageNumber="487">
<emphasis id="5FF7EAB8FFADFFBDFF79F94D8B0C6ABD" bold="true" box="[151,366,1708,1732]" pageId="36" pageNumber="487">FIGURES 115120.</emphasis>
<taxonomicName id="AA834D29FFADFFBDFE9BF94D88416ABD" authority="Wood, 2021" authorityName="Wood" authorityYear="2021" box="[373,547,1708,1732]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="487" phylum="Arthropoda" rank="species" species="zagrosa" status="sp. nov.">
<emphasis id="5FF7EAB8FFADFFBDFE9BF94D88416ABD" box="[373,547,1708,1732]" italics="true" pageId="36" pageNumber="487">Andrena zagrosa</emphasis>
</taxonomicName>
<emphasis id="5FF7EAB8FFADFFBDFDC5F94C88F66ABC" bold="true" box="[555,660,1709,1733]" pageId="36" pageNumber="487">
<taxonomicNameLabel id="44C457C3FFADFFBDFDC5F94C88F66ABC" box="[555,660,1709,1733]" pageId="36" pageNumber="487" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
115. male profile, 116. male face, 117. male scutum, 118. male terga, 119. male genitalia, 120. male genitalia detail.
</paragraph>
</caption>
<paragraph id="6D3C36AAFFADFFBCFF29F8F68E036E18" blockId="36.[151,1436,1814,2021]" lastBlockId="37.[151,1437,150,609]" lastPageId="37" lastPageNumber="488" pageId="36" pageNumber="487">
<emphasis id="5FF7EAB8FFADFFBDFF29F8F68B6F6B48" bold="true" box="[199,269,1815,1841]" pageId="36" pageNumber="487">Male:</emphasis>
Body length 1010.5 mm (
<figureCitation id="F5B82A2FFFADFFBDFDBBF8F688BA6B48" box="[597,728,1815,1841]" captionStart="FIGURES 131136" captionStartId="39.[151,264,1670,1694]" captionTargetBox="[151,1436,201,1646]" captionTargetId="figure-15@39.[151,1436,201,1646]" captionTargetPageId="39" captionText="FIGURES 131136. Andrena syriensis spec. nov. 131. male profile, 132. male clypeus detail, 133. male antenna detail, 134. male terga, 135. male genitalia detail, 136. male genitalia detail, dorsal view, arrow indicating lateral finger-like projections of the penis valve." figureDoi="http://doi.org/10.5281/zenodo.4556598" httpUri="https://zenodo.org/record/4556598/files/figure.png" pageId="36" pageNumber="487">Figure 131</figureCitation>
).
<emphasis id="5FF7EAB8FFADFFBDFD1EF8F689546B48" box="[752,822,1815,1841]" italics="true" pageId="36" pageNumber="487">Head:</emphasis>
Black, 1.2 times wider than long. Clypeus coloured yellow with exception of two lateral black marks, yellow colouration extending onto lower paraocular areas (
<figureCitation id="F5B82A2FFFADFFBDFA82F8DA8A936B00" captionStart="FIGURES 131136" captionStartId="39.[151,264,1670,1694]" captionTargetBox="[151,1436,201,1646]" captionTargetId="figure-15@39.[151,1436,201,1646]" captionTargetPageId="39" captionText="FIGURES 131136. Andrena syriensis spec. nov. 131. male profile, 132. male clypeus detail, 133. male antenna detail, 134. male terga, 135. male genitalia detail, 136. male genitalia detail, dorsal view, arrow indicating lateral finger-like projections of the penis valve." figureDoi="http://doi.org/10.5281/zenodo.4556598" httpUri="https://zenodo.org/record/4556598/files/figure.png" pageId="36" pageNumber="487">Figure 132</figureCitation>
). Clypeus regularly punctate, punctures laterally separated by 0.51 puncture diameter, becoming sparser centrally, here separated by 12 puncture diameters; underlying surface shagreened, weakly shining. Process of labrum trapezoidal, fore margin rounded emarginate. Gena equalling width of compound eye; ocelloccipital distance equalling width of lateral ocellus. Gena, vertex, face, and scape with long white hairs, not exceeding length of scape. Antenna dark, A4 apically, A513 lightened orange below, A3 exceeds A4, shorter than A4+5 (
<figureCitation id="F5B82A2FFFACFFBCFAE1FF768FF26CC8" box="[1295,1424,151,177]" captionStart="FIGURES 131136" captionStartId="39.[151,264,1670,1694]" captionTargetBox="[151,1436,201,1646]" captionTargetId="figure-15@39.[151,1436,201,1646]" captionTargetPageId="39" captionText="FIGURES 131136. Andrena syriensis spec. nov. 131. male profile, 132. male clypeus detail, 133. male antenna detail, 134. male terga, 135. male genitalia detail, 136. male genitalia detail, dorsal view, arrow indicating lateral finger-like projections of the penis valve." figureDoi="http://doi.org/10.5281/zenodo.4556598" httpUri="https://zenodo.org/record/4556598/files/figure.png" pageId="37" pageNumber="488">Figure 133</figureCitation>
).
<taxonomicName id="AA834D29FFACFFBCFF79FF5A8B156CAC" authority=": Scutum" authorityName="Scutum" box="[151,375,187,213]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="37" pageNumber="488" phylum="Arthropoda" rank="genus">
<emphasis id="5FF7EAB8FFACFFBCFF79FF5A8B7A6CAD" box="[151,280,187,212]" italics="true" pageId="37" pageNumber="488">Mesosoma:</emphasis>
Scutum
</taxonomicName>
and scutellum consistently punctate, punctures separated by 0.51 puncture diameter, underlying surface shagreened, weakly shining, shagreenation becoming weaker centrally, here shining more strongly. Pronotum rounded, without humeral angle. Episternum and propodeum microreticulate, weakly shining, becoming weakly rugose on propodeum, propodeal triangle indicated by change in surface sculpture, surface finely microreticulate. Episternum, propodeum, scutum, and scutellum covered with long whitish hairs, clearly exceeding length of scape. Legs dark, apical tarsal segment and hind basitarsi lightened orange, pubescence whitish. Wings hyaline, venation and stigma orange, nervulus strongly antefurcal.
<emphasis id="5FF7EAB8FFACFFBCFCF4FE7289FA6DD5" box="[794,920,403,428]" italics="true" pageId="37" pageNumber="488">Metasoma:</emphasis>
Terga dark, apical margins narrowly lightened yellow, partly hyaline (
<figureCitation id="F5B82A2FFFACFFBCFE73FE56887C6DA8" box="[413,542,439,465]" captionStart="FIGURES 131136" captionStartId="39.[151,264,1670,1694]" captionTargetBox="[151,1436,201,1646]" captionTargetId="figure-15@39.[151,1436,201,1646]" captionTargetPageId="39" captionText="FIGURES 131136. Andrena syriensis spec. nov. 131. male profile, 132. male clypeus detail, 133. male antenna detail, 134. male terga, 135. male genitalia detail, 136. male genitalia detail, dorsal view, arrow indicating lateral finger-like projections of the penis valve." figureDoi="http://doi.org/10.5281/zenodo.4556598" httpUri="https://zenodo.org/record/4556598/files/figure.png" pageId="37" pageNumber="488">Figure 134</figureCitation>
). Tergal discs finely and regularly punctate, punctures separated by 2 puncture diameters; terga covered with loose, long whitish hairs, forming denser hairbands on apical margins of T24, obscuring underlying surface. Genital capsule compact, gonocoxae with short, weakly pointed teeth, these diverging widely; gonostyli with apical blades flattened, triangular, outer face weakly concave, with long plumose hairs. Penis valve broad, laterally with pair of lamellate, finger-like projections (
<figureCitation id="F5B82A2FFFACFFBCFC7FFDA68E376E18" box="[913,1109,582,609]" captionStart="FIGURES 131136" captionStartId="39.[151,264,1670,1694]" captionTargetBox="[151,1436,201,1646]" captionTargetId="figure-15@39.[151,1436,201,1646]" captionTargetPageId="39" captionText="FIGURES 131136. Andrena syriensis spec. nov. 131. male profile, 132. male clypeus detail, 133. male antenna detail, 134. male terga, 135. male genitalia detail, 136. male genitalia detail, dorsal view, arrow indicating lateral finger-like projections of the penis valve." figureDoi="http://doi.org/10.5281/zenodo.4556598" httpUri="https://zenodo.org/record/4556598/files/figure.png" pageId="37" pageNumber="488">Figures 135136</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="39FC6622FFACFFBCFF79F9B48AA46AE8" ID-DOI="http://doi.org/10.5281/zenodo.4556594" ID-Zenodo-Dep="4556594" httpUri="https://zenodo.org/record/4556594/files/figure.png" pageId="37" pageNumber="488" startId="37.[151,264,1621,1645]" targetBox="[151,1436,633,1598]" targetPageId="37">
<paragraph id="6D3C36AAFFACFFBCFF79F9B48AA46AE8" blockId="37.[151,1436,1621,1681]" pageId="37" pageNumber="488">
<emphasis id="5FF7EAB8FFACFFBCFF79F9B48B0C6A14" bold="true" box="[151,366,1621,1645]" pageId="37" pageNumber="488">FIGURES 121124.</emphasis>
<taxonomicName id="AA834D29FFACFFBCFE9AF9B4884C6A14" authority="Wood, 2021" authorityName="Wood" authorityYear="2021" box="[372,558,1621,1645]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="37" pageNumber="488" phylum="Arthropoda" rank="species" species="inaquosa" status="sp. nov.">
<emphasis id="5FF7EAB8FFACFFBCFE9AF9B4884C6A14" box="[372,558,1621,1645]" italics="true" pageId="37" pageNumber="488">Andrena inaquosa</emphasis>
</taxonomicName>
<emphasis id="5FF7EAB8FFACFFBCFDDBF9B488FE6A14" bold="true" box="[565,668,1621,1645]" pageId="37" pageNumber="488">
<taxonomicNameLabel id="44C457C3FFACFFBCFDDBF9B488FE6A14" box="[565,668,1621,1645]" pageId="37" pageNumber="488" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
121. male profile, 122. male face, 123. male antenna detail, 124. male scutum.
</paragraph>
</caption>
<subSubSection id="25996521FFACFFBFFF29F95E88F96DD7" lastPageId="38" lastPageNumber="489" pageId="37" pageNumber="488" type="diagnosis">
<paragraph id="6D3C36AAFFACFFBCFF29F95E8B7A6BAF" blockId="37.[151,1436,1727,2042]" pageId="37" pageNumber="488">
<emphasis id="5FF7EAB8FFACFFBCFF29F95E8B266AA0" bold="true" box="[199,324,1727,1753]" pageId="37" pageNumber="488">Diagnosis:</emphasis>
Placement of
<taxonomicName id="AA834D29FFACFFBCFE0CF92188006AA0" authorityName="Wood" authorityYear="2021" box="[482,610,1728,1754]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="37" pageNumber="488" phylum="Arthropoda" rank="species" species="syriensis">
<emphasis id="5FF7EAB8FFACFFBCFE0CF92188006AA0" box="[482,610,1728,1754]" italics="true" pageId="37" pageNumber="488">A. syriensis</emphasis>
</taxonomicName>
within a subgenus is not immediately clear. The combination of very short malar space, granulose propodeal triangle with clear dorsal area (not declivous), rounded (uncarinate) pronotum, large body size (&gt;
<quantity id="AA7B9B4FFFACFFBCFE8EF8E68BC16B5B" box="[352,419,1799,1826]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="37" pageNumber="488" unit="mm" value="8.0">8 mm</quantity>
), and yellow clypeus with colouration extending onto the lower paraocular area would suggest placement in the
<taxonomicName id="AA834D29FFACFFBCFE7FF8CA887E6B3C" authorityName="Warncke" authorityYear="1968" box="[401,540,1835,1861]" class="Insecta" family="Andrenidae" genus="Nobandrena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="37" pageNumber="488" phylum="Arthropoda" rank="genus">
<emphasis id="5FF7EAB8FFACFFBCFE7FF8CA887E6B3C" box="[401,540,1835,1861]" italics="true" pageId="37" pageNumber="488">Nobandrena</emphasis>
</taxonomicName>
. However,
<taxonomicName id="AA834D29FFACFFBCFD4EF8CD89416B3C" authorityName="Wood" authorityYear="2021" box="[672,803,1836,1862]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="37" pageNumber="488" phylum="Arthropoda" rank="species" species="syriensis">
<emphasis id="5FF7EAB8FFACFFBCFD4EF8CD89416B3C" box="[672,803,1836,1862]" italics="true" pageId="37" pageNumber="488">A. syriensis</emphasis>
</taxonomicName>
lacks depressed tergal margins and the typical elongate genital capsule of
<taxonomicName id="AA834D29FFACFFBCFE8AF8AE8B8D6B10" authorityName="Warncke" authorityYear="1968" box="[356,495,1871,1897]" class="Insecta" family="Andrenidae" genus="Nobandrena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="37" pageNumber="488" phylum="Arthropoda" rank="genus">
<emphasis id="5FF7EAB8FFACFFBCFE8AF8AE8B8D6B10" box="[356,495,1871,1897]" italics="true" pageId="37" pageNumber="488">Nobandrena</emphasis>
</taxonomicName>
, making
<taxonomicName id="AA834D29FFACFFBCFDB6F8AE88946B10" box="[600,758,1871,1897]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="37" pageNumber="490" phylum="Arthropoda" rank="subGenus" subGenus="Truncandrena">
<emphasis id="5FF7EAB8FFACFFBCFDB6F8AE88946B10" box="[600,758,1871,1897]" italics="true" pageId="37" pageNumber="488">Truncandrena</emphasis>
</taxonomicName>
a better fit where, though less common, yellow facial markings can extend onto the lower paraocular area, and the overall more beak-like shape of the clypeus is a better fit, though it should be noted that the antefurcal nervulus is unusual for the
<taxonomicName id="AA834D29FFACFFBCFBECF8768EC26BC8" box="[1026,1184,1943,1969]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="37" pageNumber="490" phylum="Arthropoda" rank="subGenus" subGenus="Truncandrena">
<emphasis id="5FF7EAB8FFACFFBCFBECF8768EC26BC8" box="[1026,1184,1943,1969]" italics="true" pageId="37" pageNumber="488">Truncandrena</emphasis>
</taxonomicName>
(usually interstitial to postfurcal).
</paragraph>
<paragraph id="6D3C36AAFFACFFBFFF29F83E88F96DD7" blockId="37.[151,1436,1727,2042]" lastBlockId="38.[151,1437,151,430]" lastPageId="38" lastPageNumber="489" pageId="37" pageNumber="488">
Within the
<taxonomicName id="AA834D29FFACFFBCFEA9F83E8B876B80" box="[327,485,2015,2041]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="37" pageNumber="490" phylum="Arthropoda" rank="subGenus" subGenus="Truncandrena">
<emphasis id="5FF7EAB8FFACFFBCFEA9F83E8B876B80" box="[327,485,2015,2041]" italics="true" pageId="37" pageNumber="488">Truncandrena</emphasis>
</taxonomicName>
, based on genital construction
<taxonomicName id="AA834D29FFACFFBCFCA6F80189A86B80" authorityName="Wood" authorityYear="2021" box="[840,970,2016,2042]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="37" pageNumber="488" phylum="Arthropoda" rank="species" species="syriensis">
<emphasis id="5FF7EAB8FFACFFBCFCA6F80189A86B80" box="[840,970,2016,2042]" italics="true" pageId="37" pageNumber="488">A. syriensis</emphasis>
</taxonomicName>
can be placed closest to
<taxonomicName id="AA834D29FFACFFBFFB07F8018B296CC8" authority="Morawitz, 1878" authorityName="Morawitz" authorityYear="1878" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" lastPageId="38" lastPageNumber="489" order="Hymenoptera" pageId="37" pageNumber="488" phylum="Arthropoda" rank="species" species="rotundilabris">
<emphasis id="5FF7EAB8FFACFFBCFB07F8018FFE6B80" box="[1257,1436,2015,2042]" italics="true" pageId="37" pageNumber="488">A. rotundilabris</emphasis>
Morawitz, 1878
</taxonomicName>
(
<collectingCountry id="1594763AFFAFFFBFFEB4FF768BFE6CC8" box="[346,412,151,177]" name="Israel" pageId="38" pageNumber="489">Israel</collectingCountry>
,
<collectingCountry id="1594763AFFAFFFBFFE46FF76886F6CC8" box="[424,525,151,177]" name="Lebanon" pageId="38" pageNumber="489">Lebanon</collectingCountry>
,
<collectingCountry id="1594763AFFAFFFBFFDF6FF7688086CCB" box="[536,618,151,178]" name="Turkey" pageId="38" pageNumber="489">Turkey</collectingCountry>
) and
<taxonomicName id="AA834D29FFAFFFBFFD47FF7689D16CC8" authority="Warncke, 1969" authorityName="Warncke" authorityYear="1969" box="[681,947,151,177]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="489" phylum="Arthropoda" rank="species" species="ulula">
<emphasis id="5FF7EAB8FFAFFFBFFD47FF7689616CC8" box="[681,771,151,177]" italics="true" pageId="38" pageNumber="489">A. ulula</emphasis>
Warncke, 1969
</taxonomicName>
(
<collectingCountry id="1594763AFFAFFFBFFC2CFF768E666CC8" box="[962,1028,151,177]" name="Israel" pageId="38" pageNumber="489">Israel</collectingCountry>
,
<collectingCountry id="1594763AFFAFFFBFFBFEFF768E006CCB" box="[1040,1122,151,178]" name="Turkey" pageId="38" pageNumber="489">Turkey</collectingCountry>
) because of the broad penis valve that occupies most of the space between the gonostyli, and the broad apically triangular gonostyli. As well as different facial markings (both with yellow markings restricted to the clypeus), structurally
<taxonomicName id="AA834D29FFAFFFBFFB27FF3E8F296C81" authorityName="Wood" authorityYear="2021" box="[1225,1355,223,249]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="489" phylum="Arthropoda" rank="species" species="syriensis">
<emphasis id="5FF7EAB8FFAFFFBFFB27FF3E8F296C81" box="[1225,1355,223,249]" italics="true" pageId="38" pageNumber="489">A. syriensis</emphasis>
</taxonomicName>
differs in the position of the antefurcal nervulus (both with a postfurcal nervulus), the centrally weak shagreenation of the scutum (both with dull scuta, shagreenation uninform, not centrally shining), and most noticeably in the form of the gonocoxal teeth which are short, pointed, and broadly splayed laterally (pointed but only diverging weakly in
<taxonomicName id="AA834D29FFAFFFBFFA6AFEAA8B486DF0" authorityName="Morawitz" authorityYear="1878" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="489" phylum="Arthropoda" rank="species" species="rotundilabris">
<emphasis id="5FF7EAB8FFAFFFBFFA6AFEAA8B486DF0" italics="true" pageId="38" pageNumber="489">A. rotundilabris</emphasis>
</taxonomicName>
, apically rounded in
<taxonomicName id="AA834D29FFAFFFBFFDF9FE8E88126DF0" authorityName="Warncke" authorityYear="1969" box="[535,624,367,393]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="489" phylum="Arthropoda" rank="species" species="ulula">
<emphasis id="5FF7EAB8FFAFFFBFFDF9FE8E88126DF0" box="[535,624,367,393]" italics="true" pageId="38" pageNumber="489">A. ulula</emphasis>
</taxonomicName>
) and of the penis valve, which has laterally projecting, lamellate, fingerlike projections (absent in both other species).
</paragraph>
</subSubSection>
<caption id="39FC6622FFAFFFBFFF79F864890A6B9C" ID-DOI="http://doi.org/10.5281/zenodo.4556596" ID-Zenodo-Dep="4556596" httpUri="https://zenodo.org/record/4556596/files/figure.png" pageId="38" pageNumber="489" startId="38.[151,264,1925,1949]" targetBox="[151,1436,453,1886]" targetPageId="38">
<paragraph id="6D3C36AAFFAFFFBFFF79F864890A6B9C" blockId="38.[151,1436,1925,2022]" pageId="38" pageNumber="489">
<emphasis id="5FF7EAB8FFAFFFBFFF79F8648B0D6BE4" bold="true" box="[151,367,1925,1949]" pageId="38" pageNumber="489">FIGURES 125130.</emphasis>
<taxonomicName id="AA834D29FFAFFFBFFE99F86488506BE4" authority="Wood, 2021" authorityName="Wood" authorityYear="2021" box="[375,562,1925,1949]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="489" phylum="Arthropoda" rank="species" species="inaquosa" status="sp. nov.">
<emphasis id="5FF7EAB8FFAFFFBFFE99F86488506BE4" box="[375,562,1925,1949]" italics="true" pageId="38" pageNumber="489">Andrena inaquosa</emphasis>
</taxonomicName>
<emphasis id="5FF7EAB8FFAFFFBFFDD4F86788C16BE7" bold="true" box="[570,675,1926,1950]" pageId="38" pageNumber="489">
<taxonomicNameLabel id="44C457C3FFAFFFBFFDD4F86788C16BE7" box="[570,675,1926,1950]" pageId="38" pageNumber="489" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
125. male terga, 126. male sternite 8 profile, 127. male sternite 8 ventral view, 128. male genitalia, arrow indicating bulges on surface of penis valve.
<taxonomicName id="AA834D29FFAFFFBFFC75F8488E946BBB" authority="Dours, 1872" authorityName="Dours" authorityYear="1872" box="[923,1270,1961,1986]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="489" phylum="Arthropoda" rank="species" species="planiventris">
<emphasis id="5FF7EAB8FFAFFFBFFC75F8488E126BB8" box="[923,1136,1961,1985]" italics="true" pageId="38" pageNumber="489">Andrena planiventris</emphasis>
Dours, 1872
</taxonomicName>
; 129, male genitalia.
<taxonomicName id="AA834D29FFAFFFBFFF21F82C88F76B9C" authority="Warncke, 1969" authorityName="Warncke" authorityYear="1969" box="[207,661,1997,2021]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="489" phylum="Arthropoda" rank="subSpecies" species="cyanomicans" subSpecies="mirna">
<emphasis id="5FF7EAB8FFAFFFBFFF21F82C8B966B9C" box="[207,500,1997,2021]" italics="true" pageId="38" pageNumber="489">Andrena cyanomicans mirna</emphasis>
Warncke, 1969
</taxonomicName>
; 130. male genitalia.
</paragraph>
</caption>
<caption id="39FC6622FFAEFFBEFF79F9678B306A9F" ID-DOI="http://doi.org/10.5281/zenodo.4556598" ID-Zenodo-Dep="4556598" httpUri="https://zenodo.org/record/4556598/files/figure.png" pageId="39" pageNumber="490" startId="39.[151,264,1670,1694]" targetBox="[151,1436,201,1646]" targetPageId="39">
<paragraph id="6D3C36AAFFAEFFBEFF79F9678B306A9F" blockId="39.[151,1437,1670,1766]" pageId="39" pageNumber="490">
<emphasis id="5FF7EAB8FFAEFFBEFF79F9678B0D6AE7" bold="true" box="[151,367,1670,1694]" pageId="39" pageNumber="490">FIGURES 131136.</emphasis>
<taxonomicName id="AA834D29FFAEFFBEFE98F967884E6AE7" authority="Wood, 2021" authorityName="Wood" authorityYear="2021" box="[374,556,1670,1694]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="39" pageNumber="490" phylum="Arthropoda" rank="species" species="syriensis" status="sp. nov.">
<emphasis id="5FF7EAB8FFAEFFBEFE98F967884E6AE7" box="[374,556,1670,1694]" italics="true" pageId="39" pageNumber="490">Andrena syriensis</emphasis>
</taxonomicName>
<emphasis id="5FF7EAB8FFAEFFBEFDDDF96788F96AE7" bold="true" box="[563,667,1670,1694]" pageId="39" pageNumber="490">
<taxonomicNameLabel id="44C457C3FFAEFFBEFDDDF96788F96AE7" box="[563,667,1670,1694]" pageId="39" pageNumber="490" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
131. male profile, 132. male clypeus detail, 133. male antenna detail, 134. male terga, 135. male genitalia detail, 136. male genitalia detail, dorsal view, arrow indicating lateral finger-like projections of the penis valve.
</paragraph>
</caption>
<subSubSection id="25996521FFAEFFB1FF29F8F58B7D6D64" lastPageId="40" lastPageNumber="491" pageId="39" pageNumber="490" type="discussion">
<paragraph id="6D3C36AAFFAEFFB1FF29F8F58B7D6D64" blockId="39.[151,1437,1812,2019]" lastBlockId="40.[151,1436,151,321]" lastPageId="40" lastPageNumber="491" pageId="39" pageNumber="490">
<emphasis id="5FF7EAB8FFAEFFBEFF29F8F58B2D6B57" bold="true" box="[199,335,1812,1838]" pageId="39" pageNumber="490">Discussion:</emphasis>
The subgenus
<taxonomicName id="AA834D29FFAEFFBEFDEEF8F588FC6B57" box="[512,670,1812,1838]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="39" pageNumber="490" phylum="Arthropoda" rank="subGenus" subGenus="Truncandrena">
<emphasis id="5FF7EAB8FFAEFFBEFDEEF8F588FC6B57" box="[512,670,1812,1838]" italics="true" pageId="39" pageNumber="490">Truncandrena</emphasis>
</taxonomicName>
has its greatest diversity in the Eastern Mediterranean, and so the presence of another undescribed species here is not surprising. The formation of the genital capsule with reduced and laterally splayed gonocoxal teeth and the lateral extensions to the penis valve has not previously been described within the
<taxonomicName id="AA834D29FFAEFFBEFEE1F8618BCF6BE3" box="[271,429,1920,1946]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="39" pageNumber="490" phylum="Arthropoda" rank="subGenus" subGenus="Truncandrena">
<emphasis id="5FF7EAB8FFAEFFBEFEE1F8618BCF6BE3" box="[271,429,1920,1946]" italics="true" pageId="39" pageNumber="490">Truncandrena</emphasis>
</taxonomicName>
, and the antefurcal position of the nervulus is aberrant within the subgenus. However, the Middle East is home to some remarkable
<taxonomicName id="AA834D29FFAEFFBEFD89F84588A46BC7" box="[615,710,1956,1982]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="39" pageNumber="490" phylum="Arthropoda" rank="genus">
<emphasis id="5FF7EAB8FFAEFFBEFD89F84588A46BC7" box="[615,710,1956,1982]" italics="true" pageId="39" pageNumber="490">Andrena</emphasis>
</taxonomicName>
diversity that does not fit well with existing subgeneric concepts and traits, even when they can be placed in subgenera at all (e.g.
<bibRefCitation id="09124B5BFFAEFFBEFC8BF8298E556B9B" author="Pisanty, G. &amp; Scheuchl, E. &amp; Dorchin, N." box="[869,1079,1992,2019]" pageId="39" pageNumber="490" pagination="485 - 515" refId="ref23712" refString="Pisanty, G., Scheuchl, E. &amp; Dorchin, N. (2016) Eight new species of Andrena Fabricius (Hymenoptera: Apoidea: Andrenidae) from Israel - a Mediterranean hotspot for wild bees. Zootaxa, 4189 (3), 485 - 515. https: // doi. org / 10.11646 / zootaxa. 4189.3.3" type="journal article" year="2016">
Pisanty
<emphasis id="5FF7EAB8FFAEFFBEFC53F828898C6B9B" box="[957,1006,1992,2018]" italics="true" pageId="39" pageNumber="490">et al</emphasis>
. 2016
</bibRefCitation>
;
<bibRefCitation id="09124B5BFFAEFFBEFBAFF8288E1E6B9A" author="Pisanty, G. &amp; Scheuchl, E. &amp; Dorchin, N." box="[1089,1148,1993,2019]" pageId="39" pageNumber="490" pagination="151 - 188" refId="ref23775" refString="Pisanty, G., Scheuchl, E. &amp; Dorchin, N. (2018) Taxonomic review of the subgenus Andrena (Poecilandrena) (Hymenoptera: Andrenidae) in Israel and the Levant. Zootaxa, 4374 (2), 151 - 188. https: // doi. org / 10.11646 / zootaxa. 4374.2.1" type="journal article" year="2018">2018</bibRefCitation>
;
<bibRefCitation id="09124B5BFFAEFFBEFB69F8288EA16B9A" author="Pisanty, G. &amp; Richter, R. &amp; Martin, T. &amp; Dettman, J. &amp; Cardinal, S." box="[1159,1219,1993,2019]" pageId="39" pageNumber="490" refId="ref23835" refString="Pisanty, G., Richter, R., Martin, T., Dettman, J. &amp; Cardinal, S. (2020) Molecular phylogeny and historical biogeography of an- drenine bees (Hymenoptera: Andrenidae). bioRxiv. [published online] https: // doi. org / 10.1101 / 2020.06.09.103556" type="book" year="2020">2020</bibRefCitation>
; Wood 2020;
<bibRefCitation id="09124B5BFFAEFFB1FAB4F8298B466CCB" author="Wood, T. J. &amp; Boustani, M. &amp; Rasmont, P." lastPageId="40" lastPageNumber="491" pageId="39" pageNumber="490" pagination="279 - 312" refId="ref24553" refString="Wood, T. J., Boustani, M. &amp; Rasmont, P. (2020 b) A revision of the Andrena (Hymenoptera: Andrenidae) of Lebanon with the description of six new species. Annales de la Societe entomologique de France, 56, 279 - 312. https: // doi. org / 10.1080 / 00379271.2020.1794960" type="journal article" year="2020">
Wood
<emphasis id="5FF7EAB8FFA1FFB1FF79FF768AA86CC8" box="[151,202,151,177]" italics="true" pageId="40" pageNumber="491">et al</emphasis>
. 2020b
</bibRefCitation>
). The diversity of new taxa and their morphological peculiarities described both here and elsewhere recently (Wood 2020) from arid desertic regions of
<collectingCountry id="1594763AFFA1FFB1FD04FF5A89476CAC" box="[746,805,187,213]" name="Syria" pageId="40" pageNumber="491">Syria</collectingCountry>
suggests that molecular study of this fauna could shed further light of our understanding of the evolution of this genus, given its likely East Mediterranean origin (
<bibRefCitation id="09124B5BFFA1FFB1FAA6FF3E8B716D64" author="Pisanty, G. &amp; Richter, R. &amp; Martin, T. &amp; Dettman, J. &amp; Cardinal, S." pageId="40" pageNumber="491" refId="ref23835" refString="Pisanty, G., Richter, R., Martin, T., Dettman, J. &amp; Cardinal, S. (2020) Molecular phylogeny and historical biogeography of an- drenine bees (Hymenoptera: Andrenidae). bioRxiv. [published online] https: // doi. org / 10.1101 / 2020.06.09.103556" type="book" year="2020">
Pisanty
<emphasis id="5FF7EAB8FFA1FFB1FF79FEE28AAA6D64" box="[151,200,259,285]" italics="true" pageId="40" pageNumber="491">et al</emphasis>
. 2020
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="25996521FFA1FFB1FF29FEC689146D38" box="[199,886,295,321]" pageId="40" pageNumber="491" type="etymology">
<paragraph id="6D3C36AAFFA1FFB1FF29FEC689146D38" blockId="40.[151,1436,151,321]" box="[199,886,295,321]" pageId="40" pageNumber="491">
<emphasis id="5FF7EAB8FFA1FFB1FF29FEC68B306D38" bold="true" box="[199,338,295,321]" pageId="40" pageNumber="491">Etymology:</emphasis>
This species is named after the country of
<collectingCountry id="1594763AFFA1FFB1FCDDFEC689106D38" box="[819,882,295,321]" name="Syria" pageId="40" pageNumber="491">Syria</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>