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<document id="3071F4918CC2145ABD8B42D8C0E85639" ID-CLB-Dataset="7784" ID-DOI="10.11646/zootaxa.4933.4.1" ID-GBIF-Dataset="2ffdac24-b2ae-448c-b88e-b4f0dd1a9547" ID-ISSN="1175-5326" ID-Zenodo-Dep="4556546" ID-ZooBank="1FC0D2E0-888E-4F79-ABFE-BC7E91ADEECE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1614066053014" checkinUser="plazi" docAuthor="Wood, Thomas James" docDate="2021" docId="E52A87BCFF95FF84FF79FF7689036BD4" docLanguage="en" docName="zootaxa.4933.4.1.pdf" docOrigin="Zootaxa 4933 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Andrena (Orandrena) densissima Wood 2021, spec. nov." docType="treatment" docUuid="BEA79392-12BB-4796-872A-8CDD9C876A4A" docUuidSource="ZooBank" docVersion="7" lastPageNumber="480" masterDocId="1913FFC4FF89FF99FFEEFFE18A626C79" masterDocTitle="Fifteen new Andrena species from little-visited arid, Mediterranean, and mountainous parts of the Old World (Hymenoptera: Andrenidae)" masterLastPageNumber="492" masterPageNumber="451" pageNumber="479" updateTime="1698930763140" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="BE650535C6723742DC28EB457B374E4C">Fifteen new Andrena species from little-visited arid, Mediterranean, and mountainous parts of the Old World (Hymenoptera: Andrenidae)</mods:title>
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<mods:namePart id="69CA111E07AFDD1E3054C1544F64FD86">Wood, Thomas James</mods:namePart>
<mods:affiliation id="C6AA5F01446C391922DB60B51E286F8B">Laboratoire de Zoologie, Université de Mons, Mons, 7000, Belgium</mods:affiliation>
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<mods:date id="AB8B9308018721B403AC46FB186A6A30">2021</mods:date>
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<treatment id="E52A87BCFF95FF84FF79FF7689036BD4" ID-DOI="http://doi.org/10.5281/zenodo.4599194" ID-GBIF-Taxon="180326865" ID-Zenodo-Dep="4599194" LSID="urn:lsid:zoobank.org:act:BEA79392-12BB-4796-872A-8CDD9C876A4A" httpUri="http://treatment.plazi.org/id/E52A87BCFF95FF84FF79FF7689036BD4" lastPageId="29" lastPageNumber="480" pageId="28" pageNumber="479">
<subSubSection id="25996521FF95FF85FF79FF7689CF6CB7" pageId="28" pageNumber="479" type="nomenclature">
<paragraph id="6D3C36AAFF95FF85FF79FF7688CE6CC8" blockId="28.[151,941,151,206]" box="[151,684,151,178]" pageId="28" pageNumber="479">
<heading id="367481C6FF95FF85FF79FF7688CE6CC8" bold="true" box="[151,684,151,178]" fontSize="11" level="1" pageId="28" pageNumber="479" reason="1">
<emphasis id="5FF7EAB8FF95FF85FF79FF7688CE6CC8" bold="true" box="[151,684,151,178]" pageId="28" pageNumber="479">
<taxonomicName id="AA834D29FF95FF85FF79FF76884F6CCB" authority="Wood, 2021" authorityName="Wood" authorityYear="2021" box="[151,557,151,178]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="479" phylum="Arthropoda" rank="species" species="densissima" status="sp. nov." subGenus="Orandrena">
<emphasis id="5FF7EAB8FF95FF85FF79FF768B636CCB" bold="true" box="[151,257,151,178]" italics="true" pageId="28" pageNumber="479">Andrena</emphasis>
(
<emphasis id="5FF7EAB8FF95FF85FEFCFF768BFA6CCB" bold="true" box="[274,408,151,178]" italics="true" pageId="28" pageNumber="479">Orandrena</emphasis>
)
<emphasis id="5FF7EAB8FF95FF85FE47FF76884F6CCB" bold="true" box="[425,557,151,178]" italics="true" pageId="28" pageNumber="479">densissima</emphasis>
</taxonomicName>
<taxonomicNameLabel id="44C457C3FF95FF85FDDAFF7688CE6CC8" box="[564,684,151,177]" pageId="28" pageNumber="479" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="6D3C36AAFF95FF85FF79FF5789CF6CB7" blockId="28.[151,941,151,206]" box="[151,941,182,206]" pageId="28" pageNumber="479">
<uri id="19123AA8FF95FF85FF79FF5789CF6CB7" box="[151,941,182,206]" pageId="28" pageNumber="479">
urn:lsid:zoobank.org:act:
<uuid id="19250C7FFF95FF85FE74FF5789CF6CB7" box="[410,941,182,206]" pageId="28" pageNumber="479">BEA79392-12BB-4796-872A-8CDD9C876A4A</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="25996521FF95FF85FF79FF1D8E5B6D60" box="[151,1081,252,281]" pageId="28" pageNumber="479" type="materials_examined">
<paragraph id="6D3C36AAFF95FF85FF79FF1D8E5B6D60" blockId="28.[151,1437,252,2043]" box="[151,1081,252,281]" pageId="28" pageNumber="479">
<materialsCitation id="DDEB3CF7FF95FF85FF79FF1D8E5B6D60" ID-GBIF-Occurrence="3050682332" box="[151,1081,252,281]" collectingDate="1996-04-04" collectingDateMax="1996-04-05" collectingDateMin="1996-04-04" collectionCode="OOLM" collectorName="K. Denes" country="Tunisia" location="Ksar Hadada" pageId="28" pageNumber="479" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<emphasis id="5FF7EAB8FF95FF85FF79FF1D8BA66D6F" bold="true" box="[151,452,252,278]" pageId="28" pageNumber="479">
<typeStatus id="B2388808FF95FF85FF79FF1D8B586D6F" box="[151,314,252,278]" pageId="28" pageNumber="479" type="holotype">HOLOTYPE</typeStatus>
:
<collectingCountry id="1594763AFF95FF85FEA8FF1D8BDD6D6F" box="[326,447,252,278]" name="Tunisia" pageId="28" pageNumber="479">TUNISIA</collectingCountry>
:
</emphasis>
<location id="685C6071FF95FF85FE25FF1C883D6D6F" LSID="urn:lsid:plazi:treatment:E52A87BCFF95FF84FF79FF7689036BD4:685C6071FF95FF85FE25FF1C883D6D6F" box="[459,607,252,279]" country="Tunisia" name="Ksar Hadada" pageId="28" pageNumber="479">Ksar Hadada</location>
,
<date id="193D106AFF95FF85FD84FF1D88936D6E" box="[618,753,252,279]" pageId="28" pageNumber="479" value="1996-04-04" valueMax="1996-04-05" valueMin="1996-04-04">
<collectingDate id="0979E982FF95FF85FD84FF1D88936D6E" box="[618,753,252,279]" pageId="28" pageNumber="479" value="1996-04-04" valueMax="1996-04-05" valueMin="1996-04-04">45.iv.1996</collectingDate>
</date>
,
<specimenCount id="7B85FD23FF95FF85FD12FF1C897D6D6E" box="[764,799,253,279]" pageId="28" pageNumber="479" type="female">1♀</specimenCount>
, leg.
<collectorName id="C076537CFF95FF85FCB3FF1C89A76D6E" box="[861,965,253,279]" pageId="28" pageNumber="479">K. Deneš</collectorName>
(
<collectionCode id="0B92AE6FFF95FF85FC3AFF1F8E4F6D60" box="[980,1069,254,281]" pageId="28" pageNumber="479">OÖLM</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="25996521FF95FF85FF29FEC18BE8682B" pageId="28" pageNumber="479" type="description">
<paragraph id="6D3C36AAFF95FF85FF29FEC1898A6857" blockId="28.[151,1437,252,2043]" pageId="28" pageNumber="479">
<emphasis id="5FF7EAB8FF95FF85FF29FEC18BA36D42" bold="true" box="[199,449,288,315]" pageId="28" pageNumber="479">Description: Female:</emphasis>
Body length
<quantity id="AA7B9B4FFF95FF85FDB9FEC088F56D42" box="[599,663,289,315]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="28" pageNumber="479" unit="mm" value="8.0">8 mm</quantity>
(
<figureCitation id="F5B82A2FFF95FF85FD4AFEC189776D42" box="[676,789,288,315]" captionStart="FIGURES 95100" captionStartId="29.[151,264,1653,1677]" captionTargetBox="[151,1436,181,1629]" captionTargetId="figure-15@29.[151,1436,181,1629]" captionTargetPageId="29" captionText="FIGURES 95100. Andrena densissima spec. nov. 95. female profile, 96. female face, 97. female scutum, 98. female hind femur; 99. female terga. Andrena monilia Warncke, 1967 (holotype); 100. female terga." figureDoi="http://doi.org/10.5281/zenodo.4556586" httpUri="https://zenodo.org/record/4556586/files/figure.png" pageId="28" pageNumber="479">Figure 95</figureCitation>
).
<emphasis id="5FF7EAB8FF95FF85FCC6FEC1890F6D43" box="[808,877,288,314]" italics="true" pageId="28" pageNumber="479">Head:</emphasis>
Black, 1.1 times wider than long. Clypeus weakly domed, shallowly and irregularly punctate, punctures separated by 0.53 puncture diameters; narrow longitudinal impunctate central line present, underlying surface strongly microreticulate, dull (
<figureCitation id="F5B82A2FFF95FF85FBACFE898ED56DFA" box="[1090,1207,360,387]" captionStart="FIGURES 95100" captionStartId="29.[151,264,1653,1677]" captionTargetBox="[151,1436,181,1629]" captionTargetId="figure-15@29.[151,1436,181,1629]" captionTargetPageId="29" captionText="FIGURES 95100. Andrena densissima spec. nov. 95. female profile, 96. female face, 97. female scutum, 98. female hind femur; 99. female terga. Andrena monilia Warncke, 1967 (holotype); 100. female terga." figureDoi="http://doi.org/10.5281/zenodo.4556586" httpUri="https://zenodo.org/record/4556586/files/figure.png" pageId="28" pageNumber="479">Figure 96</figureCitation>
). Process of labrum short and wide, apically evenly rounded, dorsal surface microreticulate, dull. Gena subequal to width of compound eye; ocelloccipital distance equals ½ width of lateral ocellus. Fovea dorsally occupying ½ distance between lateral ocellus and compound eye, strongly narrowed ventrally, narrower than width of flagellum. Gena, vertex, face, and scape with long whitish to yellowish hairs, longest equalling length of scape. Antenna dark, A4 apically, A512 lightened orange below, A3 exceeds A4+5, shorter than A4+5+6.
<taxonomicName id="AA834D29FF95FF85FDDDFDA089726E22" authority=": Scutum" authorityName="Scutum" box="[563,784,577,603]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="28" pageNumber="479" phylum="Arthropoda" rank="genus">
<emphasis id="5FF7EAB8FF95FF85FDDDFDA088D66E23" box="[563,692,577,602]" italics="true" pageId="28" pageNumber="479">Mesosoma:</emphasis>
Scutum
</taxonomicName>
and scutellum irregularly punctate, punctures separated by 0.53 puncture diameters, underlying surface shagreened, weakly shining (
<figureCitation id="F5B82A2FFF95FF85FC0FFD858E316E06" box="[993,1107,612,639]" captionStart="FIGURES 95100" captionStartId="29.[151,264,1653,1677]" captionTargetBox="[151,1436,181,1629]" captionTargetId="figure-15@29.[151,1436,181,1629]" captionTargetPageId="29" captionText="FIGURES 95100. Andrena densissima spec. nov. 95. female profile, 96. female face, 97. female scutum, 98. female hind femur; 99. female terga. Andrena monilia Warncke, 1967 (holotype); 100. female terga." figureDoi="http://doi.org/10.5281/zenodo.4556586" httpUri="https://zenodo.org/record/4556586/files/figure.png" pageId="28" pageNumber="479">Figure 97</figureCitation>
). Pronotum rounded, without humeral angle. Episternum and lateral and dorsolateral parts of propodeum shallowly punctate with large punctures, separated by 0.51 puncture diameter, underlying surface shagreened, weakly shining, propodeal triangle indicated by change in sculpturing, internal area finely shagreened with weak central rugosity. Episternum and propodeum with long whitish hairs, scutum and scutellum almost hairless, laterally with short sparse light brownish hairs. Legs dark, apical tarsal segments lightened brown, pubescence white, scopa white; hind face of hind femur with 6 strongly produced semi-translucent orange pointed spines, clearly projecting perpendicularly (
<figureCitation id="F5B82A2FFF95FF85FB81FCDD8E836F2E" box="[1135,1249,828,855]" captionStart="FIGURES 95100" captionStartId="29.[151,264,1653,1677]" captionTargetBox="[151,1436,181,1629]" captionTargetId="figure-15@29.[151,1436,181,1629]" captionTargetPageId="29" captionText="FIGURES 95100. Andrena densissima spec. nov. 95. female profile, 96. female face, 97. female scutum, 98. female hind femur; 99. female terga. Andrena monilia Warncke, 1967 (holotype); 100. female terga." figureDoi="http://doi.org/10.5281/zenodo.4556586" httpUri="https://zenodo.org/record/4556586/files/figure.png" pageId="28" pageNumber="479">Figure 98</figureCitation>
). Wings hyaline, venation and stigma orange, nervulus interstitial.
<emphasis id="5FF7EAB8FF95FF85FD2CFC8089226F03" box="[706,832,865,890]" italics="true" pageId="28" pageNumber="479">Metasoma:</emphasis>
Tergal discs dark, apical margins occupying 2/5ths to ½ visible area, lightened orange-yellow, hyaline (
<figureCitation id="F5B82A2FFF95FF85FD2DFC6589576FE6" box="[707,821,900,927]" captionStart="FIGURES 95100" captionStartId="29.[151,264,1653,1677]" captionTargetBox="[151,1436,181,1629]" captionTargetId="figure-15@29.[151,1436,181,1629]" captionTargetPageId="29" captionText="FIGURES 95100. Andrena densissima spec. nov. 95. female profile, 96. female face, 97. female scutum, 98. female hind femur; 99. female terga. Andrena monilia Warncke, 1967 (holotype); 100. female terga." figureDoi="http://doi.org/10.5281/zenodo.4556586" httpUri="https://zenodo.org/record/4556586/files/figure.png" pageId="28" pageNumber="479">Figure 99</figureCitation>
), sterna lightened dark orange-red. Terga regularly but superficially punctate, punctures separated by 1 puncture diameter; underlying surface of T1 smooth and shining, T24 shagreened, weakly shining. T14 with thick, long apical hair bands that obscure underlying surface, widely interrupted on T12, continuous on T34, apical fringe of T5 and hairs flanking pygidial plate golden; pygidial plate triangular, apex pointed, dorsal surface apically weakly domed, impunctate.
</paragraph>
<paragraph id="6D3C36AAFF95FF85FF29FBD88BE8682B" blockId="28.[151,1437,252,2043]" box="[199,394,1080,1107]" pageId="28" pageNumber="479">
<emphasis id="5FF7EAB8FF95FF85FF29FBD88B6F682A" bold="true" box="[199,269,1081,1107]" pageId="28" pageNumber="479">Male:</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="25996521FF95FF85FF29FBBD8F076A32" pageId="28" pageNumber="479" type="diagnosis">
<paragraph id="6D3C36AAFF95FF85FF29FBBD8F076A32" blockId="28.[151,1437,252,2043]" pageId="28" pageNumber="479">
<emphasis id="5FF7EAB8FF95FF85FF29FBBD8B26680F" bold="true" box="[199,324,1116,1142]" pageId="28" pageNumber="479">Diagnosis:</emphasis>
The subgenus
<taxonomicName id="AA834D29FF95FF85FE03FBBD880A680F" box="[493,616,1116,1142]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="479" phylum="Arthropoda" rank="subGenus" subGenus="Orandrena">
<emphasis id="5FF7EAB8FF95FF85FE03FBBD880A680F" box="[493,616,1116,1142]" italics="true" pageId="28" pageNumber="479">Orandrena</emphasis>
</taxonomicName>
is easily recognised in the female sex by the combination of pronounced spines on the hind face of the hind femur, simple scopal hairs, and foveae that are strongly narrowed ventrally. Note, this subgenus should exclude
<taxonomicName id="AA834D29FF95FF85FE0CFB44896168C6" authority="Warncke, 1965" authorityName="Warncke" authorityYear="1965" box="[482,771,1188,1215]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="479" phylum="Arthropoda" rank="species" species="garrula">
<emphasis id="5FF7EAB8FF95FF85FE0CFB44883468C7" box="[482,598,1188,1215]" italics="true" pageId="28" pageNumber="479">A. garrula</emphasis>
<bibRefCitation id="09124B5BFF95FF85FDB5FB45896168C6" author="Warncke, K." box="[603,771,1188,1215]" pageId="28" pageNumber="479" pagination="27 - 76" refId="ref24173" refString="Warncke, K. (1965) Beitrag zur Kenntnis der Bienengattung Andrena Fabricius in Griechenland. Beitrage zur Entomologie, 15, 27 - 76." type="journal article" year="1965">Warncke, 1965</bibRefCitation>
</taxonomicName>
which falls outside the true
<taxonomicName id="AA834D29FF95FF85FBD5FB458ED468C7" box="[1083,1206,1188,1214]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="479" phylum="Arthropoda" rank="subGenus" subGenus="Orandrena">
<emphasis id="5FF7EAB8FF95FF85FBD5FB458ED468C7" box="[1083,1206,1188,1214]" italics="true" pageId="28" pageNumber="479">Orandrena</emphasis>
</taxonomicName>
(
<bibRefCitation id="09124B5BFF95FF85FB2DFB458FF768C6" author="Pisanty, G. &amp; Richter, R. &amp; Martin, T. &amp; Dettman, J. &amp; Cardinal, S." box="[1219,1429,1188,1215]" pageId="28" pageNumber="479" refId="ref23835" refString="Pisanty, G., Richter, R., Martin, T., Dettman, J. &amp; Cardinal, S. (2020) Molecular phylogeny and historical biogeography of an- drenine bees (Hymenoptera: Andrenidae). bioRxiv. [published online] https: // doi. org / 10.1101 / 2020.06.09.103556" type="book" year="2020">
Pisanty
<emphasis id="5FF7EAB8FF95FF85FAF0FB448F2C68C7" box="[1310,1358,1188,1214]" italics="true" pageId="28" pageNumber="479">et al</emphasis>
. 2020
</bibRefCitation>
) and which can easily be separated in the female sex by its lack of large punctures on the episternum. With the exception of
<taxonomicName id="AA834D29FF95FF85FF05FB0C8B8F697E" authority="Warncke, 1974" authorityName="Warncke" authorityYear="1974" box="[235,493,1260,1287]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="479" phylum="Arthropoda" rank="species" species="pela">
<emphasis id="5FF7EAB8FF95FF85FF05FB0C8B59697F" box="[235,315,1260,1287]" italics="true" pageId="28" pageNumber="479">A. pela</emphasis>
Warncke, 1974
</taxonomicName>
from
<collectingCountry id="1594763AFF95FF85FDDAFB0C88FB697E" box="[564,665,1261,1287]" name="Morocco" pageId="28" pageNumber="479">Morocco</collectingCountry>
that is easily separated by its strongly depressed tergal margins, female
<taxonomicName id="AA834D29FF95FF85FF3AFAF18B2D6953" box="[212,335,1296,1322]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="479" phylum="Arthropoda" rank="subGenus" subGenus="Orandrena">
<emphasis id="5FF7EAB8FF95FF85FF3AFAF18B2D6953" box="[212,335,1296,1322]" italics="true" pageId="28" pageNumber="479">Orandrena</emphasis>
</taxonomicName>
fall into two groups: those with a dull clypeus (
<taxonomicName id="AA834D29FF95FF85FC88FAF08EEF6952" authority="Warncke, 1967" authorityName="Warncke" authorityYear="1967" box="[870,1165,1296,1323]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="479" phylum="Arthropoda" rank="species" species="monilia">
<emphasis id="5FF7EAB8FF95FF85FC88FAF089B96953" box="[870,987,1296,1323]" italics="true" pageId="28" pageNumber="479">A. monilia</emphasis>
<bibRefCitation id="09124B5BFF95FF85FC0FFAF18EEF6952" author="Warncke K." box="[993,1165,1296,1323]" pageId="28" pageNumber="479" pagination="171 - 318" refId="ref24200" refString="Warncke K. (1967) Beitrag zur Klarung palaarktischer Andrena - Arten. Eos, 43, 171 - 318." type="journal article" year="1967">Warncke, 1967</bibRefCitation>
</taxonomicName>
), and the remaining five species (
<tableCitation id="20010311FF95FF85FF19FAD58B2E6936" box="[247,332,1332,1359]" captionStart="TABLE 2" captionStartId="30.[151,239,151,175]" captionText="TABLE 2. Andrena (Orandrena) species and their global distributions. Note, following Pisanty et al. (2020), Andrena garrula Warncke, 1965 is excluded from the subgenus." pageId="28" pageNumber="479">Table 2</tableCitation>
) that have clypei with clear longitudinal striations.
<taxonomicName id="AA834D29FF95FF85FC61FAD58E126937" authorityName="Wood" authorityYear="2021" box="[911,1136,1332,1358]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="479" phylum="Arthropoda" rank="species" species="densissima">
<emphasis id="5FF7EAB8FF95FF85FC61FAD58E126937" box="[911,1136,1332,1358]" italics="true" pageId="28" pageNumber="479">Andrena densissima</emphasis>
</taxonomicName>
has a dull clypeus without striations, placing it close to
<taxonomicName id="AA834D29FF95FF85FE37FAB8882C690B" authorityName="Warncke" authorityYear="1967" box="[473,590,1368,1395]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="479" phylum="Arthropoda" rank="species" species="monilia">
<emphasis id="5FF7EAB8FF95FF85FE37FAB8882C690B" box="[473,590,1368,1395]" italics="true" pageId="28" pageNumber="479">A. monilia</emphasis>
</taxonomicName>
, but it can be easily separated (
<taxonomicName id="AA834D29FF95FF85FC40FAB88E41690B" authorityName="Warncke" authorityYear="1967" box="[942,1059,1368,1395]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="479" phylum="Arthropoda" rank="species" species="monilia">
<emphasis id="5FF7EAB8FF95FF85FC40FAB88E41690B" box="[942,1059,1368,1395]" italics="true" pageId="28" pageNumber="479">A. monilia</emphasis>
</taxonomicName>
character state in parentheses) by the process of the labrum which is wide, short, and shallowly rounded, not forming a clear apical point (forming a clear triangular point), by its smaller size of
<quantity id="AA7B9B4FFF95FF85FD67FA4088A969C2" box="[649,715,1441,1467]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="28" pageNumber="479" unit="mm" value="8.0">8 mm</quantity>
(larger,
<quantity id="AA7B9B4FFF95FF85FCC6FA4089C769C2" box="[808,933,1441,1467]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" metricValueMax="1.1" metricValueMin="1.0" pageId="28" pageNumber="479" unit="mm" value="10.5" valueMax="11.0" valueMin="10.0">1011 mm</quantity>
), by the shagreened and only weakly shining scutum (scutum with very weak shagreenation, shining. Note, this only applies to western
<taxonomicName id="AA834D29FF95FF85FB75FA248F7069A7" authorityName="Warncke" authorityYear="1967" box="[1179,1298,1476,1503]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="479" phylum="Arthropoda" rank="species" species="monilia">
<emphasis id="5FF7EAB8FF95FF85FB75FA248F7069A7" box="[1179,1298,1476,1503]" italics="true" pageId="28" pageNumber="479">A. monilia</emphasis>
</taxonomicName>
populations from
<collectingCountry id="1594763AFF95FF85FF3BFA098B776A7B" box="[213,277,1512,1538]" name="Spain" pageId="28" pageNumber="479">Spain</collectingCountry>
to
<collectingCountry id="1594763AFF95FF85FED7FA098BF26A7B" box="[313,400,1512,1538]" name="Tunisia" pageId="28" pageNumber="479">Tunisia</collectingCountry>
. Eastern populations in
<collectingCountry id="1594763AFF95FF85FD72FA0988B86A7B" box="[668,730,1512,1538]" name="Israel" pageId="28" pageNumber="479">Israel</collectingCountry>
and
<collectingCountry id="1594763AFF95FF85FCFCFA09893F6A7B" box="[786,861,1512,1538]" name="Jordan" pageId="28" pageNumber="479">Jordan</collectingCountry>
have similarly shagreened scuta; these populations may deserve species status in future revisions), and by the much wider tergal hair bands that clearly exceed the width of the tergal margins (hairbands not clearly exceeding the width of the tergal margins,
<figureCitation id="F5B82A2FFF95FF85FB4BF9D18F3B6A32" box="[1189,1369,1584,1611]" captionStart="FIGURES 95100" captionStartId="29.[151,264,1653,1677]" captionTargetBox="[151,1436,181,1629]" captionTargetId="figure-15@29.[151,1436,181,1629]" captionTargetPageId="29" captionText="FIGURES 95100. Andrena densissima spec. nov. 95. female profile, 96. female face, 97. female scutum, 98. female hind femur; 99. female terga. Andrena monilia Warncke, 1967 (holotype); 100. female terga." figureDoi="http://doi.org/10.5281/zenodo.4556586" httpUri="https://zenodo.org/record/4556586/files/figure.png" pageId="28" pageNumber="479">Figures 99100</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="25996521FF95FF85FF29F9B58BB76BCB" pageId="28" pageNumber="479" type="discussion">
<paragraph id="6D3C36AAFF95FF85FF29F9B58BB76BCB" blockId="28.[151,1437,252,2043]" pageId="28" pageNumber="479">
<emphasis id="5FF7EAB8FF95FF85FF29F9B58B2D6A17" bold="true" box="[199,335,1620,1646]" pageId="28" pageNumber="479">Discussion:</emphasis>
As a subgenus,
<taxonomicName id="AA834D29FF95FF85FDECF9B5881F6A17" box="[514,637,1620,1646]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="479" phylum="Arthropoda" rank="subGenus" subGenus="Orandrena">
<emphasis id="5FF7EAB8FF95FF85FDECF9B5881F6A17" box="[514,637,1620,1646]" italics="true" pageId="28" pageNumber="479">Orandrena</emphasis>
</taxonomicName>
shows a Mediterranean distribution, but individual species have strongly restricted distributions, most notably those found in
<collectingCountry id="1594763AFF95FF85FD11F999892D6AEB" box="[767,847,1656,1682]" name="Turkey" pageId="28" pageNumber="479">Turkey</collectingCountry>
which is the clear centre of diversity, most of which were described by Warncke (
<tableCitation id="20010311FF95FF85FE31F97D885A6ACE" box="[479,568,1692,1719]" captionStart="TABLE 2" captionStartId="30.[151,239,151,175]" captionText="TABLE 2. Andrena (Orandrena) species and their global distributions. Note, following Pisanty et al. (2020), Andrena garrula Warncke, 1965 is excluded from the subgenus." pageId="28" pageNumber="479">Table 2</tableCitation>
). The exception to this rule is
<taxonomicName id="AA834D29FF95FF85FC7DF97C8E6B6ACF" authorityName="Warncke" authorityYear="1967" box="[915,1033,1692,1719]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="479" phylum="Arthropoda" rank="species" species="monilia">
<emphasis id="5FF7EAB8FF95FF85FC7DF97C8E6B6ACF" box="[915,1033,1692,1719]" italics="true" pageId="28" pageNumber="479">A. monilia</emphasis>
</taxonomicName>
which is found across the Mediterranean basin, though it is variable throughout this range, and populations in the Middle East potentially deserve species status because of variation in integumental sculpturing and male clypeal colouration.
<taxonomicName id="AA834D29FF95FF85FB43F9058F086A87" authorityName="Warncke" authorityYear="1967" box="[1197,1386,1764,1790]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="479" phylum="Arthropoda" rank="species" species="monilia">
<emphasis id="5FF7EAB8FF95FF85FB43F9058F086A87" box="[1197,1386,1764,1790]" italics="true" pageId="28" pageNumber="479">Andrena monilia</emphasis>
</taxonomicName>
was reported from the northern part of
<collectingCountry id="1594763AFF95FF85FDFAF8E988056B5B" box="[532,615,1800,1826]" name="Tunisia" pageId="28" pageNumber="479">Tunisia</collectingCountry>
some years after its original description (
<bibRefCitation id="09124B5BFF95FF85FBDFF8E98EBA6B5A" author="Warncke, K." box="[1073,1240,1800,1827]" pageId="28" pageNumber="479" pagination="86 - 87" refId="ref24288" refString="Warncke, K. (1980) Zur Verbreitung der Bienengattung Andrena F. in Tunesien. Mitteilungen der Munchner Entomologischen Gesellschaft, 70, 86 - 87." type="journal article" year="1980">Warncke 1980</bibRefCitation>
), and its presence in the country can be confirmed by more recently examined material (see below). In contrast,
<taxonomicName id="AA834D29FF95FF85FB22F8CC8F0A6B3F" authorityName="Wood" authorityYear="2021" box="[1228,1384,1836,1863]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="479" phylum="Arthropoda" rank="species" species="densissima">
<emphasis id="5FF7EAB8FF95FF85FB22F8CC8F0A6B3F" box="[1228,1384,1836,1863]" italics="true" pageId="28" pageNumber="479">A. densissima</emphasis>
</taxonomicName>
was collected from southern
<collectingCountry id="1594763AFF95FF85FE43F8B188626B13" box="[429,512,1872,1898]" name="Tunisia" pageId="28" pageNumber="479">Tunisia</collectingCountry>
and clearly differs morphologically. Like most members of the subgenus, it may have a highly restricted range, and similar desert habitats in
<collectingCountry id="1594763AFF95FF85FCC0F89589E06BF7" box="[814,898,1908,1934]" name="Algeria" pageId="28" pageNumber="479">Algeria</collectingCountry>
and
<collectingCountry id="1594763AFF95FF85FC59F895899A6BF7" box="[951,1016,1908,1934]" name="Libya" pageId="28" pageNumber="479">Libya</collectingCountry>
should be searched to see if it is more widespread in these regions.
</paragraph>
</subSubSection>
<subSubSection id="25996521FF95FF84FF29F85C89036BD4" lastPageId="29" lastPageNumber="480" pageId="28" pageNumber="479" type="etymology">
<paragraph id="6D3C36AAFF95FF85FF29F85C8F3A6B83" blockId="28.[151,1437,252,2043]" pageId="28" pageNumber="479">
<emphasis id="5FF7EAB8FF95FF85FF29F85C8B306BAE" bold="true" box="[199,338,1981,2007]" pageId="28" pageNumber="479">Etymology:</emphasis>
The name is the feminine superlative degree of
<emphasis id="5FF7EAB8FF95FF85FC6AF85D89A76BAF" box="[900,965,1980,2006]" italics="true" pageId="28" pageNumber="479">denso</emphasis>
(dense, thick, therefore very dense, very thick) because of the pronounced hair bands in the female sex that set it apart from other
<taxonomicName id="AA834D29FF95FF85FB93F8018E9A6B83" box="[1149,1272,2016,2042]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="479" phylum="Arthropoda" rank="subGenus" subGenus="Orandrena">
<emphasis id="5FF7EAB8FF95FF85FB93F8018E9A6B83" box="[1149,1272,2016,2042]" italics="true" pageId="28" pageNumber="479">Orandrena</emphasis>
</taxonomicName>
species.
</paragraph>
<caption id="39FC6622FF94FF84FF79F9948E6C6ACB" ID-DOI="http://doi.org/10.5281/zenodo.4556586" ID-Zenodo-Dep="4556586" httpUri="https://zenodo.org/record/4556586/files/figure.png" pageId="29" pageNumber="480" startId="29.[151,264,1653,1677]" targetBox="[151,1436,181,1629]" targetPageId="29">
<paragraph id="6D3C36AAFF94FF84FF79F9948E6C6ACB" blockId="29.[151,1436,1653,1714]" pageId="29" pageNumber="480">
<emphasis id="5FF7EAB8FF94FF84FF79F9948B016AF4" bold="true" box="[151,355,1653,1677]" pageId="29" pageNumber="480">FIGURES 95100.</emphasis>
<taxonomicName id="AA834D29FF94FF84FE85F994885B6AF4" authority="Wood, 2021" authorityName="Wood" authorityYear="2021" box="[363,569,1653,1677]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="480" phylum="Arthropoda" rank="species" species="densissima" status="sp. nov.">
<emphasis id="5FF7EAB8FF94FF84FE85F994885B6AF4" box="[363,569,1653,1677]" italics="true" pageId="29" pageNumber="480">Andrena densissima</emphasis>
</taxonomicName>
<emphasis id="5FF7EAB8FF94FF84FDACF99488C96AF4" bold="true" box="[578,683,1653,1677]" pageId="29" pageNumber="480">
<taxonomicNameLabel id="44C457C3FF94FF84FDACF99488C96AF4" box="[578,683,1653,1677]" pageId="29" pageNumber="480" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
95. female profile, 96. female face, 97. female scutum, 98. female hind femur; 99. female terga.
<taxonomicName id="AA834D29FF94FF84FE7FF97888B86AC8" authority="Warncke, 1967" authorityName="Warncke" authorityYear="1967" box="[401,730,1689,1713]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="480" phylum="Arthropoda" rank="species" species="monilia">
<emphasis id="5FF7EAB8FF94FF84FE7FF97888596AC8" box="[401,571,1689,1713]" italics="true" pageId="29" pageNumber="480">Andrena monilia</emphasis>
<bibRefCitation id="09124B5BFF94FF84FDAFF97888B86AC8" author="Warncke K." box="[577,730,1689,1713]" pageId="29" pageNumber="480" pagination="171 - 318" refId="ref24200" refString="Warncke K. (1967) Beitrag zur Klarung palaarktischer Andrena - Arten. Eos, 43, 171 - 318." type="journal article" year="1967">Warncke, 1967</bibRefCitation>
</taxonomicName>
(holotype); 100. female terga.
</paragraph>
</caption>
<paragraph id="6D3C36AAFF94FF84FF29F93E89036BD4" blockId="29.[151,1436,1759,1968]" pageId="29" pageNumber="480">
<materialsCitation id="DDEB3CF7FF94FF84FF29F93E8B956B67" ID-GBIF-Occurrence="3050682337" collectionCode="ZMHB" collectorName="Bytinski-Salz" country="Israel" location="Mishmar HaNegev" pageId="29" pageNumber="480" specimenCount="1" specimenCount-female="1">
<emphasis id="5FF7EAB8FF94FF84FF29F93E88626A80" bold="true" box="[199,512,1759,1785]" pageId="29" pageNumber="480">Other material examined:</emphasis>
(
<taxonomicName id="AA834D29FF94FF84FDFCF93E88B26A80" authorityName="Warncke" authorityYear="1967" box="[530,720,1759,1785]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="480" phylum="Arthropoda" rank="species" species="monilia">
<emphasis id="5FF7EAB8FF94FF84FDFCF93E88B26A80" box="[530,720,1759,1785]" italics="true" pageId="29" pageNumber="480">Andrena monilia</emphasis>
</taxonomicName>
):
<emphasis id="5FF7EAB8FF94FF84FD07F93E89386A80" bold="true" box="[745,858,1759,1785]" pageId="29" pageNumber="480">
<collectingCountry id="1594763AFF94FF84FD07F93E89346A80" box="[745,854,1759,1785]" name="Israel" pageId="29" pageNumber="480">ISRAEL</collectingCountry>
:
</emphasis>
<location id="685C6071FF94FF84FC8DF93E8E596A83" LSID="urn:lsid:plazi:treatment:E52A87BCFF95FF84FF79FF7689036BD4:685C6071FF94FF84FC8DF93E8E596A83" box="[867,1083,1759,1786]" country="Israel" name="Mishmar HaNegev" pageId="29" pageNumber="480">Mishmar HaNegev</location>
, 24.iii.19xx [year unspecified],
<specimenCount id="7B85FD23FF94FF84FF79F8E28AD86B64" box="[151,186,1795,1821]" pageId="29" pageNumber="480" type="female">1♀</specimenCount>
, leg.
<collectorName id="C076537CFF94FF84FF15F8E28BF46B64" box="[251,406,1795,1821]" pageId="29" pageNumber="480">Bytinski-Salz</collectorName>
,
<collectionCode id="0B92AE6FFF94FF84FE4DF8E58B956B67" box="[419,503,1796,1822]" pageId="29" pageNumber="480">ZMHB</collectionCode>
</materialsCitation>
;
<materialsCitation id="DDEB3CF7FF94FF84FDEAF8E28E3E6B64" ID-GBIF-Occurrence="3050682352" box="[516,1116,1795,1821]" collectingDate="2010-03-23" collectionCode="SMNHTAU" collectorName="G. Pisanty" country="Israel" location="Gal'on" pageId="29" pageNumber="480" specimenCount="1" specimenCount-female="1">
<location id="685C6071FF94FF84FDEAF8E288366B64" LSID="urn:lsid:plazi:treatment:E52A87BCFF95FF84FF79FF7689036BD4:685C6071FF94FF84FDEAF8E288366B64" box="[516,596,1795,1821]" country="Israel" name="Gal'on" pageId="29" pageNumber="480">Galon</location>
,
<date id="193D106AFF94FF84FD8FF8E288BE6B64" box="[609,732,1795,1821]" pageId="29" pageNumber="480" value="2010-03-23">
<collectingDate id="0979E982FF94FF84FD8FF8E288BE6B64" box="[609,732,1795,1821]" pageId="29" pageNumber="480" value="2010-03-23">23.iii.2010</collectingDate>
</date>
,
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, leg.
<collectorName id="C076537CFF94FF84FCA3F8E289A76B64" box="[845,965,1795,1821]" pageId="29" pageNumber="480">G. Pisanty</collectorName>
,
<collectionCode id="0B92AE6FFF94FF84FC3EF8E28E3E6B64" box="[976,1116,1795,1821]" pageId="29" pageNumber="480">SMNHTAU</collectionCode>
</materialsCitation>
;
<materialsCitation id="DDEB3CF7FF94FF84FB87F8E288906B3D" ID-GBIF-Occurrence="3050682318" collectingDate="1996-05-02" collectionCode="OOLM" collectorName="Ma. Halada" country="Jordan" location="At-Tafilah" pageId="29" pageNumber="480" specimenCount="4" specimenCount-female="4" stateProvince="At Tafilah">
<emphasis id="5FF7EAB8FF94FF84FB87F8E28E856B64" bold="true" box="[1129,1255,1795,1821]" pageId="29" pageNumber="480">
<collectingCountry id="1594763AFF94FF84FB87F8E28E806B64" box="[1129,1250,1795,1821]" name="Jordan" pageId="29" pageNumber="480">JORDAN</collectingCountry>
:
</emphasis>
<locationDeviation id="7C8E2F88FF94FF84FB01F8E28FFE6B64" box="[1263,1436,1795,1822]" location="Tafila" pageId="29" pageNumber="480">
<quantity id="AA7B9B4FFF94FF84FB01F8E28F556B64" box="[1263,1335,1795,1821]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="29" pageNumber="480" unit="km" value="30.0">30 km</quantity>
N
<collectingRegion id="AF47F848FF94FF84FAB5F8E28FFE6B64" box="[1371,1436,1795,1821]" country="Jordan" name="At Tafilah" pageId="29" pageNumber="480">Tafila</collectingRegion>
</locationDeviation>
[
<location id="685C6071FF94FF84FF73F8C68B786B3B" LSID="urn:lsid:plazi:treatment:E52A87BCFF95FF84FF79FF7689036BD4:685C6071FF94FF84FF73F8C68B786B3B" box="[157,282,1831,1858]" country="Jordan" name="At-Tafilah" pageId="29" pageNumber="480" stateProvince="At Tafilah">At-Tafilah</location>
],
<date id="193D106AFF94FF84FEC1F8C68BF36B38" box="[303,401,1831,1857]" pageId="29" pageNumber="480" value="1996-05-02">
<collectingDate id="0979E982FF94FF84FEC1F8C68BF36B38" box="[303,401,1831,1857]" pageId="29" pageNumber="480" value="1996-05-02">2.v.1996</collectingDate>
</date>
,
<specimenCount id="7B85FD23FF94FF84FE71F8C68BA66B38" box="[415,452,1831,1857]" pageId="29" pageNumber="480" type="female">4♀</specimenCount>
, leg.
<collectorName id="C076537CFF94FF84FDE8F8C988EC6B38" box="[518,654,1831,1858]" pageId="29" pageNumber="480">Ma. Halada</collectorName>
,
<collectionCode id="0B92AE6FFF94FF84FD72F8C888906B3D" box="[668,754,1833,1860]" pageId="29" pageNumber="480">OÖLM</collectionCode>
</materialsCitation>
;
<materialsCitation id="DDEB3CF7FF94FF84FCEEF8C68A8F6B11" ID-GBIF-Occurrence="3050682361" collectingDate="2013-04-06" collectionCode="OOLM" collectorName="M. Snizek" country="Jordan" location="Al Karak" pageId="29" pageNumber="480" specimenCount="1" specimenCount-female="1" stateProvince="Al Karak">
<collectingRegion id="AF47F848FF94FF84FCEEF8C6890B6B38" box="[768,873,1831,1857]" country="Jordan" name="Al Karak" pageId="29" pageNumber="480">Al Karak</collectingRegion>
env [
<collectingRegion id="AF47F848FF94FF84FC42F8C6899A6B3B" box="[940,1016,1831,1858]" country="Jordan" name="Al Karak" pageId="29" pageNumber="480">Kerak</collectingRegion>
],
<date id="193D106AFF94FF84FBE3F8C68E1A6B38" box="[1037,1144,1831,1857]" pageId="29" pageNumber="480" value="2013-04-06">
<collectingDate id="0979E982FF94FF84FBE3F8C68E1A6B38" box="[1037,1144,1831,1857]" pageId="29" pageNumber="480" value="2013-04-06">6.iv.2013</collectingDate>
</date>
, 1³,
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, leg.
<collectorName id="C076537CFF94FF84FAF1F8C98FFA6B38" box="[1311,1432,1831,1858]" pageId="29" pageNumber="480">M. Snižek</collectorName>
,
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</materialsCitation>
;
<materialsCitation id="DDEB3CF7FF94FF84FF12F8AA89A86B11" ID-GBIF-Occurrence="3050682363" box="[252,970,1867,1896]" collectingDate="1998-04-13" collectionCode="OOLM" collectorName="K. Denes" country="Tunisia" location="Kasserine" pageId="29" pageNumber="480" specimenCount="1" stateProvince="Kasserine">
<emphasis id="5FF7EAB8FF94FF84FF12F8AA8B186B1C" bold="true" box="[252,378,1867,1893]" pageId="29" pageNumber="480">
<collectingCountry id="1594763AFF94FF84FF12F8AA8B176B1C" box="[252,373,1867,1893]" name="Tunisia" pageId="29" pageNumber="480">TUNISIA</collectingCountry>
:
</emphasis>
<collectingRegion id="AF47F848FF94FF84FE6BF8AA8B946B1C" box="[389,502,1867,1893]" country="Tunisia" name="Kasserine" pageId="29" pageNumber="480">Kasserine</collectingRegion>
,
<date id="193D106AFF94FF84FDEBF8AA881F6B1C" box="[517,637,1867,1893]" pageId="29" pageNumber="480" value="1998-04-13">
<collectingDate id="0979E982FF94FF84FDEBF8AA881F6B1C" box="[517,637,1867,1893]" pageId="29" pageNumber="480" value="1998-04-13">13.iv.1998</collectingDate>
</date>
, 2³, leg.
<collectorName id="C076537CFF94FF84FD18F8AD89076B1F" box="[758,869,1868,1894]" pageId="29" pageNumber="480">K. Deneš</collectorName>
,
<collectionCode id="0B92AE6FFF94FF84FC9AF8AC89A86B11" box="[884,970,1869,1896]" pageId="29" pageNumber="480">OÖLM</collectionCode>
</materialsCitation>
;
<materialsCitation id="DDEB3CF7FF94FF84FC37F8AA8A8F6BF5" ID-GBIF-Occurrence="3050682330" collectingDate="1998-04-12" collectionCode="OOLM" collectorName="K. Denes" country="Tunisia" location="Sbeitla" pageId="29" pageNumber="480" specimenCount="1" specimenCount-female="1">
<location id="685C6071FF94FF84FC37F8AA8E486B1C" LSID="urn:lsid:plazi:treatment:E52A87BCFF95FF84FF79FF7689036BD4:685C6071FF94FF84FC37F8AA8E486B1C" box="[985,1066,1867,1893]" country="Tunisia" name="Sbeitla" pageId="29" pageNumber="480">Sbeitla</location>
,
<date id="193D106AFF94FF84FBD7F8AA8ED36B1C" box="[1081,1201,1867,1893]" pageId="29" pageNumber="480" value="1998-04-12">
<collectingDate id="0979E982FF94FF84FBD7F8AA8ED36B1C" box="[1081,1201,1867,1893]" pageId="29" pageNumber="480" value="1998-04-12">12.iv.1998</collectingDate>
</date>
,
<specimenCount id="7B85FD23FF94FF84FB2EF8AA8E816B1C" box="[1216,1251,1867,1893]" pageId="29" pageNumber="480" type="female">1♀</specimenCount>
, leg.
<collectorName id="C076537CFF94FF84FAC7F8AD8FFA6B1F" box="[1321,1432,1868,1894]" pageId="29" pageNumber="480">K. Deneš</collectorName>
,
<collectionCode id="0B92AE6FFF94FF84FF79F8908A8F6BF5" box="[151,237,1905,1932]" pageId="29" pageNumber="480">OÖLM</collectionCode>
</materialsCitation>
;
<materialsCitation id="DDEB3CF7FF94FF84FF16F88E8F7F6BF0" ID-GBIF-Occurrence="3050682357" box="[248,1309,1903,1932]" collectingDate="1933-05-10" collectionCode="OOLM" collectorName="J. M. Dusmet &amp; Alonso" country="Spain" location="Montarco" pageId="29" pageNumber="480" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<emphasis id="5FF7EAB8FF94FF84FF16F88E8B366BF0" bold="true" box="[248,340,1903,1929]" pageId="29" pageNumber="480">
<collectingCountry id="1594763AFF94FF84FF16F88E8B2C6BF0" box="[248,334,1903,1929]" name="Spain" pageId="29" pageNumber="480">SPAIN</collectingCountry>
:
</emphasis>
<location id="685C6071FF94FF84FEB5F8918BA96BF3" LSID="urn:lsid:plazi:treatment:E52A87BCFF95FF84FF79FF7689036BD4:685C6071FF94FF84FEB5F8918BA96BF3" box="[347,459,1904,1930]" country="Spain" name="Montarco" pageId="29" pageNumber="480">Montarco</location>
,
<date id="193D106AFF94FF84FE38F88E88246BF0" box="[470,582,1903,1929]" pageId="29" pageNumber="480" value="1933-05-10">
<collectingDate id="0979E982FF94FF84FE38F88E88246BF0" box="[470,582,1903,1929]" pageId="29" pageNumber="480" value="1933-05-10">10.v.1933</collectingDate>
</date>
,
<specimenCount id="7B85FD23FF94FF84FDBCF88E88176BF0" box="[594,629,1903,1929]" pageId="29" pageNumber="480" type="female">1♀</specimenCount>
, leg.
<collectorName id="C076537CFF94FF84FD5DF89189286BF3" box="[691,842,1904,1930]" pageId="29" pageNumber="480">J. M. Dusmet</collectorName>
y
<collectorName id="C076537CFF94FF84FC8AF88E89DA6BF0" box="[868,952,1903,1929]" pageId="29" pageNumber="480">Alonso</collectorName>
,
<collectionCode id="0B92AE6FFF94FF84FC2DF8908E776BF5" box="[963,1045,1905,1932]" pageId="29" pageNumber="480">OÖLM</collectionCode>
(
<typeStatus id="B2388808FF94FF84FBCAF88E8EE86BF0" box="[1060,1162,1903,1929]" pageId="29" pageNumber="480" type="holotype">holotype</typeStatus>
,
<figureCitation id="F5B82A2FFF94FF84FB78F88E8F766BF0" box="[1174,1300,1903,1929]" captionStart="FIGURES 95100" captionStartId="29.[151,264,1653,1677]" captionTargetBox="[151,1436,181,1629]" captionTargetId="figure-15@29.[151,1436,181,1629]" captionTargetPageId="29" captionText="FIGURES 95100. Andrena densissima spec. nov. 95. female profile, 96. female face, 97. female scutum, 98. female hind femur; 99. female terga. Andrena monilia Warncke, 1967 (holotype); 100. female terga." figureDoi="http://doi.org/10.5281/zenodo.4556586" httpUri="https://zenodo.org/record/4556586/files/figure.png" pageId="29" pageNumber="480">Figure 100</figureCitation>
)
</materialsCitation>
;
<materialsCitation id="DDEB3CF7FF94FF84FAC6F89188BD6BC9" ID-GBIF-Occurrence="3050682333" collectingDate="1924-04-28" collectionCode="OOLM" collectorName="J. M. Dusmet &amp; Alonso" country="Spain" location="Montarco" pageId="29" pageNumber="480" specimenCount="1">
<location id="685C6071FF94FF84FAC6F8918FFA6BF3" LSID="urn:lsid:plazi:treatment:E52A87BCFF95FF84FF79FF7689036BD4:685C6071FF94FF84FAC6F8918FFA6BF3" box="[1320,1432,1904,1930]" country="Spain" name="Montarco" pageId="29" pageNumber="480">Montarco</location>
,
<date id="193D106AFF94FF84FF79F8728B6D6BD4" box="[151,271,1939,1965]" pageId="29" pageNumber="480" value="1924-04-28">
<collectingDate id="0979E982FF94FF84FF79F8728B6D6BD4" box="[151,271,1939,1965]" pageId="29" pageNumber="480" value="1924-04-28">28.iv.1924</collectingDate>
</date>
, 1³, leg.
<collectorName id="C076537CFF94FF84FE92F87588716BD7" box="[380,531,1940,1966]" pageId="29" pageNumber="480">J. M. Dusmet</collectorName>
y
<collectorName id="C076537CFF94FF84FDC0F87288E06BD4" box="[558,642,1939,1965]" pageId="29" pageNumber="480">Alonso</collectorName>
,
<collectionCode id="0B92AE6FFF94FF84FD63F87488BD6BC9" box="[653,735,1941,1968]" pageId="29" pageNumber="480">OÖLM</collectionCode>
</materialsCitation>
(
<typeStatus id="B2388808FF94FF84FD00F87289376BD4" box="[750,853,1939,1965]" pageId="29" pageNumber="480" type="paratype">paratype</typeStatus>
).
</paragraph>
</subSubSection>
</treatment>
</document>