377 lines
48 KiB
XML
377 lines
48 KiB
XML
<document id="068D815C8074EA986C2FCA1652249BCF" ID-CLB-Dataset="7784" ID-DOI="10.11646/zootaxa.4933.4.1" ID-GBIF-Dataset="2ffdac24-b2ae-448c-b88e-b4f0dd1a9547" ID-ISSN="1175-5326" ID-Zenodo-Dep="4556546" ID-ZooBank="1FC0D2E0-888E-4F79-ABFE-BC7E91ADEECE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1614066053014" checkinUser="plazi" docAuthor="Wood, Thomas James" docDate="2021" docId="E52A87BCFF8BFF9CFF79FF3E899B6DD4" docLanguage="en" docName="zootaxa.4933.4.1.pdf" docOrigin="Zootaxa 4933 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Andrena (Aciandrena) badiyah Wood 2021, spec. nov." docType="treatment" docUuid="7EED4526-5105-47EF-8EFB-102B70A7E89A" docUuidSource="ZooBank" docVersion="9" lastPageNumber="456" masterDocId="1913FFC4FF89FF99FFEEFFE18A626C79" masterDocTitle="Fifteen new Andrena species from little-visited arid, Mediterranean, and mountainous parts of the Old World (Hymenoptera: Andrenidae)" masterLastPageNumber="492" masterPageNumber="451" pageNumber="453" updateTime="1698930763140" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="4D1702B85742586EE7220FBA958199E6">Fifteen new Andrena species from little-visited arid, Mediterranean, and mountainous parts of the Old World (Hymenoptera: Andrenidae)</mods:title>
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<mods:name id="227AE1A7479FD5363EAC2E6DEF50ADB5" type="personal">
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<mods:namePart id="6C9BCFBBAA17BDF00A622331F045B9CE">Wood, Thomas James</mods:namePart>
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<mods:affiliation id="938280F0D4402EEBB8BD9470010A9C41">Laboratoire de Zoologie, Université de Mons, Mons, 7000, Belgium</mods:affiliation>
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<mods:title id="21697AC0F36F0371D884F66FC9EB79C9">Zootaxa</mods:title>
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<mods:part id="D18F47A779C00A12344BFF38B3DCE236">
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<mods:date id="E860D8D0EE8AAB38AF709A9D9801028F">2021</mods:date>
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<mods:number id="2781DE810991A7395443E184739E2141">2021-02-23</mods:number>
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<mods:identifier id="1DC4D1FDAE7123948B3308CFEF52A596" type="DOI">10.11646/zootaxa.4933.4.1</mods:identifier>
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<treatment id="E52A87BCFF8BFF9CFF79FF3E899B6DD4" ID-DOI="http://doi.org/10.5281/zenodo.4599184" ID-GBIF-Taxon="180326869" ID-Zenodo-Dep="4599184" LSID="urn:lsid:zoobank.org:act:7EED4526-5105-47EF-8EFB-102B70A7E89A" httpUri="http://treatment.plazi.org/id/E52A87BCFF8BFF9CFF79FF3E899B6DD4" lastPageId="5" lastPageNumber="456" pageId="2" pageNumber="453">
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<subSubSection id="25996521FF8BFF9BFF79FF3E89FF6D6F" pageId="2" pageNumber="453" type="nomenclature">
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<paragraph id="6D3C36AAFF8BFF9BFF79FF3E88FD6C80" blockId="2.[151,925,223,278]" box="[151,671,223,250]" pageId="2" pageNumber="453">
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<heading id="367481C6FF8BFF9BFF79FF3E88FD6C80" bold="true" box="[151,671,223,250]" fontSize="11" level="1" pageId="2" pageNumber="453" reason="1">
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<emphasis id="5FF7EAB8FF8BFF9BFF79FF3E88FD6C80" bold="true" box="[151,671,223,250]" pageId="2" pageNumber="453">
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<taxonomicName id="AA834D29FF8BFF9BFF79FF3E88426C83" authority="Wood, 2021" authorityName="Wood" authorityYear="2021" box="[151,544,223,250]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="456" phylum="Arthropoda" rank="species" species="badiyah" status="sp. nov." subGenus="Aciandrena">
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||
<emphasis id="5FF7EAB8FF8BFF9BFF79FF3E8B636C83" bold="true" box="[151,257,223,250]" italics="true" pageId="2" pageNumber="453">Andrena</emphasis>
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||
(
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||
<emphasis id="5FF7EAB8FF8BFF9BFEFCFF3E8BCD6C83" bold="true" box="[274,431,223,250]" italics="true" pageId="2" pageNumber="453">?Aciandrena</emphasis>
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||
)
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<emphasis id="5FF7EAB8FF8BFF9BFE2EFF3E88426C83" bold="true" box="[448,544,223,250]" italics="true" pageId="2" pageNumber="453">badiyah</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="44C457C3FF8BFF9BFDC9FF3E88FD6C80" box="[551,671,223,249]" pageId="2" pageNumber="453" rank="species">spec. nov.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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||
<paragraph id="6D3C36AAFF8BFF9BFF79FF1F89FF6D6F" blockId="2.[151,925,223,278]" box="[151,925,254,278]" pageId="2" pageNumber="453">
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||
<uri id="19123AA8FF8BFF9BFF79FF1F89FF6D6F" box="[151,925,254,278]" pageId="2" pageNumber="453">
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||
urn:lsid:zoobank.org:act:
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||
<uuid id="19250C7FFF8BFF9BFE75FF1F89FF6D6F" box="[411,925,254,278]" pageId="2" pageNumber="453">7EED4526-5105-47EF-8EFB-102B70A7E89A</uuid>
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</uri>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="25996521FF8BFF9BFF79FEA58F606DFA" pageId="2" pageNumber="453" type="materials_examined">
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<paragraph id="6D3C36AAFF8BFF9BFF79FEA58E866D18" blockId="2.[151,1437,324,2043]" box="[151,1252,324,353]" pageId="2" pageNumber="453">
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||
<materialsCitation id="DDEB3CF7FF8BFF9BFF79FEA58E866D18" ID-GBIF-Occurrence="3050682306" box="[151,1252,324,353]" collectingDate="1996-04-23" collectionCode="OOLM" collectorName="Marek Halada" country="Jordan" location="Rawayshid" pageId="2" pageNumber="453" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
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||
<emphasis id="5FF7EAB8FF8BFF9BFF79FEA58BA66D27" bold="true" box="[151,452,324,350]" pageId="2" pageNumber="453">
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||
<typeStatus id="B2388808FF8BFF9BFF79FEA58B586D27" box="[151,314,324,350]" pageId="2" pageNumber="453" type="holotype">HOLOTYPE</typeStatus>
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||
:
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||
<collectingCountry id="1594763AFF8BFF9BFEA8FEA58BDD6D27" box="[326,447,324,350]" name="Jordan" pageId="2" pageNumber="453">JORDAN</collectingCountry>
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||
:
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||
</emphasis>
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||
<location id="685C6071FF8BFF9BFE25FEA588256D27" LSID="urn:lsid:plazi:treatment:E52A87BCFF8BFF9CFF79FF3E899B6DD4:685C6071FF8BFF9BFE25FEA588256D27" box="[459,583,324,350]" country="Jordan" name="Rawayshid" pageId="2" pageNumber="453">Rawayshid</location>
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||
[Ruwaished],
|
||
<date id="193D106AFF8BFF9BFD02FEA589066D27" box="[748,868,324,350]" pageId="2" pageNumber="453" value="1996-04-23">
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||
<collectingDate id="0979E982FF8BFF9BFD02FEA589066D27" box="[748,868,324,350]" pageId="2" pageNumber="453" value="1996-04-23">23.iv.1996</collectingDate>
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||
</date>
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||
,
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||
<specimenCount id="7B85FD23FF8BFF9BFC9EFEA489F16D26" box="[880,915,325,351]" pageId="2" pageNumber="453" type="female">1♀</specimenCount>
|
||
, leg.
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||
<collectorName id="C076537CFF8BFF9BFC3FFEA58E126D27" box="[977,1136,324,350]" pageId="2" pageNumber="453">Marek Halada</collectorName>
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||
(
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||
<collectionCode id="0B92AE6FFF8BFF9BFB91FEA78EBA6D18" box="[1151,1240,326,353]" pageId="2" pageNumber="453">OÖLM</collectionCode>
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||
).
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||
</materialsCitation>
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||
</paragraph>
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||
<paragraph id="6D3C36AAFF8BFF9BFF29FE898F606DFA" blockId="2.[151,1437,324,2043]" box="[199,1282,360,389]" pageId="2" pageNumber="453">
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<materialsCitation id="DDEB3CF7FF8BFF9BFF29FE8988976DFC" ID-GBIF-Occurrence="3050682313" box="[199,757,360,389]" country="Jordan" location="JORDAN" pageId="2" pageNumber="453" specimenCount="2" typeStatus="paratype">
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<emphasis id="5FF7EAB8FF8BFF9BFF29FE898B9B6DFB" bold="true" box="[199,505,360,386]" pageId="2" pageNumber="453">
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<typeStatus id="B2388808FF8BFF9BFF29FE898B0D6DFB" box="[199,367,360,386]" pageId="2" pageNumber="453" type="paratype">PARATYPES</typeStatus>
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:
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<collectingCountry id="1594763AFF8BFF9BFE95FE898B966DFB" box="[379,500,360,386]" name="Jordan" pageId="2" pageNumber="453">JORDAN</collectingCountry>
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||
:
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||
</emphasis>
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||
Same as
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||
<typeStatus id="B2388808FF8BFF9BFD8AFE8988AB6DFB" box="[612,713,360,386]" pageId="2" pageNumber="453" type="holotype">holotype</typeStatus>
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||
, 1³
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||
</materialsCitation>
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||
,
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<materialsCitation id="DDEB3CF7FF8BFF9BFCEBFE8889F56DFC" ID-GBIF-Occurrence="3050682335" box="[773,919,361,389]" collectionCode="OOLM" country="Jordan" pageId="2" pageNumber="453" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
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||
<specimenCount id="7B85FD23FF8BFF9BFCEBFE88894A6DFA" box="[773,808,361,387]" pageId="2" pageNumber="453" type="female">1♀</specimenCount>
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(
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||
<collectionCode id="0B92AE6FFF8BFF9BFCD9FE8B89ED6DFC" box="[823,911,362,389]" pageId="2" pageNumber="453">OÖLM</collectionCode>
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||
)
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||
</materialsCitation>
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||
,
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||
<materialsCitation id="DDEB3CF7FF8BFF9BFC4CFE888E496DFA" ID-GBIF-Occurrence="3050682312" box="[930,1067,361,387]" collectionCode="TJWC" country="Jordan" pageId="2" pageNumber="453" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
|
||
<specimenCount id="7B85FD23FF8BFF9BFC4CFE8889A76DFA" box="[930,965,361,387]" pageId="2" pageNumber="453" type="female">1♀</specimenCount>
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||
(
|
||
<collectionCode id="0B92AE6FFF8BFF9BFC3AFE888E416DFA" box="[980,1059,361,387]" pageId="2" pageNumber="453">TJWC</collectionCode>
|
||
)
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||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="DDEB3CF7FF8BFF9BFBD8FE888E9D6DFA" ID-GBIF-Occurrence="3050682343" box="[1078,1279,361,387]" collectionCode="SMNHTAU" country="Jordan" pageId="2" pageNumber="453" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
|
||
<specimenCount id="7B85FD23FF8BFF9BFBD8FE888E3B6DFA" box="[1078,1113,361,387]" pageId="2" pageNumber="453" type="female">1♀</specimenCount>
|
||
(
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||
<collectionCode id="0B92AE6FFF8BFF9BFB89FE888E946DFA" box="[1127,1270,361,387]" pageId="2" pageNumber="453">SMNHTAU</collectionCode>
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||
)
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||
</materialsCitation>
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||
.
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||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="25996521FF8BFF9BFF29FE6D8F726A16" pageId="2" pageNumber="453" type="description">
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<paragraph id="6D3C36AAFF8BFF9BFF29FE6D8E2068E2" blockId="2.[151,1437,324,2043]" pageId="2" pageNumber="453">
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||
<emphasis id="5FF7EAB8FF8BFF9BFF29FE6D8BA06DDE" bold="true" box="[199,450,396,423]" pageId="2" pageNumber="453">Description: Female:</emphasis>
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Body length
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<quantity id="AA7B9B4FFF8BFF9BFDB7FE6C88F86DDE" box="[601,666,397,423]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="2" pageNumber="453" unit="mm" value="7.0">7 mm</quantity>
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(
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<figureCitation id="F5B82A2FFF8BFF9BFD46FE6D89656DDE" box="[680,775,396,423]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1958,1982]" captionTargetBox="[151,1436,489,1935]" captionTargetId="figure-166@3.[151,1436,489,1935]" captionTargetPageId="3" captionText="FIGURES 1–6. Andrena badiyah spec. nov. 1. female profile, 2. female face, 3. female face detail, 4. female scutum, 5. female terga, 6. female terga detail." figureDoi="http://doi.org/10.5281/zenodo.4556548" httpUri="https://zenodo.org/record/4556548/files/figure.png" pageId="2" pageNumber="453">Figure 1</figureCitation>
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).
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||
<emphasis id="5FF7EAB8FF8BFF9BFCF1FE6D89076DDF" box="[799,869,396,422]" italics="true" pageId="2" pageNumber="453">Head:</emphasis>
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Black, 1.3 times wider than long (
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<figureCitation id="F5B82A2FFF8BFF9BFB08FE6D8F2A6DDE" box="[1254,1352,396,423]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1958,1982]" captionTargetBox="[151,1436,489,1935]" captionTargetId="figure-166@3.[151,1436,489,1935]" captionTargetPageId="3" captionText="FIGURES 1–6. Andrena badiyah spec. nov. 1. female profile, 2. female face, 3. female face detail, 4. female scutum, 5. female terga, 6. female terga detail." figureDoi="http://doi.org/10.5281/zenodo.4556548" httpUri="https://zenodo.org/record/4556548/files/figure.png" pageId="2" pageNumber="453">Figure 2</figureCitation>
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). Clypeus domed, irregularly punctate, punctures separated by 0.5–2 puncture diameters; longitudinal impunctate central line present, underlying surface weakly shagreened, stronger basally, becoming weaker apically. Process of labrum trapezoidal, corners rounded, shiny (
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<figureCitation id="F5B82A2FFF8BFF9BFDD7FE1988FC6E6A" box="[569,670,504,531]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1958,1982]" captionTargetBox="[151,1436,489,1935]" captionTargetId="figure-166@3.[151,1436,489,1935]" captionTargetPageId="3" captionText="FIGURES 1–6. Andrena badiyah spec. nov. 1. female profile, 2. female face, 3. female face detail, 4. female scutum, 5. female terga, 6. female terga detail." figureDoi="http://doi.org/10.5281/zenodo.4556548" httpUri="https://zenodo.org/record/4556548/files/figure.png" pageId="2" pageNumber="453">Figure 3</figureCitation>
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). Gena moderately broad, equalling width of compound eye; ocelloccipital distance subequal to width of lateral ocellus. Fovea moderately broad, dorsally occupying ½ of area between lateral ocellus and compound eye, slightly narrowed below to level of antennal insertions, not separated from inner margin of compound eye. Gena, vertex, face, and scape with white hairs, longest not exceeding half of length of scape. Antenna dark, A3 apically, A4–12 lightened orange below, A3 slightly exceeds A4+5, shorter than A4+5+6.
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<taxonomicName id="AA834D29FF8BFF9BFED4FD4C887B6EBE" authority=": Scutum" authorityName="Scutum" box="[314,537,685,711]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="453" phylum="Arthropoda" rank="genus">
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<emphasis id="5FF7EAB8FF8BFF9BFED4FD4C8BD96EBF" box="[314,443,685,710]" italics="true" pageId="2" pageNumber="453">Mesosoma:</emphasis>
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Scutum
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</taxonomicName>
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and scutellum finely and regularly punctate, punctures separated by 2 puncture diameters, underlying surface completely smooth and strongly shining (
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<figureCitation id="F5B82A2FFF8BFF9BFC23FD318E506E92" box="[973,1074,720,747]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1958,1982]" captionTargetBox="[151,1436,489,1935]" captionTargetId="figure-166@3.[151,1436,489,1935]" captionTargetPageId="3" captionText="FIGURES 1–6. Andrena badiyah spec. nov. 1. female profile, 2. female face, 3. female face detail, 4. female scutum, 5. female terga, 6. female terga detail." figureDoi="http://doi.org/10.5281/zenodo.4556548" httpUri="https://zenodo.org/record/4556548/files/figure.png" pageId="2" pageNumber="453">Figure 4</figureCitation>
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). Pronotum with weak humeral angle, dorsolaterally slightly angulate, inconspicuous. Episternum and propodeum finely reticulate, weakly shining, propodeal triangle granulose, indicated by change in reticulation, slightly duller, basally with very slight short rugosity, without ridges or rugae. Episternum and propodeum with moderately long white hairs, propodeal corbicula well-formed dorsally, lateral faces of propodeum regularly but sparsely haired, all hairs not exceeding length of scape. Scutum and scutellum with short, semi-squamous light brown hairs, very sparse centrally, dense along lateral margins of discs. Legs dark, apical tarsal segments and hind basitarsi lightened orange, pubescence whitish, scopa white. Hind tarsal claws without inner teeth. Wings hyaline, venation and stigma orange, nervulus strongly antefurcal.
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<emphasis id="5FF7EAB8FF8BFF9BFF2BFC108B216873" box="[197,323,1009,1034]" italics="true" pageId="2" pageNumber="453">Metasoma:</emphasis>
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Terga dark, apical margins lightened yellow and broadly hyaline (
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<figureCitation id="F5B82A2FFF8BFF9BFBF1FC118E1D6872" box="[1055,1151,1008,1035]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1958,1982]" captionTargetBox="[151,1436,489,1935]" captionTargetId="figure-166@3.[151,1436,489,1935]" captionTargetPageId="3" captionText="FIGURES 1–6. Andrena badiyah spec. nov. 1. female profile, 2. female face, 3. female face detail, 4. female scutum, 5. female terga, 6. female terga detail." figureDoi="http://doi.org/10.5281/zenodo.4556548" httpUri="https://zenodo.org/record/4556548/files/figure.png" pageId="2" pageNumber="453">Figure 5</figureCitation>
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), tergal discs with sparse, fine, inconspicuous hair-bearing punctures, becoming denser laterally, here separated by 2–3 puncture diameters (
|
||
<figureCitation id="F5B82A2FFF8BFF9BFF71FBD98B63682A" box="[159,257,1080,1107]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1958,1982]" captionTargetBox="[151,1436,489,1935]" captionTargetId="figure-166@3.[151,1436,489,1935]" captionTargetPageId="3" captionText="FIGURES 1–6. Andrena badiyah spec. nov. 1. female profile, 2. female face, 3. female face detail, 4. female scutum, 5. female terga, 6. female terga detail." figureDoi="http://doi.org/10.5281/zenodo.4556548" httpUri="https://zenodo.org/record/4556548/files/figure.png" pageId="2" pageNumber="453">Figure 6</figureCitation>
|
||
). Tergal discs with underlying surface microreticulate, weakly shining; T2–4 with thick white hair bands that obscure underlying surface, on T2 medially interrupted, T3–4 complete. Apical fringe of T5 and hairs flanking pygidial plate golden; pygidial plate flattened, centrally with fine mesh-like pattern.
|
||
</paragraph>
|
||
<paragraph id="6D3C36AAFF8BFF9BFF29FB448EB569C2" blockId="2.[151,1437,324,2043]" pageId="2" pageNumber="453">
|
||
<emphasis id="5FF7EAB8FF8BFF9BFF29FB448B6F68C6" bold="true" box="[199,269,1189,1215]" pageId="2" pageNumber="453">Male:</emphasis>
|
||
Body length
|
||
<quantity id="AA7B9B4FFF8BFF9BFE5EFB458B9668C6" box="[432,500,1188,1215]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="2" pageNumber="453" unit="mm" value="6.0">6 mm</quantity>
|
||
(
|
||
<figureCitation id="F5B82A2FFF8BFF9BFDE8FB45880E68C6" box="[518,620,1188,1215]" captionStart="FIGURES 7–12" captionStartId="4.[151,264,1648,1672]" captionTargetBox="[151,1436,181,1625]" captionTargetId="figure-20@4.[151,1436,181,1625]" captionTargetPageId="4" captionText="FIGURES 7–12. Andrena badiyah spec. nov. 7. male profile, 8. male face, 9. male scutum, 10. male terga, 11. male genitalia. Andrena spolata Warncke, 1968; 12. male genitalia." figureDoi="http://doi.org/10.5281/zenodo.4556552" httpUri="https://zenodo.org/record/4556552/files/figure.png" pageId="2" pageNumber="453">Figure 7</figureCitation>
|
||
).
|
||
<emphasis id="5FF7EAB8FF8BFF9BFD6BFB4588A868C7" box="[645,714,1188,1214]" italics="true" pageId="2" pageNumber="453">Head:</emphasis>
|
||
Black, 1.4 times wider than long (
|
||
<figureCitation id="F5B82A2FFF8BFF9BFB88FB458EAE68C6" box="[1126,1228,1188,1215]" captionStart="FIGURES 7–12" captionStartId="4.[151,264,1648,1672]" captionTargetBox="[151,1436,181,1625]" captionTargetId="figure-20@4.[151,1436,181,1625]" captionTargetPageId="4" captionText="FIGURES 7–12. Andrena badiyah spec. nov. 7. male profile, 8. male face, 9. male scutum, 10. male terga, 11. male genitalia. Andrena spolata Warncke, 1968; 12. male genitalia." figureDoi="http://doi.org/10.5281/zenodo.4556552" httpUri="https://zenodo.org/record/4556552/files/figure.png" pageId="2" pageNumber="453">Figure 8</figureCitation>
|
||
). Clypeus weakly domed, coloured yellow with exception of two lateral black marks, evenly punctate, punctures separated by 0.5–1 puncture diameter; weak longitudinal impunctate line present, underlying surface shagreened, weakly shining. Gena moderately broad, equalling width of compound eye; ocelloccipital distance subequal to width of lateral ocellus. Gena, vertex, face, and scape with white hairs, longest not exceeding half of length of scape. Antenna dark, A4–12 lightened orange below, A3 longer than A4 which is clearly wider than long, A3 shorter than A4+5.
|
||
<taxonomicName id="AA834D29FF8BFF9BFAF5FAB88A8F69EE" authority=": Scutum" authorityName="Scutum" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="453" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="5FF7EAB8FF8BFF9BFAF5FAB88FFE690B" box="[1307,1436,1369,1394]" italics="true" pageId="2" pageNumber="453">Mesosoma:</emphasis>
|
||
Scutum
|
||
</taxonomicName>
|
||
and scutellum sparsely and irregularly punctate, punctures separated by 2–3 puncture diameters, underlying surface smooth and strongly shining, shagreenation restricted to extreme disc margins (
|
||
<figureCitation id="F5B82A2FFF8BFF9BFB89FA418EAB69C2" box="[1127,1225,1440,1467]" captionStart="FIGURES 7–12" captionStartId="4.[151,264,1648,1672]" captionTargetBox="[151,1436,181,1625]" captionTargetId="figure-20@4.[151,1436,181,1625]" captionTargetPageId="4" captionText="FIGURES 7–12. Andrena badiyah spec. nov. 7. male profile, 8. male face, 9. male scutum, 10. male terga, 11. male genitalia. Andrena spolata Warncke, 1968; 12. male genitalia." figureDoi="http://doi.org/10.5281/zenodo.4556552" httpUri="https://zenodo.org/record/4556552/files/figure.png" pageId="2" pageNumber="453">Figure 9</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="6D3C36AAFF8BFF9BFF29FA248F726A16" blockId="2.[151,1437,324,2043]" pageId="2" pageNumber="453">
|
||
Pronotum, episternum and propodeum as in female. Episternum, propodeum, scutum, and scutellum with long white hairs, longest equalling length of scape. Legs dark, apexes of tibiae and tarsi lightened orange. Wings hyaline, venation and stigma orange, nervulus strongly antefurcal.
|
||
<emphasis id="5FF7EAB8FF8BFF9BFCDCF9EC89D26A5F" box="[818,944,1549,1574]" italics="true" pageId="2" pageNumber="453">Metasoma:</emphasis>
|
||
Terga structurally as in female, hair bands reduced (
|
||
<figureCitation id="F5B82A2FFF8BFF9BFEEEF9D18B116A32" box="[256,371,1584,1611]" captionStart="FIGURES 7–12" captionStartId="4.[151,264,1648,1672]" captionTargetBox="[151,1436,181,1625]" captionTargetId="figure-20@4.[151,1436,181,1625]" captionTargetPageId="4" captionText="FIGURES 7–12. Andrena badiyah spec. nov. 7. male profile, 8. male face, 9. male scutum, 10. male terga, 11. male genitalia. Andrena spolata Warncke, 1968; 12. male genitalia." figureDoi="http://doi.org/10.5281/zenodo.4556552" httpUri="https://zenodo.org/record/4556552/files/figure.png" pageId="2" pageNumber="453">Figure 10</figureCitation>
|
||
). Genitalia simple, gonocoxae with weak teeth, gonostyli with straight outer margin, apically rounded, penis valve basally relatively broad and parallel-sided before narrowing apically (
|
||
<figureCitation id="F5B82A2FFF8BFF9BFB7DF9B58F606A16" box="[1171,1282,1620,1647]" captionStart="FIGURES 7–12" captionStartId="4.[151,264,1648,1672]" captionTargetBox="[151,1436,181,1625]" captionTargetId="figure-20@4.[151,1436,181,1625]" captionTargetPageId="4" captionText="FIGURES 7–12. Andrena badiyah spec. nov. 7. male profile, 8. male face, 9. male scutum, 10. male terga, 11. male genitalia. Andrena spolata Warncke, 1968; 12. male genitalia." figureDoi="http://doi.org/10.5281/zenodo.4556552" httpUri="https://zenodo.org/record/4556552/files/figure.png" pageId="2" pageNumber="453">Figure 11</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="25996521FF8BFF9CFF29F9998E7D6D1C" lastPageId="5" lastPageNumber="456" pageId="2" pageNumber="453" type="diagnosis">
|
||
<paragraph id="6D3C36AAFF8BFF9BFF29F9998B1A6B13" blockId="2.[151,1437,324,2043]" pageId="2" pageNumber="453">
|
||
<emphasis id="5FF7EAB8FF8BFF9BFF29F9998B266AEB" bold="true" box="[199,324,1656,1682]" pageId="2" pageNumber="453">Diagnosis:</emphasis>
|
||
The placement of small black
|
||
<taxonomicName id="AA834D29FF8BFF9BFD47F999896A6AEB" box="[681,776,1656,1682]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="453" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="5FF7EAB8FF8BFF9BFD47F999896A6AEB" box="[681,776,1656,1682]" italics="true" pageId="2" pageNumber="453">Andrena</emphasis>
|
||
</taxonomicName>
|
||
is highly challenging because of the variation within this group and the polyphyly of the existing subgeneric concepts (
|
||
<bibRefCitation id="09124B5BFF8BFF9BFCB6F97D8E576ACE" author="Pisanty, G. & Richter, R. & Martin, T. & Dettman, J. & Cardinal, S." box="[856,1077,1692,1719]" pageId="2" pageNumber="453" refId="ref23835" refString="Pisanty, G., Richter, R., Martin, T., Dettman, J. & Cardinal, S. (2020) Molecular phylogeny and historical biogeography of an- drenine bees (Hymenoptera: Andrenidae). bioRxiv. [published online] https: // doi. org / 10.1101 / 2020.06.09.103556" type="book" year="2020">
|
||
Pisanty
|
||
<emphasis id="5FF7EAB8FF8BFF9BFC5BF97C898A6ACF" box="[949,1000,1692,1718]" italics="true" pageId="2" pageNumber="453">et al</emphasis>
|
||
. 2020
|
||
</bibRefCitation>
|
||
). Recognising which characters can be used to consistently identify good monophyletic groups is therefore difficult. The distinctive characters of
|
||
<taxonomicName id="AA834D29FF8BFF9BFF79F9048B736A87" authorityName="Wood" authorityYear="2021" box="[151,273,1764,1791]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="453" phylum="Arthropoda" rank="species" species="badiyah">
|
||
<emphasis id="5FF7EAB8FF8BFF9BFF79F9048B736A87" box="[151,273,1764,1791]" italics="true" pageId="2" pageNumber="453">A. badiyah</emphasis>
|
||
</taxonomicName>
|
||
are the strongly shiny and sparsely punctate scutum, the presence of short semi-squamous hairs on the thoracic dorsum, the antefurcal nervulus, the absence of sculpturing on the propodeal triangle, the orange hind basitarsi, the simple male genitalia, and the domed and striation-free clypeus in the female and the yellow-marked clypeus in the male.
|
||
</paragraph>
|
||
<paragraph id="6D3C36AAFF8BFF9AFF29F8958AAF6DA8" blockId="2.[151,1437,324,2043]" lastBlockId="3.[151,1437,151,465]" lastPageId="3" lastPageNumber="454" pageId="2" pageNumber="453">
|
||
The sparsely punctate and shiny scutum in combination with short semi-squamous hairs in the female sex is highly suggestive of an association with the red-marked
|
||
<taxonomicName id="AA834D29FF8BFF9BFCE7F8788EE26BCA" authority="Schmiedeknecht, 1900" authorityName="Schmiedeknecht" authorityYear="1900" box="[777,1152,1944,1971]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="453" phylum="Arthropoda" rank="species" species="arsinoe">
|
||
<emphasis id="5FF7EAB8FF8BFF9BFCE7F87889196BCB" box="[777,891,1945,1971]" italics="true" pageId="2" pageNumber="453">A. arsinoe</emphasis>
|
||
Schmiedeknecht, 1900
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="AA834D29FF8BFF9BFB5AF8788AAD6BAF" authority="Warncke, 1967" authorityName="Warncke" authorityYear="1967" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="453" phylum="Arthropoda" rank="species" species="amicula">
|
||
<emphasis id="5FF7EAB8FF8BFF9BFB5AF8788F4F6BCB" box="[1204,1325,1944,1971]" italics="true" pageId="2" pageNumber="453">A. amicula</emphasis>
|
||
<bibRefCitation id="09124B5BFF8BFF9BFADCF8798AAD6BAF" author="Warncke K." pageId="2" pageNumber="453" pagination="171 - 318" refId="ref24200" refString="Warncke K. (1967) Beitrag zur Klarung palaarktischer Andrena - Arten. Eos, 43, 171 - 318." type="journal article" year="1967">Warncke, 1967</bibRefCitation>
|
||
</taxonomicName>
|
||
which were placed in the
|
||
<emphasis id="5FF7EAB8FF8BFF9BFDE9F85D88CB6BAF" box="[519,681,1980,2006]" italics="true" pageId="2" pageNumber="453">Graecandrena</emphasis>
|
||
(
|
||
<bibRefCitation id="09124B5BFF8BFF9BFD54F85D89066BAE" author="Warncke, K." box="[698,868,1980,2007]" pageId="2" pageNumber="453" pagination="1 - 110" refId="ref24222" refString="Warncke, K. (1968) Die Untergattungen der westpalaarktischen Bienengattung Andrena F. Memorias e Estudos do Museu Zoologico da Universidade de Coimbra, 307, 1 - 110." type="journal article" year="1968">Warncke 1968</bibRefCitation>
|
||
), but these have strongly different male genitalia (gonostyli with strongly constricted, finger-like apexes) and have been shown to fall far from the other clades of small black
|
||
<taxonomicName id="AA834D29FF8AFF9AFECEFF7688126CC8" authority="(Pisanty et al. 2020)" baseAuthorityName="Pisanty" baseAuthorityYear="2020" box="[288,624,151,177]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="454" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="5FF7EAB8FF8AFF9AFECEFF768B1D6CC8" box="[288,383,151,177]" italics="true" pageId="3" pageNumber="454">Andrena</emphasis>
|
||
(
|
||
<bibRefCitation id="09124B5BFF8AFF9AFE60FF76880A6CC8" author="Pisanty, G. & Richter, R. & Martin, T. & Dettman, J. & Cardinal, S." box="[398,616,151,177]" pageId="3" pageNumber="454" refId="ref23835" refString="Pisanty, G., Richter, R., Martin, T., Dettman, J. & Cardinal, S. (2020) Molecular phylogeny and historical biogeography of an- drenine bees (Hymenoptera: Andrenidae). bioRxiv. [published online] https: // doi. org / 10.1101 / 2020.06.09.103556" type="book" year="2020">
|
||
Pisanty
|
||
<emphasis id="5FF7EAB8FF8AFF9AFE04FF76887E6CC8" box="[490,540,151,177]" italics="true" pageId="3" pageNumber="454">et al</emphasis>
|
||
. 2020
|
||
</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. The granulose (lacking ridges or rugae) propodeal triangle in combination with the moderately broad foveae should place it in the true
|
||
<taxonomicName id="AA834D29FF8AFF9AFC85FF5A898F6CAC" box="[875,1005,187,213]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="456" phylum="Arthropoda" rank="subGenus" subGenus="Aciandrena">
|
||
<emphasis id="5FF7EAB8FF8AFF9AFC85FF5A898F6CAC" box="[875,1005,187,213]" italics="true" pageId="3" pageNumber="454">Aciandrena</emphasis>
|
||
</taxonomicName>
|
||
clade (the species around
|
||
<taxonomicName id="AA834D29FF8AFF9AFAFBFF5A8B2F6C83" authority="Morawitz, 1886" authorityName="Morawitz" authorityYear="1886" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" species="aciculata">
|
||
<emphasis id="5FF7EAB8FF8AFF9AFAFBFF5A8FFE6CAC" box="[1301,1436,187,213]" italics="true" pageId="3" pageNumber="454">A. aciculata</emphasis>
|
||
Morawitz, 1886
|
||
</taxonomicName>
|
||
) as it lacks a propodeal triangle well-marked by carinae or rugosity (
|
||
<emphasis id="5FF7EAB8FF8AFF9AFB8DFF3E8E946C80" box="[1123,1270,223,249]" italics="true" pageId="3" pageNumber="454">Micrandrena</emphasis>
|
||
,
|
||
<taxonomicName id="AA834D29FF8AFF9AFAE8FF3E8FF46C80" authorityName="Warncke" authorityYear="1975" box="[1286,1430,223,249]" class="Insecta" genus="Fumandrena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="454" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="5FF7EAB8FF8AFF9AFAE8FF3E8FF46C80" box="[1286,1430,223,249]" italics="true" pageId="3" pageNumber="454">Fumandrena</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="AA834D29FF8AFF9AFF79FEE28B526D64" authorityName="Schmid-Egger" authorityYear="2005" box="[151,304,259,285]" class="Insecta" family="Andrenidae" genus="Proxiandrena" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="454" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="5FF7EAB8FF8AFF9AFF79FEE28B526D64" box="[151,304,259,285]" italics="true" pageId="3" pageNumber="454">Proxiandrena</emphasis>
|
||
</taxonomicName>
|
||
), a clypeus with longitudinal striations (
|
||
<emphasis id="5FF7EAB8FF8AFF9AFD14FEE289E76D64" box="[762,901,259,285]" italics="true" pageId="3" pageNumber="454">Distandrena</emphasis>
|
||
partim), or foveae narrowed below to the point of linearity (
|
||
<emphasis id="5FF7EAB8FF8AFF9AFECDFEC68BCC6D38" box="[291,430,295,321]" italics="true" pageId="3" pageNumber="454">Distandrena</emphasis>
|
||
partim), and it can be separated from the true
|
||
<emphasis id="5FF7EAB8FF8AFF9AFC5DFEC68E376D38" box="[947,1109,295,321]" italics="true" pageId="3" pageNumber="454">Graecandrena</emphasis>
|
||
by the absence of rugosity at the base of the propodeal triangle, by the yellow clypeus of the male, and by the male genitalia which do not have the typical
|
||
<emphasis id="5FF7EAB8FF8AFF9AFEFAFE8E8BD46DF0" box="[276,438,367,393]" italics="true" pageId="3" pageNumber="454">Graecandrena</emphasis>
|
||
form. However, because of the shiny scutal integument which is not known from otherp
|
||
<taxonomicName id="AA834D29FF8AFF9AFF79FE728B7B6DD4" box="[151,281,403,429]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="456" phylum="Arthropoda" rank="subGenus" subGenus="Aciandrena">
|
||
<emphasis id="5FF7EAB8FF8AFF9AFF79FE728B7B6DD4" box="[151,281,403,429]" italics="true" pageId="3" pageNumber="454">Aciandrena</emphasis>
|
||
</taxonomicName>
|
||
species, this placement should be considered provisional until further, preferably genetic, data are available.
|
||
</paragraph>
|
||
<caption id="39FC6622FF8AFF9AFF79F8478BD16B9B" ID-DOI="http://doi.org/10.5281/zenodo.4556548" ID-Zenodo-Dep="4556548" httpUri="https://zenodo.org/record/4556548/files/figure.png" pageId="3" pageNumber="454" startId="3.[151,264,1958,1982]" targetBox="[151,1436,489,1935]" targetPageId="3">
|
||
<paragraph id="6D3C36AAFF8AFF9AFF79F8478BD16B9B" blockId="3.[151,1436,1958,2019]" pageId="3" pageNumber="454">
|
||
<emphasis id="5FF7EAB8FF8AFF9AFF79F8478B586BC7" bold="true" box="[151,314,1958,1982]" pageId="3" pageNumber="454">FIGURES 1–6.</emphasis>
|
||
<taxonomicName id="AA834D29FF8AFF9AFEAEF8478B8C6BC7" authority="Wood, 2021" authorityName="Wood" authorityYear="2021" box="[320,494,1958,1982]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" species="badiyah" status="sp. nov.">
|
||
<emphasis id="5FF7EAB8FF8AFF9AFEAEF8478B8C6BC7" box="[320,494,1958,1982]" italics="true" pageId="3" pageNumber="454">Andrena badiyah</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="5FF7EAB8FF8AFF9AFE1DF84688386BC6" bold="true" box="[499,602,1959,1983]" pageId="3" pageNumber="454">
|
||
<taxonomicNameLabel id="44C457C3FF8AFF9AFE1DF84688386BC6" box="[499,602,1959,1983]" pageId="3" pageNumber="454" rank="species">spec. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
1. female profile, 2. female face, 3. female face detail, 4. female scutum, 5. female terga, 6. female terga detail.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="39FC6622FF8DFF9DFF79F99188C56AD4" ID-DOI="http://doi.org/10.5281/zenodo.4556552" ID-Zenodo-Dep="4556552" httpUri="https://zenodo.org/record/4556552/files/figure.png" pageId="4" pageNumber="455" startId="4.[151,264,1648,1672]" targetBox="[151,1436,181,1625]" targetPageId="4">
|
||
<paragraph id="6D3C36AAFF8DFF9DFF79F99188C56AD4" blockId="4.[151,1437,1648,1709]" pageId="4" pageNumber="455">
|
||
<emphasis id="5FF7EAB8FF8DFF9DFF79F9918B2A6AF0" bold="true" box="[151,328,1648,1673]" pageId="4" pageNumber="455">FIGURES 7–12.</emphasis>
|
||
<taxonomicName id="AA834D29FF8DFF9DFEA0F9908B9F6AF0" authority="Wood, 2021" authorityName="Wood" authorityYear="2021" box="[334,509,1649,1673]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="badiyah" status="sp. nov.">
|
||
<emphasis id="5FF7EAB8FF8DFF9DFEA0F9908B9F6AF0" box="[334,509,1649,1673]" italics="true" pageId="4" pageNumber="455">Andrena badiyah</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="5FF7EAB8FF8DFF9DFDEDF99088096AF0" bold="true" box="[515,619,1649,1673]" pageId="4" pageNumber="455">
|
||
<taxonomicNameLabel id="44C457C3FF8DFF9DFDEDF99088096AF0" box="[515,619,1649,1673]" pageId="4" pageNumber="455" rank="species">spec. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
7. male profile, 8. male face, 9. male scutum, 10. male terga, 11. male genitalia.
|
||
<taxonomicName id="AA834D29FF8DFF9DFF79F9748B826AD4" authority="Warncke, 1968" authorityName="Warncke" authorityYear="1968" box="[151,480,1685,1709]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="spolata">
|
||
<emphasis id="5FF7EAB8FF8DFF9DFF79F9748B5D6AD4" box="[151,319,1685,1709]" italics="true" pageId="4" pageNumber="455">Andrena spolata</emphasis>
|
||
<bibRefCitation id="09124B5BFF8DFF9DFEAAF9748B826AD4" author="Warncke, K." box="[324,480,1685,1709]" pageId="4" pageNumber="455" pagination="1 - 110" refId="ref24222" refString="Warncke, K. (1968) Die Untergattungen der westpalaarktischen Bienengattung Andrena F. Memorias e Estudos do Museu Zoologico da Universidade de Coimbra, 307, 1 - 110." type="journal article" year="1968">Warncke, 1968</bibRefCitation>
|
||
</taxonomicName>
|
||
; 12. male genitalia.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="6D3C36AAFF8DFF9CFF29F93A8E7D6D1C" blockId="4.[151,1437,1755,2034]" lastBlockId="5.[151,1437,151,430]" lastPageId="5" lastPageNumber="456" pageId="4" pageNumber="455">
|
||
For direct species diagnosis, in the female sex the combination of an almost entirely shiny scutum and scutellum together with short semi-squamous hairs and without red-marked terga (excluding
|
||
<taxonomicName id="AA834D29FF8DFF9DFB80F91E8E806B60" authorityName="Schmiedeknecht" authorityYear="1900" box="[1134,1250,1791,1817]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="arsinoe">
|
||
<emphasis id="5FF7EAB8FF8DFF9DFB80F91E8E806B60" box="[1134,1250,1791,1817]" italics="true" pageId="4" pageNumber="455">A. arsinoe</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="AA834D29FF8DFF9DFAF7F91E8FF16B60" authorityName="Warncke" authorityYear="1967" box="[1305,1427,1791,1817]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="amicula">
|
||
<emphasis id="5FF7EAB8FF8DFF9DFAF7F91E8FF16B60" box="[1305,1427,1791,1817]" italics="true" pageId="4" pageNumber="455">A. amicula</emphasis>
|
||
</taxonomicName>
|
||
) is distinctive, as similar
|
||
<taxonomicName id="AA834D29FF8DFF9DFE4AF8C288446B44" box="[420,550,1827,1853]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="456" phylum="Arthropoda" rank="subGenus" subGenus="Aciandrena">
|
||
<emphasis id="5FF7EAB8FF8DFF9DFE4AF8C288446B44" box="[420,550,1827,1853]" italics="true" pageId="4" pageNumber="455">Aciandrena</emphasis>
|
||
</taxonomicName>
|
||
species with similar short semi-squamous hairs have clear shagreenation on the scutal disc (e.g.
|
||
<taxonomicName id="AA834D29FF8DFF9DFEA2F8A6880E6B18" authority="Warncke, 1968" authorityName="Warncke" authorityYear="1968" box="[332,620,1863,1889]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="hillana">
|
||
<emphasis id="5FF7EAB8FF8DFF9DFEA2F8A68BDB6B18" box="[332,441,1863,1889]" italics="true" pageId="4" pageNumber="455">A. hillana</emphasis>
|
||
<bibRefCitation id="09124B5BFF8DFF9DFE2EF8A6880E6B18" author="Warncke, K." box="[448,620,1863,1889]" pageId="4" pageNumber="455" pagination="1 - 110" refId="ref24222" refString="Warncke, K. (1968) Die Untergattungen der westpalaarktischen Bienengattung Andrena F. Memorias e Estudos do Museu Zoologico da Universidade de Coimbra, 307, 1 - 110." type="journal article" year="1968">Warncke, 1968</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="AA834D29FF8DFF9DFD98F8A689C06B18" authority="Warncke, 1974" authorityName="Warncke" authorityYear="1974" box="[630,930,1863,1889]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="pavonia">
|
||
<emphasis id="5FF7EAB8FF8DFF9DFD98F8A6888D6B18" box="[630,751,1863,1889]" italics="true" pageId="4" pageNumber="455">A. pavonia</emphasis>
|
||
Warncke, 1974
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="AA834D29FF8DFF9DFC32F8A68F746B18" authority="Warncke, 1974" authorityName="Warncke" authorityYear="1974" box="[988,1302,1863,1889]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="pellucida">
|
||
<emphasis id="5FF7EAB8FF8DFF9DFC32F8A68E016B18" box="[988,1123,1863,1889]" italics="true" pageId="4" pageNumber="455">A. pellucida</emphasis>
|
||
Warncke, 1974
|
||
</taxonomicName>
|
||
), and where such shagreenation is weak (e.g.
|
||
<taxonomicName id="AA834D29FF8DFF9DFDE4F88A88F06BFC" authorityName="Warncke" authorityYear="1974" box="[522,658,1899,1925]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="pellucida">
|
||
<emphasis id="5FF7EAB8FF8DFF9DFDE4F88A88F06BFC" box="[522,658,1899,1925]" italics="true" pageId="4" pageNumber="455">A. pellucida</emphasis>
|
||
</taxonomicName>
|
||
) it can be further separated by the broader foveae that are wider than the width of a flagellum (as wide as a flagellum in
|
||
<taxonomicName id="AA834D29FF8DFF9DFD36F86E89026BD0" authorityName="Warncke" authorityYear="1974" box="[728,864,1935,1961]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="pellucida">
|
||
<emphasis id="5FF7EAB8FF8DFF9DFD36F86E89026BD0" box="[728,864,1935,1961]" italics="true" pageId="4" pageNumber="455">A. pellucida</emphasis>
|
||
</taxonomicName>
|
||
). It can be separated from
|
||
<taxonomicName id="AA834D29FF8DFF9DFB7EF86E8F406BD0" authority="Wood, 2021" authorityName="Wood" authorityYear="2021" box="[1168,1314,1935,1961]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="palmyriae" status="sp. nov.">
|
||
<emphasis id="5FF7EAB8FF8DFF9DFB7EF86E8F406BD0" box="[1168,1314,1935,1961]" italics="true" pageId="4" pageNumber="455">A. palmyriae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="5FF7EAB8FF8DFF9DFAC7F86E8FFE6BD0" bold="true" box="[1321,1436,1935,1961]" pageId="4" pageNumber="455">
|
||
<taxonomicNameLabel id="44C457C3FF8DFF9DFAC7F86E8FFE6BD0" box="[1321,1436,1935,1961]" pageId="4" pageNumber="455" rank="species">spec. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(see below) because the process of the labrum is entire (not apically notched), the hind tibiae are dark (not orange), the nervulus is antefurcal (interstitial), and the tergal hair bands are shorter and less dense. In the male sex, complete diagnosis is difficult because many of the desert-living
|
||
<taxonomicName id="AA834D29FF8CFF9CFCE9FF7689EB6CC8" box="[775,905,151,177]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="456" phylum="Arthropoda" rank="subGenus" subGenus="Aciandrena">
|
||
<emphasis id="5FF7EAB8FF8CFF9CFCE9FF7689EB6CC8" box="[775,905,151,177]" italics="true" pageId="5" pageNumber="456">Aciandrena</emphasis>
|
||
</taxonomicName>
|
||
have been described only from the female sex, but the combination of yellow clypeus, granulose propodeal triangle, shiny scutum and scutellum, antefurcal nervulus, orange hind basitarsi, and genitalia with weakly produced gonocoxal teeth should be sufficient to facilitate identification, as most
|
||
<taxonomicName id="AA834D29FF8CFF9CFE7AFEE288746D64" box="[404,534,259,285]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="456" phylum="Arthropoda" rank="subGenus" subGenus="Aciandrena">
|
||
<emphasis id="5FF7EAB8FF8CFF9CFE7AFEE288746D64" box="[404,534,259,285]" italics="true" pageId="5" pageNumber="456">Aciandrena</emphasis>
|
||
</taxonomicName>
|
||
males have more strongly produced gonocoxal teeth such as
|
||
<taxonomicName id="AA834D29FF8CFF9CFB52FEE28AAD6D38" authority="Warncke, 1968" authorityName="Warncke" authorityYear="1968" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="456" phylum="Arthropoda" rank="species" species="spolata">
|
||
<emphasis id="5FF7EAB8FF8CFF9CFB52FEE28F4E6D64" box="[1212,1324,259,285]" italics="true" pageId="5" pageNumber="456">A. spolata</emphasis>
|
||
<bibRefCitation id="09124B5BFF8CFF9CFADCFEE28AAD6D38" author="Warncke, K." pageId="5" pageNumber="456" pagination="1 - 110" refId="ref24222" refString="Warncke, K. (1968) Die Untergattungen der westpalaarktischen Bienengattung Andrena F. Memorias e Estudos do Museu Zoologico da Universidade de Coimbra, 307, 1 - 110." type="journal article" year="1968">Warncke, 1968</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="F5B82A2FFF8CFF9CFF30FEC68B2C6D38" box="[222,334,295,321]" captionStart="FIGURES 7–12" captionStartId="4.[151,264,1648,1672]" captionTargetBox="[151,1436,181,1625]" captionTargetId="figure-20@4.[151,1436,181,1625]" captionTargetPageId="4" captionText="FIGURES 7–12. Andrena badiyah spec. nov. 7. male profile, 8. male face, 9. male scutum, 10. male terga, 11. male genitalia. Andrena spolata Warncke, 1968; 12. male genitalia." figureDoi="http://doi.org/10.5281/zenodo.4556552" httpUri="https://zenodo.org/record/4556552/files/figure.png" pageId="5" pageNumber="456">Figure 12</figureCitation>
|
||
, see also photographs and notes in
|
||
<bibRefCitation id="09124B5BFF8CFF9CFD37FEC689CC6D3B" author="Pisanty, G. & Scheuchl, E. & Dorchin, N." box="[729,942,295,322]" pageId="5" pageNumber="456" pagination="485 - 515" refId="ref23712" refString="Pisanty, G., Scheuchl, E. & Dorchin, N. (2016) Eight new species of Andrena Fabricius (Hymenoptera: Apoidea: Andrenidae) from Israel - a Mediterranean hotspot for wild bees. Zootaxa, 4189 (3), 485 - 515. https: // doi. org / 10.11646 / zootaxa. 4189.3.3" type="journal article" year="2016">
|
||
Pisanty
|
||
<emphasis id="5FF7EAB8FF8CFF9CFCDCFEC689016D38" box="[818,867,295,321]" italics="true" pageId="5" pageNumber="456">et al</emphasis>
|
||
. 2016
|
||
</bibRefCitation>
|
||
). The genitalia are extremely close to
|
||
<taxonomicName id="AA834D29FF8CFF9CFAB6FEC68AB76D1D" authorityName="Warncke" authorityYear="1974" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="456" phylum="Arthropoda" rank="species" species="pavonia">
|
||
<emphasis id="5FF7EAB8FF8CFF9CFAB6FEC68AB76D1D" italics="true" pageId="5" pageNumber="456">A. pavonia</emphasis>
|
||
</taxonomicName>
|
||
, but this species has a black clypeus (see
|
||
<bibRefCitation id="09124B5BFF8CFF9CFD44FEAA8E716D1C" author="Gusenleitner, F. & Schwarz, M." box="[682,1043,331,357]" pageId="5" pageNumber="456" pagination="1 - 1280" refId="ref23399" refString="Gusenleitner, F. & Schwarz, M. (2002) Weltweite Checkliste der Bienengattung Andrena mit Bemerkungen und Erganzungen zu palaarktischen Arten (Hymenoptera, Apidae, Andreninae, Andrena). Entomofauna, Supplement 10, 1 - 1280." type="journal article" year="2002">Gusenleitner and Schwarz 2002</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="25996521FF8CFF9CFF29FE8E899B6DD4" pageId="5" pageNumber="456" type="etymology">
|
||
<paragraph id="6D3C36AAFF8CFF9CFF29FE8E899B6DD4" blockId="5.[151,1437,151,430]" pageId="5" pageNumber="456">
|
||
<emphasis id="5FF7EAB8FF8CFF9CFF29FE8E8B306DF0" bold="true" box="[199,338,367,393]" pageId="5" pageNumber="456">Etymology:</emphasis>
|
||
The name is derived from the Arabic name for the Syrian desert, the Badia (Arabic: Bâdiyah Ash-Shâm) that stretches over large parts of
|
||
<collectingCountry id="1594763AFF8CFF9CFDBAFE7288F06DD4" box="[596,658,403,429]" name="Syria" pageId="5" pageNumber="456">Syria</collectingCountry>
|
||
,
|
||
<collectingCountry id="1594763AFF8CFF9CFD73FE7288896DD4" box="[669,747,403,429]" name="Jordan" pageId="5" pageNumber="456">Jordan</collectingCountry>
|
||
,
|
||
<collectingCountry id="1594763AFF8CFF9CFD18FE7289E86DD4" box="[758,906,403,429]" name="Saudi Arabia" pageId="5" pageNumber="456">Saudi Arabia</collectingCountry>
|
||
, and
|
||
<collectingCountry id="1594763AFF8CFF9CFC2BFE7289946DD4" box="[965,1014,403,429]" name="Iraq" pageId="5" pageNumber="456">Iraq</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |