388 lines
69 KiB
XML
388 lines
69 KiB
XML
<document ID-CLB-Dataset="100959" ID-DOI="http://dx.doi.org/10.3897/asp.80.e82644" ID-GBIF-Dataset="1d579114-3554-4e38-bd81-ec33a8c44b9f" ID-Pensoft-Pub="1864-8312-80-279" ID-Pensoft-UUID="42DA6B89695151758680B79C0CF21997" ID-ZooBank="1D57CB0C56C44DA591E1CF7E9BB40E74" ModsDocID="1864-8312-80-279" checkinTime="1658318833579" checkinUser="pensoft" docAuthor="Yin, Zi-Wei, Tihelka, Erik, Lozano-Fernandez, Jesus & Cai, Chen-Yang" docDate="2022" docId="E28824C68DB052F68688C5F1D149E573" docLanguage="en" docName="ArthropodSystPhyl 80: 279-294" docOrigin="Arthropod Systematics & amp; Phylogeny 80" docPubDate="2022-07-19" docSource="http://dx.doi.org/10.3897/asp.80.e82644" docTitle="Europharinodes Yin & Cai 2022, gen. nov." docType="treatment" docUuid="0B631F8C-D490-4174-89F9-EE6274592F92" docUuidSource="ZooBank" docVersion="5" id="42DA6B89695151758680B79C0CF21997" lastPageNumber="279" masterDocId="42DA6B89695151758680B79C0CF21997" masterDocTitle="The first fossil Hybocephalini (Coleoptera: Staphylinidae: Pselaphinae) from the middle Eocene of Europe and its evolutionary and biogeographic implications" masterLastPageNumber="294" masterPageNumber="279" pageNumber="279" updateTime="1732844240309" updateUser="ExternalLinkService">
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<mods:titleInfo id="B3545FCD96B2E6CB39E3140D31ED9F19">
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<mods:title id="03E9CCFFACF396E4779DF56636BEA0AE">The first fossil Hybocephalini (Coleoptera: Staphylinidae: Pselaphinae) from the middle Eocene of Europe and its evolutionary and biogeographic implications</mods:title>
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<mods:namePart id="6BEEDCECD13FD71D37F2FD7EA5C11811">Yin, Zi-Wei</mods:namePart>
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<mods:affiliation id="56E99FC20AF50E7C66351EA622299542">Laboratory of Systematic Entomology, College of Life and Environmental Sciences, Shanghai Normal University, Shanghai, 200234, China</mods:affiliation>
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<mods:nameIdentifier id="009DB89263C5B2F456B55245B9A41FD0" type="email">pselaphinae@gmail.com</mods:nameIdentifier>
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<mods:namePart id="E26DAAC0B8C5F0CA8ACAE1F8295608F1">Tihelka, Erik</mods:namePart>
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<mods:nameIdentifier id="E785DF4C21A1C90038FBC4D6A5C3B4E4" type="ORCID">https://orcid.org/0000-0002-5048-5355</mods:nameIdentifier>
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<mods:affiliation id="8B99A4B743A654FCB9E57309CCACF94A">State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, and Center for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, Nanjing 210008, China; cycai @ nigpas. ac. cn & School of Earth Sciences, University of Bristol, Life Sciences Building, Tyndall Avenue, Bristol, BS 8 1 TQ, UK</mods:affiliation>
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<mods:namePart id="686D3BEBB4650E568BAF71F861F1D278">Lozano-Fernandez, Jesus</mods:namePart>
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<mods:affiliation id="EE797A7947F850CC290EAEA0AC8B41D7">Department of Genetics, Microbiology and Statistics, Biodiversity Research Institute (IRBio), University of Barcelona, Avd. Diagonal 643, 08028</mods:affiliation>
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<mods:namePart id="4E937499F9E2666B8466F869605D7DD4">Cai, Chen-Yang</mods:namePart>
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<mods:affiliation id="F088BA02A5BE163641682127EA69D35B">State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, and Center for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, Nanjing 210008, China; cycai @ nigpas. ac. cn & School of Earth Sciences, University of Bristol, Life Sciences Building, Tyndall Avenue, Bristol, BS 8 1 TQ, UK</mods:affiliation>
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<mods:title id="2E548EFCA7A6E8A862D157186B389D11">Arthropod Systematics & amp; Phylogeny</mods:title>
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<mods:date id="95853CE903C44A8DD36BD6F11E67F999">2022</mods:date>
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<mods:number id="65B5DE7545606E1B568CA689D2025BC7">2022-07-19</mods:number>
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<mods:number id="A70C06A0588A03AB237257FBAB0075AF">80</mods:number>
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<mods:start id="A8F65F41BB282ABDF566F28A93F3E84F">279</mods:start>
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<mods:identifier id="21134C69C5F23D5A9C96DE8D5DF9268E" type="DOI">http://dx.doi.org/10.3897/asp.80.e82644</mods:identifier>
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<treatment id="E28824C68DB052F68688C5F1D149E573" ID-GBIF-Taxon="197824438" LSID="urn:lsid:zoobank.org:act:0B631F8C-D490-4174-89F9-EE6274592F92" httpUri="http://treatment.plazi.org/id/E28824C68DB052F68688C5F1D149E573" lastPageNumber="279" pageId="0" pageNumber="279" scope_family="Staphylinidae" scope_order="Coleoptera" scope_subFamily="Pselaphinae" scope_tribe="Hybocephalini">
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<subSubSection id="49FF63C7FB8285D8460186EBC58E54BC" pageId="0" pageNumber="279" type="nomenclature">
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<paragraph id="1530D9F4F287B71923ABE4596183762C" pageId="0" pageNumber="279">
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<taxonomicName id="E57E56EC9EE815AF4F2C13E2C96A9D52" ID-CoL="9GDKX" LSID="http://zoobank.org/0B631F8C-D490-4174-89F9-EE6274592F92" authority="Yin & Cai" authorityName="Yin & Cai" authorityYear="2022" class="Insecta" family="Staphylinidae" genus="Europharinodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Europharinodes" order="Coleoptera" pageId="0" pageNumber="279" phylum="Arthropoda" rank="genus" status="gen. nov.">Europharinodes Yin & Cai</taxonomicName>
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<taxonomicNameLabel id="45419052D5A8504C34302E132C158CBD" pageId="0" pageNumber="279">gen. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="CCFCF85D94CFD9249536D4AE603A4DB3" pageId="0" pageNumber="279" type="description">
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<paragraph id="E90477F1FF52B8CB2ABC92AC66B8C9E7" pageId="0" pageNumber="279">
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<figureCitation id="0A076201B31119312F52654EE1CB1E3E" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Amber piece (A) containing Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102, and its morphological details (B-G) of (A, B, G: under incident light; C-F: X-ray microtomographic reconstruction). A Original amber piece containing the holotype before cutting and polishing. B, C Left lateral habitus. D Right lateral habitus. E Head in lateral view, show tentorial arms and pharynx. F Right antennal club. G Pro- and metatarsus and respective pretarsal claws. Abbreviations: aed, aedeagus; hw, hind wing; ph, pharynx; ta, tentorial arm. Scale bars: 0.5 mm in B-D, 0.2 mm in E, F." figureDoi="10.3897/asp.80.e82644.figure2" httpUri="https://binary.pensoft.net/fig/717124" pageId="0" pageNumber="279">Figs 2</figureCitation>
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<figureCitation id="707D414C2A1C364E317E9B8D74B51899" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Morphology of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102 (A, C, D, I: under incident light; B, E-H: X-ray microtomographic reconstruction). A, B Dorsal habitus. C Left half of pronotum, showing squamous setae. D Lateral portion of the abdominal base, showing squamous setae. E Head dorsum. F Head venter. G Head, in antero-lateral view. H, I Left maxillary palpus. Abbreviations: ce, compound eye; cl, clypeus; ff, frontal fovea; gf, gular fovea; la, labrum; ma, mandible; p 2 - 4, maxillary palpomere 2 - 4; vf, vertexal fovea. Scale bars: 0.5 mm in A, B, 0.2 mm in E-I." figureDoi="10.3897/asp.80.e82644.figure3" httpUri="https://binary.pensoft.net/fig/717125" pageId="0" pageNumber="279">, 3</figureCitation>
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<figureCitation id="78BF80ABA591B108D44975D65C075D93" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. X-ray microtomographic reconstruction of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102. A - C: prothorax, dorsal (A), lateral (B) and ventral (C). D Right elytron. E, F Meso- and metathorax, ventral (E) and lateral (F). G - I Abdomen, dorsal (G), lateral (H) and ventral (I). Abbreviations: ahf, antero-hypomeral fovea; bef, basal elytral fovea; blf, basal lateral fovea; ds, discal stria; laf, lateral antebasal fovea; lmcf, lateral metaventral fovea; lpcf, lateral procoxal fovea; maf, median antebasal fovea; mmsf, median mesoventral fovea; mc, marginal carina; mmtf, median metaventral fovea; mn, median nodule; ms, median split; pt 1 - 3, paratergite 1 - 3; s 2 - 6, sternite 2 - 6 (IV-VIII); t 1 - 5, tergite 1 - 5 (IV-VIII). Scale bars: 0.2 mm." figureDoi="10.3897/asp.80.e82644.figure4" httpUri="https://binary.pensoft.net/fig/717126" pageId="0" pageNumber="279">, 4</figureCitation>
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<figureCitation id="08DBD0CEB2695C7A2FC42F4478C2831A" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Morphology of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102 (A-H: X-ray microtomographic reconstruction; I-K: schematic). A Right foreleg. B Right middle leg. C Left hindleg. D - F, I - K Aedeagus, dorsal (D, I), lateral (E, J) and ventral (F, K). G, H endophallus, dorsal (G) and lateral (H). Abbreviations: bc, basal capsule; bp, basoventral projection; dd, dorsal diaphragm; en, endophallus; ml, median lobe; pa, paramere. Scale bars: 0.3 mm in A-C; 0.1 mm in D-K." figureDoi="10.3897/asp.80.e82644.figure5" httpUri="https://binary.pensoft.net/fig/717127" pageId="0" pageNumber="279">, 5</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="D40FE597CC3FB042FF3909319E8AEE22" pageId="0" pageNumber="279" type="type species">
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<paragraph id="4D8546E662B7BB3A5873D7706206D7D0" pageId="0" pageNumber="279">Type species.</paragraph>
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<paragraph id="86C296703E8261EF8B98E078860F25E6" pageId="0" pageNumber="279">
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<taxonomicName id="272D43D2A43977DD2107F0E4ECDE79F9" authorityName="Yin & Cai" authorityYear="2022" class="Insecta" family="Staphylinidae" genus="Europharinodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Europharinodes schaufussi" order="Coleoptera" pageId="0" pageNumber="279" phylum="Arthropoda" rank="species" species="schaufussi">
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<emphasis id="35656E0462AB3FFF37EFC904F6036ED6" italics="true" pageId="0" pageNumber="279">Europharinodes schaufussi</emphasis>
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</taxonomicName>
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<emphasis id="20D9D5ECBD199944AB2FC0967ED4D9C8" bold="true" pageId="0" pageNumber="279">sp. nov.</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="57F5F27A8DA64C4DF5216FB5365CF215" pageId="0" pageNumber="279" type="diagnosis">
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<paragraph id="F40C51E71D9FCF22D1782EC1DF3628A2" pageId="0" pageNumber="279">Diagnosis.</paragraph>
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<paragraph id="1FD81E4282BB1D7D92FC2D5C4A01106B" pageId="0" pageNumber="279">Most of body covered with squamous setae; body generally compact; head, pronotum and femora coarsely punctate. Head roundly-triangular, with short and narrow frontal rostrum; large vertexal and frontal foveae present; antennal insertions close, antennomeres slightly elongate to sub-moniliform, clubs formed by apical three enlarged antennomeres; lacking ocular-mandibular carinae; clypeus sharply sloping, anterior margin carinate and rounded, lateral margins straight to eyes; maxillary palpi small, four-segmented, with short apical palpal cone. Pronotum with median and lateral antebasal foveae obscured by squamous setae; with antero-hypomeral foveae, lacking hypomeral carinae. Elytra each with two large basal foveae and two distinct discal striae. Each tarsus plesiomorphically with two subequal claws. Abdomen with tergite 1 (IV) slightly longer than 2 (V), tergites 1-4 (IV-VII) broadly sulcate at bases; paratergites moderately broad and laterally protruding. Aedeagus symmetrical, basal capsule enlarged, paired parameres elongate.</paragraph>
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</subSubSection>
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<subSubSection id="088297026EE08859ECB105E802C88891" pageId="0" pageNumber="279" type="description">
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<paragraph id="9287AE756757E2A49FCFF14E75C7A095" pageId="0" pageNumber="279">Description.</paragraph>
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<paragraph id="BD5EBD2DE8429A9C784548FA8DA55924" pageId="0" pageNumber="279">
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Habitus (Figs
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<figureCitation id="053B57932AF60DB333CFC3742F263C8E" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Amber piece (A) containing Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102, and its morphological details (B-G) of (A, B, G: under incident light; C-F: X-ray microtomographic reconstruction). A Original amber piece containing the holotype before cutting and polishing. B, C Left lateral habitus. D Right lateral habitus. E Head in lateral view, show tentorial arms and pharynx. F Right antennal club. G Pro- and metatarsus and respective pretarsal claws. Abbreviations: aed, aedeagus; hw, hind wing; ph, pharynx; ta, tentorial arm. Scale bars: 0.5 mm in B-D, 0.2 mm in E, F." figureDoi="10.3897/asp.80.e82644.figure2" httpUri="https://binary.pensoft.net/fig/717124" pageId="0" pageNumber="279">2B-D</figureCitation>
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;
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<figureCitation id="D5A6F3F0DACC7143A6E25DEF9FCE5A26" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Morphology of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102 (A, C, D, I: under incident light; B, E-H: X-ray microtomographic reconstruction). A, B Dorsal habitus. C Left half of pronotum, showing squamous setae. D Lateral portion of the abdominal base, showing squamous setae. E Head dorsum. F Head venter. G Head, in antero-lateral view. H, I Left maxillary palpus. Abbreviations: ce, compound eye; cl, clypeus; ff, frontal fovea; gf, gular fovea; la, labrum; ma, mandible; p 2 - 4, maxillary palpomere 2 - 4; vf, vertexal fovea. Scale bars: 0.5 mm in A, B, 0.2 mm in E-I." figureDoi="10.3897/asp.80.e82644.figure3" httpUri="https://binary.pensoft.net/fig/717125" pageId="0" pageNumber="279">3A, B</figureCitation>
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) stout, compact; most of body with squamous setae. Head (Fig.
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<figureCitation id="0AC2B9A80186FE6BD1FFF3B975EA8835" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Morphology of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102 (A, C, D, I: under incident light; B, E-H: X-ray microtomographic reconstruction). A, B Dorsal habitus. C Left half of pronotum, showing squamous setae. D Lateral portion of the abdominal base, showing squamous setae. E Head dorsum. F Head venter. G Head, in antero-lateral view. H, I Left maxillary palpus. Abbreviations: ce, compound eye; cl, clypeus; ff, frontal fovea; gf, gular fovea; la, labrum; ma, mandible; p 2 - 4, maxillary palpomere 2 - 4; vf, vertexal fovea. Scale bars: 0.5 mm in A, B, 0.2 mm in E-I." figureDoi="10.3897/asp.80.e82644.figure3" httpUri="https://binary.pensoft.net/fig/717125" pageId="0" pageNumber="279">3E</figureCitation>
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) roundly triangular; vertex coarsely punctate, vertexal foveae (= dorsal tentorial pits; Fig.
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<figureCitation id="9280846BD969ABE1A1381A6729FDF03F" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Morphology of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102 (A, C, D, I: under incident light; B, E-H: X-ray microtomographic reconstruction). A, B Dorsal habitus. C Left half of pronotum, showing squamous setae. D Lateral portion of the abdominal base, showing squamous setae. E Head dorsum. F Head venter. G Head, in antero-lateral view. H, I Left maxillary palpus. Abbreviations: ce, compound eye; cl, clypeus; ff, frontal fovea; gf, gular fovea; la, labrum; ma, mandible; p 2 - 4, maxillary palpomere 2 - 4; vf, vertexal fovea. Scale bars: 0.5 mm in A, B, 0.2 mm in E-I." figureDoi="10.3897/asp.80.e82644.figure3" httpUri="https://binary.pensoft.net/fig/717125" pageId="0" pageNumber="279">3E</figureCitation>
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;
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<emphasis id="7A15E09B0093A497BEE9F81CC290D1B3" italics="true" pageId="0" pageNumber="279">vf</emphasis>
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) relatively large, obscured by squamous setae; frontal fovea (Fig.
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<figureCitation id="17688833BE44D66939F56F665CCA175E" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Morphology of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102 (A, C, D, I: under incident light; B, E-H: X-ray microtomographic reconstruction). A, B Dorsal habitus. C Left half of pronotum, showing squamous setae. D Lateral portion of the abdominal base, showing squamous setae. E Head dorsum. F Head venter. G Head, in antero-lateral view. H, I Left maxillary palpus. Abbreviations: ce, compound eye; cl, clypeus; ff, frontal fovea; gf, gular fovea; la, labrum; ma, mandible; p 2 - 4, maxillary palpomere 2 - 4; vf, vertexal fovea. Scale bars: 0.5 mm in A, B, 0.2 mm in E-I." figureDoi="10.3897/asp.80.e82644.figure3" httpUri="https://binary.pensoft.net/fig/717125" pageId="0" pageNumber="279">3E</figureCitation>
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;
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<emphasis id="8965B1DCA80E6851BF132A608574D423" italics="true" pageId="0" pageNumber="279">ff</emphasis>
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) distinct, rostrum short and narrow, antennal insertions located at ventral surface of rostrum, tubercles barely raised; compound eyes (Fig.
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<figureCitation id="7D505613FA35A3C6CBEDC53E6F27FBE1" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Morphology of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102 (A, C, D, I: under incident light; B, E-H: X-ray microtomographic reconstruction). A, B Dorsal habitus. C Left half of pronotum, showing squamous setae. D Lateral portion of the abdominal base, showing squamous setae. E Head dorsum. F Head venter. G Head, in antero-lateral view. H, I Left maxillary palpus. Abbreviations: ce, compound eye; cl, clypeus; ff, frontal fovea; gf, gular fovea; la, labrum; ma, mandible; p 2 - 4, maxillary palpomere 2 - 4; vf, vertexal fovea. Scale bars: 0.5 mm in A, B, 0.2 mm in E-I." figureDoi="10.3897/asp.80.e82644.figure3" httpUri="https://binary.pensoft.net/fig/717125" pageId="0" pageNumber="279">3G</figureCitation>
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;
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<emphasis id="0298687963A4E1409221966113DDB580" italics="true" pageId="0" pageNumber="279">ce</emphasis>
|
|
) large, prominent; lacking ocular-mandibular carina; clypeus (Fig.
|
|
<figureCitation id="39BCE5B9DB44540A37745891392E7387" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Morphology of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102 (A, C, D, I: under incident light; B, E-H: X-ray microtomographic reconstruction). A, B Dorsal habitus. C Left half of pronotum, showing squamous setae. D Lateral portion of the abdominal base, showing squamous setae. E Head dorsum. F Head venter. G Head, in antero-lateral view. H, I Left maxillary palpus. Abbreviations: ce, compound eye; cl, clypeus; ff, frontal fovea; gf, gular fovea; la, labrum; ma, mandible; p 2 - 4, maxillary palpomere 2 - 4; vf, vertexal fovea. Scale bars: 0.5 mm in A, B, 0.2 mm in E-I." figureDoi="10.3897/asp.80.e82644.figure3" httpUri="https://binary.pensoft.net/fig/717125" pageId="0" pageNumber="279">3G</figureCitation>
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;
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<emphasis id="D63F809C51A725E2BFAD4A0F1FCF98D1" italics="true" pageId="0" pageNumber="279">cl</emphasis>
|
|
) sharply sloping, with broadly rounded, ridged anterior margin; labrum (Fig.
|
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<figureCitation id="BECDAB3AB73C2F0F33F929836044CF1E" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Morphology of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102 (A, C, D, I: under incident light; B, E-H: X-ray microtomographic reconstruction). A, B Dorsal habitus. C Left half of pronotum, showing squamous setae. D Lateral portion of the abdominal base, showing squamous setae. E Head dorsum. F Head venter. G Head, in antero-lateral view. H, I Left maxillary palpus. Abbreviations: ce, compound eye; cl, clypeus; ff, frontal fovea; gf, gular fovea; la, labrum; ma, mandible; p 2 - 4, maxillary palpomere 2 - 4; vf, vertexal fovea. Scale bars: 0.5 mm in A, B, 0.2 mm in E-I." figureDoi="10.3897/asp.80.e82644.figure3" httpUri="https://binary.pensoft.net/fig/717125" pageId="0" pageNumber="279">3G</figureCitation>
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;
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<emphasis id="91FAB360DC93BBDA8AD4D80F17BB5DA5" italics="true" pageId="0" pageNumber="279">la</emphasis>
|
|
) transverse, subtrapezoidal; mandible (Fig.
|
|
<figureCitation id="7947AB8827583CAC6AC44E031191BDFA" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Morphology of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102 (A, C, D, I: under incident light; B, E-H: X-ray microtomographic reconstruction). A, B Dorsal habitus. C Left half of pronotum, showing squamous setae. D Lateral portion of the abdominal base, showing squamous setae. E Head dorsum. F Head venter. G Head, in antero-lateral view. H, I Left maxillary palpus. Abbreviations: ce, compound eye; cl, clypeus; ff, frontal fovea; gf, gular fovea; la, labrum; ma, mandible; p 2 - 4, maxillary palpomere 2 - 4; vf, vertexal fovea. Scale bars: 0.5 mm in A, B, 0.2 mm in E-I." figureDoi="10.3897/asp.80.e82644.figure3" httpUri="https://binary.pensoft.net/fig/717125" pageId="0" pageNumber="279">3G</figureCitation>
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;
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<emphasis id="7F57E41AF41C72382174FDF70AFD65C5" italics="true" pageId="0" pageNumber="279">ma</emphasis>
|
|
) with single distinct apical and preapical teeth; maxillary palpus small (Fig.
|
|
<figureCitation id="9180CABFC1FEA3610FF4B84687CA3C35" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Morphology of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102 (A, C, D, I: under incident light; B, E-H: X-ray microtomographic reconstruction). A, B Dorsal habitus. C Left half of pronotum, showing squamous setae. D Lateral portion of the abdominal base, showing squamous setae. E Head dorsum. F Head venter. G Head, in antero-lateral view. H, I Left maxillary palpus. Abbreviations: ce, compound eye; cl, clypeus; ff, frontal fovea; gf, gular fovea; la, labrum; ma, mandible; p 2 - 4, maxillary palpomere 2 - 4; vf, vertexal fovea. Scale bars: 0.5 mm in A, B, 0.2 mm in E-I." figureDoi="10.3897/asp.80.e82644.figure3" httpUri="https://binary.pensoft.net/fig/717125" pageId="0" pageNumber="279">3H, I</figureCitation>
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), four-segmented, with palpomere 1 small, indistinct, 2 (Fig.
|
|
<figureCitation id="701E1FF9F2DBAED55C269F3AF0367A33" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Morphology of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102 (A, C, D, I: under incident light; B, E-H: X-ray microtomographic reconstruction). A, B Dorsal habitus. C Left half of pronotum, showing squamous setae. D Lateral portion of the abdominal base, showing squamous setae. E Head dorsum. F Head venter. G Head, in antero-lateral view. H, I Left maxillary palpus. Abbreviations: ce, compound eye; cl, clypeus; ff, frontal fovea; gf, gular fovea; la, labrum; ma, mandible; p 2 - 4, maxillary palpomere 2 - 4; vf, vertexal fovea. Scale bars: 0.5 mm in A, B, 0.2 mm in E-I." figureDoi="10.3897/asp.80.e82644.figure3" httpUri="https://binary.pensoft.net/fig/717125" pageId="0" pageNumber="279">3H</figureCitation>
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;
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<emphasis id="1450955EFF94568FEEAFC25AD950C4A8" italics="true" pageId="0" pageNumber="279">p2</emphasis>
|
|
) pedunculate basally and broadened toward apex, 3 (Fig.
|
|
<figureCitation id="7D1611117D87F7A8A3E185AE6A6F286C" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Morphology of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102 (A, C, D, I: under incident light; B, E-H: X-ray microtomographic reconstruction). A, B Dorsal habitus. C Left half of pronotum, showing squamous setae. D Lateral portion of the abdominal base, showing squamous setae. E Head dorsum. F Head venter. G Head, in antero-lateral view. H, I Left maxillary palpus. Abbreviations: ce, compound eye; cl, clypeus; ff, frontal fovea; gf, gular fovea; la, labrum; ma, mandible; p 2 - 4, maxillary palpomere 2 - 4; vf, vertexal fovea. Scale bars: 0.5 mm in A, B, 0.2 mm in E-I." figureDoi="10.3897/asp.80.e82644.figure3" httpUri="https://binary.pensoft.net/fig/717125" pageId="0" pageNumber="279">3H</figureCitation>
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;
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<emphasis id="68E805653761FD341D810FE6CBA03FAF" italics="true" pageId="0" pageNumber="279">p3</emphasis>
|
|
) roundly triangular, 4 (Fig.
|
|
<figureCitation id="3C3D85C8E8C5C11DD25898BC6A1B30E9" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Morphology of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102 (A, C, D, I: under incident light; B, E-H: X-ray microtomographic reconstruction). A, B Dorsal habitus. C Left half of pronotum, showing squamous setae. D Lateral portion of the abdominal base, showing squamous setae. E Head dorsum. F Head venter. G Head, in antero-lateral view. H, I Left maxillary palpus. Abbreviations: ce, compound eye; cl, clypeus; ff, frontal fovea; gf, gular fovea; la, labrum; ma, mandible; p 2 - 4, maxillary palpomere 2 - 4; vf, vertexal fovea. Scale bars: 0.5 mm in A, B, 0.2 mm in E-I." figureDoi="10.3897/asp.80.e82644.figure3" httpUri="https://binary.pensoft.net/fig/717125" pageId="0" pageNumber="279">3H</figureCitation>
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;
|
|
<emphasis id="38A80451E18D3A22CFF39706A92167AF" italics="true" pageId="0" pageNumber="279">p4</emphasis>
|
|
) sub-fusiform, with short palpal cone at apex. Venter with small, widely separated gular foveae (= posterior tentorial pits; Fig.
|
|
<figureCitation id="3372EDA5F28ECA7A22097C3446E2DFEC" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Morphology of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102 (A, C, D, I: under incident light; B, E-H: X-ray microtomographic reconstruction). A, B Dorsal habitus. C Left half of pronotum, showing squamous setae. D Lateral portion of the abdominal base, showing squamous setae. E Head dorsum. F Head venter. G Head, in antero-lateral view. H, I Left maxillary palpus. Abbreviations: ce, compound eye; cl, clypeus; ff, frontal fovea; gf, gular fovea; la, labrum; ma, mandible; p 2 - 4, maxillary palpomere 2 - 4; vf, vertexal fovea. Scale bars: 0.5 mm in A, B, 0.2 mm in E-I." figureDoi="10.3897/asp.80.e82644.figure3" httpUri="https://binary.pensoft.net/fig/717125" pageId="0" pageNumber="279">3F</figureCitation>
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;
|
|
<emphasis id="2DE2AAFF8B2ABEC7FA58E15256369C65" italics="true" pageId="0" pageNumber="279">gf</emphasis>
|
|
) in shared transverse impression, gular moderately raised along middle, weakly impressed admesally; neck region broad. Paired tentorial arms (Fig.
|
|
<figureCitation id="8C94EC8ADC2191438305C7BC2F4D6989" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Morphology of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102 (A, C, D, I: under incident light; B, E-H: X-ray microtomographic reconstruction). A, B Dorsal habitus. C Left half of pronotum, showing squamous setae. D Lateral portion of the abdominal base, showing squamous setae. E Head dorsum. F Head venter. G Head, in antero-lateral view. H, I Left maxillary palpus. Abbreviations: ce, compound eye; cl, clypeus; ff, frontal fovea; gf, gular fovea; la, labrum; ma, mandible; p 2 - 4, maxillary palpomere 2 - 4; vf, vertexal fovea. Scale bars: 0.5 mm in A, B, 0.2 mm in E-I." figureDoi="10.3897/asp.80.e82644.figure3" httpUri="https://binary.pensoft.net/fig/717125" pageId="0" pageNumber="279">3E</figureCitation>
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;
|
|
<emphasis id="2E9380732958D16EBCC8D60722B25DEC" italics="true" pageId="0" pageNumber="279">ta</emphasis>
|
|
) V-shaped, each branched at base and extending anteriorly to reach inner wall of clypeus (right arm broken at apex in holotype). Pharynx (Fig.
|
|
<figureCitation id="501665473198A49AE96647F20B37A1F7" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Morphology of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102 (A, C, D, I: under incident light; B, E-H: X-ray microtomographic reconstruction). A, B Dorsal habitus. C Left half of pronotum, showing squamous setae. D Lateral portion of the abdominal base, showing squamous setae. E Head dorsum. F Head venter. G Head, in antero-lateral view. H, I Left maxillary palpus. Abbreviations: ce, compound eye; cl, clypeus; ff, frontal fovea; gf, gular fovea; la, labrum; ma, mandible; p 2 - 4, maxillary palpomere 2 - 4; vf, vertexal fovea. Scale bars: 0.5 mm in A, B, 0.2 mm in E-I." figureDoi="10.3897/asp.80.e82644.figure3" httpUri="https://binary.pensoft.net/fig/717125" pageId="0" pageNumber="279">3E</figureCitation>
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;
|
|
<emphasis id="F8133888B87A300ECA539E4E149CE89A" italics="true" pageId="0" pageNumber="279">ph</emphasis>
|
|
) about 2/3 of head length, posteriorly broadened. Antenna 11-segmented, moderately elongate, extending to approximately half of elytral length when extending posteriorly, antennomeres each slightly elongate to sub-moniliform, club (Fig.
|
|
<figureCitation id="AB089C9CCBFEE1B66A78FA272E21AB27" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Morphology of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102 (A, C, D, I: under incident light; B, E-H: X-ray microtomographic reconstruction). A, B Dorsal habitus. C Left half of pronotum, showing squamous setae. D Lateral portion of the abdominal base, showing squamous setae. E Head dorsum. F Head venter. G Head, in antero-lateral view. H, I Left maxillary palpus. Abbreviations: ce, compound eye; cl, clypeus; ff, frontal fovea; gf, gular fovea; la, labrum; ma, mandible; p 2 - 4, maxillary palpomere 2 - 4; vf, vertexal fovea. Scale bars: 0.5 mm in A, B, 0.2 mm in E-I." figureDoi="10.3897/asp.80.e82644.figure3" httpUri="https://binary.pensoft.net/fig/717125" pageId="0" pageNumber="279">3F</figureCitation>
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|
) formed by enlarged apical three antennomeres.
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|
</paragraph>
|
|
<caption id="058F5596DB0AE2473DA1B3BF764E709A" doi="10.3897/asp.80.e82644.figure2" httpUri="https://binary.pensoft.net/fig/717124" pageId="0" pageNumber="279" start="Figure 2" startId="F2">
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|
<paragraph id="40B3B375547F7890416A74B2A942AA8D" pageId="0" pageNumber="279">
|
|
<emphasis id="C4543F4C37437FA8AA7570B149A48C0B" bold="true" pageId="0" pageNumber="279">Figure 2.</emphasis>
|
|
Amber piece (A) containing
|
|
<taxonomicName id="E0F05C4749FD3F2511A59F73EF3643A1" authorityName="Yin & Cai" authorityYear="2022" class="Insecta" family="Staphylinidae" genus="Europharinodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Europharinodes schaufussi" order="Coleoptera" pageId="0" pageNumber="279" phylum="Arthropoda" rank="species" species="schaufussi">
|
|
<emphasis id="EED50857036425F2F5EDA63BB5B80A7F" italics="true" pageId="0" pageNumber="279">Europharinodes schaufussi</emphasis>
|
|
</taxonomicName>
|
|
<emphasis id="33370E78691290B5FAE0B105C759A946" bold="true" pageId="0" pageNumber="279">gen. et sp. nov.</emphasis>
|
|
, holotype, SNUC-Paleo-0102, and its morphological details (B-G) of (A, B, G: under incident light; C-F: X-ray microtomographic reconstruction).
|
|
<emphasis id="8A68BBB82738CAC0DF5E0E8B58BC69AC" bold="true" pageId="0" pageNumber="279">A</emphasis>
|
|
Original amber piece containing the holotype before cutting and polishing.
|
|
<emphasis id="8B888DE62D0A5817BC024208BE5293A1" bold="true" pageId="0" pageNumber="279">B</emphasis>
|
|
,
|
|
<emphasis id="01E0DD0A9D10E2493A475FFDD51F1A5E" bold="true" pageId="0" pageNumber="279">C</emphasis>
|
|
Left lateral habitus.
|
|
<emphasis id="EB10157D3434232C1B549DE21CC84677" bold="true" pageId="0" pageNumber="279">D</emphasis>
|
|
Right lateral habitus.
|
|
<emphasis id="E04FFA1F11FB5A583E1DE3D7B35C3039" bold="true" pageId="0" pageNumber="279">E</emphasis>
|
|
Head in lateral view, show tentorial arms and pharynx.
|
|
<emphasis id="83FBFDFD105FE4A454828F60A3A37C54" bold="true" pageId="0" pageNumber="279">F</emphasis>
|
|
Right antennal club.
|
|
<emphasis id="26943A554B684A5D43291850AFCD08BA" bold="true" pageId="0" pageNumber="279">G</emphasis>
|
|
Pro- and metatarsus and respective pretarsal claws. Abbreviations: aed, aedeagus; hw, hind wing; ph, pharynx; ta, tentorial arm. Scale bars: 0.5 mm in B-D, 0.2 mm in E, F.
|
|
</paragraph>
|
|
</caption>
|
|
<caption id="89E405F58F706ABE13959190C29AB76C" doi="10.3897/asp.80.e82644.figure3" httpUri="https://binary.pensoft.net/fig/717125" pageId="0" pageNumber="279" start="Figure 3" startId="F3">
|
|
<paragraph id="5199800A861E3D291720A70463A65FAE" pageId="0" pageNumber="279">
|
|
<emphasis id="667C3FB1A71EBDF299292384572DFCE3" bold="true" pageId="0" pageNumber="279">Figure 3.</emphasis>
|
|
Morphology of
|
|
<taxonomicName id="3805FA4B69082E538A363EF12DC56371" authorityName="Yin & Cai" authorityYear="2022" class="Insecta" family="Staphylinidae" genus="Europharinodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Europharinodes schaufussi" order="Coleoptera" pageId="0" pageNumber="279" phylum="Arthropoda" rank="species" species="schaufussi">
|
|
<emphasis id="0BFD7AC527D7A3C28FA80A61C2C4E869" italics="true" pageId="0" pageNumber="279">Europharinodes schaufussi</emphasis>
|
|
</taxonomicName>
|
|
<emphasis id="70266B2B6B984F94DF31704E7B0A0E97" bold="true" pageId="0" pageNumber="279">gen. et sp. nov.</emphasis>
|
|
, holotype, SNUC-Paleo-0102 (A, C, D, I: under incident light; B, E-H: X-ray microtomographic reconstruction).
|
|
<emphasis id="F0322053B3C5485A97F17B0716867489" bold="true" pageId="0" pageNumber="279">A</emphasis>
|
|
,
|
|
<emphasis id="DB9FA565E0CC781CD2A27880F965F7B8" bold="true" pageId="0" pageNumber="279">B</emphasis>
|
|
Dorsal habitus.
|
|
<emphasis id="580CD7787E2A9FA9AE8BC3ECF784C293" bold="true" pageId="0" pageNumber="279">C</emphasis>
|
|
Left half of pronotum, showing squamous setae.
|
|
<emphasis id="FDCA8476B698D0416D9304D5C6FF5B53" bold="true" pageId="0" pageNumber="279">D</emphasis>
|
|
Lateral portion of the abdominal base, showing squamous setae.
|
|
<emphasis id="D44AFD82527DF2270359E6FE9D313E4E" bold="true" pageId="0" pageNumber="279">E</emphasis>
|
|
Head dorsum.
|
|
<emphasis id="F7956661F0A939F1CD5AC2C50D3AE433" bold="true" pageId="0" pageNumber="279">F</emphasis>
|
|
Head venter.
|
|
<emphasis id="BAA13862883988F15CAC34C0ECB745BB" bold="true" pageId="0" pageNumber="279">G</emphasis>
|
|
Head, in antero-lateral view.
|
|
<emphasis id="EDBDE1CE67EE1181010FF9993C77CEA3" bold="true" pageId="0" pageNumber="279">H</emphasis>
|
|
,
|
|
<emphasis id="EB77E4E5746C3C0F34734168345059FE" bold="true" pageId="0" pageNumber="279">I</emphasis>
|
|
Left maxillary palpus. Abbreviations: ce, compound eye; cl, clypeus; ff, frontal fovea; gf, gular fovea; la, labrum; ma, mandible; p2-4, maxillary palpomere 2-4; vf, vertexal fovea. Scale bars: 0.5 mm in A, B, 0.2 mm in E-I.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph id="31AC5BD75C1CB6311D2914189A6C423F" pageId="0" pageNumber="279">
|
|
Pronotum (Fig.
|
|
<figureCitation id="979F515BD4A9C91D1C544E837FA5C773" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. X-ray microtomographic reconstruction of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102. A - C: prothorax, dorsal (A), lateral (B) and ventral (C). D Right elytron. E, F Meso- and metathorax, ventral (E) and lateral (F). G - I Abdomen, dorsal (G), lateral (H) and ventral (I). Abbreviations: ahf, antero-hypomeral fovea; bef, basal elytral fovea; blf, basal lateral fovea; ds, discal stria; laf, lateral antebasal fovea; lmcf, lateral metaventral fovea; lpcf, lateral procoxal fovea; maf, median antebasal fovea; mmsf, median mesoventral fovea; mc, marginal carina; mmtf, median metaventral fovea; mn, median nodule; ms, median split; pt 1 - 3, paratergite 1 - 3; s 2 - 6, sternite 2 - 6 (IV-VIII); t 1 - 5, tergite 1 - 5 (IV-VIII). Scale bars: 0.2 mm." figureDoi="10.3897/asp.80.e82644.figure4" httpUri="https://binary.pensoft.net/fig/717126" pageId="0" pageNumber="279">4A</figureCitation>
|
|
) approximately as long as broad, sides rounded, broadest at middle, anteriorly and posteriorly narrowed, surface with dense squamous setae (Fig.
|
|
<figureCitation id="A554EFBDBB539144C560366053D00E97" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Morphology of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102 (A, C, D, I: under incident light; B, E-H: X-ray microtomographic reconstruction). A, B Dorsal habitus. C Left half of pronotum, showing squamous setae. D Lateral portion of the abdominal base, showing squamous setae. E Head dorsum. F Head venter. G Head, in antero-lateral view. H, I Left maxillary palpus. Abbreviations: ce, compound eye; cl, clypeus; ff, frontal fovea; gf, gular fovea; la, labrum; ma, mandible; p 2 - 4, maxillary palpomere 2 - 4; vf, vertexal fovea. Scale bars: 0.5 mm in A, B, 0.2 mm in E-I." figureDoi="10.3897/asp.80.e82644.figure3" httpUri="https://binary.pensoft.net/fig/717125" pageId="0" pageNumber="279">3C</figureCitation>
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|
); disc coarsely punctate, with squamous setae all over surface, covering median and lateral antebasal foveae (Fig.
|
|
<figureCitation id="2B8622971E947504AA87A1360607C50D" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. X-ray microtomographic reconstruction of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102. A - C: prothorax, dorsal (A), lateral (B) and ventral (C). D Right elytron. E, F Meso- and metathorax, ventral (E) and lateral (F). G - I Abdomen, dorsal (G), lateral (H) and ventral (I). Abbreviations: ahf, antero-hypomeral fovea; bef, basal elytral fovea; blf, basal lateral fovea; ds, discal stria; laf, lateral antebasal fovea; lmcf, lateral metaventral fovea; lpcf, lateral procoxal fovea; maf, median antebasal fovea; mmsf, median mesoventral fovea; mc, marginal carina; mmtf, median metaventral fovea; mn, median nodule; ms, median split; pt 1 - 3, paratergite 1 - 3; s 2 - 6, sternite 2 - 6 (IV-VIII); t 1 - 5, tergite 1 - 5 (IV-VIII). Scale bars: 0.2 mm." figureDoi="10.3897/asp.80.e82644.figure4" httpUri="https://binary.pensoft.net/fig/717126" pageId="0" pageNumber="279">4A, B</figureCitation>
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;
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<emphasis id="FEF2E97DCCCD449421B830DA8E5A0387" italics="true" pageId="0" pageNumber="279">maf</emphasis>
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,
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<emphasis id="A4D7445B00F9C1B084593632CF7463F8" italics="true" pageId="0" pageNumber="279">laf</emphasis>
|
|
), lacking sulcus or carina. Hypomeron extended, lacking carina, with anterior hypomeral fovea (Fig.
|
|
<figureCitation id="D1D2544C2BA1E9A773C4615738135E5D" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. X-ray microtomographic reconstruction of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102. A - C: prothorax, dorsal (A), lateral (B) and ventral (C). D Right elytron. E, F Meso- and metathorax, ventral (E) and lateral (F). G - I Abdomen, dorsal (G), lateral (H) and ventral (I). Abbreviations: ahf, antero-hypomeral fovea; bef, basal elytral fovea; blf, basal lateral fovea; ds, discal stria; laf, lateral antebasal fovea; lmcf, lateral metaventral fovea; lpcf, lateral procoxal fovea; maf, median antebasal fovea; mmsf, median mesoventral fovea; mc, marginal carina; mmtf, median metaventral fovea; mn, median nodule; ms, median split; pt 1 - 3, paratergite 1 - 3; s 2 - 6, sternite 2 - 6 (IV-VIII); t 1 - 5, tergite 1 - 5 (IV-VIII). Scale bars: 0.2 mm." figureDoi="10.3897/asp.80.e82644.figure4" httpUri="https://binary.pensoft.net/fig/717126" pageId="0" pageNumber="279">4B, C</figureCitation>
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;
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<emphasis id="D7930624A37F40C95B6D8438ED73E877" italics="true" pageId="0" pageNumber="279">ahf</emphasis>
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). Prosternum (Fig.
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<figureCitation id="80FA5DF83EA18A56C11C535490FA04BC" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. X-ray microtomographic reconstruction of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102. A - C: prothorax, dorsal (A), lateral (B) and ventral (C). D Right elytron. E, F Meso- and metathorax, ventral (E) and lateral (F). G - I Abdomen, dorsal (G), lateral (H) and ventral (I). Abbreviations: ahf, antero-hypomeral fovea; bef, basal elytral fovea; blf, basal lateral fovea; ds, discal stria; laf, lateral antebasal fovea; lmcf, lateral metaventral fovea; lpcf, lateral procoxal fovea; maf, median antebasal fovea; mmsf, median mesoventral fovea; mc, marginal carina; mmtf, median metaventral fovea; mn, median nodule; ms, median split; pt 1 - 3, paratergite 1 - 3; s 2 - 6, sternite 2 - 6 (IV-VIII); t 1 - 5, tergite 1 - 5 (IV-VIII). Scale bars: 0.2 mm." figureDoi="10.3897/asp.80.e82644.figure4" httpUri="https://binary.pensoft.net/fig/717126" pageId="0" pageNumber="279">4C</figureCitation>
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) with widely separated lateral procoxal foveae (Fig.
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<figureCitation id="679C455C71EACC77F2A73E4FE4C96517" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. X-ray microtomographic reconstruction of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102. A - C: prothorax, dorsal (A), lateral (B) and ventral (C). D Right elytron. E, F Meso- and metathorax, ventral (E) and lateral (F). G - I Abdomen, dorsal (G), lateral (H) and ventral (I). Abbreviations: ahf, antero-hypomeral fovea; bef, basal elytral fovea; blf, basal lateral fovea; ds, discal stria; laf, lateral antebasal fovea; lmcf, lateral metaventral fovea; lpcf, lateral procoxal fovea; maf, median antebasal fovea; mmsf, median mesoventral fovea; mc, marginal carina; mmtf, median metaventral fovea; mn, median nodule; ms, median split; pt 1 - 3, paratergite 1 - 3; s 2 - 6, sternite 2 - 6 (IV-VIII); t 1 - 5, tergite 1 - 5 (IV-VIII). Scale bars: 0.2 mm." figureDoi="10.3897/asp.80.e82644.figure4" httpUri="https://binary.pensoft.net/fig/717126" pageId="0" pageNumber="279">4C</figureCitation>
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;
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<emphasis id="80D190289A453BE348083056AC06864D" italics="true" pageId="0" pageNumber="279">lpcf</emphasis>
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), anterior part shorter than coxal part, margins of coxal cavity moderately carinate.
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</paragraph>
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<caption id="8CBE2083F9B185A9B35C517A646E1E25" doi="10.3897/asp.80.e82644.figure4" httpUri="https://binary.pensoft.net/fig/717126" pageId="0" pageNumber="279" start="Figure 4" startId="F4">
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<paragraph id="7BB4C5EE7FF3AF227FD4469D24FDB111" pageId="0" pageNumber="279">
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<emphasis id="6EF01A3DD2B7833BFF0342DCAAAB1303" bold="true" pageId="0" pageNumber="279">Figure 4.</emphasis>
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X-ray microtomographic reconstruction of
|
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<taxonomicName id="05A1BB881CB0D4074C8E837B29B938DD" authorityName="Yin & Cai" authorityYear="2022" class="Insecta" family="Staphylinidae" genus="Europharinodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Europharinodes schaufussi" order="Coleoptera" pageId="0" pageNumber="279" phylum="Arthropoda" rank="species" species="schaufussi">
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<emphasis id="B5ABD8C0BBC3FC3F4290831D844E2D00" italics="true" pageId="0" pageNumber="279">Europharinodes schaufussi</emphasis>
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</taxonomicName>
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<emphasis id="FBE5AB6C49289F063762C69B36832160" bold="true" pageId="0" pageNumber="279">gen. et sp. nov.</emphasis>
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, holotype, SNUC-Paleo-0102.
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<emphasis id="7A62791FF52760D618DADE4C53BE815B" bold="true" pageId="0" pageNumber="279">A</emphasis>
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-
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<emphasis id="EB06885FE7D17E4AAA8AE47899DC9F0A" bold="true" pageId="0" pageNumber="279">C</emphasis>
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: prothorax, dorsal (A), lateral (B) and ventral (C).
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<emphasis id="1FB2DED8EF9910E9DE1627E7E80AA9E1" bold="true" pageId="0" pageNumber="279">D</emphasis>
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Right elytron.
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<emphasis id="7698998C7686A6742A32212DFD4B15BD" bold="true" pageId="0" pageNumber="279">E</emphasis>
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,
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<emphasis id="C8F337D9A335BE8371ACB56FA383BEDA" bold="true" pageId="0" pageNumber="279">F</emphasis>
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Meso- and metathorax, ventral (E) and lateral (F).
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<emphasis id="FCD8AFB737E569CD971F015A056CE426" bold="true" pageId="0" pageNumber="279">G</emphasis>
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-
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<emphasis id="390510B5E1986E8A80CAEA08C709E087" bold="true" pageId="0" pageNumber="279">I</emphasis>
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Abdomen, dorsal (G), lateral (H) and ventral (I). Abbreviations: ahf, antero-hypomeral fovea; bef, basal elytral fovea; blf, basal lateral fovea; ds, discal stria; laf, lateral antebasal fovea; lmcf, lateral metaventral fovea; lpcf, lateral procoxal fovea; maf, median antebasal fovea; mmsf, median mesoventral fovea; mc, marginal carina; mmtf, median metaventral fovea; mn, median nodule; ms, median split; pt1-3, paratergite 1-3; s2-6, sternite 2-6 (IV-VIII); t1-5, tergite 1-5 (IV-VIII). Scale bars: 0.2 mm.
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</paragraph>
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</caption>
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<paragraph id="ECDDCDAA2E7191AB7767E7E011784BD9" pageId="0" pageNumber="279">
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|
Elytra broadly truncate at bases, each elytron with two large basal foveae (Fig.
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|
<figureCitation id="39F21F364FE028D099FA3DB14434B4CC" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. X-ray microtomographic reconstruction of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102. A - C: prothorax, dorsal (A), lateral (B) and ventral (C). D Right elytron. E, F Meso- and metathorax, ventral (E) and lateral (F). G - I Abdomen, dorsal (G), lateral (H) and ventral (I). Abbreviations: ahf, antero-hypomeral fovea; bef, basal elytral fovea; blf, basal lateral fovea; ds, discal stria; laf, lateral antebasal fovea; lmcf, lateral metaventral fovea; lpcf, lateral procoxal fovea; maf, median antebasal fovea; mmsf, median mesoventral fovea; mc, marginal carina; mmtf, median metaventral fovea; mn, median nodule; ms, median split; pt 1 - 3, paratergite 1 - 3; s 2 - 6, sternite 2 - 6 (IV-VIII); t 1 - 5, tergite 1 - 5 (IV-VIII). Scale bars: 0.2 mm." figureDoi="10.3897/asp.80.e82644.figure4" httpUri="https://binary.pensoft.net/fig/717126" pageId="0" pageNumber="279">4D</figureCitation>
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;
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<emphasis id="CFD653742235E53397F21482672E74DC" italics="true" pageId="0" pageNumber="279">bef</emphasis>
|
|
), with two long discal striae (Fig.
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|
<figureCitation id="F605497CB0BBDBFB0219C2E2CA4A3805" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. X-ray microtomographic reconstruction of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102. A - C: prothorax, dorsal (A), lateral (B) and ventral (C). D Right elytron. E, F Meso- and metathorax, ventral (E) and lateral (F). G - I Abdomen, dorsal (G), lateral (H) and ventral (I). Abbreviations: ahf, antero-hypomeral fovea; bef, basal elytral fovea; blf, basal lateral fovea; ds, discal stria; laf, lateral antebasal fovea; lmcf, lateral metaventral fovea; lpcf, lateral procoxal fovea; maf, median antebasal fovea; mmsf, median mesoventral fovea; mc, marginal carina; mmtf, median metaventral fovea; mn, median nodule; ms, median split; pt 1 - 3, paratergite 1 - 3; s 2 - 6, sternite 2 - 6 (IV-VIII); t 1 - 5, tergite 1 - 5 (IV-VIII). Scale bars: 0.2 mm." figureDoi="10.3897/asp.80.e82644.figure4" httpUri="https://binary.pensoft.net/fig/717126" pageId="0" pageNumber="279">4G</figureCitation>
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;
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<emphasis id="5334C99E92772CA54A29DCB6781CB1FF" italics="true" pageId="0" pageNumber="279">ds</emphasis>
|
|
) extending from foveae to posterior elytral margin, lacking subhumeral fovea or marginal sulcus. Metathoracic (hind) wings (Fig.
|
|
<figureCitation id="FC4AE574090A4A75F80486C7430FA317" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Amber piece (A) containing Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102, and its morphological details (B-G) of (A, B, G: under incident light; C-F: X-ray microtomographic reconstruction). A Original amber piece containing the holotype before cutting and polishing. B, C Left lateral habitus. D Right lateral habitus. E Head in lateral view, show tentorial arms and pharynx. F Right antennal club. G Pro- and metatarsus and respective pretarsal claws. Abbreviations: aed, aedeagus; hw, hind wing; ph, pharynx; ta, tentorial arm. Scale bars: 0.5 mm in B-D, 0.2 mm in E, F." figureDoi="10.3897/asp.80.e82644.figure2" httpUri="https://binary.pensoft.net/fig/717124" pageId="0" pageNumber="279">2D</figureCitation>
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;
|
|
<emphasis id="78CD0D30C086DA2AB2E0EF933D31BEF0" italics="true" pageId="0" pageNumber="279">hw</emphasis>
|
|
) fully developed.
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|
</paragraph>
|
|
<paragraph id="B59211BE6C0E49577BF5BD225C643FCA" pageId="0" pageNumber="279">
|
|
Mesoventrite with median foveae (Fig.
|
|
<figureCitation id="51F1DDAD4096C124DF6BB5E5E5A9F114" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. X-ray microtomographic reconstruction of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102. A - C: prothorax, dorsal (A), lateral (B) and ventral (C). D Right elytron. E, F Meso- and metathorax, ventral (E) and lateral (F). G - I Abdomen, dorsal (G), lateral (H) and ventral (I). Abbreviations: ahf, antero-hypomeral fovea; bef, basal elytral fovea; blf, basal lateral fovea; ds, discal stria; laf, lateral antebasal fovea; lmcf, lateral metaventral fovea; lpcf, lateral procoxal fovea; maf, median antebasal fovea; mmsf, median mesoventral fovea; mc, marginal carina; mmtf, median metaventral fovea; mn, median nodule; ms, median split; pt 1 - 3, paratergite 1 - 3; s 2 - 6, sternite 2 - 6 (IV-VIII); t 1 - 5, tergite 1 - 5 (IV-VIII). Scale bars: 0.2 mm." figureDoi="10.3897/asp.80.e82644.figure4" httpUri="https://binary.pensoft.net/fig/717126" pageId="0" pageNumber="279">4E</figureCitation>
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;
|
|
<emphasis id="4EDA48EF7C34DA5DEF9DE8D4FBBD4466" italics="true" pageId="0" pageNumber="279">mmsf</emphasis>
|
|
) originating from shared transverse impression, moderately separated, with pairs of large lateral and distinct anterolateral mesoventral foveae (not shown in figure); metaventrite with large, setose lateral coxal foveae (Fig.
|
|
<figureCitation id="EA720DE4BDB2850495F31F65564E942A" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. X-ray microtomographic reconstruction of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102. A - C: prothorax, dorsal (A), lateral (B) and ventral (C). D Right elytron. E, F Meso- and metathorax, ventral (E) and lateral (F). G - I Abdomen, dorsal (G), lateral (H) and ventral (I). Abbreviations: ahf, antero-hypomeral fovea; bef, basal elytral fovea; blf, basal lateral fovea; ds, discal stria; laf, lateral antebasal fovea; lmcf, lateral metaventral fovea; lpcf, lateral procoxal fovea; maf, median antebasal fovea; mmsf, median mesoventral fovea; mc, marginal carina; mmtf, median metaventral fovea; mn, median nodule; ms, median split; pt 1 - 3, paratergite 1 - 3; s 2 - 6, sternite 2 - 6 (IV-VIII); t 1 - 5, tergite 1 - 5 (IV-VIII). Scale bars: 0.2 mm." figureDoi="10.3897/asp.80.e82644.figure4" httpUri="https://binary.pensoft.net/fig/717126" pageId="0" pageNumber="279">4F</figureCitation>
|
|
;
|
|
<emphasis id="5E34903CED71E9E023E87203B86A0F93" italics="true" pageId="0" pageNumber="279">lmcf</emphasis>
|
|
), with single median metaventral fovea (Fig.
|
|
<figureCitation id="48A5D8ADDDBFA8A615F36E183606EBA8" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. X-ray microtomographic reconstruction of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102. A - C: prothorax, dorsal (A), lateral (B) and ventral (C). D Right elytron. E, F Meso- and metathorax, ventral (E) and lateral (F). G - I Abdomen, dorsal (G), lateral (H) and ventral (I). Abbreviations: ahf, antero-hypomeral fovea; bef, basal elytral fovea; blf, basal lateral fovea; ds, discal stria; laf, lateral antebasal fovea; lmcf, lateral metaventral fovea; lpcf, lateral procoxal fovea; maf, median antebasal fovea; mmsf, median mesoventral fovea; mc, marginal carina; mmtf, median metaventral fovea; mn, median nodule; ms, median split; pt 1 - 3, paratergite 1 - 3; s 2 - 6, sternite 2 - 6 (IV-VIII); t 1 - 5, tergite 1 - 5 (IV-VIII). Scale bars: 0.2 mm." figureDoi="10.3897/asp.80.e82644.figure4" httpUri="https://binary.pensoft.net/fig/717126" pageId="0" pageNumber="279">4E</figureCitation>
|
|
;
|
|
<emphasis id="C71EE2869630B9CFA9795AEBB9B05D7D" italics="true" pageId="0" pageNumber="279">mmtf</emphasis>
|
|
), posterior margin with small, narrow split at middle (Fig.
|
|
<figureCitation id="A74D1A8B729C67293DF1D0CA84C191E7" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. X-ray microtomographic reconstruction of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102. A - C: prothorax, dorsal (A), lateral (B) and ventral (C). D Right elytron. E, F Meso- and metathorax, ventral (E) and lateral (F). G - I Abdomen, dorsal (G), lateral (H) and ventral (I). Abbreviations: ahf, antero-hypomeral fovea; bef, basal elytral fovea; blf, basal lateral fovea; ds, discal stria; laf, lateral antebasal fovea; lmcf, lateral metaventral fovea; lpcf, lateral procoxal fovea; maf, median antebasal fovea; mmsf, median mesoventral fovea; mc, marginal carina; mmtf, median metaventral fovea; mn, median nodule; ms, median split; pt 1 - 3, paratergite 1 - 3; s 2 - 6, sternite 2 - 6 (IV-VIII); t 1 - 5, tergite 1 - 5 (IV-VIII). Scale bars: 0.2 mm." figureDoi="10.3897/asp.80.e82644.figure4" httpUri="https://binary.pensoft.net/fig/717126" pageId="0" pageNumber="279">4E</figureCitation>
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;
|
|
<emphasis id="A34AB8CADBD742B178DF044F95BF4F3F" italics="true" pageId="0" pageNumber="279">ms</emphasis>
|
|
); metacoxae broadly separated. Marginal carina of meso- and metaventrite complete (Fig.
|
|
<figureCitation id="4E8671296548BF57CB65DDB7F9AC2E09" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. X-ray microtomographic reconstruction of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102. A - C: prothorax, dorsal (A), lateral (B) and ventral (C). D Right elytron. E, F Meso- and metathorax, ventral (E) and lateral (F). G - I Abdomen, dorsal (G), lateral (H) and ventral (I). Abbreviations: ahf, antero-hypomeral fovea; bef, basal elytral fovea; blf, basal lateral fovea; ds, discal stria; laf, lateral antebasal fovea; lmcf, lateral metaventral fovea; lpcf, lateral procoxal fovea; maf, median antebasal fovea; mmsf, median mesoventral fovea; mc, marginal carina; mmtf, median metaventral fovea; mn, median nodule; ms, median split; pt 1 - 3, paratergite 1 - 3; s 2 - 6, sternite 2 - 6 (IV-VIII); t 1 - 5, tergite 1 - 5 (IV-VIII). Scale bars: 0.2 mm." figureDoi="10.3897/asp.80.e82644.figure4" httpUri="https://binary.pensoft.net/fig/717126" pageId="0" pageNumber="279">4F</figureCitation>
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;
|
|
<emphasis id="34E0D0F987DF13A6477332900E745D8B" italics="true" pageId="0" pageNumber="279">mc</emphasis>
|
|
).
|
|
</paragraph>
|
|
<paragraph id="B09E81C6082B5A85A9DE3E4DA5C6B9AA" pageId="0" pageNumber="279">
|
|
Legs (Fig.
|
|
<figureCitation id="272F942E527F7A711DF6C465C8F48F9F" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Morphology of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102 (A-H: X-ray microtomographic reconstruction; I-K: schematic). A Right foreleg. B Right middle leg. C Left hindleg. D - F, I - K Aedeagus, dorsal (D, I), lateral (E, J) and ventral (F, K). G, H endophallus, dorsal (G) and lateral (H). Abbreviations: bc, basal capsule; bp, basoventral projection; dd, dorsal diaphragm; en, endophallus; ml, median lobe; pa, paramere. Scale bars: 0.3 mm in A-C; 0.1 mm in D-K." figureDoi="10.3897/asp.80.e82644.figure5" httpUri="https://binary.pensoft.net/fig/717127" pageId="0" pageNumber="279">5A-C</figureCitation>
|
|
) moderately elongate; femora roughly punctate; with short tarsomere 1 and long tarsomere 2 and 3, 2 slightly longer than 3, each tarsus with two subequal pretarsal claws (Fig.
|
|
<figureCitation id="AFDDE77EBD693800864E1D2097BECF53" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Amber piece (A) containing Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102, and its morphological details (B-G) of (A, B, G: under incident light; C-F: X-ray microtomographic reconstruction). A Original amber piece containing the holotype before cutting and polishing. B, C Left lateral habitus. D Right lateral habitus. E Head in lateral view, show tentorial arms and pharynx. F Right antennal club. G Pro- and metatarsus and respective pretarsal claws. Abbreviations: aed, aedeagus; hw, hind wing; ph, pharynx; ta, tentorial arm. Scale bars: 0.5 mm in B-D, 0.2 mm in E, F." figureDoi="10.3897/asp.80.e82644.figure2" httpUri="https://binary.pensoft.net/fig/717124" pageId="0" pageNumber="279">2G</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<caption id="324D8A791E4845FB1B9026F47E2FBAAF" doi="10.3897/asp.80.e82644.figure5" httpUri="https://binary.pensoft.net/fig/717127" pageId="0" pageNumber="279" start="Figure 5" startId="F5">
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|
<paragraph id="A2D8BD4EAFECFC8105D8CC3B5D148F79" pageId="0" pageNumber="279">
|
|
<emphasis id="FA672107AD2D6A02E23FFD3659A0744B" bold="true" pageId="0" pageNumber="279">Figure 5.</emphasis>
|
|
Morphology of
|
|
<taxonomicName id="F460B834E98313F2C9B39CD81A6C5C3E" authorityName="Yin & Cai" authorityYear="2022" class="Insecta" family="Staphylinidae" genus="Europharinodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Europharinodes schaufussi" order="Coleoptera" pageId="0" pageNumber="279" phylum="Arthropoda" rank="species" species="schaufussi">
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<emphasis id="686C60FF10B550FEED2A2FF285264386" italics="true" pageId="0" pageNumber="279">Europharinodes schaufussi</emphasis>
|
|
</taxonomicName>
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<emphasis id="1AD97C1A6C986A2F8A74794810AB9E93" bold="true" pageId="0" pageNumber="279">gen. et sp. nov.</emphasis>
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, holotype, SNUC-Paleo-0102 (A-H: X-ray microtomographic reconstruction; I-K: schematic).
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<emphasis id="A0157060C00BA6B59C142C4C42841E32" bold="true" pageId="0" pageNumber="279">A</emphasis>
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Right foreleg.
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<emphasis id="56205C012B998353C3CD03BFED12FA73" bold="true" pageId="0" pageNumber="279">B</emphasis>
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Right middle leg.
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<emphasis id="7DA38B068C9FFA7E4A43D6C366F8C8A1" bold="true" pageId="0" pageNumber="279">C</emphasis>
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Left hindleg.
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<emphasis id="051C9D7824CAFBE85DAC7901440FC872" bold="true" pageId="0" pageNumber="279">D</emphasis>
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-
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<emphasis id="078941F407E5D1E1925B78F482EF0560" bold="true" pageId="0" pageNumber="279">F</emphasis>
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,
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<emphasis id="868A0DAEDA6009DAC6D9B72FFA934066" bold="true" pageId="0" pageNumber="279">I</emphasis>
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-
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<emphasis id="03F278883F8FECA02413655B1C9DC3B5" bold="true" pageId="0" pageNumber="279">K</emphasis>
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Aedeagus, dorsal (D, I), lateral (E, J) and ventral (F, K).
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<emphasis id="71E2761B7BCFEEE5B96251FB5A252233" bold="true" pageId="0" pageNumber="279">G</emphasis>
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,
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<emphasis id="EBE69EDF5C1E2715A385442A9A208CD6" bold="true" pageId="0" pageNumber="279">H</emphasis>
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endophallus, dorsal (G) and lateral (H). Abbreviations: bc, basal capsule; bp, basoventral projection; dd, dorsal diaphragm; en, endophallus; ml, median lobe; pa, paramere. Scale bars: 0.3 mm in A-C; 0.1 mm in D-K.
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</paragraph>
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</caption>
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<paragraph id="CEB45EC769CCC1A9C4C390DC39BED826" pageId="0" pageNumber="279">
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Abdomen constricted at base, with dense squamous setae (Fig.
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<figureCitation id="0F7E28FACE50A165D3D2016A00608487" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Morphology of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102 (A, C, D, I: under incident light; B, E-H: X-ray microtomographic reconstruction). A, B Dorsal habitus. C Left half of pronotum, showing squamous setae. D Lateral portion of the abdominal base, showing squamous setae. E Head dorsum. F Head venter. G Head, in antero-lateral view. H, I Left maxillary palpus. Abbreviations: ce, compound eye; cl, clypeus; ff, frontal fovea; gf, gular fovea; la, labrum; ma, mandible; p 2 - 4, maxillary palpomere 2 - 4; vf, vertexal fovea. Scale bars: 0.5 mm in A, B, 0.2 mm in E-I." figureDoi="10.3897/asp.80.e82644.figure3" httpUri="https://binary.pensoft.net/fig/717125" pageId="0" pageNumber="279">3D</figureCitation>
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); tergite 1 (IV) slightly longer than 2 (V), 2-4 (V-VII) subequal in length, tergites 1-4 (Fig.
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<figureCitation id="26FD4287106FD539EE40995C49EF1C0E" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. X-ray microtomographic reconstruction of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102. A - C: prothorax, dorsal (A), lateral (B) and ventral (C). D Right elytron. E, F Meso- and metathorax, ventral (E) and lateral (F). G - I Abdomen, dorsal (G), lateral (H) and ventral (I). Abbreviations: ahf, antero-hypomeral fovea; bef, basal elytral fovea; blf, basal lateral fovea; ds, discal stria; laf, lateral antebasal fovea; lmcf, lateral metaventral fovea; lpcf, lateral procoxal fovea; maf, median antebasal fovea; mmsf, median mesoventral fovea; mc, marginal carina; mmtf, median metaventral fovea; mn, median nodule; ms, median split; pt 1 - 3, paratergite 1 - 3; s 2 - 6, sternite 2 - 6 (IV-VIII); t 1 - 5, tergite 1 - 5 (IV-VIII). Scale bars: 0.2 mm." figureDoi="10.3897/asp.80.e82644.figure4" httpUri="https://binary.pensoft.net/fig/717126" pageId="0" pageNumber="279">4G, H</figureCitation>
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;
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<emphasis id="7F9C503F013051B1EA71C7C36BC400F6" italics="true" pageId="0" pageNumber="279">t1-4</emphasis>
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) each deeply and broadly sulcate at base, at lease with one pair of basolateral foveae (Fig.
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<figureCitation id="1976760BC463682E5C74169701655818" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. X-ray microtomographic reconstruction of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102. A - C: prothorax, dorsal (A), lateral (B) and ventral (C). D Right elytron. E, F Meso- and metathorax, ventral (E) and lateral (F). G - I Abdomen, dorsal (G), lateral (H) and ventral (I). Abbreviations: ahf, antero-hypomeral fovea; bef, basal elytral fovea; blf, basal lateral fovea; ds, discal stria; laf, lateral antebasal fovea; lmcf, lateral metaventral fovea; lpcf, lateral procoxal fovea; maf, median antebasal fovea; mmsf, median mesoventral fovea; mc, marginal carina; mmtf, median metaventral fovea; mn, median nodule; ms, median split; pt 1 - 3, paratergite 1 - 3; s 2 - 6, sternite 2 - 6 (IV-VIII); t 1 - 5, tergite 1 - 5 (IV-VIII). Scale bars: 0.2 mm." figureDoi="10.3897/asp.80.e82644.figure4" httpUri="https://binary.pensoft.net/fig/717126" pageId="0" pageNumber="279">4G</figureCitation>
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;
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<emphasis id="02098066DDA8CA4FA03E78E55FBE12D7" italics="true" pageId="0" pageNumber="279">blf</emphasis>
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|
), tergite 5 (VIII) (Fig.
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|
<figureCitation id="50CB42DF902E84454313EED0E6B53C76" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. X-ray microtomographic reconstruction of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102. A - C: prothorax, dorsal (A), lateral (B) and ventral (C). D Right elytron. E, F Meso- and metathorax, ventral (E) and lateral (F). G - I Abdomen, dorsal (G), lateral (H) and ventral (I). Abbreviations: ahf, antero-hypomeral fovea; bef, basal elytral fovea; blf, basal lateral fovea; ds, discal stria; laf, lateral antebasal fovea; lmcf, lateral metaventral fovea; lpcf, lateral procoxal fovea; maf, median antebasal fovea; mmsf, median mesoventral fovea; mc, marginal carina; mmtf, median metaventral fovea; mn, median nodule; ms, median split; pt 1 - 3, paratergite 1 - 3; s 2 - 6, sternite 2 - 6 (IV-VIII); t 1 - 5, tergite 1 - 5 (IV-VIII). Scale bars: 0.2 mm." figureDoi="10.3897/asp.80.e82644.figure4" httpUri="https://binary.pensoft.net/fig/717126" pageId="0" pageNumber="279">4H</figureCitation>
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;
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|
<emphasis id="950D10C2ACB93D04D52E54964A2196CE" italics="true" pageId="0" pageNumber="279">t5</emphasis>
|
|
) roundly triangular, transverse, medially roundly emarginate at posterior margin; sternite 2-5 (IV-VII) (Fig.
|
|
<figureCitation id="04CFF7491E88377983E0D837348C42C4" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. X-ray microtomographic reconstruction of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102. A - C: prothorax, dorsal (A), lateral (B) and ventral (C). D Right elytron. E, F Meso- and metathorax, ventral (E) and lateral (F). G - I Abdomen, dorsal (G), lateral (H) and ventral (I). Abbreviations: ahf, antero-hypomeral fovea; bef, basal elytral fovea; blf, basal lateral fovea; ds, discal stria; laf, lateral antebasal fovea; lmcf, lateral metaventral fovea; lpcf, lateral procoxal fovea; maf, median antebasal fovea; mmsf, median mesoventral fovea; mc, marginal carina; mmtf, median metaventral fovea; mn, median nodule; ms, median split; pt 1 - 3, paratergite 1 - 3; s 2 - 6, sternite 2 - 6 (IV-VIII); t 1 - 5, tergite 1 - 5 (IV-VIII). Scale bars: 0.2 mm." figureDoi="10.3897/asp.80.e82644.figure4" httpUri="https://binary.pensoft.net/fig/717126" pageId="0" pageNumber="279">4H, I</figureCitation>
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;
|
|
<emphasis id="477B2AF8253B0446DBC8BF64D759A912" italics="true" pageId="0" pageNumber="279">s2-5</emphasis>
|
|
) each broadly sulcate at base, with one pair of basolateral foveae (Fig.
|
|
<figureCitation id="6AA1EC0A2ECB8FA7B9D8DDCB1521182F" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Morphology of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102 (A, C, D, I: under incident light; B, E-H: X-ray microtomographic reconstruction). A, B Dorsal habitus. C Left half of pronotum, showing squamous setae. D Lateral portion of the abdominal base, showing squamous setae. E Head dorsum. F Head venter. G Head, in antero-lateral view. H, I Left maxillary palpus. Abbreviations: ce, compound eye; cl, clypeus; ff, frontal fovea; gf, gular fovea; la, labrum; ma, mandible; p 2 - 4, maxillary palpomere 2 - 4; vf, vertexal fovea. Scale bars: 0.5 mm in A, B, 0.2 mm in E-I." figureDoi="10.3897/asp.80.e82644.figure3" httpUri="https://binary.pensoft.net/fig/717125" pageId="0" pageNumber="279">3H</figureCitation>
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;
|
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<emphasis id="6A30000CB2353B7DC14259FBF46A9462" italics="true" pageId="0" pageNumber="279">blf</emphasis>
|
|
), 6 (VIII), sternite 6 (Fig.
|
|
<figureCitation id="B8C5C107407201B0A38E2560F2082769" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. X-ray microtomographic reconstruction of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102. A - C: prothorax, dorsal (A), lateral (B) and ventral (C). D Right elytron. E, F Meso- and metathorax, ventral (E) and lateral (F). G - I Abdomen, dorsal (G), lateral (H) and ventral (I). Abbreviations: ahf, antero-hypomeral fovea; bef, basal elytral fovea; blf, basal lateral fovea; ds, discal stria; laf, lateral antebasal fovea; lmcf, lateral metaventral fovea; lpcf, lateral procoxal fovea; maf, median antebasal fovea; mmsf, median mesoventral fovea; mc, marginal carina; mmtf, median metaventral fovea; mn, median nodule; ms, median split; pt 1 - 3, paratergite 1 - 3; s 2 - 6, sternite 2 - 6 (IV-VIII); t 1 - 5, tergite 1 - 5 (IV-VIII). Scale bars: 0.2 mm." figureDoi="10.3897/asp.80.e82644.figure4" httpUri="https://binary.pensoft.net/fig/717126" pageId="0" pageNumber="279">4H, I</figureCitation>
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|
;
|
|
<emphasis id="6007ACE13ECA2F4E275FDA1E9BCD952B" italics="true" pageId="0" pageNumber="279">s6</emphasis>
|
|
) transverse.
|
|
</paragraph>
|
|
<paragraph id="A43351D362B3252A21C187CD9556BB58" pageId="0" pageNumber="279">
|
|
Males have modified sternites 4 and 5 (VI and VII). Aedeagus (Fig.
|
|
<figureCitation id="91416F1314B8AEE284F7564ACA1B0A05" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Morphology of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102 (A-H: X-ray microtomographic reconstruction; I-K: schematic). A Right foreleg. B Right middle leg. C Left hindleg. D - F, I - K Aedeagus, dorsal (D, I), lateral (E, J) and ventral (F, K). G, H endophallus, dorsal (G) and lateral (H). Abbreviations: bc, basal capsule; bp, basoventral projection; dd, dorsal diaphragm; en, endophallus; ml, median lobe; pa, paramere. Scale bars: 0.3 mm in A-C; 0.1 mm in D-K." figureDoi="10.3897/asp.80.e82644.figure5" httpUri="https://binary.pensoft.net/fig/717127" pageId="0" pageNumber="279">5D-F, I-K</figureCitation>
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|
) dorso-ventrally symmetrical.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="7B56231E5525ED833F21CC3D1914738F" pageId="0" pageNumber="279" type="etymology">
|
|
<paragraph id="475D0B389A6C8739E23AF4F1378FEA67" pageId="0" pageNumber="279">Etymology.</paragraph>
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<paragraph id="44034A0B6681C9BDF64697A2E20F0386" pageId="0" pageNumber="279">
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|
The new generic name is a combination of Latin '
|
|
<emphasis id="766B8586FDAAB17C65A33066B3579CA7" italics="true" pageId="0" pageNumber="279">
|
|
<normalizedToken id="F54C8132E599649DC5079A8805CD7DF5" originalValue="Eurōpa">Eurōpa</normalizedToken>
|
|
</emphasis>
|
|
(Europe)' and genus
|
|
<taxonomicName id="B8CC8DDB17C7FD5546A020613DC28436" authorityName="Raffray" authorityYear="1890" class="Insecta" family="Staphylinidae" genus="Apharinodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Apharinodes" order="Coleoptera" pageId="0" pageNumber="279" phylum="Arthropoda" rank="genus">
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<emphasis id="815F0E61BE71539BAAE3614C22EF7599" italics="true" pageId="0" pageNumber="279">Apharinodes</emphasis>
|
|
</taxonomicName>
|
|
, referring to the origin of the fossil in Baltic amber and its affinity with
|
|
<taxonomicName id="7C88BD396DE63D0D4E4F9017D1711055" authorityName="Raffray" authorityYear="1890" class="Insecta" family="Staphylinidae" genus="Apharinodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Apharinodes" order="Coleoptera" pageId="0" pageNumber="279" phylum="Arthropoda" rank="genus">
|
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<emphasis id="026D402C187C404CFAF07A343E3BE128" italics="true" pageId="0" pageNumber="279">Apharinodes</emphasis>
|
|
</taxonomicName>
|
|
. The gender is feminine.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
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</document> |