239 lines
20 KiB
XML
239 lines
20 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.763.21252" ID-GBIF-Dataset="6162ab6f-6b50-40ee-b28e-c72eee01e99e" ID-PMC="PMC5996013" ID-Pensoft-Pub="1313-2970-763-1" ID-PubMed="29896045" ID-ZBK="90B772554C5E436CA793D924892B5B14" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1313-2970-763-1" ModsDocOrigin="ZooKeys 763" ModsDocTitle="Integrative taxonomy of a new and highly-diverse genus of onchidiid slugs from the Coral Triangle (Gastropoda, Pulmonata, Onchidiidae)" checkinTime="1528130281812" checkinUser="pensoft" docAuthor="Goulding, Tricia C., Khalil, Munawar, Tan, Shau Hwai & Dayrat, Benoit" docDate="2018" docId="DF9DB794E7B0C4BB9CA15740C33BEECF" docLanguage="en" docName="ZooKeys 763: 1-111" docOrigin="ZooKeys 763" docSource="http://dx.doi.org/10.3897/zookeys.763.21252" docTitle="Wallaconchis comendadori Goulding & Dayrat, sp. n." docType="treatment" docUuid="F96331E4-4A09-425E-B103-62811C10816D" docUuidSource="ZooBank" docVersion="5" lastPageNumber="1" masterDocId="FFF78434FFEAFFF5FFBBF264B1300C6B" masterDocTitle="Integrative taxonomy of a new and highly-diverse genus of onchidiid slugs from the Coral Triangle (Gastropoda, Pulmonata, Onchidiidae)" masterLastPageNumber="111" masterPageNumber="1" pageNumber="1" updateTime="1668165860460" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Integrative taxonomy of a new and highly-diverse genus of onchidiid slugs from the Coral Triangle (Gastropoda, Pulmonata, Onchidiidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Goulding, Tricia C.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Khalil, Munawar</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Tan, Shau Hwai</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Dayrat, Benoit</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2018</mods:date>
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<mods:detail type="volume">
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<mods:number>763</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>111</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.763.21252</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.763.21252</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-763-1</mods:identifier>
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<mods:identifier type="ZBK">90B772554C5E436CA793D924892B5B14</mods:identifier>
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<mods:identifier type="ZooBank">90B772554C5E436CA793D924892B5B14</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="144234916" LSID="urn:lsid:zoobank.org:act:F96331E4-4A09-425E-B103-62811C10816D" httpUri="http://treatment.plazi.org/id/DF9DB794E7B0C4BB9CA15740C33BEECF" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName LSID="http://zoobank.org/F96331E4-4A09-425E-B103-62811C10816D" authority="Goulding & Dayrat" class="Gastropoda" family="Onchidiidae" genus="Wallaconchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wallaconchis comendadori" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="comendadori">Wallaconchis comendadori Goulding & Dayrat</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="1">sp. n.</taxonomicNameLabel>
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Figs 54, 55, 56, 57, 58
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type locality">
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<paragraph pageId="0" pageNumber="1">Type locality.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Philippines, Bohol, Guindulman,
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<geoCoordinate direction="north" orientation="latitude" precision="9" value="9.734333">09°44.06'N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="9" value="124.4605">124°27.63'E</geoCoordinate>
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, st 197, rocks and coral rubble near a few
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<taxonomicName class="Magnoliopsida" family="Acanthaceae" genus="Avicennia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Avicennia" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Avicennia</taxonomicName>
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trees.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type material">
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<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="1">Holotype: 12/11 mm [3626], designated here (PNM 041232).</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="materials_examined">
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<paragraph pageId="0" pageNumber="1">Additional material examined.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Indonesia, Sulawesi, Mantehage Island,
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<geoCoordinate direction="north" orientation="latitude" precision="9" value="1.698">01°41.88'N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="9" value="124.779">124°46.74'E</geoCoordinate>
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, 1 specimen 15/7 mm [2315], st 91, rocks behind a mangrove of
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<taxonomicName class="Cephalopoda" family="Hoplitidae" genus="Sonneratia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sonneratia" order="Ammonoidea" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Sonneratia</taxonomicName>
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and
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<taxonomicName class="Magnoliopsida" family="Rhizophoraceae" genus="Rhizophora" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Rhizophora" order="Malpighiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Rhizophora</taxonomicName>
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(UMIZ 00060); Ambon, Haruku Island,
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<geoCoordinate direction="south" orientation="latitude" precision="9" value="-3.6086667">03°36.52'S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="9" value="128.41783">128°25.07'E</geoCoordinate>
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, 2 specimens 12/7 mm [2725] and 7/6 mm [3539], st 127, rocky
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<taxonomicName class="Cephalopoda" family="Hoplitidae" genus="Sonneratia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sonneratia" order="Ammonoidea" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Sonneratia</taxonomicName>
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mangrove with coral rubble (UMIZ 00061); Lombok, Don Don,
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<geoCoordinate direction="south" orientation="latitude" precision="9" value="-8.909">08°54.54'S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="9" value="116.35833">116°21.50'E</geoCoordinate>
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, 2 specimens 18/12 mm [2985] and 7/6 mm [2983], st 149, old
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<taxonomicName class="Magnoliopsida" family="Acanthaceae" genus="Avicennia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Avicennia" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Avicennia</taxonomicName>
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forest with coral rubble (UMIZ 00063); Bali, Pemuteran, Labuhan Lalang Harbor,
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<geoCoordinate direction="south" orientation="latitude" precision="9" value="-8.1435">08°08.61'S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="9" value="114.53883">114°32.33'E</geoCoordinate>
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, 2 specimens 16/8 mm [3133] and 13/9 mm [3131], st 157, coral rubble, rocks and a few
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<taxonomicName class="Magnoliopsida" family="Acanthaceae" genus="Avicennia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Avicennia" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Avicennia</taxonomicName>
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(UMIZ 00064). Philippines, Bohol, Guindulman,
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<geoCoordinate direction="north" orientation="latitude" precision="9" value="9.734333">09°44.06'N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="9" value="124.4605">124°27.63'E</geoCoordinate>
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, 2 specimens 30/11 mm [3627] and 19/15 mm [3630], st 197, rocks and coral rubble near a few
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<taxonomicName class="Magnoliopsida" family="Acanthaceae" genus="Avicennia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Avicennia" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Avicennia</taxonomicName>
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trees (PNM 041233); Bohol, Maribojoc,
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="9.733666">09°44.020'N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="9" value="123.79083">123°47.45'E</geoCoordinate>
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, 2 specimens 22/13 mm [3385] and 21/16 mm [3400], st 200, coral rubble with sand, low tide at night (PNM 041234).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="distribution">
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<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="1">Indonesia: Ambon, Bali, Lombok, and Sulawesi. Philippines: Bohol.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="habitat">
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<paragraph pageId="0" pageNumber="1">Habitat</paragraph>
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<paragraph pageId="0" pageNumber="1">
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(Fig. 54, Table 3).
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<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Wallaconchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wallaconchis comendadori" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="comendadori">Wallaconchis comendadori</taxonomicName>
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is found in the rocky intertidal, on rocks or large pieces of coral rubble, generally covered by a thin mat of algae.
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</paragraph>
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<caption pageId="0" pageNumber="1">
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<paragraph pageId="0" pageNumber="1">
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Figure 54. Habitats,
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<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Wallaconchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wallaconchis comendadori" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="comendadori">Wallaconchis comendadori</taxonomicName>
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. A Type locality, Philippines, Bohol, Guindulman, rocks and coral rubble near a few
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<taxonomicName class="Magnoliopsida" family="Acanthaceae" genus="Avicennia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Avicennia" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Avicennia</taxonomicName>
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trees (st 197) B View of the shore (same locality as A) C Indonesia, Bali, Pemuteran, Labuhan Lalang Harbor, coral rubble, rocks and a few
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<taxonomicName class="Magnoliopsida" family="Acanthaceae" genus="Avicennia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Avicennia" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Avicennia</taxonomicName>
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trees (st 157) D Indonesia, Lombok, Don Don, old
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<taxonomicName class="Magnoliopsida" family="Acanthaceae" genus="Avicennia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Avicennia" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Avicennia</taxonomicName>
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forest, with coral rubble (st 149).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="etymology">
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<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="1">This species is dedicated to Joseph Comendador, from the National Museum of the Philippines. Our expedition in the Philippines would not have been possible without his help with logistics, and we enjoyed exploring mangroves with him.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="diagnosis">
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<paragraph pageId="0" pageNumber="1">Diagnosis</paragraph>
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<paragraph pageId="0" pageNumber="1">
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(Table 5). Externally,
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<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Wallaconchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wallaconchis comendadori" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="comendadori">Wallaconchis comendadori</taxonomicName>
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cannot be distinguished from brown or black specimens of other
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<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Wallaconchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wallaconchis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Wallaconchis</taxonomicName>
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species, although a bright yellow, longitudinal band on the dorsum is more commonly found in this species. Internally, the narrow penis, a free oviduct (not attached to the body wall by fibers), and the apple-shaped spermatheca distinguishes
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<taxonomicName lsidName="W. comendadori" pageId="0" pageNumber="1" rank="species" species="comendadori">W. comendadori</taxonomicName>
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from most
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<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Wallaconchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wallaconchis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Wallaconchis</taxonomicName>
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species, except
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<taxonomicName lsidName="W. melanesiensis" pageId="0" pageNumber="1" rank="species" species="melanesiensis">W. melanesiensis</taxonomicName>
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, from which it differs by having a longer penis and a more highly convoluted deferent duct.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="color and morphology of live animals">
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<paragraph pageId="0" pageNumber="1">Color and morphology of live animals</paragraph>
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<paragraph pageId="0" pageNumber="1">(Fig. 55). Animals are not frequently brightly colored. The dorsal notum is predominantly brown or black, which may be mixed with subtle patches of yellow, green, red, and grey. A median, longitudinal yellow band was observed in a few individuals. The ocular tentacles are brown or grey. The hyponotum is light or bright yellow. The foot is bright yellow or light yellow-grey.</paragraph>
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<caption pageId="0" pageNumber="1">
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<paragraph pageId="0" pageNumber="1">
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Figure 55. Live specimens,
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<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Wallaconchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wallaconchis comendadori" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="comendadori">Wallaconchis comendadori</taxonomicName>
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. A Dorsal view, 16 mm long [3133], Indonesia, Bali (UMIZ 00064) B Dorsal view, 18 mm long [2985], Indonesia, Lombok (UMIZ 00063) C Dorsal view, 13 mm long [3131], Indonesia, Bali (UMIZ 00064) D Dorsal view, 12 mm long [2725], Indonesia, Ambon (UMIZ 00061) E Dorsal view, 7 mm long [2983], Indonesia, Lombok (UMIZ 00063) F Dorsal view, holotype, 12 mm long [3626], Philippines, Bohol (PNM 041232) G Ventral view, same as B H Ventral view, 15 mm long [2315], Indonesia, Sulawesi (UMIZ 00060) I Ventral view, 22 mm long [3385], Philippines, Bohol (PNM 041234).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="external morphology">
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<paragraph pageId="0" pageNumber="1">External morphology.</paragraph>
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<paragraph pageId="0" pageNumber="1">Between five and seven papillae bear eyes (with three or four eyes per papilla) but more may be retracted. There is a retractable papilla with eyes in the center of the dorsal notum, which is not raised above the other papillae.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="digestive system">
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<paragraph pageId="0" pageNumber="1">Digestive system</paragraph>
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<paragraph pageId="0" pageNumber="1">
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(Fig. 56, Table 4). Examples of radular formulae are presented in Table 4. The length of the rachidian teeth is approximately 15-22
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<normalizedToken originalValue="µm">µm</normalizedToken>
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, significantly smaller than that of the lateral teeth. The length of the lateral teeth (from 30 to 60
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<normalizedToken originalValue="µm">µm</normalizedToken>
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) gradually increases from the inner to the outer teeth (excluding the innermost and outermost lateral teeth which are significantly smaller). The intestinal loops are of type I.
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</paragraph>
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<caption pageId="0" pageNumber="1">
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<paragraph pageId="0" pageNumber="1">
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Figure 56. Radula,
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<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Wallaconchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wallaconchis comendadori" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="comendadori">Wallaconchis comendadori</taxonomicName>
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. A Rachidian and innermost lateral teeth, Philippines, Bohol, scale bar 20
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<normalizedToken originalValue="μm">μm</normalizedToken>
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[3385] (PNM 041234) B Rachidian and innermost lateral teeth, Indonesia, Lombok, scale bar 20
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<normalizedToken originalValue="μm">μm</normalizedToken>
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[2985] (UMIZ 00063) C Rachidian and inner lateral teeth, scale bar 30
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<normalizedToken originalValue="μm">μm</normalizedToken>
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, same as B D Transition between inner lateral teeth and outer lateral teeth, arrow shows basal lateral spine, scale bar 30
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<normalizedToken originalValue="μm">μm</normalizedToken>
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, same as B E Transition between inner lateral teeth and outer lateral teeth, scale bar 30
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<normalizedToken originalValue="μm">μm</normalizedToken>
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, same as B F Outermost lateral teeth, Indonesia, Ambon, scale bar 20
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<normalizedToken originalValue="μm">μm</normalizedToken>
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[3539] (UMIZ 00061).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="reproductive system">
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<paragraph pageId="0" pageNumber="1">Reproductive system</paragraph>
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<paragraph pageId="0" pageNumber="1">(Fig. 57A). The oviduct is narrow, approximately the same width as the deferent duct. The spermatheca is apple-shaped, with two lobes, and joins the oviduct through a short duct.</paragraph>
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<caption pageId="0" pageNumber="1">
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<paragraph pageId="0" pageNumber="1">
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Figure 57. Reproductive system,
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<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Wallaconchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wallaconchis comendadori" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="comendadori">Wallaconchis comendadori</taxonomicName>
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. A Hermaphroditic (female), posterior parts, Indonesia, Lombok, scale bar 3 mm [2985] (UMIZ 00063) B Anterior, copulatory parts, Philippines, Bohol, scale bar 2 mm [3385] (PNM 041234). Abbreviations: dd deferent duct fgm female gland mass hd hermaphroditic duct hg hermaphroditic gland ov oviduct ps penial sheath rm retractor muscle rs receptaculum seminis sp spermatheca.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="copulatory apparatus">
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<paragraph pageId="0" pageNumber="1">Copulatory apparatus</paragraph>
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<paragraph pageId="0" pageNumber="1">
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(Figs 57B, 58). The penis (more than 1.5 mm long) is narrow (approximately 100
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<normalizedToken originalValue="µm">µm</normalizedToken>
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) and smooth with no hooks (Fig. 58). The penial sheath is narrow proximally, and gradually widens distally into a vestibule. The penis is within the narrow region of the vestibule. The deferent duct is extremely convoluted (Fig. 57B), but slightly less convoluted in immature specimens. The length of the penial sheath equals approximately two thirds to three quarters the length of the body cavity. The deferent duct is approximately the same width as the penial vestibule. The retractor muscle is narrow and inserts at the posterior end of the body cavity, near the rectum.
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</paragraph>
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<caption pageId="0" pageNumber="1">
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<paragraph pageId="0" pageNumber="1">
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Figure 58. Penis,
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<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Wallaconchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wallaconchis comendadori" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="comendadori">Wallaconchis comendadori</taxonomicName>
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, Philippines, Bohol [3385] (PNM 041234). A View of entire penis, scale bar 0.5 mm B Distal tip of penis, scale bar 50
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<normalizedToken originalValue="μm">μm</normalizedToken>
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.
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</paragraph>
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</caption>
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</subSubSection>
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</treatment>
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</document> |