133 lines
12 KiB
XML
133 lines
12 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.863.34662" ID-GBIF-Dataset="af839023-d05c-4769-b2a1-f9f9f8561df7" ID-PMC="PMC6639351" ID-Pensoft-Pub="1313-2970-863-35" ID-PubMed="31341392" ID-ZBK="763EDE2B5F0C414D8289D37765E993E4" ModsDocAuthor="" ModsDocDate="2019" ModsDocID="1313-2970-863-35" ModsDocOrigin="ZooKeys 863" ModsDocTitle="Unexpected palaeodiversity of omaliine rove beetles in Eocene Baltic amber (Coleoptera, Staphylinidae, Omaliinae)" checkinTime="1562893176132" checkinUser="pensoft" docAuthor="Shavrin, Alexey V. & Yamamoto, Shuhei" docDate="2019" docId="DEBCE552E5B763507BEF3A47A634F59F" docLanguage="en" docName="ZooKeys 863: 35-83" docOrigin="ZooKeys 863" docSource="http://dx.doi.org/10.3897/zookeys.863.34662" docTitle="Eusphalerum sp. 2." docType="treatment" docVersion="4" lastPageNumber="35" masterDocId="FFE7E644384D885DFFF3E730B1230775" masterDocTitle="Unexpected palaeodiversity of omaliine rove beetles in Eocene Baltic amber (Coleoptera, Staphylinidae, Omaliinae)" masterLastPageNumber="83" masterPageNumber="35" pageNumber="35" updateTime="1668167561321" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Unexpected palaeodiversity of omaliine rove beetles in Eocene Baltic amber (Coleoptera, Staphylinidae, Omaliinae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Shavrin, Alexey V.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Yamamoto, Shuhei</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2019</mods:date>
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<mods:detail type="volume">
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<mods:number>863</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>35</mods:start>
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<mods:end>83</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.863.34662</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.863.34662</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-863-35</mods:identifier>
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<mods:identifier type="ZBK">763EDE2B5F0C414D8289D37765E993E4</mods:identifier>
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<mods:identifier type="ZooBank">763EDE2B5F0C414D8289D37765E993E4</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="158605282" LSID="urn:lsid:plazi:treatment:DEBCE552E5B763507BEF3A47A634F59F" httpUri="http://treatment.plazi.org/id/DEBCE552E5B763507BEF3A47A634F59F" lastPageNumber="35" pageId="0" pageNumber="35">
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<subSubSection pageId="0" pageNumber="35" type="nomenclature">
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<paragraph pageId="0" pageNumber="35">
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<taxonomicName authority="sp. 2." class="Insecta" family="Staphylinidae" genus="Eusphalerum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eusphalerum" order="Coleoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="genus">Eusphalerum sp. 2.</taxonomicName>
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Figures 12, 13, 81-82, 83-85
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="35" type="materials_examined">
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<paragraph pageId="0" pageNumber="35">Materials examined.</paragraph>
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<paragraph pageId="0" pageNumber="35">
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One male, complete specimen as an inclusion in a piece of yellow Baltic amber 35.4 mm
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<normalizedToken originalValue="×">x</normalizedToken>
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21.5 mm
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<normalizedToken originalValue="×">x</normalizedToken>
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7.5 mm in size (Figs 12, 13), with glued very small quadrate paper on plastic bag labeled "AWI | 045", with the following labels: "AWI-045 | Phyllodrepa (?) | 3 spec." <rectangular handwritten label>, "Dr. Vitalii Alekseev's | Collection" <rectangular handwritten label>, "
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<taxonomicName class="Insecta" family="Staphylinidae" genus="Eusphalerum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eusphalerum" order="Coleoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="genus">Eusphalerum</taxonomicName>
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sp. 2 | Shavrin A.V. det. 2018" <rectangular label, printed>. The specimen is deposited in the private collection of Vitalii I. Alekseev (Kaliningrad, Russia), registered as AWI-045.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="35" type="preservation">
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<paragraph pageId="0" pageNumber="35">Preservation.</paragraph>
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<paragraph pageId="0" pageNumber="35">
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The single specimen is a male located close to the outer surface of the piece of amber, with many details visible in both dorsal and ventral surfaces. The elytra are somewhat deformed and seem flattened, and the right elytron is depressed into the thorax. Additionally, the piece of the amber contains two males of
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<taxonomicName genus="Eu." lsidName="Eu." pageId="0" pageNumber="35" rank="genus">Eu.</taxonomicName>
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sp. 3 and
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<taxonomicName genus="Eu." lsidName="Eu." pageId="0" pageNumber="35" rank="genus">Eu.</taxonomicName>
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sp. 4 (see below), and a syninclusion located near the narrowest side of the amber: nymph of small mite about 0.50 mm in length (Figs 12, 13).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="35" type="description">
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<paragraph pageId="0" pageNumber="35">Description.</paragraph>
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<paragraph pageId="0" pageNumber="35">
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Measurements: HW: 0.53; HL: 0.20; OL: 0.11; AL: 0.69; PML
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<normalizedToken originalValue="×">x</normalizedToken>
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PMW (III, IV): III: 0.03
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<normalizedToken originalValue="×">x</normalizedToken>
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0.01, IV: 0.07
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<normalizedToken originalValue="×">x</normalizedToken>
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0.02; PL: 0.43; PW: 0.67; ESL: 0.85; EW: 0.81; MTbL: 0.36; MTrL: 0.27 (
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<normalizedToken originalValue="I–IV">I-IV</normalizedToken>
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: 0.14; V: 0.13); AW: 0.79; TL: 2.06. Antennomeres with lengths
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<normalizedToken originalValue="×">x</normalizedToken>
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widths: 1: 0.12
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<normalizedToken originalValue="×">x</normalizedToken>
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0.02; 2: 0.07
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<normalizedToken originalValue="×">x</normalizedToken>
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0.02; 3: 0.06
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<normalizedToken originalValue="×">x</normalizedToken>
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0.02; 4: 0.05
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<normalizedToken originalValue="×">x</normalizedToken>
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0.02; 5-6: 0.05
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<normalizedToken originalValue="×">x</normalizedToken>
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0.03; 7: 0.04
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<normalizedToken originalValue="×">x</normalizedToken>
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0.04; 8: 0.05
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<normalizedToken originalValue="×">x</normalizedToken>
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0.04; 9-10: 0.05
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<normalizedToken originalValue="×">x</normalizedToken>
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0.05; 11: 0.10
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<normalizedToken originalValue="×">x</normalizedToken>
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0.04.
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</paragraph>
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<paragraph pageId="0" pageNumber="35">Body elongate, somewhat flattened (Fig. 81), glossy and glabrous, without visible setation. Body appears dark-brown, with basal portions of pronotum and legs reddish-brown. Body dorsolaterally as in Figure 82 and ventrally as in Figure 83.</paragraph>
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<caption pageId="0" pageNumber="35">
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<paragraph pageId="0" pageNumber="35">
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Figures 81, 82. Habitus of
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<taxonomicName class="Insecta" family="Staphylinidae" genus="Eusphalerum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eusphalerum" order="Coleoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="genus">Eusphalerum</taxonomicName>
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sp. 2 81 oblique dorsal view 82 dorsolateral view. Scale bars: 1.0 mm.
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</paragraph>
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</caption>
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<paragraph pageId="0" pageNumber="35">Head strongly transverse, distinctly more than twice as wide as long, with slightly convex middle portion and posterior parts of infraorbital ridges, without visible grooves in front of ocelli and postocular carina (Fig. 84). Head with indistinct, small and sparse punctation, with moderately coarse and large isodiametric microsculpture between punctures becoming more transverse toward middle part of neck. Eyes large, strongly protruding laterad. Ocelli small, convex, situated very close to infraorbital ridges at level of about middle length of eyes, distance between ocelli about twice as long as distance between ocellus and margin of eye (Figs 82, 84). Labrum transverse. Apical segment of maxillary palp elongate, slightly wider in middle than penultimate segment, from apical third gradually narrowed toward moderately acute apex (Figs 82, 84). Gular sutures not fully visible, widely separated from each other (Fig. 83). Antenna moderately long, distinctly exceeding shoulders of elytra, with elongate setae, distinctly longer on antennomeres 6-11; basal antennomere wide, gradually widened apicad, antennomere 2 similar in width, swollen and elongate, 3 with thin basal portion, slightly widened apicad, 4 slightly shorter than 3, 5, and 6 slightly wider than 4, 7 short and moderately rounded, 8-10 slightly transverse, apical antennomere more than twice as long as broad, from about middle strongly narrowed toward acute apex (Figs 81-84).</paragraph>
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<paragraph pageId="0" pageNumber="35">Pronotum 1.5 times as wide as long, slightly broader than head, widest in middle, markedly more narrowed posterad than anterad; apical margin slightly and widely rounded, about as broad as posterior margin, anterior and posterior angles widely rounded; laterobasal margins slightly concaved, with very indistinct small crenulation; lateral margins narrowly explanate; lateral portions with indistinct semioval impression about middle (Figs 81, 84). Pronotum with somewhat regular small and sparse punctation and with isodiametric ground sculpture slightly coarser than that on head (Figs 81, 84). Prosternum with moderately wide and protruded prosternal process (Fig. 83). Scutellum large and wide (Fig. 81).</paragraph>
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<paragraph pageId="0" pageNumber="35">Elytra little longer than wide, about twice as long as pronotum, gradually widened apicad, reaching basal to apical margins of abdominal tergite IV, with widely rounded apicolateral angles; shoulders moderately widely rounded; lateral edges narrowly explanate (Fig. 81). Punctation of elytra invisible in details but appears slightly denser and deeper than that on pronotum.</paragraph>
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<paragraph pageId="0" pageNumber="35">Legs moderately long and slender, femora markedly widened in middle, tibiae moderately short and thin, gradually widened apicad, slightly shorter than femora, covered by elongate setae, with a few strong setae on apical margins near apex; tarsomeres 1-4 distinctly wide, with dense and long setae; apical metatarsomere long, yet slightly shorter than length of preceding tarsomeres together; tarsal claws simple, elongate (Figs 81-83).</paragraph>
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<paragraph pageId="0" pageNumber="35">Abdomen (Fig. 85) slightly narrower than elytra; abdominal tergites with sparse and moderately small punctures, no wing-folding patches are visible.</paragraph>
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<paragraph pageId="0" pageNumber="35">Male. Apical margin of abdominal tergite VIII slightly rounded. Apical margin of abdominal sternite VIII slightly sinuate.</paragraph>
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<paragraph pageId="0" pageNumber="35">Female unknown.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="35" type="remarks">
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<paragraph pageId="0" pageNumber="35">Remarks.</paragraph>
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<paragraph pageId="0" pageNumber="35">As in the previous species, this specimen has very long and moderately dense setae on lateral portions of tarsomeres 1-4, distinctly deformed body (especially elytra) and unusually strongly protruded eyes.</paragraph>
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<caption pageId="0" pageNumber="35">
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<paragraph pageId="0" pageNumber="35">
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Figures 83-85.
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<taxonomicName class="Insecta" family="Staphylinidae" genus="Eusphalerum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eusphalerum" order="Coleoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="genus">Eusphalerum</taxonomicName>
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sp. 2 83 habitus, ventral view 84 left antenna, head and pronotum, dorsal view 85 abdomen, dorsal view. Scale bars: 1.0 mm (83), 0.4 mm (84, 85).
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</paragraph>
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</caption>
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</subSubSection>
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</treatment>
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</document> |