192 lines
15 KiB
XML
192 lines
15 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1146.97517" ID-Pensoft-Pub="1313-2970-1146-43" ID-Pensoft-UUID="B1CCD44D55C25FFD8D9DE752139E864F" ID-ZooBank="01E76F6AB9914F339BD6991090F07E80" ModsDocID="1313-2970-1146-43" checkinTime="1675846382090" checkinUser="pensoft" docAuthor="Zamani, Alireza, Marusik, Yuri M. & Szűts, Tamas" docDate="2023" docId="AFD3A047BC9C5FB69377ABA45FFAB955" docLanguage="en" docName="ZooKeys 1146: 43-86" docOrigin="ZooKeys 1146" docPubDate="2023-02-07" docSource="http://dx.doi.org/10.3897/zookeys.1146.97517" docTitle="Dysdera verkana Zamani & Marusik & Szűts 2023, sp. nov." docType="treatment" docUuid="2E18F484-6E0E-4296-B253-285A0E98E268" docUuidSource="ZooBank" docVersion="1" id="B1CCD44D55C25FFD8D9DE752139E864F" lastPageNumber="43" masterDocId="B1CCD44D55C25FFD8D9DE752139E864F" masterDocTitle="A survey of the spider genus Dysdera Latreille, 1804 (Araneae, Dysderidae) in Iran, with fourteen new species and notes on two fossil genera" masterLastPageNumber="86" masterPageNumber="43" pageNumber="43" updateTime="1675846382090" updateUser="pensoft">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A survey of the spider genus Dysdera Latreille, 1804 (Araneae, Dysderidae) in Iran, with fourteen new species and notes on two fossil genera</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Zamani, Alireza</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-8084-9666</mods:nameIdentifier>
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<mods:affiliation>Zoological Museum, Biodiversity Unit, FI- 20014 University of Turku, Turku, Finland</mods:affiliation>
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<mods:nameIdentifier type="email">zamani.alireza5@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Marusik, Yuri M.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-4499-5148</mods:nameIdentifier>
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<mods:affiliation>Department of Zoology & Entomology, University of the Free State, Bloemfontein 9300, South Africa</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Szűts, Tamas</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-8954-0641</mods:nameIdentifier>
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<mods:affiliation>Department of Ecology, University of Veterinary Medicine Budapest, Rottenbiller u. 50, Budapest, 1077, Hungary</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-02-07</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1146</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>43</mods:start>
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<mods:end>86</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1146.97517</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1146.97517</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1146-43</mods:identifier>
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<mods:identifier type="ZooBank">01E76F6AB9914F339BD6991090F07E80</mods:identifier>
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<mods:identifier type="Pensoft-UUID">B1CCD44D55C25FFD8D9DE752139E864F</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:zoobank.org:act:2E18F484-6E0E-4296-B253-285A0E98E268" httpUri="http://treatment.plazi.org/id/AFD3A047BC9C5FB69377ABA45FFAB955" lastPageNumber="43" pageId="0" pageNumber="43">
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<subSubSection pageId="0" pageNumber="43" type="nomenclature">
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<paragraph pageId="0" pageNumber="43">
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<taxonomicName LSID="https://zoobank.org/2E18F484-6E0E-4296-B253-285A0E98E268" authority="Zamani & Marusik & Szűts, 2023" authorityName="Zamani & Marusik & Szűts" authorityYear="2023" class="Arachnida" family="Dysderidae" genus="Dysdera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dysdera verkana" order="Araneae" pageId="0" pageNumber="43" phylum="Arthropoda" rank="species" species="verkana" status="sp. nov.">Dysdera verkana</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="43">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="43" type="description">
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<paragraph pageId="0" pageNumber="43">
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<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Male of Dysdera verkana sp. nov., habitus A dorsal view B ventral view C lateral view. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1146.97517.figure21" httpUri="https://binary.pensoft.net/fig/806632" pageId="0" pageNumber="43">Figs 21A-C</figureCitation>
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<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Male of Dysdera verkana sp. nov., bulb A retrolateral view B prolateral view C anterior view D posterior view. Scale bars: 0.25 mm." figureDoi="10.3897/zookeys.1146.97517.figure22" httpUri="https://binary.pensoft.net/fig/806633" pageId="0" pageNumber="43">, 22A-D</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="43" type="type material">
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<paragraph pageId="0" pageNumber="43">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="43">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="43">Holotype</emphasis>
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♂ (ZMUT), Iran: Golestan Province: Azadshahr County, Khosh Yeylaq,
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<geoCoordinate degrees="36" direction="north" minutes="49" orientation="latitude" precision="925" value="36.816666">36°49'N</geoCoordinate>
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,
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<geoCoordinate degrees="55" direction="east" minutes="20" orientation="longitude" precision="925" value="55.333332">55°20'E</geoCoordinate>
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, 15.06.2016 (D. Kasatkin).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="43" type="etymology">
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<paragraph pageId="0" pageNumber="43">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="43">The specific epithet is a noun in apposition, referring to an Old Persian word for the Gorgan region, meaning "land of wolves".</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="43" type="diagnosis">
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<paragraph pageId="0" pageNumber="43">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="43">
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The male of the new species is most similar to that of
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<taxonomicName lsidName="D. sagartia" pageId="0" pageNumber="43" rank="species" species="sagartia">
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<emphasis italics="true" pageId="0" pageNumber="43">D. sagartia</emphasis>
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</taxonomicName>
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sp. nov., but differs by the more rounded median crest, the posterior apophysis not bent on right angle (cf. Fig.
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<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Male of Dysdera verkana sp. nov., bulb A retrolateral view B prolateral view C anterior view D posterior view. Scale bars: 0.25 mm." figureDoi="10.3897/zookeys.1146.97517.figure22" httpUri="https://binary.pensoft.net/fig/806633" pageId="0" pageNumber="43">22A</figureCitation>
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and Fig.
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<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Male of Dysdera sagartia sp. nov., bulb A retrolateral view B prolateral view C anterior view D posterior view. Scale bars: 0.25 mm." figureDoi="10.3897/zookeys.1146.97517.figure19" httpUri="https://binary.pensoft.net/fig/806630" pageId="0" pageNumber="43">19A</figureCitation>
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), and the relatively longer psembolus (i.e., length of psembolus/length of tegulum = 1.66 in
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<taxonomicName lsidName="D. verkana" pageId="0" pageNumber="43" rank="species" species="verkana">
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<emphasis italics="true" pageId="0" pageNumber="43">D. verkana</emphasis>
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</taxonomicName>
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sp. nov., vs. 1.44 in
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<taxonomicName lsidName="D. sagartia" pageId="0" pageNumber="43" rank="species" species="sagartia">
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<emphasis italics="true" pageId="0" pageNumber="43">D. sagartia</emphasis>
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</taxonomicName>
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sp. nov.).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="43" type="description">
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<paragraph pageId="0" pageNumber="43">Description.</paragraph>
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<paragraph pageId="0" pageNumber="43">
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<emphasis bold="true" pageId="0" pageNumber="43">Male.</emphasis>
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Habitus as in Fig.
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<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Male of Dysdera verkana sp. nov., habitus A dorsal view B ventral view C lateral view. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1146.97517.figure21" httpUri="https://binary.pensoft.net/fig/806632" pageId="0" pageNumber="43">21A-C</figureCitation>
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. Total length 10.8. Carapace 5.65 long, 4.26 wide. Eye diameters: AME 0.24, PME 0.23, PLE 0.21. Carapace, sternum, chelicerae, labium, and maxillae reddish brown. Legs orange. Abdomen cream-coloured, without any pattern. Spinnerets uniformly dark yellowish. Measurements of legs: I: 13.04 (3.90, 2.23, 3.10, 3.06, 0.75), II: 11.76 (3.39, 1.98, 2.91, 2.71, 0.77), III: 8.56 (2.64, 1.26, 1.69, 2.33, 0.64), IV: 11.46 (3.32, 1.76, 2.48, 3.20, 0.70). Spination: I: Fe: 2pl. II: Fe: 1pl. III: Fe: 1d, 2pl; Ti: 4pl, 2rl, 5v; Mt: 3pl, 2rl, 6v. IV: Fe: 6d, 1pl; Ti: 4pl, 4rl, 5v; Mt: 4pl, 2rl, 6v.
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</paragraph>
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<caption doi="10.3897/zookeys.1146.97517.figure21" httpUri="https://binary.pensoft.net/fig/806632" pageId="0" pageNumber="43" start="Figure 21" startId="F21">
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<paragraph pageId="0" pageNumber="43">
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<emphasis bold="true" pageId="0" pageNumber="43">Figure 21.</emphasis>
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Male of
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<taxonomicName authorityName="Zamani & Marusik & Szűts" authorityYear="2023" class="Arachnida" family="Dysderidae" genus="Dysdera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dysdera verkana" order="Araneae" pageId="0" pageNumber="43" phylum="Arthropoda" rank="species" species="verkana">
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<emphasis italics="true" pageId="0" pageNumber="43">Dysdera verkana</emphasis>
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</taxonomicName>
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sp. nov., habitus
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<emphasis bold="true" pageId="0" pageNumber="43">A</emphasis>
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dorsal view
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<emphasis bold="true" pageId="0" pageNumber="43">B</emphasis>
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ventral view
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<emphasis bold="true" pageId="0" pageNumber="43">C</emphasis>
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lateral view. Scale bars: 1.0 mm.
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</paragraph>
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</caption>
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<paragraph pageId="0" pageNumber="43">
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Palp as in Fig.
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<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Male of Dysdera verkana sp. nov., bulb A retrolateral view B prolateral view C anterior view D posterior view. Scale bars: 0.25 mm." figureDoi="10.3897/zookeys.1146.97517.figure22" httpUri="https://binary.pensoft.net/fig/806633" pageId="0" pageNumber="43">22A-D</figureCitation>
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; bulb ca. 2.2
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<normalizedToken originalValue="×">x</normalizedToken>
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longer than wide; tegulum bell-shaped, almost as long as wide; psembolus 1.66
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<normalizedToken originalValue="×">x</normalizedToken>
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longer than tegulum; median crest rounded, ca. 2.3
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<normalizedToken originalValue="×">x</normalizedToken>
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shorter than length of psembolus, ca. 2.7
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<normalizedToken originalValue="×">x</normalizedToken>
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wider than high; posterior apophysis broad; incision between tegulum and psembolus present; retrolateral crest roundly bent, forming right angle.
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</paragraph>
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<caption doi="10.3897/zookeys.1146.97517.figure22" httpUri="https://binary.pensoft.net/fig/806633" pageId="0" pageNumber="43" start="Figure 22" startId="F22">
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<paragraph pageId="0" pageNumber="43">
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<emphasis bold="true" pageId="0" pageNumber="43">Figure 22.</emphasis>
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Male of
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<taxonomicName authorityName="Zamani & Marusik & Szűts" authorityYear="2023" class="Arachnida" family="Dysderidae" genus="Dysdera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dysdera verkana" order="Araneae" pageId="0" pageNumber="43" phylum="Arthropoda" rank="species" species="verkana">
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<emphasis italics="true" pageId="0" pageNumber="43">Dysdera verkana</emphasis>
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</taxonomicName>
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sp. nov., bulb
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<emphasis bold="true" pageId="0" pageNumber="43">A</emphasis>
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retrolateral view
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<emphasis bold="true" pageId="0" pageNumber="43">B</emphasis>
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prolateral view
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<emphasis bold="true" pageId="0" pageNumber="43">C</emphasis>
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anterior view
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<emphasis bold="true" pageId="0" pageNumber="43">D</emphasis>
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posterior view. Scale bars: 0.25 mm.
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</paragraph>
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</caption>
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<paragraph pageId="0" pageNumber="43">
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<emphasis bold="true" pageId="0" pageNumber="43">Female.</emphasis>
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Unknown.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="43" type="distribution">
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<paragraph pageId="0" pageNumber="43">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="43">
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Known only from the type locality in Golestan Province, northern Iran (Fig.
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<figureCitation captionStart="Figure 35" captionStartId="F35" captionText="Figure 35. Distribution records of Dysdera spp. in Iran 1 D. achaemenes sp. nov. 2 D. bakhtiari sp. nov. 3 D. hormuzensis sp. nov. 4 D. iranica sp. nov. 5 D. isfahanica sp. nov. 6 D. mazeruni sp. nov. 7 D. persica sp. nov. 8 D. pococki 9 D. sagartia sp. nov. and D. medes sp. nov. 10 D. verkana sp. nov. 11 D. xerxesi sp. nov. 12 D. damavandica sp. nov. 13 D. tapuria sp. nov. 14 D. genoensis sp. nov." figureDoi="10.3897/zookeys.1146.97517.figure35" httpUri="https://binary.pensoft.net/fig/806646" pageId="0" pageNumber="43">35</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="43" type="crocata species group">
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<paragraph pageId="0" pageNumber="43">
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<taxonomicName authorityName="C. L. Koch" authorityYear="1838" class="Arachnida" family="Dysderidae" genus="Dysdera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dysdera crocata" order="Araneae" pageId="0" pageNumber="43" phylum="Arthropoda" rank="species" species="crocata">
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<emphasis italics="true" pageId="0" pageNumber="43">Dysdera crocata</emphasis>
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</taxonomicName>
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species group
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</paragraph>
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<paragraph pageId="0" pageNumber="43">
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<emphasis bold="true" pageId="0" pageNumber="43">Diagnosis.</emphasis>
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This group can be diagnosed by a combination of the following characters: the chelicerae straight or anteriorly convergent and longer than half of the length of the carapace, carapace broad and flat, and bulb with small or no lateral projection (
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<bibRefCitation author="Deeleman-Reinhold, CL" journalOrPublisher="Tijdschrift voor Entomologie" pageId="0" pageNumber="43" pagination="141 - 269" refId="B5" refString="Deeleman-Reinhold, CL, Deeleman, PR, 1988. Revision des Dysderinae (Araneae, Dysderidae), les especes mediterraneennes occidentales exceptees. Tijdschrift voor Entomologie 131: 141 - 269" title="Revision des Dysderinae (Araneae, Dysderidae), les especes mediterraneennes occidentales exceptees." volume="131" year="1988">Deeleman-Reinhold and Deeleman 1988</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |