237 lines
17 KiB
XML
237 lines
17 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.67.6703" ID-PMC="PMC4972245" ID-Pensoft-Pub="1314-2003-67-1" ID-Pensoft-UUID="D846EB3F7746FFFE4A469751FFEF3B22" ID-PubMed="27536193" ID-Zenodo-Dep="133270" ModsDocID="1314-2003-67-1" checkinTime="1469054247116" checkinUser="pensoft" docAuthor="Kriebel, Ricardo" docDate="2016" docId="ADB2ABA42EF05FA48370463051F6A740" docLanguage="en" docName="PhytoKeys 67: 1-326" docOrigin="PhytoKeys 67" docPubDate="2016-07-20" docSource="http://dx.doi.org/10.3897/phytokeys.67.6703" docTitle="Conostegia pendula Kriebel 2016, comb. nov." docType="treatment" docVersion="3" id="D846EB3F7746FFFE4A469751FFEF3B22" lastPageNumber="209" masterDocId="D846EB3F7746FFFE4A469751FFEF3B22" masterDocTitle="A Monograph of Conostegia (Melastomataceae, Miconieae)" masterLastPageNumber="326" masterPageNumber="1" pageNumber="208" updateTime="1668141747722" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A Monograph of Conostegia (Melastomataceae, Miconieae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Kriebel, Ricardo</mods:namePart>
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<mods:affiliation>Department of Botany, University of Wisconsin-Madison, 430 Lincoln Drive Madison, Wisconsin 53706, USA</mods:affiliation>
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<mods:nameIdentifier type="email">kriebelr@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>PhytoKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2016</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2016-07-20</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>67</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>326</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/phytokeys.67.6703</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.67.6703</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1314-2003-67-1</mods:identifier>
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<mods:identifier type="Pensoft-UUID">D846EB3F7746FFFE4A469751FFEF3B22</mods:identifier>
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<mods:identifier type="Zenodo-Dep">133270</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="182226934" LSID="urn:lsid:plazi:treatment:ADB2ABA42EF05FA48370463051F6A740" httpUri="http://treatment.plazi.org/id/ADB2ABA42EF05FA48370463051F6A740" lastPageId="209" lastPageNumber="209" pageId="207" pageNumber="208">
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<subSubSection pageId="207" pageNumber="208" type="nomenclature">
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<paragraph pageId="207" pageNumber="208">
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<taxonomicName LSID="ADB2ABA4-2EF0-5FA4-8370-463051F6A740" authority="(Umana & Almeda) Kriebel" authorityName="Kriebel" authorityYear="2016" baseAuthorityName="Umana & Almeda" baseAuthorityYear="2016" class="Magnoliopsida" family="Melastomataceae" genus="Conostegia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Conostegia pendula" order="Myrtales" pageId="207" pageNumber="208" phylum="Tracheophyta" rank="species" species="pendula" status="comb. nov.">
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Conostegia pendula (
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<normalizedToken originalValue="Umaña">Umana</normalizedToken>
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& Almeda) Kriebel
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</taxonomicName>
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<taxonomicNameLabel pageId="207" pageNumber="208">comb. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="207" pageNumber="208" type="reference_group">
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<paragraph pageId="207" pageNumber="208">
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<taxonomicName authorityName="Kriebel" authorityYear="2016" baseAuthorityName="Umana & Almeda" baseAuthorityYear="2016" class="Magnoliopsida" family="Melastomataceae" genus="Conostegia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Conostegia pendula" order="Myrtales" pageId="207" pageNumber="208" phylum="Tracheophyta" rank="species" species="pendula" status="comb. nov.">Conostegia pendula</taxonomicName>
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(
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<normalizedToken originalValue="Umaña">Umana</normalizedToken>
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& Almeda) Kriebel. Basionym:
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<taxonomicName authorityName="Kriebel" authorityYear="2016" class="Magnoliopsida" family="Melastomataceae" genus="Miconia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Miconia pendula" order="Myrtales" pageId="207" pageNumber="208" phylum="Tracheophyta" rank="species" species="pendula" status="comb. nov.">Miconia pendula</taxonomicName>
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<normalizedToken originalValue="Umaña">Umana</normalizedToken>
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& Almeda, Novon 3(1): 8, f. 2. 1993. Type: Costa Rica. Cartago: Refugio Nacional de Vida Silvestre
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<normalizedToken originalValue="Tapantí">Tapanti</normalizedToken>
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, orilla de Sendero Los Palmitos, 1300-1400 m, 0944'00" N, 8347'00"W, 2 August 1990, G.
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<normalizedToken originalValue="Umaña">Umana</normalizedToken>
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, Kennedy, H., Nilson, V., & R.
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<normalizedToken originalValue="Chacón">Chacon</normalizedToken>
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391 (holotype: CR!, isotypes: BM, BR, CAS!, COL, F!, MEXU!, MO!, NY!, PMA, US!, USJ).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="207" pageNumber="208" type="description">
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<paragraph pageId="207" pageNumber="208">Description.</paragraph>
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<paragraph pageId="207" pageNumber="208">
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Shrub to small tree 1.5-5 m tall with somewhat tetragonal stems in newer branches densely covered with a lanate indument of variable dendritic trichomes; the nodal line present. Petioles 1-7 cm. Leaves at a node equal or somewhat unequal at a node. Leaf blades 6.5-28
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<normalizedToken originalValue="×">x</normalizedToken>
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3.5-13.5 cm, 7-plinerved, with the inner pairs of innermost veins arising 1-5 cm above the base mostly in alternate fashion, elliptic to elliptic-ovate, the base obtuse to asymmetrical, the apex acuminate, the margin entire to inconspicuously denticulate, the adaxial surface strigose or lightly lanate to glabrous, lanate on the primary veins abaxially, and with furfuraceous hairs on higher order veins. Inflorescence a pendant modified cyme branching well above the base, 4-12 cm long, the flowers clustered at the end of the branches, accessory branches absent, rachis lanate, bracteoles 0.2-0.3 mm, persistent. Pedicel 0.5-2.5 mm. Flower buds 3-4
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<normalizedToken originalValue="×">x</normalizedToken>
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2-3 mm. Flowers 5-merous, calyx not calyptrate but closed in bud and crowned by an apiculum, rupturing irregularly at anthesis into 5 hyaline irregular lobes 0.25-0.5 mm long, exterior calyx teeth narrowly triangular, 0.9-1.3 mm long, torus glabrous; the hypanthium 2-3
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<normalizedToken originalValue="×">x</normalizedToken>
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1-2 mm, campanulate. Petals 2.9-3.5
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<normalizedToken originalValue="×">x</normalizedToken>
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1 mm, pale pink, linear-oblong, not observed at anthesis, glabrous, the apex obtuse to rounded. Stamens 10, ca. 3-4 mm, radially arranged around the style, the filaments ca. 2 mm long, geniculate near the apex, white, anthers 1-1.5
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<normalizedToken originalValue="×">x</normalizedToken>
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0.35-0.5 mm, linear-oblong, somewhat recurved, yellow, the connective thickened dorsally, the pore 0.1 mm wide, slightly dorsally inclined. Ovary 5 locular, 2/3 inferior, slightly fluted and with glandular hairs on the apex. Style reportedly 2.5-5 mm long, not observed at anthesis but notes on the type suggest it is exserted, stigma truncate and somewhat dilated. Berry 4-5
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<normalizedToken originalValue="×">x</normalizedToken>
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4-5 mm, pink turning purple black when mature. Seeds 0.4-0.6 mm long, angular pyramidate to somewhat cescent shaped in profile, the testa smooth.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="208" pageNumber="209" type="distribution">
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<paragraph pageId="208" pageNumber="209">
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<pageBreakToken pageId="208" pageNumber="209" start="start">Distribution</pageBreakToken>
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</paragraph>
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<paragraph pageId="208" pageNumber="209">
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(Fig.
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<figureCitation captionStart="Figure 184" captionStartId="F184" captionText="Figure 184. Distribution of Conostegia pendula." figureDoi="10.3897/phytokeys.67.6703.figure184" httpUri="https://binary.pensoft.net/fig/96542" pageId="208" pageNumber="209">184</figureCitation>
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). Endemic to cloud forests of the Caribbean slope of the Central and Talamanca Cordilleras of Costa Rica, 1200-1800 m in elevation.
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</paragraph>
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<caption ID-Zenodo-Dep="974025" doi="10.3897/phytokeys.67.6703.figure184" httpUri="https://binary.pensoft.net/fig/96542" pageId="208" pageNumber="209" start="Figure 184" startId="F184">
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<paragraph pageId="208" pageNumber="209">
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<emphasis bold="true" pageId="208" pageNumber="209">Figure 184.</emphasis>
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Distribution of
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<taxonomicName authorityName="Kriebel" authorityYear="2016" baseAuthorityName="Umana & Almeda" baseAuthorityYear="2016" class="Magnoliopsida" family="Melastomataceae" genus="Conostegia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Conostegia pendula" order="Myrtales" pageId="208" pageNumber="209" phylum="Tracheophyta" rank="species" species="pendula">
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<emphasis italics="true" pageId="208" pageNumber="209">Conostegia pendula</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</caption>
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<paragraph pageId="208" pageNumber="209">
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The description of this species is for the most part based on the original one by
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<bibRefCitation author="Almeda, F" journalOrPublisher="Novon" pageId="240" pageNumber="241" pagination="5 - 10" publicationUrl="10.2307/3391407" refId="B11" refString="Almeda, F, Umana Dodero, G, 1993. New species of Miconia (Melastomataceae) from Costa Rica. Novon 3: 5 - 10, DOI: 10.2307/3391407" title="New species of Miconia (Melastomataceae) from Costa Rica." url="10.2307/3391407" volume="3" year="1993">
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Almeda and
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<normalizedToken originalValue="Umaña">Umana</normalizedToken>
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Dodero (1993)
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</bibRefCitation>
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. Few recent collections have been made since its description but as
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<bibRefCitation author="Almeda, F" journalOrPublisher="Novon" pageId="240" pageNumber="241" pagination="5 - 10" publicationUrl="10.2307/3391407" refId="B11" refString="Almeda, F, Umana Dodero, G, 1993. New species of Miconia (Melastomataceae) from Costa Rica. Novon 3: 5 - 10, DOI: 10.2307/3391407" title="New species of Miconia (Melastomataceae) from Costa Rica." url="10.2307/3391407" volume="3" year="1993">
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Almeda and
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<normalizedToken originalValue="Umaña">Umana</normalizedToken>
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Dodero (1993)
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</bibRefCitation>
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predicted, the species has been collected in the Cordillera de Talamanca in Fila Matama,
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<normalizedToken originalValue="Limón">Limon</normalizedToken>
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. The notes on the type specimen describe the anthers around the style which suggests the style is exserted as in its close relative like
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<taxonomicName authorityName="Kriebel" authorityYear="2016" baseAuthorityName="Almeda & Umana" baseAuthorityYear="2016" class="Magnoliopsida" family="Melastomataceae" genus="Conostegia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Conostegia friedmaniorum" order="Myrtales" pageId="208" pageNumber="209" phylum="Tracheophyta" rank="species" species="friedmaniorum">
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<emphasis italics="true" pageId="208" pageNumber="209">Conostegia friedmaniorum</emphasis>
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</taxonomicName>
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.
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<bibRefCitation author="Almeda, F" journalOrPublisher="Novon" pageId="240" pageNumber="241" pagination="5 - 10" publicationUrl="10.2307/3391407" refId="B11" refString="Almeda, F, Umana Dodero, G, 1993. New species of Miconia (Melastomataceae) from Costa Rica. Novon 3: 5 - 10, DOI: 10.2307/3391407" title="New species of Miconia (Melastomataceae) from Costa Rica." url="10.2307/3391407" volume="3" year="1993">
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Almeda and
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<normalizedToken originalValue="Umaña">Umana</normalizedToken>
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Dodero (1993)
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</bibRefCitation>
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also noted the filament geniculation in this species, something not commonly done. They mention strong herbivory on these plants as is frequently noted in species of section
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<taxonomicName class="Magnoliopsida" family="Melastomataceae" genus="Conostegia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Myrtales" pageId="208" pageNumber="209" phylum="Tracheophyta" rank="section" section="Geniculatae">
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<emphasis italics="true" pageId="208" pageNumber="209">Geniculatae</emphasis>
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</taxonomicName>
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. The label on
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<emphasis italics="true" pageId="208" pageNumber="209">Almeda et al. 7245</emphasis>
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(CAS) records a pleasant acidic flavor of the berries of this species.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="209" lastPageNumber="210" pageId="208" pageNumber="209" type="materials_examined">
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<paragraph pageId="208" pageNumber="209">Specimens examined.</paragraph>
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<paragraph lastPageId="209" lastPageNumber="210" pageId="208" pageNumber="209">
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<materialsCitation collectingDate="2016-01-01" collectingDateMax="2016-12-31" collectingDateMin="2016-01-01" collectorName="P. N. Tapanti" country="COSTA RICA" county="Refugio Nacional de Vida Silvestre Tapanti" location="Rio Grande de Orosi" municipality="Tapanti Reserve" specimenCount="1" stateProvince="Cartago">
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<emphasis bold="true" pageId="208" pageNumber="209">
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<collectingCountry name="Costa Rica">COSTA RICA</collectingCountry>
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.
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<collectingRegion country="Costa Rica" name="Cartago">Cartago</collectingRegion>
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</emphasis>
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:
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<collectingCounty>
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Refugio Nacional de Vida Silvestre
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<normalizedToken originalValue="Tapantí">Tapanti</normalizedToken>
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</collectingCounty>
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, sendero Palmito about
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<quantity metricMagnitude="3" metricUnit="m" metricValue="6.0" unit="km" value="6.0">6 km</quantity>
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from main gate to refuge, Almeda et al.
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<pageBreakToken pageId="209" pageNumber="210" start="start">7245</pageBreakToken>
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(CAS, CR);
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<collectingMunicipality>
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<normalizedToken originalValue="Tapantí">Tapanti</normalizedToken>
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Reserve
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</collectingMunicipality>
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ca
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<quantity metricMagnitude="3" metricUnit="m" metricValue="1.0" unit="km" value="1.0">1 km</quantity>
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S of jct of Quebrada Salto and
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<location LSID="urn:lsid:plazi:treatment:ADB2ABA42EF05FA48370463051F6A740:01494D3335315797E9D16E58824608B9" country="COSTA RICA" county="Refugio Nacional de Vida Silvestre Tapanti" municipality="Tapanti Reserve" name="Rio Grande de Orosi" stateProvince="Cartago">
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<normalizedToken originalValue="Río">Rio</normalizedToken>
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Grande de
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<normalizedToken originalValue="Orosí">Orosi</normalizedToken>
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</location>
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, Croat and Grayum 68255 (CAS); in forest along creek debouching into
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<location LSID="urn:lsid:plazi:treatment:ADB2ABA42EF05FA48370463051F6A740:A4C0654C05A573539BC6AA34482D5892" country="COSTA RICA" county="Refugio Nacional de Vida Silvestre Tapanti" municipality="Tapanti Reserve" name="Rio Grande de Orosi" stateProvince="Cartago">
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<normalizedToken originalValue="Río">Rio</normalizedToken>
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Grande de Orosi
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</location>
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, from the east, ca.
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<quantity metricMagnitude="3" metricUnit="m" metricValue="1.0" unit="km" value="1.0">1 km</quantity>
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upstream from confluence of Quebrada Salta,
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<normalizedToken originalValue="Tapantí">Tapanti</normalizedToken>
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, Grayum and Sleeper 3486 (CAS, CR);
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<collectorName>
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P. N.
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<normalizedToken originalValue="Tapantí">Tapanti</normalizedToken>
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</collectorName>
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, sendero Palmito,
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<normalizedToken originalValue="Umaña">Umana</normalizedToken>
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, Valerio and
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<normalizedToken originalValue="Chacón">Chacon</normalizedToken>
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378 (CAS, CR)
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</materialsCitation>
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.
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<materialsCitation collectingDate="2016-01-01" collectingDateMax="2016-12-31" collectingDateMin="2016-01-01" collectorName="La Legua" country="Costa Rica" county="Canton de Barva" location="Rio San Fernando" municipality="Rio Volcan" specimenCount="1" stateProvince="Heredia">
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<emphasis bold="true" pageId="209" pageNumber="210">
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<collectingRegion country="Costa Rica" name="Heredia">Heredia</collectingRegion>
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</emphasis>
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:
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<collectingCounty>
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<normalizedToken originalValue="Cantón">Canton</normalizedToken>
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de Barva
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</collectingCounty>
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,
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<collectorName>La Legua</collectorName>
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, Finca Montreal, ridge between headwaters of
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<collectingMunicipality>
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<normalizedToken originalValue="Río">Rio</normalizedToken>
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<normalizedToken originalValue="Volcán">Volcan</normalizedToken>
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</collectingMunicipality>
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and
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<location LSID="urn:lsid:plazi:treatment:ADB2ABA42EF05FA48370463051F6A740:CDDC22503F7E98CB970EFA3D68C011A0" country="Costa Rica" county="Canton de Barva" municipality="Rio Volcan" name="Rio San Fernando" stateProvince="Heredia">
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<normalizedToken originalValue="Río">Rio</normalizedToken>
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San Fernando
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</location>
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, Boyle et al. 1168 (CAS, CR, INB)
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</materialsCitation>
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.
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<materialsCitation collectingDate="2016-01-01" collectingDateMax="2016-12-31" collectingDateMin="2016-01-01" collectorName="P. N. La Amistad, de Aguas Zarcas" country="Costa Rica" location="Limon" specimenCount="1" stateProvince="Limon">
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<emphasis bold="true" pageId="209" pageNumber="210">
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<normalizedToken originalValue="Limón">
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<collectingRegion country="Costa Rica" name="Limon">Limon</collectingRegion>
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</normalizedToken>
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</emphasis>
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:
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<collectorName>P. N. La Amistad</collectorName>
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, Fila Matama, cerca de
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<quantity metricMagnitude="4" metricUnit="m" metricValue="1.1" unit="km" value="11.0">11 km</quantity>
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SW del pueblo
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<collectorName>de Aguas Zarcas</collectorName>
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,
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<normalizedToken originalValue="Santamaría">Santamaria</normalizedToken>
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et al. 6672 (INB, NY)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |