212 lines
27 KiB
XML
212 lines
27 KiB
XML
<document id="2243A5D20EE43D100A13B3AFC43A614A" ENCODING="UTF-8" ID-GBIF-Dataset="a3a089ba-4f85-4373-8813-4b9c6a99d6c1" ModsDocID="z01096p001" checkinTime="1247049491658" checkinUser="thomas" docAuthor="Douglass F. Hoese & Helen K. Larson" docDate="2005" docId="A92DCF0694E4C7E790987AE1F2B833CE" docLanguage="en" docName="2005_Hoese_Larson_gg1.xml" docOrigin="Zootaxa 1096" docSource="http://www.zoobank.org/urn:lsid:zoobank.org:pub:5090B868-375F-42B9-A4E4-ED7739E68786" docTitle="Hetereleotris" docType="treatment" docVersion="13" lastPageNumber="4" masterDocId="66DFBA1B650F5042A23215D98F336892" masterDocTitle="Description of two new species of Hetereleotris (Gobiidae) from the south Pacific, with a revised key to species and synonymization of the genus Pascua with Hetereleotris." masterLastPageNumber="16" masterPageNumber="1" pageNumber="1" updateTime="1701325942412" updateUser="ExternalLinkService">
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<mods:titleInfo id="00F563DBD7FAB63F3298EB12DBCD2DDB">
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<mods:title id="6C15AE67AD39E743F3E65581BDC7A84A">Description of two new species of Hetereleotris (Gobiidae) from the south Pacific, with a revised key to species and synonymization of the genus Pascua with Hetereleotris.</mods:title>
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<mods:roleTerm id="6D2A894C1F8A8A3542C9D39F136CC32E">Author</mods:roleTerm>
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<mods:namePart id="38F11540B70E89C6D6DA797EF4A83397">Douglass F. Hoese</mods:namePart>
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<mods:namePart id="DCCE7627D7DA2D6314A09ABC90E493AF">Helen K. Larson</mods:namePart>
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<mods:typeOfResource id="0F88166E5995D2F1C7831784A3B65823">text</mods:typeOfResource>
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<mods:title id="9978DAFE7EBA1B99D70BE4922B6F4176">Zootaxa</mods:title>
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<mods:date id="16E63E4938F101556E982912782EF36A">2005</mods:date>
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<mods:number id="A24A1B8A42015DECFD25C85B56F41CC4">1096</mods:number>
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<mods:identifier id="059D6A00B3D69924562114CC03136DDB" type="Plazi-Custom">z01096p001</mods:identifier>
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<treatment id="A92DCF0694E4C7E790987AE1F2B833CE" ID-DOI="http://doi.org/10.5281/zenodo.6264613" ID-GBIF-Taxon="100121598" ID-Zenodo-Dep="6264613" LSID="urn:lsid:plazi:treatment:A92DCF0694E4C7E790987AE1F2B833CE" httpUri="http://treatment.plazi.org/id/A92DCF0694E4C7E790987AE1F2B833CE" lastPageNumber="4" pageNumber="1">
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<subSubSection id="120EDEFAAC189A892C7E0FD7FE6695F7" type="nomenclature">
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<paragraph id="B346F3622434BB8609140C7A3E81EF04">
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[[ Genus
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<taxonomicName id="E61AE691277FB4D5F08BD51A24006288" ID-CoL="62V3V" ID-ENA="2067727" LSID-ZBK="urn:lsid:zoobank.org:act:D5E53DC5-5480-4F01-9C4C-2D9BF239B442" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Hetereleotris Bleeker 1874:306" order="Perciformes" pageNumber="1" phylum="Chordata" rank="genus">Hetereleotris</taxonomicName>
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]]
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</paragraph>
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</subSubSection>
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<subSubSection id="0CB2A2EB2AEB38F0F0107DD3445DE745" type="discussion">
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<paragraph id="B156E30B5459B2F0C53291387F2783F5" pageNumber="1">
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The gobiid genus
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<taxonomicName id="16398D3137C1623AAC6616BEF2FA840D" LSID-ZBK="urn:lsid:zoobank.org:act:D5E53DC5-5480-4F01-9C4C-2D9BF239B442" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Hetereleotris Bleeker 1874:306" order="Perciformes" pageNumber="1" phylum="Chordata" rank="genus">Hetereleotris</taxonomicName>
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is speciose in the western Indian Ocean. Smith (1958) placed a number of species in
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<taxonomicName id="A6FB02F36C37EAA421B951F362DFAD89" LSID-ZBK="urn:lsid:zoobank.org:act:E59F9F13-515E-4908-BF3D-E8A0D425C382" family="Gobiidae" genus="Monishia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Monishia Smith 1959:206" order="Perciformes" pageNumber="1" phylum="Chordata" rank="genus">Monishia</taxonomicName>
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or in various genera, now regarded by Akihito & Meguro (1981) and Hoese (1986) as junior synonyms of
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<taxonomicName id="FCC4FE30B9E48222649CF373ED0AC0C1" LSID-ZBK="urn:lsid:zoobank.org:act:D5E53DC5-5480-4F01-9C4C-2D9BF239B442" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Hetereleotris Bleeker 1874:306" order="Perciformes" pageNumber="1" phylum="Chordata" rank="genus">Hetereleotris</taxonomicName>
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. Hoese & Winterbottom (1979) treated some species from South Africa, while Hoese (1986) revised the genus and recognized 13 species and synonymized several nominal genera. Subsequently Gill (1998) described an additional species from Mauritius. The present paper describes two new species from the south Pacific and redescribes
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<taxonomicName id="6606D44BE1CE24FCCFD7B4A23403CB34" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="1" phylum="Chordata" rank="species" species="caudilinea">Hetereleotris caudilinea (Randall)</taxonomicName>
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, in order to compare its characteristics with the two new taxa. Previously only a single species of
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<taxonomicName id="22E438A34099797F56A597295B291667" LSID-ZBK="urn:lsid:zoobank.org:act:D5E53DC5-5480-4F01-9C4C-2D9BF239B442" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Hetereleotris Bleeker 1874:306" order="Perciformes" pageNumber="1" phylum="Chordata" rank="genus">Hetereleotris</taxonomicName>
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had been known from outside the Indian Ocean.
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</paragraph>
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<paragraph id="C73BD75ACD1C4C38B7EE413C77A6BBBE" pageNumber="2">
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Tentatively the species are placed here in
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<taxonomicName id="7351615433E1CDB438A62F9F56BB2657" LSID-ZBK="urn:lsid:zoobank.org:act:D5E53DC5-5480-4F01-9C4C-2D9BF239B442" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Hetereleotris Bleeker 1874:306" order="Perciformes" pageNumber="2" phylum="Chordata" rank="genus">Hetereleotris</taxonomicName>
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based on the presence of 27 vertebrae and the closure of the first gill slit. Other genera with 27 vertebrae from the Indo-Pacific region include
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<taxonomicName id="D427E282108AFABEF653B670BA780BF8" LSID-ZBK="urn:lsid:zoobank.org:act:ADAF83BF-42AD-4A07-8D23-0365879A2423" family="Gobiidae" genus="Psammogobius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Psammogobius Smith 1935:215" order="Perciformes" pageNumber="2" phylum="Chordata" rank="genus">Psammogobius</taxonomicName>
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,
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<taxonomicName id="1C6D7B6E8617D82422B409B56DD2A5EA" LSID-ZBK="urn:lsid:zoobank.org:act:4DC3E52C-04FA-4AEB-B7F4-9941C410D753" family="Gobiidae" genus="Glossogobius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Glossogobius Gill 1859:46" order="Perciformes" pageNumber="2" phylum="Chordata" rank="genus">Glossogobius</taxonomicName>
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,
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<taxonomicName id="671FD70EB70F6B07B98EC6B3D12532F0" LSID-ZBK="urn:lsid:zoobank.org:act:E608AAC1-2634-4D90-B4F5-3F6F4BD78FBB" family="Gobiidae" genus="Bathygobius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Bathygobius Bleeker 1878:54" order="Perciformes" pageNumber="2" phylum="Chordata" rank="genus">Bathygobius</taxonomicName>
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,
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<taxonomicName id="34C2F5D3DE9396A18199B6FCC00E0B2D" LSID-ZBK="urn:lsid:zoobank.org:act:97FDC767-1FD7-483E-8184-8BED94A7B9AD" family="Gobiidae" genus="Palutrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Palutrus Smith 1959:208" order="Perciformes" pageNumber="2" phylum="Chordata" rank="genus">Palutrus</taxonomicName>
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and
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<taxonomicName id="F44627070B4D489B699AC654F1B7CF21" LSID-ZBK="urn:lsid:zoobank.org:act:2B81E71A-6677-4E07-9CEB-8947612B603B" family="Gobiidae" genus="Coryogalops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Coryogalops Smith 1958:144" order="Perciformes" pageNumber="2" phylum="Chordata" rank="genus">Coryogalops</taxonomicName>
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. The vertebral counts are highly consistent within these genera, except for some dwarf species of
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<taxonomicName id="C88C4968BD4D9CE092946446347D7A96" LSID-ZBK="urn:lsid:zoobank.org:act:97FDC767-1FD7-483E-8184-8BED94A7B9AD" family="Gobiidae" genus="Palutrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Palutrus Smith 1959:208" order="Perciformes" pageNumber="2" phylum="Chordata" rank="genus">Palutrus</taxonomicName>
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and
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<taxonomicName id="FB8EAC06E7BBE1F9982BCA8E25FC64A6" LSID-ZBK="urn:lsid:zoobank.org:act:4DC3E52C-04FA-4AEB-B7F4-9941C410D753" family="Gobiidae" genus="Glossogobius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Glossogobius Gill 1859:46" order="Perciformes" pageNumber="2" phylum="Chordata" rank="genus">Glossogobius</taxonomicName>
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which have 26 vertebrae and some species of
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<taxonomicName id="D033286C1780A6E9E81DE13A488275B3" LSID-ZBK="urn:lsid:zoobank.org:act:4DC3E52C-04FA-4AEB-B7F4-9941C410D753" family="Gobiidae" genus="Glossogobius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Glossogobius Gill 1859:46" order="Perciformes" pageNumber="2" phylum="Chordata" rank="genus">Glossogobius</taxonomicName>
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, known from high altitudes of greater than 500m from New Guinea, which have 28 or 29 vertebrae. All of these also share a lobed mental frenum (highly modified and sometimes absent in
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<taxonomicName id="A3B94F04BFC03B8E39EDB99896E41C6D" LSID-ZBK="urn:lsid:zoobank.org:act:4DC3E52C-04FA-4AEB-B7F4-9941C410D753" family="Gobiidae" genus="Glossogobius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Glossogobius Gill 1859:46" order="Perciformes" pageNumber="2" phylum="Chordata" rank="genus">Glossogobius</taxonomicName>
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).
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</paragraph>
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<paragraph id="8FD2BEC7ED494C8B77A5D8E5F7B3CADB" pageNumber="2">
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All three species described here share a number of characters not commonly found in
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<taxonomicName id="FE5A012CE1D883A9A8C7CBBC30E10465" LSID-ZBK="urn:lsid:zoobank.org:act:D5E53DC5-5480-4F01-9C4C-2D9BF239B442" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Hetereleotris Bleeker 1874:306" order="Perciformes" pageNumber="2" phylum="Chordata" rank="genus">Hetereleotris</taxonomicName>
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, particularly the flattened and elongate urogenital papilla of the males, the modified basicaudal scales (Fig. 1), the posterior nostril a simple pore or with only a slightly elevated margin anteriorly, two papillae just behind the mental frenum and the reduced transverse papilla pattern (Fig. 2). The modified urogenital papilla of males is similar to that described as the nonfimbriate condition for many species of
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<taxonomicName id="DFB7835D584C55CDB059BB4E3CBC45E4" LSID-ZBK="urn:lsid:zoobank.org:act:8AC2C2C8-4045-4563-9189-6C090E61ACAB" family="Gobiidae" genus="Eviota" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Eviota Jenkins 1903:501" order="Perciformes" pageNumber="2" phylum="Chordata" rank="genus">Eviota</taxonomicName>
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(Lachner & Karnella 1980), a genus which has 25 or 26 vertebrae. The modified basicaudal scale is found in a number of species of New World gobies (
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<normalizedToken id="74DA85B072B66766EF1DFD61215250B5" originalValue="Böhlke">Boehlke</normalizedToken>
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& Robins 1968), which also have 27 vertebrae, but which have 11 precaudal vertebrae, rather than 10 as in the Indo-Pacific species. The closure of the first gill slit is rare, but also found in the eleotrid
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<taxonomicName id="DDE51A61935330621AE4D806AB40A1D2" LSID-ZBK="urn:lsid:zoobank.org:act:86A153A0-8DED-4BB0-91E8-8E3D7E7B92A0" family="Thalasseleotrididae" genus="Thalasseleotris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Thalasseleotris Hoese & Larson 1987:44" order="Perciformes" pageNumber="2" phylum="Chordata" rank="genus">Thalasseleotris</taxonomicName>
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(Hoese & Larson, 1987), which bears a remarkable overall external similarity to the present species, but has 6 branchiostegal rays. Other species of
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<taxonomicName id="DE778C97A165C8A8DC11F224ACDA1DEB" LSID-ZBK="urn:lsid:zoobank.org:act:D5E53DC5-5480-4F01-9C4C-2D9BF239B442" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Hetereleotris Bleeker 1874:306" order="Perciformes" pageNumber="2" phylum="Chordata" rank="genus">Hetereleotris</taxonomicName>
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usually have the posterior nostril with an elevated rim or at the end of a short tube, two parallel rows of papillae behind the mental frenum, although in
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<taxonomicName id="8898C8A57AF30617023229B9A2BE3B8A" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="2" phylum="Chordata" rank="species" species="zanzibariensis">H. zanzibariensis</taxonomicName>
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, the number of papillae in each row can be as few as 3. A transverse and reduced transverse papilla pattern, similar to the new
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<taxonomicName id="BF4A9E93946C550560C7C8E7DECA3430" LSID-ZBK="urn:lsid:zoobank.org:act:D5E53DC5-5480-4F01-9C4C-2D9BF239B442" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Hetereleotris Bleeker 1874:306" order="Perciformes" pageNumber="2" phylum="Chordata" rank="genus">Hetereleotris</taxonomicName>
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species, also occur in
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<taxonomicName id="C1C2F7ED072DF448A52E52C39A710582" family="Gobiidae" genus="Priolepis" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="2" phylum="Chordata" rank="genus">Priolepis</taxonomicName>
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(Winterbottom & Burridge, 1992, 1993).
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</paragraph>
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<paragraph id="304259D08FED7DBCB6A660E5917E4B75" pageNumber="3">
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Despite these differences from typical
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<taxonomicName id="3B954C423E0EE45171894AC106313C79" LSID-ZBK="urn:lsid:zoobank.org:act:D5E53DC5-5480-4F01-9C4C-2D9BF239B442" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Hetereleotris Bleeker 1874:306" order="Perciformes" pageNumber="3" phylum="Chordata" rank="genus">Hetereleotris</taxonomicName>
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, the species are included here in that genus until a more thorough phylogenetic analysis of the Indo-Pacific genera (particularly those with 27 vertebrae) can be carried out. Examination of the osteology of one species described below shows it to be typical of other small gobiids and consistent with that described for
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<taxonomicName id="F0307FC7D4CB50A2CE998FC04FC550AE" LSID-ZBK="urn:lsid:zoobank.org:act:D5E53DC5-5480-4F01-9C4C-2D9BF239B442" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Hetereleotris Bleeker 1874:306" order="Perciformes" pageNumber="3" phylum="Chordata" rank="genus">Hetereleotris</taxonomicName>
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by Hoese (1986). Randall (2005) defined
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<taxonomicName id="CDFCCA411F538502C77B3D2054575D1F" LSID-ZBK="urn:lsid:zoobank.org:act:A528131D-AC68-4546-B0D0-6E94AC65DA5B" family="Gobiidae" genus="Pascua" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Pascua Randall 2005:20" order="Perciformes" pageNumber="3" phylum="Chordata" rank="genus">Pascua</taxonomicName>
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based on several characters. These are:
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</paragraph>
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<paragraph id="1DBE5410B8C6B21C8953C8978F3A2BFB" pageNumber="3">
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1 head width less than body depth (common to many gobioids and some species of
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<taxonomicName id="B5248E73090DE50DC68218B5849EDAD2" LSID-ZBK="urn:lsid:zoobank.org:act:D5E53DC5-5480-4F01-9C4C-2D9BF239B442" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Hetereleotris Bleeker 1874:306" order="Perciformes" pageNumber="3" phylum="Chordata" rank="genus">Hetereleotris</taxonomicName>
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, such as
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<taxonomicName id="1A7ACC3477A47808388EB416C3AE87DF" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="tentaculata">H. tentaculata</taxonomicName>
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, and therefore not useful as a generic character);
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</paragraph>
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<paragraph id="38FC9596153D2E7BA3265049D8041748" pageNumber="3">
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2 1st gill slit closed by membrane - shared with all
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<taxonomicName id="16D310803D4DE5B7E9503989CB881970" LSID-ZBK="urn:lsid:zoobank.org:act:D5E53DC5-5480-4F01-9C4C-2D9BF239B442" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Hetereleotris Bleeker 1874:306" order="Perciformes" pageNumber="3" phylum="Chordata" rank="genus">Hetereleotris</taxonomicName>
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;
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</paragraph>
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<paragraph id="3DB315F7F07C031E05BB4C2E19B01B43" pageNumber="3">
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3 gill opening ending at level of ventral edge of pectoral-fin base - shared with all species of
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<taxonomicName id="C71CCDD4E8F308D7F72D78ED3525E98C" LSID-ZBK="urn:lsid:zoobank.org:act:D5E53DC5-5480-4F01-9C4C-2D9BF239B442" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Hetereleotris Bleeker 1874:306" order="Perciformes" pageNumber="3" phylum="Chordata" rank="genus">Hetereleotris</taxonomicName>
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and numerous other gobiid genera;
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</paragraph>
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<paragraph id="3D40E9C1A99AFA01F2230CD100CD9047" pageNumber="3">
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4 posterior nostril not tubular as in other
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<taxonomicName id="C18FD879092E776D68873603DD3A1F06" LSID-ZBK="urn:lsid:zoobank.org:act:D5E53DC5-5480-4F01-9C4C-2D9BF239B442" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Hetereleotris Bleeker 1874:306" order="Perciformes" pageNumber="3" phylum="Chordata" rank="genus">Hetereleotris</taxonomicName>
|
|
;
|
|
</paragraph>
|
|
<paragraph id="29E28A4EFA41BDDDB8917E4C2B757B46" pageNumber="3">
|
|
5 a complete cephalic sensory system - complete is a misleading term, because many gobiid species have more head pores than this species and all
|
|
<taxonomicName id="05509FF09120EEC293D034616638C9DC" LSID-ZBK="urn:lsid:zoobank.org:act:D5E53DC5-5480-4F01-9C4C-2D9BF239B442" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Hetereleotris Bleeker 1874:306" order="Perciformes" pageNumber="3" phylum="Chordata" rank="genus">Hetereleotris</taxonomicName>
|
|
lack the posterior portion of the oculoscapular canal (above the operculum) and some species lack pores entirely;
|
|
</paragraph>
|
|
<paragraph id="B9A01C7B88964C876F49FCF532893856" pageNumber="3">
|
|
6 reduced sensory papillae on cheek - species in other genera such as
|
|
<taxonomicName id="2A69F3C5C69B53558DCD048608AF2F69" family="Gobiidae" genus="Priolepis" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="genus">Priolepis</taxonomicName>
|
|
show a similar variation from transverse to reduced transverse;
|
|
</paragraph>
|
|
<paragraph id="9ACC0DF8A1DDD76675628AF6569C3760" lastPageNumber="4" pageNumber="3">
|
|
7 dorsal fins not completely separate - common to some other species of
|
|
<taxonomicName id="BAD59962E93B91D1E0DAC1E73CAC1793" LSID-ZBK="urn:lsid:zoobank.org:act:D5E53DC5-5480-4F01-9C4C-2D9BF239B442" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Hetereleotris Bleeker 1874:306" order="Perciformes" pageNumber="3" phylum="Chordata" rank="genus">Hetereleotris</taxonomicName>
|
|
; in material of the species treated here the first dorsal fin connects to the base of the second dorsal fin, varying from attaching to the back just before the second dorsal fin to the basal one fifth of the spine of the second dorsal fin, which is similar to species of
|
|
<taxonomicName id="1186C39BEE17C4E37E8A36D945D1F9D7" LSID-ZBK="urn:lsid:zoobank.org:act:D5E53DC5-5480-4F01-9C4C-2D9BF239B442" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Hetereleotris Bleeker 1874:306" order="Perciformes" pageNumber="3" phylum="Chordata" rank="genus">Hetereleotris</taxonomicName>
|
|
, such as
|
|
<taxonomicName id="347C32DF1292E0F90D284209BC1D9B02" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="vulgaris">H. vulgaris</taxonomicName>
|
|
and
|
|
<taxonomicName id="7E3ED8B49CAA52DBA7C7331D7ABAD164" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="aporia">H. aporia</taxonomicName>
|
|
as reported by Hoese and Winterbottom, 1979.
|
|
</paragraph>
|
|
<paragraph id="3434F6D4CF40E4A05201348B5E35F24F" pageNumber="4">
|
|
8 dorsal fin pterygiophores as in
|
|
<taxonomicName id="4D356F5A89FC5A1215381EC9B43FDF9C" LSID-ZBK="urn:lsid:zoobank.org:act:E608AAC1-2634-4D90-B4F5-3F6F4BD78FBB" family="Gobiidae" genus="Bathygobius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Bathygobius Bleeker 1878:54" order="Perciformes" pageNumber="4" phylum="Chordata" rank="genus">Bathygobius</taxonomicName>
|
|
- common to most gobiines;
|
|
</paragraph>
|
|
<paragraph id="49C8B92FF11C3197400AB43E4BC9C637" pageNumber="4">
|
|
9 17 segmented caudal rays (14 branched) - 17 segmented rays is common to most gobioid fishes, we find 13-15 branched caudal rays for
|
|
<taxonomicName id="CEFF11C3DE1CB2A298D9FEFB0680D6B0" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="4" phylum="Chordata" rank="species" species="caudilinea">caudilinea</taxonomicName>
|
|
and the species described here;
|
|
</paragraph>
|
|
<paragraph id="F4F8C5011A52B2A09486C9320CB980CC" pageNumber="4">
|
|
10 pectoral rays 20 or 21 - we find 18-21 rays for
|
|
<taxonomicName id="7A05A490A45FB972E243697B04B3FB20" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="4" phylum="Chordata" rank="species" species="caudilinea">caudilinea</taxonomicName>
|
|
and 17-21 for other species treated here and Hoese (1986) reported counts of 15 to 19 pectoral rays for Indian Ocean species of
|
|
<taxonomicName id="4B770751818A6E62DD955BCDEE4B1FA9" LSID-ZBK="urn:lsid:zoobank.org:act:D5E53DC5-5480-4F01-9C4C-2D9BF239B442" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Hetereleotris Bleeker 1874:306" order="Perciformes" pageNumber="4" phylum="Chordata" rank="genus">Hetereleotris</taxonomicName>
|
|
;
|
|
</paragraph>
|
|
<paragraph id="1509EA25DC0376A3B76F604F8408E66E" pageNumber="4">
|
|
11 pelvic rays I,5 - common to most gobioid fishes, but some species of
|
|
<taxonomicName id="C7F40AF26F032E56677918D942920722" LSID-ZBK="urn:lsid:zoobank.org:act:D5E53DC5-5480-4F01-9C4C-2D9BF239B442" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Hetereleotris Bleeker 1874:306" order="Perciformes" pageNumber="4" phylum="Chordata" rank="genus">Hetereleotris</taxonomicName>
|
|
have I,4;
|
|
</paragraph>
|
|
<paragraph id="4AA516F681D4E507B6785E4B778C8ACC" pageNumber="4">
|
|
12 fins completely separate, without frenum - common to most
|
|
<taxonomicName id="C2164550F2BB86BFF2D7C698F8CE5676" LSID-ZBK="urn:lsid:zoobank.org:act:D5E53DC5-5480-4F01-9C4C-2D9BF239B442" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Hetereleotris Bleeker 1874:306" order="Perciformes" pageNumber="4" phylum="Chordata" rank="genus">Hetereleotris</taxonomicName>
|
|
;
|
|
</paragraph>
|
|
<paragraph id="25CFF6DB12C8DA8E205D14A36509F7D5" pageNumber="4">
|
|
13 fifth pelvic ray unbranched and clearly shorter than fourth ray - we find the fifth ray sometimes with a branch at the tip, the shorter fifth ray is common to all
|
|
<taxonomicName id="0F823443538CBF9AB3F98AD60ECC3D04" LSID-ZBK="urn:lsid:zoobank.org:act:D5E53DC5-5480-4F01-9C4C-2D9BF239B442" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Hetereleotris Bleeker 1874:306" order="Perciformes" pageNumber="4" phylum="Chordata" rank="genus">Hetereleotris</taxonomicName>
|
|
that have the fifth ray;
|
|
</paragraph>
|
|
<paragraph id="F759236307A911BA2CC3597FF2EB6037" pageNumber="4">
|
|
14 scales ctenoid, except a few anteriorly and on abdomen - common to numerous gobioid genera and some species of
|
|
<taxonomicName id="4DDACCE5B7541E8B1B0FA1F85B4ED078" LSID-ZBK="urn:lsid:zoobank.org:act:D5E53DC5-5480-4F01-9C4C-2D9BF239B442" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Hetereleotris Bleeker 1874:306" order="Perciformes" pageNumber="4" phylum="Chordata" rank="genus">Hetereleotris</taxonomicName>
|
|
;
|
|
</paragraph>
|
|
<paragraph id="2477CB26C9AFD41266B7C7890F1FE069" pageNumber="4">
|
|
15 scales 24-29 in longitudinal series - we find 23-26 for
|
|
<taxonomicName id="55C1BB5F841111D9E57307153165E403" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="4" phylum="Chordata" rank="species" species="caudilinea">H. caudilinea</taxonomicName>
|
|
and 22-26 for all species studied here,
|
|
<taxonomicName id="5E3D8A13A027C595926F8E842EF6865F" LSID-ZBK="urn:lsid:zoobank.org:act:D5E53DC5-5480-4F01-9C4C-2D9BF239B442" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Hetereleotris Bleeker 1874:306" order="Perciformes" pageNumber="4" phylum="Chordata" rank="genus">Hetereleotris</taxonomicName>
|
|
species vary from no scales to 52 in the longitudinal series;
|
|
</paragraph>
|
|
<paragraph id="56DDF3BDFA07BC55F7E98CFF5CDA6086" pageNumber="4">
|
|
16 no scales on head, nape, prepectoral area or chest - common to numerous gobiine genera and all other species of
|
|
<taxonomicName id="C222D2C7171551289C0EF4DB28081A39" LSID-ZBK="urn:lsid:zoobank.org:act:D5E53DC5-5480-4F01-9C4C-2D9BF239B442" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Hetereleotris Bleeker 1874:306" order="Perciformes" pageNumber="4" phylum="Chordata" rank="genus">Hetereleotris</taxonomicName>
|
|
.
|
|
</paragraph>
|
|
<paragraph id="1CD813C086E0625581E3E85CF78D9DCB" pageNumber="4">
|
|
Consequently only one of the characters reported appears to be consistent for the south Pacific species (no elevated posterior nasal tube). Evidence is presented below for other specialisations, but because of the variation of characteristics in
|
|
<taxonomicName id="638268DC5ED2360B0F640A31221427F4" LSID-ZBK="urn:lsid:zoobank.org:act:D5E53DC5-5480-4F01-9C4C-2D9BF239B442" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Hetereleotris Bleeker 1874:306" order="Perciformes" pageNumber="4" phylum="Chordata" rank="genus">Hetereleotris</taxonomicName>
|
|
we do not consider that the Pacific species warrant generic separation. The genus needs to be thoroughly revised to take into account the range of characters and species.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |