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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.756.24397" ID-GBIF-Dataset="4f06fa64-12e5-416c-8261-d8e1b330e8cb" ID-PMC="PMC5956031" ID-Pensoft-Pub="1313-2970-756-1" ID-PubMed="29773959" ID-ZBK="83CE3672A4E14990A54C5D712D09974E" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1313-2970-756-1" ModsDocOrigin="ZooKeys 756" ModsDocTitle="Conservation systematics of the shield-backed trapdoor spiders of the nigrum-group (Mygalomorphae, Idiopidae, Idiosoma): integrative taxonomy reveals a diverse and threatened fauna from south-western Australia" checkinTime="1525927788324" checkinUser="pensoft" docAuthor="Rix, Michael G., Huey, Joel A., Cooper, Steven J. B., Austin, Andrew D. &amp; Harvey, Mark S." docDate="2018" docId="9E1290C72479A042287D33F335A1077A" docLanguage="en" docName="ZooKeys 756: 1-121" docOrigin="ZooKeys 756" docSource="http://dx.doi.org/10.3897/zookeys.756.24397" docTitle="Idiosoma intermedium Rix &amp; Harvey, sp. n." docType="treatment" docUuid="6EB66C6F-97ED-45E9-9821-C533120D26A7" docUuidSource="ZooBank" docVersion="5" lastPageNumber="43" masterDocId="F86B8B0DFFFA8657FFF5FFC4FFCDFFAD" masterDocTitle="Conservation systematics of the shield-backed trapdoor spiders of the nigrum-group (Mygalomorphae, Idiopidae, Idiosoma): integrative taxonomy reveals a diverse and threatened fauna from south-western Australia" masterLastPageNumber="121" masterPageNumber="1" pageNumber="40" updateTime="1668165771019" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Conservation systematics of the shield-backed trapdoor spiders of the nigrum-group (Mygalomorphae, Idiopidae, Idiosoma): integrative taxonomy reveals a diverse and threatened fauna from south-western Australia</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Rix, Michael G.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Huey, Joel A.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Cooper, Steven J. B.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Austin, Andrew D.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Harvey, Mark S.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2018</mods:date>
<mods:detail type="volume">
<mods:number>756</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>121</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.756.24397</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.756.24397</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-756-1</mods:identifier>
<mods:identifier type="ZBK">83CE3672A4E14990A54C5D712D09974E</mods:identifier>
<mods:identifier type="ZooBank">83CE3672A4E14990A54C5D712D09974E</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="143929132" LSID="urn:lsid:zoobank.org:act:6EB66C6F-97ED-45E9-9821-C533120D26A7" httpUri="http://treatment.plazi.org/id/9E1290C72479A042287D33F335A1077A" lastPageId="43" lastPageNumber="43" pageId="39" pageNumber="40">
<subSubSection pageId="39" pageNumber="40" type="nomenclature">
<paragraph pageId="39" pageNumber="40">
<taxonomicName LSID="http://zoobank.org/6EB66C6F-97ED-45E9-9821-C533120D26A7" authority="Rix &amp; Harvey" class="Arachnida" family="Idiopidae" genus="Idiosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Idiosoma intermedium" order="Araneae" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="intermedium">Idiosoma intermedium Rix &amp; Harvey</taxonomicName>
<taxonomicNameLabel pageId="39" pageNumber="40">sp. n.</taxonomicNameLabel>
Figs 25, 206-215, 216-218, 219-227, 376
</paragraph>
</subSubSection>
<subSubSection pageId="40" pageNumber="41" type="type material">
<paragraph pageId="40" pageNumber="41">
<pageBreakToken pageId="40" pageNumber="41" start="start">Type</pageBreakToken>
material.
</paragraph>
<paragraph pageId="40" pageNumber="41">
Holotype male. Bodallin (IBRA_AVW), Western Australia, Australia,
<geoCoordinate direction="south" orientation="latitude" precision="925" value="-31.366667">31°22'S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="925" value="118.85">118°51'E</geoCoordinate>
, 26 June 1970, L.C. Birlles (WAM T139520).
</paragraph>
</subSubSection>
<subSubSection pageId="40" pageNumber="41" type="materials_examined">
<paragraph pageId="40" pageNumber="41">Other material examined.</paragraph>
<paragraph pageId="40" pageNumber="41">
AUSTRALIA: Western Australia: 1 ♂, Billiburning Rock (IBRA_COO),
<geoCoordinate direction="south" orientation="latitude" precision="925" value="-30.166666">30°10'S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="925" value="117.916664">117°55'E</geoCoordinate>
, 19 May 1985, B.Y. Main (WAM T139494); 1 ♀, same data (WAM T144851); ♀, same data (WAM T144854); 1 ♀, ca. 80 km NE. of Bullfinch, north of J5 (IBRA_COO),
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-30.305555">30°18'20&quot;S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="15" value="119.60166">119°36'06&quot;E</geoCoordinate>
, hand collected, 20 November 2015, M.K. Curran, D. Harms (WAM T140940DNA_Voucher_NCB_005); 1 juvenile, same data except NE. of J5,
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-30.25639">30°15'23&quot;S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="15" value="119.40083">119°24'03&quot;E</geoCoordinate>
(WAM T140939); 1 juvenile, same data except J5 deposit,
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-30.365278">30°21'55&quot;S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="15" value="119.612495">119°36'45&quot;E</geoCoordinate>
(WAM T140938); 1 ♀, Bungalbin Hill, ca. 48.2 km NNE. of Koolyanobbing (IBRA_COO),
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-30.4975">30°29'51&quot;S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="15" value="119.599724">119°35'59&quot;E</geoCoordinate>
, 9-17 October 2012, N. Dight (WAM T127934); 1 ♀, 5.5 km SE. of Koolyanobbing (IBRA_COO),
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-30.851667">30°51'06&quot;S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="15" value="119.56195">119°33'43&quot;E</geoCoordinate>
, dug from burrow, 23 August 2009, R. Teale (WAM T99749DNA_Voucher_133); 1 juvenile, Mt Manning area, site CR4 (IBRA_COO),
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-30.48139">30°28'53&quot;S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="15" value="119.99528">119°59'43&quot;E</geoCoordinate>
, open tall eucalypt woodland with mixed shrubs, 20 June 2008, J, Francesconi et al. (WAM T92079DNA_Voucher_310); 1 ♂, Mungarri Nature Reserve, south, site BE13 (IBRA_AVW),
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-30.348612">30°20'55&quot;S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="15" value="117.75806">117°45'29&quot;E</geoCoordinate>
, wet pitfall traps, 15 September 1998-25 October 1999, L. King, CALM Survey (WAM T139517DNA_Voucher_NCB_011); 1 ♂, Warrachuppin North Road, site MN2 (IBRA_AVW),
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-31.001667">31°00'06&quot;S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="15" value="118.69195">118°41'31&quot;E</geoCoordinate>
, wet pitfall traps, 21
<normalizedToken originalValue="May">May-</normalizedToken>
22 September 1998, N. Guthrie, CALM Survey (WAM T139519DNA_Voucher_NCB_010).
</paragraph>
</subSubSection>
<subSubSection pageId="40" pageNumber="41" type="etymology">
<paragraph pageId="40" pageNumber="41">Etymology.</paragraph>
<paragraph pageId="40" pageNumber="41">
The specific epithet is derived from the Latin intermedius (adjective: 'in
<normalizedToken originalValue="between">between'</normalizedToken>
,
<normalizedToken originalValue="intermediate">'intermediate'</normalizedToken>
; see Brown 1956), in reference to the intermediate size of the sigilla and relatively unsclerotised abdominal morphology of this species.
</paragraph>
</subSubSection>
<subSubSection lastPageId="41" lastPageNumber="42" pageId="40" pageNumber="41" type="diagnosis">
<paragraph pageId="40" pageNumber="41">Diagnosis.</paragraph>
<paragraph lastPageId="41" lastPageNumber="42" pageId="40" pageNumber="41">
<taxonomicName class="Arachnida" family="Idiopidae" genus="Idiosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Idiosoma intermedium" order="Araneae" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="intermedium">Idiosoma intermedium</taxonomicName>
is one of nine south-western Australian species in the intermedium- and sigillatum-clades which does not belong to the distinctive 'sigillate
<normalizedToken originalValue="complex">complex'</normalizedToken>
(Fig. 25); these nine species can be distinguished from those 'sigillate
<normalizedToken originalValue="complex">complex'</normalizedToken>
taxa (i.e.,
<taxonomicName lsidName="I. arenaceum" pageId="40" pageNumber="41" rank="species" species="arenaceum">I. arenaceum</taxonomicName>
,
<taxonomicName lsidName="I. clypeatum" pageId="40" pageNumber="41" rank="species" species="clypeatum">I. clypeatum</taxonomicName>
,
<taxonomicName lsidName="I. dandaragan" pageId="40" pageNumber="41" rank="species" species="dandaragan">I. dandaragan</taxonomicName>
,
<taxonomicName lsidName="I. kopejtkaorum" pageId="40" pageNumber="41" rank="species" species="kopejtkaorum">I. kopejtkaorum</taxonomicName>
,
<taxonomicName lsidName="I. kwongan" pageId="40" pageNumber="41" rank="species" species="kwongan">I. kwongan</taxonomicName>
,
<taxonomicName lsidName="I. nigrum" pageId="40" pageNumber="41" rank="species" species="nigrum">I. nigrum</taxonomicName>
and
<taxonomicName lsidName="I. schoknechtorum" pageId="40" pageNumber="41" rank="species" species="schoknechtorum">I. schoknechtorum</taxonomicName>
) by the absence of well-defined lateral sclerotic strips on the male abdomen (e.g., Figs 151, 212, 234), and by the significantly
<pageBreakToken pageId="41" pageNumber="42" start="start">less</pageBreakToken>
sclerotised morphology of the female abdomen (which may be strongly corrugate but never leathery and
<normalizedToken originalValue="shield-like">'shield-like'</normalizedToken>
) (e.g., Figs 4, 7, 8, 159, 220, 242). Males of
<taxonomicName lsidName="I. intermedium" pageId="41" pageNumber="42" rank="species" species="intermedium">I. intermedium</taxonomicName>
can be further distinguished from those of
<taxonomicName lsidName="I. gutharuka" pageId="41" pageNumber="42" rank="species" species="gutharuka">I. gutharuka</taxonomicName>
and
<taxonomicName lsidName="I. incomptum" pageId="41" pageNumber="42" rank="species" species="incomptum">I. incomptum</taxonomicName>
by the presence of enlarged (i.e., clearly visible) SP4 sclerites (Fig. 212; cf. Figs 186, 199); from
<taxonomicName lsidName="I. jarrah" pageId="41" pageNumber="42" rank="species" species="jarrah">I. jarrah</taxonomicName>
and
<taxonomicName lsidName="I. mcclementsorum" pageId="41" pageNumber="42" rank="species" species="mcclementsorum">I. mcclementsorum</taxonomicName>
by the colour of the legs, which do not have strongly contrasting bright yellow or orange-yellow femora (Fig. 213; cf. Figs 235, 292); from
<taxonomicName lsidName="I. gardneri" pageId="41" pageNumber="42" rank="species" species="gardneri">I. gardneri</taxonomicName>
and
<taxonomicName lsidName="I. sigillatum" pageId="41" pageNumber="42" rank="species" species="sigillatum">I. sigillatum</taxonomicName>
by the absence of well-defined dorso-lateral abdominal corrugations or striations (Figs 207, 212, Key pane 9.2; cf. Figs 168, 173, 352, 357, Key pane 9.1); from
<taxonomicName lsidName="I. formosum" pageId="41" pageNumber="42" rank="species" species="formosum">I. formosum</taxonomicName>
by the shape of tibia I, which is longer (with the prolateral clasping spurs occupying the distal third of the segment) (Fig. 213; cf. Fig. 152), and by the colour of the abdomen, which is more uniformly coloured dorsally (Figs 207, 212; cf. Figs 146, 151); and from
<taxonomicName lsidName="I. mcnamarai" pageId="41" pageNumber="42" rank="species" species="mcnamarai">I. mcnamarai</taxonomicName>
by the smaller SP4 sclerites (Fig. 212; cf. Fig. 313), and by the morphology of the SP3 sclerites, each of which may have a laterally or postero-laterally directed triangular
<normalizedToken originalValue="corner">'corner'</normalizedToken>
laterally (as opposed to an antero-laterally directed triangular
<normalizedToken originalValue="corner">'corner'</normalizedToken>
) (Fig. 212, Key pane 12.3; cf. Fig. 313, Key panes 12.1, 12.2).
</paragraph>
<paragraph pageId="41" pageNumber="42">
Females can be distinguished from those of
<taxonomicName lsidName="I. mcclementsorum" pageId="41" pageNumber="42" rank="species" species="mcclementsorum">I. mcclementsorum</taxonomicName>
and
<taxonomicName lsidName="I. sigillatum" pageId="41" pageNumber="42" rank="species" species="sigillatum">I. sigillatum</taxonomicName>
by the absence of reinforced, sclerotised ridges on the abdomen (Figs 220, 223; cf. Figs 299, 302, 365, 368); from
<taxonomicName lsidName="I. formosum" pageId="41" pageNumber="42" rank="species" species="formosum">I. formosum</taxonomicName>
and
<taxonomicName lsidName="I. mcnamarai" pageId="41" pageNumber="42" rank="species" species="mcnamarai">I. mcnamarai</taxonomicName>
by the size of the SP4 sclerites, which are not significantly larger than the SP2 sclerites (Fig. 223; cf. Figs 162, 324); and from
<taxonomicName lsidName="I. jarrah" pageId="41" pageNumber="42" rank="species" species="jarrah">I. jarrah</taxonomicName>
by the slightly larger size of the SP3 and SP4 sclerites (Fig. 223; cf. Fig. 245) [NB. females of
<taxonomicName lsidName="I. gardneri" pageId="41" pageNumber="42" rank="species" species="gardneri">I. gardneri</taxonomicName>
,
<taxonomicName lsidName="I. gutharuka" pageId="41" pageNumber="42" rank="species" species="gutharuka">I. gutharuka</taxonomicName>
and
<taxonomicName lsidName="I. incomptum" pageId="41" pageNumber="42" rank="species" species="incomptum">I. incomptum</taxonomicName>
are unknown].
</paragraph>
<paragraph pageId="41" pageNumber="42">
This species can also be distinguished from
<taxonomicName lsidName="I. corrugatum" pageId="41" pageNumber="42" rank="species" species="corrugatum">I. corrugatum</taxonomicName>
(from the Eyre Peninsula of South Australia) by the shape of the prolateral clasping spurs on the male tibia I, which are oriented longitudinally (Fig. 214; cf. Fig. 109), and by the shape of the female eye group, which is broadly trapezoidal (Fig. 222; cf. Fig. 117).
</paragraph>
</subSubSection>
<subSubSection lastPageId="42" lastPageNumber="43" pageId="41" pageNumber="42" type="description">
<paragraph pageId="41" pageNumber="42">Description (male holotype).</paragraph>
<paragraph lastPageId="42" lastPageNumber="43" pageId="41" pageNumber="42">
Total length 19.8. Carapace 9.7 long, 7.3 wide. Abdomen 8.2 long, 5.4 wide. Carapace (Fig. 206) dark tan, with darker ocular region; lateral margins with uniformly-spaced fringe of porrect black setae; fovea procurved. Eye group (Fig. 209) trapezoidal (anterior eye row strongly procurved), 0.6
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide,
<normalizedToken originalValue="PLEPLE/ALEALE">PLE-PLE/ALE-ALE</normalizedToken>
ratio 2.3; ALE almost contiguous; AME separated by less than their own diameter; PME separated by 5.3
<normalizedToken originalValue="×">x</normalizedToken>
their own diameter; PME and PLE separated by slightly more than diameter of PME, PME positioned in line with level of PLE. Maxillae with field of small cuspules confined to inner corner; labium without cuspules. Abdomen (Figs 207, 212) oval, charcoal-brown in dorsal view with tan mottling and assortment of stiff, porrect black setae, each with slightly raised, dark brown sclerotic base. Posterior abdomen moderately sigillate (Figs 207, 212); SP2 sclerites comma-shaped spots; SP3 sclerites circular with irregular margins, each with unsclerotised triangular
<normalizedToken originalValue="corner">'corner'</normalizedToken>
laterally; SP4 sclerites oval, each surrounded by chevron-like pad of unsclerotised cuticle laterally; SP5 obscured. Legs (Figs 213-215) variable shades of dark tan, with light scopulae on tarsi
<normalizedToken originalValue="III">I-II</normalizedToken>
; distal tibia I with pair of large prolateral clasping spurs oriented longitudinally. Leg I: femur 8.0; patella 4.1; tibia 5.7; metatarsus 6.1; tarsus 3.7; total 27.5. Leg I
<normalizedToken originalValue="femurtarsus">femur-tarsus</normalizedToken>
/carapace length
<pageBreakToken pageId="42" pageNumber="43" start="start">ratio</pageBreakToken>
2.8. Pedipalpal tibia (Figs 216-218) 2.4
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide; RTA burr-like, with conical basal protuberance and field of retroventral spinules; digital process porrect, unmodified. Cymbium (Figs 216-218) setose, with field of spinules disto-dorsally. Embolus (Figs 216-218) broadly twisted and sharply tapering distally, with prominent longitudinal flange and triangular (sub-distal) embolic apophysis.
</paragraph>
</subSubSection>
<subSubSection pageId="42" pageNumber="43" type="description">
<paragraph pageId="42" pageNumber="43">Description (female WAM T99749).</paragraph>
<paragraph pageId="42" pageNumber="43">
Total length 25.0. Carapace 10.5 long, 7.3 wide. Abdomen 11.6 long, 8.9 wide. Carapace (Fig. 219) dark tan, with darker ocular region; fovea procurved. Eye group (Fig. 222) trapezoidal (anterior eye row strongly procurved), 0.6
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide,
<normalizedToken originalValue="PLEPLE/ALEALE">PLE-PLE/ALE-ALE</normalizedToken>
ratio 2.4; ALE almost contiguous; AME separated by approximately their own diameter; PME separated by 1.9
<normalizedToken originalValue="×">x</normalizedToken>
their own diameter; PME and PLE separated by more than diameter of PME, PME positioned in line with level of PLE. Maxillae with field of cuspules confined to inner corner (Fig. 224); labium without cuspules. Abdomen (Figs 220, 223) broadly oval, dark grey-brown in dorsal view with tan mottling. Posterior abdomen moderately sigillate (Figs 220, 223); SP2 sclerites irregular, comma-shaped spots; SP3 sclerites subcircular with irregular margins, each surrounded by pad of unsclerotised cuticle; SP4 sclerites subcircular with irregular margins, each surrounded by chevron-like pad of unsclerotised cuticle laterally; SP5 obscured. Legs (Figs 225-226) variable shades of dark tan; scopulae present on tarsi and metatarsi
<normalizedToken originalValue="III">I-II</normalizedToken>
; tibia I with one stout pro-distal macroseta (broken at base) and row of seven longer retroventral macrosetae; metatarsus I with nine stout macrosetae; tarsus I with distal cluster of short macrosetae. Leg I: femur 6.4; patella 4.2; tibia 4.1; metatarsus 3.0; tarsus 2.4; total 20.1. Leg I
<normalizedToken originalValue="femurtarsus">femur-tarsus</normalizedToken>
/carapace length ratio 1.9. Pedipalp dark tan, spinose on tibia and tarsus, with thick tarsal scopula. Genitalia (Fig. 227) with pair of short spermathecae on broad
<normalizedToken originalValue="stems">'stems'</normalizedToken>
, each bearing dense field of glandular vesicles distally, and more sparsely distributed glandular field sub-distally.
</paragraph>
</subSubSection>
<subSubSection pageId="42" pageNumber="43" type="distribution">
<paragraph pageId="42" pageNumber="43">Distribution and remarks.</paragraph>
<paragraph pageId="42" pageNumber="43">
<taxonomicName class="Arachnida" family="Idiopidae" genus="Idiosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Idiosoma intermedium" order="Araneae" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="intermedium">Idiosoma intermedium</taxonomicName>
(formerly known by WAM identification code
<normalizedToken originalValue="MYG475">'MYG475'</normalizedToken>
), the nominate member of the intermedium-clade (Fig. 25), has a relatively widespread albeit poorly defined distribution in the eastern Wheatbelt and north-western Coolgardie bioregions of south-western Western Australia (Fig. 376). Its known range extends from Bodallin north to Billiburning Rock in the eastern Wheatbelt, and east to near the Helena-Aurora Range, Mount Manning, and Koolyanobbing in the Coolgardie bioregion. Like
<taxonomicName lsidName="I. jarrah" pageId="42" pageNumber="43" rank="species" species="jarrah">I. jarrah</taxonomicName>
,
<taxonomicName lsidName="I. formosum" pageId="42" pageNumber="43" rank="species" species="formosum">I. formosum</taxonomicName>
and
<taxonomicName lsidName="I. mcnamarai" pageId="42" pageNumber="43" rank="species" species="mcnamarai">I. mcnamarai</taxonomicName>
,
<taxonomicName lsidName="I. intermedium" pageId="42" pageNumber="43" rank="species" species="intermedium">I. intermedium</taxonomicName>
exhibits a transitional morphology between other species in the intermedium-clade (i.e.
<taxonomicName lsidName="I. incomptum" pageId="42" pageNumber="43" rank="species" species="incomptum">I. incomptum</taxonomicName>
and
<taxonomicName lsidName="I. gutharuka" pageId="42" pageNumber="43" rank="species" species="gutharuka">I. gutharuka</taxonomicName>
), and the more obviously phragmotic taxa in the clypeatum- and sigillatum-clades. Little is known of the biology of this species, other than that males have been collected wandering in search of females in late autumn and winter.
</paragraph>
</subSubSection>
<subSubSection lastPageId="43" lastPageNumber="44" pageId="42" pageNumber="43" type="conservation assessment">
<paragraph pageId="42" pageNumber="43">Conservation assessment.</paragraph>
<paragraph lastPageId="43" lastPageNumber="44" pageId="42" pageNumber="43">
In 2017,
<taxonomicName class="Arachnida" family="Idiopidae" genus="Idiosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Idiosoma intermedium" order="Araneae" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="intermedium">Idiosoma intermedium</taxonomicName>
was formally assessed as 'priority 3' fauna; this assessment incorporated the latest taxonomic, geographic and genetic data summarised in the current study (with a number of additional records also identified subsequently). It has a known extent of occurrence of nearly 14,500 km2 [14,102 km2] (a likely underestimate due to limited survey effort), and while it
<pageBreakToken pageId="43" pageNumber="44" start="start">therefore</pageBreakToken>
cannot be considered threatened under Criterion B, a 'priority 3' recommendation was made due to the occurrence of this species in areas prospective for mining and mineral resources. Further close assessment under both Criteria A and B is warranted in the future.
</paragraph>
</subSubSection>
</treatment>
</document>