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<document id="2B1A9B94D3228792C4AE2865E2BF1C8D" ID-CLB-Dataset="26662" ID-DOI="10.11646/zootaxa.4612.1.12" ID-GBIF-Dataset="d1f4c672-a490-4c05-8124-803b5d99a165" ID-ISSN="1175-5326" ID-Zenodo-Dep="3233559" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1559035637844" checkinUser="plazi" docAuthor="Kocian, Matúš &amp; Hlaváč, Peter" docDate="2019" docId="997A8799FFA0FFC3AEDBFF4DF20EF860" docLanguage="en" docName="zootaxa.4612.1.12.pdf" docOrigin="Zootaxa 4612 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Atanygnathus poussereaui Kocian &amp; Hlaváč 2019, sp. n." docType="treatment" docVersion="8" lastPageNumber="142" masterDocId="6543FFE1FFA1FFC2AE4CFFF0F668FFDB" masterDocTitle="The new species of the genus Atanygnathus Jacobson, 1909 (Coleoptera: Styphylinidae: Staphylininae) from Réunion Island" masterLastPageNumber="144" masterPageNumber="141" pageNumber="142" updateTime="1698732116163" updateUser="ExternalLinkService">
<mods:mods id="33C6F98793C4EA7B87B131A26B83C4FB" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="6DA01C9DF3548C92E35FB57DEE60E66E">
<mods:title id="40F28ECE4234D6B5D912C2B31D7C249F">The new species of the genus Atanygnathus Jacobson, 1909 (Coleoptera: Styphylinidae: Staphylininae) from Réunion Island</mods:title>
</mods:titleInfo>
<mods:name id="80B7B461E135C6825EA1FAF100FB0CD6" type="personal">
<mods:role id="8822773ADC467A642FC7F201F5C04D97">
<mods:roleTerm id="C00F67A0B46F973AD75A786665AC3E82">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1DEE13E0BBCEACECB29006CBCDB4BEC9">Kocian, Matúš</mods:namePart>
</mods:name>
<mods:name id="F9236268438D7342EFE2EE21176F28BD" type="personal">
<mods:role id="8702CC3F2112B4BF7255E65DA4E09950">
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<mods:namePart id="FA4717AEF2D9CFCDE38BA3E6BE4910B7">Hlaváč, Peter</mods:namePart>
</mods:name>
<mods:typeOfResource id="7B2C762288F9866425D9A70C1DDA007D">text</mods:typeOfResource>
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<mods:title id="D2CF61773D9A644A3509662117FAF461">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="5EF8CB01D921BE7B5C5788C148974319">
<mods:date id="27D65F7A846C2880C5D1413510C2DEDF">2019</mods:date>
<mods:detail id="7BFF42BC121D74D841031D909D7FBA44" type="pubDate">
<mods:number id="4296DF09012453060258AAB671CF9D8C">2019-05-28</mods:number>
</mods:detail>
<mods:detail id="7436A3541FC123CD9C63AC4B6D758B2F" type="volume">
<mods:number id="F9E9F7F2BDE1BB471C8352DD09B8DDA4">4612</mods:number>
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<mods:number id="92DFA75E5C53B0C41DA0AAC9F1AFD75E">1</mods:number>
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<mods:classification id="0800F63FD7BF47D9C1644F1FC1FC72F1">journal article</mods:classification>
<mods:identifier id="F75BC0289A12E4E1730419F534303438" type="CLB-Dataset">26662</mods:identifier>
<mods:identifier id="DEA6E98CA0E7B9197378E37E2170EB6F" type="DOI">10.11646/zootaxa.4612.1.12</mods:identifier>
<mods:identifier id="4F790240126AAF0909492728C93A8270" type="GBIF-Dataset">d1f4c672-a490-4c05-8124-803b5d99a165</mods:identifier>
<mods:identifier id="EB54A0E79AC0C29D0F120CE85C2A66E2" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="A7B8248504A4F0EEE8CD1A1D9A9B05CF" type="Zenodo-Dep">3233559</mods:identifier>
<mods:identifier id="8BBFC68E0DA4B9B26F798446E10209D4" type="ZooBank">13356FA3-ACBA-40BC-9695-9502174A5BEA</mods:identifier>
</mods:mods>
<treatment id="997A8799FFA0FFC3AEDBFF4DF20EF860" ID-DOI="http://doi.org/10.5281/zenodo.5945351" ID-GBIF-Taxon="157292278" ID-Zenodo-Dep="5945351" LSID="urn:lsid:plazi:treatment:997A8799FFA0FFC3AEDBFF4DF20EF860" httpUri="http://treatment.plazi.org/id/997A8799FFA0FFC3AEDBFF4DF20EF860" lastPageNumber="142" pageId="1" pageNumber="142">
<subSubSection id="59C96504FFA0FFC3AEDBFF4DF5E4FF0C" box="[151,908,189,216]" pageId="1" pageNumber="142" type="nomenclature">
<paragraph id="116C368FFFA0FFC3AEDBFF4DF5E4FF0C" blockId="1.[151,1437,154,1979]" box="[151,908,189,216]" pageId="1" pageNumber="142">
<heading id="4A2481E3FFA0FFC3AEDBFF4DF5E4FF0C" bold="true" box="[151,908,189,216]" fontSize="11" level="1" pageId="1" pageNumber="142" reason="1">
<emphasis id="23A7EA9DFFA0FFC3AEDBFF4DF5E4FF0C" bold="true" box="[151,908,189,216]" pageId="1" pageNumber="142">
<taxonomicName id="D6D34D0CFFA0FFC3AEDBFF4DF4D3FF03" ID-CoL="8XV6D" authority="Kocian &amp; Hlavac" authorityName="Kocian &amp; Hlaváč" authorityYear="2019" box="[151,699,189,216]" class="Insecta" family="Staphylinidae" genus="Atanygnathus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="142" phylum="Arthropoda" rank="species" species="poussereaui" status="sp. nov.">
<emphasis id="23A7EA9DFFA0FFC3AEDBFF4DF7B1FF0C" bold="true" box="[151,473,189,216]" italics="true" pageId="1" pageNumber="142">Atanygnathus poussereaui</emphasis>
Kocian &amp; Hlaváč
</taxonomicName>
,
<taxonomicNameLabel id="389457E6FFA0FFC3AC84FF4EF562FF03" box="[712,778,190,216]" pageId="1" pageNumber="142" rank="species">sp. n.</taxonomicNameLabel>
(
<figureCitation id="89E82A0AFFA0FFC3AD55FF4DF5EBFF0C" box="[793,899,189,215]" captionStart="FIGURES 1–4" captionStartId="2.[151,258,1644,1666]" captionTargetBox="[151,1436,185,1600]" captionTargetId="figure@2.[151,1436,175,1603]" captionTargetPageId="2" captionText="FIGURES 1–4. Atanygnathus poussereaui sp. n. Male sternite VIII (1); aedeagus from ventral view (2); habitus (3); type locality, Plaine des Palmisters (4)." httpUri="https://zenodo.org/record/3233561/files/figure.png" pageId="1" pageNumber="142">Figs 1–4</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="59C96504FFA0FFC3AEDBFF13F52BFC10" pageId="1" pageNumber="142" type="description">
<paragraph id="116C368FFFA0FFC3AEDBFF13F34FFE98" blockId="1.[151,1437,154,1979]" pageId="1" pageNumber="142">
<emphasis id="23A7EA9DFFA0FFC3AEDBFF13F74AFF20" bold="true" box="[151,290,227,251]" pageId="1" pageNumber="142">Description.</emphasis>
Body (
<figureCitation id="89E82A0AFFA0FFC3AF3EFF13F7DAFF20" box="[370,434,227,251]" captionStart="FIGURES 1–4" captionStartId="2.[151,258,1644,1666]" captionTargetBox="[151,1436,185,1600]" captionTargetId="figure@2.[151,1436,175,1603]" captionTargetPageId="2" captionText="FIGURES 1–4. Atanygnathus poussereaui sp. n. Male sternite VIII (1); aedeagus from ventral view (2); habitus (3); type locality, Plaine des Palmisters (4)." httpUri="https://zenodo.org/record/3233561/files/figure.png" pageId="1" pageNumber="142">Fig. 3</figureCitation>
) rufobrunneous, brunneous to piceous. Pronotal margins, scutellum, elytra around scutellum, along suture and in posterior margin paler, brownish-yellow. Abdomen uniformly rufobrunneous, only anterior part of tergites VII and VIII darker. Legs, palpi and antennae yellowish, antennomeres 2
<emphasis id="23A7EA9DFFA0FFC3ADB9FEDDF269FE99" bold="true" box="[1013,1025,301,322]" pageId="1" pageNumber="142">–</emphasis>
9 slightly darker, brownish.
</paragraph>
<paragraph id="116C368FFFA0FFC3AE8BFEBFF312FE74" blockId="1.[151,1437,154,1979]" pageId="1" pageNumber="142">
Head
<quantity id="D62B9B6AFFA0FFC3AF44FEBFF7FFFEBC" box="[264,407,335,359]" metricMagnitude="-4" metricUnit="m" metricValue="4.75" metricValueMax="5.0" metricValueMin="4.5" pageId="1" pageNumber="142" unit="mm" value="0.475" valueMax="0.5" valueMin="0.45">0,45–0,5 mm</quantity>
wide, 0,46–
<quantity id="D62B9B6AFFA0FFC3AC6CFEBFF4E9FEBC" box="[544,641,335,359]" metricMagnitude="-4" metricUnit="m" metricValue="4.6" pageId="1" pageNumber="142" unit="mm" value="0.46">0,46 mm</quantity>
long. Surface shiny, but with well perspicuous, extremely fine and shallow microsculpture formed by polygonal to slightly transverse meshes. Eyes small, approximately 1,2 times shorter than temples, formed by about 30 ommatidies. Antennae
<quantity id="D62B9B6AFFA0FFC3AC89FE67F537FE74" box="[709,863,407,431]" metricMagnitude="-3" metricUnit="m" metricValue="1.305" metricValueMax="1.36" metricValueMin="1.25" pageId="1" pageNumber="142" unit="mm" value="1.305" valueMax="1.36" valueMin="1.25">1,25–1,36 mm</quantity>
long, filiform, all antennomeres longer than wide.
</paragraph>
<paragraph id="116C368FFFA0FFC3AE8BFE4CF682FDB8" blockId="1.[151,1437,154,1979]" pageId="1" pageNumber="142">
Pronotum
<quantity id="D62B9B6AFFA0FFC3AF7BFE4BF7ADFE08" box="[311,453,443,467]" metricMagnitude="-4" metricUnit="m" metricValue="8.45" metricValueMax="9.0" metricValueMin="7.9" pageId="1" pageNumber="142" unit="mm" value="0.845" valueMax="0.9" valueMin="0.79">0,79–0,9 mm</quantity>
wide,
<quantity id="D62B9B6AFFA0FFC3AC42FE4BF4C0FE08" box="[526,680,443,467]" metricMagnitude="-4" metricUnit="m" metricValue="6.8999999999999995" metricValueMax="7.3" metricValueMin="6.5" pageId="1" pageNumber="142" unit="mm" value="0.69" valueMax="0.73" valueMin="0.65">0,65–0,73 mm</quantity>
long, in average 0,9 times as wide as long. Surface shiny, with similar microsculpture as in head. Inner pair of anterior punctures separated from pronotum margin by distance 1,5 times longer than puncture diameter. Discal area with one pair of punctures, situated almost in middle of length of pronotum or (more usually) slightly before the middle. Outer anterior, lateral and posterior punctures situated precisely on the pronotum margin.
</paragraph>
<paragraph id="116C368FFFA0FFC3AE8BFD9FF563FD28" blockId="1.[151,1437,154,1979]" pageId="1" pageNumber="142">
Elytra short, distinctly widened posteriad, broadest in posterior angles, 0,85–1.0 mm wide, in suture
<quantity id="D62B9B6AFFA0FFC3AB4DFD9FF3F4FD5C" box="[1281,1436,623,647]" metricMagnitude="-4" metricUnit="m" metricValue="3.8" metricValueMax="4.1" metricValueMin="3.5" pageId="1" pageNumber="142" unit="mm" value="0.38" valueMax="0.41" valueMin="0.35">0,35–0,41 mm</quantity>
long, slightly wider than pronotum, at suture approximately 0,6 times shorter than pronotum. Surface, including scutellum smooth, lacking microsculpture, with dense, tight and light pubescent. Distance between punctures slightly longer than puncture diameter. Wings atrophied, with visible residuals.
</paragraph>
<paragraph id="116C368FFFA0FFC3AE8BFD0FF469FC84" blockId="1.[151,1437,154,1979]" pageId="1" pageNumber="142">Abdomen at base parallel-sided, as wide or slightly wider than elytra, tapering toward apex from visible terigite V. Tergites densely puntate and pubescent. Surface with fine transverse microsculpture. Apical margin of visible tergite VII with rudiments of palisade fringe.</paragraph>
<paragraph id="116C368FFFA0FFC3AE8BFC9BF32EFC7C" blockId="1.[151,1437,154,1979]" pageId="1" pageNumber="142">
Male: Apical margin of sternite VIII slightly concave (
<figureCitation id="89E82A0AFFA0FFC3AD5FFC9BF53BFC58" box="[787,851,875,899]" captionStart="FIGURES 1–4" captionStartId="2.[151,258,1644,1666]" captionTargetBox="[151,1436,185,1600]" captionTargetId="figure@2.[151,1436,175,1603]" captionTargetPageId="2" captionText="FIGURES 1–4. Atanygnathus poussereaui sp. n. Male sternite VIII (1); aedeagus from ventral view (2); habitus (3); type locality, Plaine des Palmisters (4)." httpUri="https://zenodo.org/record/3233561/files/figure.png" pageId="1" pageNumber="142">Fig. 1</figureCitation>
). Aedeagus with broadly rounded apex with two pairs of shorter subapical setae and two pairs of longer apical setae. Structures of internal sac of aedeagus as in
<figureCitation id="89E82A0AFFA0FFC3AB4DFC7FF32AFC7C" box="[1281,1346,911,935]" captionStart="FIGURES 1–4" captionStartId="2.[151,258,1644,1666]" captionTargetBox="[151,1436,185,1600]" captionTargetId="figure@2.[151,1436,175,1603]" captionTargetPageId="2" captionText="FIGURES 1–4. Atanygnathus poussereaui sp. n. Male sternite VIII (1); aedeagus from ventral view (2); habitus (3); type locality, Plaine des Palmisters (4)." httpUri="https://zenodo.org/record/3233561/files/figure.png" pageId="1" pageNumber="142">Fig. 2</figureCitation>
.
</paragraph>
<paragraph id="116C368FFFA0FFC3AE8BFC43F52BFC10" blockId="1.[151,1437,154,1979]" box="[199,835,947,971]" pageId="1" pageNumber="142">Female: Female lacking modifications of sexual characters.</paragraph>
</subSubSection>
<subSubSection id="59C96504FFA0FFC3AE8BFC27F7D6FBEC" pageId="1" pageNumber="142" type="materials_examined">
<paragraph id="116C368FFFA0FFC3AE8BFC27F7D6FBEC" blockId="1.[151,1437,154,1979]" pageId="1" pageNumber="142">
<emphasis id="23A7EA9DFFA0FFC3AE8BFC27F707FC34" bold="true" box="[199,367,983,1007]" pageId="1" pageNumber="142">Type material.</emphasis>
<materialsCitation id="A1BB3CD2FFA0FFC3AF3BFC27F5B6FBC8" ID-GBIF-Occurrence="2244393169" collectingDate="2018-02-11" collectingDateMax="2018-02-21" collectingDateMin="2018-02-11" collectionCode="NMP" collectorName="Hlavac P. &amp; Kocian M." country="Réunion" elevation="740" latitude="-21.108196" location="Plaine des Palmisters-Pont Payet" longLatPrecision="1" longitude="55.66328" pageId="1" pageNumber="142" specimenCount="1" typeStatus="holotype">
<typeStatus id="CE68882DFFA0FFC3AF3BFC27F7B5FC34" box="[375,477,983,1007]" pageId="1" pageNumber="142" type="holotype">Holotype</typeStatus>
, male:
<collectingCountry id="69C4761FFFA0FFC3AC61FC27F4BBFC34" box="[557,723,981,1007]" name="Réunion" pageId="1" pageNumber="142">LA RÉUNION</collectingCountry>
,
<collectingDate id="7529E9A7FFA0FFC3AC93FC27F516FC34" box="[735,894,983,1007]" pageId="1" pageNumber="142" value="2018-02-11" valueMax="2018-02-21" valueMin="2018-02-11">
<date id="656D104FFFA0FFC3AC93FC27F516FC34" box="[735,894,983,1007]" pageId="1" pageNumber="142" value="2018-02-11" valueMax="2018-02-21" valueMin="2018-02-11">11.–21.II.2018</date>
</collectingDate>
, forest between
<location id="140C6054FFA0FFC3AA7EFC27F3F0FC2B" LSID="urn:lsid:plazi:treatment:997A8799FFA0FFC3AEDBFF4DF20EF860:140C6054FFA0FFC3AA7EFC27F3F0FC2B" box="[1074,1432,983,1008]" country="Réunion" latitude="-21.108196" longLatPrecision="1" longitude="55.66328" name="Plaine des Palmisters-Pont Payet" pageId="1" pageNumber="142">Plaine des Palmisters-Pont Payet</location>
, sifting,
<elevation id="9AFED1BCFFA0FFC3AEA7FC0BF743FBC8" box="[235,299,1019,1043]" metricMagnitude="2" metricUnit="m" metricValue="7.4" pageId="1" pageNumber="142" unit="m" value="740.0">
<quantity id="D62B9B6AFFA0FFC3AEA7FC0BF743FBC8" box="[235,299,1019,1043]" metricMagnitude="2" metricUnit="m" metricValue="7.4" pageId="1" pageNumber="142" unit="m" value="740.0">740m</quantity>
</elevation>
, -
<geoCoordinate id="74E75048FFA0FFC3AF72FC0BF7AAFBC8" box="[318,450,1019,1043]" degrees="21.108196" direction="south" orientation="latitude" pageId="1" pageNumber="142" precision="1" value="-21.108196">21.108196S</geoCoordinate>
;
<geoCoordinate id="74E75048FFA0FFC3AF89FC0BF424FBC8" box="[453,588,1019,1043]" degrees="55.663279" direction="east" orientation="longitude" pageId="1" pageNumber="142" precision="1" value="55.66328">55.663279E</geoCoordinate>
,
<collectorName id="BC265359FFA0FFC3AC15FC0BF4A8FBC8" box="[601,704,1019,1043]" pageId="1" pageNumber="142">Hlaváč P.</collectorName>
&amp;
<collectorName id="BC265359FFA0FFC3ACAAFC0BF533FBC8" box="[742,859,1019,1043]" pageId="1" pageNumber="142">Kocian M.</collectorName>
lgt. (
<collectionCode id="77C2AE4AFFA0FFC3ADDAFC0BF5BEFBC8" box="[918,982,1019,1043]" country="Czech Republic" httpUri="http://grbio.org/cool/fx6k-enrh" name="National Museum (Prague)" pageId="1" pageNumber="142">NMP</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="A1BB3CD2FFA0FFC3ADA6FC0CF7D2FBEC" ID-GBIF-Occurrence="2244393170" collectingDate="2018-02-11" collectingDateMax="2018-02-21" collectingDateMin="2018-02-11" collectionCode="NMP, MHNR" collectorName="Hlavac P. &amp; Kocian M." country="Réunion" elevation="740" latitude="-21.108196" location="Plaine des Palmisters-Pont Payet" longLatPrecision="1" longitude="55.66328" pageId="1" pageNumber="142" specimenCount="1" typeStatus="paratype">
<typeStatus id="CE68882DFFA0FFC3ADA6FC0CF23AFBCF" box="[1002,1106,1020,1044]" pageId="1" pageNumber="142" type="paratype">Paratypes</typeStatus>
(17), same data as HT (
<collectionCode id="77C2AE4AFFA0FFC3AB11FC0BF3F0FBC8" box="[1373,1432,1019,1043]" country="Czech Republic" httpUri="http://grbio.org/cool/fx6k-enrh" name="National Museum (Prague)" pageId="1" pageNumber="142">NMP</collectionCode>
,
<collectionCode id="77C2AE4AFFA0FFC3AEDBFBD0F682FBEC" box="[151,234,1056,1079]" pageId="1" pageNumber="142">MHNR</collectionCode>
, cJan, cBan, cKoc)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="59C96504FFA0FFC3AE8BFBB3F7AAFB80" box="[199,450,1091,1115]" pageId="1" pageNumber="142" type="distribution">
<paragraph id="116C368FFFA0FFC3AE8BFBB3F7AAFB80" blockId="1.[151,1437,154,1979]" box="[199,450,1091,1115]" pageId="1" pageNumber="142">
<emphasis id="23A7EA9DFFA0FFC3AE8BFBB3F73AFB80" bold="true" box="[199,338,1091,1115]" pageId="1" pageNumber="142">Distribution</emphasis>
:
<collectingCountry id="69C4761FFFA0FFC3AF2DFBB3F7D7FB80" box="[353,447,1091,1115]" name="Réunion" pageId="1" pageNumber="142">Réunion</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="59C96504FFA0FFC3AE8BFB97F473F8F0" pageId="1" pageNumber="142" type="etymology">
<paragraph id="116C368FFFA0FFC3AE8BFB97F54BFB78" blockId="1.[151,1437,154,1979]" pageId="1" pageNumber="142">
<emphasis id="23A7EA9DFFA0FFC3AE8BFB97F723FBA4" bold="true" box="[199,331,1127,1151]" pageId="1" pageNumber="142">Etymology:</emphasis>
The species is dedicated to Jacques Poussereau, specialist on
<taxonomicName id="D6D34D0CFFA0FFC3ADBDFB97F20EFBA4" authorityName="Linnaeus" authorityYear="1758" box="[1009,1126,1127,1151]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="142" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
of
<collectingCountry id="69C4761FFFA0FFC3AAC1FB97F28FFBA4" box="[1165,1255,1127,1151]" name="Réunion" pageId="1" pageNumber="142">Réunion</collectingCountry>
to recognize his kind help with the organization of our short trip to the island.
</paragraph>
<paragraph id="116C368FFFA0FFC3AE8BFB5FF7DDFA3C" blockId="1.[151,1437,154,1979]" pageId="1" pageNumber="142">
<emphasis id="23A7EA9DFFA0FFC3AE8BFB5FF429FB1C" bold="true" box="[199,577,1199,1223]" pageId="1" pageNumber="142">Comparative notes and remarks</emphasis>
. Both known species from
<collectingCountry id="69C4761FFFA0FFC3AD3AFB5FF5B8FB1C" box="[886,976,1199,1223]" name="Réunion" pageId="1" pageNumber="142">Réunion</collectingCountry>
island,
<taxonomicName id="D6D34D0CFFA0FFC3AA66FB5FF2E4FB1C" authorityName="Lecoq" authorityYear="1990" box="[1066,1164,1199,1223]" class="Insecta" family="Staphylinidae" genus="Atanygnathus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="142" phylum="Arthropoda" rank="species" species="gomyi">
<emphasis id="23A7EA9DFFA0FFC3AA66FB5FF2E4FB1C" box="[1066,1164,1199,1223]" italics="true" pageId="1" pageNumber="142">A. gomyi</emphasis>
</taxonomicName>
and
<taxonomicName id="D6D34D0CFFA0FFC3AA8AFB5FF330FB1C" authorityName="Lecoq" authorityYear="1990" box="[1222,1368,1199,1223]" class="Insecta" family="Staphylinidae" genus="Atanygnathus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="142" phylum="Arthropoda" rank="species" species="variegatus">
<emphasis id="23A7EA9DFFA0FFC3AA8AFB5FF330FB1C" box="[1222,1368,1199,1223]" italics="true" pageId="1" pageNumber="142">A. variegatus</emphasis>
</taxonomicName>
differ remarkably from
<taxonomicName id="D6D34D0CFFA0FFC3AF1FFB23F799FB30" authority="Kocian &amp; Hlaváč, 2019" authorityName="Kocian &amp; Hlaváč" authorityYear="2019" box="[339,497,1235,1259]" class="Insecta" family="Staphylinidae" genus="Atanygnathus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="142" phylum="Arthropoda" rank="species" species="poussereaui" status="sp. nov.">
<emphasis id="23A7EA9DFFA0FFC3AF1FFB23F799FB30" box="[339,497,1235,1259]" italics="true" pageId="1" pageNumber="142">A. poussereaui</emphasis>
</taxonomicName>
<taxonomicNameLabel id="389457E6FFA0FFC3AFB4FB24F45AFB30" box="[504,562,1236,1259]" pageId="1" pageNumber="142" rank="species">sp. n.</taxonomicNameLabel>
The differences in habitus are present especially in the colouration, form of body and wings. Both known species are conspicuously bi-coloured, with dark brunneous to piceous head and anterior part of elytra, rufotestaceous and vaguely anteriorly darkened pronotum, and rufotestaceous, sharply distinguished posterior part of elytra, the colouration of
<taxonomicName id="D6D34D0CFFA0FFC3AF87FACFF402FA8C" authority="Kocian &amp; Hlaváč, 2019" authorityName="Kocian &amp; Hlaváč" authorityYear="2019" box="[459,618,1343,1367]" class="Insecta" family="Staphylinidae" genus="Atanygnathus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="142" phylum="Arthropoda" rank="species" species="poussereaui" status="sp. nov.">
<emphasis id="23A7EA9DFFA0FFC3AF87FACFF402FA8C" box="[459,618,1343,1367]" italics="true" pageId="1" pageNumber="142">A. poussereaui</emphasis>
</taxonomicName>
<taxonomicNameLabel id="389457E6FFA0FFC3AC3EFAB0F4C5FA8C" box="[626,685,1344,1367]" pageId="1" pageNumber="142" rank="species">sp. n.</taxonomicNameLabel>
is more uniform, generally brunneous without any part of body with paler colour. Body of
<taxonomicName id="D6D34D0CFFA0FFC3AFC9FA93F44CFAA0" authority="Kocian &amp; Hlaváč, 2019" authorityName="Kocian &amp; Hlaváč" authorityYear="2019" box="[389,548,1379,1403]" class="Insecta" family="Staphylinidae" genus="Atanygnathus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="142" phylum="Arthropoda" rank="species" species="poussereaui" status="sp. nov.">
<emphasis id="23A7EA9DFFA0FFC3AFC9FA93F44CFAA0" box="[389,548,1379,1403]" italics="true" pageId="1" pageNumber="142">A. poussereaui</emphasis>
</taxonomicName>
<taxonomicNameLabel id="389457E6FFA0FFC3AC60FA94F40FFAA0" box="[556,615,1380,1403]" pageId="1" pageNumber="142" rank="species">sp. n.</taxonomicNameLabel>
is smaller and slender, elytra are in shoulders narrower and generally much shorter. The surface of elytra is more coarse, punctures are larger and slightly rasper. Abdomen of
<taxonomicName id="D6D34D0CFFA0FFC3AA84FA77F332FA44" authority="Kocian &amp; Hlaváč, 2019" authorityName="Kocian &amp; Hlaváč" authorityYear="2019" box="[1224,1370,1415,1439]" class="Insecta" family="Staphylinidae" genus="Atanygnathus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="142" phylum="Arthropoda" rank="species" species="possereaui" status="sp. nov.">
<emphasis id="23A7EA9DFFA0FFC3AA84FA77F332FA44" box="[1224,1370,1415,1439]" italics="true" pageId="1" pageNumber="142">A. possereaui</emphasis>
</taxonomicName>
<taxonomicNameLabel id="389457E6FFA0FFC3AB2EFA78F3F4FA44" box="[1378,1436,1416,1439]" pageId="1" pageNumber="142" rank="species">sp. n.</taxonomicNameLabel>
is more parallel-sided and wider basally.
<taxonomicName id="D6D34D0CFFA0FFC3AC1CFA5BF48CFA18" authority="Kocian &amp; Hlaváč, 2019" authorityName="Kocian &amp; Hlaváč" authorityYear="2019" box="[592,740,1451,1475]" class="Insecta" family="Staphylinidae" genus="Atanygnathus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="142" phylum="Arthropoda" rank="species" species="pousereaui" status="sp. nov.">
<emphasis id="23A7EA9DFFA0FFC3AC1CFA5BF48CFA18" box="[592,740,1451,1475]" italics="true" pageId="1" pageNumber="142">A. pousereaui</emphasis>
</taxonomicName>
<taxonomicNameLabel id="389457E6FFA0FFC3ACA7FA5BF540FA18" box="[747,808,1451,1475]" pageId="1" pageNumber="142" rank="species">
<emphasis id="23A7EA9DFFA0FFC3ACA7FA5BF549FA18" bold="true" box="[747,801,1451,1475]" pageId="1" pageNumber="142">sp. n</emphasis>
.
</taxonomicNameLabel>
is apterous species while
<taxonomicName id="D6D34D0CFFA0FFC3AA0CFA5BF2F7FA18" authorityName="Lecoq" authorityYear="1990" box="[1088,1183,1451,1475]" class="Insecta" family="Staphylinidae" genus="Atanygnathus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="142" phylum="Arthropoda" rank="species" species="gomyi">
<emphasis id="23A7EA9DFFA0FFC3AA0CFA5BF2F7FA18" box="[1088,1183,1451,1475]" italics="true" pageId="1" pageNumber="142">A. gomyi</emphasis>
</taxonomicName>
and
<taxonomicName id="D6D34D0CFFA0FFC3AA98FA5BF30BFA18" authorityName="Lecoq" authorityYear="1990" box="[1236,1379,1451,1475]" class="Insecta" family="Staphylinidae" genus="Atanygnathus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="142" phylum="Arthropoda" rank="species" species="variegatus">
<emphasis id="23A7EA9DFFA0FFC3AA98FA5BF30BFA18" box="[1236,1379,1451,1475]" italics="true" pageId="1" pageNumber="142">A. variegatus</emphasis>
</taxonomicName>
have normally developer wings.
</paragraph>
<paragraph id="116C368FFFA0FFC3AE8BFA03F708F940" blockId="1.[151,1437,154,1979]" pageId="1" pageNumber="142">
Clear differences are also in the shape of aedeagus. Aedeagi of
<taxonomicName id="D6D34D0CFFA0FFC3AD2FFA03F5A9F9D0" authorityName="Lecoq" authorityYear="1990" box="[867,961,1523,1547]" class="Insecta" family="Staphylinidae" genus="Atanygnathus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="142" phylum="Arthropoda" rank="species" species="gomyi">
<emphasis id="23A7EA9DFFA0FFC3AD2FFA03F5A9F9D0" box="[867,961,1523,1547]" italics="true" pageId="1" pageNumber="142">A. gomyi</emphasis>
</taxonomicName>
and
<taxonomicName id="D6D34D0CFFA0FFC3ADB8FA03F2EAF9D0" authorityName="Lecoq" authorityYear="1990" box="[1012,1154,1523,1547]" class="Insecta" family="Staphylinidae" genus="Atanygnathus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="142" phylum="Arthropoda" rank="species" species="variegatus">
<emphasis id="23A7EA9DFFA0FFC3ADB8FA03F2EAF9D0" box="[1012,1154,1523,1547]" italics="true" pageId="1" pageNumber="142">A. variegatus</emphasis>
</taxonomicName>
were illustrated by Lecoq, (1990).
<taxonomicName id="D6D34D0CFFA0FFC3AEA7F9E7F712F9F4" authorityName="Lecoq" authorityYear="1990" box="[235,378,1559,1583]" class="Insecta" family="Staphylinidae" genus="Atanygnathus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="142" phylum="Arthropoda" rank="species" species="variegatus">
<emphasis id="23A7EA9DFFA0FFC3AEA7F9E7F712F9F4" box="[235,378,1559,1583]" italics="true" pageId="1" pageNumber="142">A. variegatus</emphasis>
</taxonomicName>
has aedeagus very slender, strongly narrowed apically. On the contrary, the aedeagus of
<taxonomicName id="D6D34D0CFFA0FFC3AB6AF9E7F3ECF9F4" authorityName="Lecoq" authorityYear="1990" box="[1318,1412,1559,1583]" class="Insecta" family="Staphylinidae" genus="Atanygnathus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="142" phylum="Arthropoda" rank="species" species="gomyi">
<emphasis id="23A7EA9DFFA0FFC3AB6AF9E7F3ECF9F4" box="[1318,1412,1559,1583]" italics="true" pageId="1" pageNumber="142">A. gomyi</emphasis>
</taxonomicName>
is short, basally wide, apically with broadly rounded apex and three pairs of long apical setae. Aedeagus of
<taxonomicName id="D6D34D0CFFA0FFC3AAB2F9CBF3F4F988" authority="Kocian &amp; Hlaváč, 2019" authorityName="Kocian &amp; Hlaváč" authorityYear="2019" box="[1278,1436,1595,1619]" class="Insecta" family="Staphylinidae" genus="Atanygnathus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="142" phylum="Arthropoda" rank="species" species="poussereaui" status="sp. nov.">
<emphasis id="23A7EA9DFFA0FFC3AAB2F9CBF3F4F988" box="[1278,1436,1595,1619]" italics="true" pageId="1" pageNumber="142">A. poussereaui</emphasis>
</taxonomicName>
<taxonomicNameLabel id="389457E6FFA0FFC3AEDBF9AFF6BCF9AC" box="[151,212,1631,1655]" pageId="1" pageNumber="142" rank="species">
<emphasis id="23A7EA9DFFA0FFC3AEDBF9AFF6A5F9AC" bold="true" box="[151,205,1631,1655]" pageId="1" pageNumber="142">sp. n</emphasis>
.
</taxonomicNameLabel>
is also rounded apically, but in contrast to
<taxonomicName id="D6D34D0CFFA0FFC3ACD2F9AFF495F9AC" authorityName="Lecoq" authorityYear="1990" box="[670,765,1631,1655]" class="Insecta" family="Staphylinidae" genus="Atanygnathus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="142" phylum="Arthropoda" rank="species" species="gomyi">
<emphasis id="23A7EA9DFFA0FFC3ACD2F9AFF495F9AC" box="[670,765,1631,1655]" italics="true" pageId="1" pageNumber="142">A. gomyi</emphasis>
</taxonomicName>
is more slender, with completely different inner structures and apical pubescence.
</paragraph>
<paragraph id="116C368FFFA0FFC3AE8BF958F473F8F0" blockId="1.[151,1437,154,1979]" pageId="1" pageNumber="142">
In comparison to Himalayan species described by
<bibRefCitation id="75424B7EFFA0FFC3AC91F957F516F964" author="Smetana, A." box="[733,894,1703,1727]" pageId="1" pageNumber="142" pagination="163 - 464" refId="ref2228" refString="Smetana, A. (1988) Revision of the tribes Quediini and Atanygnathini. Part II. The Himalayan region (Coleoptera: Staphylinidae). Quaestiones Entomologicae, 24, 163 - 464." type="journal article" year="1988">Smetana, 1988</bibRefCitation>
, the male genitalia of
<taxonomicName id="D6D34D0CFFA0FFC3AA24F957F36DF964" authority="Kocian &amp; Hlaváč, 2019" authorityName="Kocian &amp; Hlaváč" authorityYear="2019" box="[1128,1285,1703,1727]" class="Insecta" family="Staphylinidae" genus="Atanygnathus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="142" phylum="Arthropoda" rank="species" species="poussereaui" status="sp. nov.">
<emphasis id="23A7EA9DFFA0FFC3AA24F957F36DF964" box="[1128,1285,1703,1727]" italics="true" pageId="1" pageNumber="142">A. poussereaui</emphasis>
</taxonomicName>
<taxonomicNameLabel id="389457E6FFA0FFC3AB47F957F32FF964" box="[1291,1351,1703,1727]" pageId="1" pageNumber="142" rank="species">
<emphasis id="23A7EA9DFFA0FFC3AB47F957F329F964" bold="true" box="[1291,1345,1703,1727]" pageId="1" pageNumber="142">sp. n</emphasis>
.
</taxonomicNameLabel>
is similar to
<taxonomicName id="D6D34D0CFFA0FFC3AE9FF93BF7BFF938" authority="Smetana, 1988" authorityName="Smetana" authorityYear="1988" box="[211,471,1739,1763]" class="Insecta" family="Staphylinidae" genus="Atanygnathus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="142" phylum="Arthropoda" rank="species" species="purba">
<emphasis id="23A7EA9DFFA0FFC3AE9FF93BF758F938" box="[211,304,1739,1763]" italics="true" pageId="1" pageNumber="142">A. purba</emphasis>
<bibRefCitation id="75424B7EFFA0FFC3AF79F93BF7BFF938" author="Smetana, A." box="[309,471,1739,1763]" pageId="1" pageNumber="142" pagination="163 - 464" refId="ref2228" refString="Smetana, A. (1988) Revision of the tribes Quediini and Atanygnathini. Part II. The Himalayan region (Coleoptera: Staphylinidae). Quaestiones Entomologicae, 24, 163 - 464." type="journal article" year="1988">Smetana, 1988</bibRefCitation>
</taxonomicName>
. Both species are similar in shape of aedeagus, similarly organised structures of internal sac of aedeagus, and theirs bodies are similarly uniformly coloured. In contrast,
<taxonomicName id="D6D34D0CFFA0FFC3AD82F91FF244F8DC" authorityName="Smetana" authorityYear="1988" box="[974,1068,1775,1799]" class="Insecta" family="Staphylinidae" genus="Atanygnathus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="142" phylum="Arthropoda" rank="species" species="purba">
<emphasis id="23A7EA9DFFA0FFC3AD82F91FF244F8DC" box="[974,1068,1775,1799]" italics="true" pageId="1" pageNumber="142">A. purba</emphasis>
</taxonomicName>
is macropterous, identically as all other Asian and Reunionian species.
</paragraph>
</subSubSection>
<subSubSection id="59C96504FFA0FFC3AE8BF8C7F20EF860" pageId="1" pageNumber="142" type="biology_ecology">
<paragraph id="116C368FFFA0FFC3AE8BF8C7F20EF860" blockId="1.[151,1437,154,1979]" pageId="1" pageNumber="142">
<emphasis id="23A7EA9DFFA0FFC3AE8BF8C7F74EF894" bold="true" box="[199,294,1847,1871]" pageId="1" pageNumber="142">Biology:</emphasis>
<taxonomicName id="D6D34D0CFFA0FFC3AF60F8C7F7A1F894" authority="Kocian &amp; Hlaváč, 2019" authorityName="Kocian &amp; Hlaváč" authorityYear="2019" box="[300,457,1847,1871]" class="Insecta" family="Staphylinidae" genus="Atanygnathus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="142" phylum="Arthropoda" rank="species" species="poussereaui" status="sp. nov.">
<emphasis id="23A7EA9DFFA0FFC3AF60F8C7F7A1F894" box="[300,457,1847,1871]" italics="true" pageId="1" pageNumber="142">A. poussereaui</emphasis>
</taxonomicName>
<taxonomicNameLabel id="389457E6FFA0FFC3AF82F8C7F462F894" box="[462,522,1847,1871]" pageId="1" pageNumber="142" rank="species">
<emphasis id="23A7EA9DFFA0FFC3AF82F8C7F46BF894" bold="true" box="[462,515,1847,1871]" pageId="1" pageNumber="142">sp. n</emphasis>
.
</taxonomicNameLabel>
was collected in unique isolated locality in central-western part of the island, in Plaine des Palmisters (
<figureCitation id="89E82A0AFFA0FFC3AF0EF8ABF7EDF8A8" box="[322,389,1883,1907]" captionStart="FIGURES 1–4" captionStartId="2.[151,258,1644,1666]" captionTargetBox="[151,1436,185,1600]" captionTargetId="figure@2.[151,1436,175,1603]" captionTargetPageId="2" captionText="FIGURES 1–4. Atanygnathus poussereaui sp. n. Male sternite VIII (1); aedeagus from ventral view (2); habitus (3); type locality, Plaine des Palmisters (4)." httpUri="https://zenodo.org/record/3233561/files/figure.png" pageId="1" pageNumber="142">Fig. 4</figureCitation>
), in medium altitude, by sifting of wet humus and leaf litter. The
<typeStatus id="CE68882DFFA0FFC3AA03F8ACF215F8AF" box="[1103,1149,1884,1908]" pageId="1" pageNumber="142">type</typeStatus>
locality is very dense rain forest situated in most rainy part of
<collectingCountry id="69C4761FFFA0FFC3AC59F88FF41BF84C" box="[533,627,1919,1943]" name="Réunion" pageId="1" pageNumber="142">Réunion</collectingCountry>
. It seems, that this species is in locality very abundant and dominant. It was found in each sifting-sample made there and occurrence of other staphylinids was minimal.
</paragraph>
</subSubSection>
</treatment>
</document>
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