311 lines
32 KiB
XML
311 lines
32 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.230.105878" ID-Pensoft-Pub="1314-2003-230-157" ID-Pensoft-UUID="2E95E2BCAF7955F8B2ABBD7F472DD9B0" ModsDocID="1314-2003-230-157" checkinTime="1691771898489" checkinUser="pensoft" docAuthor="Vigosa-Mercado, J. Luis, Delgado-Salinas, Alfonso, Alvarado Cardenas, Leonardo O. & Eguiarte, Luis E." docDate="2023" docId="948A2A637B9E5E8E800D50278004588E" docLanguage="en" docName="PhytoKeys 230: 157-256" docOrigin="PhytoKeys 230" docPubDate="2023-08-11" docSource="http://dx.doi.org/10.3897/phytokeys.230.105878" docTitle="Agrostis subrepens (Hitchc.) Hitchc., in Britton, N. Amer. Fl. 17 (7): 525. 1937." docType="treatment" docVersion="1" id="2E95E2BCAF7955F8B2ABBD7F472DD9B0" lastPageNumber="157" masterDocId="2E95E2BCAF7955F8B2ABBD7F472DD9B0" masterDocTitle="Revision of the genus Agrostis (Poaceae, Pooideae, Poeae) in Megamexico" masterLastPageNumber="256" masterPageNumber="157" pageNumber="157" updateTime="1691771898489" updateUser="pensoft">
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<mods:mods id="B5177809AFF57700482F15E0938A1C80" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="9CD7F5AB209240906ED30317C7BD9D68">
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<mods:title id="A5E4C8217685304FDBA83BB5BFE68FDE">Revision of the genus Agrostis (Poaceae, Pooideae, Poeae) in Megamexico</mods:title>
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</mods:titleInfo>
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<mods:name id="668DD68DA1BAD436FAACB8A61ED1482E" type="personal">
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<mods:namePart id="329E3FA06A74977B8C3CE924E4FEBE26">Vigosa-Mercado, J. Luis</mods:namePart>
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<mods:nameIdentifier id="F7DB98DA8F9157A3D77F57817EC5C5F5" type="ORCID">https://orcid.org/0000-0002-9999-0272</mods:nameIdentifier>
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<mods:affiliation id="254C7899845E93FA5DCDA05E31BF5244">Posgrado en Ciencias Biologicas, Universidad Nacional Autonoma de Mexico, Ciudad Universitaria, Av. Universidad 3000, Coyoacan, 04510, Cd. Mx., Mexico</mods:affiliation>
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<mods:nameIdentifier id="D4E1B9984E0B0F6817C5B39D3B8679FE" type="email">luis_vigosa@ciencias.unam.mx</mods:nameIdentifier>
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<mods:roleTerm id="015D9EB14BF3BC7FF1A1403FF8B87110">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="1A69092E5393416849B19A931330BF54">Delgado-Salinas, Alfonso</mods:namePart>
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<mods:nameIdentifier id="DF24D0E9B1972D4E67EDEB3F22B2D5FD" type="ORCID">https://orcid.org/0000-0002-9322-9968</mods:nameIdentifier>
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<mods:affiliation id="F56D7260A1574A08DA3C0CC1BB24B9B9">Posgrado en Ciencias Biologicas, Universidad Nacional Autonoma de Mexico, Ciudad Universitaria, Av. Universidad 3000, Coyoacan, 04510, Cd. Mx., Mexico</mods:affiliation>
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</mods:name>
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<mods:name id="BF29F5DEF0FC1F4C502FC1ED87807410" type="personal">
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<mods:role id="055BF5FE5020746C459E5092CB19E73C">
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<mods:roleTerm id="A7F934337C8E3A53D264CC5F55DBFA68">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="239451C6C7AD069945D25D8944A8D12E">Alvarado Cardenas, Leonardo O.</mods:namePart>
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<mods:nameIdentifier id="22F1DFFD764DDE9FADB8EF0C22487B88" type="ORCID">https://orcid.org/0000-0002-4938-8339</mods:nameIdentifier>
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<mods:affiliation id="2024DE582BF93000CA1D989960824D7F">Posgrado en Ciencias Biologicas, Universidad Nacional Autonoma de Mexico, Ciudad Universitaria, Av. Universidad 3000, Coyoacan, 04510, Cd. Mx., Mexico</mods:affiliation>
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</mods:name>
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<mods:name id="4DD6EA22EA0579D037F3C879AC66D84C" type="personal">
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<mods:roleTerm id="C682223941B85C0E442D859ECDA9093D">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="7C63BEBF90B14FC130FCDEB179888728">Eguiarte, Luis E.</mods:namePart>
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<mods:nameIdentifier id="1DA7F3F1E93F5E516DE0E615D691A747" type="ORCID">https://orcid.org/0000-0002-5906-9737</mods:nameIdentifier>
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<mods:affiliation id="0A79B1C74CEDCEE1A1A86D49C4CA351D">Posgrado en Ciencias Biologicas, Universidad Nacional Autonoma de Mexico, Ciudad Universitaria, Av. Universidad 3000, Coyoacan, 04510, Cd. Mx., Mexico</mods:affiliation>
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<mods:typeOfResource id="BDA688FA5618EACC8549E911DF4F54DB">text</mods:typeOfResource>
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<mods:titleInfo id="72777874E86F387E1A24DC10D0E3E86C">
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<mods:title id="A5F0D27981B1E15A0CD2EBA1A5658765">PhytoKeys</mods:title>
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</mods:titleInfo>
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<mods:part id="FFA76CDD5A59CAE661AEC4582BA67955">
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<mods:date id="22FC12DAF6290AA36E717B57BFA357FD">2023</mods:date>
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<mods:detail id="716A698AB23F86DCF50D5C35D053B025" type="pubDate">
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<mods:number id="CD3588D32A10B59198857DACBC744188">2023-08-11</mods:number>
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</mods:detail>
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<mods:detail id="15C2989FED83ED8B24FC1CF9507DFB47" type="volume">
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<mods:number id="75040D4DED104AE8800FD80B51754847">230</mods:number>
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</mods:detail>
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<mods:start id="9F6F681E2A900E6F9824AC0C0E35EB4A">157</mods:start>
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<mods:end id="8EA61014C7C4024866EC571F9CB82857">256</mods:end>
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<mods:url id="7FA217C72AECFD26503BBF2060AA7004">http://dx.doi.org/10.3897/phytokeys.230.105878</mods:url>
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</mods:location>
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<mods:classification id="4A1B6A96092BE41AAEC69C3FDDC2706D">journal article</mods:classification>
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<mods:identifier id="B6EA3816287A4D50916AAAAB14F4CF99" type="DOI">http://dx.doi.org/10.3897/phytokeys.230.105878</mods:identifier>
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<mods:identifier id="8C49FD5E9C72946D0C0158800FF79B65" type="Pensoft-Pub">1314-2003-230-157</mods:identifier>
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<mods:identifier id="AD744EDFBD4953459120392FD93568AB" type="Pensoft-UUID">2E95E2BCAF7955F8B2ABBD7F472DD9B0</mods:identifier>
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</mods:mods>
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<treatment id="948A2A637B9E5E8E800D50278004588E" LSID="urn:lsid:plazi:treatment:948A2A637B9E5E8E800D50278004588E" httpUri="http://treatment.plazi.org/id/948A2A637B9E5E8E800D50278004588E" lastPageNumber="157" pageId="0" pageNumber="157">
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<subSubSection id="7EC95C1583B78063DDD3DD5BF6467A71" pageId="0" pageNumber="157" type="nomenclature">
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<paragraph id="AEA2D3214F70DFC5FA41943BEFC3E7C6" pageId="0" pageNumber="157">
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17.
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<taxonomicName id="B32BA9A992ECAD8CAC9620BBC7119C01" ID-CoL="65TH4" LSID="948A2A63-7B9E-5E8E-800D-50278004588E" authority="(Hitchc.) Hitchc., in Britton, N. Amer. Fl. 17 (7): 525. 1937." authorityName="(Hitchc.) Hitchc., in Britton, N. Amer. Fl. 17 (7): 525. 1937." class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis subrepens" order="Poales" pageId="0" pageNumber="157" phylum="Tracheophyta" rank="species" species="subrepens">Agrostis subrepens (Hitchc.) Hitchc., in Britton, N. Amer. Fl. 17(7): 525. 1937.</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="29117D3F99CE6D2CD3077E7A4045F1CE" pageId="0" pageNumber="157" type="description">
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<paragraph id="762B81909829908939F8ECC10513DD0A" pageId="0" pageNumber="157">
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<figureCitation id="5EF15A6DFD665CC42F801121F941C858" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Florets of Agrostis species observed with SEM A A. bourgaei B A. calderoniae C A. capillaris D A. elliottiana E A. exarata F A. ghiesbreghtii G A. gigantea H A. hyemalis I A. laxissima J A. microphylla K A. pallens L A. perennans M A. scabra N A. stolonifera O A. subpatens P A. subrepens Q A. tolucensis R A. turrialbae S A. variabilis. Scale bars: 0.3 mm." figureDoi="10.3897/phytokeys.230.105878.figure4" httpUri="https://binary.pensoft.net/fig/891239" pageId="0" pageNumber="157">Figs 4P</figureCitation>
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<figureCitation id="2CE28826F9213E0942F632492F9BBDEC" captionStart="Figure 30" captionStartId="F30" captionText="Figure 30. Calluses of Agrostis species observed with SEM A A. scabra B A. stolonifera C A. subpatens D A. subrepens E A. tolucensis F A. turrialbae G A. variabilis. Scale bars: 50 μm." figureDoi="10.3897/phytokeys.230.105878.figure30" httpUri="https://binary.pensoft.net/fig/891265" pageId="0" pageNumber="157">, 30D</figureCitation>
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<figureCitation id="A32B65EE3DCBFB0A4402360F17A647A8" captionStart="Figure 35" captionStartId="F35" captionText="Figure 35. Agrostis subrepens A whole plant B detail of the pseudostolons C ligular area D spikelet E floret, abaxial view F floret, adaxial view. Based on Reeder et al. 3535 (MEXU). Scale bars: 3 cm (A); 1 cm (B); 1 mm (C); 0.5 mm (D); 0.3 mm (E, F)." figureDoi="10.3897/phytokeys.230.105878.figure35" httpUri="https://binary.pensoft.net/fig/891270" pageId="0" pageNumber="157">, 35</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="9429F3E36C4ADD7D2060359C22105B0F" pageId="0" pageNumber="157" type="reference_group">
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<paragraph id="030A62506740DB553028A00AC3AF3DC1" pageId="0" pageNumber="157">
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<taxonomicName id="F90A6750A806F49BB6198392CC4F4D15" class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis hyemalis" order="Poales" pageId="0" pageNumber="157" phylum="Tracheophyta" rank="species" species="hyemalis">Agrostis hyemalis</taxonomicName>
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(Walter) Britton, Sterns & Poggenb. var.
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<taxonomicName id="3A199E2E6C7ABF73CEAAF02A47FD7292" class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis hyemalis var. subrepens" order="Poales" pageId="0" pageNumber="157" phylum="Tracheophyta" rank="variety" species="hyemalis" variety="subrepens">Agrostis hyemalis subrepens</taxonomicName>
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Hitchc., U.S.D.A. Bur. Pl. Industr. Bull. 68: 44. 1905. Type: Mexico. Chihuahua: in wet places, pine plains, base of Sierra Madre Mountains, 28 Sep 1887, C.G. Pringle 1420 (holotype: US (US00131756 [image!]); isotypes: F (F-104784 [image!], F-2108718 [image!]), K (K000308371 [image!]), NY (NY-327645 [image!], NY-327646 [image!], US (US00131757 [image!])).
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</paragraph>
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</subSubSection>
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<subSubSection id="7F108CC0A01618187B093BE3AD00C012" pageId="0" pageNumber="157" type="materials_examined">
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<paragraph id="17C6BCA083982115BF569F5068D5D05B" pageId="0" pageNumber="157">
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<typeStatus id="97D664CACB7F39F5308481A7C77D04B1">Type</typeStatus>
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.
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</paragraph>
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<paragraph id="42256412E921F418D44C5059A16C18A2" pageId="0" pageNumber="157">
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Based on
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<taxonomicName id="3051FF11CE7380DA55BC12A69E9485DC" class="Liliopsida" family="Gramineae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis hyemalis" order="Poales" pageId="0" pageNumber="157" phylum="Tracheophyta" rank="species" species="hyemalis">
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<emphasis id="6EB3DB1B390E9F7EF942A1CA110D5832" italics="true" pageId="0" pageNumber="157">Agrostis hyemalis</emphasis>
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</taxonomicName>
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(Walter) Britton, Sterns & Poggenb. var.
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<taxonomicName id="97A5C5973F0E35945EB1D35FA80B7769" class="Liliopsida" family="Gramineae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis hyemalis var. subrepens" order="Poales" pageId="0" pageNumber="157" phylum="Tracheophyta" rank="variety" species="hyemalis" variety="subrepens">Agrostis hyemalis subrepens</taxonomicName>
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Hitchc.
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</paragraph>
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</subSubSection>
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<subSubSection id="4438AD7C92940B167199A5BC4D763BE7" pageId="0" pageNumber="157" type="description">
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<paragraph id="A19775AE0644DEEABF2E720EB62BE7E8" pageId="0" pageNumber="157">Description.</paragraph>
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<paragraph id="A63856CA67FB7C75D7188C05348E4C97" pageId="0" pageNumber="157">
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<emphasis id="1ECDBB63AE1ECAAFA73E942F7F87D63A" bold="true" italics="true" pageId="0" pageNumber="157">Plants</emphasis>
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perennial, rhizomatous or developing pseudostolons.
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<emphasis id="97C2ED96C485B82B21FB34EA23D839D6" bold="true" italics="true" pageId="0" pageNumber="157">Tillers</emphasis>
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extravaginal, with cataphylls.
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<emphasis id="AED82A2A86C7F0D21C2A35B377118792" bold="true" italics="true" pageId="0" pageNumber="157">Rhizomes and pseudostolons</emphasis>
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up to 5 cm long.
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<emphasis id="A83C821CE1F8ADB1842A9F21017933E8" bold="true" italics="true" pageId="0" pageNumber="157">Culms</emphasis>
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0.6-1 m long, erect, decumbent at the base, nodes 2-4, glabrous, internodes glabrous.
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<emphasis id="5FA59780C7279802E8F31BDF1FE28FA2" bold="true" italics="true" pageId="0" pageNumber="157">Leaves</emphasis>
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mostly cauline; sheaths 3.5-7 cm long, usually shorter than the internodes, glabrous; ligules 1-2 mm long, longer than wide, dorsally scaberulous, apices truncate, erose; blades 3-5 cm long, 1-1.5 mm wide, linear, flat or involute, scaberulous on both surfaces.
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<emphasis id="6FE7EABC3787D4A4142B5323E71879BA" bold="true" italics="true" pageId="0" pageNumber="157">Panicles</emphasis>
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9-22 cm long, 5-10 cm wide, open, lax, pyramidal, long-exserted from the upper sheaths; branches spreading, rebranching about mid-length, scaberulous, without spikelets near their base, inferior branches 2-8 cm long; pedicels 1-3 mm long, ascending to spreading, scaberulous.
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<emphasis id="A95F3008ED8B1B768C450BA5FF83774F" bold="true" italics="true" pageId="0" pageNumber="157">Spikelets</emphasis>
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1.8-2.8 mm long, purplish; glumes subequal, lanceolate, apices acute, 1-veined, scaberulous on the keel, lower glume 1.8-2.8 mm long, upper glume 1.7-2.7 mm long; callus glabrous or with a few trichomes, Inconspicuous; lemmas 1.3-2 mm long, elliptic, apices entire, acute, 5-veined, veins prominent, unawned; paleas absent; anthers 3, 1-1.3 mm long.
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<emphasis id="34304635C7F79B962BC2401FEE0002C9" bold="true" italics="true" pageId="0" pageNumber="157">Caryopsis</emphasis>
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ca. 1.2 mm long, elliptic; endosperm solid. 2n= unknown.
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</paragraph>
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<caption id="EA81E967553E74AACC4CD1A6C7AAA848" doi="10.3897/phytokeys.230.105878.figure35" httpUri="https://binary.pensoft.net/fig/891270" pageId="0" pageNumber="157" start="Figure 35" startId="F35">
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<paragraph id="FB4FA7AF5BEAAD823A7A8E4B7EE5231D" pageId="0" pageNumber="157">
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<emphasis id="3D6D6ED552D09424B5FFA96E48469C33" bold="true" pageId="0" pageNumber="157">Figure 35.</emphasis>
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<taxonomicName id="60AF467C7955CDB6B616B95E78644974" class="Liliopsida" family="Gramineae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis subrepens" order="Poales" pageId="0" pageNumber="157" phylum="Tracheophyta" rank="species" species="subrepens">
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<emphasis id="CABC1A549D05986108EA100F99DBE97D" italics="true" pageId="0" pageNumber="157">Agrostis subrepens</emphasis>
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</taxonomicName>
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<emphasis id="8694299F5BC9411AF8038F8241423BFC" bold="true" pageId="0" pageNumber="157">A</emphasis>
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whole plant
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<emphasis id="9A8BC663D0617801ED41D7A492D931E1" bold="true" pageId="0" pageNumber="157">B</emphasis>
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detail of the pseudostolons
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<emphasis id="2D510898D4A6DE275E2115DA1D6FEDFC" bold="true" pageId="0" pageNumber="157">C</emphasis>
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ligular area
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<emphasis id="5FC1AB91222698902C09B70B23742A45" bold="true" pageId="0" pageNumber="157">D</emphasis>
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spikelet
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<emphasis id="3EEA31C3CDC9B41F1D267FA60C184508" bold="true" pageId="0" pageNumber="157">E</emphasis>
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floret, abaxial view
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<emphasis id="2E948218AA8E8CD2DC41730FE9A23D58" bold="true" pageId="0" pageNumber="157">F</emphasis>
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floret, adaxial view. Based on Reeder et al. 3535 (MEXU). Scale bars: 3 cm (
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<emphasis id="4F9A581B371C2AAADC45086C9981EE1B" bold="true" pageId="0" pageNumber="157">A</emphasis>
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); 1 cm (
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<emphasis id="2B34030815C02B96A03F97F88E58F325" bold="true" pageId="0" pageNumber="157">B</emphasis>
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); 1 mm (
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<emphasis id="3583A02499D79C0F85C2605C53A3D226" bold="true" pageId="0" pageNumber="157">C</emphasis>
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); 0.5 mm (
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<emphasis id="EB0074DFA8B4F23A34F43FAE81A9B774" bold="true" pageId="0" pageNumber="157">D</emphasis>
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); 0.3 mm (
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<emphasis id="65AB4D94078539C646C2401B27D154E6" bold="true" pageId="0" pageNumber="157">E, F</emphasis>
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).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection id="4807CE0132CF60CB86E5645EAAA63752" pageId="0" pageNumber="157" type="anatomy and micromorphology">
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<paragraph id="C54D70C10773806F74C0B483B20A355F" pageId="0" pageNumber="157">Anatomy and micromorphology.</paragraph>
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<paragraph id="5F628B60D86CEE124B4967A00B6F70BD" pageId="0" pageNumber="157">
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Leaf blades flat in transversal section; adaxial furrows deep, narrow; adaxial ribs square to triangular; keel absent; first order bundles circular in outline, sheath not interrupted, abaxial and adaxial sclerenchyma in strands; second order bundles circular in outline, sheath not interrupted, abaxial and adaxial sclerenchyma in strands; intercostal sclerenchyma absent; leaf margins with well-developed sclerenchyma caps, rounded; colorless cells absent (Fig.
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<figureCitation id="AC760CD40799DC36E5842B75530FC0F8" captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Leaf blade anatomy in transversal section of Agrostis species, in general view, and details of lateral bundles. A - C A. scabra D - E A. stolonifera F - G A. subpatens H - I A. subrepens. Scale bars: 0.1 mm." figureDoi="10.3897/phytokeys.230.105878.figure31" httpUri="https://binary.pensoft.net/fig/891266" pageId="0" pageNumber="157">31H, I</figureCitation>
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). Lemmas with transversal thickenings irregular, wider than the unthickened portion of the wall; prickle hairs abundant (Fig.
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<figureCitation id="175ED474DB6254844A30F72CCE7E4573" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Lemma surface of Agrostis observed with SEM A A. scabra B A. stolonifera C A. subpatens D A. subrepens E A. tolucensis F A. turrialbae G A. variabilis. Scale bars: 15 μm." figureDoi="10.3897/phytokeys.230.105878.figure32" httpUri="https://binary.pensoft.net/fig/891267" pageId="0" pageNumber="157">32D</figureCitation>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="17026FBF15F65E8A86BF8AA6221A4A40" pageId="0" pageNumber="157" type="distribution">
|
|
<paragraph id="CA81C576C6773915C6BDB805721AA1E7" pageId="0" pageNumber="157">Distribution and habitat.</paragraph>
|
|
<paragraph id="40E1B4F9E75CA64C422BD96F0F581451" pageId="0" pageNumber="157">
|
|
<taxonomicName id="2ABBABFD70B95B527B4E46ACCC1023DC" class="Liliopsida" family="Gramineae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis subrepens" order="Poales" pageId="0" pageNumber="157" phylum="Tracheophyta" rank="species" species="subrepens">
|
|
<emphasis id="F45DF8D7A0C96CEFB27F612F070B895B" italics="true" pageId="0" pageNumber="157">Agrostis subrepens</emphasis>
|
|
</taxonomicName>
|
|
was described from the Sierra Madre Occidental, in Chihuahua, Mexico (Fig.
|
|
<figureCitation id="81880370817EF2B26F54D77D0B07CCCD" captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Map of known geographic distribution of Agrostis species, based on herbarium specimen data A A. perennans B A. scabra C A. stolonifera D A. subpatens E A. subrepens F A. tolucensis." figureDoi="10.3897/phytokeys.230.105878.figure26" httpUri="https://binary.pensoft.net/fig/891261" pageId="0" pageNumber="157">26E</figureCitation>
|
|
). It has also been reported from South America, in Bolivia, Ecuador, Colombia, Paraguay,
|
|
<normalizedToken id="A9212E7B276C95F03011E9E04F39D5D4" originalValue="Perú">Peru</normalizedToken>
|
|
and Venezuela (
|
|
<bibRefCitation id="B6909AA8C0430E4423242C92FF68E3CF" author="Soreng, RJ" editor="Soreng, RJ" journalOrPublisher="Contributions from the United States National Herbarium" pageId="0" pageNumber="157" pagination="42 - 89" publicationUrl="https://archive.org/details/mobot39088011697067" refId="B64" refString="Soreng, RJ, Peterson, PM, 2003. Agrostis. In: Soreng, RJ, Peterson, PM, Davidse, G, Judziewicz, EJ, Zuloaga, FO, Filgueiras, TS, Morrone, O, Eds., Catalogue of New World Grasses (Poaceae): IV. Subfamily Pooideae. Contributions from the United States National Herbarium 48: 42 - 89, https://archive.org/details/mobot39088011697067" title="Agrostis." url="https://archive.org/details/mobot39088011697067" volume="48" volumeTitle="Catalogue of New World Grasses (Poaceae): IV. Subfamily Pooideae." year="2003">Soreng and Peterson 2003</bibRefCitation>
|
|
;
|
|
<bibRefCitation id="B24885CBE5E55FE2B8846A549D15AA1E" author="Idarraga-Piedrahita, A" journalOrPublisher="17 (7 )). New York Botanical Garden, New York" pageId="0" pageNumber="157" publicationUrl="https://www.tropicos.org/projectimages/Antioquia/Flora%20Antioquia%20Vol%20II.pdf" refId="B30" refString="Idarraga-Piedrahita, A, Ortiz, RC, Callejas-Posada, R, Merello, M [Eds], 2011. Listado de las plantas vasculares del departamento de Antioquia. In: Flora de Antioquia: Catalogo de las Plantas Vasculares (Vol. 2) Universidad de Antioquia, Medellin, 127-923. https://www.tropicos.org/projectimages/Antioquia/Flora%20Antioquia%20Vol%20II.pdf" title="Listado de las plantas vasculares del departamento de Antioquia. In: Flora de Antioquia: Catalogo de las Plantas Vasculares (Vol. 2) Universidad de Antioquia, Medellin, 127 - 923." url="https://www.tropicos.org/projectimages/Antioquia/Flora%20Antioquia%20Vol%20II.pdf" year="2011">
|
|
<normalizedToken id="04BF6AFD842CC1B463B1E0D2A2E9F675" originalValue="Idárraga-Piedrahita">Idarraga-Piedrahita</normalizedToken>
|
|
et al. 2011
|
|
</bibRefCitation>
|
|
), but the specimens from these countries were not seen, and could represent misidentifications of other species. In the study zone,
|
|
<taxonomicName id="AB197EA5640FA290A54FC2EB3A47013E" family="Poaceae" kingdom="Plantae" lsidName="A. subrepens" order="Poales" pageId="0" pageNumber="157" rank="species" species="subrepens">
|
|
<emphasis id="1394787D3A7B43F64F7D7937FB57E1DC" italics="true" pageId="0" pageNumber="157">A. subrepens</emphasis>
|
|
</taxonomicName>
|
|
grows in wet areas, in forests with
|
|
<taxonomicName id="DE5D24534F06B80A012470812A441298" class="Pinopsida" family="Pinaceae" genus="Pinus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Pinus" order="Pinales" pageId="0" pageNumber="157" phylum="Coniferophyta" rank="genus">
|
|
<emphasis id="995C03552E5C3093BE6D9BABF0FFC49A" italics="true" pageId="0" pageNumber="157">Pinus</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="ACF9812DB94790AF783F1B6F4B27D1BC" class="Dicotyledoneae" family="Fagaceae" genus="Quercus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Quercus" order="Fagales" pageId="0" pageNumber="157" phylum="Angiospermae" rank="genus">
|
|
<emphasis id="6239A9135B28849AA02C869E9EDF6C06" italics="true" pageId="0" pageNumber="157">Quercus</emphasis>
|
|
</taxonomicName>
|
|
, between 2000-2168 m in elevation (Fig.
|
|
<figureCitation id="7FFF6E6EAAD9ED930DAE1ED5A283D2EA" captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Elevation histograms of Agrostis species A A. perennans B A. scabra C A. stolonifera D A. subpatens E A. subrepens F A. tolucensis G A. turrialbae H A. variabilis." figureDoi="10.3897/phytokeys.230.105878.figure27" httpUri="https://binary.pensoft.net/fig/891262" pageId="0" pageNumber="157">27E</figureCitation>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="E0DF50AD32142D64CEEBF76F24039756" pageId="0" pageNumber="157" type="phenology">
|
|
<paragraph id="8C9EC3CBCEBA364EE2F4C660B261E5DB" pageId="0" pageNumber="157">Phenology.</paragraph>
|
|
<paragraph id="5BA34AB972C50A509A8244D3D4D48F79" pageId="0" pageNumber="157">
|
|
Specimens with spikelets have been collected from August to September (Fig.
|
|
<figureCitation id="47E54D7C9AF50BBBAAF703A02563FC39" captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Phenology histograms of Agrostis species A A. perennans B A. scabra C A. stolonifera D A. subpatens E A. subrepens F A. tolucensis G A. turrialbae H A. variabilis." figureDoi="10.3897/phytokeys.230.105878.figure28" httpUri="https://binary.pensoft.net/fig/891263" pageId="0" pageNumber="157">28E</figureCitation>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="CC21F2510F952425F2D6517A085FBEDF" pageId="0" pageNumber="157" type="comments">
|
|
<paragraph id="6A1319B273790457D79CB6F00F6776FF" pageId="0" pageNumber="157">Commentaries.</paragraph>
|
|
<paragraph id="DDBFF24EA5D13584CAB37C8E6D6D9DE8" pageId="0" pageNumber="157">
|
|
The status of
|
|
<taxonomicName id="951C5B46023211A00992955FBA9CC3C8" family="Poaceae" kingdom="Plantae" lsidName="A. subrepens" order="Poales" pageId="0" pageNumber="157" rank="species" species="subrepens">
|
|
<emphasis id="6D2A6A4826CE9E7AC078437E8020292D" italics="true" pageId="0" pageNumber="157">A. subrepens</emphasis>
|
|
</taxonomicName>
|
|
as a distinct species and its distribution has been put in doubt recently (
|
|
<bibRefCitation id="C47042C3E667891428A3FB6643296DBC" DOI="https://doi.org/10.3897/phytokeys.151.50538" author="Sylvester, SP" journalOrPublisher="PhytoKeys" pageId="0" pageNumber="157" pagination="107 - 160" refId="B66" refString="Sylvester, SP, Cuta-Alarcon, LE, Bravo-Pedraza, WJ, Soreng, RJ, 2020a. Agrostis and Podagrostis from paramos of Boyaca, Colombia: Synoptic taxonomy including a key to Colombian species. PhytoKeys 151: 107 - 160, DOI: https://doi.org/10.3897/phytokeys.151.50538" title="Agrostis and Podagrostis from paramos of Boyaca, Colombia: Synoptic taxonomy including a key to Colombian species." url="https://doi.org/10.3897/phytokeys.151.50538" volume="151" year="2020 a">Sylvester et al. 2020a</bibRefCitation>
|
|
). This species is very similar to
|
|
<taxonomicName id="88FAE089755BECFDE1A513B1C33E43C6" family="Poaceae" kingdom="Plantae" lsidName="A. perennans" order="Poales" pageId="0" pageNumber="157" rank="species" species="perennans">
|
|
<emphasis id="3EF6DE99BEEBAE90A422725130574D74" italics="true" pageId="0" pageNumber="157">A. perennans</emphasis>
|
|
</taxonomicName>
|
|
sensu lato and other awnless species of the study zone, but differs from them in the presence of pseudostolons, leaf blades with square to triangular adaxial ribs, and lemmas with irregular thickenings (vs. caespitose plants, leaf blades with rounded adaxial ribs, lemmas with usually polygonal thickenings). Some individuals of
|
|
<taxonomicName id="8BB9FCD64971864C18F7DA5D78E76446" family="Poaceae" kingdom="Plantae" lsidName="A. perennans" order="Poales" pageId="0" pageNumber="157" rank="species" species="perennans">
|
|
<emphasis id="E9B54FEE18A5D7C8055DDC042A29BDC5" italics="true" pageId="0" pageNumber="157">A. perennans</emphasis>
|
|
</taxonomicName>
|
|
sensu lato sometimes develop pseudostolons, but despite the leaf anatomy, there are few differences between the two taxa. We recognise
|
|
<taxonomicName id="BC15FA49AD0D8B27EA90E102459B298A" family="Poaceae" kingdom="Plantae" lsidName="A. subrepens" order="Poales" pageId="0" pageNumber="157" rank="species" species="subrepens">
|
|
<emphasis id="D219CF5174945B55056B81A344AE6E5F" italics="true" pageId="0" pageNumber="157">A. subrepens</emphasis>
|
|
</taxonomicName>
|
|
as a distinct species, until more evidence is available.
|
|
</paragraph>
|
|
<paragraph id="4677E60ABBA4FEA33A3EA5BE5695B6E7" pageId="0" pageNumber="157">
|
|
It could also be confused with
|
|
<taxonomicName id="F484BF000354EB74CE89802DEB9778F0" family="Poaceae" kingdom="Plantae" lsidName="A. pallens" order="Poales" pageId="0" pageNumber="157" rank="species" species="pallens">
|
|
<emphasis id="B1319B28495768A504AE37D8CA361C72" italics="true" pageId="0" pageNumber="157">A. pallens</emphasis>
|
|
</taxonomicName>
|
|
from California and Baja California, but it is distinguished in the more open panicles, and unawned lemmas (vs. panicles often contracted, often awned lemmas), as well as the leaf anatomy.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="FD5C83E20FBF6E09EDF90BD21274795A" pageId="0" pageNumber="157" type="conservation status">
|
|
<paragraph id="DFD6FCE2C75217686616B1A995D7773F" pageId="0" pageNumber="157">Conservation status.</paragraph>
|
|
<paragraph id="D446CEC7BE8741EAC3F429D79D4D9129" pageId="0" pageNumber="157">
|
|
<taxonomicName id="CE2CC221E8A2E7284D3B4CD744424E9D" class="Liliopsida" family="Gramineae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis subrepens" order="Poales" pageId="0" pageNumber="157" phylum="Tracheophyta" rank="species" species="subrepens">
|
|
<emphasis id="D478F0AD54F542C224C50B632CCB08F6" italics="true" pageId="0" pageNumber="157">Agrostis subrepens</emphasis>
|
|
</taxonomicName>
|
|
is known in the study zone from a few localities in Chihuahua, Mexico. It is represented by five collections, with no populations occurring in protected areas. The EOO is 74 km2 and the AOO is 12 km2. Following the IUCN criteria, the preliminary assessment category is Endangered (EN).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="E45576B15E639869ED9898954520EE29" pageId="0" pageNumber="157" type="materials_examined">
|
|
<paragraph id="A19944DCFEF80FCB961F54EBBC05041A" pageId="0" pageNumber="157">Specimens examined.</paragraph>
|
|
<paragraph id="90248EFC0662A577CC6C52422A1974A9" pageId="0" pageNumber="157">
|
|
<materialsCitation id="90D6FDB55B31E4D7834BB4AEBB38EEF3" collectingDate="1899-08-01" collectingDateMax="1960-09-18" collectingDateMin="1899-08-01" collectorName="J. Reeder, H. LeSueur, E. W. Nelson" country="Mexico" county="Municipio Casas Grandes" elevation="2000" latitude="29.9833" location="Sierra Madre Occidental" longLatPrecision="48" longitude="-108.333" municipality="Casas Grandes" specimenCount="1" stateProvince="Chihuahua">
|
|
<emphasis id="7700C16B11EA64E2D1B28EB2AF43D975" bold="true" pageId="0" pageNumber="157">
|
|
<collectingCountry id="E04630458D3BCA9FDB309090B1EE8D69" name="Mexico">Mexico</collectingCountry>
|
|
.
|
|
</emphasis>
|
|
<collectingRegion id="91409A403BF95A320AD6BAAF38BB3684" country="Mexico" name="Chihuahua">Chihuahua</collectingRegion>
|
|
:
|
|
<collectingCounty id="5018D09F2A7CFA56A0F818F830DF2920">
|
|
Municipio
|
|
<collectingMunicipality id="B6FB0B2A154DA9835641093996BAB4F1">Casas Grandes</collectingMunicipality>
|
|
</collectingCounty>
|
|
,
|
|
<location id="AA815473945D2004FB997778542D5597" LSID="urn:lsid:plazi:treatment:948A2A637B9E5E8E800D50278004588E:AA815473945D2004FB997778542D5597" country="Mexico" county="Municipio Casas Grandes" latitude="29.9833" longLatPrecision="48" longitude="-108.333" municipality="Casas Grandes" name="Sierra Madre Occidental" stateProvince="Chihuahua">Sierra Madre Occidental</location>
|
|
,
|
|
<locationDeviation id="8DB342BBCEE1F44194FE2BBBDE47E3BF" location="Casas Grandes">W of Casas Grandes</locationDeviation>
|
|
, just
|
|
<locationDeviation id="9671A7BFB62C31A1917B09730401A292" location="Hernandez">
|
|
S of
|
|
<normalizedToken id="162DCDCD9E9F12F7C1151384D82D5667" originalValue="Hernández">Hernandez</normalizedToken>
|
|
</locationDeviation>
|
|
[
|
|
<geoCoordinate id="E9585378942B5F0B93B6D94360336153" degrees="30.04186415" direction="north" orientation="latitude" precision="1" value="30.041864">30.04186415°N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="FAB59C9E611A99A73030B112D789E0E5" degrees="108.2901759" direction="west" orientation="longitude" precision="1" value="-108.29018">108.2901759°W</geoCoordinate>
|
|
],
|
|
<quantity id="3B881A817D17D48E144EC5483493FCF6" metricMagnitude="3" metricUnit="m" metricValue="2.0" unit="m" value="2000.0">
|
|
<elevation id="21F4F3AD0E91962108A7C58E511B4DD5" metricMagnitude="3" metricUnit="m" metricValue="2.0" unit="m" value="2000.0">2000 m</elevation>
|
|
</quantity>
|
|
alt.,
|
|
<collectingDate id="691132A6AD44608F8124AF61726FEBC9" value="1960-09-18">18 Sep 1960</collectingDate>
|
|
,
|
|
<collectorName id="D9BC3E9A0E36935764166BFA6625FB70">J. Reeder</collectorName>
|
|
et al. 3535 (MEXU [*, **], US).
|
|
<emphasis id="9015C56A28F9600F9AE7106A15E3CBB1" bold="true" pageId="0" pageNumber="157">Municipio Madera</emphasis>
|
|
, Chuhuichupa, [
|
|
<geoCoordinate id="72BBA5AF8C35BDCA361F37E7975790D6" degrees="29.60543842" direction="north" orientation="latitude" precision="1" value="29.605438">29.60543842°N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="C87068E8E8C070CDB0FAF21C740EA07F" degrees="108.3736947" direction="west" orientation="longitude" precision="1" value="-108.373695">108.3736947°W</geoCoordinate>
|
|
,
|
|
<quantity id="CA55D26B7590452D45CA9D7B24B8FAEB" metricMagnitude="3" metricUnit="m" metricValue="2.168" unit="m" value="2168.0">
|
|
<elevation id="A99AC5E22E8103D38CCBD1035FA66906" metricMagnitude="3" metricUnit="m" metricValue="2.168" unit="m" value="2168.0">2168 m</elevation>
|
|
</quantity>
|
|
alt.], Aug-Sep 1936,
|
|
<collectorName id="9B8232E192D42A3B08500405347958F2">H. LeSueur</collectorName>
|
|
87 (US), 198 (US), near Colonia Garcia, in the Sierra Madre, [
|
|
<geoCoordinate id="EC47C35267653A69054DD390BB14967F" degrees="29.9833" direction="north" orientation="latitude" precision="5" value="29.9833">29.9833°N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="D8C85C3D90CDE463F9F099EC9092B699" degrees="108.333" direction="west" orientation="longitude" precision="55" value="-108.333">108.333°W</geoCoordinate>
|
|
,
|
|
<quantity id="E273630A230958A6A60A658383289157" metricMagnitude="3" metricUnit="m" metricValue="2.149" unit="m" value="2149.0">
|
|
<elevation id="08D403FF3AD7E0C116B5FF83851DA537" metricMagnitude="3" metricUnit="m" metricValue="2.149" unit="m" value="2149.0">2149 m</elevation>
|
|
</quantity>
|
|
alt.],
|
|
<collectingDate id="AC7A70CB4C4C12B4CC51C6F9FB68E5E8" value="1899-08-01">1 Aug 1899</collectingDate>
|
|
,
|
|
<collectorName id="DD7B108EA716D1010EB4EFF2F9E1A660">E.W. Nelson</collectorName>
|
|
6195 (US)
|
|
</materialsCitation>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |