264 lines
42 KiB
XML
264 lines
42 KiB
XML
<document id="BBAB06F240DFAEA14AD1EBB92091E449" ID="10.11646/zootaxa.4254.2.3SLASH10792" ID-CLB-Dataset="33238" ID-DOI="10.11646/zootaxa.4254.2.3" ID-GBIF-Dataset="07060c15-139e-413a-8b41-e123e4232696" ID-ISSN="1175-5326" ID-Zenodo-Dep="545877" ID-ZooBank="88B33DE9-BCF2-4AE4-A1B4-F0D39DCDF5C3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1492068723912" checkinUser="plazi" docAuthor="Knight-Jones, Phyllis, Darbyshire, Teresa, Petersen, Mary E. & Tovar-Hernández, María Ana" docDate="2017" docId="747D7A68FFD9061AFF41F903FE29F791" docLanguage="en" docName="zootaxa.4254.2.3SLASH10792.pdf" docOrigin="Zootaxa 4254 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pseudopotamilla aspersa Kroyer 1856" docType="treatment" docVersion="7" lastPageNumber="210" masterDocId="88440210FFDE0613FFD6FFAEFFCAF171" masterDocTitle="What is Pseudopotamilla reniformis (Sabellidae)? Comparisons of populations from Britain, Iceland and Canada with comments on Eudistylia and Schizobranchia" masterLastPageNumber="220" masterPageNumber="201" pageNumber="208" updateTime="1698438879117" updateUser="ExternalLinkService">
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<mods:title id="CDB5A5E6E71F803D3C636E1FF146A490">What is Pseudopotamilla reniformis (Sabellidae)? Comparisons of populations from Britain, Iceland and Canada with comments on Eudistylia and Schizobranchia</mods:title>
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<mods:namePart id="EF51AE58D4B4F57B93F868817992A64E">Knight-Jones, Phyllis</mods:namePart>
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<mods:namePart id="928F8DB4AEFB9465EB4EE54BEC7BB3C2">Darbyshire, Teresa</mods:namePart>
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<mods:namePart id="5123D73F80233B202E69DD0BDDBF0BD0">Petersen, Mary E.</mods:namePart>
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<mods:namePart id="6838E8BE968A7A41A50B73138D6FBBE6">Tovar-Hernández, María Ana</mods:namePart>
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<mods:title id="5827D544F8C0C5AC9B8C82E75A10C472">Zootaxa</mods:title>
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<mods:date id="4AF0BC919DF0C4B53FFD5C84431AA653">2017</mods:date>
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<mods:number id="45237CFC028B1A6ADCA72AA6EBBD025D">4254</mods:number>
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<treatment id="747D7A68FFD9061AFF41F903FE29F791" ID-DOI="http://doi.org/10.5281/zenodo.6040991" ID-GBIF-Taxon="128527388" ID-Zenodo-Dep="6040991" LSID="urn:lsid:plazi:treatment:747D7A68FFD9061AFF41F903FE29F791" httpUri="http://treatment.plazi.org/id/747D7A68FFD9061AFF41F903FE29F791" lastPageId="9" lastPageNumber="210" pageId="7" pageNumber="208">
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<subSubSection id="B4CE98F5FFD90614FF41F903FF26F799" pageId="7" pageNumber="208" type="nomenclature">
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<paragraph id="FC6BCB7EFFD90614FF41F903FCC9F7B6" blockId="7.[151,771,1709,1768]" box="[151,771,1709,1735]" pageId="7" pageNumber="208">
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<heading id="A7237C12FFD90614FF41F903FCC9F7B6" bold="true" box="[151,771,1709,1735]" fontSize="11" level="1" pageId="7" pageNumber="208" reason="1">
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<emphasis id="CEA0176CFFD90614FF41F903FCC9F7B6" bold="true" box="[151,771,1709,1735]" pageId="7" pageNumber="208">
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<taxonomicName id="3BD4B0FDFFD90614FF41F903FDB5F7B6" ID-CoL="4P8TH" authority="Kroyer, 1856" authorityName="Kroyer" authorityYear="1856" box="[151,639,1709,1735]" class="Polychaeta" family="Sabellidae" genus="Pseudopotamilla" kingdom="Animalia" order="Sabellida" pageId="7" pageNumber="208" phylum="Annelida" rank="species" species="aspersa">
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<emphasis id="CEA0176CFFD90614FF41F903FE09F7B6" bold="true" box="[151,451,1709,1735]" italics="true" pageId="7" pageNumber="208">Pseudopotamilla aspersa</emphasis>
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(
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<bibRefCitation id="9845B68FFFD90614FE04F903FDBDF7B6" author="Kroyer" box="[466,631,1709,1735]" pageId="7" pageNumber="208" refString="Kroyer, H. (1856) Bidrag til Kundskab af Sabellerne. Oversigt over det Kongelige Danske videnskabernes selskabs forhandlinger, 1856, 1 - 36." type="journal article" year="1856">Krøyer, 1856</bibRefCitation>
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)
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</taxonomicName>
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reinstated
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</emphasis>
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||
</heading>
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||
</paragraph>
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<paragraph id="FC6BCB7EFFD90614FF41F961FF26F799" blockId="7.[151,771,1709,1768]" box="[151,236,1743,1768]" pageId="7" pageNumber="208">
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<emphasis id="CEA0176CFFD90614FF41F961FF6AF799" bold="true" box="[151,160,1743,1768]" pageId="7" pageNumber="208">(</emphasis>
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<figureCitation id="64EFD7FBFFD90614FF76F961FF29F799" box="[160,227,1743,1768]" captionStart="FIGURE 4" captionStartId="8.[151,250,1830,1852]" captionTargetBox="[230,1365,207,1794]" captionTargetId="figure@8.[215,1365,193,1804]" captionTargetPageId="8" captionText="FIGURE 4. Pseudopotamilla aspersa (Krøyer, 1856). A – J & P, N: Type ZMUC POL- 00523; K, L, M: lost Syntype, after Krøyer: A) ‘ whole’ worm, left crown missing, posterior damaged; B) antero-lateral detail of A; C) anterior thorax, dorsal view; D) same, ventral view; E) superior thoracic chaeta; F) inferior thoracic chaeta, front view; G) same, side view; H) side views of thoracic uncinus (left) and companion chaeta (right); J) view of companion chaeta showing teardrop-shaped ‘ blade’; K) uncinus and companion chaeta (after Krøyer); L) same, but part of torus; M) left side of crown; N) and P) two views of abdominal chaetae. Not scaled: H, J. Krøyer gave no scales for K – M." httpUri="https://zenodo.org/record/545881/files/figure.png" pageId="7" pageNumber="208">Fig. 4</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="B4CE98F5FFD90614FF41F8B6FDD2F65F" box="[151,536,1816,1838]" pageId="7" pageNumber="208" type="reference_group">
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<paragraph id="FC6BCB7EFFD90614FF41F8B6FDD2F65F" blockId="7.[151,536,1816,1838]" box="[151,536,1816,1838]" pageId="7" pageNumber="208">
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<treatmentCitationGroup id="DCC4EC50FFD90614FF41F8B6FDD2F65F" box="[151,536,1816,1838]" pageId="7" pageNumber="208">
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<taxonomicName id="3BD4B0FDFFD90614FF41F8B6FE20F65F" ID-CoL="78YT2" authority="Kroyer, 1856: 19" authorityName="Kroyer" authorityPageNumber="19" authorityYear="1856" box="[151,490,1816,1838]" class="Polychaeta" family="Sabellidae" genus="Sabella" kingdom="Animalia" order="Sabellida" pageId="7" pageNumber="208" phylum="Annelida" rank="species" species="aspersa">
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<emphasis id="CEA0176CFFD90614FF41F8B6FEF2F65F" box="[151,312,1816,1838]" italics="true" pageId="7" pageNumber="208">Sabella aspersa</emphasis>
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<treatmentCitation id="7D75ED6FFFD90614FEE8F8B7FE20F65F" author="Kroyer" box="[318,490,1816,1838]" page="19" pageId="7" pageNumber="208" year="1856">
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<bibRefCitation id="9845B68FFFD90614FEE8F8B7FE02F65F" author="Kroyer" box="[318,456,1816,1838]" pageId="7" pageNumber="208" refString="Kroyer, H. (1856) Bidrag til Kundskab af Sabellerne. Oversigt over det Kongelige Danske videnskabernes selskabs forhandlinger, 1856, 1 - 36." type="journal article" year="1856">Krøyer, 1856</bibRefCitation>
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: 19
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</treatmentCitation>
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</taxonomicName>
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–20.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="B4CE98F5FFD90614FF41F8F4FD23F6CA" pageId="7" pageNumber="208" type="materials_examined">
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<paragraph id="FC6BCB7EFFD90614FF41F8F4FD23F6CA" blockId="7.[151,1436,1882,1979]" pageId="7" pageNumber="208">
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<emphasis id="CEA0176CFFD90614FF41F8F4FE48F602" bold="true" box="[151,386,1882,1907]" pageId="7" pageNumber="208">Material examined.</emphasis>
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<typeStatus id="236F75DCFFD90614FE5CF8F5FE08F602" box="[394,450,1883,1907]" pageId="7" pageNumber="208">Type</typeStatus>
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material:
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<typeStatus id="236F75DCFFD90614FDEEF8F5FDBAF602" box="[568,624,1883,1907]" pageId="7" pageNumber="208">Type</typeStatus>
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(
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<collectionCode id="9AC553BBFFD90614FDA9F8F5FD1AF602" box="[639,720,1883,1907]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="7" pageNumber="208">ZMUC</collectionCode>
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–
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<collectionCode id="9AC553BBFFD90614FD09F8F5FCDFF602" box="[735,789,1883,1907]" pageId="7" pageNumber="208">POL</collectionCode>
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00423),
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<collectingCountry id="84C38BEEFFD90614FCAAF8F4FC39F602" box="[892,1011,1882,1907]" name="Greenland" pageId="7" pageNumber="208">Greenland</collectingCountry>
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, by default as two other
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<typeStatus id="236F75DCFFD90614FAD9F8F5FABBF602" box="[1295,1393,1883,1907]" pageId="7" pageNumber="208" type="syntype">syntypes</typeStatus>
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are missing. In the description, other data in brackets comes from unpublished drawings and notes of one of Krøyer’s missing
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<typeStatus id="236F75DCFFD90614FF23F80DFE9DF6CA" box="[245,343,1955,1979]" pageId="7" pageNumber="208" type="syntype">syntypes</typeStatus>
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(
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<collectionCode id="9AC553BBFFD90614FEB0F80DFE6BF6CA" box="[358,417,1955,1979]" pageId="7" pageNumber="208">RLC</collectionCode>
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, New Royal collection, 322).
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</paragraph>
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</subSubSection>
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||
<caption id="A8AB9BF6FFD6061BFF41F888FC9CF6A7" httpUri="https://zenodo.org/record/545881/files/figure.png" pageId="8" pageNumber="209" targetBox="[230,1365,207,1794]" targetPageId="8">
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<paragraph id="FC6BCB7EFFD6061BFF41F888FC9CF6A7" blockId="8.[151,1436,1830,2006]" pageId="8" pageNumber="209">
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<emphasis id="CEA0176CFFD6061BFF41F888FEDFF64D" bold="true" box="[151,277,1830,1852]" pageId="8" pageNumber="209">FIGURE 4.</emphasis>
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<taxonomicName id="3BD4B0FDFFD6061BFEC8F888FD0DF64D" authority="Kroyer, 1856" authorityName="Kroyer" authorityYear="1856" box="[286,711,1830,1853]" class="Polychaeta" family="Sabellidae" genus="Pseudopotamilla" kingdom="Animalia" order="Sabellida" pageId="8" pageNumber="209" phylum="Annelida" rank="species" species="aspersa">
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<emphasis id="CEA0176CFFD6061BFEC8F888FDE8F64C" box="[286,546,1830,1853]" italics="true" pageId="8" pageNumber="209">Pseudopotamilla aspersa</emphasis>
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(Krøyer, 1856)
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</taxonomicName>
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. A–J & P, N: Type ZMUC POL-00523; K, L, M: lost Syntype, after Krøyer: A) ‘whole’ worm, left crown missing, posterior damaged; B) antero-lateral detail of A; C) anterior thorax, dorsal view; D) same, ventral view; E) superior thoracic chaeta; F) inferior thoracic chaeta, front view; G) same, side view; H) side views of thoracic uncinus (left) and companion chaeta (right); J) view of companion chaeta showing teardrop-shaped ‘blade’; K) uncinus and companion chaeta (after Krøyer); L) same, but part of torus; M) left side of crown; N) and P) two views of abdominal chaetae. Not scaled: H, J. Krøyer gave no scales for K–M.
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</paragraph>
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</caption>
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<subSubSection id="B4CE98F5FFD7061AFF11FF39FE83F189" pageId="9" pageNumber="210" type="diagnosis">
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<paragraph id="FC6BCB7EFFD7061AFF11FF39FE83F189" blockId="9.[151,1437,151,1760]" pageId="9" pageNumber="210">
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<emphasis id="CEA0176CFFD7061AFF11FF39FE8BF1C1" bold="true" box="[199,321,151,176]" pageId="9" pageNumber="210">Diagnosis.</emphasis>
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Collar with low, rounded lappets dorsally; dorsal collar margin convex; peristomium slightly exposed laterally; lateral margins of collar V-shaped; handles of companion chaetae twice length of handles of adjacent uncini.
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</paragraph>
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</subSubSection>
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<subSubSection id="B4CE98F5FFD7061AFF11FEAAFBFFF2B9" pageId="9" pageNumber="210" type="description">
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||
<paragraph id="FC6BCB7EFFD7061AFF11FEAAFD25F2F1" blockId="9.[151,1437,151,1760]" pageId="9" pageNumber="210">
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<emphasis id="CEA0176CFFD7061AFF11FEAAFE93F06C" bold="true" box="[199,345,260,285]" pageId="9" pageNumber="210">Description.</emphasis>
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Body length (without radiolar crown) about 31 (48) mm and
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<quantity id="3B2C669BFFD7061AFBEBFEAAFB49F06D" box="[1085,1155,260,284]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="210" unit="mm" value="2.0">2 mm</quantity>
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wide (
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<figureCitation id="64EFD7FBFFD7061AFB0FFEAAFAEBF06D" box="[1241,1313,260,285]" captionStart="FIGURE 4" captionStartId="8.[151,250,1830,1852]" captionTargetBox="[230,1365,207,1794]" captionTargetId="figure@8.[215,1365,193,1804]" captionTargetPageId="8" captionText="FIGURE 4. Pseudopotamilla aspersa (Krøyer, 1856). A – J & P, N: Type ZMUC POL- 00523; K, L, M: lost Syntype, after Krøyer: A) ‘ whole’ worm, left crown missing, posterior damaged; B) antero-lateral detail of A; C) anterior thorax, dorsal view; D) same, ventral view; E) superior thoracic chaeta; F) inferior thoracic chaeta, front view; G) same, side view; H) side views of thoracic uncinus (left) and companion chaeta (right); J) view of companion chaeta showing teardrop-shaped ‘ blade’; K) uncinus and companion chaeta (after Krøyer); L) same, but part of torus; M) left side of crown; N) and P) two views of abdominal chaetae. Not scaled: H, J. Krøyer gave no scales for K – M." httpUri="https://zenodo.org/record/545881/files/figure.png" pageId="9" pageNumber="210">Fig. 4</figureCitation>
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A). Crown damaged, but Krøyer’s unpublished figures of a lost
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<typeStatus id="236F75DCFFD7061AFD28FE86FC9FF031" box="[766,853,296,320]" pageId="9" pageNumber="210" type="syntype">syntype</typeStatus>
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shows crown length equal to length of 8 thoracic segments and
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<figureCitation id="64EFD7FBFFD7061AFEEEFEE2FEB1F015" box="[312,379,332,357]" captionStart="FIGURE 4" captionStartId="8.[151,250,1830,1852]" captionTargetBox="[230,1365,207,1794]" captionTargetId="figure@8.[215,1365,193,1804]" captionTargetPageId="8" captionText="FIGURE 4. Pseudopotamilla aspersa (Krøyer, 1856). A – J & P, N: Type ZMUC POL- 00523; K, L, M: lost Syntype, after Krøyer: A) ‘ whole’ worm, left crown missing, posterior damaged; B) antero-lateral detail of A; C) anterior thorax, dorsal view; D) same, ventral view; E) superior thoracic chaeta; F) inferior thoracic chaeta, front view; G) same, side view; H) side views of thoracic uncinus (left) and companion chaeta (right); J) view of companion chaeta showing teardrop-shaped ‘ blade’; K) uncinus and companion chaeta (after Krøyer); L) same, but part of torus; M) left side of crown; N) and P) two views of abdominal chaetae. Not scaled: H, J. Krøyer gave no scales for K – M." httpUri="https://zenodo.org/record/545881/files/figure.png" pageId="9" pageNumber="210">Fig. 4</figureCitation>
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M (copy of Krøyer’s figure) shows a detached crown with one or two single compound eyes per radiole, except dorsal-most. Dorsal lip with bifid appearance, but difficult to see (
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<figureCitation id="64EFD7FBFFD7061AFB80FEC1FB54F0F9" box="[1110,1182,367,392]" captionStart="FIGURE 4" captionStartId="8.[151,250,1830,1852]" captionTargetBox="[230,1365,207,1794]" captionTargetId="figure@8.[215,1365,193,1804]" captionTargetPageId="8" captionText="FIGURE 4. Pseudopotamilla aspersa (Krøyer, 1856). A – J & P, N: Type ZMUC POL- 00523; K, L, M: lost Syntype, after Krøyer: A) ‘ whole’ worm, left crown missing, posterior damaged; B) antero-lateral detail of A; C) anterior thorax, dorsal view; D) same, ventral view; E) superior thoracic chaeta; F) inferior thoracic chaeta, front view; G) same, side view; H) side views of thoracic uncinus (left) and companion chaeta (right); J) view of companion chaeta showing teardrop-shaped ‘ blade’; K) uncinus and companion chaeta (after Krøyer); L) same, but part of torus; M) left side of crown; N) and P) two views of abdominal chaetae. Not scaled: H, J. Krøyer gave no scales for K – M." httpUri="https://zenodo.org/record/545881/files/figure.png" pageId="9" pageNumber="210">Figs 4</figureCitation>
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B–D). Body (damaged posteriorly) with about 60 (90) abdominal segments and 10 thoracic ones. First thoracic segment about twice length of following ones (
|
||
<figureCitation id="64EFD7FBFFD7061AFEB9FE19FE7EF0A1" box="[367,436,439,464]" captionStart="FIGURE 4" captionStartId="8.[151,250,1830,1852]" captionTargetBox="[230,1365,207,1794]" captionTargetId="figure@8.[215,1365,193,1804]" captionTargetPageId="8" captionText="FIGURE 4. Pseudopotamilla aspersa (Krøyer, 1856). A – J & P, N: Type ZMUC POL- 00523; K, L, M: lost Syntype, after Krøyer: A) ‘ whole’ worm, left crown missing, posterior damaged; B) antero-lateral detail of A; C) anterior thorax, dorsal view; D) same, ventral view; E) superior thoracic chaeta; F) inferior thoracic chaeta, front view; G) same, side view; H) side views of thoracic uncinus (left) and companion chaeta (right); J) view of companion chaeta showing teardrop-shaped ‘ blade’; K) uncinus and companion chaeta (after Krøyer); L) same, but part of torus; M) left side of crown; N) and P) two views of abdominal chaetae. Not scaled: H, J. Krøyer gave no scales for K – M." httpUri="https://zenodo.org/record/545881/files/figure.png" pageId="9" pageNumber="210">Fig. 4</figureCitation>
|
||
B). Dorsal collar margins convex with rounded, low lappets extending above junction of crown and thorax. Oblique lateral collar margins sloping posteriorly, forming each side a wide V–shaped notch showing peristomium (
|
||
<figureCitation id="64EFD7FBFFD7061AFE75FE51FE26F369" box="[419,492,511,536]" captionStart="FIGURE 4" captionStartId="8.[151,250,1830,1852]" captionTargetBox="[230,1365,207,1794]" captionTargetId="figure@8.[215,1365,193,1804]" captionTargetPageId="8" captionText="FIGURE 4. Pseudopotamilla aspersa (Krøyer, 1856). A – J & P, N: Type ZMUC POL- 00523; K, L, M: lost Syntype, after Krøyer: A) ‘ whole’ worm, left crown missing, posterior damaged; B) antero-lateral detail of A; C) anterior thorax, dorsal view; D) same, ventral view; E) superior thoracic chaeta; F) inferior thoracic chaeta, front view; G) same, side view; H) side views of thoracic uncinus (left) and companion chaeta (right); J) view of companion chaeta showing teardrop-shaped ‘ blade’; K) uncinus and companion chaeta (after Krøyer); L) same, but part of torus; M) left side of crown; N) and P) two views of abdominal chaetae. Not scaled: H, J. Krøyer gave no scales for K – M." httpUri="https://zenodo.org/record/545881/files/figure.png" pageId="9" pageNumber="210">Fig. 4</figureCitation>
|
||
B). Highest ventral collar margin distant from midline incision in front of small ventral sacs (
|
||
<figureCitation id="64EFD7FBFFD7061AFEE5FD8AFEB0F34D" box="[307,378,548,573]" captionStart="FIGURE 4" captionStartId="8.[151,250,1830,1852]" captionTargetBox="[230,1365,207,1794]" captionTargetId="figure@8.[215,1365,193,1804]" captionTargetPageId="8" captionText="FIGURE 4. Pseudopotamilla aspersa (Krøyer, 1856). A – J & P, N: Type ZMUC POL- 00523; K, L, M: lost Syntype, after Krøyer: A) ‘ whole’ worm, left crown missing, posterior damaged; B) antero-lateral detail of A; C) anterior thorax, dorsal view; D) same, ventral view; E) superior thoracic chaeta; F) inferior thoracic chaeta, front view; G) same, side view; H) side views of thoracic uncinus (left) and companion chaeta (right); J) view of companion chaeta showing teardrop-shaped ‘ blade’; K) uncinus and companion chaeta (after Krøyer); L) same, but part of torus; M) left side of crown; N) and P) two views of abdominal chaetae. Not scaled: H, J. Krøyer gave no scales for K – M." httpUri="https://zenodo.org/record/545881/files/figure.png" pageId="9" pageNumber="210">Fig. 4</figureCitation>
|
||
B–D). Anterior margin of first ventral shield indented medially, following thoracic shields rectangular, some with indistinct transverse grooves (
|
||
<figureCitation id="64EFD7FBFFD7061AFD2DFDE9FC88F311" box="[763,834,583,608]" captionStart="FIGURE 4" captionStartId="8.[151,250,1830,1852]" captionTargetBox="[230,1365,207,1794]" captionTargetId="figure@8.[215,1365,193,1804]" captionTargetPageId="8" captionText="FIGURE 4. Pseudopotamilla aspersa (Krøyer, 1856). A – J & P, N: Type ZMUC POL- 00523; K, L, M: lost Syntype, after Krøyer: A) ‘ whole’ worm, left crown missing, posterior damaged; B) antero-lateral detail of A; C) anterior thorax, dorsal view; D) same, ventral view; E) superior thoracic chaeta; F) inferior thoracic chaeta, front view; G) same, side view; H) side views of thoracic uncinus (left) and companion chaeta (right); J) view of companion chaeta showing teardrop-shaped ‘ blade’; K) uncinus and companion chaeta (after Krøyer); L) same, but part of torus; M) left side of crown; N) and P) two views of abdominal chaetae. Not scaled: H, J. Krøyer gave no scales for K – M." httpUri="https://zenodo.org/record/545881/files/figure.png" pageId="9" pageNumber="210">Fig. 4</figureCitation>
|
||
D). Thoracic tori with gap between ventral ends and lateral margins of shields (
|
||
<figureCitation id="64EFD7FBFFD7061AFE68FDC2FDC8F3F5" box="[446,514,620,645]" captionStart="FIGURE 4" captionStartId="8.[151,250,1830,1852]" captionTargetBox="[230,1365,207,1794]" captionTargetId="figure@8.[215,1365,193,1804]" captionTargetPageId="8" captionText="FIGURE 4. Pseudopotamilla aspersa (Krøyer, 1856). A – J & P, N: Type ZMUC POL- 00523; K, L, M: lost Syntype, after Krøyer: A) ‘ whole’ worm, left crown missing, posterior damaged; B) antero-lateral detail of A; C) anterior thorax, dorsal view; D) same, ventral view; E) superior thoracic chaeta; F) inferior thoracic chaeta, front view; G) same, side view; H) side views of thoracic uncinus (left) and companion chaeta (right); J) view of companion chaeta showing teardrop-shaped ‘ blade’; K) uncinus and companion chaeta (after Krøyer); L) same, but part of torus; M) left side of crown; N) and P) two views of abdominal chaetae. Not scaled: H, J. Krøyer gave no scales for K – M." httpUri="https://zenodo.org/record/545881/files/figure.png" pageId="9" pageNumber="210">Fig. 4</figureCitation>
|
||
B). Thoracic tori with about nine avicular uncini each with numerous teeth on crest covering third of main fang, and handle not much longer than length from breast to crest, but adjacent companion chaetae with handle at least twice as long (
|
||
<figureCitation id="64EFD7FBFFD7061AFD5DFD1AFD1CF3BD" box="[651,726,692,717]" captionStart="FIGURE 4" captionStartId="8.[151,250,1830,1852]" captionTargetBox="[230,1365,207,1794]" captionTargetId="figure@8.[215,1365,193,1804]" captionTargetPageId="8" captionText="FIGURE 4. Pseudopotamilla aspersa (Krøyer, 1856). A – J & P, N: Type ZMUC POL- 00523; K, L, M: lost Syntype, after Krøyer: A) ‘ whole’ worm, left crown missing, posterior damaged; B) antero-lateral detail of A; C) anterior thorax, dorsal view; D) same, ventral view; E) superior thoracic chaeta; F) inferior thoracic chaeta, front view; G) same, side view; H) side views of thoracic uncinus (left) and companion chaeta (right); J) view of companion chaeta showing teardrop-shaped ‘ blade’; K) uncinus and companion chaeta (after Krøyer); L) same, but part of torus; M) left side of crown; N) and P) two views of abdominal chaetae. Not scaled: H, J. Krøyer gave no scales for K – M." httpUri="https://zenodo.org/record/545881/files/figure.png" pageId="9" pageNumber="210">Figs 4</figureCitation>
|
||
J–L). First thoracic fascicle with two rows of broadly hooded chaetae, hood a little wider than handle. Superior notochaetae of other thoracic fascicles narrowly hooded (about 5), most longer than inferior ones that are paleate (about 17) with a distal mucro (
|
||
<figureCitation id="64EFD7FBFFD7061AFBB5FD52FB60F265" box="[1123,1194,764,789]" captionStart="FIGURE 4" captionStartId="8.[151,250,1830,1852]" captionTargetBox="[230,1365,207,1794]" captionTargetId="figure@8.[215,1365,193,1804]" captionTargetPageId="8" captionText="FIGURE 4. Pseudopotamilla aspersa (Krøyer, 1856). A – J & P, N: Type ZMUC POL- 00523; K, L, M: lost Syntype, after Krøyer: A) ‘ whole’ worm, left crown missing, posterior damaged; B) antero-lateral detail of A; C) anterior thorax, dorsal view; D) same, ventral view; E) superior thoracic chaeta; F) inferior thoracic chaeta, front view; G) same, side view; H) side views of thoracic uncinus (left) and companion chaeta (right); J) view of companion chaeta showing teardrop-shaped ‘ blade’; K) uncinus and companion chaeta (after Krøyer); L) same, but part of torus; M) left side of crown; N) and P) two views of abdominal chaetae. Not scaled: H, J. Krøyer gave no scales for K – M." httpUri="https://zenodo.org/record/545881/files/figure.png" pageId="9" pageNumber="210">Fig. 4</figureCitation>
|
||
F–G). All abdominal chaetae elongate broadly hooded, each with hood twice width of handle (
|
||
<figureCitation id="64EFD7FBFFD7061AFC37FCB1FBEDF249" box="[993,1063,799,824]" captionStart="FIGURE 4" captionStartId="8.[151,250,1830,1852]" captionTargetBox="[230,1365,207,1794]" captionTargetId="figure@8.[215,1365,193,1804]" captionTargetPageId="8" captionText="FIGURE 4. Pseudopotamilla aspersa (Krøyer, 1856). A – J & P, N: Type ZMUC POL- 00523; K, L, M: lost Syntype, after Krøyer: A) ‘ whole’ worm, left crown missing, posterior damaged; B) antero-lateral detail of A; C) anterior thorax, dorsal view; D) same, ventral view; E) superior thoracic chaeta; F) inferior thoracic chaeta, front view; G) same, side view; H) side views of thoracic uncinus (left) and companion chaeta (right); J) view of companion chaeta showing teardrop-shaped ‘ blade’; K) uncinus and companion chaeta (after Krøyer); L) same, but part of torus; M) left side of crown; N) and P) two views of abdominal chaetae. Not scaled: H, J. Krøyer gave no scales for K – M." httpUri="https://zenodo.org/record/545881/files/figure.png" pageId="9" pageNumber="210">Fig. 4</figureCitation>
|
||
N, P). Abdominal uncini smaller than those of thorax (shorter handles,
|
||
<figureCitation id="64EFD7FBFFD7061AFD83FCEAFD54F22D" box="[597,670,836,861]" captionStart="FIGURE 4" captionStartId="8.[151,250,1830,1852]" captionTargetBox="[230,1365,207,1794]" captionTargetId="figure@8.[215,1365,193,1804]" captionTargetPageId="8" captionText="FIGURE 4. Pseudopotamilla aspersa (Krøyer, 1856). A – J & P, N: Type ZMUC POL- 00523; K, L, M: lost Syntype, after Krøyer: A) ‘ whole’ worm, left crown missing, posterior damaged; B) antero-lateral detail of A; C) anterior thorax, dorsal view; D) same, ventral view; E) superior thoracic chaeta; F) inferior thoracic chaeta, front view; G) same, side view; H) side views of thoracic uncinus (left) and companion chaeta (right); J) view of companion chaeta showing teardrop-shaped ‘ blade’; K) uncinus and companion chaeta (after Krøyer); L) same, but part of torus; M) left side of crown; N) and P) two views of abdominal chaetae. Not scaled: H, J. Krøyer gave no scales for K – M." httpUri="https://zenodo.org/record/545881/files/figure.png" pageId="9" pageNumber="210">Figs 4</figureCitation>
|
||
K–L). Pygidium bilobed. Tube “leathery, tenacious, rather pale or yellowish, covered with sand or bare” (
|
||
<bibRefCitation id="9845B68FFFD7061AFD98FCC6FD29F2F1" author="Kroyer" box="[590,739,871,896]" pageId="9" pageNumber="210" refString="Kroyer, H. (1856) Bidrag til Kundskab af Sabellerne. Oversigt over det Kongelige Danske videnskabernes selskabs forhandlinger, 1856, 1 - 36." type="journal article" year="1856">Krøyer 1856</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="FC6BCB7EFFD7061AFF11FC22FBFFF2B9" blockId="9.[151,1437,151,1760]" pageId="9" pageNumber="210">
|
||
<emphasis id="CEA0176CFFD7061AFF11FC22FED5F2D4" bold="true" box="[199,287,908,933]" pageId="9" pageNumber="210">Colour.</emphasis>
|
||
“Yellow or orange with the lateral margins of the first four to ten segments purple” “purple spots on paler radioles (with a single eyespot or two.)” (
|
||
<bibRefCitation id="9845B68FFFD7061AFD73FC1EFCF5F2B9" author="Kroyer" box="[677,831,943,968]" pageId="9" pageNumber="210" refString="Kroyer, H. (1856) Bidrag til Kundskab af Sabellerne. Oversigt over det Kongelige Danske videnskabernes selskabs forhandlinger, 1856, 1 - 36." type="journal article" year="1856">Krøyer, 1856</bibRefCitation>
|
||
). Preserved
|
||
<typeStatus id="236F75DCFFD7061AFC1EFC1EFC32F2B9" box="[968,1016,944,968]" pageId="9" pageNumber="210">type</typeStatus>
|
||
pale.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B4CE98F5FFD7061AFF11FC7AFE80F7CD" pageId="9" pageNumber="210" type="discussion">
|
||
<paragraph id="FC6BCB7EFFD7061AFF11FC7AFC04F50C" blockId="9.[151,1437,151,1760]" pageId="9" pageNumber="210">
|
||
<emphasis id="CEA0176CFFD7061AFF11FC7AFEF1F29C" bold="true" box="[199,315,980,1005]" pageId="9" pageNumber="210">Remarks.</emphasis>
|
||
The type is one of three specimens of
|
||
<taxonomicName id="3BD4B0FDFFD7061AFCDAFC7AFC08F29D" box="[780,962,980,1005]" class="Polychaeta" family="Sabellidae" genus="Sabella" kingdom="Animalia" order="Sabellida" pageId="9" pageNumber="210" phylum="Annelida" rank="species" species="aspersa">
|
||
<emphasis id="CEA0176CFFD7061AFCDAFC7AFC08F29D" box="[780,962,980,1005]" italics="true" pageId="9" pageNumber="210">Sabella aspersa</emphasis>
|
||
</taxonomicName>
|
||
listed in Krøyer’s catalogue of sabellids (RLC). Some of his material was passed on to the Zoological Museum of the University of Copenhagen. Only one specimen remains in the ZMUC collection, which can be regarded as type. Excellent drawings exist (RLC) of a lost syntype and Krøyer shows the unusually long handles of the companion chaetae compared with those of the adjacent uncini (
|
||
<figureCitation id="64EFD7FBFFD7061AFE86FBCAFE59F50D" box="[336,403,1124,1149]" captionStart="FIGURE 4" captionStartId="8.[151,250,1830,1852]" captionTargetBox="[230,1365,207,1794]" captionTargetId="figure@8.[215,1365,193,1804]" captionTargetPageId="8" captionText="FIGURE 4. Pseudopotamilla aspersa (Krøyer, 1856). A – J & P, N: Type ZMUC POL- 00523; K, L, M: lost Syntype, after Krøyer: A) ‘ whole’ worm, left crown missing, posterior damaged; B) antero-lateral detail of A; C) anterior thorax, dorsal view; D) same, ventral view; E) superior thoracic chaeta; F) inferior thoracic chaeta, front view; G) same, side view; H) side views of thoracic uncinus (left) and companion chaeta (right); J) view of companion chaeta showing teardrop-shaped ‘ blade’; K) uncinus and companion chaeta (after Krøyer); L) same, but part of torus; M) left side of crown; N) and P) two views of abdominal chaetae. Not scaled: H, J. Krøyer gave no scales for K – M." httpUri="https://zenodo.org/record/545881/files/figure.png" pageId="9" pageNumber="210">Fig. 4</figureCitation>
|
||
K–L), the main diagnostic character of the species.
|
||
</paragraph>
|
||
<paragraph id="FC6BCB7EFFD7061AFF11FB27FC3DF409" blockId="9.[151,1437,151,1760]" pageId="9" pageNumber="210">
|
||
Krøyer mysteriously mentioned “Buccal cirri very minute almost rudimentary, but more numerous than usual (5–6 each side)”. There should be only four, if one counts the enlarged pinnule attached to each dorsal lip (e. g.
|
||
<figureCitation id="64EFD7FBFFD7061AFAA6FB02FF68F599" captionStart="FIGURE 1" captionStartId="6.[151,250,1869,1891]" captionTargetBox="[212,1376,203,1844]" captionTargetId="figure@6.[194,1388,193,1853]" captionTargetPageId="6" captionText="FIGURE 1. Pseudopotamilla reniformis (Bruguiére, 1789), Neotype (NMW. Z. 2001.042.0001). A) Whole worm; B) radiolar compound eye; C) cross section of radiole; D) dorsal view of anterior thorax and crown base; E) same, ventral view; F) dorsal lip and dorsal radiolar appendage; G – H, J) collar chaetae; K) superior thoracic chaeta; L) inferior thoracic chaeta; M) abdominal chaeta; N) thoracic uncinus (right) and companion chaeta (left); P) different view of companion chaeta; Q) abdominal uncinus; R) tube showing distal enrolling and the branch of an offspring. Not scaled: B – C, Q & R." httpUri="https://zenodo.org/record/545878/files/figure.png" pageId="9" pageNumber="210">Fig. 1</figureCitation>
|
||
F). The other ‘cirri’ may have been digitiform regeneration of radioles after damage similar to the protuberances in
|
||
<figureCitation id="64EFD7FBFFD7061AFF61FB5AFF36F47D" box="[183,252,1268,1293]" captionStart="FIGURE 4" captionStartId="8.[151,250,1830,1852]" captionTargetBox="[230,1365,207,1794]" captionTargetId="figure@8.[215,1365,193,1804]" captionTargetPageId="8" captionText="FIGURE 4. Pseudopotamilla aspersa (Krøyer, 1856). A – J & P, N: Type ZMUC POL- 00523; K, L, M: lost Syntype, after Krøyer: A) ‘ whole’ worm, left crown missing, posterior damaged; B) antero-lateral detail of A; C) anterior thorax, dorsal view; D) same, ventral view; E) superior thoracic chaeta; F) inferior thoracic chaeta, front view; G) same, side view; H) side views of thoracic uncinus (left) and companion chaeta (right); J) view of companion chaeta showing teardrop-shaped ‘ blade’; K) uncinus and companion chaeta (after Krøyer); L) same, but part of torus; M) left side of crown; N) and P) two views of abdominal chaetae. Not scaled: H, J. Krøyer gave no scales for K – M." httpUri="https://zenodo.org/record/545881/files/figure.png" pageId="9" pageNumber="210">Fig. 4</figureCitation>
|
||
D, where the left side of the crown has been shed. Krøyer also noted that the faecal groove continues longitudinally across the thoracic ventral shields of one of his specimens. This is not uncommon, and represents a thorax once regenerated from abdominal segments after predation (or scissiparity). He notes that the species is rare so probably not found together in a clump, often an indication of scissiparity.
|
||
</paragraph>
|
||
<paragraph id="FC6BCB7EFFD7061AFF11FA2AFBAAF4B1" blockId="9.[151,1437,151,1760]" pageId="9" pageNumber="210">
|
||
<bibRefCitation id="9845B68FFFD7061AFF11FA2AFDE3F4ED" author="Malmgren" box="[199,553,1412,1437]" pageId="9" pageNumber="210" refString="Malmgren, A. J. (1867) Annulata Polychaeta Spetsbergiae, Groenlandiae, Islandiae et Scandinaviae hactenus cognita. Helsingforsiae. Ofversigt af Koniglich Vetenskapsakademiens forhandlingar, Stockholm, 24, 127 - 235." type="journal article" year="1867">
|
||
Malmgren (1867 as
|
||
<taxonomicName id="3BD4B0FDFFD7061AFE67FA2AFDD5F4EC" box="[433,543,1412,1437]" class="Polychaeta" family="Sabellidae" genus="Potamilla" kingdom="Animalia" order="Sabellida" pageId="9" pageNumber="210" phylum="Annelida" rank="genus">
|
||
<emphasis id="CEA0176CFFD7061AFE67FA2AFDD5F4EC" box="[433,543,1412,1437]" italics="true" pageId="9" pageNumber="210">Potamilla</emphasis>
|
||
</taxonomicName>
|
||
)
|
||
</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="9845B68FFFD7061AFDB3FA2AFC74F4ED" author="McIntosh" box="[613,958,1412,1437]" pageId="9" pageNumber="210" refString="McIntosh, W. C. (1923) A monograph of the British marine annelids. Polychaeta, Sabellidae to Serpulidae. With additions to the British marine Polychaeta during the publication of the monograph. Royal Society of London, 4, 251 - 538." type="journal article" year="1923">
|
||
McIntosh (1923 as
|
||
<taxonomicName id="3BD4B0FDFFD7061AFC90FA2AFC7EF4EC" box="[838,948,1412,1437]" class="Polychaeta" family="Sabellidae" genus="Potamilla" kingdom="Animalia" order="Sabellida" pageId="9" pageNumber="210" phylum="Annelida" rank="genus">
|
||
<emphasis id="CEA0176CFFD7061AFC90FA2AFC7EF4EC" box="[838,948,1412,1437]" italics="true" pageId="9" pageNumber="210">Potamilla</emphasis>
|
||
</taxonomicName>
|
||
)
|
||
</bibRefCitation>
|
||
wrongly synonymised
|
||
<taxonomicName id="3BD4B0FDFFD7061AFB1AFA2BFA88F4ED" box="[1228,1346,1413,1436]" class="Polychaeta" family="Sabellidae" genus="Sabella" kingdom="Animalia" order="Sabellida" pageId="9" pageNumber="210" phylum="Annelida" rank="species" species="aspersa">
|
||
<emphasis id="CEA0176CFFD7061AFB1AFA2BFA88F4ED" box="[1228,1346,1413,1436]" italics="true" pageId="9" pageNumber="210">P. aspersa</emphasis>
|
||
</taxonomicName>
|
||
with
|
||
<taxonomicName id="3BD4B0FDFFD7061AFA51FA2BFEC1F4B1" class="Polychaeta" family="Sabellidae" genus="Pseudopotamilla" kingdom="Animalia" order="Sabellida" pageId="9" pageNumber="210" phylum="Annelida" rank="species" species="reniformis">
|
||
<emphasis id="CEA0176CFFD7061AFA51FA2BFEC1F4B1" italics="true" pageId="9" pageNumber="210">P. reniformis</emphasis>
|
||
</taxonomicName>
|
||
, but they probably did not see Krøyer’s material and unpublished drawings.
|
||
</paragraph>
|
||
<paragraph id="FC6BCB7EFFD7061AFF11FA62FC73F721" blockId="9.[151,1437,151,1760]" pageId="9" pageNumber="210">
|
||
The collar of
|
||
<taxonomicName id="3BD4B0FDFFD7061AFE8BFA63FE05F495" box="[349,463,1485,1508]" class="Polychaeta" family="Sabellidae" genus="Sabella" kingdom="Animalia" order="Sabellida" pageId="9" pageNumber="210" phylum="Annelida" rank="species" species="aspersa">
|
||
<emphasis id="CEA0176CFFD7061AFE8BFA63FE05F495" box="[349,463,1485,1508]" italics="true" pageId="9" pageNumber="210">P. aspersa</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="3BD4B0FDFFD7061AFDB4FA63FD3AF495" box="[610,752,1484,1508]" class="Polychaeta" family="Sabellidae" genus="Pseudopotamilla" kingdom="Animalia" order="Sabellida" pageId="9" pageNumber="210" phylum="Annelida" rank="species" species="reniformis">
|
||
<emphasis id="CEA0176CFFD7061AFDB4FA63FD3AF495" box="[610,752,1484,1508]" italics="true" pageId="9" pageNumber="210">P. reniformis</emphasis>
|
||
</taxonomicName>
|
||
in being low ventrally; in having low, rounded dorsal lappets slightly exposing the peristomium; and dorso-lateral margins with deeper notches. The most consistent difference between
|
||
<taxonomicName id="3BD4B0FDFFD7061AFF2DF9BBFE41F75D" box="[251,395,1556,1580]" class="Polychaeta" family="Sabellidae" genus="Pseudopotamilla" kingdom="Animalia" order="Sabellida" pageId="9" pageNumber="210" phylum="Annelida" rank="species" species="reniformis">
|
||
<emphasis id="CEA0176CFFD7061AFF2DF9BBFE41F75D" box="[251,395,1556,1580]" italics="true" pageId="9" pageNumber="210">P. reniformis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="3BD4B0FDFFD7061AFE14F9BBFDFEF75D" box="[450,564,1557,1580]" class="Polychaeta" family="Sabellidae" genus="Sabella" kingdom="Animalia" order="Sabellida" pageId="9" pageNumber="210" phylum="Annelida" rank="species" species="aspersa">
|
||
<emphasis id="CEA0176CFFD7061AFE14F9BBFDFEF75D" box="[450,564,1557,1580]" italics="true" pageId="9" pageNumber="210">P. aspersa</emphasis>
|
||
</taxonomicName>
|
||
is that the latter differs in having each companion chaeta with a handle about twice as long as that of its adjacent uncinus (measured from the breast).
|
||
</paragraph>
|
||
<paragraph id="FC6BCB7EFFD7061AFF11F9F2FE80F7CD" blockId="9.[151,1437,151,1760]" pageId="9" pageNumber="210">
|
||
<emphasis id="CEA0176CFFD7061AFF11F9F2FEE0F704" bold="true" box="[199,298,1628,1653]" pageId="9" pageNumber="210">Habitat.</emphasis>
|
||
Krøyer does not note the habitat of
|
||
<taxonomicName id="3BD4B0FDFFD7061AFCC5F9F2FBF3F705" box="[787,1081,1628,1653]" class="Polychaeta" family="Sabellidae" genus="Pseudopotamilla" kingdom="Animalia" order="Sabellida" pageId="9" pageNumber="210" phylum="Annelida" rank="species" species="aspersa">
|
||
<emphasis id="CEA0176CFFD7061AFCC5F9F2FBF3F705" box="[787,1081,1628,1653]" italics="true" pageId="9" pageNumber="210">Pseudopotamilla aspersa</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="CEA0176CFFD7061AFB84F9F2FA56F705" box="[1106,1436,1628,1653]" italics="true" pageId="9" pageNumber="210">
|
||
<taxonomicName id="3BD4B0FDFFD7061AFB84F9F2FA52F705" box="[1106,1432,1628,1653]" class="Polychaeta" family="Sabellidae" genus="Pseudopotamilla" kingdom="Animalia" order="Sabellida" pageId="9" pageNumber="210" phylum="Annelida" rank="species" species="reniformis">Pseudopotamilla reniformis</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
commonly found on the shore, is also found in
|
||
<collectingCountry id="84C38BEEFFD7061AFD72F9D1FCD6F7E9" box="[676,796,1663,1688]" name="Greenland" pageId="9" pageNumber="210">Greenland</collectingCountry>
|
||
, so niche separation may indicate that
|
||
<taxonomicName id="3BD4B0FDFFD7061AFB1FF92FFAF1F7E9" box="[1225,1339,1665,1688]" class="Polychaeta" family="Sabellidae" genus="Pseudopotamilla" kingdom="Animalia" order="Sabellida" pageId="9" pageNumber="210" phylum="Annelida" rank="species" species="aspersa">
|
||
<emphasis id="CEA0176CFFD7061AFB1FF92FFAF1F7E9" box="[1225,1339,1665,1688]" italics="true" pageId="9" pageNumber="210">P. aspersa</emphasis>
|
||
</taxonomicName>
|
||
is found in deeper water.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B4CE98F5FFD7061AFF11F969FE29F791" box="[199,483,1735,1760]" pageId="9" pageNumber="210" type="distribution">
|
||
<paragraph id="FC6BCB7EFFD7061AFF11F969FE29F791" blockId="9.[151,1437,151,1760]" box="[199,483,1735,1760]" pageId="9" pageNumber="210">
|
||
<emphasis id="CEA0176CFFD7061AFF11F969FEAAF791" bold="true" box="[199,352,1735,1760]" pageId="9" pageNumber="210">Distribution.</emphasis>
|
||
<collectingCountry id="84C38BEEFFD7061AFEB1F969FE15F791" box="[359,479,1735,1760]" name="Greenland" pageId="9" pageNumber="210">Greenland</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |