219 lines
27 KiB
XML
219 lines
27 KiB
XML
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<mods:title id="88C30009885F0DCA1AFDB299AA1BF58F">The genus Symmela Erichson, 1835 (Coleoptera, Scarabaeidae, Sericini): taxonomy and phylogenetic analysis</mods:title>
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<mods:namePart id="F445270431AC0CAB0168717EADDC93C2">Pacheco, Thaynara L.</mods:namePart>
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<mods:namePart id="3C0B391E5338B6285F436FABF2FD66D0">Wipfler, Benedict</mods:namePart>
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<mods:namePart id="914C48D76286C0334FDE48B416C596CD">Monné, Marcela L.</mods:namePart>
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<mods:namePart id="5A2C431E2BF144B06DAF62EAC5B643E7">Ahrens, Dirk</mods:namePart>
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<mods:title id="007743EA3439789A8B31B392643F4637">Journal of Natural History</mods:title>
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<mods:date id="B2F06E0A2E5A956C2E7F9C41A028B24D">2022</mods:date>
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<mods:number id="496A3FD44B1B402C84C62821F86F9A86">2022-07-11</mods:number>
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<mods:number id="4A56339B61E17476BA46801FF9B7A790">56</mods:number>
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<mods:number id="B67182B2CEE12012AB9B3476D4A3FC17">9 - 12</mods:number>
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<mods:identifier id="32E48B64CB658F92BDFCDC537355623A" type="DOI">10.1080/00222933.2022.2084649</mods:identifier>
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<treatment id="5D554F4F217DFFAAFEBCFA843A45FB27" ID-DOI="http://doi.org/10.5281/zenodo.7019067" ID-GBIF-Taxon="199728119" ID-Zenodo-Dep="7019067" LSID="urn:lsid:plazi:treatment:5D554F4F217DFFAAFEBCFA843A45FB27" httpUri="http://treatment.plazi.org/id/5D554F4F217DFFAAFEBCFA843A45FB27" lastPageId="78" lastPageNumber="685" pageId="75" pageNumber="682">
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<subSubSection id="9DE6ADD2217DFFAFFEBCFA843D2EFA2A" box="[262,1053,1385,1411]" pageId="75" pageNumber="682" type="nomenclature">
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<paragraph id="D543FE59217DFFAFFEBCFA843D2EFA2A" blockId="75.[262,1053,1385,1411]" box="[262,1053,1385,1411]" pageId="75" pageNumber="682">
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<heading id="8E0B4935217DFFAFFEBCFA843D2EFA2A" bold="true" box="[262,1053,1385,1411]" centered="true" fontSize="11" level="2" pageId="75" pageNumber="682" reason="3">
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||
<emphasis id="E788224B217DFFAFFEBCFA843D2EFA2A" bold="true" box="[262,1053,1385,1411]" pageId="75" pageNumber="682">
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<taxonomicName id="12FC85DA217DFFAFFEBCFA843A80FA2A" authority="Pacheco, Wipfler, Monne and Ahrens" authorityName="Pacheco, Wipfler, Monne and Ahrens" authorityYear="2022" box="[262,947,1385,1411]" class="Insecta" family="Melolonthidae" genus="Symmela" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="682" phylum="Arthropoda" rank="species" species="reischei" status="sp. nov.">
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||
<emphasis id="E788224B217DFFAFFEBCFA8438E9FA2A" bold="true" box="[262,474,1385,1411]" italics="true" pageId="75" pageNumber="682">Symmela reischei</emphasis>
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Pacheco, Wipfler, Monné and Ahrens
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||
</taxonomicName>
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||
<taxonomicNameLabel id="FCBB9F30217DFFAFFC01FA843D2EFA2A" box="[955,1053,1385,1411]" pageId="75" pageNumber="682" rank="species">sp. nov.</taxonomicNameLabel>
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||
</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="9DE6ADD2217DFFAFFD96FA4D3BC4FA13" box="[556,759,1440,1466]" pageId="75" pageNumber="682" type="description">
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<paragraph id="D543FE59217DFFAFFD96FA4D3BC4FA13" blockId="75.[556,759,1440,1466]" box="[556,759,1440,1466]" pageId="75" pageNumber="682">
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||
(
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<figureCitation id="4DC7E2DC217DFFAFFD8EFA4D3B9DFA13" box="[564,686,1440,1466]" captionStart="Figure 25" captionStartId="77.[160,225,1516,1539]" captionTargetBox="[168,1149,142,1498]" captionTargetId="figure-3@77.[167,1150,142,1499]" captionTargetPageId="77" captionText="Figure 25. Symmela reischei sp. nov., (a, b) paratype ♂, (c–e) holotype ♂. (a) Habitus, dorsal view; (b) habitus, lateral view; (c) aedeagus, lateral view; (d) aedeagus, dorsal view; (e) parameres, dorsal view. Scale bars: a, b = 1 mm; c–e = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7012576" httpUri="https://zenodo.org/record/7012576/files/figure.png" pageId="75" pageNumber="682">Figures 25</figureCitation>
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,
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<figureCitation id="4DC7E2DC217DFFAFFD06FA4D3BE9FA13" box="[700,730,1440,1466]" captionStart="Figure 31" captionStartId="88.[160,225,1283,1306]" captionTargetBox="[163,1153,142,1266]" captionTargetId="figure-83@88.[163,1153,142,1266]" captionTargetPageId="88" captionText="Figure 31. Heads of Symmela species. (a) S. martinezi sp. nov.; (b) S. mutabilis; (c) S. nitida; (d) S. nitidicollis; (e) S. nunesorum sp. nov.; (f) S. opaca; (g) S. paschoali sp. nov.; (h) S. pseudopaca sp. nov.; (i) S. reischei sp. nov.; (j) S. tarsalis; (k) S. terezae sp. nov.; (l) S. unidentata sp. nov. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7012588" httpUri="https://zenodo.org/record/7012588/files/figure.png" pageId="75" pageNumber="682">31</figureCitation>
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(i))
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="9DE6ADD2217DFFAFFF1AF9EA3D1AF9CF" pageId="75" pageNumber="682" type="materials_examined">
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||
<paragraph id="D543FE59217DFFAFFF1AF9EA388CF988" blockId="75.[160,1156,1543,1638]" box="[160,447,1543,1569]" pageId="75" pageNumber="682">
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||
<emphasis id="E788224B217DFFAFFF1AF9EA388CF988" bold="true" box="[160,447,1543,1569]" italics="true" pageId="75" pageNumber="682">Type material examined</emphasis>
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||
</paragraph>
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||
<paragraph id="D543FE59217DFFAFFF1AF9C43D1AF9CF" blockId="75.[160,1156,1543,1638]" pageId="75" pageNumber="682">
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||
<materialsCitation id="6594F404217DFFAFFF1AF9C43DB3F9EA" ID-GBIF-Occurrence="3892866343" box="[160,1152,1577,1603]" collectionCode="MNHN" country="Brazil" location="Reische" municipality="Neotropical Sericini" pageId="75" pageNumber="682" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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||
<emphasis id="E788224B217DFFAFFF1AF9C43821F9EA" bold="true" box="[160,274,1577,1603]" italics="true" pageId="75" pageNumber="682">
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||
<typeStatus id="0A4740FB217DFFAFFF1AF9C4383EF9EA" box="[160,269,1577,1603]" pageId="75" pageNumber="682" type="holotype">Holotype</typeStatus>
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||
.
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||
</emphasis>
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||
<specimenCount id="C3FA35D0217DFFAFFE9BF9C63806F9EB" box="[289,309,1579,1602]" pageId="75" pageNumber="682" type="male">♂</specimenCount>
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‘
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<taxonomicName id="12FC85DA217DFFAFFE84F9C73B30F9EA" authorityName="Erichson" authorityYear="1835" box="[318,515,1578,1603]" class="Insecta" family="Melolonthidae" genus="Symmela" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="682" phylum="Arthropoda" rank="species" species="nitidula">Symmela nitidula</taxonomicName>
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<collectingCountry id="ADEBBEC9217DFFAFFDB3F9C73B76F9EA" box="[521,581,1578,1603]" name="Brazil" pageId="75" pageNumber="682">Brésil</collectingCountry>
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||
<location id="D023A882217DFFAFFDF0F9C73BACF9EA" LSID="urn:lsid:plazi:treatment:5D554F4F217DFFAAFEBCFA843A45FB27:D023A882217DFFAFFDF0F9C73BACF9EA" box="[586,671,1578,1603]" country="Brazil" municipality="Neotropical Sericini" name="Reische" pageId="75" pageNumber="682">Reische</location>
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||
2/2/152
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<collectingMunicipality id="35276423217DFFAFFCB8F9C73AE8F9EA" box="[770,987,1578,1603]" pageId="75" pageNumber="682">Neotropical Sericini</collectingMunicipality>
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spec.’ (
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<collectionCode id="B3ED669C217DFFAFFB91F9C73D48F9EA" box="[1067,1147,1578,1603]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="75" pageNumber="682">MNHN</collectionCode>
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)
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</materialsCitation>
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||
.
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<typeStatus id="0A4740FB217DFFAFFF1AF9A03836F9CF" box="[160,261,1613,1638]" pageId="75" pageNumber="682" type="paratype">Paratype</typeStatus>
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:
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<specimenCount id="C3FA35D0217DFFAFFEAAF9A0380AF9CD" box="[272,313,1613,1638]" pageId="75" pageNumber="682" type="male">1 ♂</specimenCount>
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‘Brasilia/
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<taxonomicName id="12FC85DA217DFFAFFE1EF9A03B45F9CF" authority=", Er." authorityName="Er." box="[420,630,1613,1638]" class="Insecta" family="Scarabaeidae" genus="Symela" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="682" phylum="Arthropoda" rank="species" species="longula">
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<emphasis id="E788224B217DFFAFFE1EF9A03B7FF9CF" box="[420,588,1613,1638]" italics="true" pageId="75" pageNumber="682">Symela longula</emphasis>
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, Er.
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</taxonomicName>
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/ 79 Neotropical Sericini spec.’ (
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<collectionCode id="B3ED669C217DFFAFFC6FF9A03D2CF9CF" box="[981,1055,1613,1638]" pageId="75" pageNumber="682">ZMHB</collectionCode>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="9DE6ADD2217AFFAAFF1AFF633859FDC3" lastPageId="78" lastPageNumber="685" pageId="76" pageNumber="683" type="description">
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<paragraph id="D543FE59217AFFA8FF1AFF633815FF01" blockId="76.[160,1156,142,1312]" box="[160,294,142,168]" pageId="76" pageNumber="683">
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<emphasis id="E788224B217AFFA8FF1AFF633815FF01" bold="true" box="[160,294,142,168]" italics="true" pageId="76" pageNumber="683">Description</emphasis>
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</paragraph>
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<paragraph id="D543FE59217AFFA8FF1AFF5F3BB3FEFC" blockId="76.[160,1156,142,1312]" pageId="76" pageNumber="683">
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Elytral length:
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<quantity id="120453BC217AFFA8FEF3FF5F38A2FF62" box="[329,401,178,203]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="76" pageNumber="683" unit="mm" value="4.0">4 mm</quantity>
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. Width:
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<quantity id="120453BC217AFFA8FE49FF5F3B7DFF62" box="[499,590,178,203]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="76" pageNumber="683" unit="mm" value="3.3">3.3 mm</quantity>
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.
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<taxonomicName id="12FC85DA217AFFA8FDE7FF5F3BFFFF62" box="[605,716,178,203]" class="Trilobita" family="Styginidae" genus="Scutellum" kingdom="Animalia" order="Corynexochida" pageId="76" pageNumber="613" phylum="Arthropoda" rank="genus">Scutellum</taxonomicName>
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||
without punctures. Elytra bicoloured, yellowish brown in major part, margins blackish; surface shiny, without setae and microsculpture; striae and intervals distinct; punctation between striae sparse, diameter of punctures compared to those of striae smaller; intervals flat; posterior edge convex; apical margin chitinous, without microtrichomes.
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</paragraph>
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<paragraph id="D543FE59217AFFA8FF7AFEB23BE6FD8C" blockId="76.[160,1156,142,1312]" pageId="76" pageNumber="683">
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Abdomen: sternites flat in sagittal plane; lateral carina absent; first visible ventrite with row of setae on posterior edge; second visible ventrite with row of setae in medial region; basal ventrites medially with half length as laterally; ultimate and penultimate apical ventrites simple and flat; apical ventrite very short (half as long as preceding ventrite); penultimate apical ventrite with row of setae like preceding one; beside the robust row of setae, almost glabrous.
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<taxonomicName id="12FC85DA217AFFA8FE10FDE13B27FD8C" box="[426,532,524,549]" class="Actinopterygii" family="Trichomycteridae" genus="Pygidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="76" pageNumber="613" phylum="Chordata" rank="genus">Pygidium</taxonomicName>
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strongly convex.
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</paragraph>
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<paragraph id="D543FE59217AFFA8FF7AFDC23BEFFB72" blockId="76.[160,1156,142,1312]" pageId="76" pageNumber="683">Legs yellow. Ratio of length of metepisternum/metacoxa: 1/1.03. Metacoxa entirely plain, without elevated lateral longitudinal carina; without longer lateral bristles, with setae on basal surface, posterior margin glabrous. Metafemur with posterior margin straight. Metatibia widest at apex; ratio of width/length: 1/3.14; dorsal margin carinated, smooth; interior surface more or less glabrous; median longitudinal punctation present on interior surface; interior margin of apex sharply and deeply truncate towards ventral margin; punctation on entire lateral face present; beside dorsal margin without a row of robust setae along entire length; basal group of spines comprising at least two robust spines at same level of metatibial length; distal groups of spines disposed at two-thirds of metatibia length; ventral margin not serrated. Dorsal surface of tarsi glabrous, with dense punctures. Length ratio of dorsal tibial spur/first metatarsomere: 1/2. Metatarsi at apex weakly widened; first tarsomere straight, its length subequal to second and third tarsomeres combined; laterally not carinate; ventral margin serrated; ventral margin glabrous; medial face glabrous. Protibia with two distinct teeth; spur straight. First protarsomere straight, its length subequal to second and third combined; width ratio of first protarsomere vs second protarsomere: 1.5/1; second protarsomere narrow and straight; third and fourth protarsomere elongate and narrow (distinctly longer than wide); fifth protarsomere circular in cross section, a little shorter than third and fourth protarsomere combined. Protarsal claws asymmetrical, shape of inner protarsal claw bifid, apex of basal tooth of inner protarsal claw convex.</paragraph>
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<paragraph id="D543FE59217AFFA8FF7AFB0938D2FB54" blockId="76.[160,1156,142,1312]" box="[192,481,1252,1277]" pageId="76" pageNumber="683">
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Aedeagus:
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<figureCitation id="4DC7E2DC217AFFA8FE86FB093894FB54" box="[316,423,1252,1277]" captionStart="Figure 25" captionStartId="77.[160,225,1516,1539]" captionTargetBox="[168,1149,142,1498]" captionTargetId="figure-3@77.[167,1150,142,1499]" captionTargetPageId="77" captionText="Figure 25. Symmela reischei sp. nov., (a, b) paratype ♂, (c–e) holotype ♂. (a) Habitus, dorsal view; (b) habitus, lateral view; (c) aedeagus, lateral view; (d) aedeagus, dorsal view; (e) parameres, dorsal view. Scale bars: a, b = 1 mm; c–e = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7012576" httpUri="https://zenodo.org/record/7012576/files/figure.png" pageId="76" pageNumber="683">Figure 25</figureCitation>
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(c–e).
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</paragraph>
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<paragraph id="D543FE59217AFFA8FF7AFAEA38BAFA89" blockId="76.[160,1156,142,1312]" box="[192,393,1287,1312]" pageId="76" pageNumber="683">Female unknown.</paragraph>
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<paragraph id="D543FE59217AFFA8FF1AFAA1383DFACF" blockId="76.[160,1156,1356,1728]" box="[160,270,1356,1382]" pageId="76" pageNumber="683">
|
||
<emphasis id="E788224B217AFFA8FF1AFAA1383DFACF" bold="true" box="[160,270,1356,1382]" italics="true" pageId="76" pageNumber="683">Variation</emphasis>
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||
</paragraph>
|
||
<paragraph id="D543FE59217AFFAAFF1AFA823AE2FEA6" blockId="76.[160,1156,1356,1728]" lastBlockId="78.[160,1156,142,618]" lastPageId="78" lastPageNumber="685" pageId="76" pageNumber="683">
|
||
Length:
|
||
<quantity id="120453BC217AFFA8FF41FA823863FA21" box="[251,336,1391,1416]" metricMagnitude="-3" metricUnit="m" metricValue="7.1" pageId="76" pageNumber="683" unit="mm" value="7.1">7.1 mm</quantity>
|
||
. Elytral length:
|
||
<quantity id="120453BC217AFFA8FE42FA823B5AFA21" box="[504,617,1391,1416]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" metricValueMax="4.2" metricValueMin="4.0" pageId="76" pageNumber="683" unit="mm" value="4.1" valueMax="4.2" valueMin="4.0">4–4.2 mm</quantity>
|
||
. Width:
|
||
<quantity id="120453BC217AFFA8FD7BFA823A76FA21" box="[705,837,1391,1416]" metricMagnitude="-3" metricUnit="m" metricValue="3.55" metricValueMax="3.8" metricValueMin="3.3" pageId="76" pageNumber="683" unit="mm" value="3.55" valueMax="3.8" valueMin="3.3">3.3–3.8 mm</quantity>
|
||
. Head blackish brown, shiny, with erect, sparse setae. Labroclypeus as wide as frons, subtrapezoidal; surface convex; plain, without an anterior concavity on each side; anterior margin straight, weakly reflexed, with a rim of sparse setae behind anterior margin; lateral margin not incised anteriorly; punctation of surface posteriorly dense; suture between labrum and clypeus invisible. Smooth area in front of eye distinctly separated from frons by a suture. Frons anteriorly flat, punctation irregular, dense. Ocular canthus with terminal setae, ratio of length of ocular canthus/eye diameter: 1/3. Eyes evenly spherical. Antenna with eight antennomeres. Antennal club with three antennomeres, blackish, straight; width of antennal lamella equal to width of antennomeres of funiculus; apical lamella subequal to basal and median ones; length ratio of antennal club vs remaining antennomeres combined: 1/1.1; length ratio of third vs fourth antennomere: 1/2; fourth antennomere without lateral projection; length ratio of fifth vs fourth antennomere: 1/3. Labrum anteriorly without blunt processes. Setae on mentum moderately dense.
|
||
</paragraph>
|
||
<caption id="8183AED1217BFFA9FF1AFA013B1AF997" ID-DOI="http://doi.org/10.5281/zenodo.7012576" ID-Zenodo-Dep="7012576" httpUri="https://zenodo.org/record/7012576/files/figure.png" pageId="77" pageNumber="684" startId="77.[160,225,1516,1539]" targetBox="[168,1149,142,1498]" targetPageId="77">
|
||
<paragraph id="D543FE59217BFFA9FF1AFA013B1AF997" blockId="77.[160,1158,1516,1598]" pageId="77" pageNumber="684">
|
||
<emphasis id="E788224B217BFFA9FF1AFA013834F9AA" bold="true" box="[160,263,1516,1539]" pageId="77" pageNumber="684">Figure 25.</emphasis>
|
||
<taxonomicName id="12FC85DA217BFFA9FEB7FA013882F9AA" authority="Pacheco & Wipfler & Monné & Ahrens, 2022" authorityName="Pacheco & Wipfler & Monné & Ahrens" authorityYear="2022" box="[269,433,1516,1539]" class="Insecta" family="Melolonthidae" genus="Symmela" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="684" phylum="Arthropoda" rank="species" species="reischei" status="sp. nov.">
|
||
<emphasis id="E788224B217BFFA9FEB7FA013882F9AA" box="[269,433,1516,1539]" italics="true" pageId="77" pageNumber="684">Symmela reischei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="FCBB9F30217BFFA9FE02FA013B34F9AA" box="[440,519,1516,1539]" pageId="77" pageNumber="684" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, (a, b) paratype ♂, (c–e) holotype ♂. (a) Habitus, dorsal view; (b) habitus, lateral view; (c) aedeagus, lateral view; (d) aedeagus, dorsal view; (e) parameres, dorsal view. Scale bars: a, b = 1 mm; c–e = 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="D543FE592178FFAAFF7AFEF43859FDC3" blockId="78.[160,1156,142,618]" pageId="78" pageNumber="685">Head and pronotum without copper iridescent shine. Pronotum bicoloured, shiny, blackish brown; widest at base, width subequal to width of elytral base at humerus; anterior marginal line not interrupted medially, posterior marginal line absent; lateral margin not serrated, generally bent at middle, otherwise straight, behind anterior angles convex, posteriorly straight; setae of lateral margin sparse; basal margin evenly curved; transverse impression beside basal margin absent; anterior angles produced, sharply pointed; antero-lateral and postero-lateral angles not flattened dorso-ventrally; posterior angles blunt; surface without median longitudinal impression, transversal furrows or setae; punctation on disc moderately dense, simple and fine, without well-defined impunctate areas.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9DE6ADD22178FFAAFF1AFD723AFCFCEA" pageId="78" pageNumber="685" type="diagnosis">
|
||
<paragraph id="D543FE592178FFAAFF1AFD723827FD10" blockId="78.[160,1156,671,835]" box="[160,276,671,697]" pageId="78" pageNumber="685">
|
||
<emphasis id="E788224B2178FFAAFF1AFD723827FD10" bold="true" box="[160,276,671,697]" italics="true" pageId="78" pageNumber="685">Diagnosis</emphasis>
|
||
</paragraph>
|
||
<paragraph id="D543FE592178FFAAFF1AFD2F3AFCFCEA" blockId="78.[160,1156,671,835]" pageId="78" pageNumber="685">
|
||
<taxonomicName id="12FC85DA2178FFAAFF1AFD2F386FFD72" authority="Pacheco & Wipfler & Monné & Ahrens, 2022" authorityName="Pacheco & Wipfler & Monné & Ahrens" authorityYear="2022" box="[160,348,706,731]" class="Insecta" family="Melolonthidae" genus="Symmela" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="685" phylum="Arthropoda" rank="species" species="reischei" status="sp. nov.">
|
||
<emphasis id="E788224B2178FFAAFF1AFD2F386FFD72" box="[160,348,706,731]" italics="true" pageId="78" pageNumber="685">Symmela reischei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="FCBB9F302178FFAAFEDEFD2F388FFD72" box="[356,444,706,731]" pageId="78" pageNumber="685" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is in external appearance similar to
|
||
<taxonomicName id="12FC85DA2178FFAAFCE3FD2F39E4FD57" authority="Erichson, 1835" authorityName="Erichson" authorityYear="1835" class="Insecta" family="Melolonthidae" genus="Symmela" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="685" phylum="Arthropoda" rank="species" species="longula">
|
||
<emphasis id="E788224B2178FFAAFCE3FD2F3D25FD72" box="[857,1046,706,731]" italics="true" pageId="78" pageNumber="685">Symmela longula</emphasis>
|
||
<bibRefCitation id="B16D83A82178FFAAFBA5FD2F39E4FD57" author="Erichson WF" pageId="78" pageNumber="685" pagination="256 - 270" refId="ref42712" refString="Erichson WF. 1835. Neue Sudamerikanische Kafergattungen aus der Familie der Blatterhorner. Arch Naturgeschichte. 1: 256 - 270." type="journal article" year="1835">Erichson, 1835</bibRefCitation>
|
||
</taxonomicName>
|
||
, but the body of this new species is significantly larger, and the parameres are stout and have a complicated shape, which differentiates them from all other
|
||
<taxonomicName id="12FC85DA2178FFAAFC79FCE53DB3FC88" box="[963,1152,776,801]" pageId="78" pageNumber="685">
|
||
<emphasis id="E788224B2178FFAAFC79FCE53D15FC88" box="[963,1062,776,801]" italics="true" pageId="78" pageNumber="685">Symmela</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
: at the apex they have a sharp dorsal process which is reflexed interiorly.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9DE6ADD22178FFAAFF1AFC953872FC71" pageId="78" pageNumber="685" type="etymology">
|
||
<paragraph id="D543FE592178FFAAFF1AFC95382EFC3B" blockId="78.[160,1155,888,984]" box="[160,285,888,914]" pageId="78" pageNumber="685">
|
||
<emphasis id="E788224B2178FFAAFF1AFC95382EFC3B" bold="true" box="[160,285,888,914]" italics="true" pageId="78" pageNumber="685">Etymology</emphasis>
|
||
</paragraph>
|
||
<paragraph id="D543FE592178FFAAFF1AFC713872FC71" blockId="78.[160,1155,888,984]" pageId="78" pageNumber="685">
|
||
The new species is named after Reische, collector of the
|
||
<typeStatus id="0A4740FB2178FFAAFC86FC713A93FC1C" box="[828,928,924,949]" pageId="78" pageNumber="685" type="holotype">holotype</typeStatus>
|
||
specimen (noun in genitive case).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9DE6ADD22178FFAAFF1AFBE03A45FB27" pageId="78" pageNumber="685" type="discussion">
|
||
<paragraph id="D543FE592178FFAAFF1AFBE03835FB8E" blockId="78.[160,1156,1037,1166]" box="[160,262,1037,1063]" pageId="78" pageNumber="685">
|
||
<emphasis id="E788224B2178FFAAFF1AFBE03835FB8E" bold="true" box="[160,262,1037,1063]" italics="true" pageId="78" pageNumber="685">Remarks</emphasis>
|
||
</paragraph>
|
||
<paragraph id="D543FE592178FFAAFF1AFBDD3A45FB27" blockId="78.[160,1156,1037,1166]" pageId="78" pageNumber="685">
|
||
Pronotum and head of the
|
||
<typeStatus id="0A4740FB2178FFAAFE54FBDD3B67FBE0" box="[494,596,1072,1097]" pageId="78" pageNumber="685" type="holotype">holotype</typeStatus>
|
||
are missing. The parameres are broken in the
|
||
<typeStatus id="0A4740FB2178FFAAFF1AFBBE3835FBC5" box="[160,262,1107,1132]" pageId="78" pageNumber="685" type="paratype">paratype</typeStatus>
|
||
; however, based on the shape of the phallobase and the rest of the external morphology, this specimen can be clearly assigned to
|
||
<taxonomicName id="12FC85DA2178FFAAFCBCFB983A5CFB27" authorityName="Pacheco & Wipfler & Monné & Ahrens" authorityYear="2022" box="[774,879,1141,1166]" class="Insecta" family="Melolonthidae" genus="Symmela" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="685" phylum="Arthropoda" rank="species" species="reischei">
|
||
<emphasis id="E788224B2178FFAAFCBCFB983A5CFB27" box="[774,879,1141,1166]" italics="true" pageId="78" pageNumber="685">S. reischei</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |