158 lines
16 KiB
XML
158 lines
16 KiB
XML
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<mods:title id="5FBD8A515AFB933EFAD26DA70C4BDA91">A molecular and conchological dissection of the " scaly " Georissa of Malaysian Borneo (Gastropoda, Neritimorpha, Hydrocenidae)</mods:title>
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<mods:namePart id="69AB0BECBC784E9429B7A82B77C47A3C">Khalik, Mohd Zacaery</mods:namePart>
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<mods:namePart id="929968DC5C112A4F94FB327358866E86">Hendriks, Kasper</mods:namePart>
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<mods:namePart id="801EB01523F5EC26DD083CBEAC995D9B">Vermeulen, Jaap J.</mods:namePart>
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<mods:namePart id="1986C76E92B008A2A4734AC6762A6AF4">Schilthuizen, Menno</mods:namePart>
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<mods:date id="2ACC6790EC93767B6125321A6F428EBF">2018</mods:date>
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<mods:number id="2482BE7887D248A0E4FB05185FFDDE82">773</mods:number>
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<mods:identifier id="AEA581DD8939B438F3528DFD5051C3BD" type="DOI">http://dx.doi.org/10.3897/zookeys.773.24878</mods:identifier>
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<treatment id="52CEE790316365EEBF506FD11C98E6C7" ID-GBIF-Taxon="146053688" LSID="urn:lsid:plazi:treatment:52CEE790316365EEBF506FD11C98E6C7" httpUri="http://treatment.plazi.org/id/52CEE790316365EEBF506FD11C98E6C7" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection id="111D3109CBEA62623F948060BD8DB7FA" pageId="0" pageNumber="1" type="nomenclature">
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<paragraph id="805753A6FE2AF82AB98443FC8CA4459E" pageId="0" pageNumber="1">
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<taxonomicName id="EEF38ADBE16A983AD25365DA337F2F2D" ID-CoL="3FRYF" authority="van Benthem-Jutting, 1966" authorityName="van Benthem-Jutting" authorityYear="1966" class="Gastropoda" family="Hydrocenidae" genus="Georissa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Georissa scalinella" order="Cycloneritida" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="scalinella">Georissa scalinella (van Benthem-Jutting, 1966)</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="74A318E7A9FF25C2F55F5F0FC70D61D9" pageId="0" pageNumber="1" type="reference_group">
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<paragraph id="1DBB85E6B7C880F9D419E51286C4588D" pageId="0" pageNumber="1">
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<taxonomicName id="0458BDFD57DA859AC17B23D8A968FBFD" class="Gastropoda" family="Hydrocenidae" genus="Hydrocena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydrocena scalinella" order="Cycloneritida" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="scalinella">Hydrocena scalinella</taxonomicName>
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van Benthem-Jutting, 1966: 39, fig. 1;
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<bibRefCitation id="E66E32F3080AF3BE0F9C506C09C9A933" author="Saul, M" journalOrPublisher="Sabah Society Journal" pageId="0" pageNumber="1" pagination="105 - 110" title="Shell collecting in the limestone caves of Borneo." volume="3" year="1967">Saul 1967</bibRefCitation>
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: 108.
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</paragraph>
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<paragraph id="7376459EEB7A398B022A62914D088F20" pageId="0" pageNumber="1">
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<taxonomicName id="6B5CE0521757E14D0B8A838EE917D2A2" class="Gastropoda" family="Hydrocenidae" genus="Georissa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Georissa scalinella" order="Cycloneritida" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="scalinella">Georissa scalinella</taxonomicName>
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(van Benthem-Jutting):
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<bibRefCitation id="0D339145A1012C225C5B62E22364E6A1" author="Thompson, FG" journalOrPublisher="Bulletin of the Florida State Museum Biological Sciences" pageId="0" pageNumber="1" pagination="101 - 152" title="Non-marine mollusks of Borneo. II Pulmonata: Pupillidae, Clausiliidae. III Prosobranchia: Hydrocenidae, Helicinidae." url="https://www.floridamuseum.ufl.edu/files/2114/7180/1931/Vol-29-No-3.PDF" volume="29" year="1983">Thompson and Dance 1983</bibRefCitation>
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: 119;
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<bibRefCitation id="CB6938CD1DE2B7142BD66ADCF5C89623" author="Phung, CC" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="49 - 104" title="A checklist of land snails from the west coast islands of Sabah, Borneo (Mollusca, Gastropoda)." url="https://doi.org/10.3897/zookeys.673.12422" volume="673" year="2017">Phung et al. 2017</bibRefCitation>
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: 68, fig. 8B.
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</paragraph>
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</subSubSection>
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<subSubSection id="9F7AC433F6C85847A775DF7DD5A24288" pageId="0" pageNumber="1" type="type locality">
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<paragraph id="33FDCE7BD371A778970FD3C641B6AE63" pageId="0" pageNumber="1">Type locality.</paragraph>
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<paragraph id="BA4E3DDB4984F2E1FE8482B102747F3E" pageId="0" pageNumber="1">Lahad Datu Caves on Teck Guan Estate, Sabah.</paragraph>
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</subSubSection>
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<subSubSection id="B99A937F1835C06E5DF3007577EC7675" pageId="0" pageNumber="1" type="type material">
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<paragraph id="F486EC88C0BB507FB91C01907D16D815" pageId="0" pageNumber="1">Type material.</paragraph>
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<paragraph id="568A8D5C7CBD8B2576E4B08A019088C3" pageId="0" pageNumber="1">Holotype. Lahad Datu Caves on Teck Guan Estate, Sabah: ZMA/MOLL 135736 (seen). Paratypes. Lahad Datu Caves on Teck Guan Estate, Sabah: ZMA/MOLL 135735 (seen), ZMA/MOLL 315596 (seen).</paragraph>
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</subSubSection>
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<subSubSection id="19E1102AD38F00EC69FFCB8CBAB0ABA1" pageId="0" pageNumber="1" type="description">
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<paragraph id="97238FC24F66E82393B882AC8D05C937" pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph id="6F8967A3135F3FA671E6D0088F38C869" pageId="0" pageNumber="1">
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Protoconch. Color: orange to red. Sculpture: smooth to meshed - semi oval mesh to undefined mesh pattern. Mesh width: 7-17
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<normalizedToken id="6EF89238F9D83C4AF3CC4A81129E6ADF" originalValue="µm">µm</normalizedToken>
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. Teleoconch. Color: orange. First whorl: flat at the shoulder. Subsequent whorls: flat above, slightly rounded below the periphery. Total number of whorls: 2
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<normalizedToken id="1C252F3D440CAC3CED5ACC97E43DCA16" originalValue="¼">1/4</normalizedToken>
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-2
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<normalizedToken id="FDE41FE1A1EFF3C3B9AE1511A762BCFC" originalValue="½">1/2</normalizedToken>
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. SH: 1.56-1.80 mm, SW: 1.46-1.65 mm, SI: 1.03-1.15. Shell sculpture. Radial sculpture: absent, only weak to strong growth lines are visible. Spiral sculpture: present, and strongly sculpted, with continuous and discontinuous ribbing. Scales: a series of acute scales, low to highly projected, and regularly spaced. Intercept between growth lines and spiral ribbings form small pointed scale structures throughout the length of the body whorl. Aperture. Shape: oval. Basal side: rounded, angular at the columellar region. Parietal side: straight, palatal edge attached to the body whorl. AH: 0.78-0.94 mm, AW: 0.97-1.12 mm, AI: 0.75-0.89. Holotype dimension. SH: 1.88 mm, SW: 1.72 mm, AH: 0.84 mm, AW: 1.18 mm.
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</paragraph>
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<caption id="02C7625B537A8037E8FA429DC74D4266" pageId="0" pageNumber="1">
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<paragraph id="61976F1CEF6A854D1B3E74E90353FA9F" pageId="0" pageNumber="1">
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Figure 5.
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<taxonomicName id="2DE437A2B155966EBE2CA84B6E6B83C5" class="Gastropoda" family="Hydrocenidae" genus="Georissa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Georissa scalinella" order="Cycloneritida" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="scalinella">Georissa scalinella</taxonomicName>
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(van Benthem-Jutting, 1966).
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<normalizedToken id="664D50B476DC8687BD0370C27A09B833" originalValue="A–C">A-C</normalizedToken>
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Holotype: ZMA/MOL/ 135736
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<normalizedToken id="84714DD079E18DE73AB73503DF0EAF23" originalValue="D–K">D-K</normalizedToken>
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Paratypes: ZMA/MOLL 135735. A, D Shell apertural view B Shell side view C Shell rear view
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<normalizedToken id="2674BF49EB3DDB4A955E9E0F4260E58C" originalValue="E–F">E-F</normalizedToken>
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Shell cross-section from 3D model
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<normalizedToken id="15A419CE48AF83D66497D5DBF60B2739" originalValue="G–H">G-H</normalizedToken>
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Operculum frontal and ventral view I Shell top view J Protoconch side view K Close up of protoconch from top at 1000
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<normalizedToken id="DB65C4EE71ADB076B8DC5A538334B7AA" originalValue="×">x</normalizedToken>
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magnification. Scale bars: 500
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<normalizedToken id="C580EBB9A8834A3C757AE8036A0A7695" originalValue="µm">µm</normalizedToken>
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(
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<normalizedToken id="DA10FFAA5956B93D6ABE9B7E001D6366" originalValue="A–I">A-I</normalizedToken>
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); 200
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<normalizedToken id="EC1E06D7E80C5EFC81358290CE1848CA" originalValue="µm">µm</normalizedToken>
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(J); 10
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<normalizedToken id="41D60E5A0616B54E8720162F0DD9993A" originalValue="µm">µm</normalizedToken>
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(K).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection id="7DF71DFBF355892F7217A668A69BDD7A" pageId="0" pageNumber="1" type="cross diagnosis">
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<paragraph id="60A0CA1D44ECDF4045C472D42D84F176" pageId="0" pageNumber="1">Cross diagnosis.</paragraph>
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<paragraph id="1AAE8EF2CDC45557BCD2A87ED176A404" pageId="0" pageNumber="1">
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<taxonomicName id="B2779DB253E44A9BF652CA0599FD32D3" class="Gastropoda" family="Hydrocenidae" genus="Georissa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Georissa scalinella" order="Cycloneritida" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="scalinella">Georissa scalinella</taxonomicName>
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has a series of scales at the shoulder. In habitus and scale characters, it resembles
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<taxonomicName id="3A143D8BCF8E2693FFE9F719FFA28CB7" lsidName="G. pyrrhoderma" pageId="0" pageNumber="1" rank="species" species="pyrrhoderma">G. pyrrhoderma</taxonomicName>
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from Gunung Silabur, Sarawak. The angular shoulder and small scale-like nodular structure at the intersection of strong spiral ribbings and growth lines are diagnostic for
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<taxonomicName id="83153102136EC03FF4DC1FCAEC4DC70A" lsidName="G. scalinella" pageId="0" pageNumber="1" rank="species" species="scalinella">G. scalinella</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="68005BED40F9310A763FF9785515516F" pageId="0" pageNumber="1" type="distribution">
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<paragraph id="671528556FA1FB12FCAC5F1565685BC9" pageId="0" pageNumber="1">Distribution.</paragraph>
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<paragraph id="CE4CDC3DBF86F2716F4828979F1BE775" pageId="0" pageNumber="1">
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Known only from the type locality, Teck Guan Estate, Lahad Datu, Sabah, and also reported by
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<bibRefCitation id="C04543C19204CB830AD62FE6B6C5AC6C" author="Phung, CC" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="49 - 104" title="A checklist of land snails from the west coast islands of Sabah, Borneo (Mollusca, Gastropoda)." url="https://doi.org/10.3897/zookeys.673.12422" volume="673" year="2017">Phung et al. (2017)</bibRefCitation>
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at Pulau Tiga, Sandakan, Sabah. However, this may also refer to one of the other "scaly group" species from Sabah.
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</paragraph>
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</subSubSection>
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<subSubSection id="C5A999A97006BC4FA5DA2991D2425B8D" pageId="0" pageNumber="1" type="discussion">
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<paragraph id="F01DA496E88C3237D2C36188E78D29E3" pageId="0" pageNumber="1">Discussion.</paragraph>
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<paragraph id="EFE7C17D681A85202539498ACD0505F6" pageId="0" pageNumber="1">
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<taxonomicName id="0D2EB4580A0E1128AB95D2691C110023" class="Gastropoda" family="Hydrocenidae" genus="Georissa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Georissa scalinella" order="Cycloneritida" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="scalinella">Georissa scalinella</taxonomicName>
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was first described as
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<taxonomicName id="49E4D485F36D3A5E842FCF626C99E95D" class="Gastropoda" family="Hydrocenidae" genus="Hydrocena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydrocena scalinella" order="Cycloneritida" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="scalinella">Hydrocena scalinella</taxonomicName>
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van Benthem-Jutting, 1966, before reclassified as
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<taxonomicName id="37B7E4049287278498234B8DCE95E9B5" class="Gastropoda" family="Hydrocenidae" genus="Georissa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Georissa" order="Cycloneritida" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Georissa</taxonomicName>
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by
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<bibRefCitation id="697EB2F9F48128E5F1DF81205A05710B" author="Thompson, FG" journalOrPublisher="Bulletin of the Florida State Museum Biological Sciences" pageId="0" pageNumber="1" pagination="101 - 152" title="Non-marine mollusks of Borneo. II Pulmonata: Pupillidae, Clausiliidae. III Prosobranchia: Hydrocenidae, Helicinidae." url="https://www.floridamuseum.ufl.edu/files/2114/7180/1931/Vol-29-No-3.PDF" volume="29" year="1983">Thompson and Dance (1983)</bibRefCitation>
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.
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<bibRefCitation id="51A19EBDD0F722B906EC750BD70F2BC9" author="van Benthem-Jutting, WSS" journalOrPublisher="Journal of Conchology" pageId="0" pageNumber="1" pagination="39 - 41" title="Two new species of Hydrocena (Neritacea) from Sabah, Borneo." volume="26" year="1966">van Benthem-Jutting (1966)</bibRefCitation>
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described
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<taxonomicName id="BBC18B9D57319FF36D2F3FBF5774B22B" lsidName="G. scalinella" pageId="0" pageNumber="1" rank="species" species="scalinella">G. scalinella</taxonomicName>
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as having strong spiral ribbing and multiple lines of scales.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |