treatments-xml/data/51/76/88/517688F4269755A0804D7256372126A0.xml

<document ID-DOI="http://dx.doi.org/10.3897/jhr.80.56178" ID-GBIF-Dataset="c03ce5c7-7805-4f32-95db-d4ce50995274" ID-Pensoft-Pub="1314-2607-80-99" ID-Pensoft-UUID="BCA40A8EFADF5E8A9E51A75C7C3C0D1C" ID-Zenodo-Dep="4420570" ID-ZooBank="F9DF28B5BF7545F5860B3FB234679C6D" ModsDocID="1314-2607-80-99" checkinTime="1609325082188" checkinUser="pensoft" docAuthor="Komeda, Yoto, Mita, Toshiharu, Hirose, Yoshimi &amp; Yamagishi, Kenzo" docDate="2020" docId="517688F4269755A0804D7256372126A0" docLanguage="en" docName="JourHymenoptRes 80: 99-135" docOrigin="Journal of Hymenoptera Research 80" docPubDate="2020-12-29" docSource="http://dx.doi.org/10.3897/jhr.80.56178" docTitle="Gryon shisa Komeda &amp; Mita 2020, sp. nov." docType="treatment" docUuid="45003564-D0AB-4E7B-A91A-E6495DE3ADB1" docUuidSource="ZooBank" docVersion="5" id="BCA40A8EFADF5E8A9E51A75C7C3C0D1C" lastPageNumber="99" masterDocId="BCA40A8EFADF5E8A9E51A75C7C3C0D1C" masterDocTitle="Taxonomic revision of charon -, floridanum - and muscaeforme - groups of Gryon Haliday, 1833 (Hymenoptera, Scelionidae) from Japan, with descriptions of two new species and host information" masterLastPageNumber="135" masterPageNumber="99" pageNumber="99" updateTime="1678754838156" updateUser="pensoft">
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<mods:title>Taxonomic revision of charon -, floridanum - and muscaeforme - groups of Gryon Haliday, 1833 (Hymenoptera, Scelionidae) from Japan, with descriptions of two new species and host information</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Komeda, Yoto</mods:namePart>
<mods:affiliation>Entomological Laboratory, Graduate School of Bioresource and Bioenvironmental Sciences, Kyushu University, Motooka 744, Fukuoka, 819 - 0395, Japan</mods:affiliation>
<mods:nameIdentifier type="email">kome123k123@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Mita, Toshiharu</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-8322-6045</mods:nameIdentifier>
<mods:affiliation>Entomological Laboratory, Graduate School of Bioresource and Bioenvironmental Sciences, Kyushu University, Motooka 744, Fukuoka, 819 - 0395, Japan</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hirose, Yoshimi</mods:namePart>
<mods:affiliation>Entomological Laboratory, Graduate School of Bioresource and Bioenvironmental Sciences, Kyushu University, Motooka 744, Fukuoka, 819 - 0395, Japan</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Yamagishi, Kenzo</mods:namePart>
<mods:affiliation>Entomological Laboratory, Faculty of Agriculture, Kyushu University, Motooka 744, Fukuoka, 819 - 0395, Japan</mods:affiliation>
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<mods:title>Journal of Hymenoptera Research</mods:title>
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<mods:date>2020</mods:date>
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<mods:number>2020-12-29</mods:number>
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<mods:detail type="volume">
<mods:number>80</mods:number>
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<mods:start>99</mods:start>
<mods:end>135</mods:end>
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<mods:url>http://dx.doi.org/10.3897/jhr.80.56178</mods:url>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/jhr.80.56178</mods:identifier>
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<treatment ID-GBIF-Taxon="176345127" LSID="urn:lsid:zoobank.org:act:45003564-D0AB-4E7B-A91A-E6495DE3ADB1" httpUri="http://treatment.plazi.org/id/517688F4269755A0804D7256372126A0" lastPageNumber="99" pageId="0" pageNumber="99">
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<paragraph pageId="0" pageNumber="99">
<taxonomicName LSID="http://zoobank.org/45003564-D0AB-4E7B-A91A-E6495DE3ADB1" authority="Komeda &amp; Mita" authorityName="Komeda &amp; Mita" authorityYear="2020" class="Insecta" family="Scelionidae" genus="Gryon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gryon shisa" order="Hymenoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="shisa" status="sp. nov.">Gryon shisa Komeda &amp; Mita</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="99">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="99" type="description">
<paragraph pageId="0" pageNumber="99">
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Japanese Gryon spp., dorsal view A G. fulvicoxa sp. nov., holotype B G. japonicum C G. yamagishii D G. philippinense E G. shisa sp. nov., holotype F G. pennsylvanicum. Scale bars: 1 mm." figureDoi="10.3897/jhr.80.56178.figure1" httpUri="https://binary.pensoft.net/fig/490334" pageId="0" pageNumber="99">Figs 1E</figureCitation>
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Japanese Gryon spp., lateral views A G. fulvicoxa sp. nov., holotype B G. japonicum C G. yamagishii D G. philippinense E G. shisa sp. nov., paratype F G. pennsylvanicum. Scale bars: 1 mm." figureDoi="10.3897/jhr.80.56178.figure2" httpUri="https://binary.pensoft.net/fig/490335" pageId="0" pageNumber="99">, 2E</figureCitation>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Frons of Japanese Gryon spp. A G. fulvicoxa sp. nov. B G. japonicum C G. yamagishii D G. philippinense E G. shisa sp. nov. F G. pennsylvanicum." figureDoi="10.3897/jhr.80.56178.figure3" httpUri="https://binary.pensoft.net/fig/490336" pageId="0" pageNumber="99">, 3E</figureCitation>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Occiput of Japanese Gryon spp. A G. fulvicoxa sp. nov. B G. japonicum C G. yamagishii D G. philippinense E G. shisa sp. nov. F G. pennsylvanicum." figureDoi="10.3897/jhr.80.56178.figure4" httpUri="https://binary.pensoft.net/fig/490337" pageId="0" pageNumber="99">, 4E</figureCitation>
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Antennae of Japanese Gryon spp. A G. fulvicoxa sp. nov., female B G. japonicum, female C G. yamagishii, female D G. philippinense, female E G. shisa sp. nov., female F G. pennsylvanicum, female G G. fulvicoxa sp. nov., male H G. japonicum, male I G. yamagishii, male J G. philippinense, male K G. shisa sp. nov., male L G. pennsylvanicum, male." figureDoi="10.3897/jhr.80.56178.figure5" httpUri="https://binary.pensoft.net/fig/490338" pageId="0" pageNumber="99">, 5E, K</figureCitation>
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Mesosoma of Japanese Gryon spp. A G. japonicum, dorsal view B lateral view C G. philippinense, dorsal view D lateral view E G. pennsylvanicum, dorsal view F lateral view." figureDoi="10.3897/jhr.80.56178.figure7" httpUri="https://binary.pensoft.net/fig/490340" pageId="0" pageNumber="99">, 7E</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="99" type="diagnosis">
<paragraph pageId="0" pageNumber="99">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="99">Horizontal portion of occipital carina curved, reaching central longitudinal line of lateral ocelli. Postgena smooth; postgenal sulcus straight; postgenal bridge smooth.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="99" type="description">
<paragraph pageId="0" pageNumber="99">Description.</paragraph>
<paragraph pageId="0" pageNumber="99">
<emphasis bold="true" pageId="0" pageNumber="99">Female.</emphasis>
Length = 1.6-1.7 mm.
</paragraph>
<paragraph pageId="0" pageNumber="99">
<emphasis bold="true" pageId="0" pageNumber="99">
<emphasis italics="true" pageId="0" pageNumber="99">Color</emphasis>
.
</emphasis>
(Figs 
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Japanese Gryon spp., dorsal view A G. fulvicoxa sp. nov., holotype B G. japonicum C G. yamagishii D G. philippinense E G. shisa sp. nov., holotype F G. pennsylvanicum. Scale bars: 1 mm." figureDoi="10.3897/jhr.80.56178.figure1" httpUri="https://binary.pensoft.net/fig/490334" pageId="0" pageNumber="99">1E</figureCitation>
, 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Japanese Gryon spp., lateral views A G. fulvicoxa sp. nov., holotype B G. japonicum C G. yamagishii D G. philippinense E G. shisa sp. nov., paratype F G. pennsylvanicum. Scale bars: 1 mm." figureDoi="10.3897/jhr.80.56178.figure2" httpUri="https://binary.pensoft.net/fig/490335" pageId="0" pageNumber="99">2E</figureCitation>
). 
<emphasis bold="true" italics="true" pageId="0" pageNumber="99">Body</emphasis>
mainly black. Mandibles brown. A1-6 and legs (excluding coxae) yellow.
</paragraph>
<paragraph pageId="0" pageNumber="99">
<emphasis bold="true" pageId="0" pageNumber="99">
<emphasis italics="true" pageId="0" pageNumber="99">Head</emphasis>
.
</emphasis>
FCI = 1.15-1.22; LCI = 1.51-1.62; DCI = 1.75-1.88; HW/IOS = 1.88-2.14; head about 1.2 times as wide as mesosoma (HW/TSL = 1.10-1.19). Frons reticulate with setae, with transverse carina between eye and frontal depression; central carina weakly present in lower half of frontal depression, absent in upper half of frontal depression, present between enclosing carina of frontal depression and anterior ocellus; frontal depression weakly developed, with enclosing carina; antennal depression about 1.3 times wider than distance between eye and antennal depression (WAD/OAD = 1.19-1.40). Vertex reticulate with setae; interocellar space reticulate-granulate; hyperoccipital carina present; POL about six times as long as OOL (POL/OOL = 5.54-6.55); OOL about 0.3 times as long as LOL (OOL/LOL = 0.26-0.32). Clypeus trapezoidal, with rounded corners. Gena coriaceous with setae; medial genal carina present. Occiput transversely semi-elliptically costate, with setae; occipital carina complete; angular point of occipital carina developed; horizontal portion of occipital carina curved, reaching central longitudinal line of lateral ocelli; postoccipital carina present laterally; postgena smooth; postgenal sulcus straight; postgenal bridge smooth. Antennae (Fig. 
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Antennae of Japanese Gryon spp. A G. fulvicoxa sp. nov., female B G. japonicum, female C G. yamagishii, female D G. philippinense, female E G. shisa sp. nov., female F G. pennsylvanicum, female G G. fulvicoxa sp. nov., male H G. japonicum, male I G. yamagishii, male J G. philippinense, male K G. shisa sp. nov., male L G. pennsylvanicum, male." figureDoi="10.3897/jhr.80.56178.figure5" httpUri="https://binary.pensoft.net/fig/490338" pageId="0" pageNumber="99">5E</figureCitation>
) clavate; A1 about 4.8 times longer than radicle, about 1.2 times longer than clava; clava with six segments; claval sensilla formula A7-12/1-2-2-2-2-1; claval length about 4.1 times longer than width. Mandibles thin, tridentate, anterior tooth longer than other teeth.
</paragraph>
<paragraph pageId="0" pageNumber="99">
<emphasis bold="true" pageId="0" pageNumber="99">
<emphasis italics="true" pageId="0" pageNumber="99">Mesosoma</emphasis>
.
</emphasis>
Cervical pronotal area costate dorsad, smooth-imbricate ventrad, with sparse setae; epomial carina strongly present, not reaching dorsal edge; pronotal suprahumeral sulcus foveolate with setae, unclear mesad; lateral pronotal area with transverse dense carina; pronotal cervical sulcus foveolete. Propleuron weakly transversely costate. Mesoscutum about 1.5 times as wide as long (TSL/ML = 1.44-1.61), reticulate; parascutal carina absent; notaulus absent. Mesoscutellum about 2.1 times as wide as long (SW/SL = 1.84-2.22), reticulate with setae, slightly produced posteriorly. Mesopleuron mesopleuron costate-reticulate above mesopleural canina, reticulate with setae below mesopleural canina; prespecular and upper mesepisternal sulci foveolate; prespecular sulcus with setae; mesopleural carina strongly present; postacetabular sulcus foveolate. Metascutellum weakly produced, striate. Metapleuron foveolate anteriorly, rugulose with dense setae posteriorly, with longitudinal irregular carina; anterior part of metapleural sulcus and upper paracoxal sulcus with setae. Propodeum foveolate, with setae laterad. Fore wing (Fig. 
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Mesosoma of Japanese Gryon spp. A G. japonicum, dorsal view B lateral view C G. philippinense, dorsal view D lateral view E G. pennsylvanicum, dorsal view F lateral view." figureDoi="10.3897/jhr.80.56178.figure7" httpUri="https://binary.pensoft.net/fig/490340" pageId="0" pageNumber="99">7E</figureCitation>
); stigmal vein about four times longer than marginal vein; postmarginal vein about 7.3 times longer than marginal vein.
</paragraph>
<paragraph pageId="0" pageNumber="99">
<emphasis bold="true" pageId="0" pageNumber="99">
<emphasis italics="true" pageId="0" pageNumber="99">Metasoma</emphasis>
.
</emphasis>
T1 longitudinally striate, setose laterally. S1 longitudinally striate. T2 reticulate with setae laterally. S2 reticulate-granulate, with setae. T3 reticulate with setae. T4 reticulate-rugose with setae. T5-6 rugose with setae. S3-6 punctate with setae
</paragraph>
<paragraph pageId="0" pageNumber="99">
<emphasis bold="true" pageId="0" pageNumber="99">Male.</emphasis>
Almost same as female, but antennae (Fig. 
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Antennae of Japanese Gryon spp. A G. fulvicoxa sp. nov., female B G. japonicum, female C G. yamagishii, female D G. philippinense, female E G. shisa sp. nov., female F G. pennsylvanicum, female G G. fulvicoxa sp. nov., male H G. japonicum, male I G. yamagishii, male J G. philippinense, male K G. shisa sp. nov., male L G. pennsylvanicum, male." figureDoi="10.3897/jhr.80.56178.figure5" httpUri="https://binary.pensoft.net/fig/490338" pageId="0" pageNumber="99">5K</figureCitation>
) filiform; A1-11 yellow.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="99" type="host">
<paragraph pageId="0" pageNumber="99">Host.</paragraph>
<paragraph pageId="0" pageNumber="99">
<taxonomicName authorityName="Leach" authorityYear="1815" class="Insecta" family="Coreidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Hemiptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="family">Coreidae</taxonomicName>
: 
<taxonomicName authorityName="Hsiao" authorityYear="1963" class="Insecta" family="Coreidae" genus="Paradasynus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Paradasynus spinosus" order="Hemiptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="spinosus">
<emphasis italics="true" pageId="0" pageNumber="99">Paradasynus spinosus</emphasis>
</taxonomicName>
Hsiao, 1963.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="99" type="materials_examined">
<paragraph pageId="0" pageNumber="99">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="99">
<materialsCitation ID-GBIF-Occurrence="3026168303" collectingDate="2002-07-30" collectingDateMax="1833-12-31" collectingDateMin="1833-01-01" collectionCode="ELKU" collectorName="Yasutsune Sadoyama" county="Ogimi" location="Okinawa" municipality="Kunigami Dist." specimenCount="1" specimenCount-female="1" typeStatus="Holotype">
<emphasis bold="true" italics="true" pageId="0" pageNumber="99">
<typeStatus>Holotype</typeStatus>
</emphasis>
: 
<collectingMunicipality>
<location LSID="urn:lsid:plazi:treatment:517688F4269755A0804D7256372126A0:38443DAD33115F1B7A6D19996F6CE8E0" county="Ogimi" municipality="Kunigami Dist." name="Okinawa">Okinawa</location>
pref
</collectingMunicipality>
., 
<location LSID="urn:lsid:plazi:treatment:517688F4269755A0804D7256372126A0:9E8DB428790C7AB2BDB2FD13ADD8436C" county="Ogimi" municipality="Kunigami Dist." name="Okinawa Isl.">Okinawa Isl.</location>
, 
<collectingMunicipality>
<location LSID="urn:lsid:plazi:treatment:517688F4269755A0804D7256372126A0:F59206112A76C05C2A0DCECE22EBA2AB" municipality="Okinawa pref" name="Kunigami Dist">Kunigami Dist</location>
.
</collectingMunicipality>
, 
<normalizedToken originalValue="Ôgimi">
<collectingCounty>
<location LSID="urn:lsid:plazi:treatment:517688F4269755A0804D7256372126A0:730D318C4B6969514B725DF426371AE3" municipality="Okinawa pref" name="Ogimi">Ogimi</location>
</collectingCounty>
</normalizedToken>
vill., 
<location LSID="urn:lsid:plazi:treatment:517688F4269755A0804D7256372126A0:72F366CE9C512EB8FB1955FF91CFECCA" municipality="Okinawa pref" name="Janagusuku">Janagusuku</location>
. 
<collectingDate value="2002-07-30">30.VII.2002</collectingDate>
, 
<collectorName>Yasutsune Sadoyama</collectorName>
leg. (emergence from an egg of 
<taxonomicName authorityName="Hsiao" authorityYear="1963" class="Insecta" family="Coreidae" genus="Paradasynus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Paradasynus spinosus" order="Hemiptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="spinosus">
<emphasis italics="true" pageId="0" pageNumber="99">Paradasynus spinosus</emphasis>
</taxonomicName>
) 
<specimenCount type="female">1♀</specimenCount>
[
<collectionCode>ELKU</collectionCode>
]
</materialsCitation>
. 
<materialsCitation collectingDate="2020-01-01" collectingDateMax="2020-12-31" collectingDateMin="2020-01-01" collectorName="Yasutsune Sadoyama" county="Ogimi" location="Okinawa" municipality="Kunigami Dist." specimenCount="10" specimenCount-female="7" specimenCount-male="3" typeStatus="Paratypes">
<emphasis bold="true" pageId="0" pageNumber="99">
<emphasis italics="true" pageId="0" pageNumber="99">
<typeStatus>Paratypes</typeStatus>
</emphasis>
.
</emphasis>
Same data as holotype. 
<specimenCount type="male">3♂</specimenCount>
<specimenCount type="female">7♀</specimenCount>
[ELKU]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="99" type="distribution">
<paragraph pageId="0" pageNumber="99">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="99">Japan (Ryukyus; Okinawa Is.)</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="99" type="etymology">
<paragraph pageId="0" pageNumber="99">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="99">
The species name refers to 
<normalizedToken originalValue="Shīsā">Shīsa</normalizedToken>
, the Okinawan traditional statue of the guardian lion, because this species defends the shequasar (
<taxonomicName authorityName="Hayata" authorityYear="1919" class="Magnoliopsida" family="Rutaceae" genus="Citrus" higherTaxonomySource="GBIF" isHybrid="true" kingdom="Plantae" lsidName="Citrus depressa" order="Sapindales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="species" species="depressa">
Citrus 
<normalizedToken originalValue="×">x</normalizedToken>
depressa
</taxonomicName>
Hayata), a kind of citrus fruit, from the important pest (
<taxonomicName authorityName="Hsiao" authorityYear="1963" class="Insecta" family="Coreidae" genus="Paradasynus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Paradasynus spinosus" order="Hemiptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="spinosus">
<emphasis italics="true" pageId="0" pageNumber="99">Paradasynus spinosus</emphasis>
</taxonomicName>
) on Okinawa Island. (
<bibRefCitation author="Zukeyama, H" journalOrPublisher="Plant Protection" pageId="0" pageNumber="99" pagination="430 - 434" publicationUrl="http://jppa.or.jp/archive/pdf/61_08_24.pdf" refId="B79" refString="Zukeyama, H, Sadoyama, Y, Teruya, T, 2007. Life history of Paradasynus spinosus (Heteroptera: Coreidae) in Okinawa, Japan. Plant Protection 61 (8): 430 - 434, http://jppa.or.jp/archive/pdf/61_08_24.pdf" title="Life history of Paradasynus spinosus (Heteroptera: Coreidae) in Okinawa, Japan." url="http://jppa.or.jp/archive/pdf/61_08_24.pdf" volume="61" year="2007">Zukeyama et al. 2007</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="99" type="remarks">
<paragraph pageId="0" pageNumber="99">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="99">
Among the species of the 
<taxonomicName authorityName="- group (Mineo" authorityYear="1983" class="Insecta" family="Scelionidae" genus="Gryon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gryon charon" order="Hymenoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="charon">
<emphasis italics="true" pageId="0" pageNumber="99">Gryon charon</emphasis>
</taxonomicName>
-group, 
<taxonomicName class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. shisa" order="Hymenoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="shisa">
<emphasis italics="true" pageId="0" pageNumber="99">G. shisa</emphasis>
</taxonomicName>
is the only species without sculpture on the postgena, except for the postgenal sulcus (Fig. 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Occiput of Japanese Gryon spp. A G. fulvicoxa sp. nov. B G. japonicum C G. yamagishii D G. philippinense E G. shisa sp. nov. F G. pennsylvanicum." figureDoi="10.3897/jhr.80.56178.figure4" httpUri="https://binary.pensoft.net/fig/490337" pageId="0" pageNumber="99">4E</figureCitation>
). This species also differs from 
<taxonomicName class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. philippinense" order="Hymenoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="philippinense">
<emphasis italics="true" pageId="0" pageNumber="99">G. philippinense</emphasis>
</taxonomicName>
in the shape and length of the postgenal sulcus (
<taxonomicName class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. shisa" order="Hymenoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="shisa">
<emphasis italics="true" pageId="0" pageNumber="99">G. shisa</emphasis>
</taxonomicName>
(Fig. 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Occiput of Japanese Gryon spp. A G. fulvicoxa sp. nov. B G. japonicum C G. yamagishii D G. philippinense E G. shisa sp. nov. F G. pennsylvanicum." figureDoi="10.3897/jhr.80.56178.figure4" httpUri="https://binary.pensoft.net/fig/490337" pageId="0" pageNumber="99">4E</figureCitation>
): straight; 
<taxonomicName class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. philippinense" order="Hymenoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="philippinense">
<emphasis italics="true" pageId="0" pageNumber="99">G. philippinense</emphasis>
</taxonomicName>
(Fig. 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Occiput of Japanese Gryon spp. A G. fulvicoxa sp. nov. B G. japonicum C G. yamagishii D G. philippinense E G. shisa sp. nov. F G. pennsylvanicum." figureDoi="10.3897/jhr.80.56178.figure4" httpUri="https://binary.pensoft.net/fig/490337" pageId="0" pageNumber="99">4D</figureCitation>
): curved toward hypostoma) and the length of horizontal portion of occipital carina (
<taxonomicName class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. shisa" order="Hymenoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="shisa">
<emphasis italics="true" pageId="0" pageNumber="99">G. shisa</emphasis>
</taxonomicName>
(Fig. 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Occiput of Japanese Gryon spp. A G. fulvicoxa sp. nov. B G. japonicum C G. yamagishii D G. philippinense E G. shisa sp. nov. F G. pennsylvanicum." figureDoi="10.3897/jhr.80.56178.figure4" httpUri="https://binary.pensoft.net/fig/490337" pageId="0" pageNumber="99">4E</figureCitation>
): long, reaching central longitudinal line of lateral ocelli; 
<taxonomicName class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. philippinense" order="Hymenoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="philippinense">
<emphasis italics="true" pageId="0" pageNumber="99">G. philippinense</emphasis>
</taxonomicName>
(Fig. 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Occiput of Japanese Gryon spp. A G. fulvicoxa sp. nov. B G. japonicum C G. yamagishii D G. philippinense E G. shisa sp. nov. F G. pennsylvanicum." figureDoi="10.3897/jhr.80.56178.figure4" httpUri="https://binary.pensoft.net/fig/490337" pageId="0" pageNumber="99">4D</figureCitation>
): short, reaching longitudinal extension line of outer margin of lateral ocelli or shorter). Among Vietnamese species, four species, 
<taxonomicName class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. ancinla" order="Hymenoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="ancinla">
<emphasis italics="true" pageId="0" pageNumber="99">G. ancinla</emphasis>
</taxonomicName>
Kozlov &amp; 
<normalizedToken originalValue="Lê">Le</normalizedToken>
, 1996, 
<taxonomicName class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. clavaerus" order="Hymenoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="clavaerus">
<emphasis italics="true" pageId="0" pageNumber="99">G. clavaerus</emphasis>
</taxonomicName>
Kozlov &amp; 
<normalizedToken originalValue="Lê">Le</normalizedToken>
, 1996, 
<taxonomicName class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. drunoris" order="Hymenoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="drunoris">
<emphasis italics="true" pageId="0" pageNumber="99">G. drunoris</emphasis>
</taxonomicName>
Kozlov &amp; 
<normalizedToken originalValue="Lê">Le</normalizedToken>
, 1996, and 
<taxonomicName class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. sponus" order="Hymenoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="sponus">
<emphasis italics="true" pageId="0" pageNumber="99">G. sponus</emphasis>
</taxonomicName>
Kozlov &amp; 
<normalizedToken originalValue="Lê">Le</normalizedToken>
, 1996 seem to belong to 
<taxonomicName authorityName="- group (Mineo" authorityYear="1983" class="Insecta" family="Scelionidae" genus="Gryon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gryon charon" order="Hymenoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="charon">
<emphasis italics="true" pageId="0" pageNumber="99">Gryon charon</emphasis>
</taxonomicName>
group, but their horizontal portions of occipital carinae is also short like 
<taxonomicName class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. philippinense" order="Hymenoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="philippinense">
<emphasis italics="true" pageId="0" pageNumber="99">G. philippinense</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>