266 lines
35 KiB
XML
266 lines
35 KiB
XML
<document ID-CLB-Dataset="21464" ID-DOI="http://dx.doi.org/10.3897/zookeys.949.51269" ID-GBIF-Dataset="e4d2c425-4596-487c-8c4f-b09330b48203" ID-PMC="PMC7381721" ID-Pensoft-Pub="1313-2970-949-1" ID-Pensoft-UUID="303D64593B3B550DBB7CE01AA547409D" ID-PubMed="32765176" ID-ZooBank="93BFA448BB7343CDBC4EA86090DA63D5" ModsDocID="1313-2970-949-1" checkinTime="1594832489974" checkinUser="pensoft" docAuthor="Salata, Sebastian & Fisher, Brian L." docDate="2020" docId="333BE617A4F557CA8050326F1FBCB9F6" docLanguage="en" docName="ZooKeys 949: 1-185" docOrigin="ZooKeys 949" docSource="http://dx.doi.org/10.3897/zookeys.949.51269" docTitle="Pheidole itremo Salata & Fisher 2020, sp. nov." docType="treatment" docUuid="3E990650-E613-475B-8C4B-E16C43898146" docUuidSource="ZooBank" docVersion="6" id="303D64593B3B550DBB7CE01AA547409D" lastPageNumber="1" masterDocId="303D64593B3B550DBB7CE01AA547409D" masterDocTitle="Taxonomic revision of the Pheidole sikorae species group (Hymenoptera, Formicidae) from Madagascar" masterLastPageNumber="185" masterPageNumber="1" pageNumber="1" updateTime="1732651982216" updateUser="ExternalLinkService">
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<mods:titleInfo id="5C96A2AA6C505385DCC22F5D54BFE064">
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<mods:title id="E11952FEE00FF8F988BA0784FFB7E3A1">Taxonomic revision of the Pheidole sikorae species group (Hymenoptera, Formicidae) from Madagascar</mods:title>
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<mods:name id="3D6BB6C9D8A7F46AB6254364DCC4E363" type="personal">
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<mods:roleTerm id="211A86ACE09CF3F4BBC73AB2F77CA650">Author</mods:roleTerm>
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<mods:namePart id="87AD9363E915AA7D3E5AB4CE20216CB2">Salata, Sebastian</mods:namePart>
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<mods:affiliation id="2F7D637E68E63165553271B15CD9879A">Department of Entomology, California Academy of Sciences, San Francisco, CA 94118, USA</mods:affiliation>
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<mods:nameIdentifier id="623244F67541269680A4ABEF28928233" type="orcid">https://orcid.org/0000-0003-0811-2309</mods:nameIdentifier>
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<mods:nameIdentifier id="68CB8FB5A4D419DDDAED8DEF177D46E4" type="email">sdsalata@gmail.com</mods:nameIdentifier>
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<mods:namePart id="E3857419FFFA53B7BB7BC89F7E95BD19">Fisher, Brian L.</mods:namePart>
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<mods:affiliation id="345BA609D69CC078636C78349550080D">Department of Entomology, California Academy of Sciences, San Francisco, CA 94118, USA</mods:affiliation>
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<mods:nameIdentifier id="B7BF51DF19F2096D69C746AC462DEC90" type="orcid">https://orcid.org/0000-0002-4653-3270</mods:nameIdentifier>
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<mods:title id="D3ED4D972EAD52F2F64E10E7016E22D3">ZooKeys</mods:title>
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<mods:part id="98FDDE85EF24D0D2A31031F6C6F3DEE3">
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<mods:date id="55CB27FC90CC4DD61936955BCB27AF67">2020</mods:date>
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<mods:detail id="DE5D7E967A2E8D7810DC2F60B311F91D" type="volume">
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<mods:number id="56B1B2EC18E36D78B0FD1DFB4DC62A50">949</mods:number>
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<mods:url id="3A5E727E984731C9F10D8222954DC699">http://dx.doi.org/10.3897/zookeys.949.51269</mods:url>
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<mods:classification id="660774938A155DF041451933FFD33443">journal article</mods:classification>
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<mods:identifier id="6DCCB2023737AFB71E9EA9F250570118" type="DOI">http://dx.doi.org/10.3897/zookeys.949.51269</mods:identifier>
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<treatment id="333BE617A4F557CA8050326F1FBCB9F6" ID-GBIF-Taxon="165090190" LSID="urn:lsid:zoobank.org:act:3E990650-E613-475B-8C4B-E16C43898146" httpUri="http://treatment.plazi.org/id/333BE617A4F557CA8050326F1FBCB9F6" lastPageNumber="1" pageId="0" pageNumber="1" scope_family="Formicidae" scope_order="Hymenoptera">
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<subSubSection id="7091A83CFF4343AB0313C3DA10A00012" pageId="0" pageNumber="1" type="nomenclature">
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<paragraph id="68BF5393613C38596B051F54ECDC128B" pageId="0" pageNumber="1">
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<taxonomicName id="42019DB5C37EA2CF62FF1AD7897451C2" ID-CoL="CMLB9" LSID="333BE617-A4F5-57CA-8050-326F1FBCB9F6" authority="Salata & Fisher, 2020" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole itremo" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="itremo" status="sp. nov.">Pheidole itremo</taxonomicName>
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<taxonomicNameLabel id="2E176FDEA84B97FC3C0DF938EB4AF631" pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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<figureCitation id="E2D78EC229806246956173B4C11D969D" captionStart="Figure 34" captionStartId="F34" captionText="Figure 34. Pheidole itremo sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0491586) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0491584)." figureDoi="10.3897/zookeys.949.51269.figure34" httpUri="https://binary.pensoft.net/fig/432203" pageId="0" pageNumber="1">Figs 34A-F</figureCitation>
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<figureCitation id="ABFCA02B2DFE769EA5789AB8055E23E3" captionStart="Figure 63" captionStartId="F63" captionText="Figure 63. Major worker, hypostomal teeth. Pheidole alina sp. nov. (A). Pheidole ambohimanga sp. nov. (B). Pheidole analavelona sp. nov. (C). Pheidole andohahela sp. nov. (D). Pheidole anomala sp. nov. (E). Pheidole anosyenne sp. nov. (F). Pheidole antranohofa sp. nov. (G). Pheidole beanka sp. nov. (H). Pheidole befotaka sp. nov. (I). Pheidole dasos sp. nov. (J). Pheidole flavominuta sp. nov. (K). Pheidole gracilis sp. nov. (L). Pheidole haboka sp. nov. (M). Pheidole havoana sp. nov. (N). Pheidole hazo sp. nov. (O). Pheidole itremo sp. nov. (P). Pheidole joffreville sp. nov. (Q). Pheidole kely sp. nov. (R). Pheidole lavasoa sp. nov. (S). Pheidole litigiosa Forel (T). Pheidole mahamavo sp. nov. (U). Pheidole mainty sp. nov. (V). Pheidole mamiratra sp. nov. (W). Pheidole manantenina sp. nov. (X)." figureDoi="10.3897/zookeys.949.51269.figure63" httpUri="https://binary.pensoft.net/fig/432232" pageId="0" pageNumber="1">, 63P</figureCitation>
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<figureCitation id="F95D8C9E785FB7845C42540847FBB306" captionStart="Figure 65" captionStartId="F65" captionText="Figure 65. Distribution. Pheidole alina sp. nov. (A). Pheidole ambohimanga sp. nov. (B). Pheidole analavelona sp. nov. (C). Pheidole andohahela sp. nov. (D). Pheidole anomala sp. nov. (E). Pheidole anosyenne sp. nov. (F). Pheidole antranohofa sp. nov. (G). Pheidole beanka sp. nov. (H). Pheidole befotaka sp. nov. (I). Pheidole dasos sp. nov. (J). Pheidole flavominuta sp. nov. (K). Pheidole gracilis sp. nov. (L). Pheidole haboka sp. nov. (M). Pheidole havoana sp. nov. (N). Pheidole hazo sp. nov. (O). Pheidole itremo sp. nov. (P). Pheidole joffreville sp. nov. (Q). Pheidole kely sp. nov. (R). Pheidole lavasoa sp. nov. (S). Pheidole litigiosa Forel (T)." figureDoi="10.3897/zookeys.949.51269.figure65" httpUri="https://binary.pensoft.net/fig/432234" pageId="0" pageNumber="1">, 65P</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="A8A2AB1DC3AC2940DA5CD7D88B3AA7FD" pageId="0" pageNumber="1" type="type material">
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<paragraph id="AEEF802124AAD2332B68C4F814B317A1" pageId="0" pageNumber="1">Type material.</paragraph>
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<paragraph id="086258149B72F4EF7F4CD614D3C80618" pageId="0" pageNumber="1">
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<emphasis id="383ADDC612DD0740D7548840A258AD7C" bold="true" italics="true" pageId="0" pageNumber="1">Holotype.</emphasis>
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Madagascar. • 1 major worker; Fianarantsoa;
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<normalizedToken id="04CFDA49A9FE2DE15B5C590B79F23F75" originalValue="Forêt">Foret</normalizedToken>
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<normalizedToken id="5EB6DBF9468C85D546C1EA151FA5694E" originalValue="d’Atsirakambiaty">d'Atsirakambiaty</normalizedToken>
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, 7.6 km 285°WNW Itremo;
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<geoCoordinate id="7A312B2E83A7DADBCD86A12F577F11B2" degrees="20.59333" direction="south" orientation="latitude" precision="1" value="-20.59333">-20.59333</geoCoordinate>
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,
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<geoCoordinate id="41968837854614B7A4C0546DC1E71E20" degrees="46.56333" direction="east" orientation="longitude" precision="1" value="46.56333">46.56333</geoCoordinate>
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; alt. 1550 m; 22 Jan 2003; Fisher et al. leg.; montane rainforest, ex rotten log; BLF07251; CASENT0491584, top specimen on the pin (CASC).
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<emphasis id="A993D49CC5BF908FE59166AFB004E0A4" bold="true" italics="true" pageId="0" pageNumber="1">Paratypes.</emphasis>
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• 6w., 1s.; same data as for holotype, CASENT0872249, CASENT0491586, CASENT0491585 (CASC, MHNG, PBZT).
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</paragraph>
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<caption id="79F034B3741A62F3127CAE952E74F181" doi="10.3897/zookeys.949.51269.figure34" httpUri="https://binary.pensoft.net/fig/432203" pageId="0" pageNumber="1" start="Figure 34" startId="F34">
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<paragraph id="CF0F3CF2E8E7A3C5E16B0AFCDA1972EE" pageId="0" pageNumber="1">
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<emphasis id="FD45324147258328F88596A0F9D8EA07" bold="true" pageId="0" pageNumber="1">Figure 34.</emphasis>
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<taxonomicName id="AE8AEC2C38297CA9655DD736333FDF88" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole itremo" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="itremo">
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<emphasis id="F73AA5DD0F15E4EB852BA4D460E01374" italics="true" pageId="0" pageNumber="1">Pheidole itremo</emphasis>
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</taxonomicName>
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sp. nov., full-face view (
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<emphasis id="D585408B05ABEC412B73879810C5836B" bold="true" pageId="0" pageNumber="1">A</emphasis>
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), profile (
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<emphasis id="9BECD8D51DB138C3F2C0241087B7E4C7" bold="true" pageId="0" pageNumber="1">C</emphasis>
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), and dorsal view (
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<emphasis id="5B4960038C0D7FB365C82F7821A2F53E" bold="true" pageId="0" pageNumber="1">E</emphasis>
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) of paratype minor worker (CASENT0491586) and full-face view (
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<emphasis id="CA7983332AE8D74F6F13C1A8F8F0169E" bold="true" pageId="0" pageNumber="1">B</emphasis>
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), profile (
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<emphasis id="88083273FA8F650A071585303BDC8555" bold="true" pageId="0" pageNumber="1">D</emphasis>
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), and dorsal view (
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<emphasis id="FAE7431DAAA949F649C91E094F769034" bold="true" pageId="0" pageNumber="1">F</emphasis>
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) of holotype major worker (CASENT0491584).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection id="EA253EDC384600E5A8C6A75C6035A475" pageId="0" pageNumber="1" type="materials_examined">
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<paragraph id="D52B37A29B1D6F0919720C706D6B5377" pageId="0" pageNumber="1">Other material.</paragraph>
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<paragraph id="04396A35B48C5B5DB12BC5E6595DDEFA" pageId="0" pageNumber="1">
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Madagascar. -
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<emphasis id="BF2E521CB3269C28BB9EB1CC4BEDE7FB" bold="true" pageId="0" pageNumber="1">Antananarivo</emphasis>
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: • 1w., 2s.;
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<normalizedToken id="6F361EF1E8B2BF2956BA2C46BE35F242" originalValue="Réserve">Reserve</normalizedToken>
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||
<normalizedToken id="F87F73563772A9471CA15A10E0B312DF" originalValue="Spéciale">Speciale</normalizedToken>
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<normalizedToken id="4DA231FB34880EE1715FCFACF49B9FD0" originalValue="d’Ambohitantely">d'Ambohitantely</normalizedToken>
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,
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<normalizedToken id="4784EAA34D9E43E121947363B155588F" originalValue="Forêt">Foret</normalizedToken>
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d Ambohitantely, 20.9 km 72°NE d Ankazobe;
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<geoCoordinate id="C282605EC9EC3F8721B658908FA9D23D" degrees="18.22528" direction="south" orientation="latitude" precision="1" value="-18.22528">-18.22528</geoCoordinate>
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,
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<geoCoordinate id="1D04D35B56D1B9C6A96E5B62959A6B73" degrees="47.28683" direction="east" orientation="longitude" precision="1" value="47.28683">47.28683</geoCoordinate>
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; alt. 1410 m; 17 Apr 2001; Fisher et al. leg.; montane rainforest, ex root mat, ground layer; BLF03700 (CASC). • 3s.;
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<normalizedToken id="46B742ADCE79820182A5AD49D3D12FC4" originalValue="Réserve">Reserve</normalizedToken>
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<normalizedToken id="657CB92A745FB2908C79D44AFC148630" originalValue="d’Ambohitantely">d'Ambohitantely</normalizedToken>
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,
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<normalizedToken id="7FD9239C5DC2598449F2B82768B99BFA" originalValue="Forêt">Foret</normalizedToken>
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d Ambohitantely, Jardin Botanique, 24.1km 59°NE d Ankazobe;
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<geoCoordinate id="4CC703428C78F25E0FD9864D5CB08663" degrees="18.17139" direction="south" orientation="latitude" precision="1" value="-18.17139">-18.17139</geoCoordinate>
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,
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<geoCoordinate id="337E7D891B99024ED7D11696D8F84059" degrees="47.28182" direction="east" orientation="longitude" precision="1" value="47.28182">47.28182</geoCoordinate>
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; alt. 1620 m; 17 Apr 2001; Fisher et al. leg.; montane rainforest, sifted litter; BLF03720 (CASC). • 2w.;
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<normalizedToken id="D2CDCCA5BBBB6EB9A3944435AFB5A067" originalValue="Réserve">Reserve</normalizedToken>
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Speciale
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<normalizedToken id="1A622400339C83D04F566B463235584B" originalValue="d’Ambohitantely">d'Ambohitantely</normalizedToken>
|
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;
|
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<geoCoordinate id="B6BC6587B24387E152040F98075C1AD5" degrees="18.18762" direction="south" orientation="latitude" precision="1" value="-18.18762">-18.18762</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="26A9F4CB67902043664646FE4BBFA2A9" degrees="47.28576" direction="east" orientation="longitude" precision="1" value="47.28576">47.28576</geoCoordinate>
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; alt. 1580 m; 8 Mar 2012; Fisher et al. leg.; montane forest, ex rotten log; BLF28200 (CASC). • 3w., 1q.;
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<normalizedToken id="066D6278E6C0DECFA5A2BA39E615D40C" originalValue="Réserve">Reserve</normalizedToken>
|
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Speciale
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<normalizedToken id="3AFBF72B16D1791444435CF3A100A93F" originalValue="d’Ambohitantely">d'Ambohitantely</normalizedToken>
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;
|
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<geoCoordinate id="915B56E1A473CFC4E5535031651DC388" degrees="18.18762" direction="south" orientation="latitude" precision="1" value="-18.18762">-18.18762</geoCoordinate>
|
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,
|
||
<geoCoordinate id="9CD93F6BC87AC4105B00797711913FFE" degrees="47.28576" direction="east" orientation="longitude" precision="1" value="47.28576">47.28576</geoCoordinate>
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; alt. 1580 m; 8 Mar 2012; Fisher et al. leg.; montane forest, ex rotten log; BLF28206 (CASC). • 2w.;
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<normalizedToken id="A0B813084C9A7745FE46AD92622F254A" originalValue="Réserve">Reserve</normalizedToken>
|
||
Speciale
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||
<normalizedToken id="35CEC5DAFAC4EE2B8F6A5CD69761619E" originalValue="d’Ambohitantely">d'Ambohitantely</normalizedToken>
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||
;
|
||
<geoCoordinate id="0001DA9611951F197B5C227D088AFD26" degrees="18.18762" direction="south" orientation="latitude" precision="1" value="-18.18762">-18.18762</geoCoordinate>
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,
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||
<geoCoordinate id="B8E3E1555E2CE49D6F4D3144C58FB0A2" degrees="47.28576" direction="east" orientation="longitude" precision="1" value="47.28576">47.28576</geoCoordinate>
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; alt. 1580 m; 8 Mar 2012; Fisher et al. leg.; montane forest, ex rotten log; BLF28218 (CASC). • 3w., 1q.;
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<normalizedToken id="5E03717BAE059B9BFC0967F9B59DE94C" originalValue="Réserve">Reserve</normalizedToken>
|
||
Speciale
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||
<normalizedToken id="0714910BBC9935E0009B9D6A22C80F0A" originalValue="d’Ambohitantely">d'Ambohitantely</normalizedToken>
|
||
;
|
||
<geoCoordinate id="AD39ABEF3924B76777904FABBA5106A9" degrees="18.18762" direction="south" orientation="latitude" precision="1" value="-18.18762">-18.18762</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="781E16C83F92CEC49748E918E2F28525" degrees="47.28576" direction="east" orientation="longitude" precision="1" value="47.28576">47.28576</geoCoordinate>
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; alt. 1580 m; 8 Mar 2012; Fisher et al. leg.; montane forest, ex rotten log; BLF28227 (CASC).
|
||
</paragraph>
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</subSubSection>
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<subSubSection id="DC5DBFC77BE68DE62122ACBC2AAA7410" pageId="0" pageNumber="1" type="diagnosis">
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<paragraph id="311692ECB0F80A34C0396D8D7E735943" pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph id="9C4486D9545878250D0E83768A9DF85D" pageId="0" pageNumber="1">
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Moderately large species.
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<emphasis id="F33CE1F635EE851CC38E066E6E9288F2" bold="true" italics="true" pageId="0" pageNumber="1">Major workers.</emphasis>
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||
Head in full-face view sub-oval, widening posteriorly, with anterior and posterior sides convex, in lateral view sub-oval; ventral and dorsal faces convex; sides of the head with moderately dense, moderately long, suberect to erect pilosity; medial part of frons with thick, dense, longitudinal, and interrupted rugae and smooth interspaces, posteromedial frons with more irregular rugae directed slightly outward; lateral sides with thick and dense rugae, interspaces shiny and smooth or indistinctly rugulate; rugae longitudinal on anterolateral sides and more irregular on posterolateral sides; area posterolateral from eyes without smooth notches; scape, when laid back, exceeding the midlength of head by one-fifth of its length; inner hypostomal teeth distinct, large, closely spaced, triangular, with rounded apex directed upward; outer hypostomal teeth lobe-like, wider than inner teeth and approximately the same height; inner and outer hypostomal teeth closely spaced and connected by indistinct concavity; mesosoma with fine rugofoveolae; pronotum with rugofoveolae sparser and additional thin and transverse rugae; gaster smooth; body brown.
|
||
<emphasis id="BD8BFF7D71A8F4EED4670E3E639C04EE" bold="true" italics="true" pageId="0" pageNumber="1">Minor workers.</emphasis>
|
||
Head smooth; posterolateral sides of frons with sparse and indistinct foveolae; scape, when laid back, surpassing the posterior head margin by two-fifths of its length; promesonotum moderately high and short, arched; promesonotal groove absent; propodeal spines minute and triangular; mesosoma smooth and shiny; body yellowish brown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DA906705A56F59BE4AC99D47F8726295" pageId="0" pageNumber="1" type="description">
|
||
<paragraph id="C62D27390A3A3903731D95C9E753206B" pageId="0" pageNumber="1">Description.</paragraph>
|
||
<paragraph id="CA77EF7CC7B4ECBB37F58295ED1AC0DE" pageId="0" pageNumber="1">
|
||
<emphasis id="675F96EFFDCEC1486EC18125FBB4ADDF" bold="true" pageId="0" pageNumber="1">Major workers.</emphasis>
|
||
Measurements (
|
||
<emphasis id="E6A9D1608E02D6F57171DB3108162600" italics="true" pageId="0" pageNumber="1">N</emphasis>
|
||
= 6): HL: 1.14-1.24 (1.18); HW: 1.16-1.21 (1.17); SL: 0.64-0.69 (0.66); EL: 0.12-0.14 (0.13); WL: 0.99-1.06 (1.03); PSL: 0.18-0.2 (0.19); MTL: 0.65-0.68 (0.66); PNW: 0.51-0.54 (0.52); PTW: 0.15-0.17 (0.16); PPW: 0.28-0.34 (0.31); CI: 98.3-103.3 (100.8); SI: 53.7-58.8 (56.0); PSLI: 15.5-16.9 (16.1); PPI: 43.0-56.6 (51.5); PNI: 42.7-46.2 (44.3); MTI: 54.6-57.7 (56.6).
|
||
</paragraph>
|
||
<paragraph id="7FED728701FE4F583EE0193953D5FDEE" pageId="0" pageNumber="1">
|
||
<emphasis id="5B85B22B33CD5F91B0742FF53CC1D7CD" bold="true" italics="true" pageId="0" pageNumber="1">Head.</emphasis>
|
||
In full-face view sub-oval, widening posteriorly, with anterior and posterior sides convex (Fig.
|
||
<figureCitation id="4941B21DB6F15674A6B7E6EC5FB0C3C4" captionStart="Figure 34" captionStartId="F34" captionText="Figure 34. Pheidole itremo sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0491586) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0491584)." figureDoi="10.3897/zookeys.949.51269.figure34" httpUri="https://binary.pensoft.net/fig/432203" pageId="0" pageNumber="1">34B</figureCitation>
|
||
). In lateral view sub-oval; ventral and dorsal faces convex; inner hypostomal teeth visible. Sides of the head with moderately dense, moderately long, suberect to erect pilosity; whole head with dense, long, decumbent to erect pilosity. Medial part of frons with thick, dense, longitudinal, and interrupted rugae and smooth interspaces, posteromedial frons with rugae more irregular and directed slightly outward; lateral sides with thick and dense rugae, interspaces shiny and smooth or indistinctly rugulate; rugae longitudinal on anterolateral sides and more irregular on posterolateral sides. Occipital lobes with thinner, irregular rugae and smooth to indistinctly rugoreticulate interspaces. Area posterolateral from eyes with irregular to longitudinal, relatively thick to thin rugulae and rugoreticulate interspaces. Gena with relatively dense, thick, longitudinal rugae and smooth to indistinctly rugoreticulate interspaces. Centre of clypeus smooth and shiny, lateral sides with indistinct rugulae; median notch present, moderately wide, and shallow; median longitudinal carina present; lateral longitudinal carinae absent. Scape, when laid back, exceeding the midlength of head by one-fifth of its length; pilosity subdecumbent to erect (Fig.
|
||
<figureCitation id="382AB7EBA7597289A349317CE815B9E8" captionStart="Figure 34" captionStartId="F34" captionText="Figure 34. Pheidole itremo sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0491586) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0491584)." figureDoi="10.3897/zookeys.949.51269.figure34" httpUri="https://binary.pensoft.net/fig/432203" pageId="0" pageNumber="1">34B, D</figureCitation>
|
||
). Inner hypostomal teeth distinct, large, closely spaced, triangular, with rounded apex directed upward; outer hypostomal teeth lobe-like, wider than inner teeth and approximately the same height; inner and outer hypostomal teeth closely spaced and connected by indistinct concavity (Fig.
|
||
<figureCitation id="1CF0B95582FDC06629AD154AF2114BD4" captionStart="Figure 63" captionStartId="F63" captionText="Figure 63. Major worker, hypostomal teeth. Pheidole alina sp. nov. (A). Pheidole ambohimanga sp. nov. (B). Pheidole analavelona sp. nov. (C). Pheidole andohahela sp. nov. (D). Pheidole anomala sp. nov. (E). Pheidole anosyenne sp. nov. (F). Pheidole antranohofa sp. nov. (G). Pheidole beanka sp. nov. (H). Pheidole befotaka sp. nov. (I). Pheidole dasos sp. nov. (J). Pheidole flavominuta sp. nov. (K). Pheidole gracilis sp. nov. (L). Pheidole haboka sp. nov. (M). Pheidole havoana sp. nov. (N). Pheidole hazo sp. nov. (O). Pheidole itremo sp. nov. (P). Pheidole joffreville sp. nov. (Q). Pheidole kely sp. nov. (R). Pheidole lavasoa sp. nov. (S). Pheidole litigiosa Forel (T). Pheidole mahamavo sp. nov. (U). Pheidole mainty sp. nov. (V). Pheidole mamiratra sp. nov. (W). Pheidole manantenina sp. nov. (X)." figureDoi="10.3897/zookeys.949.51269.figure63" httpUri="https://binary.pensoft.net/fig/432232" pageId="0" pageNumber="1">63P</figureCitation>
|
||
).
|
||
<emphasis id="20437D5F9684CBC8E09989C439487BFD" bold="true" italics="true" pageId="0" pageNumber="1">Mesosoma.</emphasis>
|
||
In lateral view, promesonotum short, angular, and moderately low, posterior mesonotum moderately steep, mesonotal process very indistinct, tubercle-like; promesonotal groove absent; metanotal groove absent; propodeal spines moderately long, with moderately narrow base and acute apex; humeral area laterally weakly produced (Fig.
|
||
<figureCitation id="B91A2822F6A9797671E1A14D11746F2F" captionStart="Figure 34" captionStartId="F34" captionText="Figure 34. Pheidole itremo sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0491586) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0491584)." figureDoi="10.3897/zookeys.949.51269.figure34" httpUri="https://binary.pensoft.net/fig/432203" pageId="0" pageNumber="1">34D</figureCitation>
|
||
). Surface shiny with fine rugofoveolae; pronotum with rugofoveolae sparser and additional thin and transverse rugae. Pilosity moderately sparse, long, and erect (Fig.
|
||
<figureCitation id="11C71768DBB043DEC608D0BFB66473B9" captionStart="Figure 34" captionStartId="F34" captionText="Figure 34. Pheidole itremo sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0491586) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0491584)." figureDoi="10.3897/zookeys.949.51269.figure34" httpUri="https://binary.pensoft.net/fig/432203" pageId="0" pageNumber="1">34D, F</figureCitation>
|
||
).
|
||
<emphasis id="9BE93720FB7C33FD385E5AAC87218DFE" bold="true" italics="true" pageId="0" pageNumber="1">Petiole.</emphasis>
|
||
Shiny with fine foveolae; node smooth to finely foveolate, triangular, with rounded and thick apex, in rear view node dorsoventrally straight to slightly convex; pilosity moderately sparse and erect (Fig.
|
||
<figureCitation id="075E65E79EA66E36C680A63D49FE5FC3" captionStart="Figure 34" captionStartId="F34" captionText="Figure 34. Pheidole itremo sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0491586) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0491584)." figureDoi="10.3897/zookeys.949.51269.figure34" httpUri="https://binary.pensoft.net/fig/432203" pageId="0" pageNumber="1">34D, F</figureCitation>
|
||
).
|
||
<emphasis id="8B2E1290BBD039EFC7EA0A79CA230534" bold="true" italics="true" pageId="0" pageNumber="1">Postpetiole.</emphasis>
|
||
Shiny and foveolate; dorsum with reduced sculpture and smooth notch; in dorsal view oval, lateral margins medially with two dentate projections; pilosity long, moderately sparse, and erect (Fig.
|
||
<figureCitation id="2D6DF65F5E604734D5E1DB5388020A79" captionStart="Figure 34" captionStartId="F34" captionText="Figure 34. Pheidole itremo sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0491586) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0491584)." figureDoi="10.3897/zookeys.949.51269.figure34" httpUri="https://binary.pensoft.net/fig/432203" pageId="0" pageNumber="1">34D, F</figureCitation>
|
||
).
|
||
<emphasis id="A4076CCF273BC178276D33299A57B4D5" bold="true" italics="true" pageId="0" pageNumber="1">Gaster.</emphasis>
|
||
Shiny and smooth; pilosity dense, long, and erect (Fig.
|
||
<figureCitation id="3F4BF2FECBFE0AFAA8A7DC9AA9902D57" captionStart="Figure 34" captionStartId="F34" captionText="Figure 34. Pheidole itremo sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0491586) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0491584)." figureDoi="10.3897/zookeys.949.51269.figure34" httpUri="https://binary.pensoft.net/fig/432203" pageId="0" pageNumber="1">34D, F</figureCitation>
|
||
).
|
||
<emphasis id="DE1535A01A15507136B2767E52A3460A" bold="true" italics="true" pageId="0" pageNumber="1">Colour.</emphasis>
|
||
Brown; legs yellow (Fig.
|
||
<figureCitation id="A12D01184AC1AB650D290E932AF74F53" captionStart="Figure 34" captionStartId="F34" captionText="Figure 34. Pheidole itremo sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0491586) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0491584)." figureDoi="10.3897/zookeys.949.51269.figure34" httpUri="https://binary.pensoft.net/fig/432203" pageId="0" pageNumber="1">34D, F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="5B0AAFDF4990AA8BD7691409518AC212" pageId="0" pageNumber="1">
|
||
<emphasis id="C698B015F82B52610432D5366BAFA382" bold="true" pageId="0" pageNumber="1">Minor workers.</emphasis>
|
||
Measurements (
|
||
<emphasis id="65FF9CAF57A8E6D48B557CC920C45081" italics="true" pageId="0" pageNumber="1">N</emphasis>
|
||
= 10): HL: 0.54-0.59 (0.56); HW: 0.46-0.5 (0.48); SL: 0.59-0.63 (0.6); EL: 0.1-0.13 (0.11); WL: 0.68-0.74 (0.71); PSL: 0.06-0.1 (0.08); MTL: 0.45-0.5 (0.47); PNW: 0.3-0.35 (0.32); PTW: 0.08-0.1 (0.08); PPW: 0.13-0.16 (0.14); CI: 115.7-121.8 (117.9); SI: 120.4-131.4 (125.6); PSLI: 11.8-17.0 (14.0); PPI: 49.4-67.0 (58.5); PNI: 65.8-71.0 (68.2); MTI: 94.5-101.7 (99.2).
|
||
</paragraph>
|
||
<paragraph id="15C7D56BC7650C5C87C1A1ACF0E4A0C9" pageId="0" pageNumber="1">
|
||
<emphasis id="4FDE9EBED2D7AE1C9E91D5B14E63945F" bold="true" italics="true" pageId="0" pageNumber="1">Head.</emphasis>
|
||
Cephalic margin indistinctly convex (Fig.
|
||
<figureCitation id="8BA8FCB418A0401AEE447E0710AD63DF" captionStart="Figure 34" captionStartId="F34" captionText="Figure 34. Pheidole itremo sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0491586) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0491584)." figureDoi="10.3897/zookeys.949.51269.figure34" httpUri="https://binary.pensoft.net/fig/432203" pageId="0" pageNumber="1">34A</figureCitation>
|
||
). Pilosity relatively sparse, long, decumbent to suberect. Sculpture shiny and smooth; posterolateral sides of frons with sparse and indistinct foveolae; antennal sockets with few thick, curved outward rugae and smooth interspaces. Clypeus with median longitudinal carina absent; two lateral longitudinal carinae absent. Scape, when laid back, surpassing the posterior head margin by two-fifths of its length; pilosity dense, subdecumbent to erect (Fig.
|
||
<figureCitation id="AC77C1EAFA3EA968981ADCF019519D0F" captionStart="Figure 34" captionStartId="F34" captionText="Figure 34. Pheidole itremo sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0491586) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0491584)." figureDoi="10.3897/zookeys.949.51269.figure34" httpUri="https://binary.pensoft.net/fig/432203" pageId="0" pageNumber="1">34A, C</figureCitation>
|
||
).
|
||
<emphasis id="2D185C491E06E617A7EBBEAFB6A26DA3" bold="true" italics="true" pageId="0" pageNumber="1">Mesosoma.</emphasis>
|
||
In lateral view, promesonotum moderately high and short, arched; promesonotal groove absent; metanotal groove distinct; propodeal spines minute and triangular (Fig.
|
||
<figureCitation id="237014AB7112D80EECB2E84936F51057" captionStart="Figure 34" captionStartId="F34" captionText="Figure 34. Pheidole itremo sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0491586) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0491584)." figureDoi="10.3897/zookeys.949.51269.figure34" httpUri="https://binary.pensoft.net/fig/432203" pageId="0" pageNumber="1">34C</figureCitation>
|
||
). Sculpture shiny and smooth. Pilosity moderately sparse, long, and erect (Fig.
|
||
<figureCitation id="B4A88CF8ECA113207855AA4E40D855F8" captionStart="Figure 34" captionStartId="F34" captionText="Figure 34. Pheidole itremo sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0491586) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0491584)." figureDoi="10.3897/zookeys.949.51269.figure34" httpUri="https://binary.pensoft.net/fig/432203" pageId="0" pageNumber="1">34C, E</figureCitation>
|
||
).
|
||
<emphasis id="FAE42CBDDB91E3BC41B18674B08A9F89" bold="true" italics="true" pageId="0" pageNumber="1">Gaster.</emphasis>
|
||
With sparse, erect pilosity (Fig.
|
||
<figureCitation id="15AEA22780213AA7CE1615174578CAB6" captionStart="Figure 34" captionStartId="F34" captionText="Figure 34. Pheidole itremo sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0491586) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0491584)." figureDoi="10.3897/zookeys.949.51269.figure34" httpUri="https://binary.pensoft.net/fig/432203" pageId="0" pageNumber="1">34C, E</figureCitation>
|
||
).
|
||
<emphasis id="94655C52B8CAF06DC7DEC70BFF506B0C" bold="true" italics="true" pageId="0" pageNumber="1">Colour.</emphasis>
|
||
Yellowish brown (Fig.
|
||
<figureCitation id="02D6677A5669D70C41806A5B284DC79E" captionStart="Figure 34" captionStartId="F34" captionText="Figure 34. Pheidole itremo sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0491586) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0491584)." figureDoi="10.3897/zookeys.949.51269.figure34" httpUri="https://binary.pensoft.net/fig/432203" pageId="0" pageNumber="1">34C, E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="30323DB4CC0BCFF4B2291B227D5AC038" pageId="0" pageNumber="1" type="etymology">
|
||
<paragraph id="BE9BDB261EEDA33363CCF17E7BF90EC9" pageId="0" pageNumber="1">Etymology.</paragraph>
|
||
<paragraph id="4318B030DDB467CDC83AFC970EBC175B" pageId="0" pageNumber="1">From the type locality.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A816F86774EA313C42C2BF9ABCBD7EB4" pageId="0" pageNumber="1" type="biology_ecology">
|
||
<paragraph id="074AF4A99FFA7A873FFDEBC2D7F5B194" pageId="0" pageNumber="1">Biology.</paragraph>
|
||
<paragraph id="F25D752F0B4589FC18477BC602B178F5" pageId="0" pageNumber="1">The species was collected between 1410-1620 m in elevation, in montane forest and montane rainforest. Nests were located in rotten logs and root mats.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="269B67983256B037FB2FCC3F763BBEB6" pageId="0" pageNumber="1" type="comments">
|
||
<paragraph id="2E33C652158E3065CC8F4A22C94C051A" pageId="0" pageNumber="1">Comments.</paragraph>
|
||
<paragraph id="7FC1D807BAD7B2DF8E29B56A90D41B04" pageId="0" pageNumber="1">
|
||
<taxonomicName id="00E9F86A9709191E3B372238A9B371A6" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole itremo" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="itremo">
|
||
<emphasis id="432426D28D2A5EA0AAE9C3881A2745E3" italics="true" pageId="0" pageNumber="1">Pheidole itremo</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., known from
|
||
<normalizedToken id="D494F3480F1C174376286732592C29E1" originalValue="Forêt">Foret</normalizedToken>
|
||
<normalizedToken id="ECC790530D0656AB1B65EEF0627BAE21" originalValue="d’Atsirakambiaty">d'Atsirakambiaty</normalizedToken>
|
||
in Fianarantsoa and
|
||
<normalizedToken id="EE481496EF74D171C44274AA43E57F55" originalValue="Forêt">Foret</normalizedToken>
|
||
<normalizedToken id="0C731385BCF445DD8C1DDEF7B94BE74D" originalValue="d’Ambohitantely">d'Ambohitantely</normalizedToken>
|
||
in Antananarivo, is most similar to
|
||
<taxonomicName id="CEFF5BEDE45157B95118973921C1102D" lsidName="P. sava" pageId="0" pageNumber="1" rank="species" species="sava">
|
||
<emphasis id="6A146232BF2231F0BE1D0EA79B0B42BF" italics="true" pageId="0" pageNumber="1">P. sava</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., described from Parc National de Marojejy in Antsiranana. Major workers of
|
||
<taxonomicName id="22BE6C1265DA66977ACE0D24026CD2CC" lsidName="P. itremo" pageId="0" pageNumber="1" rank="species" species="itremo">
|
||
<emphasis id="E038894E1FF284620D844CFE484126D5" italics="true" pageId="0" pageNumber="1">P. itremo</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. can be separated based on darker body colouration, presence of moderately dense, moderately long, suberect to erect pilosity on sides of the head, medial frons with smooth interspaces, finely rugofoveolae mesosoma lacking smooth notches; minor workers can be separated based on predominantly smooth head with posterolateral sides of frons with sparse and indistinct foveolae, and entirely smooth mesosoma.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |