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<document id="377E30EF2412B09FF518089391A40B37" ID="10.11646/zootaxa.4254.3.3SLASH10849" ID-CLB-Dataset="33209" ID-DOI="10.11646/zootaxa.4254.3.3" ID-GBIF-Dataset="5f00a3f5-ecce-427f-b83e-4756bab5f231" ID-ISSN="1175-5326" ID-Zenodo-Dep="546827" ID-ZooBank="88F93D05-41AD-4726-A16C-76CDE023B505" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1492501084686" checkinUser="plazi" docAuthor="Riedel, Alexander &amp; Kilmaskossu, Agust" docDate="2017" docId="291D9614A964B723D9D5F9E1FD13FCDB" docLanguage="en" docName="zootaxa.4254.3.3SLASH10849.pdf" docOrigin="Zootaxa 4254 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Gymnopholus (Niphetoscapha) inexspectatus Riedel, sp. n." docType="treatment" docVersion="7" lastPageNumber="351" masterDocId="D524EE6CA96EB72FD942FFCCFFD5FF86" masterDocTitle="Revision of the subgenus Niphetoscapha Heller of Gymnopholus Heller (Coleoptera, Curculionoidea, Entiminae, Eupholini) and a new species with epizoic symbiosis from West New Guinea" masterLastPageNumber="356" masterPageNumber="339" pageNumber="349" updateTime="1698438923602" updateUser="ExternalLinkService">
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<mods:title id="31541E69BA4E656740DAE01BB5783F5C">Revision of the subgenus Niphetoscapha Heller of Gymnopholus Heller (Coleoptera, Curculionoidea, Entiminae, Eupholini) and a new species with epizoic symbiosis from West New Guinea</mods:title>
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<mods:namePart id="2A50F8824C60A7F512519EF6613DA4F3">Riedel, Alexander</mods:namePart>
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<treatment id="291D9614A964B723D9D5F9E1FD13FCDB" ID-DOI="http://doi.org/10.5281/zenodo.6008759" ID-GBIF-Taxon="128535131" ID-Zenodo-Dep="6008759" LSID="urn:lsid:plazi:treatment:291D9614A964B723D9D5F9E1FD13FCDB" httpUri="http://treatment.plazi.org/id/291D9614A964B723D9D5F9E1FD13FCDB" lastPageId="12" lastPageNumber="351" pageId="10" pageNumber="349">
<subSubSection id="E9AE7489A964B725D9D5F9E1FCDDF9EF" pageId="10" pageNumber="349" type="nomenclature">
<paragraph id="A10B2702A964B725D9D5F9E1FC8FF9C1" blockId="10.[151,858,1581,1641]" box="[151,858,1581,1607]" pageId="10" pageNumber="349">
<heading id="FA43906EA964B725D9D5F9E1FC8FF9C1" bold="true" box="[151,858,1581,1607]" fontSize="11" level="1" pageId="10" pageNumber="349" reason="1">
<emphasis id="93C0FB10A964B725D9D5F9E1FC8FF9C1" bold="true" box="[151,858,1581,1607]" pageId="10" pageNumber="349">
<taxonomicName id="66B45C81A964B725D9D5F9E1FCC4F9C1" ID-CoL="3HT5Q" authority="Riedel" authorityName="Riedel" box="[151,785,1581,1607]" class="Insecta" family="Curculionidae" genus="Gymnopholus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="349" phylum="Arthropoda" rank="species" species="inexspectatus" status="sp. nov." subGenus="Niphetoscapha">
<emphasis id="93C0FB10A964B725D9D5F9E1FE95F9C1" bold="true" box="[151,320,1581,1607]" italics="true" pageId="10" pageNumber="349">Gymnopholus</emphasis>
(
<emphasis id="93C0FB10A964B725D813F9E1FDD0F9C1" bold="true" box="[337,517,1581,1607]" italics="true" pageId="10" pageNumber="349">Niphetoscapha</emphasis>
)
<emphasis id="93C0FB10A964B725DB55F9E1FD6CF9C1" bold="true" box="[535,697,1581,1607]" italics="true" pageId="10" pageNumber="349">inexspectatus</emphasis>
Riedel
</taxonomicName>
<taxonomicNameLabel id="88F3466BA964B725DA5AF9E1FC8FF9C1" box="[792,858,1581,1607]" pageId="10" pageNumber="349" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="A10B2702A964B725D9D5F99CFCDDF9EF" blockId="10.[151,858,1581,1641]" box="[151,776,1616,1641]" pageId="10" pageNumber="349">
(
<figureCitation id="398F3B87A964B725D9DDF99CFEDFF9EF" box="[159,266,1616,1641]" captionStart="FIGURES 5 8" captionStartId="3.[151,264,1905,1928]" captionTargetBox="[247,1348,197,1869]" captionTargetId="figure@3.[228,1358,193,1885]" captionTargetPageId="3" captionText="FIGURES 5 8. Habitus of Gymnopholus (Niphetoscapha) spp., dorsal aspect: (5) G. inexspectatus sp. n., holotype; (6) G. inexspectatus sp. n., allotype; (7) G. wichmanni, paralectotype, male; (8) G. wichmanni, lectotype, female." httpUri="https://zenodo.org/record/546829/files/figure.png" pageId="10" pageNumber="349">Figs. 56</figureCitation>
,
<figureCitation id="398F3B87A964B725D854F99DFEE6F9EF" box="[278,307,1617,1641]" captionStart="FIGURES 9 12" captionStartId="4.[151,264,1672,1695]" captionTargetBox="[299,1273,206,1652]" captionTargetId="figure@4.[299,1288,193,1652]" captionTargetPageId="4" captionText="FIGURES 9 12. Head and rostrum of Gymnopholus (Niphetoscapha) spp.: (9) G. audax; (10) G. nitidus; (11) G. inexspectatus sp. n., holotype; (12) G. wichmanni, lectotype." httpUri="https://zenodo.org/record/546830/files/figure.png" pageId="10" pageNumber="349">11</figureCitation>
,
<figureCitation id="398F3B87A964B725D87DF99CFE88F9EF" box="[319,349,1616,1641]" captionStart="FIGURES 13 16" captionStartId="5.[151,264,1926,1949]" captionTargetBox="[251,1335,481,1906]" captionTargetId="figure@5.[251,1335,481,1906]" captionTargetPageId="5" captionText="FIGURES 13 16. Pronotum of Gymnopholus (Niphetoscapha) spp., female: (13) G. audax (14) G. nitidus; (15) G. inexspectatus sp. n., allotype; (16) G. wichmanni, lectotype." httpUri="https://zenodo.org/record/546831/files/figure.png" pageId="10" pageNumber="349">15</figureCitation>
,
<figureCitation id="398F3B87A964B725D82BF99DFE52F9EF" box="[361,391,1617,1641]" captionStart="FIGURES 17 20" captionStartId="6.[151,264,1873,1896]" captionTargetBox="[277,1315,194,1852]" captionTargetId="figure@6.[272,1315,193,1852]" captionTargetPageId="6" captionText="FIGURES 17 20. Elytral apex of Gymnopholus (Niphetoscapha) spp., female; in dorsoapical aspect (left), in profile (right): (17) G. audax; (18) G. nitidus; (19) G. wichmanni, lectotype; (20) G. inexspectatus sp. n., allotype." httpUri="https://zenodo.org/record/546832/files/figure.png" pageId="10" pageNumber="349">20</figureCitation>
,
<figureCitation id="398F3B87A964B725D8D1F99DFE64F9EF" box="[403,433,1617,1641]" captionStart="FIGURES 21 24" captionStartId="7.[151,264,1510,1533]" captionTargetBox="[200,1389,199,1489]" captionTargetId="figure@7.[192,1395,193,1489]" captionTargetPageId="7" captionText="FIGURES 21 24. Gymnopholus (Niphetoscapha) spp.; Male genitalia; body of penis in dorsal aspect (left), in profile (right): (21) G. audax; (22) G. nitidus; (23) G. inexspectatus sp. n., holotype; (24) G. wichmanni, paralectotype." httpUri="https://zenodo.org/record/546833/files/figure.png" pageId="10" pageNumber="349">23</figureCitation>
,
<figureCitation id="398F3B87A964B725D8FFF99CFD78F9EF" box="[445,685,1616,1641]" captionStart-0="FIGURES 25 32" captionStart-1="FIGURES 33 38" captionStartId-0="8.[151,264,1626,1649]" captionStartId-1="9.[151,264,1654,1677]" captionTargetBox-0="[194,1395,196,1592]" captionTargetBox-1="[237,1370,369,1632]" captionTargetId-0="figure@8.[188,1398,193,1605]" captionTargetId-1="figure@9.[217,1370,369,1633]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="FIGURES 25 32. Gymnopholus (Niphetoscapha) spp.; female terminalia: (25) G. audax; (26) G. inexspectatus sp. n., allotype; (27) sternite VIII of G. audax; (28) sternite VIII of G. inexspectatus sp. n., allotype; (28) spermatheca of G. audax; (29) spermatheca of G. inexspectatus sp. n., allotype; (31) hemisternite of G. audax; (32) hemisternite of G. inexspectatus sp. n., allotype." captionText-1="FIGURES 33 38. Gymnopholus spp.: (33) Transfer apparatus of G. inexspectatus sp. n., holotype; (34) Transfer apparatus of G. nitidus; (35) Left wing of G. (Symbiopholus) lichenifer; (36) Left wing of G. (Niphetoscapha) inexspectatus sp. n., allotype; (37) labels of G. wichmanni, lectotype; (38) labels of G. wichmanni, paralectotype." httpUri-0="https://zenodo.org/record/546834/files/figure.png" httpUri-1="https://zenodo.org/record/546835/files/figure.png" pageId="10" pageNumber="349">26, 28, 29, 3233, 36</figureCitation>
, 40, 41)
</paragraph>
</subSubSection>
<subSubSection id="E9AE7489A964B725D9D5F954FC2BF953" pageId="10" pageNumber="349" type="diagnosis">
<paragraph id="A10B2702A964B725D9D5F954FC2BF953" blockId="10.[151,1436,1688,2001]" pageId="10" pageNumber="349">
<emphasis id="93C0FB10A964B725D9D5F954FEDFF937" bold="true" box="[151,266,1688,1713]" pageId="10" pageNumber="349">Diagnosis</emphasis>
. Elytron and pronotum with irregular rough sculpture composed of wrinkles, pits and tubercles; densely covered with scales and setae; partly with greenish to brownish encrustations.
</paragraph>
</subSubSection>
<subSubSection id="E9AE7489A964B723D985F92CFB7EFF7E" lastPageId="12" lastPageNumber="351" pageId="10" pageNumber="349" type="description">
<paragraph id="A10B2702A964B725D985F92CFACBF97E" blockId="10.[151,1436,1688,2001]" box="[199,1310,1760,1785]" pageId="10" pageNumber="349">
<emphasis id="93C0FB10A964B725D985F92CFE8CF97F" bold="true" box="[199,345,1760,1785]" pageId="10" pageNumber="349">Description.</emphasis>
<typeStatus id="7E0F99A0A964B725D81DF92CFE1FF97F" box="[351,458,1760,1785]" pageId="10" pageNumber="349" type="holotype">Holotype</typeStatus>
, male (
<figureCitation id="398F3B87A964B725DB59F92CFD8AF97F" box="[539,607,1760,1785]" captionStart="FIGURES 5 8" captionStartId="3.[151,264,1905,1928]" captionTargetBox="[247,1348,197,1869]" captionTargetId="figure@3.[228,1358,193,1885]" captionTargetPageId="3" captionText="FIGURES 5 8. Habitus of Gymnopholus (Niphetoscapha) spp., dorsal aspect: (5) G. inexspectatus sp. n., holotype; (6) G. inexspectatus sp. n., allotype; (7) G. wichmanni, paralectotype, male; (8) G. wichmanni, lectotype, female." httpUri="https://zenodo.org/record/546829/files/figure.png" pageId="10" pageNumber="349">Fig. 5</figureCitation>
). Length. Total length
<quantity id="664C8AE7A964B725DA22F92CFC12F97E" box="[864,967,1760,1785]" metricMagnitude="-2" metricUnit="m" metricValue="2.35" pageId="10" pageNumber="349" unit="mm" value="23.5">23.5 mm</quantity>
; pronotum + elytron
<quantity id="664C8AE7A964B725DDF1F92CFACFF97E" box="[1203,1306,1760,1785]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" pageId="10" pageNumber="349" unit="mm" value="18.5">18.5 mm</quantity>
.
</paragraph>
<paragraph id="A10B2702A964B725D985F8C8FF36F8E2" blockId="10.[151,1436,1688,2001]" pageId="10" pageNumber="349">Color of integument black, pronotum and elytron densely setose-squamose with encrustations of algae or lichens, in live specimens greenish, in dry specimens brownish; venter shining, legs and head dull and sparsely setose.</paragraph>
<paragraph id="A10B2702A964B725D985F8BCFF36F857" blockId="10.[151,1436,1688,2001]" pageId="10" pageNumber="349">
Head (
<figureCitation id="398F3B87A964B725D856F8BCFEB3F80F" box="[276,358,1904,1929]" captionStart="FIGURES 9 12" captionStartId="4.[151,264,1672,1695]" captionTargetBox="[299,1273,206,1652]" captionTargetId="figure@4.[299,1288,193,1652]" captionTargetPageId="4" captionText="FIGURES 9 12. Head and rostrum of Gymnopholus (Niphetoscapha) spp.: (9) G. audax; (10) G. nitidus; (11) G. inexspectatus sp. n., holotype; (12) G. wichmanni, lectotype." httpUri="https://zenodo.org/record/546830/files/figure.png" pageId="10" pageNumber="349">Fig. 11</figureCitation>
) dull, densely punctate, microreticulate, sparsely setose; between eyes ca. as wide as vertical diameter of eye (1.01 X). Eyes markedly prominent from outline of head. Gena 1.09 X as long as vertical diameter of eye.
</paragraph>
<caption id="F5CB778AA965B724D9D5FD07FEC6FC98" httpUri="https://zenodo.org/record/546836/files/figure.png" pageId="11" pageNumber="350" targetBox="[191,1396,193,693]" targetPageId="11">
<paragraph id="A10B2702A965B724D9D5FD07FEC6FC98" blockId="11.[151,1436,714,798]" pageId="11" pageNumber="350">
<emphasis id="93C0FB10A965B724D9D5FD07FEFDFD66" bold="true" box="[151,296,715,737]" pageId="11" pageNumber="350">FIGURE 39.</emphasis>
Elytral base of
<taxonomicName id="66B45C81A965B724D8A8FD07FC57FD67" box="[490,898,715,737]" class="Insecta" family="Curculionidae" genus="Gymnopholus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="350" phylum="Arthropoda" rank="species" species="nitidus" subGenus="Niphetoscapha">
<emphasis id="93C0FB10A965B724D8A8FD07FC57FD67" box="[490,898,715,737]" italics="true" pageId="11" pageNumber="350">Gymnopholus (Niphetoscapha) nitidus</emphasis>
</taxonomicName>
with exudates.
<emphasis id="93C0FB10A965B724DD78FD06FAD4FD66" bold="true" box="[1082,1281,714,737]" pageId="11" pageNumber="350">FIGURES 40ab.</emphasis>
<taxonomicName id="66B45C81A965B724DC4DFD07FE03FD79" class="Insecta" family="Curculionidae" genus="Gymnopholus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="350" phylum="Arthropoda" rank="species" species="inexspectatus" status="sp. nov." subGenus="Niphetoscapha">
<emphasis id="93C0FB10A965B724DC4DFD07FA4EFD67" box="[1295,1435,715,737]" italics="true" pageId="11" pageNumber="350">Gymnopholus</emphasis>
(
<emphasis id="93C0FB10A965B724D9DDFD25FEE2FD79" box="[159,311,745,767]" italics="true" pageId="11" pageNumber="350">Niphetoscapha</emphasis>
)
<emphasis id="93C0FB10A965B724D80CFD26FE03FD79" box="[334,470,746,767]" italics="true" pageId="11" pageNumber="350">inexspectatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="88F3466BA965B724D8A7FD26FDF2FD79" box="[485,551,746,767]" pageId="11" pageNumber="350" rank="species">
<emphasis id="93C0FB10A965B724D8A7FD26FDF2FD79" bold="true" box="[485,551,746,767]" pageId="11" pageNumber="350">sp. n.</emphasis>
</taxonomicNameLabel>
, live specimen with incrustrations of algae and lichens; photographs M. Wild, Mokndoma.
</paragraph>
</caption>
<paragraph id="A10B2702A965B724D985FC8BFDE6FC20" blockId="11.[151,1437,837,2038]" pageId="11" pageNumber="350">
Rostrum (
<figureCitation id="398F3B87A965B724D875FC89FE55FCD8" box="[311,384,837,862]" captionStart="FIGURES 9 12" captionStartId="4.[151,264,1672,1695]" captionTargetBox="[299,1273,206,1652]" captionTargetId="figure@4.[299,1288,193,1652]" captionTargetPageId="4" captionText="FIGURES 9 12. Head and rostrum of Gymnopholus (Niphetoscapha) spp.: (9) G. audax; (10) G. nitidus; (11) G. inexspectatus sp. n., holotype; (12) G. wichmanni, lectotype." httpUri="https://zenodo.org/record/546830/files/figure.png" pageId="11" pageNumber="350">Fig.11</figureCitation>
) 2.50 X longer than wide at base; maximum width in front of antennal insertion 1.42 X width at base. Dorsum with marked median impression; pair of distinct dorsolateral furrows and antennal scrobes clothed with scales and brown encrustations.
</paragraph>
<paragraph id="A10B2702A965B724D985FC7EFBF3FC4C" blockId="11.[151,1437,837,2038]" box="[199,1062,945,970]" pageId="11" pageNumber="350">Antenna. Funicle + club 1.66 X length of scape; club 4.08 X as long as wide.</paragraph>
<paragraph id="A10B2702A965B724D985FC19FE86FBF8" blockId="11.[151,1437,837,2038]" pageId="11" pageNumber="350">Thorax. Pronotum 1.14 X longer than wide; sides gently sinuate, widest in apical half; vestiture of dense recumbent scales and sparse suberect scales, partly covered by encrustations; disc with irregular tubercles, some clothed by vestiture, others nude; in basal half with narrow, median impression, in anterior half with wider, roundish pit. Thoracic venter setose with thin setae not concealing surface; mesoventrite and mesocoxae with dense recumbent setae.</paragraph>
<paragraph id="A10B2702A965B724D985FB45FE51FAD0" blockId="11.[151,1437,837,2038]" pageId="11" pageNumber="350">
Elytron 4.10 X longer than wide between humeri; at greatest width 1.12 X wider than between humeri; sides subparallel; surface with vestiture of dense recumbent scales and sparse suberect scales, partly covered by encrustations; with coarse irregular transverse wrinkles obscuring indistinct striae; without longitudinal ridges; epipleural edge with indistinct fringe of setae; at apical declivity (
<figureCitation id="398F3B87A965B724DAD9FB39FC25FA88" box="[923,1008,1269,1294]" captionStart="FIGURES 17 20" captionStartId="6.[151,264,1873,1896]" captionTargetBox="[277,1315,194,1852]" captionTargetId="figure@6.[272,1315,193,1852]" captionTargetPageId="6" captionText="FIGURES 17 20. Elytral apex of Gymnopholus (Niphetoscapha) spp., female; in dorsoapical aspect (left), in profile (right): (17) G. audax; (18) G. nitidus; (19) G. wichmanni, lectotype; (20) G. inexspectatus sp. n., allotype." httpUri="https://zenodo.org/record/546832/files/figure.png" pageId="11" pageNumber="350">Fig. 20</figureCitation>
) intervals 13 with transverse ridge; subapically intervals 3 and 9 with longitudinal ridges meeting at acute angle; apical declivity without distinct protrusions or knobs.
</paragraph>
<paragraph id="A10B2702A965B724D985FAAEFB5FFA18" blockId="11.[151,1437,837,2038]" pageId="11" pageNumber="350">Legs. Surface dull, microreticulate, punctate; with minute, recumbent setae and with long, yellowish, suberect bristles. Femora relatively long, parallel-sided; metafemur in repose reaching elytral apex.</paragraph>
<paragraph id="A10B2702A965B724D985FA65FA4CFA60" blockId="11.[151,1437,837,2038]" pageId="11" pageNumber="350">
Abdomen. Ventrite 12 medially concave; with sparse, long setae; ventrite 5 flat, subapically densely setose. Penis (
<figureCitation id="398F3B87A965B724D9A1FA01FEE1FA60" box="[227,308,1485,1510]" captionStart="FIGURES 21 24" captionStartId="7.[151,264,1510,1533]" captionTargetBox="[200,1389,199,1489]" captionTargetId="figure@7.[192,1395,193,1489]" captionTargetPageId="7" captionText="FIGURES 21 24. Gymnopholus (Niphetoscapha) spp.; Male genitalia; body of penis in dorsal aspect (left), in profile (right): (21) G. audax; (22) G. nitidus; (23) G. inexspectatus sp. n., holotype; (24) G. wichmanni, paralectotype." httpUri="https://zenodo.org/record/546833/files/figure.png" pageId="11" pageNumber="350">Fig. 23</figureCitation>
) with asymmetrical apical tip slender. Transfer apparatus (
<figureCitation id="398F3B87A965B724DA87FA01FBC2FA60" box="[965,1047,1485,1510]" captionStart="FIGURES 33 38" captionStartId="9.[151,264,1654,1677]" captionTargetBox="[237,1370,369,1632]" captionTargetId="figure@9.[217,1370,369,1633]" captionTargetPageId="9" captionText="FIGURES 33 38. Gymnopholus spp.: (33) Transfer apparatus of G. inexspectatus sp. n., holotype; (34) Transfer apparatus of G. nitidus; (35) Left wing of G. (Symbiopholus) lichenifer; (36) Left wing of G. (Niphetoscapha) inexspectatus sp. n., allotype; (37) labels of G. wichmanni, lectotype; (38) labels of G. wichmanni, paralectotype." httpUri="https://zenodo.org/record/546835/files/figure.png" pageId="11" pageNumber="350">Fig. 33</figureCitation>
) ca. 1.5 X as long as wide at base.
</paragraph>
<paragraph id="A10B2702A965B724D985FA3DFAB6F9A8" blockId="11.[151,1437,837,2038]" pageId="11" pageNumber="350">
<typeStatus id="7E0F99A0A965B724D985FA3DFEF9F98C" box="[199,300,1521,1546]" pageId="11" pageNumber="350" type="allotype">Allotype</typeStatus>
, female (
<figureCitation id="398F3B87A965B724D8D6FA3DFE0CF98C" box="[404,473,1521,1546]" captionStart="FIGURES 5 8" captionStartId="3.[151,264,1905,1928]" captionTargetBox="[247,1348,197,1869]" captionTargetId="figure@3.[228,1358,193,1885]" captionTargetPageId="3" captionText="FIGURES 5 8. Habitus of Gymnopholus (Niphetoscapha) spp., dorsal aspect: (5) G. inexspectatus sp. n., holotype; (6) G. inexspectatus sp. n., allotype; (7) G. wichmanni, paralectotype, male; (8) G. wichmanni, lectotype, female." httpUri="https://zenodo.org/record/546829/files/figure.png" pageId="11" pageNumber="350">Fig. 6</figureCitation>
). Same as
<typeStatus id="7E0F99A0A965B724DB16FA3DFD63F98C" box="[596,694,1521,1546]" pageId="11" pageNumber="350" type="holotype">holotype</typeStatus>
except: Length. Total length
<quantity id="664C8AE7A965B724DD41FA3EFB83F98F" box="[1027,1110,1522,1546]" metricMagnitude="-2" metricUnit="m" metricValue="2.7" pageId="11" pageNumber="350" unit="mm" value="27.0">27 mm</quantity>
; pronotum + elytron
<quantity id="664C8AE7A965B724DC04FA3EFA4CF98F" box="[1350,1433,1522,1546]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" pageId="11" pageNumber="350" unit="mm" value="22.0">22 mm</quantity>
. Elytron wider, 3.88 X longer than wide between humeri, at greatest width 1.10 X wider than between humeri.
</paragraph>
<paragraph id="A10B2702A965B723D985F9F7FB7EFF7E" blockId="11.[151,1437,837,2038]" lastBlockId="12.[151,1437,151,861]" lastPageId="12" lastPageNumber="351" pageId="11" pageNumber="350">
Intraspecific variation. Length. Total length of males
<quantity id="664C8AE7A965B724DA5AF9F7FC6EF9D5" box="[792,955,1595,1620]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" metricValueMax="2.75" metricValueMin="2.25" pageId="11" pageNumber="350" unit="mm" value="25.0" valueMax="27.5" valueMin="22.5">22.527.5 mm</quantity>
(n=6,
<emphasis id="93C0FB10A965B724DD45F9F1FBC0F9DF" box="[1031,1045,1597,1625]" italics="true" pageId="11" pageNumber="350">x</emphasis>
=24.3 ± 1.8), of females 27.030.0 mm (n=6,
<emphasis id="93C0FB10A965B724D850F9AFFEF5F9F9" box="[274,288,1635,1663]" italics="true" pageId="11" pageNumber="350">x</emphasis>
=28.6 ± 1.3); pronotum + elytron of males 18.522.0 mm (n=6,
<emphasis id="93C0FB10A965B724DD42F9AFFBDBF9F9" box="[1024,1038,1635,1663]" italics="true" pageId="11" pageNumber="350">x</emphasis>
=19.8 ± 1.5), of females 22.025.0 mm (n=6,
<emphasis id="93C0FB10A965B724D853F945FECAF923" box="[273,287,1673,1701]" italics="true" pageId="11" pageNumber="350">x</emphasis>
=23.4 ± 1.2). Coloration of life specimens usually green and dominated by algal encrustations turning brown in dry collection specimens; amount of algal or lichen encrustations variable; recumbent scales of elytral grooves visible or fully covered by encrustations. Rostrum of males 2.502.71 X longer than wide at base (n=6,
<emphasis id="93C0FB10A965B724D9D9F934FF7CF892" box="[155,169,1784,1812]" italics="true" pageId="11" pageNumber="350">x</emphasis>
=2.59 ± 0.08), of females 2.052.40 X longer than wide at base (n=6,
<emphasis id="93C0FB10A965B724DA86F934FC07F892" box="[964,978,1784,1812]" italics="true" pageId="11" pageNumber="350">x</emphasis>
=2.24 ± 0.11); maximum width in males 1.421.56 X width at base (n=6,
<emphasis id="93C0FB10A965B724DB5EF8D2FDFFF8BC" box="[540,554,1822,1850]" italics="true" pageId="11" pageNumber="350">x</emphasis>
=1.52 ± 0.05), in females 1.451.47 X width at base (n=6,
<emphasis id="93C0FB10A965B724DDA6F8D2FB27F8BC" box="[1252,1266,1822,1850]" italics="true" pageId="11" pageNumber="350">x</emphasis>
=1.46 ± 0.01). Antenna. Funicle + club 1.451.71 X length of scape (n=12,
<emphasis id="93C0FB10A965B724DA2EF888FCAFF8E6" box="[876,890,1860,1888]" italics="true" pageId="11" pageNumber="350">x</emphasis>
=1.56 ± 0.08). Pronotum with shallow median groove wide or narrow, more or less distinct; submedian tubercles more or less numerous, more or less distinct. Elytron of males 3.924.55 X longer than wide between humeri (n=6,
<emphasis id="93C0FB10A965B724DAF3F842FC6AF82C" box="[945,959,1934,1962]" italics="true" pageId="11" pageNumber="350">x</emphasis>
=4.21 ± 0.21), at greatest width 1.041.15 X wider than between humeri (n=6,
<emphasis id="93C0FB10A965B724DB03F878FD9AF856" box="[577,591,1972,2000]" italics="true" pageId="11" pageNumber="350">x</emphasis>
=1.11 ± 0.04); elytron of females 3.884.16 X longer than wide between humeri (n=6,
<emphasis id="93C0FB10A965B724D87BF816FE92F870" box="[313,327,2010,2038]" italics="true" pageId="11" pageNumber="350">x</emphasis>
=3.96 ± 0.19), at greatest width 1.091.20 X wider than between humeri (n=6,
<emphasis id="93C0FB10A965B724DDA6F816FB27F870" box="[1252,1266,2010,2038]" italics="true" pageId="11" pageNumber="350">x</emphasis>
=1.13 ± 0.04); elytral sculpture more distinct in specimens from Bilogay in which transverse wrinkles and grooves are deep, partly tuberculate; elytral transverse wrinkles in specimens from other localities more gentle, in some specimens almost absent; shape of suberect elytral scales relatively short, almond-shaped, or longer, lanceolate.
</paragraph>
</subSubSection>
<subSubSection id="E9AE7489A962B723D985FEC8FE91FD2E" pageId="12" pageNumber="351" type="materials_examined">
<paragraph id="A10B2702A962B723D985FEC8FBD3FDE6" blockId="12.[151,1437,151,861]" pageId="12" pageNumber="351">
<materialsCitation id="11DC2D5FA962B723D985FEC8FE04FEC6" ID-GBIF-Occurrence="1456819364" collectingDate="1995-12-14" collectingDateMax="1995-12-17" collectingDateMin="1995-12-14" collectionCode="WEST, NEW, MZB" collectorName="Material &amp; Paniai Reg. &amp; Sinak" country="Guinea-Bissau" elevation="2100" latitude="-3.8333333" location="GUINEA" longLatPrecision="1307" longitude="137.85" pageId="12" pageNumber="351" specimenCount="1" typeStatus="holotype">
<emphasis id="93C0FB10A962B723D985FEC8FE7CFE9B" bold="true" box="[199,425,260,285]" pageId="12" pageNumber="351">
<collectorName id="0C4142D4A962B723D985FEC8FEFAFE9B" box="[199,303,260,285]" pageId="12" pageNumber="351">Material</collectorName>
examined
</emphasis>
.
<typeStatus id="7E0F99A0A962B723D8F4FEC8FDF6FE9B" box="[438,547,260,285]" pageId="12" pageNumber="351" type="holotype">Holotype</typeStatus>
:
<collectionCode id="C7A5BFC7A962B723DB6FFEC8FDACFE9A" box="[557,633,260,284]" pageId="12" pageNumber="351">WEST</collectionCode>
<collectionCode id="C7A5BFC7A962B723DBC2FEC9FD15FE9A" box="[640,704,261,284]" country="Australia" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="12" pageNumber="351">NEW</collectionCode>
<collectingCountry id="D9A36792A962B723DB84FEC8FCE1FE9A" box="[710,820,260,284]" name="Guinea-Bissau" pageId="12" pageNumber="351">GUINEA</collectingCountry>
,
<collectorName id="0C4142D4A962B723DA7DFEC8FC16FE9B" box="[831,963,260,285]" pageId="12" pageNumber="351">Paniai Reg.</collectorName>
,
<collectorName id="0C4142D4A962B723DA8CFEC8FBDBFE9B" box="[974,1038,260,285]" pageId="12" pageNumber="351">Sinak</collectorName>
[
<geoCoordinate id="C48041C5A962B723DD5EFEC9FB42FE9B" box="[1052,1175,260,285]" direction="east" orientation="longitude" pageId="12" pageNumber="351" precision="925" value="137.85">E 137° 51´</geoCoordinate>
<geoCoordinate id="C48041C5A962B723DDDCFEC8FADEFE9B" box="[1182,1291,260,285]" direction="south" orientation="latitude" pageId="12" pageNumber="351" precision="925" value="-3.8333333">S 03° 50´</geoCoordinate>
],
<elevation id="2A99C031A962B723DC5CFEC8FF7AFEB9" metricMagnitude="3" metricUnit="m" metricValue="2.1" metricValueMax="2.2" metricValueMin="2.0" pageId="12" pageNumber="351" unit="m" value="2100.0" valueMax="2200.0" valueMin="2000.0">
<quantity id="664C8AE7A962B723DC5CFEC8FF7AFEB9" metricMagnitude="3" metricUnit="m" metricValue="2.1" metricValueMax="2.2" metricValueMin="2.0" pageId="12" pageNumber="351" unit="m" value="2100.0" valueMax="2200.0" valueMin="2000.0">20002200 m</quantity>
</elevation>
,
<collectingDate id="C54EF82AA962B723D982FEEBFEA7FEC6" box="[192,370,295,320]" pageId="12" pageNumber="351" value="1995-12-14" valueMax="1995-12-17" valueMin="1995-12-14">
<date id="D50A01C2A962B723D982FEEBFEA7FEC6" box="[192,370,295,320]" pageId="12" pageNumber="351" value="1995-12-14" valueMax="1995-12-17" valueMin="1995-12-14">1417.XII.1995</date>
</collectingDate>
(
<collectionCode id="C7A5BFC7A962B723D8C5FEE4FE1CFEC6" box="[391,457,296,320]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="12" pageNumber="351">MZB</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="11DC2D5FA962B723D8A3FEE4FD1FFEE2" ID-GBIF-Occurrence="1456819363" collectingDate="1995-12-14" collectingDateMax="1995-12-17" collectingDateMin="1995-12-14" collectionCode="WEST, NEW, ARC, SMNK" collectorName="Material &amp; Paniai Reg. &amp; Sinak" country="Guinea-Bissau" elevation="2100" latitude="-3.8333333" location="GUINEA" longLatPrecision="1307" longitude="137.85" pageId="12" pageNumber="351" specimenCount="8" specimenCount-female="5" specimenCount-male="3" typeStatus="paratype">
<typeStatus id="7E0F99A0A962B723D8A3FEE4FD86FEC6" box="[481,595,296,320]" pageId="12" pageNumber="351" type="paratype">Paratypes</typeStatus>
:
<collectionCode id="C7A5BFC7A962B723DB20FEE4FD7BFEC6" box="[610,686,296,320]" pageId="12" pageNumber="351">WEST</collectionCode>
<collectionCode id="C7A5BFC7A962B723DBF9FEE4FD2EFEB9" box="[699,763,296,319]" country="Australia" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="12" pageNumber="351">NEW</collectionCode>
<collectingCountry id="D9A36792A962B723DA45FEE4FCA0FEC6" box="[775,885,296,320]" name="Guinea-Bissau" pageId="12" pageNumber="351">GUINEA</collectingCountry>
, Paniai Reg.:
<specimenCount id="B7B2EC8BA962B723DD5DFEE4FBABFEC6" box="[1055,1150,295,320]" pageId="12" pageNumber="351" type="male">3 males</specimenCount>
,
<specimenCount id="B7B2EC8BA962B723DDCCFEEBFB20FEC6" box="[1166,1269,295,320]" pageId="12" pageNumber="351" type="female">5 female</specimenCount>
(1 marked as “
<typeStatus id="7E0F99A0A962B723D9E3FE80FED6FEE3" box="[161,259,332,357]" pageId="12" pageNumber="351" type="allotype">allotype</typeStatus>
”), same data as holotype (
<collectionCode id="C7A5BFC7A962B723DB66FE80FDB7FEE2" box="[548,610,332,356]" country="Canada" httpUri="http://grbio.org/cool/r82x-h8re" name="Atlantic Reference Centre" pageId="12" pageNumber="351">ARC</collectionCode>
,
<collectionCode id="C7A5BFC7A962B723DB2EFE80FD17FEE2" box="[620,706,332,356]" country="Germany" httpUri="http://grbio.org/cool/tc3a-fmpc" name="Staatliches Museum fuer Naturkunde Karlsruhe (State Museum of Natural History)" pageId="12" pageNumber="351">SMNK</collectionCode>
)
</materialsCitation>
; 1 female, Sinak, Tuput,
<quantity id="664C8AE7A962B723DAA2FE80FBE3FEE2" box="[992,1078,332,356]" metricMagnitude="3" metricUnit="m" metricValue="2.9" pageId="12" pageNumber="351" unit="m" value="2900.0">2900 m</quantity>
,
<date id="D50A01C2A962B723DD03FE80FB1FFEE3" box="[1089,1226,332,357]" pageId="12" pageNumber="351" value="1995-12-12">12.XII.1995</date>
(ARC); 3 males, 1 female, Sinak, trail to Ilaga,
<quantity id="664C8AE7A962B723D8A0FEA3FD57FE01" box="[482,642,367,392]" metricMagnitude="3" metricUnit="m" metricValue="2.3" metricValueMax="2.5" metricValueMin="2.1" pageId="12" pageNumber="351" unit="m" value="2300.0" valueMax="2500.0" valueMin="2100.0">21002500 m</quantity>
,
<date id="D50A01C2A962B723DBD2FEA3FCCCFE0E" box="[656,793,367,392]" pageId="12" pageNumber="351" value="1995-12-15">15.XII.1995</date>
(ARC); 3 males, 3 females, Bilogay [E 137° 01´S 03° 44´], Tigipigu,
<quantity id="664C8AE7A962B723D803FE58FE08FE2A" box="[321,477,404,428]" metricMagnitude="3" metricUnit="m" metricValue="2.25" metricValueMax="2.4" metricValueMin="2.1" pageId="12" pageNumber="351" unit="m" value="2250.0" valueMax="2400.0" valueMin="2100.0">21002400 m</quantity>
,
<date id="D50A01C2A962B723D8ABFE58FDA7FE2B" box="[489,626,404,429]" pageId="12" pageNumber="351" value="1995-12-23">23.XII.1995</date>
(ARC, SMNK); Jayawijaya Reg.: 6 males, 3 females, Mulia [E 137°57´S 03°44´], (S), Wuyuneeri,
<quantity id="664C8AE7A962B723D88BFE74FDB3FE49" box="[457,614,440,464]" metricMagnitude="3" metricUnit="m" metricValue="2.05" metricValueMax="2.2" metricValueMin="1.9" pageId="12" pageNumber="351" unit="m" value="2050.0" valueMax="2200.0" valueMin="1900.0">19002200 m</quantity>
,
<date id="D50A01C2A962B723DB33FE7BFCDDFE56" box="[625,776,439,464]" pageId="12" pageNumber="351" value="1994-07-06" valueMax="1994-07-07" valueMin="1994-07-06">67.VII.1994</date>
(ARC, SMNK). Other material not included in type series: Jayawijaya Reg.: 1 female, Tiom, Wanuga,
<quantity id="664C8AE7A962B723DBDDFE10FC94FE72" box="[671,833,476,501]" metricMagnitude="3" metricUnit="m" metricValue="2.825" metricValueMax="2.9" metricValueMin="2.75" pageId="12" pageNumber="351" unit="m" value="2825.0" valueMax="2900.0" valueMin="2750.0">27502900 m</quantity>
,
<date id="D50A01C2A962B723DA10FE10FC19FE73" box="[850,972,476,501]" pageId="12" pageNumber="351" value="1995-12-08">8.XII.1995</date>
(ARC); 1 male, Wanuga—Kwiyawagi,
<quantity id="664C8AE7A962B723D9D5FE33FEE2FD91" box="[151,311,511,536]" metricMagnitude="3" metricUnit="m" metricValue="2.875" metricValueMax="3.0" metricValueMin="2.75" pageId="12" pageNumber="351" unit="m" value="2875.0" valueMax="3000.0" valueMin="2750.0">27503000 m</quantity>
,
<date id="D50A01C2A962B723D805FE33FE17FD9E" box="[327,450,511,536]" pageId="12" pageNumber="351" value="1995-12-09">9.XII.1995</date>
(ARC);
<materialsCitation id="11DC2D5FA962B723DB69FDCCFDB2FDBA" ID-GBIF-Occurrence="1456819367" collectingDate="2010-07-20" collectionCode="ARC, SMNK" collectorName="W Wamena" country="Guinea-Bissau" elevation="2770" latitude="-4.1376" location="Lake Habbema" longLatPrecision="1" longitude="138.81749" pageId="12" pageNumber="351" specimenCode="ARC2636" specimenCount="1" typeStatus="paratype">
1 ex (
<specimenCode id="F1128F79A962B723DB33FE33FD3CFD9E" box="[625,745,511,536]" collectionCode="ARC" country="Canada" httpUri="http://grbio.org/cool/r82x-h8re" name="Atlantic Reference Centre" pageId="12" pageNumber="351">
<collectionCode id="C7A5BFC7A962B723DB33FE33FD7BFD9E" box="[625,686,511,536]" country="Canada" httpUri="http://grbio.org/cool/r82x-h8re" name="Atlantic Reference Centre" pageId="12" pageNumber="351">ARC</collectionCode>
2636
</specimenCode>
),
<collectorName id="0C4142D4A962B723DBBDFDCCFC5DFD91" box="[767,904,512,535]" pageId="12" pageNumber="351">W Wamena</collectorName>
, road to
<location id="A46B71D9A962B723DAB6FE33FB70FD9E" LSID="urn:lsid:plazi:treatment:291D9614A964B723D9D5F9E1FD13FCDB:A46B71D9A962B723DAB6FE33FB70FD9E" box="[1012,1189,511,536]" country="Guinea-Bissau" latitude="-4.1376" longLatPrecision="1" longitude="138.81749" name="Lake Habbema" pageId="12" pageNumber="351">Lake Habbema</location>
,
<geoCoordinate id="C48041C5A962B723DDF1FDCCFA9EFD9E" box="[1203,1355,511,536]" direction="south" orientation="latitude" pageId="12" pageNumber="351" precision="1" value="-4.1376">S04° 08.256´</geoCoordinate>
<geoCoordinate id="C48041C5A962B723DC14FDCCFF25FDBA" direction="east" orientation="longitude" pageId="12" pageNumber="351" precision="1" value="138.81749">E138° 49.049´</geoCoordinate>
,
<elevation id="2A99C031A962B723D9BCFDE8FE8CFDBA" box="[254,345,548,572]" metricMagnitude="3" metricUnit="m" metricValue="2.77" pageId="12" pageNumber="351" unit="m" value="2770.0">
<quantity id="664C8AE7A962B723D9BCFDE8FE8CFDBA" box="[254,345,548,572]" metricMagnitude="3" metricUnit="m" metricValue="2.77" pageId="12" pageNumber="351" unit="m" value="2770.0">2770 m</quantity>
</elevation>
,
<collectingDate id="C54EF82AA962B723D82AFDE8FE23FDBA" box="[360,502,548,572]" pageId="12" pageNumber="351" value="2010-07-20">
<date id="D50A01C2A962B723D82AFDE8FE23FDBA" box="[360,502,548,572]" pageId="12" pageNumber="351" value="2010-07-20">20-VII-2010</date>
</collectingDate>
(
<collectionCode id="C7A5BFC7A962B723DB4AFDE8FD8AFDBA" box="[520,607,548,572]" country="Germany" httpUri="http://grbio.org/cool/tc3a-fmpc" name="Staatliches Museum fuer Naturkunde Karlsruhe (State Museum of Natural History)" pageId="12" pageNumber="351">SMNK</collectionCode>
)
</materialsCitation>
; 2 exx (ARC2641, ARC2642), N Wamena, Poga, S03° 47.575´E138° 33.155´to S03° 47.473´E138° 33.163´,
<quantity id="664C8AE7A962B723DB1CFD8BFD2EFDD9" box="[606,763,583,608]" metricMagnitude="3" metricUnit="m" metricValue="2.6675" metricValueMax="2.715" metricValueMin="2.62" pageId="12" pageNumber="351" unit="m" value="2667.5" valueMax="2715.0" valueMin="2620.0">26202715 m</quantity>
,
<date id="D50A01C2A962B723DA45FD8BFC41FDE6" box="[775,916,583,608]" pageId="12" pageNumber="351" value="2010-07-15">15-VII-2010</date>
(SMNK).
</paragraph>
<paragraph id="A10B2702A962B723D985FDA0FE91FD2E" blockId="12.[151,1437,151,861]" pageId="12" pageNumber="351">
<materialsCitation id="11DC2D5FA962B723D985FDA0FE91FD2E" ID-GBIF-Occurrence="1456819368" collectionCode="WEST, NEW" country="Guinea-Bissau" latitude="-3.65" location="Mokndoma" longLatPrecision="1307" longitude="137.76666" pageId="12" pageNumber="351" specimenCount="1">
<emphasis id="93C0FB10A962B723D985FDA0FE3FFD03" bold="true" box="[199,490,620,645]" pageId="12" pageNumber="351">Other material reported</emphasis>
.
<collectionCode id="C7A5BFC7A962B723D8BAFDA0FD91FD02" box="[504,580,620,644]" pageId="12" pageNumber="351">WEST</collectionCode>
<collectionCode id="C7A5BFC7A962B723DB09FDA1FD5EFD02" box="[587,651,621,644]" country="Australia" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="12" pageNumber="351">NEW</collectionCode>
<collectingCountry id="D9A36792A962B723DBD0FDA0FCD5FD02" box="[658,768,620,644]" name="Guinea-Bissau" pageId="12" pageNumber="351">GUINEA</collectingCountry>
,
<location id="A46B71D9A962B723DA49FDA0FC5AFD03" LSID="urn:lsid:plazi:treatment:291D9614A964B723D9D5F9E1FD13FCDB:A46B71D9A962B723DA49FDA0FC5AFD03" box="[779,911,620,645]" country="Guinea-Bissau" latitude="-3.65" longLatPrecision="1307" longitude="137.76666" name="Mokndoma" pageId="12" pageNumber="351">Mokndoma</location>
,
<geoCoordinate id="C48041C5A962B723DAD9FDA0FBD0FD02" box="[923,1029,620,644]" direction="south" orientation="latitude" pageId="12" pageNumber="351" precision="925" value="-3.65">S 03° 39´</geoCoordinate>
<geoCoordinate id="C48041C5A962B723DD4EFDA1FB53FD03" box="[1036,1158,620,645]" direction="east" orientation="longitude" pageId="12" pageNumber="351" precision="925" value="137.76666">E 137° 46´</geoCoordinate>
(photographic evidence, see fig. 40ab).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="E9AE7489A962B723D985FD78FE66FD69" pageId="12" pageNumber="351" type="distribution">
<paragraph id="A10B2702A962B723D985FD78FE66FD69" blockId="12.[151,1437,151,861]" pageId="12" pageNumber="351">
<emphasis id="93C0FB10A962B723D985FD78FE8CFD4B" bold="true" box="[199,345,692,717]" pageId="12" pageNumber="351">Distribution</emphasis>
(
<figureCitation id="398F3B87A962B723D832FD78FE19FD4A" box="[368,460,692,717]" captionStart="FIGURE 41" captionStartId="12.[151,250,1614,1636]" captionTargetBox="[155,1436,916,1593]" captionTargetId="figure@12.[151,1436,916,1593]" captionTargetPageId="12" captionText="FIGURE 41. Distribution of G. (Niphetoscapha) wichmanni and G. (Niphetoscapha) inexspectatus sp. n.. Base map created with GeoMapApp (http: // www. geomapapp. org)." httpUri="https://zenodo.org/record/546837/files/figure.png" pageId="12" pageNumber="351">Fig. 41</figureCitation>
). Paniai Reg. (Sinak, Bilogay), Jayawijaya Reg. (Lake Habbema, Tiom, Mulia). Elevation:
<quantity id="664C8AE7A962B723D853FD14FE7AFD69" box="[273,431,728,752]" metricMagnitude="3" metricUnit="m" metricValue="2.55" metricValueMax="2.9" metricValueMin="2.2" pageId="12" pageNumber="351" unit="m" value="2550.0" valueMax="2900.0" valueMin="2200.0">22002900 m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="E9AE7489A962B723D985FD30FD13FCDB" pageId="12" pageNumber="351" type="etymology">
<paragraph id="A10B2702A962B723D985FD30FD13FCDB" blockId="12.[151,1437,151,861]" pageId="12" pageNumber="351">
<emphasis id="93C0FB10A962B723D985FD30FE93FC93" bold="true" box="[199,326,764,789]" pageId="12" pageNumber="351">Etymology</emphasis>
. This epithet is based on Latin participle
<taxonomicName id="66B45C81A962B723DA12FD31FC32FC92" box="[848,999,765,788]" class="Insecta" family="Curculionidae" genus="Gymnopholus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="351" phylum="Arthropoda" rank="species" species="inexspectatus">
<emphasis id="93C0FB10A962B723DA12FD31FC32FC92" box="[848,999,765,788]" italics="true" pageId="12" pageNumber="351">inexspectatus</emphasis>
</taxonomicName>
(unexpected) and refers to the first discovery of a
<taxonomicName id="66B45C81A962B723D87EFCECFE02FCBE" box="[316,471,800,824]" class="Insecta" family="Curculionidae" genus="Gymnopholus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="351" phylum="Arthropoda" rank="genus">
<emphasis id="93C0FB10A962B723D87EFCECFE02FCBE" box="[316,471,800,824]" italics="true" pageId="12" pageNumber="351">Gymnopholus</emphasis>
</taxonomicName>
species with epizoic symbiosis in West New
<collectingCountry id="D9A36792A962B723DA97FCD3FBF3FCBE" box="[981,1062,799,824]" name="Guinea-Bissau" pageId="12" pageNumber="351">Guinea</collectingCountry>
leaving a wide distributional gap to similar species in the
<collectingRegion id="6370E9E0A962B723D8E4FC89FE2FFCDA" box="[422,506,837,860]" country="American Samoa" name="Eastern" pageId="12" pageNumber="351">Eastern</collectingRegion>
half of the island.
</paragraph>
</subSubSection>
</treatment>
</document>