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<document id="2D2EE0C32C4AB232C0094998F0C841F7" ID-CLB-Dataset="55748" ID-DOI="10.5281/zenodo.5301289" ID-GBIF-Dataset="55b2b0bc-d79f-48e3-8be1-81424185dea7" ID-ISSN="0253-116X" ID-Zenodo-Dep="5301289" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630101010072" checkinUser="marcus" docAuthor="Assing, V." docDate="2013" docId="1974879EFFF3FFD1FF04EB40FD32FC3B" docLanguage="en" docName="LinzerbiolBeitr.2013.45.2.1479-1520.pdf" docOrigin="Linzer biologische Beiträge 45 (2)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Scymbalium badium" docType="treatment" docVersion="10" lastPageNumber="1487" masterDocId="E54DFFE6FFF5FFD9FFB6EA15FFE8FFA6" masterDocTitle="On the Palaearctic and Oriental species of Scymbalium and Micrillus (Coleoptera: Staphylinidae: Paederinae)" masterLastPageNumber="1520" masterPageNumber="1479" pageNumber="1485" updateTime="1699152599027" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="E517828A5EEDA462BA96A3DA0FA4489A">On the Palaearctic and Oriental species of Scymbalium and Micrillus (Coleoptera: Staphylinidae: Paederinae)</mods:title>
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<mods:namePart id="1092BE0F5A38259C35BB5CA7CB1FE7AA">Assing, V.</mods:namePart>
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<mods:title id="2E06A3C486179583D86BAE51EAEE3E7A">Linzer biologische Beiträge</mods:title>
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<mods:date id="053DFA8446DD95311463D2691784F11F">2013</mods:date>
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<mods:number id="9BD7B05E7480D1A6BE1D5A8C4ECC847C">2013-12-20</mods:number>
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<mods:classification id="D3A3654EDE60D20CF95A0E93BB6F36C1">journal article</mods:classification>
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<mods:identifier id="D37CEEA978263A0152222A96E4FB88CE" type="DOI">10.5281/zenodo.5301289</mods:identifier>
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<treatment id="1974879EFFF3FFD1FF04EB40FD32FC3B" ID-DOI="http://doi.org/10.5281/zenodo.6512311" ID-GBIF-Taxon="194955307" ID-Zenodo-Dep="6512311" LSID="urn:lsid:plazi:treatment:1974879EFFF3FFD1FF04EB40FD32FC3B" httpUri="http://treatment.plazi.org/id/1974879EFFF3FFD1FF04EB40FD32FC3B" lastPageId="8" lastPageNumber="1487" pageId="6" pageNumber="1485">
<subSubSection id="D9C76503FFF3FFDFFF04EB40FD16FECD" box="[178,766,341,363]" pageId="6" pageNumber="1485" type="nomenclature">
<paragraph id="91623688FFF3FFDFFF04EB40FD16FECD" blockId="6.[178,766,341,363]" box="[178,766,341,363]" pageId="6" pageNumber="1485">
<heading id="CA2A81E4FFF3FFDFFF04EB40FD16FECD" box="[178,766,341,363]" fontSize="8" level="2" pageId="6" pageNumber="1485" reason="2">
<taxonomicName id="56DD4D0BFFF3FFDFFF04EB40FD92FECD" authority="(MOTSCHULSKY 1858)" baseAuthorityName="Motschulsky" baseAuthorityYear="1858" box="[178,634,341,363]" class="Insecta" family="Staphylinidae" genus="Scymbalium" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1485" phylum="Arthropoda" rank="species" species="badium">
<emphasis id="A3A9EA9AFFF3FFDFFF04EB40FD92FECD" bold="true" box="[178,634,341,363]" pageId="6" pageNumber="1485">
<emphasis id="A3A9EA9AFFF3FFDFFF04EB40FE61FECD" bold="true" box="[178,393,341,363]" italics="true" pageId="6" pageNumber="1485">Scymbalium badium</emphasis>
(MOTSCHULSKY 1858)
</emphasis>
</taxonomicName>
(
<figureCitation id="09E62A0DFFF3FFDFFD31EB40FD1FFECD" box="[647,759,341,363]" captionStart="Figs 56-62" captionStartId="32.[178,218,1520,1540]" captionTargetBox="[190,1085,222,1489]" captionTargetId="figure-68@32.[178,1085,212,1497]" captionTargetPageId="32" captionText="Figs 56-62: Scymbalium badium (MOTSCHULSKY), lectotype (56-60) and S. nepalense nov.sp., holotype (61-62): (56) forebody; (57, 61) median dorsal portion of head; (58, 62) male sternite VIII; (59-60) aedeagus in lateral and in ventral view. Scale bars: 56: 1.0 mm; 58, 62: 0.5 mm; 59- 60: 0.2 mm; 57, 61: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5301311" httpUri="https://zenodo.org/record/5301311/files/figure.png" pageId="6" pageNumber="1485">Figs 56-60</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="D9C76503FFF3FFDFFF04EB68FD0BFE63" pageId="6" pageNumber="1485" type="reference_group">
<paragraph id="91623688FFF3FFDFFF04EB68FD7CFE37" blockId="6.[178,1086,381,728]" box="[178,660,381,401]" pageId="6" pageNumber="1485">
<treatmentCitationGroup id="B1CD11A6FFF3FFDFFF04EB68FD78FE37" box="[178,656,381,401]" pageId="6" pageNumber="1485">
<treatmentCitation id="107C1099FFF3FFDFFF04EB68FD78FE37" box="[178,656,381,401]" pageId="6" pageNumber="1485">
<taxonomicName id="56DD4D0BFFF3FFDFFF04EB68FD78FE37" ID-CoL="9PBMD" authority="MOTSCHULSKY 1858: 646" authorityName="MOTSCHULSKY" authorityPageNumber="646" authorityYear="1858" box="[178,656,381,401]" class="Insecta" family="Staphylinidae" genus="Lathrobomorphus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1485" phylum="Arthropoda" rank="species" species="badius">
<emphasis id="A3A9EA9AFFF3FFDFFF04EB68FE71FE37" box="[178,409,381,401]" italics="true" pageId="6" pageNumber="1485">Lathrobomorphus badius</emphasis>
MOTSCHULSKY 1858: 646
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
.
</paragraph>
<paragraph id="91623688FFF3FFDFFF04EB82FD22FE0D" blockId="6.[178,1086,381,728]" box="[178,714,407,427]" pageId="6" pageNumber="1485">
<treatmentCitationGroup id="B1CD11A6FFF3FFDFFF04EB82FD22FE0D" box="[178,714,407,427]" pageId="6" pageNumber="1485">
<treatmentCitation id="107C1099FFF3FFDFFF04EB82FD98FE0D" author="BERNHAUER M" box="[178,624,407,427]" page="13" pageId="6" pageNumber="1485" year="1928">
<taxonomicName id="56DD4D0BFFF3FFDFFF04EB82FD98FE0D" authority="BERNHAUER 1928: 13" authorityName="Bernhauer" authorityPageNumber="13" authorityYear="1928" box="[178,624,407,427]" class="Insecta" family="Staphylinidae" genus="Scymbalium" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1485" phylum="Arthropoda" rank="species" species="waageni">
<emphasis id="A3A9EA9AFFF3FFDFFF04EB82FEF5FE0D" box="[178,285,407,427]" italics="true" pageId="6" pageNumber="1485">Scimbalium</emphasis>
[sic]
<emphasis id="A3A9EA9AFFF3FFDFFEE7EB82FE76FE0D" box="[337,414,407,427]" italics="true" pageId="6" pageNumber="1485">waageni</emphasis>
<bibRefCitation id="F54C4B79FFF3FFDFFE15EB82FD98FE0D" author="BERNHAUER M" box="[419,624,407,427]" pageId="6" pageNumber="1485" pagination="1 - 28" refId="ref22774" refString="BERNHAUER M. (1928): Dr. E. Mjobergs zoological collections from Sumatra. 8. Staphylinidae. - Arkiv for Zoologi 19 A (19) [1927]: 1 - 28." type="journal article" year="1928">BERNHAUER 1928: 13</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<emphasis id="A3A9EA9AFFF3FFDFFDCCEB82FD22FE0D" bold="true" box="[634,714,407,427]" pageId="6" pageNumber="1485">nov.syn.</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="91623688FFF3FFDFFF04EBA4FD0BFE63" blockId="6.[178,1086,381,728]" box="[178,739,433,453]" pageId="6" pageNumber="1485">
<treatmentCitationGroup id="B1CD11A6FFF3FFDFFF04EBA4FD0BFE63" box="[178,739,433,453]" pageId="6" pageNumber="1485">
<treatmentCitation id="107C1099FFF3FFDFFF04EBA4FD9CFE63" author="CAMERON M" box="[178,628,433,453]" page="213" pageId="6" pageNumber="1485" year="1931">
<taxonomicName id="56DD4D0BFFF3FFDFFF04EBA4FD9CFE63" authority="CAMERON 1931: 213" authorityName=", CAMERON" authorityPageNumber="213" authorityYear="1931" box="[178,628,433,453]" class="Insecta" family="Staphylinidae" genus="Scymbalium" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1485" phylum="Arthropoda" rank="species" species="brunneum">
<emphasis id="A3A9EA9AFFF3FFDFFF04EBA4FEF5FE63" box="[178,285,433,453]" italics="true" pageId="6" pageNumber="1485">Scimbalium</emphasis>
[sic]
<emphasis id="A3A9EA9AFFF3FFDFFEE7EBA4FE45FE63" box="[337,429,433,453]" italics="true" pageId="6" pageNumber="1485">brunneum</emphasis>
<bibRefCitation id="F54C4B79FFF3FFDFFE05EBA4FD9CFE63" author="CAMERON M" box="[435,628,433,453]" pageId="6" pageNumber="1485" refId="ref22842" refString="CAMERON M (1931) The Fauna of British India including Ceylon and Burma. Coleoptera. Staphylinidae. Volume 2. - London, Taylor and Francis: viii + 257 pp." type="book" year="1931">
CAMERON 1931:
<specimenCount id="87DBFD01FFF3FFDFFDE4EBA4FD9CFE63" box="[594,628,433,453]" pageId="6" pageNumber="1485" type="generic">213</specimenCount>
</bibRefCitation>
</taxonomicName>
</treatmentCitation>
f.;
<emphasis id="A3A9EA9AFFF3FFDFFD25EBA4FD0BFE63" bold="true" box="[659,739,433,453]" pageId="6" pageNumber="1485">nov.syn.</emphasis>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="D9C76503FFF3FFDFFF04EBDFFD7AFD7E" pageId="6" pageNumber="1485" type="reference_group">
<paragraph id="91623688FFF3FFDFFF04EBDFFD30FD9F" blockId="6.[178,1086,381,728]" pageId="6" pageNumber="1485">
Type material examined
<taxonomicName id="56DD4D0BFFF3FFDFFDDDEBDFFD38FE78" baseAuthorityName="Motschulsky" baseAuthorityYear="1858" box="[619,720,458,478]" class="Insecta" family="Staphylinidae" genus="Scymbalium" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1485" phylum="Arthropoda" rank="species" species="badium">
<emphasis id="A3A9EA9AFFF3FFDFFDDDEBDFFD38FE78" box="[619,720,458,478]" italics="true" pageId="6" pageNumber="1485">S. badium</emphasis>
</taxonomicName>
:
<materialsCitation id="21B53CD5FFF3FFDFFD51EBDFFCAFFDAA" ID-GBIF-Occurrence="3760604446" collectionCode="ZMMU" determinerName="V. Assing" pageId="6" pageNumber="1485" specimenCount="3" typeStatus="lectotype">
<typeStatus id="4E66882AFFF3FFDFFD51EBDFFCAAFE78" box="[743,834,458,478]" pageId="6" pageNumber="1485" type="lectotype">Lectotype</typeStatus>
, present designation: &quot;
<taxonomicName id="56DD4D0BFFF3FFDFFF0FEBF4FDFCFE53" authority="Motch. Ind." authorityName="Motch. Ind." box="[185,532,481,501]" class="Insecta" family="Staphylinidae" genus="Luzea" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1485" phylum="Arthropoda" rank="species" species="badius">Lathrobomorphus badius Motch. Ind.</taxonomicName>
or. / Lectotypus
<taxonomicName id="56DD4D0BFFF3FFDFFD7AEBF4FBC7FE53" authority="Motschulsky" authorityName="Motschulsky" authorityYear="1895" box="[716,1071,481,501]" class="Insecta" family="Staphylinidae" genus="Luzea" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1485" phylum="Arthropoda" rank="species" species="badius">
<emphasis id="A3A9EA9AFFF3FFDFFD7AEBF4FC5CFE53" box="[716,948,481,501]" italics="true" pageId="6" pageNumber="1485">Lathrobomorphus badius</emphasis>
Motschulsky
</taxonomicName>
/
<taxonomicName id="56DD4D0BFFF3FFDFFF04EBEDFDECFDAA" authority="(Motschulsky)" baseAuthorityName="Motschulsky" baseAuthorityYear="1858" box="[178,516,504,524]" class="Insecta" family="Staphylinidae" genus="Scymbalium" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1485" phylum="Arthropoda" rank="species" species="badium">Scymbalium badium (Motschulsky)</taxonomicName>
, det.
<determinerName id="64BEDA4CFFF3FFDFFD88EBEDFD76FDAA" box="[574,670,504,524]" pageId="6" pageNumber="1485">V. Assing</determinerName>
2013&quot; (
<collectionCode id="F7CCAE4DFFF3FFDFFD46EBEDFCD4FDAA" box="[752,828,504,524]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="6" pageNumber="1485" type="Museum">ZMMU</collectionCode>
).
</materialsCitation>
<materialsCitation id="21B53CD5FFF3FFDFFCE6EBEDFD2EFD84" ID-GBIF-Occurrence="3760604371" collectionCode="ZMMU" determinerName="V. Assing" pageId="6" pageNumber="1485" specimenCount="3" typeStatus="paralectotype">
<typeStatus id="4E66882AFFF3FFDFFCE6EBEDFC30FDAA" box="[848,984,504,524]" pageId="6" pageNumber="1485" type="paralectotype">Paralectotypes</typeStatus>
: 1: with small remains of yellow label; writing illegible (ZMMU)
</materialsCitation>
;
<materialsCitation id="21B53CD5FFF3FFDFFD67E81BFD3CFD9F" ID-GBIF-Occurrence="3760604426" collectionCode="ZMMU" country="Myanmar" location="Birma" pageId="6" pageNumber="1485" specimenCount="1" typeStatus="lectotype">
1 [head and most of thorax missing]: &quot;Lathrobomorphus
<taxonomicName id="56DD4D0BFFF3FFDFFED2E830FDF9FD9F" authority="Motch. Ind." authorityName="Motch. Ind." box="[356,529,549,569]" class="Insecta" family="Staphylinidae" genus="Luzea" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1485" phylum="Arthropoda" rank="species" species="badius">badius Motch. Ind.</taxonomicName>
or.
<collectingCountry id="E9CA7618FFF3FFDFFD82E830FD87FD9F" box="[564,623,549,569]" name="Myanmar" pageId="6" pageNumber="1485">Birma</collectingCountry>
&quot; (
<collectionCode id="F7CCAE4DFFF3FFDFFD34E830FD26FD9F" box="[642,718,549,569]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="6" pageNumber="1485" type="Museum">ZMMU</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="91623688FFF3FFDFFF04E829FDB6FD32" blockId="6.[178,1086,381,728]" pageId="6" pageNumber="1485">
<taxonomicName id="56DD4D0BFFF3FFDFFF04E829FEF4FDF6" authorityName="Bernhauer" authorityYear="1928" box="[178,284,572,592]" class="Insecta" family="Staphylinidae" genus="Scymbalium" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1485" phylum="Arthropoda" rank="species" species="waageni">
<emphasis id="A3A9EA9AFFF3FFDFFF04E829FEF4FDF6" box="[178,284,572,592]" italics="true" pageId="6" pageNumber="1485">S. waageni</emphasis>
</taxonomicName>
:
<materialsCitation id="21B53CD5FFF3FFDFFE99E829FDB6FD32" ID-GBIF-Occurrence="3760604419" collectionCode="FMNH" determinerName="V. Assing" pageId="6" pageNumber="1485" specimenCount="2" typeStatus="holotype">
<typeStatus id="4E66882AFFF3FFDFFE99E829FE6BFDF6" box="[303,387,572,592]" pageId="6" pageNumber="1485" type="holotype">Holotype</typeStatus>
: &quot;Calcutta /
<collectorName id="3C28535EFFF3FFDFFD9EE829FD46FDF6" box="[552,686,572,592]" pageId="6" pageNumber="1485">Coll. Waagen</collectorName>
, don. Staudinger. / Waageni. Eppelsh. / Scimbalium / Scimbalium Waageni Brnh. Typus / Chicago NHMus,
<collectorName id="3C28535EFFF3FFDFFCFAE847FBC5FDC0" box="[844,1069,594,614]" pageId="6" pageNumber="1485">M.Bernhauer Collection</collectorName>
/ Holotypus
<taxonomicName id="56DD4D0BFFF3FFDFFE97E87CFD8AFDDB" authority="Bernhauer" authorityName="Bernhauer" authorityYear="1928" box="[289,610,617,637]" class="Insecta" family="Staphylinidae" genus="Scymbalium" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1485" phylum="Arthropoda" rank="species" species="waageni">Scymbalium waageni Bernhauer</taxonomicName>
, rev.
<collectorName id="3C28535EFFF3FFDFFD1FE87CFCE7FDDB" box="[681,783,617,637]" pageId="6" pageNumber="1485">V. Assing</collectorName>
2013 /
<taxonomicName id="56DD4D0BFFF3FFDFFCC3E87CFED1FD32" authority="(Motschulsky)" baseAuthorityName="Motschulsky" baseAuthorityYear="1858" class="Insecta" family="Staphylinidae" genus="Scymbalium" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1485" phylum="Arthropoda" rank="species" species="badium">Scymbalium badium (Motschulsky)</taxonomicName>
, det.
<determinerName id="64BEDA4CFFF3FFDFFEDFE895FE2CFD32" box="[361,452,640,660]" pageId="6" pageNumber="1485">V. Assing</determinerName>
2013&quot; (
<collectionCode id="F7CCAE4DFFF3FFDFFDBBE895FDBCFD32" box="[525,596,640,660]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="6" pageNumber="1485" type="Museum">FMNH</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="91623688FFF3FFDFFF04E882FD7AFD7E" blockId="6.[178,1086,381,728]" pageId="6" pageNumber="1485">
<taxonomicName id="56DD4D0BFFF3FFDFFF04E882FECDFD0D" authorityName=", CAMERON" authorityYear="1931" box="[178,293,663,683]" class="Insecta" family="Staphylinidae" genus="Scymbalium" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1485" phylum="Arthropoda" rank="species" species="brunneum">
<emphasis id="A3A9EA9AFFF3FFDFFF04E882FECDFD0D" box="[178,293,663,683]" italics="true" pageId="6" pageNumber="1485">S. brunneum</emphasis>
</taxonomicName>
:
<materialsCitation id="21B53CD5FFF3FFDFFE84E882FD7AFD7E" ID-GBIF-Occurrence="3760604367" pageId="6" pageNumber="1485" typeStatus="syntype">
<typeStatus id="4E66882AFFF3FFDFFE84E882FE94FD0D" box="[306,380,663,683]" pageId="6" pageNumber="1485" type="syntype">Syntype</typeStatus>
[without antennae]: &quot;
<collectingCountry id="E9CA7618FFF3FFDFFDEDE882FD63FD0D" box="[603,651,663,683]" name="India" pageId="6" pageNumber="1485">India</collectingCountry>
/ Bowring 63.47* / Scimbalium badium. Mots. / Scimb.
<taxonomicName id="56DD4D0BFFF3FFDFFEA5E8B8FE4FFD67" authority="Cam" authorityName=", CAMERON" authorityYear="1931" box="[275,423,685,705]" class="Insecta" family="Staphylinidae" genus="Scymbalium" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1485" phylum="Arthropoda" rank="species" species="brunneum">brunneum Cam</taxonomicName>
/ M.Cameron. Bequest. B.M. 1955-147. / Syntype /
<taxonomicName id="56DD4D0BFFF3FFDFFC7CE8B8FE82FD7E" authority="Cameron" authorityName=", CAMERON" authorityYear="1931" class="Insecta" family="Staphylinidae" genus="Scymbalium" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1485" phylum="Arthropoda" rank="species" species="brunneum">Scymbalium brunneum Cameron</taxonomicName>
, det. V. Assing 2013&quot; (BMNH).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="D9C76503FFF3FFD1FF04E8F3FD32FC3B" lastPageId="8" lastPageNumber="1487" pageId="6" pageNumber="1485" type="reference_group">
<paragraph id="91623688FFF3FFDFFF04E8F3FBD0FCD4" blockId="6.[178,1085,742,882]" pageId="6" pageNumber="1485">
Comment: The original description of
<taxonomicName id="56DD4D0BFFF3FFDFFDCFE8F2FD3FFD5A" authority="Motch. Ind." authorityName="Motch. Ind." box="[633,727,742,764]" class="Insecta" family="Staphylinidae" genus="Luzea" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1485" phylum="Arthropoda" rank="species" species="badius">
<emphasis id="A3A9EA9AFFF3FFDFFDCFE8F2FD3FFD5A" box="[633,727,742,764]" italics="true" pageId="6" pageNumber="1485">L. badius</emphasis>
</taxonomicName>
is based on an unspecified number of
<typeStatus id="4E66882AFFF3FFDFFF79E911FEC0FCBF" box="[207,296,772,793]" pageId="6" pageNumber="1485" type="syntype">syntypes</typeStatus>
from &quot;Indes orientales&quot; (MOTSCHULSKY 1858). Three
<typeStatus id="4E66882AFFF3FFDFFCEDE911FC5CFCBF" box="[859,948,772,793]" pageId="6" pageNumber="1485" type="syntype">syntypes</typeStatus>
were located in the Motschulsky collection at the ZMMU, a male and a female in moderate condition, and
<specimenCount id="87DBFD01FFF3FFDFFF68E92AFEA6FCF2" box="[222,334,830,852]" pageId="6" pageNumber="1485" type="female">one female</specimenCount>
in very poor condition (parts of three legs, parts of the ventral structures of the thorax, and most of the abdomen present). The male is designated as the
<typeStatus id="4E66882AFFF3FFDFFC62E949FBDCFCD4" box="[980,1076,860,882]" pageId="6" pageNumber="1485" type="lectotype">lectotype</typeStatus>
.
</paragraph>
<paragraph id="91623688FFF3FFDFFF04E994FEDFFBAA" blockId="6.[178,1085,897,1036]" pageId="6" pageNumber="1485">
<taxonomicName id="56DD4D0BFFF3FFDFFF04E994FE7AFC31" authorityName="Bernhauer" authorityYear="1928" box="[178,402,897,919]" class="Insecta" family="Staphylinidae" genus="Scymbalium" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1485" phylum="Arthropoda" rank="species" species="waageni">
<emphasis id="A3A9EA9AFFF3FFDFFF04E994FE7AFC31" box="[178,402,897,919]" italics="true" pageId="6" pageNumber="1485">Scymbalium waageni</emphasis>
</taxonomicName>
was described from a
<typeStatus id="4E66882AFFF3FFDFFD28E994FD1FFC31" box="[670,759,897,919]" pageId="6" pageNumber="1485" type="holotype">holotype</typeStatus>
(&quot;Das typische Stück&quot;) from &quot;Calcutta&quot; and a
<typeStatus id="4E66882AFFF3FFDFFED3E98AFE54FC12" box="[357,444,927,948]" pageId="6" pageNumber="1485" type="paratype">paratype</typeStatus>
(&quot;Ein zweites Stück&quot;) from &quot;
<collectingRegion id="5319F86AFFF3FFDFFD58E98BFCACFC12" box="[750,836,926,948]" country="Indonesia" name="Sumatera Utara" pageId="6" pageNumber="1485">Sumatra</collectingRegion>
(Medan)&quot; (
<bibRefCitation id="F54C4B79FFF3FFDFFC09E98BFF00FC77" author="BERNHAUER M" pageId="6" pageNumber="1485" pagination="1 - 28" refId="ref22774" refString="BERNHAUER M. (1928): Dr. E. Mjobergs zoological collections from Sumatra. 8. Staphylinidae. - Arkiv for Zoologi 19 A (19) [1927]: 1 - 28." type="journal article" year="1928">BERNHAUER 1928</bibRefCitation>
). The
<typeStatus id="4E66882AFFF3FFDFFE9CE9AEFE6DFC77" box="[298,389,955,977]" pageId="6" pageNumber="1485" type="holotype">holotype</typeStatus>
, a female, was located in the Bernhauer collection at the FMNH. It seems rather likely that the unexamined
<typeStatus id="4E66882AFFF3FFDFFDECE9CCFD59FC48" box="[602,689,985,1006]" pageId="6" pageNumber="1485" type="paratype">paratype</typeStatus>
from
<collectingRegion id="5319F86AFFF3FFDFFD42E9CCFCA0FC49" box="[756,840,985,1007]" country="Indonesia" name="Sumatera Utara" pageId="6" pageNumber="1485">Sumatra</collectingRegion>
is not conspecific with the
<typeStatus id="4E66882AFFF3FFDFFF61E9E3FEDBFBAA" box="[215,307,1014,1036]" pageId="6" pageNumber="1485" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="91623688FFF3FFDFFF04EE0EFC3DFAF0" blockId="6.[178,1086,1051,1366]" pageId="6" pageNumber="1485">
The original description of
<taxonomicName id="56DD4D0BFFF3FFDFFE67EE09FDBBFB97" authorityName=", CAMERON" authorityYear="1931" box="[465,595,1051,1073]" class="Insecta" family="Staphylinidae" genus="Scymbalium" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1485" phylum="Arthropoda" rank="species" species="brunneum">
<emphasis id="A3A9EA9AFFF3FFDFFE67EE09FDBBFB97" box="[465,595,1051,1073]" italics="true" pageId="6" pageNumber="1485">S. brunneum</emphasis>
</taxonomicName>
is based on an unspecified number of females (&quot;: unknown&quot;), possibly a unique specimen, from &quot;
<collectingCountry id="E9CA7618FFF3FFDFFD6CEE2DFCF8FBE8" box="[730,784,1080,1102]" name="India" pageId="6" pageNumber="1485">India</collectingCountry>
(without further indication)&quot; (
<bibRefCitation id="F54C4B79FFF3FFDFFF0FEE43FE8AFBCA" author="CAMERON M" box="[185,354,1110,1132]" pageId="6" pageNumber="1485" refId="ref22842" refString="CAMERON M (1931) The Fauna of British India including Ceylon and Burma. Coleoptera. Staphylinidae. Volume 2. - London, Taylor and Francis: viii + 257 pp." type="book" year="1931">CAMERON 1931</bibRefCitation>
). One
<typeStatus id="4E66882AFFF3FFDFFE1AEE42FE13FBCA" box="[428,507,1111,1132]" pageId="6" pageNumber="1485" type="syntype">syntype</typeStatus>
was located in the Cameron collection at the BMNH. When describing
<taxonomicName id="56DD4D0BFFF3FFDFFEDEEE61FD44FB2F" authority=", CAMERON (1931)" authorityName=", CAMERON" authorityYear="1931" box="[360,684,1139,1161]" class="Insecta" family="Staphylinidae" genus="Scymbalium" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1485" phylum="Arthropoda" rank="species" species="brunneum">
<emphasis id="A3A9EA9AFFF3FFDFFEDEEE61FE01FB2F" box="[360,489,1139,1161]" italics="true" pageId="6" pageNumber="1485">S. brunneum</emphasis>
,
<bibRefCitation id="F54C4B79FFF3FFDFFE4EEE66FD44FB2F" author="CAMERON M" box="[504,684,1139,1161]" pageId="6" pageNumber="1485" refId="ref22842" refString="CAMERON M (1931) The Fauna of British India including Ceylon and Burma. Coleoptera. Staphylinidae. Volume 2. - London, Taylor and Francis: viii + 257 pp." type="book" year="1931">CAMERON (1931)</bibRefCitation>
</taxonomicName>
was evidently unaware of the description of
<taxonomicName id="56DD4D0BFFF3FFDFFF4DEE84FE83FB00" authorityName="Bernhauer" authorityYear="1928" box="[251,363,1169,1190]" class="Insecta" family="Staphylinidae" genus="Scymbalium" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1485" phylum="Arthropoda" rank="species" species="waageni">
<emphasis id="A3A9EA9AFFF3FFDFFF4DEE84FE83FB00" box="[251,363,1169,1190]" italics="true" pageId="6" pageNumber="1485">S. waageni</emphasis>
</taxonomicName>
, as can be inferred from his mentioning the latter only in a preface to a subsequent work (
<bibRefCitation id="F54C4B79FFF3FFDFFEC0EEBBFDCAFB62" author="CAMERON M" box="[374,546,1198,1220]" pageId="6" pageNumber="1485" refId="ref22876" refString="CAMERON M (1932) The Fauna of British India including Ceylon and Burma. Coleoptera. Staphylinidae. Volume 3. - London, Taylor and Francis: viii + 443 pp." type="book" year="1932">CAMERON 1932</bibRefCitation>
). According to
<bibRefCitation id="F54C4B79FFF3FFDFFD66EEBBFC61FB62" author="CAMERON M" box="[720,905,1198,1220]" pageId="6" pageNumber="1485" refId="ref22842" refString="CAMERON M (1931) The Fauna of British India including Ceylon and Burma. Coleoptera. Staphylinidae. Volume 2. - London, Taylor and Francis: viii + 257 pp." type="book" year="1931">CAMERON (1931)</bibRefCitation>
,
<taxonomicName id="56DD4D0BFFF3FFDFFC2CEEBBFBF7FB62" authorityName=", CAMERON" authorityYear="1931" box="[922,1055,1198,1220]" class="Insecta" family="Staphylinidae" genus="Scymbalium" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1485" phylum="Arthropoda" rank="species" species="brunneum">
<emphasis id="A3A9EA9AFFF3FFDFFC2CEEBBFBF7FB62" box="[922,1055,1198,1220]" italics="true" pageId="6" pageNumber="1485">S. brunneum</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="56DD4D0BFFF3FFDFFECDEED9FE0BFB47" baseAuthorityName="Motschulsky" baseAuthorityYear="1858" box="[379,483,1227,1249]" class="Insecta" family="Staphylinidae" genus="Scymbalium" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1485" phylum="Arthropoda" rank="species" species="badium">
<emphasis id="A3A9EA9AFFF3FFDFFECDEED9FE0BFB47" box="[379,483,1227,1249]" italics="true" pageId="6" pageNumber="1485">S. badium</emphasis>
</taxonomicName>
by the darker coloration, a conclusion probably based on MOTSCHULSKY´ s (1858) statement that the body colour of
<taxonomicName id="56DD4D0BFFF3FFDFFCAFEEFCFC69FB58" baseAuthorityName="Motschulsky" baseAuthorityYear="1858" box="[793,897,1256,1278]" class="Insecta" family="Staphylinidae" genus="Scymbalium" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1485" phylum="Arthropoda" rank="species" species="badium">
<emphasis id="A3A9EA9AFFF3FFDFFCAFEEFCFC69FB58" box="[793,897,1256,1278]" italics="true" pageId="6" pageNumber="1485">S. badium</emphasis>
</taxonomicName>
was &quot;d'un testacé roux&quot;. The type specimens of
<taxonomicName id="56DD4D0BFFF3FFDFFE53EF13FDA3FABD" baseAuthorityName="Motschulsky" baseAuthorityYear="1858" box="[485,587,1285,1307]" class="Insecta" family="Staphylinidae" genus="Scymbalium" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1485" phylum="Arthropoda" rank="species" species="badium">
<emphasis id="A3A9EA9AFFF3FFDFFE53EF13FDA3FABD" box="[485,587,1285,1307]" italics="true" pageId="6" pageNumber="1485">S. badium</emphasis>
</taxonomicName>
, however, are all brown. A study of the types of
<taxonomicName id="56DD4D0BFFF3FFDFFF04EF36FEF5FA9F" baseAuthorityName="Motschulsky" baseAuthorityYear="1858" box="[178,285,1315,1337]" class="Insecta" family="Staphylinidae" genus="Scymbalium" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1485" phylum="Arthropoda" rank="species" species="badium">
<emphasis id="A3A9EA9AFFF3FFDFFF04EF36FEF5FA9F" box="[178,285,1315,1337]" italics="true" pageId="6" pageNumber="1485">S. badium</emphasis>
</taxonomicName>
,
<taxonomicName id="56DD4D0BFFF3FFDFFE86EF36FE4DFA9F" authorityName="Bernhauer" authorityYear="1928" box="[304,421,1315,1337]" class="Insecta" family="Staphylinidae" genus="Scymbalium" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1485" phylum="Arthropoda" rank="species" species="waageni">
<emphasis id="A3A9EA9AFFF3FFDFFE86EF36FE4DFA9F" box="[304,421,1315,1337]" italics="true" pageId="6" pageNumber="1485">S. waageni</emphasis>
</taxonomicName>
, and
<taxonomicName id="56DD4D0BFFF3FFDFFE5EEF36FD85FA9F" authorityName=", CAMERON" authorityYear="1931" box="[488,621,1315,1337]" class="Insecta" family="Staphylinidae" genus="Scymbalium" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1485" phylum="Arthropoda" rank="species" species="brunneum">
<emphasis id="A3A9EA9AFFF3FFDFFE5EEF36FD85FA9F" box="[488,621,1315,1337]" italics="true" pageId="6" pageNumber="1485">S. brunneum</emphasis>
</taxonomicName>
revealed no evidence suggesting that they should represent different species. Hence the new synonymies proposed above.
</paragraph>
<paragraph id="91623688FFF3FFDFFF04EF70FF12FA10" blockId="6.[178,1086,1381,1462]" pageId="6" pageNumber="1485">
Redescription: Body length
<quantity id="56259B6DFFF3FFDFFDE5EF70FD39FADD" box="[595,721,1381,1403]" metricMagnitude="-3" metricUnit="m" metricValue="8.9" metricValueMax="9.3" metricValueMin="8.5" pageId="6" pageNumber="1485" unit="mm" value="8.9" valueMax="9.3" valueMin="8.5">8.5-9.3 mm</quantity>
, length of forebody
<quantity id="56259B6DFFF3FFDFFC0DEF70FBD2FADD" box="[955,1082,1381,1403]" metricMagnitude="-3" metricUnit="m" metricValue="4.949999999999999" metricValueMax="5.1" metricValueMin="4.8" pageId="6" pageNumber="1485" unit="mm" value="4.949999999999999" valueMax="5.1" valueMin="4.8">4.8-5.1 mm</quantity>
. Coloration: body brown to dark-brown with reddish-brown elytra; legs brown to darkbrown.
</paragraph>
<caption id="C5A26600FFF2FFDEFF04EFCAFD81F986" ID-DOI="http://doi.org/10.5281/zenodo.5301293" ID-Zenodo-Dep="5301293" httpUri="https://zenodo.org/record/5301293/files/figure.png" pageId="7" pageNumber="1486" startId="7.[178,218,1503,1523]" targetBox="[182,1080,219,1485]" targetPageId="7">
<paragraph id="91623688FFF2FFDEFF04EFCAFD81F986" blockId="7.[178,1085,1503,1568]" pageId="7" pageNumber="1486">
<emphasis id="A3A9EA9AFFF2FFDEFF04EFCAFEE8FA55" bold="true" box="[178,256,1503,1523]" pageId="7" pageNumber="1486">Figs 2-8</emphasis>
:
<taxonomicName id="56DD4D0BFFF2FFDEFEB8EFCAFDA9FA57" ID-CoL="9QRS5" authority="CAMERON" authorityName="Cameron" authorityYear="1940" box="[270,577,1503,1523]" class="Insecta" family="Staphylinidae" genus="Scymbalium" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1486" phylum="Arthropoda" rank="species" species="persimile">
<emphasis id="A3A9EA9AFFF2FFDEFEB8EFCAFE31FA55" box="[270,473,1503,1523]" italics="true" pageId="7" pageNumber="1486">Scymbalium persimile</emphasis>
CAMERON
</taxonomicName>
, holotype (
<emphasis id="A3A9EA9AFFF2FFDEFD1AEFCAFD23FA55" bold="true" box="[684,715,1503,1523]" pageId="7" pageNumber="1486">2-4</emphasis>
) and
<taxonomicName id="56DD4D0BFFF2FFDEFCB4EFCAFC9EFA55" ID-CoL="9QRRW" authority="Assing, 2013" authorityName="Assing" authorityYear="2013" box="[770,886,1503,1523]" class="Insecta" family="Staphylinidae" genus="Scymbalium" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1486" phylum="Arthropoda" rank="species" species="nepalense" status="sp. nov.">
<emphasis id="A3A9EA9AFFF2FFDEFCB4EFCAFC9EFA55" box="[770,886,1503,1523]" italics="true" pageId="7" pageNumber="1486">S. nepalense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B89A57E1FFF2FFDEFCC8EFCAFC2AFA55" box="[894,962,1503,1523]" pageId="7" pageNumber="1486" rank="species">nov.sp.</taxonomicNameLabel>
, paratype (
<emphasis id="A3A9EA9AFFF2FFDEFB9CEFCAFF55F9AF" bold="true" pageId="7" pageNumber="1486">5- 8</emphasis>
): (
<emphasis id="A3A9EA9AFFF2FFDEFF6FEFE0FF14F9AF" bold="true" box="[217,252,1525,1545]" pageId="7" pageNumber="1486">2, 5</emphasis>
) habitus; (
<emphasis id="A3A9EA9AFFF2FFDEFED7EFE0FE64F9AF" bold="true" box="[353,396,1525,1545]" pageId="7" pageNumber="1486">3, 6)</emphasis>
forebody; (
<emphasis id="A3A9EA9AFFF2FFDEFE4CEFE0FDEDF9AF" bold="true" box="[506,517,1525,1545]" pageId="7" pageNumber="1486">4</emphasis>
) abdomen; (
<emphasis id="A3A9EA9AFFF2FFDEFDCDEFE0FD6EF9AF" bold="true" box="[635,646,1525,1545]" pageId="7" pageNumber="1486">7</emphasis>
) aedeagus in lateral view; (
<emphasis id="A3A9EA9AFFF2FFDEFC3EEFE0FC7BF9AF" bold="true" box="[904,915,1525,1545]" pageId="7" pageNumber="1486">8</emphasis>
) aedeagus in ventral view. Scale bars: 2-6: 1.0 mm; 7-8: 0.2 mm.
</paragraph>
</caption>
<paragraph id="91623688FFFDFFD1FF04EACFFD3BFEEE" blockId="8.[178,1086,218,328]" pageId="8" pageNumber="1487">
Head (
<figureCitation id="09E62A0DFFFDFFD1FF43EACFFE8CFF56" box="[245,356,218,240]" captionStart="Figs 56-62" captionStartId="32.[178,218,1520,1540]" captionTargetBox="[190,1085,222,1489]" captionTargetId="figure-68@32.[178,1085,212,1497]" captionTargetPageId="32" captionText="Figs 56-62: Scymbalium badium (MOTSCHULSKY), lectotype (56-60) and S. nepalense nov.sp., holotype (61-62): (56) forebody; (57, 61) median dorsal portion of head; (58, 62) male sternite VIII; (59-60) aedeagus in lateral and in ventral view. Scale bars: 56: 1.0 mm; 58, 62: 0.5 mm; 59- 60: 0.2 mm; 57, 61: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5301311" httpUri="https://zenodo.org/record/5301311/files/figure.png" pageId="8" pageNumber="1487">Figs 56-57</figureCitation>
) 1.00-1.04 times as long as broad; posterior angles moderately marked; punctation fine, shallow, and moderately dense; interstices with very fine transverse microsculpture. Eyes shorter than postocular region. Antenna approximately
<quantity id="56259B6DFFFDFFD1FC71EB00FBD5FE8D" box="[967,1085,277,299]" metricMagnitude="-3" metricUnit="m" metricValue="3.25" metricValueMax="3.3" metricValueMin="3.2" pageId="8" pageNumber="1487" unit="mm" value="3.25" valueMax="3.3" valueMin="3.2">3.2-3.3 mm</quantity>
long and slender; all antennomeres longer than broad.
</paragraph>
<paragraph id="91623688FFFDFFD1FF04EB4DFC80FE45" blockId="8.[178,1085,344,483]" pageId="8" pageNumber="1487">
Pronotum (
<figureCitation id="09E62A0DFFFDFFD1FE91EB4DFE9CFEC8" box="[295,372,344,366]" captionStart="Figs 56-62" captionStartId="32.[178,218,1520,1540]" captionTargetBox="[190,1085,222,1489]" captionTargetId="figure-68@32.[178,1085,212,1497]" captionTargetPageId="32" captionText="Figs 56-62: Scymbalium badium (MOTSCHULSKY), lectotype (56-60) and S. nepalense nov.sp., holotype (61-62): (56) forebody; (57, 61) median dorsal portion of head; (58, 62) male sternite VIII; (59-60) aedeagus in lateral and in ventral view. Scale bars: 56: 1.0 mm; 58, 62: 0.5 mm; 59- 60: 0.2 mm; 57, 61: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5301311" httpUri="https://zenodo.org/record/5301311/files/figure.png" pageId="8" pageNumber="1487">Fig. 56</figureCitation>
) 1.06-1.10 times as long as broad and 0.96-1.00 times as broad as head, widest near anterior angles; anterior margin weakly concave in the middle; posterior margin broadly concave; punctation very fine and dense; impunctate median band indistinct, narrow, present only in posterior half of midline, and not reaching posterior margin; interstices with or without shallow traces of microsculpture.
</paragraph>
<paragraph id="91623688FFFDFFD1FF04EBE7FBDBFDC6" blockId="8.[178,1086,498,608]" pageId="8" pageNumber="1487">
Elytra (
<figureCitation id="09E62A0DFFFDFFD1FEB6EBE7FEA3FDAE" box="[256,331,498,520]" captionStart="Figs 56-62" captionStartId="32.[178,218,1520,1540]" captionTargetBox="[190,1085,222,1489]" captionTargetId="figure-68@32.[178,1085,212,1497]" captionTargetPageId="32" captionText="Figs 56-62: Scymbalium badium (MOTSCHULSKY), lectotype (56-60) and S. nepalense nov.sp., holotype (61-62): (56) forebody; (57, 61) median dorsal portion of head; (58, 62) male sternite VIII; (59-60) aedeagus in lateral and in ventral view. Scale bars: 56: 1.0 mm; 58, 62: 0.5 mm; 59- 60: 0.2 mm; 57, 61: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5301311" httpUri="https://zenodo.org/record/5301311/files/figure.png" pageId="8" pageNumber="1487">Fig. 56</figureCitation>
) approximately as long as pronotum or slightly longer, with very dense, fine, and somewhat asperate punctation, and with distinct microsculpture, matt. Protarsomeres I-IV distinctly dilated in both sexes, without sexual dimorphism. Metatarsomere I longer than the combined length of II and III, nearly as long as the combined length of II-IV.
</paragraph>
<paragraph id="91623688FFFDFFD1FF04E87AFF18FD66" blockId="8.[178,1085,623,704]" pageId="8" pageNumber="1487">Abdomen with very fine and dense punctation, and with distinct microsculpture, nearly matt; posterior margin of tergite VIII with palisade fringe; tergite X and sternite IX very small.</paragraph>
<paragraph id="91623688FFFDFFD1FF71E8C5FDCCFC9A" blockId="8.[178,1085,719,829]" pageId="8" pageNumber="1487">
: posterior margin with concave, nearly semi-circular posterior excision (
<figureCitation id="09E62A0DFFFDFFD1FC69E8DAFBD8FD43" box="[991,1072,719,741]" captionStart="Figs 56-62" captionStartId="32.[178,218,1520,1540]" captionTargetBox="[190,1085,222,1489]" captionTargetId="figure-68@32.[178,1085,212,1497]" captionTargetPageId="32" captionText="Figs 56-62: Scymbalium badium (MOTSCHULSKY), lectotype (56-60) and S. nepalense nov.sp., holotype (61-62): (56) forebody; (57, 61) median dorsal portion of head; (58, 62) male sternite VIII; (59-60) aedeagus in lateral and in ventral view. Scale bars: 56: 1.0 mm; 58, 62: 0.5 mm; 59- 60: 0.2 mm; 57, 61: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5301311" httpUri="https://zenodo.org/record/5301311/files/figure.png" pageId="8" pageNumber="1487">Fig. 58</figureCitation>
); aedeagus (
<figureCitation id="09E62A0DFFFDFFD1FEA8E8F9FE67FCA4" box="[286,399,748,770]" captionStart="Figs 56-62" captionStartId="32.[178,218,1520,1540]" captionTargetBox="[190,1085,222,1489]" captionTargetId="figure-68@32.[178,1085,212,1497]" captionTargetPageId="32" captionText="Figs 56-62: Scymbalium badium (MOTSCHULSKY), lectotype (56-60) and S. nepalense nov.sp., holotype (61-62): (56) forebody; (57, 61) median dorsal portion of head; (58, 62) male sternite VIII; (59-60) aedeagus in lateral and in ventral view. Scale bars: 56: 1.0 mm; 58, 62: 0.5 mm; 59- 60: 0.2 mm; 57, 61: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5301311" httpUri="https://zenodo.org/record/5301311/files/figure.png" pageId="8" pageNumber="1487">Figs 59-60</figureCitation>
) small in relation to body,
<quantity id="56259B6DFFFDFFD1FD1DE8F9FCEFFCA4" box="[683,775,748,770]" metricMagnitude="-4" metricUnit="m" metricValue="6.8" pageId="8" pageNumber="1487" unit="mm" value="0.68">0.68 mm</quantity>
long, symmetric, and without distinctly sclerotized internal structures; parameres nearly completely reduced (see comment in the section on
<taxonomicName id="56DD4D0BFFFDFFD1FE2EE932FDFEFC9B" ID-CoL="9QRRW" authorityName="Assing" authorityYear="2013" box="[408,534,807,829]" class="Insecta" family="Staphylinidae" genus="Scymbalium" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1487" phylum="Arthropoda" rank="species" species="nepalense">
<emphasis id="A3A9EA9AFFFDFFD1FE2EE932FDFEFC9B" box="[408,534,807,829]" italics="true" pageId="8" pageNumber="1487">S. nepalense</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="91623688FFFDFFD1FF04E958FD32FC3B" blockId="8.[178,1085,844,925]" pageId="8" pageNumber="1487">
D i s t r i b u t i o n:
<taxonomicName id="56DD4D0BFFFDFFD1FE15E959FD9BFCC4" baseAuthorityName="Motschulsky" baseAuthorityYear="1858" box="[419,627,844,866]" class="Insecta" family="Staphylinidae" genus="Scymbalium" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1487" phylum="Arthropoda" rank="species" species="badium">
<emphasis id="A3A9EA9AFFFDFFD1FE15E959FD9BFCC4" box="[419,627,844,866]" italics="true" pageId="8" pageNumber="1487">Scymbalium badium</emphasis>
</taxonomicName>
has been reported, partly as
<taxonomicName id="56DD4D0BFFFDFFD1FC28E958FBF7FCC4" authorityName=", CAMERON" authorityYear="1931" box="[926,1055,844,866]" class="Insecta" family="Staphylinidae" genus="Scymbalium" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1487" phylum="Arthropoda" rank="species" species="brunneum">
<emphasis id="A3A9EA9AFFFDFFD1FC28E958FBF7FCC4" box="[926,1055,844,866]" italics="true" pageId="8" pageNumber="1487">S. brunneum</emphasis>
</taxonomicName>
or
<taxonomicName id="56DD4D0BFFFDFFD1FF04E97FFECFFCD9" authorityName="Bernhauer" authorityYear="1928" box="[178,295,874,895]" class="Insecta" family="Staphylinidae" genus="Scymbalium" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1487" phylum="Arthropoda" rank="species" species="waageni">
<emphasis id="A3A9EA9AFFFDFFD1FF04E97FFECFFCD9" box="[178,295,874,895]" italics="true" pageId="8" pageNumber="1487">S. waageni</emphasis>
</taxonomicName>
, from
<collectingCountry id="E9CA7618FFFDFFD1FEC3E97CFE44FCD9" box="[373,428,873,895]" name="India" pageId="8" pageNumber="1487">India</collectingCountry>
,
<collectingCountry id="E9CA7618FFFDFFD1FE0DE97CFDEFFCD9" box="[443,519,873,895]" name="Bhutan" pageId="8" pageNumber="1487">Bhutan</collectingCountry>
, and
<collectingCountry id="E9CA7618FFFDFFD1FDF0E97FFD40FCD9" box="[582,680,874,895]" name="Myanmar" pageId="8" pageNumber="1487">Myanmar</collectingCountry>
(BISWAS &amp; BISWAS 1995;
<bibRefCitation id="F54C4B79FFFDFFD1FC60E97CFF00FC3B" author="CAMERON M" pageId="8" pageNumber="1487" refId="ref22842" refString="CAMERON M (1931) The Fauna of British India including Ceylon and Burma. Coleoptera. Staphylinidae. Volume 2. - London, Taylor and Francis: viii + 257 pp." type="book" year="1931">CAMERON 1931</bibRefCitation>
;
<bibRefCitation id="F54C4B79FFFDFFD1FF44E992FE64FC3B" author="COIFFAIT H" box="[242,396,903,925]" pageId="8" pageNumber="1487" pagination="109 - 150" refId="ref23069" refString="COIFFAIT H. (1978): Ergebnisse der Bhutan-Expedition 1972 des Naturhistorischen Museums in Basel. Coleoptera: Fam. Staphylinidae Subfam. Paederinae, Euaesthetinae, Piestinae, Osoriinae et Omalinae [sic]. - Entomologica Basiliensia 3: 109 - 150." type="journal article" year="1978">COIFFAIT 1978</bibRefCitation>
;
<bibRefCitation id="F54C4B79FFFDFFD1FE20E992FDCCFC3B" author="FAUVEL A" box="[406,548,903,925]" pageId="8" pageNumber="1487" pagination="43 - 70" refId="ref23222" refString="FAUVEL A. (1904): Staphylinides de l'Hindoustan et de la Birmanie. - Revue d'Entomologie 23: 43 - 70." type="journal article" year="1904">FAUVEL 1904</bibRefCitation>
;
<bibRefCitation id="F54C4B79FFFDFFD1FD98E992FD27FC3B" author="SMETANA A" box="[558,719,903,925]" pageId="8" pageNumber="1487" pagination="579 - 624" refId="ref23296" refString="SMETANA A. (2004): Subfamily Paederinae FLEMING, 1821. - In: LOBL I. &amp; A. SMETANA (eds), Catalogue of Palaearctic Coleoptera. Volume 2. Hydrophiloidea - Histeroidea - Staphylinoidea. - Apollo Books, Stenstrup: 579 - 624." type="book chapter" year="2004">SMETANA 2004</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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