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<document id="5D69581291C9A1862CE591ED4A41E22A" ID-CLB-Dataset="30340" ID-DOI="10.5324/cjcr.v0i35.4832" ID-GBIF-Dataset="f404266e-58c5-4117-a23e-6ea2e373f22e" ID-ISSN="2387-5372" ID-Zenodo-Dep="7996326" IM.bibliography_approvedBy="juliana" IM.materialsCitations_approvedBy="felipe" IM.metadata_requiresApprovalFor="plazi" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1685471215087" checkinUser="felipe" docAuthor="Cranston, Peter S., Krosch, Matt N. &amp; Tang, Hongqu" docDate="2022" docId="136287C9FFD43159FF2227EE78B3F952" docLanguage="en" docName="ChiroJChiroRes.35.12-31.pdf" docOrigin="CHIRONOMUS Journal of Chironomidae Research 35" docStyle="DocumentStyle:B21B57A70ED88EDAB7971C4BDED0DA62.1:ChiroJChiroRes.2015-.journal_article" docStyleId="B21B57A70ED88EDAB7971C4BDED0DA62" docStyleName="ChiroJChiroRes.2015-.journal_article" docStyleVersion="1" docTitle="Nilotanypus ctenochelus Cranston &amp; Krosch &amp; Tang 2022, new species" docType="treatment" docUuid="139262E4-37D0-4487-BC82-093A9A05FBD7" docUuidSource="ZooBank" docVersion="7" lastPageNumber="25" masterDocId="EF5BFFB1FFDE3154FFC1242F793BFFF7" masterDocTitle="Verifying Australian Nilotanypus Kieffer (Chironomidae) In A Global Perspective: Molecular Phylogenetic And Temporal Analyses, New Species And Emended Generic Diagnoses" masterLastPageNumber="31" masterPageNumber="12" pageNumber="22" updateTime="1699511797472" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="77D7A3BCA9FF105F0578B174B90D4712" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="CC95F18C948BA875AC4293F311BB0008">
<mods:title id="46F5B86A8741EEB013508481A26D940F">Verifying Australian Nilotanypus Kieffer (Chironomidae) In A Global Perspective: Molecular Phylogenetic And Temporal Analyses, New Species And Emended Generic Diagnoses</mods:title>
</mods:titleInfo>
<mods:name id="0066F435B0B30BA6EDBBAABE16C89907" type="personal">
<mods:role id="9045F3EE76F327D9AF1D476733BEE9BC">
<mods:roleTerm id="5345FE9AFB16A8DC1F37EADE0CC998FC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="742F029135FD7D76E5A538D17E5D488F">Cranston, Peter S.</mods:namePart>
<mods:affiliation id="35EF9C084344823E4BC13986BD79DA02">Evolution, Ecology &amp; Genetics, Research School of Biology, Australian National University, Canberra, A. C. T. 2601, Australia.</mods:affiliation>
<mods:nameIdentifier id="3D41C0E8776B0B5B55160E80188F66AA" type="email">pscranston@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="5E79495FAE73CB4139071D789A2F0B52" type="personal">
<mods:role id="C2225DB0D8A4A1A63A8F69E31A2E5B6E">
<mods:roleTerm id="09818B302B305912D533E2846E303ED1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7A35E37A82D8FCB93891DDBED6D6943F">Krosch, Matt N.</mods:namePart>
<mods:affiliation id="1E7B123976DA3B76162DDCA07AF7324F">Forensic Services Group, Queensland Police Service, Brisbane, QLD 4000, Australia.</mods:affiliation>
<mods:nameIdentifier id="35D7C72C52BE4036D753F1ECBE96C780" type="email">krosch.mattn@police.qld.gov.au</mods:nameIdentifier>
</mods:name>
<mods:name id="48252B2AADD883126B42D78A8790A979" type="personal">
<mods:role id="AB4B58569F8B59E5FFF5D22B0E26DAC3">
<mods:roleTerm id="5221F6C99F9D7A33222B0008ABE96D88">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="493E305CFA393192F39F950737A0F126">Tang, Hongqu</mods:namePart>
<mods:affiliation id="5CB227615F65D504607CD3B5E133C795">Life Science and Technology College, Jinan University, Guangzhou 510632, China.</mods:affiliation>
<mods:nameIdentifier id="FFF8FD6F7E05CADE7F8047D12CBD16F0" type="email">townningt@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="B796408CCABD9E399270E467A03E8AB0">text</mods:typeOfResource>
<mods:relatedItem id="F22F46628B3DBE056EAE2CAFF3993F56" type="host">
<mods:titleInfo id="75119649FED2E53070277C2816DD029F">
<mods:title id="3E7A044C0548AC957AF0D7BA2EF6897C">CHIRONOMUS Journal of Chironomidae Research</mods:title>
</mods:titleInfo>
<mods:part id="06128873F5777282FA366EBDBD505C5D">
<mods:date id="526D2F036CFB97203D651838830801A5">2022</mods:date>
<mods:detail id="707EB77AC302080B0EA8A5C954AE5D3E" type="pubDate">
<mods:number id="78589967702B38D8E01BDAE4FFBF539F">2022-06-27</mods:number>
</mods:detail>
<mods:detail id="F0A32F812DD9F1667762348197527DD6" type="volume">
<mods:number id="90A7BA84C784CE299179724CF99D2355">35</mods:number>
</mods:detail>
<mods:extent id="16CAC6C798D4E9DC92E3FFF52453BD8B" unit="page">
<mods:start id="27E940877BA9EB3EFA9CF241FA0600A3">12</mods:start>
<mods:end id="B1EE2C6379559CC8F86F48E66EF9C3DC">31</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="5A7A8A84EA944783398F8B60B0212B57">journal article</mods:classification>
<mods:identifier id="189B645CD230456CB89CA43FBB3A245F" type="CLB-Dataset">30340</mods:identifier>
<mods:identifier id="36D266162639A310206C13AB0B2E9C69" type="DOI">10.5324/cjcr.v0i35.4832</mods:identifier>
<mods:identifier id="A0EDAE0A6C4E333F1B452F0F7036DA71" type="GBIF-Dataset">f404266e-58c5-4117-a23e-6ea2e373f22e</mods:identifier>
<mods:identifier id="8F75ADD0856D2B15BA5F2933E76D0960" type="ISSN">2387-5372</mods:identifier>
<mods:identifier id="58B311D5C61C37ABFBDB0B2949C1519A" type="Zenodo-Dep">7996326</mods:identifier>
<mods:identifier id="6D8BA57F80170230578A9BE1F680D6F5" type="ZooBank">94BBD58A-85FD-4E52-807E-97D27636437C</mods:identifier>
</mods:mods>
<treatment id="136287C9FFD43159FF2227EE78B3F952" ID-DOI="http://doi.org/10.5281/zenodo.7996336" ID-GBIF-Taxon="209665678" ID-Zenodo-Dep="7996336" LSID="urn:lsid:zoobank.org:act:139262E4-37D0-4487-BC82-093A9A05FBD7" httpUri="http://treatment.plazi.org/id/136287C9FFD43159FF2227EE78B3F952" lastPageId="13" lastPageNumber="25" pageId="10" pageNumber="22">
<subSubSection id="D3D16554FFD4315EFF2227EE7B40FC2E" box="[227,635,961,985]" pageId="10" pageNumber="22" type="nomenclature">
<paragraph id="9B7436DFFFD4315EFF2227EE7B40FC2E" blockId="10.[227,635,961,985]" box="[227,635,961,985]" pageId="10" pageNumber="22">
<heading id="C03C81B3FFD4315EFF2227EE7B40FC2E" bold="true" box="[227,635,961,985]" fontSize="10" level="2" pageId="10" pageNumber="22" reason="3">
<emphasis id="A9BFEACDFFD4315EFF2227EE7B40FC2E" bold="true" box="[227,635,961,985]" pageId="10" pageNumber="22">
<taxonomicName id="5CCB4D5CFFD4315EFF2227EE78CAFC2E" ID-CoL="9J66K" authority="Cranston &amp; Krosch &amp; Tang, 2022" authorityName="Cranston &amp; Krosch &amp; Tang" authorityYear="2022" box="[227,497,961,985]" class="Insecta" family="Chironomidae" genus="Nilotanypus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="22" phylum="Arthropoda" rank="species" species="ctenochelus" status="sp. nov.">
<emphasis id="A9BFEACDFFD4315EFF2227EE78CAFC2E" bold="true" box="[227,497,961,985]" italics="true" pageId="10" pageNumber="22">Nilotanypus ctenochelus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B28C57B6FFD4315EFE3927EE7B40FC2E" box="[504,635,961,985]" pageId="10" pageNumber="22" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="D3D16554FFD4315EFF2227DF78AEFBDE" pageId="10" pageNumber="22" type="description">
<paragraph id="9B7436DFFFD4315EFF2227DF78AEFBDE" blockId="10.[227,764,1008,1065]" pageId="10" pageNumber="22">
<emphasis id="A9BFEACDFFD4315EFF2227DF78AEFBDE" italics="true" pageId="10" pageNumber="22">
http://zoobank.org/
<uuid id="EF6D0C0AFFD4315EFE7927DF78AEFBDE" pageId="10" pageNumber="22">139262E4-37D0-4487-BC82- 093A9A05FBD7</uuid>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="D3D16554FFD4315EFF22206F7D1EFC0C" pageId="10" pageNumber="22" type="materials_examined">
<paragraph id="9B7436DFFFD4315EFF22206F782CFB0F" blockId="10.[227,764,1087,1272]" pageId="10" pageNumber="22">
<emphasis id="A9BFEACDFFD4315EFF22206F78A8FBA0" bold="true" box="[227,403,1087,1112]" pageId="10" pageNumber="22">Type material:</emphasis>
<materialsCitation id="2BA33C82FFD4315EFE63206F7B97FB4F" ID-GBIF-Occurrence="4122308333" collectingDate="1989-03-25" collectingDateMax="1989-03-26" collectingDateMin="1989-03-25" collectorName="Birthday Ck. &amp; Cranston" country="Australia" elevation="650" latitude="-18.983334" location="Paluma" longLatPrecision="1273" longitude="146.16667" municipality="Euparal" pageId="10" pageNumber="22" specimenCount="1" specimenCount-male="1" stateProvince="Queensland" typeStatus="holotype">
<emphasis id="A9BFEACDFFD4315EFE63206F7B3FFBAF" box="[418,516,1088,1112]" italics="true" pageId="10" pageNumber="22">
<typeStatus id="4470887DFFD4315EFE63206F7B3FFBAF" box="[418,516,1088,1112]" pageId="10" pageNumber="22" type="holotype">Holotype</typeStatus>
</emphasis>
:
<collectingCountry id="E3DC764FFFD4315EFDD8206F7B45FBAF" box="[537,638,1088,1112]" name="Australia" pageId="10" pageNumber="22">Australia</collectingCountry>
, P
<specimenCount id="8DCDFD56FFD4315EFD60206F7B8EFBA0" box="[673,693,1088,1111]" pageId="10" pageNumber="22" type="male">♂</specimenCount>
, slide mounted in
<collectingMunicipality id="7B10ACA5FFD4315EFEA8204F7884FB8F" box="[361,447,1120,1144]" pageId="10" pageNumber="22">Euparal</collectingMunicipality>
,
<collectingRegion id="590FF83DFFD4315EFE0C204F7B75FB8F" box="[461,590,1120,1144]" country="Australia" name="Queensland" pageId="10" pageNumber="22">Queensland</collectingRegion>
,
<location id="9E146004FFD4315EFD9A204F7B95FB8F" LSID="urn:lsid:plazi:treatment:136287C9FFD43159FF2227EE78B3F952:9E146004FFD4315EFD9A204F7B95FB8F" box="[603,686,1120,1144]" country="Australia" latitude="-18.983334" longLatPrecision="1273" longitude="146.16667" municipality="Euparal" name="Paluma" pageId="10" pageNumber="22" stateProvince="Queensland">Paluma</location>
,
<collectorName id="363E5309FFD4315EFD7D204F7800FB6F" pageId="10" pageNumber="22">Birthday Ck.</collectorName>
,
<geoCoordinate id="FEFF5018FFD4315EFE882050789BFB60" box="[329,416,1151,1175]" degrees="18" direction="south" minutes="59" orientation="latitude" pageId="10" pageNumber="22" precision="925" value="-18.983334">18°59’S</geoCoordinate>
<geoCoordinate id="FEFF5018FFD4315EFE6B20AF7B28FB6F" box="[426,531,1152,1176]" degrees="146" direction="east" minutes="10" orientation="longitude" pageId="10" pageNumber="22" precision="925" value="146.16667">146°10’E</geoCoordinate>
,
<elevation id="10E6D1ECFFD4315EFDE020507B98FB6F" box="[545,675,1151,1176]" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="10" pageNumber="22" unit="m" value="650.0">
<quantity id="5C339B3AFFD4315EFDE020507B5CFB60" box="[545,615,1151,1175]" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="10" pageNumber="22" unit="m" value="650.0">650 m</quantity>
a.s.l.
</elevation>
,
<date id="EF75101FFFD4315EFD7320507803FB4F" pageId="10" pageNumber="22" value="1989-03-25" valueMax="1989-03-26" valueMin="1989-03-25">
<collectingDate id="FF31E9F7FFD4315EFD7320507803FB4F" pageId="10" pageNumber="22" value="1989-03-25" valueMax="1989-03-26" valueMin="1989-03-25">25–26. iii.1989</collectingDate>
</date>
, leg.
<collectorName id="363E5309FFD4315EFEBE208F78D9FB4F" box="[383,482,1184,1208]" pageId="10" pageNumber="22">Cranston</collectorName>
, deposited ANIC
</materialsCitation>
.
<emphasis id="A9BFEACDFFD4315EFD7C208F7821FB20" italics="true" pageId="10" pageNumber="22">
<typeStatus id="4470887DFFD4315EFD7C208F7821FB20" pageId="10" pageNumber="22" type="paratype">Paratypes</typeStatus>
</emphasis>
, P
<specimenCount id="8DCDFD56FFD4315EFEFB20EF7875FB20" box="[314,334,1216,1239]" pageId="10" pageNumber="22" type="male">♂</specimenCount>
, 2P
<materialsCitation id="2BA33C82FFD4315EFEBF20EF7828FB0F" ID-GBIF-Occurrence="4122308360" collectingDate="1989-03-25" collectingDateMax="1989-03-26" collectingDateMin="1989-03-25" collectorName="Birthday Ck. &amp; Cranston" country="Australia" elevation="650" latitude="-18.983334" location="Paluma" longLatPrecision="1273" longitude="146.16667" municipality="Euparal" pageId="10" pageNumber="22" specimenCount="1" specimenCount-female="1" stateProvince="Queensland" typeStatus="holotype">
<specimenCount id="8DCDFD56FFD4315EFEBF20EF78B6FB2F" box="[382,397,1216,1240]" pageId="10" pageNumber="22" type="female">♀</specimenCount>
, on same slide as
<typeStatus id="4470887DFFD4315EFD9920EF7B83FB2F" box="[600,696,1216,1240]" pageId="10" pageNumber="22" type="holotype">holotype</typeStatus>
, same data
</materialsCitation>
.
</paragraph>
<paragraph id="9B7436DFFFD4315EFF2221207D1EFC0C" blockId="10.[227,764,1295,2023]" lastBlockId="10.[824,1361,227,1019]" pageId="10" pageNumber="22">
<emphasis id="A9BFEACDFFD4315EFF2221207B3DFAD0" box="[227,518,1295,1319]" italics="true" pageId="10" pageNumber="22">Other material examined</emphasis>
:
<materialsCitation id="2BA33C82FFD4315EFDE121207B05FA90" ID-GBIF-Occurrence="4122308354" collectingDate="1989-04-10" country="Australia" latitude="-12.666667" location="Northern Territory" longLatPrecision="1292" longitude="132.93333" pageId="10" pageNumber="22" specimenCount="1" stateProvince="Northern Territory">
<collectingCountry id="E3DC764FFFD4315EFDE121207BBEFAD0" box="[544,645,1295,1319]" name="Australia" pageId="10" pageNumber="22">Australia</collectingCountry>
,
<collectingRegion id="590FF83DFFD4315EFD5A21207878FAB0" country="Australia" name="Northern Territory" pageId="10" pageNumber="22">Northern Territory</collectingRegion>
. Kakadu N.P., Pe, Djalkmara Billabong,
<geoCoordinate id="FEFF5018FFD4315EFF2221607800FA90" box="[227,315,1359,1383]" degrees="12" direction="south" minutes="40" orientation="latitude" pageId="10" pageNumber="22" precision="925" value="-12.666667">12°40’S</geoCoordinate>
<geoCoordinate id="FEFF5018FFD4315EFE8D2160788EFA90" box="[332,437,1359,1383]" degrees="132" direction="east" minutes="56" orientation="longitude" pageId="10" pageNumber="22" precision="925" value="132.93333">132°56’E</geoCoordinate>
,
<date id="EF75101FFFD4315EFE0A21607B05FA90" box="[459,574,1359,1383]" pageId="10" pageNumber="22" value="1989-04-10">
<collectingDate id="FF31E9F7FFD4315EFE0A21607B05FA90" box="[459,574,1359,1383]" pageId="10" pageNumber="22" value="1989-04-10">10.iv.1989</collectingDate>
</date>
</materialsCitation>
; 9Pe, Rockhole Mine Ck.,
<geoCoordinate id="FEFF5018FFD4315EFE9721407896FA70" box="[342,429,1391,1415]" degrees="13" direction="south" minutes="30" orientation="latitude" pageId="10" pageNumber="22" precision="925" value="-13.5">13°30’S</geoCoordinate>
<geoCoordinate id="FEFF5018FFD4315EFE7521407B25FA70" box="[436,542,1391,1415]" degrees="132" direction="east" minutes="30" orientation="longitude" pageId="10" pageNumber="22" precision="925" value="132.5">132°30’E</geoCoordinate>
,
<date id="EF75101FFFD4315EFDE921407BB6FA70" box="[552,653,1391,1415]" pageId="10" pageNumber="22" value="1993-04-01">1.iv.1993</date>
,
<date id="EF75101FFFD4315EFD5821407BCCFA70" box="[665,759,1391,1415]" pageId="10" pageNumber="22" value="1993-05-08">8.v.1993</date>
, leg. Smith; same except P
<specimenCount id="8DCDFD56FFD4315EFDC721A07B21FA51" box="[518,538,1423,1446]" pageId="10" pageNumber="22" type="male">♂</specimenCount>
, P
<specimenCount id="8DCDFD56FFD4315EFDFD21A07B77FA51" box="[572,588,1423,1446]" pageId="10" pageNumber="22" type="female">♀</specimenCount>
, [on same slide as Pe,
<taxonomicName id="5CCB4D5CFFD4315EFEF6218078E4FA30" authorityName="Cranston &amp; Krosch &amp; Tang" authorityYear="2022" box="[311,479,1455,1479]" class="Insecta" family="Chironomidae" genus="Nilotanypus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="22" phylum="Arthropoda" rank="species" species="haplochelus">
<emphasis id="A9BFEACDFFD4315EFEF6218078E4FA30" box="[311,479,1455,1479]" italics="true" pageId="10" pageNumber="22">N. haplochelus</emphasis>
</taxonomicName>
]
<date id="EF75101FFFD4315EFE3721807B59FA30" box="[502,610,1455,1479]" pageId="10" pageNumber="22" value="1993-05-13">13.v.1993</date>
; Pe, Koolpin Ck.,
<geoCoordinate id="FEFF5018FFD4315EFEDC21E0784EFA10" box="[285,373,1487,1511]" degrees="13" direction="south" minutes="29" orientation="latitude" pageId="10" pageNumber="22" precision="925" value="-13.483334">13°29’S</geoCoordinate>
<geoCoordinate id="FEFF5018FFD4315EFE4321E078D0FA10" box="[386,491,1487,1511]" degrees="132" direction="east" minutes="35" orientation="longitude" pageId="10" pageNumber="22" precision="925" value="132.58333">132°35’E</geoCoordinate>
,
<date id="EF75101FFFD4315EFE3C21E07BAAFA10" box="[509,657,1487,1511]" pageId="10" pageNumber="22" value="1992-05-15" valueMax="1992-05-16" valueMin="1992-05-15">15–16.v.1992</date>
.
<materialsCitation id="2BA33C82FFD4315EFD6221E07BC3F9D0" ID-GBIF-Occurrence="4122308310" collectingDate="2016-10-02" collectingDateMax="2016-10-03" collectingDateMin="2016-10-02" collectorName="Krosch &amp; Cranston" country="Australia" latitude="-16.141111" location="Queensland" longLatPrecision="21" longitude="145.42696" pageId="10" pageNumber="22" specimenCount="1" stateProvince="Queensland">
<collectingRegion id="590FF83DFFD4315EFD6221E0782FF9F0" country="Australia" name="Queensland" pageId="10" pageNumber="22">Queensland</collectingRegion>
,
<collectorName id="363E5309FFD4315EFEE421C07886F9F1" box="[293,445,1519,1543]" pageId="10" pageNumber="22">Daintree N.P.</collectorName>
,
<collectorName id="363E5309FFD4315EFE0F21C078D0F9F1" box="[462,491,1519,1542]" pageId="10" pageNumber="22">Pe</collectorName>
,
<collectorName id="363E5309FFD4315EFE3C21C07B57F9F0" box="[509,620,1519,1543]" pageId="10" pageNumber="22">Noah Ck.</collectorName>
,
<geoCoordinate id="FEFF5018FFD4315EFDBC21C07BC7F9F0" box="[637,764,1519,1543]" degrees="16" direction="south" minutes="08" orientation="latitude" pageId="10" pageNumber="22" precision="15" seconds="28" value="-16.141111">16°08’28”S</geoCoordinate>
<geoCoordinate id="FEFF5018FFD4315EFF2222207849F9D0" box="[227,370,1551,1575]" degrees="145" direction="east" minutes="25" orientation="longitude" pageId="10" pageNumber="22" precision="15" seconds="37" value="145.42696">145°25’37”E</geoCoordinate>
,
<date id="EF75101FFFD4315EFEBA222078CEF9D0" box="[379,501,1551,1575]" pageId="10" pageNumber="22" value="2016-10-02" valueMax="2016-10-03" valueMin="2016-10-02">
<collectingDate id="FF31E9F7FFD4315EFEBA222078CEF9D0" box="[379,501,1551,1575]" pageId="10" pageNumber="22" value="2016-10-02" valueMax="2016-10-03" valueMin="2016-10-02">2–3.x.2016</collectingDate>
</date>
, leg. Krosch &amp; Cranston
</materialsCitation>
; 3Pe, Oliver Ck.,
<geoCoordinate id="FEFF5018FFD4315EFE52220078D1F9B0" box="[403,490,1583,1607]" degrees="16" direction="south" minutes="08" orientation="latitude" pageId="10" pageNumber="22" precision="925" value="-16.133333">16°08’S</geoCoordinate>
<geoCoordinate id="FEFF5018FFD4315EFE3322007B60F9B0" box="[498,603,1583,1607]" degrees="145" direction="east" minutes="26" orientation="longitude" pageId="10" pageNumber="22" precision="925" value="145.43333">145°26’E</geoCoordinate>
,
<date id="EF75101FFFD4315EFDA622007BC3F9B0" box="[615,760,1583,1607]" pageId="10" pageNumber="22" value="1998-09-09" valueMax="1998-09-10" valueMin="1998-09-09">9–10.ix.1998</date>
;
<materialsCitation id="2BA33C82FFD4315EFF2222607B9CF970" ID-GBIF-Occurrence="4122308325" collectingDate="2016-10-06" collectingDateMax="1997-09-09" collectingDateMin="1997-09-08" collectorName="Krosch &amp; Bryant &amp; Cranston" country="Australia" latitude="-16.253056" location="Mt Windsor" longLatPrecision="21" longitude="145.04001" pageId="10" pageNumber="22" specimenCount="1" stateProvince="Queensland">
<location id="9E146004FFD4315EFF222260784CF990" LSID="urn:lsid:plazi:treatment:136287C9FFD43159FF2227EE78B3F952:9E146004FFD4315EFF222260784CF990" box="[227,375,1615,1639]" country="Australia" latitude="-16.253056" longLatPrecision="21" longitude="145.04001" name="Mt Windsor" pageId="10" pageNumber="22" stateProvince="Queensland">Mt Windsor</location>
N.P.,
<geoCoordinate id="FEFF5018FFD4315EFE1F22607B60F990" box="[478,603,1615,1639]" degrees="16" direction="south" minutes="15" orientation="latitude" pageId="10" pageNumber="22" precision="15" seconds="11" value="-16.253056">16°15’11”S</geoCoordinate>
<geoCoordinate id="FEFF5018FFD4315EFDB422607BCCF990" box="[629,759,1615,1639]" degrees="145" direction="east" minutes="2" orientation="longitude" pageId="10" pageNumber="22" precision="15" seconds="24" value="145.04001">145°2’24”E</geoCoordinate>
,
<date id="EF75101FFFD4315EFF2222407878F970" box="[227,323,1647,1671]" pageId="10" pageNumber="22" value="2016-10-06">
<collectingDate id="FF31E9F7FFD4315EFF2222407878F970" box="[227,323,1647,1671]" pageId="10" pageNumber="22" value="2016-10-06">6.x.2016</collectingDate>
</date>
, leg.
<collectorName id="363E5309FFD4315EFEBA224078F3F970" box="[379,456,1647,1671]" pageId="10" pageNumber="22">Krosch</collectorName>
,
<collectorName id="363E5309FFD4315EFE1422407B25F970" box="[469,542,1647,1671]" pageId="10" pageNumber="22">Bryant</collectorName>
&amp;
<collectorName id="363E5309FFD4315EFD8122407B9FF970" box="[576,676,1647,1671]" pageId="10" pageNumber="22">Cranston</collectorName>
;
</materialsCitation>
<materialsCitation id="2BA33C82FFD4315EFD6F224079C2F930" ID-GBIF-Occurrence="4122308382" collectingDate="2016-10-06" collectingDateMax="1997-09-09" collectingDateMin="1997-09-08" collectorName="McKie" country="Australia" latitude="-16.253056" location="Mt Windsor" longLatPrecision="21" longitude="145.04001" pageId="10" pageNumber="22" specimenCount="1" stateProvince="Queensland">
<collectorName id="363E5309FFD4315EFD6F22407BF0F971" box="[686,715,1647,1670]" pageId="10" pageNumber="22">Pe</collectorName>
,
<location id="9E146004FFD4315EFD172240781FF950" LSID="urn:lsid:plazi:treatment:136287C9FFD43159FF2227EE78B3F952:9E146004FFD4315EFD172240781FF950" country="Australia" name="Mt. Lewis" pageId="10" pageNumber="22" stateProvince="Queensland">Mt. Lewis</location>
N.P.,
<collectorName id="363E5309FFD4315EFEA222A078CCF950" box="[355,503,1679,1703]" pageId="10" pageNumber="22">Windmill Ck.</collectorName>
,
<date id="EF75101FFFD4315EFDC322A07BB8F950" box="[514,643,1679,1703]" pageId="10" pageNumber="22" value="1997-09-08" valueMax="1997-09-09" valueMin="1997-09-08">
<collectingDate id="FF31E9F7FFD4315EFDC322A07BB8F950" box="[514,643,1679,1703]" pageId="10" pageNumber="22" value="1997-09-08" valueMax="1997-09-09" valueMin="1997-09-08">8–9.ix.1997</collectingDate>
</date>
, leg. McKie
</materialsCitation>
; Pe, nr Mareeba, Davies Ck.,
<geoCoordinate id="FEFF5018FFD4315EFDF022807BB3F930" box="[561,648,1711,1735]" degrees="17" direction="south" minutes="01" orientation="latitude" pageId="10" pageNumber="22" precision="925" value="-17.016666">17°01’S</geoCoordinate>
<geoCoordinate id="FEFF5018FFD4315EFD4F22807BCCF930" box="[654,759,1711,1735]" degrees="145" direction="east" minutes="35" orientation="longitude" pageId="10" pageNumber="22" precision="925" value="145.58333">145°35’E</geoCoordinate>
,
<date id="EF75101FFFD4315EFF2222E078B7F910" box="[227,396,1743,1767]" pageId="10" pageNumber="22" value="1997-08-27" valueMax="1997-08-28" valueMin="1997-08-27">27–28.viii.1997</date>
[same slide includes Pe,
<taxonomicName id="5CCB4D5CFFD4315EFD6922E07807F8F0" authorityName="Cranston &amp; Krosch &amp; Tang" authorityYear="2022" class="Insecta" family="Chironomidae" genus="Nilotanypus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="22" phylum="Arthropoda" rank="species" species="haplochelus">
<emphasis id="A9BFEACDFFD4315EFD6922E07807F8F0" italics="true" pageId="10" pageNumber="22">N. haplochelus</emphasis>
</taxonomicName>
];
<materialsCitation id="2BA33C82FFD4315EFE9422C07B99F8B0" ID-GBIF-Occurrence="4122308387" collectingDate="2012-08-29" collectionCode="MV" collectorName="Krosch &amp; Cranston" country="Australia" latitude="-17.45" longitude="145.48334" pageId="10" pageNumber="22" specimenCode="FNQ16, NIG15" specimenCount="1" stateProvince="Queensland">
L, Mt. Hypipamee N.P., Wondecla Ck. [=Nigger Ck.,]
<geoCoordinate id="FEFF5018FFD4315EFE48232078DAF8D0" box="[393,481,1807,1831]" degrees="17" direction="south" minutes="27" orientation="latitude" pageId="10" pageNumber="22" precision="925" value="-17.45">17°27’S</geoCoordinate>
<geoCoordinate id="FEFF5018FFD4315EFE2923207B6AF8D0" box="[488,593,1807,1831]" degrees="145" direction="east" minutes="29" orientation="longitude" pageId="10" pageNumber="22" precision="925" value="145.48334">145°29’E</geoCoordinate>
,
<date id="EF75101FFFD4315EFD9C23207BF3F8D0" box="[605,712,1807,1831]" pageId="10" pageNumber="22" value="2016-10-11">11.x.2016</date>
, leg.
<collectorName id="363E5309FFD4315EFF222300780BF8B0" box="[227,304,1839,1863]" pageId="10" pageNumber="22">Krosch</collectorName>
&amp;
<collectorName id="363E5309FFD4315EFE912300788FF8B0" box="[336,436,1839,1863]" pageId="10" pageNumber="22">Cranston</collectorName>
; (
<collectionCode id="FDDAAE1AFFD4315EFE04230078C8F8B0" box="[453,499,1839,1863]" country="0" name="University of Montana Museum" pageId="10" pageNumber="22">MV</collectionCode>
)
<specimenCode id="CB6D9EA4FFD4315EFDC023007BA5F8B0" box="[513,670,1839,1863]" collectionCode="MV" country="0" name="University of Montana Museum" pageId="10" pageNumber="22">FNQ16NIG15</specimenCode>
;
</materialsCitation>
<materialsCitation id="2BA33C82FFD4315EFD6923007B77F890" ID-GBIF-Occurrence="4122308396" collectingDate="2012-08-29" collectionCode="MV" collectorName="Krosch &amp; Cranston" country="Australia" pageId="10" pageNumber="22" specimenCode="FNQ16, NIG15" specimenCount="1" stateProvince="Queensland">
L, same except
<date id="EF75101FFFD4315EFEEF2360788AF890" box="[302,433,1871,1895]" pageId="10" pageNumber="22" value="2012-08-29">
<collectingDate id="FF31E9F7FFD4315EFEEF2360788AF890" box="[302,433,1871,1895]" pageId="10" pageNumber="22" value="2012-08-29">29.viii.2012</collectingDate>
</date>
, leg. Cranston
</materialsCitation>
;
<materialsCitation id="2BA33C82FFD4315EFD942360785FF850" ID-GBIF-Occurrence="4122308376" collectingDate="1997-04-08" collectingDateMax="1997-04-09" collectingDateMin="1997-04-08" collectorName="McKie" country="Australia" elevation="650" latitude="-17.583334" location="Pe" longLatPrecision="1278" longitude="145.7" pageId="10" pageNumber="22" specimenCount="1" stateProvince="Queensland">
Pe, Tully Gorge N.P., Pixies Ck.,
<date id="EF75101FFFD4315EFE5E23407B1AF870" box="[415,545,1903,1927]" pageId="10" pageNumber="22" value="1997-09-02" valueMax="1997-09-03" valueMin="1997-09-02">2–3.ix.1997</date>
,
<geoCoordinate id="FEFF5018FFD4315EFDEF23407BBEF870" box="[558,645,1903,1927]" degrees="17" direction="south" minutes="47" orientation="latitude" pageId="10" pageNumber="22" precision="925" value="-17.783333">17°47’S</geoCoordinate>
<geoCoordinate id="FEFF5018FFD4315EFD4F23407BCCF870" box="[654,759,1903,1927]" degrees="145" direction="east" minutes="41" orientation="longitude" pageId="10" pageNumber="22" precision="925" value="145.68333">145°41’E</geoCoordinate>
, leg.
<collectorName id="363E5309FFD4315EFED323A0785BF850" box="[274,352,1935,1959]" pageId="10" pageNumber="22">McKie</collectorName>
;
</materialsCitation>
<materialsCitation id="2BA33C82FFD4315EFEAD23A07BF1F830" ID-GBIF-Occurrence="4122308304" collectingDate="1997-04-08" collectingDateMax="1997-04-09" collectingDateMin="1997-04-08" country="Australia" elevation="650" latitude="-17.583334" location="Pe" longLatPrecision="1278" longitude="145.7" pageId="10" pageNumber="22" specimenCount="1" stateProvince="Queensland">
<location id="9E146004FFD4315EFEAD23A078B2F851" LSID="urn:lsid:plazi:treatment:136287C9FFD43159FF2227EE78B3F952:9E146004FFD4315EFEAD23A078B2F851" box="[364,393,1935,1958]" country="Australia" latitude="-17.583334" longLatPrecision="1278" longitude="145.7" name="Pe" pageId="10" pageNumber="22" stateProvince="Queensland">Pe</location>
,
<collectorName id="363E5309FFD4315EFE5723A07B7EF851" box="[406,581,1935,1959]" pageId="10" pageNumber="22">Palmerston N.P.</collectorName>
,
<collectorName id="363E5309FFD4315EFD9323A07BC3F850" box="[594,760,1935,1959]" pageId="10" pageNumber="22">Learmouth Ck.</collectorName>
,
<elevation id="10E6D1ECFFD4315EFF222380785FF830" box="[227,356,1967,1991]" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="10" pageNumber="22" unit="m" value="650.0">
<quantity id="5C339B3AFFD4315EFF2223807812F831" box="[227,297,1967,1991]" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="10" pageNumber="22" unit="m" value="650.0">650 m</quantity>
a.s.l.
</elevation>
,
<geoCoordinate id="FEFF5018FFD4315EFEB2238078F1F830" box="[371,458,1967,1991]" degrees="17" direction="south" minutes="35" orientation="latitude" pageId="10" pageNumber="22" precision="925" value="-17.583334">17°35’S</geoCoordinate>
<geoCoordinate id="FEFF5018FFD4315EFE1523807B06F830" box="[468,573,1967,1991]" degrees="145" direction="east" minutes="42" orientation="longitude" pageId="10" pageNumber="22" precision="925" value="145.7">145°42’E</geoCoordinate>
,
<date id="EF75101FFFD4315EFD8B23807BF1F830" box="[586,714,1967,1991]" pageId="10" pageNumber="22" value="1997-04-08" valueMax="1997-04-09" valueMin="1997-04-08">
<collectingDate id="FF31E9F7FFD4315EFD8B23807BF1F830" box="[586,714,1967,1991]" pageId="10" pageNumber="22" value="1997-04-08" valueMax="1997-04-09" valueMin="1997-04-08">8–9.iv.1997</collectingDate>
</date>
</materialsCitation>
,
<materialsCitation id="2BA33C82FFD4315EFD1623807C6AFEEC" ID-GBIF-Occurrence="4122308378" collectingDate="1997-06-09" collectionCode="MV" collectorName="Krosch &amp; Bryant" country="Australia" latitude="-18.185276" longLatPrecision="21" longitude="145.76666" pageId="10" pageNumber="22" specimenCode="FNQ16, RAV1.4, 1.5" specimenCount="1" stateProvince="Queensland">
3L, Koombooloomba N.P., Koombooloomba Ck., nr dam,
<geoCoordinate id="FEFF5018FFD4315EFCBD24CC7AC1FF0C" box="[892,1018,227,251]" degrees="17" direction="south" minutes="50" orientation="latitude" pageId="10" pageNumber="22" precision="15" seconds="16" value="-17.837778">17°50’16”S</geoCoordinate>
<geoCoordinate id="FEFF5018FFD4315EFBC824CC7DAEFF0C" box="[1033,1173,227,251]" degrees="145" direction="east" minutes="35" orientation="longitude" pageId="10" pageNumber="22" precision="15" seconds="16" value="-145.35">145°35’16’E</geoCoordinate>
,
<date id="EF75101FFFD4315EFB6824CC7C2DFF0C" box="[1193,1302,227,251]" pageId="10" pageNumber="22" value="2018-10-12">12.x.2018</date>
, leg.
<collectorName id="363E5309FFD4315EFCF9252C7ABEFEEC" box="[824,901,259,283]" pageId="10" pageNumber="22">Krosch</collectorName>
&amp;
<collectorName id="363E5309FFD4315EFC7D252B7D33FEEB" box="[956,1032,260,284]" pageId="10" pageNumber="22">Bryant</collectorName>
; (
<collectionCode id="FDDAAE1AFFD4315EFBE5252C7D69FEEC" box="[1060,1106,259,283]" country="0" name="University of Montana Museum" pageId="10" pageNumber="22">MV</collectionCode>
)
<specimenCode id="CB6D9EA4FFD4315EFBAB252C7C28FEEC" box="[1130,1299,259,283]" collectionCode="MV" country="0" name="University of Montana Museum" pageId="10" pageNumber="22">FNQ16RAV1.4</specimenCode>
,
<specimenCode id="CB6D9EA4FFD4315EFAE9252C7C77FEEC" box="[1320,1356,259,283]" collectionCode="MV" country="0" name="University of Montana Museum" pageId="10" pageNumber="22">1.5</specimenCode>
;
</materialsCitation>
<materialsCitation id="2BA33C82FFD4315EFCF9250C7D7FFEAC" ID-GBIF-Occurrence="4122308337" collectingDate="1997-06-09" collectionCode="MV" collectorName="McKie" country="Australia" latitude="-18.185276" longLatPrecision="21" longitude="145.76666" pageId="10" pageNumber="22" specimenCode="FNQ16, RAV1.4, 1.5" specimenCount="1" stateProvince="Queensland">
2
<collectorName id="363E5309FFD4315EFC84250C7A59FECC" box="[837,866,291,315]" pageId="10" pageNumber="22">Pe</collectorName>
,
<collectorName id="363E5309FFD4315EFCB4250C7D27FECC" box="[885,1052,291,315]" pageId="10" pageNumber="22">Yuccabine Ck.</collectorName>
,
<geoCoordinate id="FEFF5018FFD4315EFBF1250C7D96FECC" box="[1072,1197,291,315]" degrees="18" direction="south" minutes="11" orientation="latitude" pageId="10" pageNumber="22" precision="15" seconds="07" value="-18.185276">18°11’07”S</geoCoordinate>
<geoCoordinate id="FEFF5018FFD4315EFB7C250C7C77FECC" box="[1213,1356,291,315]" degrees="145" direction="east" minutes="46" orientation="longitude" pageId="10" pageNumber="22" precision="15" seconds="00" value="145.76666">145°46’00”E</geoCoordinate>
,
<date id="EF75101FFFD4315EFCF9256C7AA4FEAC" box="[824,927,323,347]" pageId="10" pageNumber="22" value="1997-06-09">
<collectingDate id="FF31E9F7FFD4315EFCF9256C7AA4FEAC" box="[824,927,323,347]" pageId="10" pageNumber="22" value="1997-06-09">9.vi.1997</collectingDate>
</date>
, leg. McKie
</materialsCitation>
; 2Pe, Yuccabine Ck.,
<date id="EF75101FFFD4315EFCF9254C7A96FE8C" box="[824,941,355,379]" pageId="10" pageNumber="22" value="1997-06-10">10.vi.1997</date>
, leg. McKie; 2P
<specimenCount id="8DCDFD56FFD4315EFB9D254C7D4BFE8C" box="[1116,1136,355,379]" pageId="10" pageNumber="22" type="male">♂</specimenCount>
, 2P
<specimenCount id="8DCDFD56FFD4315EFB5D254C7D90FE8C" box="[1180,1195,355,379]" pageId="10" pageNumber="22" type="female">♀</specimenCount>
, Paluma, Birthday Ck.,
<geoCoordinate id="FEFF5018FFD4315EFC5F25AC7ACDFE6C" box="[926,1014,387,411]" degrees="18" direction="south" minutes="59" orientation="latitude" pageId="10" pageNumber="22" precision="925" value="-18.983334">18°59’S</geoCoordinate>
<geoCoordinate id="FEFF5018FFD4315EFC3E25AC7D53FE6C" box="[1023,1128,387,411]" degrees="146" direction="east" minutes="10" orientation="longitude" pageId="10" pageNumber="22" precision="925" value="146.16667">146°10’E</geoCoordinate>
,
<quantity id="5C339B3AFFD4315EFBB725AC7D87FE6C" box="[1142,1212,387,411]" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="10" pageNumber="22" unit="m" value="650.0">650 m</quantity>
a.s.l.,
<date id="EF75101FFFD4315EFAC625AC7AB6FE4C" pageId="10" pageNumber="22" value="1989-03-25" valueMax="1989-03-26" valueMin="1989-03-25">25–26. iii.1989</date>
; 3L, same except
<date id="EF75101FFFD4315EFB88258C7D92FE4C" box="[1097,1193,419,443]" pageId="10" pageNumber="22" value="2009-10-01">1.x.2009</date>
, leg. Krosch; L, same except
<date id="EF75101FFFD4315EFC0125EC7D78FE2C" box="[960,1091,451,475]" pageId="10" pageNumber="22" value="2005-08-31">31.viii.2005</date>
, leg. Cranston; 2L, Camp Ck.,
<geoCoordinate id="FEFF5018FFD4315EFCAA25CC7AF9FE0C" box="[875,962,483,507]" degrees="18" direction="south" minutes="58" orientation="latitude" pageId="10" pageNumber="22" precision="925" value="-18.966667">18°58’S</geoCoordinate>
<geoCoordinate id="FEFF5018FFD4315EFC0825CC7D09FE0C" box="[969,1074,483,507]" degrees="146" direction="east" minutes="09" orientation="longitude" pageId="10" pageNumber="22" precision="925" value="146.15">146°09’E</geoCoordinate>
,
<date id="EF75101FFFD4315EFBFD25CC7D8AFE0C" box="[1084,1201,483,507]" pageId="10" pageNumber="22" value="2008-09-21">21.ix.2008</date>
, leg. Krosch &amp; Bryant; L, Cooloola N.P., Franki’s Gulch,
<geoCoordinate id="FEFF5018FFD4315EFB38262C7C6BFDEC" box="[1273,1360,515,539]" degrees="26" direction="south" minutes="03" orientation="latitude" pageId="10" pageNumber="22" precision="925" value="-26.05">26°03’S</geoCoordinate>
<geoCoordinate id="FEFF5018FFD4315EFCF9260C7A9AFDCC" box="[824,929,547,571]" degrees="153" direction="east" minutes="04" orientation="longitude" pageId="10" pageNumber="22" precision="925" value="153.06667">153°04’E</geoCoordinate>
,
<date id="EF75101FFFD4315EFC73260C7D23FDCC" box="[946,1048,547,571]" pageId="10" pageNumber="22" value="1996-04-06">6.iv.1996</date>
;
<materialsCitation id="2BA33C82FFD4315EFBE9260C7ABBFD8C" ID-GBIF-Occurrence="4122308344" collectingDate="1989-09-26" collectorName="Cedar Ck." country="Australia" latitude="-27.9" location="Tamborine Mt." longLatPrecision="1234" longitude="153.18333" pageId="10" pageNumber="22" specimenCount="1" stateProvince="Queensland">
3L,
<location id="9E146004FFD4315EFB98260C7C3AFDCC" LSID="urn:lsid:plazi:treatment:136287C9FFD43159FF2227EE78B3F952:9E146004FFD4315EFB98260C7C3AFDCC" box="[1113,1281,547,571]" country="Australia" latitude="-27.9" longLatPrecision="1234" longitude="153.18333" name="Tamborine Mt." pageId="10" pageNumber="22" stateProvince="Queensland">Tamborine Mt.</location>
,
<collectorName id="363E5309FFD4315EFAD0260C7A5BFDAC" pageId="10" pageNumber="22">Cedar Ck.</collectorName>
,
<geoCoordinate id="FEFF5018FFD4315EFCB0266C7AF3FDAC" box="[881,968,579,603]" degrees="27" direction="south" minutes="54" orientation="latitude" pageId="10" pageNumber="22" precision="925" value="-27.9">27°54’S</geoCoordinate>
<geoCoordinate id="FEFF5018FFD4315EFC14266C7D07FDAC" box="[981,1084,579,603]" degrees="153" direction="east" minutes="11" orientation="longitude" pageId="10" pageNumber="22" precision="925" value="153.18333">153°11’E</geoCoordinate>
,
<date id="EF75101FFFD4315EFB8F266C7DFFFDAC" box="[1102,1220,579,603]" pageId="10" pageNumber="22" value="1989-09-26">
<collectingDate id="FF31E9F7FFD4315EFB8F266C7DFFFDAC" box="[1102,1220,579,603]" pageId="10" pageNumber="22" value="1989-09-26">26.ix.1989</collectingDate>
</date>
.
<collectingRegion id="590FF83DFFD4315EFB12266B7A47FD8C" country="Australia" name="New South Wales" pageId="10" pageNumber="22">New South Wales</collectingRegion>
.
</materialsCitation>
<materialsCitation id="2BA33C82FFD4315EFC4C264C7D1FFD6C" ID-GBIF-Occurrence="4122308346" collectingDate="1989-09-26" collectorName="Cedar Ck." country="Australia" latitude="-27.9" location="Tamborine Mt." longLatPrecision="1234" longitude="153.18333" pageId="10" pageNumber="22" specimenCount="1" stateProvince="Queensland">
2L,
<collectorName id="363E5309FFD4315EFC7E264C7D12FD8C" box="[959,1065,611,635]" pageId="10" pageNumber="22">Bungonia</collectorName>
,
<location id="9E146004FFD4315EFBFA264C7DDCFD8C" LSID="urn:lsid:plazi:treatment:136287C9FFD43159FF2227EE78B3F952:9E146004FFD4315EFBFA264C7DDCFD8C" box="[1083,1255,611,635]" country="Australia" latitude="-34.783333" longLatPrecision="1197" longitude="150.0" name="Bungonia Falls" pageId="10" pageNumber="22" stateProvince="New South Wales">Bungonia Falls</location>
,
<geoCoordinate id="FEFF5018FFD4315EFB38264C7C6BFD8C" box="[1273,1360,611,635]" degrees="34" direction="south" minutes="47" orientation="latitude" pageId="10" pageNumber="22" precision="925" value="-34.783333">34°47’S</geoCoordinate>
<geoCoordinate id="FEFF5018FFD4315EFCF926AC7A9BFD6C" box="[824,928,643,667]" degrees="150" direction="east" minutes="00" orientation="longitude" pageId="10" pageNumber="22" precision="925" value="150.0">150°00’E</geoCoordinate>
,
<date id="EF75101FFFD4315EFC7126AC7D1FFD6C" box="[944,1060,643,667]" pageId="10" pageNumber="22" value="1988-11-11">
<collectingDate id="FF31E9F7FFD4315EFC7126AC7D1FFD6C" box="[944,1060,643,667]" pageId="10" pageNumber="22" value="1988-11-11">11.xi.1988</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="2BA33C82FFD4315EFBF226AC7D9FFD4C" ID-GBIF-Occurrence="4122308366" collectingDate="2001-01-04" collectionCode="R" country="Australia" latitude="-35.398335" location="Pe" longLatPrecision="19" longitude="150.02" pageId="10" pageNumber="22" specimenCount="1" stateProvince="New South Wales">
2
<collectorName id="363E5309FFD4315EFB8026AC7D65FD6C" box="[1089,1118,643,667]" pageId="10" pageNumber="22">Pe</collectorName>
, Currowan S.F., Cabbage Tree Ck.,
<geoCoordinate id="FEFF5018FFD4315EFC18268C7D0AFD4C" box="[985,1073,675,699]" degrees="35" direction="south" minutes="34" orientation="latitude" pageId="10" pageNumber="22" precision="925" value="-35.566666">35°34’S</geoCoordinate>
<geoCoordinate id="FEFF5018FFD4315EFBF7268C7D9BFD4C" box="[1078,1184,675,699]" degrees="150" direction="east" minutes="02" orientation="longitude" pageId="10" pageNumber="22" precision="925" value="150.03334">150°02’E</geoCoordinate>
;
</materialsCitation>
<materialsCitation id="2BA33C82FFD4315EFB6B268C7C77FD0C" ID-GBIF-Occurrence="4122308391" collectingDate="2001-01-04" collectionCode="R" country="Australia" latitude="-35.398335" location="Pe" longLatPrecision="19" longitude="148.7475" pageId="10" pageNumber="22" specimenCount="1" stateProvince="New South Wales">
2
<location id="9E146004FFD4315EFB76268C7DE8FD4C" LSID="urn:lsid:plazi:treatment:136287C9FFD43159FF2227EE78B3F952:9E146004FFD4315EFB76268C7DE8FD4C" box="[1207,1235,675,699]" country="Australia" latitude="-35.398335" longLatPrecision="19" longitude="148.7475" name="Pe" pageId="10" pageNumber="22" stateProvince="New South Wales">Pe</location>
[same slide includes 1Pe
<taxonomicName id="5CCB4D5CFFD4315EFC0926EB7D52FD2C" authorityName="Cranston &amp; Krosch &amp; Tang" authorityYear="2022" box="[968,1129,707,731]" class="Insecta" family="Chironomidae" genus="Nilotanypus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="22" phylum="Arthropoda" rank="species" species="haplochelus">
<emphasis id="A9BFEACDFFD4315EFC0926EB7D52FD2C" box="[968,1129,707,731]" italics="true" pageId="10" pageNumber="22">N. haplochelus</emphasis>
</taxonomicName>
]
<collectorName id="363E5309FFD4315EFBBB26EC7DC2FD2C" box="[1146,1273,707,731]" pageId="10" pageNumber="22">Brindabella</collectorName>
,
<collectorName id="363E5309FFD4315EFAC526EC7AB0FD0C" pageId="10" pageNumber="22">Goodradigbee</collectorName>
<collectionCode id="FDDAAE1AFFD4315EFC5926CB7A95FD0C" box="[920,942,740,763]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="10" pageNumber="22">R</collectionCode>
.,
<geoCoordinate id="FEFF5018FFD4315EFC0426CC7D78FD0C" box="[965,1091,739,763]" degrees="35" direction="south" minutes="23" orientation="latitude" pageId="10" pageNumber="22" precision="15" seconds="54" value="-35.398335">35°23’54”S</geoCoordinate>
<geoCoordinate id="FEFF5018FFD4315EFB9026CC7DDBFD0C" box="[1105,1248,739,763]" degrees="148" direction="east" minutes="44" orientation="longitude" pageId="10" pageNumber="22" precision="15" seconds="51" value="148.7475">148°44’51”E</geoCoordinate>
,
<date id="EF75101FFFD4315EFB3326CC7C77FD0C" box="[1266,1356,739,763]" pageId="10" pageNumber="22" value="2001-01-04">
<collectingDate id="FF31E9F7FFD4315EFB3326CC7C77FD0C" box="[1266,1356,739,763]" pageId="10" pageNumber="22" value="2001-01-04">4.i.2001</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="2BA33C82FFD4315EFCF9272B7D21FCCC" ID-GBIF-Occurrence="4122308323" collectingDate="1993-03-06" collectionCode="R" country="Australia" latitude="-35.583332" longLatPrecision="1192" longitude="149.46666" pageId="10" pageNumber="22" specimenCount="1" stateProvince="New South Wales">
7
<collectorName id="363E5309FFD4315EFC8F272B7A50FCEC" box="[846,875,772,795]" pageId="10" pageNumber="22">Pe</collectorName>
, above
<collectorName id="363E5309FFD4315EFC01272C7D69FCEC" box="[960,1106,771,795]" pageId="10" pageNumber="22">Captains Flat</collectorName>
,
<collectorName id="363E5309FFD4315EFB9E272C7DF3FCEC" box="[1119,1224,771,795]" pageId="10" pageNumber="22">Molonglo</collectorName>
<collectionCode id="FDDAAE1AFFD4315EFB10272B7DDCFCEC" box="[1233,1255,772,795]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="10" pageNumber="22">R</collectionCode>
.,
<geoCoordinate id="FEFF5018FFD4315EFB38272C7C6AFCEC" box="[1273,1361,771,795]" degrees="35" direction="south" minutes="35" orientation="latitude" pageId="10" pageNumber="22" precision="925" value="-35.583332">35°35’S</geoCoordinate>
<geoCoordinate id="FEFF5018FFD4315EFCF9270C7A9AFCCC" box="[824,929,803,827]" degrees="149" direction="east" minutes="28" orientation="longitude" pageId="10" pageNumber="22" precision="925" value="149.46666">149°28’E</geoCoordinate>
,
<date id="EF75101FFFD4315EFC71270C7D21FCCC" box="[944,1050,803,827]" pageId="10" pageNumber="22" value="1993-03-06">
<collectingDate id="FF31E9F7FFD4315EFC71270C7D21FCCC" box="[944,1050,803,827]" pageId="10" pageNumber="22" value="1993-03-06">6.iii.1993</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="2BA33C82FFD4315EFBE8270B7C1AFCAC" ID-GBIF-Occurrence="4122308330" collectingDate="1993-03-17" collectionCode="R" collectorName="Pe &amp; Mongarlowe" country="Australia" latitude="-35.4" longLatPrecision="1193" longitude="149.95" pageId="10" pageNumber="22" specimenCount="1" stateProvince="New South Wales">
<collectorName id="363E5309FFD4315EFBE8270B7D7DFCCC" box="[1065,1094,804,827]" pageId="10" pageNumber="22">Pe</collectorName>
, nr.
<collectorName id="363E5309FFD4315EFBBA270C7C3DFCCC" box="[1147,1286,803,827]" pageId="10" pageNumber="22">Mongarlowe</collectorName>
,
<collectorName id="363E5309FFD4315EFAD4270B7AB5FCAC" pageId="10" pageNumber="22">Mongarlowe</collectorName>
<collectionCode id="FDDAAE1AFFD4315EFC5D276B7A89FCAC" box="[924,946,836,859]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="10" pageNumber="22">R</collectionCode>
.,
<geoCoordinate id="FEFF5018FFD4315EFC08276C7D1AFCAC" box="[969,1057,835,859]" degrees="35" direction="south" minutes="24" orientation="latitude" pageId="10" pageNumber="22" precision="925" value="-35.4">35°24’S</geoCoordinate>
<geoCoordinate id="FEFF5018FFD4315EFBEE276C7DACFCAC" box="[1071,1175,835,859]" degrees="149" direction="east" minutes="57" orientation="longitude" pageId="10" pageNumber="22" precision="925" value="149.95">149°57’E</geoCoordinate>
,
<date id="EF75101FFFD4315EFB6B276C7C1AFCAC" box="[1194,1313,835,859]" pageId="10" pageNumber="22" value="1993-03-17">
<collectingDate id="FF31E9F7FFD4315EFB6B276C7C1AFCAC" box="[1194,1313,835,859]" pageId="10" pageNumber="22" value="1993-03-17">17.iii.1993</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="2BA33C82FFD4315EFAF2276B7DB2FC6C" ID-GBIF-Occurrence="4122308319" collectingDate="1991-07-01" collectionCode="R" collectorName="Pe &amp; Buckland" country="Australia" latitude="-36.8" location="Victoria" longLatPrecision="1185" longitude="146.85" pageId="10" pageNumber="22" specimenCount="1" stateProvince="Victoria">
L., Kosciuszko N.P., Leather Barrel Ck.,
<geoCoordinate id="FEFF5018FFD4315EFB38274C7C6AFC8C" box="[1273,1361,867,891]" degrees="36" direction="south" minutes="31" orientation="latitude" pageId="10" pageNumber="22" precision="925" value="-36.516666">36°31’S</geoCoordinate>
<geoCoordinate id="FEFF5018FFD4315EFCF927AC7A9BFC6C" box="[824,928,899,923]" degrees="148" direction="east" minutes="11" orientation="longitude" pageId="10" pageNumber="22" precision="925" value="148.18333">148°11’E</geoCoordinate>
,
<date id="EF75101FFFD4315EFC6E27AC7D24FC6C" box="[943,1055,899,923]" pageId="10" pageNumber="22" value="2010-12-04">4.xii.2010</date>
.
<collectingRegion id="590FF83DFFD4315EFBEC27AC7DBDFC6C" box="[1069,1158,899,923]" country="Australia" name="Victoria" pageId="10" pageNumber="22">Victoria</collectingRegion>
,
</materialsCitation>
<materialsCitation id="2BA33C82FFD4315EFB5527AB7C27FC4C" ID-GBIF-Occurrence="4122308370" collectingDate="1991-07-01" collectionCode="R" collectorName="Pe &amp; Buckland" country="Australia" latitude="-36.8" location="Victoria" longLatPrecision="1185" longitude="146.85" pageId="10" pageNumber="22" specimenCount="1" stateProvince="Victoria">
<collectorName id="363E5309FFD4315EFB5527AB7D8AFC6C" box="[1172,1201,900,923]" pageId="10" pageNumber="22">Pe</collectorName>
,
<collectorName id="363E5309FFD4315EFB0127AC7C1DFC6C" box="[1216,1318,899,923]" pageId="10" pageNumber="22">Buckland</collectorName>
<collectionCode id="FDDAAE1AFFD4315EFAF327AB7C73FC6C" box="[1330,1352,900,923]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="10" pageNumber="22">R</collectionCode>
.,
<geoCoordinate id="FEFF5018FFD4315EFCF9278C7AABFC4C" box="[824,912,931,955]" degrees="36" direction="south" minutes="48" orientation="latitude" pageId="10" pageNumber="22" precision="925" value="-36.8">36°48’S</geoCoordinate>
<geoCoordinate id="FEFF5018FFD4315EFC60278C7D32FC4C" box="[929,1033,931,955]" degrees="146" direction="east" minutes="51" orientation="longitude" pageId="10" pageNumber="22" precision="925" value="146.85">146°51’E</geoCoordinate>
,
<date id="EF75101FFFD4315EFBDE278C7DB6FC4C" box="[1055,1165,931,955]" pageId="10" pageNumber="22" value="1991-07-01">
<collectingDate id="FF31E9F7FFD4315EFBDE278C7DB6FC4C" box="[1055,1165,931,955]" pageId="10" pageNumber="22" value="1991-07-01">1.vii.1991</collectingDate>
</date>
, leg.
<collectorName id="363E5309FFD4315EFB1B278C7C2CFC4C" box="[1242,1303,931,955]" pageId="10" pageNumber="22">Cook</collectorName>
;
</materialsCitation>
<materialsCitation id="2BA33C82FFD4315EFAED278C7D1EFC0C" ID-GBIF-Occurrence="4122308379" collectingDate="1991-07-01" collectionCode="R" collectorName="Pe &amp; Buckland" country="Australia" latitude="-36.8" location="Victoria" longLatPrecision="1185" longitude="146.85" pageId="10" pageNumber="22" specimenCount="1" stateProvince="Victoria">
2L,
<collectorName id="363E5309FFD4315EFCF927EC7AB9FC2C" box="[824,898,963,987]" pageId="10" pageNumber="22">Tambo</collectorName>
<collectionCode id="FDDAAE1AFFD4315EFC5027EB7A9CFC2C" box="[913,935,964,987]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="10" pageNumber="22">R</collectionCode>
., south branch,
<date id="EF75101FFFD4315EFBA827EC7DDFFC2C" box="[1129,1252,963,987]" pageId="10" pageNumber="22" value="1990-12-12">
<collectingDate id="FF31E9F7FFD4315EFBA827EC7DDFFC2C" box="[1129,1252,963,987]" pageId="10" pageNumber="22" value="1990-12-12">12.xii.1990</collectingDate>
</date>
,
<geoCoordinate id="FEFF5018FFD4315EFB3827EC7C6AFC2C" box="[1273,1361,963,987]" degrees="36" direction="south" minutes="59" orientation="latitude" pageId="10" pageNumber="22" precision="925" value="-36.983334">36°59’S</geoCoordinate>
<geoCoordinate id="FEFF5018FFD4315EFCF927CC7A9BFC0C" box="[824,928,995,1019]" degrees="147" direction="east" minutes="51" orientation="longitude" pageId="10" pageNumber="22" precision="925" value="147.85">147°51’E</geoCoordinate>
, leg. Hortle.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="D3D16554FFD4315EFCF9203D7AF6FB9D" pageId="10" pageNumber="22" type="etymology">
<paragraph id="9B7436DFFFD4315EFCF9203D7AF6FB9D" blockId="10.[824,1361,1042,1130]" pageId="10" pageNumber="22">
<emphasis id="A9BFEACDFFD4315EFCF9203D7A88FBDD" bold="true" box="[824,947,1042,1066]" pageId="10" pageNumber="22">Etymology</emphasis>
: From Greek,
<emphasis id="A9BFEACDFFD4315EFB92203C7DB6FBDD" box="[1107,1165,1043,1066]" italics="true" pageId="10" pageNumber="22">cteno</emphasis>
= comb,
<emphasis id="A9BFEACDFFD4315EFB37203D7C01FBDD" box="[1270,1338,1042,1066]" italics="true" pageId="10" pageNumber="22">chelus</emphasis>
= claw, recognising the comb-like larval posterior parapod claw.
</paragraph>
</subSubSection>
<subSubSection id="D3D16554FFD4315FFCF920AE7B0FFB3C" lastPageId="11" lastPageNumber="23" pageId="10" pageNumber="22" type="diagnosis">
<paragraph id="9B7436DFFFD4315EFCF920AE7D14FB6E" blockId="10.[824,1071,1153,1177]" box="[824,1071,1153,1177]" pageId="10" pageNumber="22">
<emphasis id="A9BFEACDFFD4315EFCF920AE7D14FB6E" bold="true" box="[824,1071,1153,1177]" pageId="10" pageNumber="22">Diagnostic characters</emphasis>
</paragraph>
<paragraph id="9B7436DFFFD4315EFCF9209F7AE9F97E" blockId="10.[824,1361,1200,1673]" pageId="10" pageNumber="22">
The two new Australian species described here conform in all stages to
<taxonomicName id="5CCB4D5CFFD4315EFB9D20FF7DE6FB1F" box="[1116,1245,1232,1256]" class="Insecta" family="Chironomidae" genus="Nilotanypus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis id="A9BFEACDFFD4315EFB9D20FF7DE6FB1F" box="[1116,1245,1232,1256]" italics="true" pageId="10" pageNumber="22">Nilotanypus</emphasis>
</taxonomicName>
, with additional features noted above in an expanded generic diagnosis. Male adults may be separable by the tarsal pseudospurs:
<taxonomicName id="5CCB4D5CFFD4315EFBFA211E7DE5FABF" authority="Cranston &amp; Krosch &amp; Tang, 2022" authorityName="Cranston &amp; Krosch &amp; Tang" authorityYear="2022" box="[1083,1246,1328,1352]" class="Insecta" family="Chironomidae" genus="Nilotanypus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="22" phylum="Arthropoda" rank="species" species="haplochelus" status="sp. nov.">
<emphasis id="A9BFEACDFFD4315EFBFA211E7DE5FABF" box="[1083,1246,1328,1352]" italics="true" pageId="10" pageNumber="22">N. haplochelus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B28C57B6FFD4315EFB26211E7C1FFABF" box="[1255,1316,1329,1352]" pageId="10" pageNumber="22" rank="species">sp. n.</taxonomicNameLabel>
has a subapical pseudospur on fore tarsomere on the foreleg (P
<subScript id="074F349AFFD4315EFC6421AE7A96FA78" attach="left" box="[933,941,1409,1423]" fontSize="6" pageId="10" pageNumber="22">1</subScript>
), whereas
<taxonomicName id="5CCB4D5CFFD4315EFBE9215E7DF2FA7F" authority="Cranston &amp; Krosch &amp; Tang, 2022" authorityName="Cranston &amp; Krosch &amp; Tang" authorityYear="2022" box="[1064,1225,1392,1416]" class="Insecta" family="Chironomidae" genus="Nilotanypus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="22" phylum="Arthropoda" rank="species" species="ctenochelus" status="sp. nov.">
<emphasis id="A9BFEACDFFD4315EFBE9215E7DF2FA7F" box="[1064,1225,1392,1416]" italics="true" pageId="10" pageNumber="22">N. ctenochelus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B28C57B6FFD4315EFB12215E7C30FA7F" box="[1235,1291,1393,1416]" pageId="10" pageNumber="22" rank="species">sp. n</taxonomicNameLabel>
, lacks pseudospurs on Ta 1 of P 1. Midleg pseudospurs may distinguish but confirmation based on teneral specimens is unsafe. The gonostylus of the male genitalia can separate:
<taxonomicName id="5CCB4D5CFFD4315EFBF921DE7DE1F9FF" authority="Cranston &amp; Krosch &amp; Tang, 2022" authorityName="Cranston &amp; Krosch &amp; Tang" authorityYear="2022" box="[1080,1242,1520,1544]" class="Insecta" family="Chironomidae" genus="Nilotanypus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="22" phylum="Arthropoda" rank="species" species="ctenochelus" status="sp. nov.">
<emphasis id="A9BFEACDFFD4315EFBF921DE7DE1F9FF" box="[1080,1242,1520,1544]" italics="true" pageId="10" pageNumber="22">N. ctenochelus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B28C57B6FFD4315EFB2421DE7C19F9FF" box="[1253,1314,1521,1544]" pageId="10" pageNumber="22" rank="species">sp. n.</taxonomicNameLabel>
has few (2–3) setae and tapers to thin distal part (
<figureCitation id="03F02A5AFFD4315EFAE4223F7A6DF9BE" captionStart="Figure 1" captionStartId="7.[227,290,1809,1831]" captionTargetBox="[268,1331,400,1671]" captionTargetId="figure-87@7.[239,1348,340,1772]" captionTargetPageId="7" captionText="Figure 1. Nilotanypus Kieffer. Adult. A–D. Head, anterior view, right side, ♂, A, C. ♀, B, D.; E–F. Thorax, E. dor- sal, F. lateral; G. Mid-dorsal sensory pit; H. Wing (male); I, J. Tibial apices, I. P1, J. P3; K. Male hypopygium; L. Gonostylus; M. Female genitalia left side only; N. Anterior vaginal cavity, detail. A–B, E–K, M–N. N. haplochelus sp. n.; C–D, L. N. ctenochelus sp.n. Abbreviations: fr–frontal setae, iv–inner vertical setae, ped–pedestal setae, ov– outer vertical setae, sc-scape setae. Fig. 1G after Roback, 1986." figureDoi="http://doi.org/10.5281/zenodo.7996330" httpUri="https://zenodo.org/record/7996330/files/figure.png" pageId="10" pageNumber="22">Fig. 1L</figureCitation>
) in contrast to the more setose (7)
<taxonomicName id="5CCB4D5CFFD4315EFB06221E7A6CF99F" authority="Cranston &amp; Krosch &amp; Tang, 2022" authorityName="Cranston &amp; Krosch &amp; Tang" authorityYear="2022" class="Insecta" family="Chironomidae" genus="Nilotanypus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="22" phylum="Arthropoda" rank="species" species="haplochelus" status="sp. nov.">
<emphasis id="A9BFEACDFFD4315EFB06221E7A6CF99F" italics="true" pageId="10" pageNumber="22">N. haplochelus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B28C57B6FFD4315EFC9F227E7AA3F99F" box="[862,920,1617,1640]" pageId="10" pageNumber="22" rank="species">sp. n.</taxonomicNameLabel>
with conventional taper to broader distal part (
<figureCitation id="03F02A5AFFD4315EFCB1225F7AFEF97E" box="[880,965,1648,1673]" captionStart="Figure 1" captionStartId="7.[227,290,1809,1831]" captionTargetBox="[268,1331,400,1671]" captionTargetId="figure-87@7.[239,1348,340,1772]" captionTargetPageId="7" captionText="Figure 1. Nilotanypus Kieffer. Adult. A–D. Head, anterior view, right side, ♂, A, C. ♀, B, D.; E–F. Thorax, E. dor- sal, F. lateral; G. Mid-dorsal sensory pit; H. Wing (male); I, J. Tibial apices, I. P1, J. P3; K. Male hypopygium; L. Gonostylus; M. Female genitalia left side only; N. Anterior vaginal cavity, detail. A–B, E–K, M–N. N. haplochelus sp. n.; C–D, L. N. ctenochelus sp.n. Abbreviations: fr–frontal setae, iv–inner vertical setae, ped–pedestal setae, ov– outer vertical setae, sc-scape setae. Fig. 1G after Roback, 1986." figureDoi="http://doi.org/10.5281/zenodo.7996330" httpUri="https://zenodo.org/record/7996330/files/figure.png" pageId="10" pageNumber="22">Fig. 1K</figureCitation>
).
</paragraph>
<paragraph id="9B7436DFFFD4315EFCF9228F7D12F84F" blockId="10.[824,1361,1696,1976]" pageId="10" pageNumber="22">
The two frontal setae in the female
<taxonomicName id="5CCB4D5CFFD4315EFB72228F7C6AF94F" authority="Cranston &amp; Krosch &amp; Tang, 2022" authorityName="Cranston &amp; Krosch &amp; Tang" authorityYear="2022" box="[1203,1361,1696,1720]" class="Insecta" family="Chironomidae" genus="Nilotanypus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="22" phylum="Arthropoda" rank="species" species="ctenochelus" status="sp. nov.">
<emphasis id="A9BFEACDFFD4315EFB72228F7C6AF94F" box="[1203,1361,1696,1720]" italics="true" pageId="10" pageNumber="22">N. ctenochelus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B28C57B6FFD4315EFCF922EF7A4EF920" box="[824,885,1728,1751]" pageId="10" pageNumber="22" rank="species">sp. n.</taxonomicNameLabel>
are diagnostically stout (
<figureCitation id="03F02A5AFFD4315EFB5022EF7DD2F92F" box="[1169,1257,1728,1752]" captionStart="Figure 1" captionStartId="7.[227,290,1809,1831]" captionTargetBox="[268,1331,400,1671]" captionTargetId="figure-87@7.[239,1348,340,1772]" captionTargetPageId="7" captionText="Figure 1. Nilotanypus Kieffer. Adult. A–D. Head, anterior view, right side, ♂, A, C. ♀, B, D.; E–F. Thorax, E. dor- sal, F. lateral; G. Mid-dorsal sensory pit; H. Wing (male); I, J. Tibial apices, I. P1, J. P3; K. Male hypopygium; L. Gonostylus; M. Female genitalia left side only; N. Anterior vaginal cavity, detail. A–B, E–K, M–N. N. haplochelus sp. n.; C–D, L. N. ctenochelus sp.n. Abbreviations: fr–frontal setae, iv–inner vertical setae, ped–pedestal setae, ov– outer vertical setae, sc-scape setae. Fig. 1G after Roback, 1986." figureDoi="http://doi.org/10.5281/zenodo.7996330" httpUri="https://zenodo.org/record/7996330/files/figure.png" pageId="10" pageNumber="22">Fig. 1D</figureCitation>
), in contrast to the conventional narrower frontal setae of
<taxonomicName id="5CCB4D5CFFD4315EFCF9232F7AE2F8EF" authority="Cranston &amp; Krosch &amp; Tang, 2022" authorityName="Cranston &amp; Krosch &amp; Tang" authorityYear="2022" box="[824,985,1792,1816]" class="Insecta" family="Chironomidae" genus="Nilotanypus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="22" phylum="Arthropoda" rank="species" species="haplochelus" status="sp. nov.">
<emphasis id="A9BFEACDFFD4315EFCF9232F7AE2F8EF" box="[824,985,1792,1816]" italics="true" pageId="10" pageNumber="22">N. haplochelus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B28C57B6FFD4315EFC20232F7D20F8E0" box="[993,1051,1792,1815]" pageId="10" pageNumber="22" rank="species">sp. n.</taxonomicNameLabel>
(
<figureCitation id="03F02A5AFFD4315EFBEB232F7D45F8EF" box="[1066,1150,1792,1816]" captionStart="Figure 1" captionStartId="7.[227,290,1809,1831]" captionTargetBox="[268,1331,400,1671]" captionTargetId="figure-87@7.[239,1348,340,1772]" captionTargetPageId="7" captionText="Figure 1. Nilotanypus Kieffer. Adult. A–D. Head, anterior view, right side, ♂, A, C. ♀, B, D.; E–F. Thorax, E. dor- sal, F. lateral; G. Mid-dorsal sensory pit; H. Wing (male); I, J. Tibial apices, I. P1, J. P3; K. Male hypopygium; L. Gonostylus; M. Female genitalia left side only; N. Anterior vaginal cavity, detail. A–B, E–K, M–N. N. haplochelus sp. n.; C–D, L. N. ctenochelus sp.n. Abbreviations: fr–frontal setae, iv–inner vertical setae, ped–pedestal setae, ov– outer vertical setae, sc-scape setae. Fig. 1G after Roback, 1986." figureDoi="http://doi.org/10.5281/zenodo.7996330" httpUri="https://zenodo.org/record/7996330/files/figure.png" pageId="10" pageNumber="22">Fig. 1B</figureCitation>
). The spermathecal ducts are of even width in
<taxonomicName id="5CCB4D5CFFD4315EFBA5230F7C3FF8CF" authority="Cranston &amp; Krosch &amp; Tang, 2022" authorityName="Cranston &amp; Krosch &amp; Tang" authorityYear="2022" box="[1124,1284,1824,1848]" class="Insecta" family="Chironomidae" genus="Nilotanypus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="22" phylum="Arthropoda" rank="species" species="ctenochelus" status="sp. nov.">
<emphasis id="A9BFEACDFFD4315EFBA5230F7C3FF8CF" box="[1124,1284,1824,1848]" italics="true" pageId="10" pageNumber="22">N. ctenochelus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B28C57B6FFD4315EFACF230F7C70F8C0" box="[1294,1355,1824,1847]" pageId="10" pageNumber="22" rank="species">sp. n.</taxonomicNameLabel>
, but have a dilate section in
<taxonomicName id="5CCB4D5CFFD4315EFBA4236F7C3CF8AF" authority="Cranston &amp; Krosch &amp; Tang, 2022" authorityName="Cranston &amp; Krosch &amp; Tang" authorityYear="2022" box="[1125,1287,1856,1880]" class="Insecta" family="Chironomidae" genus="Nilotanypus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="22" phylum="Arthropoda" rank="species" species="haplochelus" status="sp. nov.">
<emphasis id="A9BFEACDFFD4315EFBA4236F7C3CF8AF" box="[1125,1287,1856,1880]" italics="true" pageId="10" pageNumber="22">N. haplochelus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B28C57B6FFD4315EFACE236F7C70F8A0" box="[1295,1355,1856,1879]" pageId="10" pageNumber="22" rank="species">sp. n.</taxonomicNameLabel>
, and seminal vesicles are small with a neck in
<taxonomicName id="5CCB4D5CFFD4315EFAF9234F7A8DF86F" authority="Cranston &amp; Krosch &amp; Tang, 2022" authorityName="Cranston &amp; Krosch &amp; Tang" authorityYear="2022" class="Insecta" family="Chironomidae" genus="Nilotanypus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="22" phylum="Arthropoda" rank="species" species="ctenochelus" status="sp. nov.">
<emphasis id="A9BFEACDFFD4315EFAF9234F7A8DF86F" italics="true" pageId="10" pageNumber="22">N. ctenochelus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B28C57B6FFD4315EFC7F23AF7AC3F860" box="[958,1016,1920,1943]" pageId="10" pageNumber="22" rank="species">sp. n.</taxonomicNameLabel>
but in
<taxonomicName id="5CCB4D5CFFD4315EFB8723AF7DDCF86F" authority="Cranston &amp; Krosch &amp; Tang, 2022" authorityName="Cranston &amp; Krosch &amp; Tang" authorityYear="2022" box="[1094,1255,1920,1944]" class="Insecta" family="Chironomidae" genus="Nilotanypus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="22" phylum="Arthropoda" rank="species" species="haplochelus" status="sp. nov.">
<emphasis id="A9BFEACDFFD4315EFB8723AF7DDCF86F" box="[1094,1255,1920,1944]" italics="true" pageId="10" pageNumber="22">N. haplochelus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B28C57B6FFD4315EFB2F23AF7C12F860" box="[1262,1321,1920,1943]" pageId="10" pageNumber="22" rank="species">sp. n.</taxonomicNameLabel>
are larger and lack a neck.
</paragraph>
<paragraph id="9B7436DFFFD5315FFF2224CC78CFFD8C" blockId="11.[227,764,227,635]" pageId="11" pageNumber="23">
The described pupae of
<taxonomicName id="5CCB4D5CFFD5315FFDC324CC7BB8FF0C" box="[514,643,227,251]" class="Insecta" family="Chironomidae" genus="Nilotanypus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="A9BFEACDFFD5315FFDC324CC7BB8FF0C" box="[514,643,227,251]" italics="true" pageId="11" pageNumber="23">Nilotanypus</emphasis>
</taxonomicName>
especially from
<bibRefCitation id="FF5A4B2EFFD5315FFEE6252C78EBFEEC" author="Roback, S. S." box="[295,464,259,283]" pageId="11" pageNumber="23" pagination="443 - 465" refId="ref17570" refString="Roback, S. S. 1986. The immature chironomids of the eastern United States VIII. Pentaneurini - genus Nilotanypus, with the description of a new species from Kansas. Proceedings of the Academy of Natural Sciences of Philadelphia 138: 443 - 465." type="journal article" year="1986">Roback (1986)</bibRefCitation>
and
<bibRefCitation id="FF5A4B2EFFD5315FFDD6252C781FFECC" author="Roback, S. S. &amp; Coffman, W. P." pageId="11" pageNumber="23" pagination="87 - 158" refId="ref17618" refString="Roback, S. S. and Coffman, W. P. 1987. Results of the Nepal Alpine zone research project, Chironomidae (Diptera). Proceedings of the Academy of Natural Sciences of Philadelphia 139: 87 - 158." type="journal article" year="1987">Roback &amp; Coffman (1987</bibRefCitation>
,
<bibRefCitation id="FF5A4B2EFFD5315FFEEF250C785EFECC" author="Roback, S. S. &amp; Coffman, W. P." box="[302,357,291,315]" pageId="11" pageNumber="23" pagination="85 - 113" refId="ref17662" refString="Roback, S. S. and Coffman, W. P. 1989. Tanypodinae Pupae from Southern India (Diptera: Chironomidae). Proceedings of the Academy of Natural Sciences of Philadelphia 141: 85 - 113." type="journal article" year="1989">1989</bibRefCitation>
) show subtle differentiation with variation in the thoracic horn and in the strength of abdominal armament. Separation of the Australian species depends upon the (unique) row of tubercles on the distal wing sheath of
<taxonomicName id="5CCB4D5CFFD5315FFDE1258B7BFBFE4C" authority="Cranston &amp; Krosch &amp; Tang, 2022" authorityName="Cranston &amp; Krosch &amp; Tang" authorityYear="2022" box="[544,704,419,443]" class="Insecta" family="Chironomidae" genus="Nilotanypus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="23" phylum="Arthropoda" rank="species" species="ctenochelus" status="sp. nov.">
<emphasis id="A9BFEACDFFD5315FFDE1258B7BFBFE4C" box="[544,704,419,443]" italics="true" pageId="11" pageNumber="23">N. ctenochelus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B28C57B6FFD5315FFD08258B7BC7FE4C" box="[713,764,420,443]" pageId="11" pageNumber="23" rank="species">sp.n.</taxonomicNameLabel>
(
<figureCitation id="03F02A5AFFD5315FFF2B25EC7879FE2C" box="[234,322,451,475]" captionStart="Figure 2" captionStartId="9.[227,290,1785,1807]" captionTargetBox="[256,1308,318,1679]" captionTargetId="figure-0@9.[239,1348,287,1775]" captionTargetPageId="9" captionText="Figure 2. Nilotanypus Kieffer. Pupa. A, B. Thoracic horn; C. Wing sheath; D–F. Abdomen, male); D. dorsal, E. ventral. Larva. F. Head capsule, left side ventral, right side dorsal; G. Dorsal head capsule; H. Antenna; I. Antennal apex, detail; J. Mandible; K. Ligula, paraligula; l. Maxilla; M. Submentum; N. Anterior parapod small comb claw; O. Posterior body; P. Posterior parapod comb claw. A, D–F, H–O. N. haplochelus sp. n.; B, C, G, P. N. ctenochelus sp. n. Abbreviations: S5 – S10 – cephalic setae, DP – dorsal pit, VP – ventral pit." figureDoi="http://doi.org/10.5281/zenodo.7996332" httpUri="https://zenodo.org/record/7996332/files/figure.png" pageId="11" pageNumber="23">Figs 2C</figureCitation>
,
<figureCitation id="03F02A5AFFD5315FFE8F25EC7849FE2C" box="[334,370,451,475]" captionStart="Figure 3" captionStartId="11.[228,291,1986,2008]" captionTargetBox="[237,1350,1444,1959]" captionTargetId="figure-579@11.[227,1361,1431,1973]" captionTargetPageId="11" captionText="Figure 3. Nilotanypus ctenochelus sp. n. A. Wing sheath, tubercle row; B. Larval posterior parapod, comb claw." figureDoi="http://doi.org/10.5281/zenodo.7996334" httpUri="https://zenodo.org/record/7996334/files/figure.png" pageId="11" pageNumber="23">3A</figureCitation>
). The two Australian species may be separable also on the thoracic horn: in
<taxonomicName id="5CCB4D5CFFD5315FFD5725CB781CFDEC" authority="Cranston &amp; Krosch &amp; Tang, 2022" authorityName="Cranston &amp; Krosch &amp; Tang" authorityYear="2022" class="Insecta" family="Chironomidae" genus="Nilotanypus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="23" phylum="Arthropoda" rank="species" species="ctenochelus" status="sp. nov.">
<emphasis id="A9BFEACDFFD5315FFD5725CB781CFDEC" italics="true" pageId="11" pageNumber="23">N. ctenochelus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B28C57B6FFD5315FFEEF262B785AFDEC" box="[302,353,516,539]" pageId="11" pageNumber="23" rank="species">sp.n.</taxonomicNameLabel>
the atrium is very narrow in the basal 1/3 and expanded from near the midpoint (
<figureCitation id="03F02A5AFFD5315FFD62260C7BC8FDCC" box="[675,755,547,571]" captionStart="Figure 2" captionStartId="9.[227,290,1785,1807]" captionTargetBox="[256,1308,318,1679]" captionTargetId="figure-0@9.[239,1348,287,1775]" captionTargetPageId="9" captionText="Figure 2. Nilotanypus Kieffer. Pupa. A, B. Thoracic horn; C. Wing sheath; D–F. Abdomen, male); D. dorsal, E. ventral. Larva. F. Head capsule, left side ventral, right side dorsal; G. Dorsal head capsule; H. Antenna; I. Antennal apex, detail; J. Mandible; K. Ligula, paraligula; l. Maxilla; M. Submentum; N. Anterior parapod small comb claw; O. Posterior body; P. Posterior parapod comb claw. A, D–F, H–O. N. haplochelus sp. n.; B, C, G, P. N. ctenochelus sp. n. Abbreviations: S5 – S10 – cephalic setae, DP – dorsal pit, VP – ventral pit." figureDoi="http://doi.org/10.5281/zenodo.7996332" httpUri="https://zenodo.org/record/7996332/files/figure.png" pageId="11" pageNumber="23">Fig. 2B</figureCitation>
) whereas in
<taxonomicName id="5CCB4D5CFFD5315FFE9D266B78C7FDAC" authorityName="Cranston &amp; Krosch &amp; Tang" authorityYear="2022" box="[348,508,579,603]" class="Insecta" family="Chironomidae" genus="Nilotanypus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="23" phylum="Arthropoda" rank="species" species="haplochelus">
<emphasis id="A9BFEACDFFD5315FFE9D266B78C7FDAC" box="[348,508,579,603]" italics="true" pageId="11" pageNumber="23">N. haplochelus</emphasis>
</taxonomicName>
sp. the atrium broadens nearer the base (
<figureCitation id="03F02A5AFFD5315FFE50264C78DCFD8C" box="[401,487,611,635]" captionStart="Figure 2" captionStartId="9.[227,290,1785,1807]" captionTargetBox="[256,1308,318,1679]" captionTargetId="figure-0@9.[239,1348,287,1775]" captionTargetPageId="9" captionText="Figure 2. Nilotanypus Kieffer. Pupa. A, B. Thoracic horn; C. Wing sheath; D–F. Abdomen, male); D. dorsal, E. ventral. Larva. F. Head capsule, left side ventral, right side dorsal; G. Dorsal head capsule; H. Antenna; I. Antennal apex, detail; J. Mandible; K. Ligula, paraligula; l. Maxilla; M. Submentum; N. Anterior parapod small comb claw; O. Posterior body; P. Posterior parapod comb claw. A, D–F, H–O. N. haplochelus sp. n.; B, C, G, P. N. ctenochelus sp. n. Abbreviations: S5 – S10 – cephalic setae, DP – dorsal pit, VP – ventral pit." figureDoi="http://doi.org/10.5281/zenodo.7996332" httpUri="https://zenodo.org/record/7996332/files/figure.png" pageId="11" pageNumber="23">Fig. 2A</figureCitation>
).
</paragraph>
<paragraph id="9B7436DFFFD5315FFF2226BD7B0FFB3C" blockId="11.[227,764,658,1227]" pageId="11" pageNumber="23">
The larvae of the two species of Australian
<taxonomicName id="5CCB4D5CFFD5315FFD6226BD782FFD3D" class="Insecta" family="Chironomidae" genus="Nilotanypus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="A9BFEACDFFD5315FFD6226BD782FFD3D" italics="true" pageId="11" pageNumber="23">Nilotanypus</emphasis>
</taxonomicName>
are differentiated by the posterior parapod in
<taxonomicName id="5CCB4D5CFFD5315FFF2226FC78BAFD1D" authority="Cranston &amp; Krosch &amp; Tang, 2022" authorityName="Cranston &amp; Krosch &amp; Tang" authorityYear="2022" box="[227,385,722,746]" class="Insecta" family="Chironomidae" genus="Nilotanypus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="23" phylum="Arthropoda" rank="species" species="ctenochelus" status="sp. nov.">
<emphasis id="A9BFEACDFFD5315FFF2226FC78BAFD1D" box="[227,385,722,746]" italics="true" pageId="11" pageNumber="23">N. ctenochelus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B28C57B6FFD5315FFE4826FC78FFFD1D" box="[393,452,723,746]" pageId="11" pageNumber="23" rank="species">sp. n.</taxonomicNameLabel>
having a long comb-toothed claw (
<figureCitation id="03F02A5AFFD5315FFEE226DD784BFCFD" box="[291,368,754,778]" captionStart="Figure 2" captionStartId="9.[227,290,1785,1807]" captionTargetBox="[256,1308,318,1679]" captionTargetId="figure-0@9.[239,1348,287,1775]" captionTargetPageId="9" captionText="Figure 2. Nilotanypus Kieffer. Pupa. A, B. Thoracic horn; C. Wing sheath; D–F. Abdomen, male); D. dorsal, E. ventral. Larva. F. Head capsule, left side ventral, right side dorsal; G. Dorsal head capsule; H. Antenna; I. Antennal apex, detail; J. Mandible; K. Ligula, paraligula; l. Maxilla; M. Submentum; N. Anterior parapod small comb claw; O. Posterior body; P. Posterior parapod comb claw. A, D–F, H–O. N. haplochelus sp. n.; B, C, G, P. N. ctenochelus sp. n. Abbreviations: S5 – S10 – cephalic setae, DP – dorsal pit, VP – ventral pit." figureDoi="http://doi.org/10.5281/zenodo.7996332" httpUri="https://zenodo.org/record/7996332/files/figure.png" pageId="11" pageNumber="23">Fig. 2P</figureCitation>
,
<figureCitation id="03F02A5AFFD5315FFEBB26DD78A2FCFD" box="[378,409,754,778]" captionStart="Figure 3" captionStartId="11.[228,291,1986,2008]" captionTargetBox="[237,1350,1444,1959]" captionTargetId="figure-579@11.[227,1361,1431,1973]" captionTargetPageId="11" captionText="Figure 3. Nilotanypus ctenochelus sp. n. A. Wing sheath, tubercle row; B. Larval posterior parapod, comb claw." figureDoi="http://doi.org/10.5281/zenodo.7996334" httpUri="https://zenodo.org/record/7996334/files/figure.png" pageId="11" pageNumber="23">3B</figureCitation>
) that is lacking in
<taxonomicName id="5CCB4D5CFFD5315FFD9D26DC7BC7FCFD" authority="Cranston &amp; Krosch &amp; Tang, 2022" authorityName="Cranston &amp; Krosch &amp; Tang" authorityYear="2022" box="[604,764,754,778]" class="Insecta" family="Chironomidae" genus="Nilotanypus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="23" phylum="Arthropoda" rank="species" species="haplochelus" status="sp. nov.">
<emphasis id="A9BFEACDFFD5315FFD9D26DC7BC7FCFD" box="[604,764,754,778]" italics="true" pageId="11" pageNumber="23">N. haplochelus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B28C57B6FFD5315FFF22273C782EFCDD" box="[227,277,787,810]" pageId="11" pageNumber="23" rank="species">sp.n.</taxonomicNameLabel>
– hence the species epithets. Other differences include dense-packed short comb-teeth claws (
<figureCitation id="03F02A5AFFD5315FFF2A277D787BFC9D" box="[235,320,850,874]" captionStart="Figure 2" captionStartId="9.[227,290,1785,1807]" captionTargetBox="[256,1308,318,1679]" captionTargetId="figure-0@9.[239,1348,287,1775]" captionTargetPageId="9" captionText="Figure 2. Nilotanypus Kieffer. Pupa. A, B. Thoracic horn; C. Wing sheath; D–F. Abdomen, male); D. dorsal, E. ventral. Larva. F. Head capsule, left side ventral, right side dorsal; G. Dorsal head capsule; H. Antenna; I. Antennal apex, detail; J. Mandible; K. Ligula, paraligula; l. Maxilla; M. Submentum; N. Anterior parapod small comb claw; O. Posterior body; P. Posterior parapod comb claw. A, D–F, H–O. N. haplochelus sp. n.; B, C, G, P. N. ctenochelus sp. n. Abbreviations: S5 – S10 – cephalic setae, DP – dorsal pit, VP – ventral pit." figureDoi="http://doi.org/10.5281/zenodo.7996332" httpUri="https://zenodo.org/record/7996332/files/figure.png" pageId="11" pageNumber="23">Fig. 2N</figureCitation>
) on the anterior parapod of
<taxonomicName id="5CCB4D5CFFD5315FFDB2277C7839FC7D" authorityName="Cranston &amp; Krosch &amp; Tang" authorityYear="2022" class="Insecta" family="Chironomidae" genus="Nilotanypus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="23" phylum="Arthropoda" rank="species" species="ctenochelus">
<emphasis id="A9BFEACDFFD5315FFDB2277C7839FC7D" italics="true" pageId="11" pageNumber="23">N. ctenochelus</emphasis>
</taxonomicName>
compared to few simple spinules in claws of
<taxonomicName id="5CCB4D5CFFD5315FFF2227BC78B3FC5D" authorityName="Cranston &amp; Krosch &amp; Tang" authorityYear="2022" box="[227,392,914,938]" class="Insecta" family="Chironomidae" genus="Nilotanypus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="23" phylum="Arthropoda" rank="species" species="haplochelus">
<emphasis id="A9BFEACDFFD5315FFF2227BC78B3FC5D" box="[227,392,914,938]" italics="true" pageId="11" pageNumber="23">N. haplochelus</emphasis>
</taxonomicName>
; and the mid-tooth of the ligula tending to be wider and to protrude further in
<taxonomicName id="5CCB4D5CFFD5315FFD22279C785FFC1D" authorityName="Cranston &amp; Krosch &amp; Tang" authorityYear="2022" class="Insecta" family="Chironomidae" genus="Nilotanypus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="23" phylum="Arthropoda" rank="species" species="haplochelus">
<emphasis id="A9BFEACDFFD5315FFD22279C785FFC1D" italics="true" pageId="11" pageNumber="23">N. haplochelus</emphasis>
</taxonomicName>
(
<figureCitation id="03F02A5AFFD5315FFEB527FD78F6FC1D" box="[372,461,978,1002]" captionStart="Figure 2" captionStartId="9.[227,290,1785,1807]" captionTargetBox="[256,1308,318,1679]" captionTargetId="figure-0@9.[239,1348,287,1775]" captionTargetPageId="9" captionText="Figure 2. Nilotanypus Kieffer. Pupa. A, B. Thoracic horn; C. Wing sheath; D–F. Abdomen, male); D. dorsal, E. ventral. Larva. F. Head capsule, left side ventral, right side dorsal; G. Dorsal head capsule; H. Antenna; I. Antennal apex, detail; J. Mandible; K. Ligula, paraligula; l. Maxilla; M. Submentum; N. Anterior parapod small comb claw; O. Posterior body; P. Posterior parapod comb claw. A, D–F, H–O. N. haplochelus sp. n.; B, C, G, P. N. ctenochelus sp. n. Abbreviations: S5 – S10 – cephalic setae, DP – dorsal pit, VP – ventral pit." figureDoi="http://doi.org/10.5281/zenodo.7996332" httpUri="https://zenodo.org/record/7996332/files/figure.png" pageId="11" pageNumber="23">Fig. 2K</figureCitation>
). The location of the dorsal cephalic seta S
<subScript id="074F349AFFD5315FFE43202D78B1FBE7" attach="left" box="[386,394,1026,1040]" fontSize="6" pageId="11" pageNumber="23">5</subScript>
relative to the dorsal pore and lateral cephalic setae S
<subScript id="074F349AFFD5315FFE7E200C78FCFBC6" attach="left" box="[447,455,1059,1073]" fontSize="6" pageId="11" pageNumber="23">7</subScript>
and S
<subScript id="074F349AFFD5315FFDCA200D7B28FBC7" attach="left" box="[523,531,1058,1072]" fontSize="6" pageId="11" pageNumber="23">8</subScript>
may inform (
<figureCitation id="03F02A5AFFD5315FFD69203D79C1FBBD" captionStart="Figure 2" captionStartId="9.[227,290,1785,1807]" captionTargetBox="[256,1308,318,1679]" captionTargetId="figure-0@9.[239,1348,287,1775]" captionTargetPageId="9" captionText="Figure 2. Nilotanypus Kieffer. Pupa. A, B. Thoracic horn; C. Wing sheath; D–F. Abdomen, male); D. dorsal, E. ventral. Larva. F. Head capsule, left side ventral, right side dorsal; G. Dorsal head capsule; H. Antenna; I. Antennal apex, detail; J. Mandible; K. Ligula, paraligula; l. Maxilla; M. Submentum; N. Anterior parapod small comb claw; O. Posterior body; P. Posterior parapod comb claw. A, D–F, H–O. N. haplochelus sp. n.; B, C, G, P. N. ctenochelus sp. n. Abbreviations: S5 – S10 – cephalic setae, DP – dorsal pit, VP – ventral pit." figureDoi="http://doi.org/10.5281/zenodo.7996332" httpUri="https://zenodo.org/record/7996332/files/figure.png" pageId="11" pageNumber="23">Fig. 2F, G</figureCitation>
). Although the head capsule of
<taxonomicName id="5CCB4D5CFFD5315FFD98201C7BC7FBBD" authorityName="Cranston &amp; Krosch &amp; Tang" authorityYear="2022" box="[601,764,1074,1098]" class="Insecta" family="Chironomidae" genus="Nilotanypus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="23" phylum="Arthropoda" rank="species" species="haplochelus">
<emphasis id="A9BFEACDFFD5315FFD98201C7BC7FBBD" box="[601,764,1074,1098]" italics="true" pageId="11" pageNumber="23">N. haplochelus</emphasis>
</taxonomicName>
is narrower (cephalic ratio ~0.5) compared to
<taxonomicName id="5CCB4D5CFFD5315FFD22207C785AFB7D" authorityName="Cranston &amp; Krosch &amp; Tang" authorityYear="2022" class="Insecta" family="Chironomidae" genus="Nilotanypus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="23" phylum="Arthropoda" rank="species" species="ctenochelus">
<emphasis id="A9BFEACDFFD5315FFD22207C785AFB7D" italics="true" pageId="11" pageNumber="23">N. ctenochelus</emphasis>
</taxonomicName>
(~0.6–0.7), the ratio varies with slide preparation. Otherwise, all mensural features ranges encompass both larval
<typeStatus id="4470887DFFD5315FFE34209C7B0AFB3C" box="[501,561,1203,1227]" pageId="11" pageNumber="23">types</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="D3D16554FFD53159FF2220CE78B3F952" lastPageId="13" lastPageNumber="25" pageId="11" pageNumber="23" type="description">
<paragraph id="9B7436DFFFD5315FFF2220CE785CFB0E" blockId="11.[227,359,1249,1273]" box="[227,359,1249,1273]" pageId="11" pageNumber="23">
<emphasis id="A9BFEACDFFD5315FFF2220CE785CFB0E" bold="true" box="[227,359,1249,1273]" pageId="11" pageNumber="23">Description</emphasis>
</paragraph>
<paragraph id="9B7436DFFFD5315FFF22213F78A1FA7F" blockId="11.[227,764,1296,1416]" pageId="11" pageNumber="23">
Male (n=1–3, all teneral). Total length ~
<quantity id="5C339B3AFFD5315FFD61213E7BCCFADF" box="[672,759,1297,1321]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="11" pageNumber="23" unit="mm" value="1.3">1.3 mm</quantity>
, wing length 750–800 μm. Overall brown throughout, legs slightly paler, abdomen with slightly paler intersegments.
</paragraph>
<paragraph id="9B7436DFFFD5315FFCF924CC7D6AFE4C" blockId="11.[824,1361,227,443]" pageId="11" pageNumber="23">
<emphasis id="A9BFEACDFFD5315FFCF924CC7AAAFF0D" box="[824,913,227,250]" italics="true" pageId="11" pageNumber="23">Antenna</emphasis>
. With 14 flagellomeres, total length 487, terminal flagellomere 40, separate but not offset from penultimate (13th) flagellomere 122, 3× length of terminal flagellomere, apical 2 flagellomeres subequal to 6.5 (6–7) preceding segments. AR 0.50; terminal seta 45–50 long. Pedicel with 1–2 setae, scape without setae.
</paragraph>
<paragraph id="9B7436DFFFD5315FFCF925FD7AE4FD9D" blockId="11.[824,1361,466,618]" pageId="11" pageNumber="23">
<emphasis id="A9BFEACDFFD5315FFCF925FD7A49FE1D" box="[824,882,466,490]" italics="true" pageId="11" pageNumber="23">Head</emphasis>
(
<figureCitation id="03F02A5AFFD5315FFCBE25FD7AEAFE1D" box="[895,977,466,490]" captionStart="Figure 1" captionStartId="7.[227,290,1809,1831]" captionTargetBox="[268,1331,400,1671]" captionTargetId="figure-87@7.[239,1348,340,1772]" captionTargetPageId="7" captionText="Figure 1. Nilotanypus Kieffer. Adult. A–D. Head, anterior view, right side, ♂, A, C. ♀, B, D.; E–F. Thorax, E. dor- sal, F. lateral; G. Mid-dorsal sensory pit; H. Wing (male); I, J. Tibial apices, I. P1, J. P3; K. Male hypopygium; L. Gonostylus; M. Female genitalia left side only; N. Anterior vaginal cavity, detail. A–B, E–K, M–N. N. haplochelus sp. n.; C–D, L. N. ctenochelus sp.n. Abbreviations: fr–frontal setae, iv–inner vertical setae, ped–pedestal setae, ov– outer vertical setae, sc-scape setae. Fig. 1G after Roback, 1986." figureDoi="http://doi.org/10.5281/zenodo.7996330" httpUri="https://zenodo.org/record/7996330/files/figure.png" pageId="11" pageNumber="23">Fig. 1C</figureCitation>
). Eye hairy with dorsomedial extension of 6 ommatidia long. Frontal setae 2, thin, at right angle to 10 uniserial temporal setae, all arising from paler field. Clypeal setae 15. Palp (2–5) 25, 47, 70, 100.
</paragraph>
<paragraph id="9B7436DFFFD5315FFCF926AE7A61FC0E" blockId="11.[824,1361,641,1017]" pageId="11" pageNumber="23">
<emphasis id="A9BFEACDFFD5315FFCF926AE7ABEFD6E" box="[824,901,641,665]" italics="true" pageId="11" pageNumber="23">Thorax</emphasis>
with uniserial tuberculose anterior margin, curved with posteromedian projecting small sense organ (half size of adjacent setal sockets); with 2–3 lateral antepronotal setae; ~17 unevenly uni-/biserial acrostichals; ~16–20 dorsocentrals, humeral cluster disorganised becoming uniserial in pale areas; isolated prescutellar, 10–12 prealars comprising anterior cluster of 4, posterior prealars disorganised; 1 supra-anal; scutellars with posteriormost row of 8 uniserial strong setae, with shorter to much smaller setae anteriorly numbering up to 22.
</paragraph>
<paragraph id="9B7436DFFFD5315FFCF9203E7DE4FB7F" blockId="11.[824,1361,1040,1161]" pageId="11" pageNumber="23">
<emphasis id="A9BFEACDFFD5315FFCF9203E7A42FBDF" box="[824,889,1041,1064]" italics="true" pageId="11" pageNumber="23">Wings</emphasis>
hyaline, veins pale, membrane and veins densely setose, submarginal apical setae dense, strong, 100–120 long. Venation as in
<taxonomicName id="5CCB4D5CFFD5315FFB06207E7A6CFB7F" authorityName="Cranston &amp; Krosch &amp; Tang" authorityYear="2022" class="Insecta" family="Chironomidae" genus="Nilotanypus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="23" phylum="Arthropoda" rank="species" species="haplochelus">
<emphasis id="A9BFEACDFFD5315FFB06207E7A6CFB7F" italics="true" pageId="11" pageNumber="23">N. haplochelus</emphasis>
</taxonomicName>
. Squama with 16–20 uniserial setae,
</paragraph>
<paragraph id="9B7436DFFFD53158FCF9208F78BFFE6C" blockId="11.[824,1361,1183,1412]" lastBlockId="12.[227,764,227,411]" lastPageId="12" lastPageNumber="24" pageId="11" pageNumber="23">
<emphasis id="A9BFEACDFFD5315FFCF9208F7A51FB40" box="[824,874,1184,1207]" italics="true" pageId="11" pageNumber="23">Legs</emphasis>
. Mensural: P
<subScript id="074F349AFFD5315FFBC2209F7D30FB49" attach="left" box="[1027,1035,1200,1214]" fontSize="6" pageId="11" pageNumber="23">1</subScript>
250–255, 212–225, 178, 75, 63, 52, 50; LR
<subScript id="074F349AFFD5315FFC2920FF7ACBFB29" attach="left" box="[1000,1008,1232,1246]" fontSize="6" pageId="11" pageNumber="23">1</subScript>
0.83, BV
<subScript id="074F349AFFD5315FFBA720FF7D55FB29" attach="left" box="[1126,1134,1232,1246]" fontSize="6" pageId="11" pageNumber="23">1</subScript>
2.68, SV
<subScript id="074F349AFFD5315FFB2020FF7DD2FB29" attach="left" box="[1249,1257,1232,1246]" fontSize="6" pageId="11" pageNumber="23">1</subScript>
2.62; P
<subScript id="074F349AFFD5315FFA8820FF7C6AFB29" attach="left" box="[1353,1361,1232,1246]" fontSize="6" pageId="11" pageNumber="23">2</subScript>
320–350, 210–275, 245, 110, 90, 60, 55 LR
<subScript id="074F349AFFD5315FFACC20DF7C2EFB09" attach="left" box="[1293,1301,1264,1278]" fontSize="6" pageId="11" pageNumber="23">2</subScript>
1.18, BV
<subScript id="074F349AFFD5315FFC9C213F7A5EFAE9" attach="left" box="[861,869,1296,1310]" fontSize="6" pageId="11" pageNumber="23">2</subScript>
2.53, SV
<subScript id="074F349AFFD5315FFC0E213F7AECFAE9" attach="left" box="[975,983,1296,1310]" fontSize="6" pageId="11" pageNumber="23">2</subScript>
2.27; P
<subScript id="074F349AFFD5315FFBEF213F7D0DFAE9" attach="left" box="[1070,1078,1296,1310]" fontSize="6" pageId="11" pageNumber="23">3</subScript>
290–295, 200, –, –, –, –, –; spurs 1, 1, 1, each narrow, slightly bent, 30–40 long with basal fine spines (‘hairs’), without lateral comb teeth; tibial comb on P
<subScript id="074F349AFFD5315FFBB021597D42FA73" attach="left" box="[1137,1145,1398,1412]" fontSize="6" pageId="11" pageNumber="23">3</subScript>
comprising slightly curved spines ~30 long. Pseudospurs 30–35 long, 2.5 wide, 2 on subapex of ta
<subScript id="074F349AFFD23158FDEB253C7B09FED6" attach="left" box="[554,562,275,289]" fontSize="6" pageId="12" pageNumber="24">2</subScript>
of midleg (P
<subScript id="074F349AFFD23158FD0D253C7BEFFED6" attach="left" box="[716,724,275,289]" fontSize="6" pageId="12" pageNumber="24">2</subScript>
), 3 slightly longer on subapex of ta
<subScript id="074F349AFFD23158FD86251C7B74FEB6" attach="left" box="[583,591,307,321]" fontSize="6" pageId="12" pageNumber="24">3</subScript>
; seemingly absent on other legs. Claws simple, gently curved, distally rounded, with strong basal rounded lobe. Pulvilli absent.
</paragraph>
<caption id="CFB46657FFD5315FFF2523ED7C29F82F" ID-DOI="http://doi.org/10.5281/zenodo.7996334" ID-Zenodo-Dep="7996334" box="[228,1298,1986,2008]" httpUri="https://zenodo.org/record/7996334/files/figure.png" pageId="11" pageNumber="23" startId="11.[228,291,1986,2008]" targetBox="[237,1350,1444,1959]" targetPageId="11" targetType="figure">
<paragraph id="9B7436DFFFD5315FFF2523ED7C29F82F" blockId="11.[228,1298,1986,2008]" box="[228,1298,1986,2008]" pageId="11" pageNumber="23">
Figure 3.
<taxonomicName id="5CCB4D5CFFD5315FFE8123ED7B17F820" authority="Cranston &amp; Krosch &amp; Tang, 2022" authorityName="Cranston &amp; Krosch &amp; Tang" authorityYear="2022" box="[320,556,1986,2007]" class="Insecta" family="Chironomidae" genus="Nilotanypus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="23" phylum="Arthropoda" rank="species" species="ctenochelus" status="sp. nov.">
<emphasis id="A9BFEACDFFD5315FFE8123ED7B17F820" box="[320,556,1986,2007]" italics="true" pageId="11" pageNumber="23">Nilotanypus ctenochelus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B28C57B6FFD5315FFDF023EC7B5EF82F" box="[561,613,1987,2008]" pageId="11" pageNumber="23" rank="species">sp. n.</taxonomicNameLabel>
A. Wing sheath, tubercle row; B. Larval posterior parapod, comb claw.
</paragraph>
</caption>
<paragraph id="9B7436DFFFD23158FF22259D7BA2FDFD" blockId="12.[227,764,434,522]" pageId="12" pageNumber="24">
<emphasis id="A9BFEACDFFD23158FF22259D7873FE3D" box="[227,328,434,458]" italics="true" pageId="12" pageNumber="24">Abdomen</emphasis>
setose with setae as long as an abdominal segment, organised into partial anterior median and lateral bands, both tergal and sternal.
</paragraph>
<paragraph id="9B7436DFFFD23158FF22260D7BFFFC0E" blockId="12.[227,764,545,1018]" pageId="12" pageNumber="24">
<emphasis id="A9BFEACDFFD23158FF22260D7850FDCE" box="[227,363,546,569]" italics="true" pageId="12" pageNumber="24">Hypopygium</emphasis>
. Tergite IX posteriorly with 6 or 8 aligned long setae; proctiger gently curved (median hyaline). Gonocoxite cylindrical, 55–60 long, maximum width 40, microtrichose, with dense dorso-laterally originating setae&gt;300 long, filling pharate pupal genital sheaths, antero-median inner surface with 4–5 medially-directed setae arising from strong tubercle bases, not coalesced to appear as a lobe. Gonostylus (
<figureCitation id="03F02A5AFFD23158FE17270E7B25FCCE" box="[470,542,801,825]" captionStart="Figure 1" captionStartId="7.[227,290,1809,1831]" captionTargetBox="[268,1331,400,1671]" captionTargetId="figure-87@7.[239,1348,340,1772]" captionTargetPageId="7" captionText="Figure 1. Nilotanypus Kieffer. Adult. A–D. Head, anterior view, right side, ♂, A, C. ♀, B, D.; E–F. Thorax, E. dor- sal, F. lateral; G. Mid-dorsal sensory pit; H. Wing (male); I, J. Tibial apices, I. P1, J. P3; K. Male hypopygium; L. Gonostylus; M. Female genitalia left side only; N. Anterior vaginal cavity, detail. A–B, E–K, M–N. N. haplochelus sp. n.; C–D, L. N. ctenochelus sp.n. Abbreviations: fr–frontal setae, iv–inner vertical setae, ped–pedestal setae, ov– outer vertical setae, sc-scape setae. Fig. 1G after Roback, 1986." figureDoi="http://doi.org/10.5281/zenodo.7996330" httpUri="https://zenodo.org/record/7996330/files/figure.png" pageId="12" pageNumber="24">Fig. 1I</figureCitation>
) 38–42 long, microtrichose, broadest near base, curved from midpoint distally tapering to narrow apex, with 2 fine outer setae, 1 internally, none adjacent to slender megaseta (1 wide, 6–7 long), continuing direction of apical gonostylus, Gc:Gs ratio 1.3–1.88. Phallapodeme strong, sternapodeme shallow arched.
</paragraph>
<paragraph id="9B7436DFFFD23158FF22203F7BDFFBBF" blockId="12.[227,764,1040,1097]" pageId="12" pageNumber="24">
Female (n=3, pharate/teneral). Total length ~
<quantity id="5C339B3AFFD23158FD2F203E7834FBBF" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="12" pageNumber="24" unit="mm" value="2.0">2 mm</quantity>
, wing length ~550–650 μm. Overall brown.
</paragraph>
<paragraph id="9B7436DFFFD23158FF22204F78BEFB20" blockId="12.[227,764,1120,1240]" pageId="12" pageNumber="24">
<emphasis id="A9BFEACDFFD23158FF22204F7807FB80" box="[227,316,1120,1143]" italics="true" pageId="12" pageNumber="24">Antenna</emphasis>
with 12 flagellomeres, total length 287, ultimate ~76–78, with blunt apex; AR 0.32–0.37; terminal seta 100 long. Pedicel with 4–5 setae, scape with 4-5.
</paragraph>
<paragraph id="9B7436DFFFD23158FF2220C07BD8FA50" blockId="12.[227,764,1263,1447]" pageId="12" pageNumber="24">
<emphasis id="A9BFEACDFFD23158FF2220C07826FAF0" box="[227,285,1263,1287]" italics="true" pageId="12" pageNumber="24">Head</emphasis>
(
<figureCitation id="03F02A5AFFD23158FEEB20C07845FAF0" box="[298,382,1263,1287]" captionStart="Figure 1" captionStartId="7.[227,290,1809,1831]" captionTargetBox="[268,1331,400,1671]" captionTargetId="figure-87@7.[239,1348,340,1772]" captionTargetPageId="7" captionText="Figure 1. Nilotanypus Kieffer. Adult. A–D. Head, anterior view, right side, ♂, A, C. ♀, B, D.; E–F. Thorax, E. dor- sal, F. lateral; G. Mid-dorsal sensory pit; H. Wing (male); I, J. Tibial apices, I. P1, J. P3; K. Male hypopygium; L. Gonostylus; M. Female genitalia left side only; N. Anterior vaginal cavity, detail. A–B, E–K, M–N. N. haplochelus sp. n.; C–D, L. N. ctenochelus sp.n. Abbreviations: fr–frontal setae, iv–inner vertical setae, ped–pedestal setae, ov– outer vertical setae, sc-scape setae. Fig. 1G after Roback, 1986." figureDoi="http://doi.org/10.5281/zenodo.7996330" httpUri="https://zenodo.org/record/7996330/files/figure.png" pageId="12" pageNumber="24">Fig. 1D</figureCitation>
). Eye hairy with dorsomedial parallel-sided extension of 6–7 ommatidia. With 2 stout lanceolate frontal setae 40 long, aligned anteroposterior, separated from 8 slender uniserial temporals, all arising from paler field. Clypeal setae 20-23, ~100 long. Palp (1–5) 30, 25; 38; 50; 75.
</paragraph>
<paragraph id="9B7436DFFFD23158FF2221917BFFF8E1" blockId="12.[227,764,1470,1814]" pageId="12" pageNumber="24">
<emphasis id="A9BFEACDFFD23158FF222191780CFA21" box="[227,311,1470,1494]" italics="true" pageId="12" pageNumber="24">Thorax.</emphasis>
With tuberculose anterior margin, without posteromedian scutal sense pit. Setal pits (and setae) in each location (ac, dc, pa, scts) bimodal, all originating from pale areas of cuticle: 2–3 lateral antepronotal setae; ~22 acrostichals +/-biserial throughout, with isolated posterior dorsocentral prescutellar, 14–15 unevenly biserial dorsocentrals; 9–10 prealars separated into anterior 2–3 and posterior cluster; 1 supra-anal; scutellum posteriorly with 8 uniserial strong setae, more anteriorly with shorter / finer setae numbering up to 30.
</paragraph>
<paragraph id="9B7436DFFFD23158FF2223017824F873" blockId="12.[227,764,1837,1924]" pageId="12" pageNumber="24">
<emphasis id="A9BFEACDFFD23158FF222301781FF8B2" box="[227,292,1838,1861]" italics="true" pageId="12" pageNumber="24">Wings</emphasis>
. Apparently as in male. Apical marginal setae up to 80 long. Squama with 17–19 uniserial setae.
</paragraph>
<paragraph id="9B7436DFFFD23158FF2223B27C14FDF2" blockId="12.[227,764,1948,2004]" lastBlockId="12.[824,1362,228,518]" pageId="12" pageNumber="24">
<emphasis id="A9BFEACDFFD23158FF2223B2782EF843" box="[227,277,1949,1972]" italics="true" pageId="12" pageNumber="24">Legs</emphasis>
. Mensural: P
<subScript id="074F349AFFD23158FE6F2383788DF84D" attach="left" box="[430,438,1964,1978]" fontSize="6" pageId="12" pageNumber="24">1</subScript>
225–250, 220–230, 150, 75, 70, 55, 75, LR 0.67, SV 2.96–3.20, BV 2.18; spur 30; P
<subScript id="074F349AFFD23158FCAF24D87A4DFEF2" attach="left" box="[878,886,247,261]" fontSize="6" pageId="12" pageNumber="24">2</subScript>
325–375, 225–238, –, –, –, –; spur 38–40; P
<subScript id="074F349AFFD23158FA8824D87C6AFEF2" attach="left" box="[1353,1361,247,261]" fontSize="6" pageId="12" pageNumber="24">3</subScript>
325–350, 300, –, –, –, –; spur 40. Tibial spurs 1, 1, 1, fine, straight, 30–40 long with basal fine spines (‘hairs’), without lateral teeth; tibial comb on P
<subScript id="074F349AFFD23158FA8825727C6AFE9C" attach="left" box="[1353,1361,349,363]" fontSize="6" pageId="12" pageNumber="24">3</subScript>
comprising 4-5 straight spines, longest 25. Paired proximate pseudospurs 38–42 long, 2.5 wide, subapical of ta
<subScript id="074F349AFFD23158FC7925927AFBFE3C" attach="left" box="[952,960,445,459]" fontSize="6" pageId="12" pageNumber="24">1</subScript>
of foreleg (P
<subScript id="074F349AFFD23158FB9D25927D5FFE3C" attach="left" box="[1116,1124,445,459]" fontSize="6" pageId="12" pageNumber="24">1</subScript>
), no others detected. Claws simple, gently curved, distally rounded, with strong basal rounded lobe. Pulvilli absent.
</paragraph>
<paragraph id="9B7436DFFFD23158FCF926337C09FDA3" blockId="12.[824,1361,540,596]" pageId="12" pageNumber="24">
<emphasis id="A9BFEACDFFD23158FCF926337AA6FDC3" box="[824,925,540,564]" italics="true" pageId="12" pageNumber="24">Abdomen</emphasis>
. Moderately dense setae more or less aligned in anterior and median transverse rows.
</paragraph>
<paragraph id="9B7436DFFFD23158FCF926447D56FC14" blockId="12.[824,1361,619,995]" pageId="12" pageNumber="24">
<emphasis id="A9BFEACDFFD23158FCF926447AA6FD74" box="[824,925,619,643]" italics="true" pageId="12" pageNumber="24">Genitalia</emphasis>
: Gonocoxapodeme VIII weak. Gonapophysis VIII simple microtrichose lobe with short setae throughout. Gonotergite IX weakly protruding, without setae. Coxosternapodeme strong, dark, curved. Notum thin, short (40–50 long) 2× seminal capsule length, posterior part of rami 45– 50 long. Three globular seminal capsules, 25–28 diameter, with distinct neck; spermathecal ducts 120–125 long, of overall even width, bare, ending uncertain. Gonocoxapodeme VIII forming continuous arc across anterior vaginal chamber. Cerci squat, small, 20–25 × 15–18.
</paragraph>
<paragraph id="9B7436DFFFD23158FCF927D47C26FBE5" blockId="12.[824,1309,1019,1043]" box="[824,1309,1019,1043]" pageId="12" pageNumber="24">
Pupa (n=10). Small, total length 2.0–
<quantity id="5C339B3AFFD23158FB0527D47C22FBE5" box="[1220,1305,1019,1043]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="12" pageNumber="24" unit="mm" value="2.7">2.7 mm</quantity>
.
</paragraph>
<paragraph id="9B7436DFFFD23158FCF920057D14FA35" blockId="12.[824,1361,1066,1474]" pageId="12" pageNumber="24">
<emphasis id="A9BFEACDFFD23158FCF920057AE3FBB5" box="[824,984,1066,1090]" italics="true" pageId="12" pageNumber="24">Cephalothorax</emphasis>
. Thoracic horn (
<figureCitation id="03F02A5AFFD23158FB4A20057DE6FBB5" box="[1163,1245,1066,1090]" captionStart="Figure 2" captionStartId="9.[227,290,1785,1807]" captionTargetBox="[256,1308,318,1679]" captionTargetId="figure-0@9.[239,1348,287,1775]" captionTargetPageId="9" captionText="Figure 2. Nilotanypus Kieffer. Pupa. A, B. Thoracic horn; C. Wing sheath; D–F. Abdomen, male); D. dorsal, E. ventral. Larva. F. Head capsule, left side ventral, right side dorsal; G. Dorsal head capsule; H. Antenna; I. Antennal apex, detail; J. Mandible; K. Ligula, paraligula; l. Maxilla; M. Submentum; N. Anterior parapod small comb claw; O. Posterior body; P. Posterior parapod comb claw. A, D–F, H–O. N. haplochelus sp. n.; B, C, G, P. N. ctenochelus sp. n. Abbreviations: S5 – S10 – cephalic setae, DP – dorsal pit, VP – ventral pit." figureDoi="http://doi.org/10.5281/zenodo.7996332" httpUri="https://zenodo.org/record/7996332/files/figure.png" pageId="12" pageNumber="24">Fig. 2B</figureCitation>
) flattenedtubular, spinulose, 130–175 long, 3–3.5 × maximum breadth, with narrow poorly-defined atrium expanded only distally (beyond 50%), with ovoid corona 75–90 long (ratio 48–51%). Thoracic comb uniserial row of 12–15 apically rounded, tubercles, 12–16 (longest) diminishing in size laterad. Basal lobe 32–50 wide, 25–30 high. resembling sharkfin. Thorax microtuberculose anteriorly and close to mid-dorsal ecdysial line. Wing sheath apicodistally with row of c. 20 small marginal tubercles aligned on anterior distal sheath (
<figureCitation id="03F02A5AFFD23158FB7621A57C2BFA55" box="[1207,1296,1418,1442]" captionStart="Figure 2" captionStartId="9.[227,290,1785,1807]" captionTargetBox="[256,1308,318,1679]" captionTargetId="figure-0@9.[239,1348,287,1775]" captionTargetPageId="9" captionText="Figure 2. Nilotanypus Kieffer. Pupa. A, B. Thoracic horn; C. Wing sheath; D–F. Abdomen, male); D. dorsal, E. ventral. Larva. F. Head capsule, left side ventral, right side dorsal; G. Dorsal head capsule; H. Antenna; I. Antennal apex, detail; J. Mandible; K. Ligula, paraligula; l. Maxilla; M. Submentum; N. Anterior parapod small comb claw; O. Posterior body; P. Posterior parapod comb claw. A, D–F, H–O. N. haplochelus sp. n.; B, C, G, P. N. ctenochelus sp. n. Abbreviations: S5 – S10 – cephalic setae, DP – dorsal pit, VP – ventral pit." figureDoi="http://doi.org/10.5281/zenodo.7996332" httpUri="https://zenodo.org/record/7996332/files/figure.png" pageId="12" pageNumber="24">Fig. 2C</figureCitation>
,
<figureCitation id="03F02A5AFFD23158FAE021A57C7FFA55" box="[1313,1348,1418,1442]" captionStart="Figure 3" captionStartId="11.[228,291,1986,2008]" captionTargetBox="[237,1350,1444,1959]" captionTargetId="figure-579@11.[227,1361,1431,1973]" captionTargetPageId="11" captionText="Figure 3. Nilotanypus ctenochelus sp. n. A. Wing sheath, tubercle row; B. Larval posterior parapod, comb claw." figureDoi="http://doi.org/10.5281/zenodo.7996334" httpUri="https://zenodo.org/record/7996334/files/figure.png" pageId="12" pageNumber="24">3A</figureCitation>
), nose shallow to strong.
</paragraph>
<paragraph id="9B7436DFFFD23159FCF921F678AEFEAC" blockId="12.[824,1361,1497,2001]" lastBlockId="13.[227,764,227,347]" lastPageId="13" lastPageNumber="25" pageId="12" pageNumber="24">
<emphasis id="A9BFEACDFFD23158FCF921F67AA6FA06" box="[824,925,1497,1521]" italics="true" pageId="12" pageNumber="24">Abdomen</emphasis>
. Armament as in
<taxonomicName id="5CCB4D5CFFD23158FB9D21F57DC6FA06" authorityName="Cranston &amp; Krosch &amp; Tang" authorityYear="2022" box="[1116,1277,1497,1521]" class="Insecta" family="Chironomidae" genus="Nilotanypus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="24" phylum="Arthropoda" rank="species" species="haplochelus">
<emphasis id="A9BFEACDFFD23158FB9D21F57DC6FA06" box="[1116,1277,1497,1521]" italics="true" pageId="12" pageNumber="24">N. haplochelus</emphasis>
</taxonomicName>
, except reduced on anterior segments to very fine scattered spinules, more microtuberculose on caudal tergites and all pleurae. Setation apparently as in
<taxonomicName id="5CCB4D5CFFD23158FB3E22157AAAF986" authorityName="Cranston &amp; Krosch &amp; Tang" authorityYear="2022" class="Insecta" family="Chironomidae" genus="Nilotanypus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="24" phylum="Arthropoda" rank="species" species="haplochelus">
<emphasis id="A9BFEACDFFD23158FB3E22157AAAF986" italics="true" pageId="12" pageNumber="24">N. haplochelus</emphasis>
</taxonomicName>
including L setae fine, short on VII; on VIII the 5 taeniate LS are distributed across caudal 70% of segment. SVIII posteriorly with subapical spinules, 3–4 long, numbering&gt;50 spinules, uni-biserial, continuous in male; multiserial, shorter and medially interrupted in female. Anal lobe in both sexes, 175–205 long, 170–190 wide, bare, without spinules on either margin, terminating with inwardly curved hyaline blunt hook; anal setae adhesive, with greatest width of anterior (AL
<subScript id="074F349AFFD23158FA8123A67C73F860" attach="left" box="[1344,1352,1929,1943]" fontSize="6" pageId="12" pageNumber="24">1</subScript>
) seta much narrower than broad posterior (AL
<subScript id="074F349AFFD23158FA8123867C73F840" attach="left" box="[1344,1352,1961,1975]" fontSize="6" pageId="12" pageNumber="24">2</subScript>
) seta (width 5–8 versus 20–25). Genital sacs sexually dimorphic, of male tapering,&gt; 400 long,&gt;2× anal lobe; in female bluntly rounded, 0.4× anal lobe length. Bases of genital sacs microspinulose more so in male.
</paragraph>
<paragraph id="9B7436DFFFD33159FF22255C7BF3FE3D" blockId="13.[227,764,370,458]" pageId="13" pageNumber="25">Larva (n=3–4). Head length 380–440, width 190– 240, cephalic index ~0.51–0.60. Yellow with mandible, ligula and occipital margin mid-brown.</paragraph>
<paragraph id="9B7436DFFFD33159FF2225CD7B70FD6E" blockId="13.[227,764,481,666]" pageId="13" pageNumber="25">
<emphasis id="A9BFEACDFFD33159FF2225CD7807FE0E" box="[227,316,482,505]" italics="true" pageId="13" pageNumber="25">Antenna</emphasis>
: basal segment 140–155 long, segment 2 30–36, segment 3 ~ 5–6, segment 4 ~4 long; style and Lauterborn organ ~5 long; blade and accessory blade subequal to flagellum; AR 2.95–3.4; Ring organ slightly protruding at ~55–70% from base. Antenna / mandible ratio 4.2–4.8.
</paragraph>
<paragraph id="9B7436DFFFD33159FF22269F7B58FCFF" blockId="13.[227,764,688,776]" pageId="13" pageNumber="25">
<emphasis id="A9BFEACDFFD33159FF22269F7872FD3F" box="[227,329,688,712]" italics="true" pageId="13" pageNumber="25">Mandible</emphasis>
. 40–47 long, seta subdentalis on welldeveloped distal molar projection (‘tooth’), proximal to distinct, rounded inner tooth.
</paragraph>
<paragraph id="9B7436DFFFD33159FF22270F7800FC8F" blockId="13.[227,764,799,888]" pageId="13" pageNumber="25">
<emphasis id="A9BFEACDFFD33159FF22270F78BAFCCF" box="[227,385,800,824]" italics="true" pageId="13" pageNumber="25">Maxillary palp</emphasis>
. 21–23 long, ring organ faint ~60% from base, longest component of apical crown 16- 20 long.
</paragraph>
<paragraph id="9B7436DFFFD33159FF2227A07B67FB90" blockId="13.[227,764,911,1127]" pageId="13" pageNumber="25">
<emphasis id="A9BFEACDFFD33159FF2227A07812FC50" box="[227,297,911,935]" italics="true" pageId="13" pageNumber="25">Ligula</emphasis>
. 35–39 long, 3–3.5 × as long as apical width, ‘waisted’, with 5 teeth, near straight with central tooth extending only slightly beyond outer teeth. Paraligula squat, bifid, 16 long, Pecten hypopharyngis with 6–8 teeth, innermost largest and directed antero-medially, remainder subequal / narrower points directed anteriorly.
</paragraph>
<paragraph id="9B7436DFFFD33159FF2220517886FAE1" blockId="13.[227,764,1150,1302]" pageId="13" pageNumber="25">
<emphasis id="A9BFEACDFFD33159FF2220517B3FFB61" box="[227,516,1150,1174]" italics="true" pageId="13" pageNumber="25">Submentum / anterior gula</emphasis>
. Ventrally V
<subScript id="074F349AFFD33159FD4E20A17BACFB6B" attach="left" box="[655,663,1166,1180]" fontSize="6" pageId="13" pageNumber="25">9</subScript>
, V
<subScript id="074F349AFFD33159FD7920A17BF3FB6B" attach="left" box="[696,712,1166,1180]" fontSize="6" pageId="13" pageNumber="25">10</subScript>
, VP, SSm as in
<taxonomicName id="5CCB4D5CFFD33159FE9920B078C2FB41" authorityName="Cranston &amp; Krosch &amp; Tang" authorityYear="2022" box="[344,505,1182,1206]" class="Insecta" family="Chironomidae" genus="Nilotanypus" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="25" phylum="Arthropoda" rank="species" species="haplochelus">
<emphasis id="A9BFEACDFFD33159FE9920B078C2FB41" box="[344,505,1182,1206]" italics="true" pageId="13" pageNumber="25">N. haplochelus</emphasis>
</taxonomicName>
(
<figureCitation id="03F02A5AFFD33159FDC820B17B62FB41" box="[521,601,1182,1206]" captionStart="Figure 2" captionStartId="9.[227,290,1785,1807]" captionTargetBox="[256,1308,318,1679]" captionTargetId="figure-0@9.[239,1348,287,1775]" captionTargetPageId="9" captionText="Figure 2. Nilotanypus Kieffer. Pupa. A, B. Thoracic horn; C. Wing sheath; D–F. Abdomen, male); D. dorsal, E. ventral. Larva. F. Head capsule, left side ventral, right side dorsal; G. Dorsal head capsule; H. Antenna; I. Antennal apex, detail; J. Mandible; K. Ligula, paraligula; l. Maxilla; M. Submentum; N. Anterior parapod small comb claw; O. Posterior body; P. Posterior parapod comb claw. A, D–F, H–O. N. haplochelus sp. n.; B, C, G, P. N. ctenochelus sp. n. Abbreviations: S5 – S10 – cephalic setae, DP – dorsal pit, VP – ventral pit." figureDoi="http://doi.org/10.5281/zenodo.7996332" httpUri="https://zenodo.org/record/7996332/files/figure.png" pageId="13" pageNumber="25">Fig. 2F</figureCitation>
, left). Dorsally with S
<subScript id="074F349AFFD33159FEE620E17814FB2B" attach="left" box="[295,303,1230,1244]" fontSize="6" pageId="13" pageNumber="25">7</subScript>
well separated from S
<subScript id="074F349AFFD33159FDDD20E17B1FFB2B" attach="left" box="[540,548,1230,1244]" fontSize="6" pageId="13" pageNumber="25">8</subScript>
with dorsal pit (DP) near midway, but closer to ecdysial line, S5 anterior to DP (
<figureCitation id="03F02A5AFFD33159FE9A20D1788AFAE1" box="[347,433,1278,1302]" captionStart="Figure 2" captionStartId="9.[227,290,1785,1807]" captionTargetBox="[256,1308,318,1679]" captionTargetId="figure-0@9.[239,1348,287,1775]" captionTargetPageId="9" captionText="Figure 2. Nilotanypus Kieffer. Pupa. A, B. Thoracic horn; C. Wing sheath; D–F. Abdomen, male); D. dorsal, E. ventral. Larva. F. Head capsule, left side ventral, right side dorsal; G. Dorsal head capsule; H. Antenna; I. Antennal apex, detail; J. Mandible; K. Ligula, paraligula; l. Maxilla; M. Submentum; N. Anterior parapod small comb claw; O. Posterior body; P. Posterior parapod comb claw. A, D–F, H–O. N. haplochelus sp. n.; B, C, G, P. N. ctenochelus sp. n. Abbreviations: S5 – S10 – cephalic setae, DP – dorsal pit, VP – ventral pit." figureDoi="http://doi.org/10.5281/zenodo.7996332" httpUri="https://zenodo.org/record/7996332/files/figure.png" pageId="13" pageNumber="25">Fig. 2G</figureCitation>
).
</paragraph>
<paragraph id="9B7436DFFFD33159FF22210278B3F952" blockId="13.[227,764,1325,1701]" pageId="13" pageNumber="25">
<emphasis id="A9BFEACDFFD33159FF2221027873FAB2" box="[227,328,1325,1349]" italics="true" pageId="13" pageNumber="25">Abdomen</emphasis>
. Anterior parapod with many small pectinate spinules (
<figureCitation id="03F02A5AFFD33159FE4D216278DBFA92" box="[396,480,1357,1381]" captionStart="Figure 2" captionStartId="9.[227,290,1785,1807]" captionTargetBox="[256,1308,318,1679]" captionTargetId="figure-0@9.[239,1348,287,1775]" captionTargetPageId="9" captionText="Figure 2. Nilotanypus Kieffer. Pupa. A, B. Thoracic horn; C. Wing sheath; D–F. Abdomen, male); D. dorsal, E. ventral. Larva. F. Head capsule, left side ventral, right side dorsal; G. Dorsal head capsule; H. Antenna; I. Antennal apex, detail; J. Mandible; K. Ligula, paraligula; l. Maxilla; M. Submentum; N. Anterior parapod small comb claw; O. Posterior body; P. Posterior parapod comb claw. A, D–F, H–O. N. haplochelus sp. n.; B, C, G, P. N. ctenochelus sp. n. Abbreviations: S5 – S10 – cephalic setae, DP – dorsal pit, VP – ventral pit." figureDoi="http://doi.org/10.5281/zenodo.7996332" httpUri="https://zenodo.org/record/7996332/files/figure.png" pageId="13" pageNumber="25">Fig. 2N</figureCitation>
) proximal to conventional claws. Posterior parapod 175–300 long, ventrally with slender spine, 130–185 long, inserted at 1/3 from base; solitary pectinate claw, 50–55 long with 16–21 internal teeth (
<figureCitation id="03F02A5AFFD33159FDCB21E27B60FA12" box="[522,603,1485,1509]" captionStart="Figure 2" captionStartId="9.[227,290,1785,1807]" captionTargetBox="[256,1308,318,1679]" captionTargetId="figure-0@9.[239,1348,287,1775]" captionTargetPageId="9" captionText="Figure 2. Nilotanypus Kieffer. Pupa. A, B. Thoracic horn; C. Wing sheath; D–F. Abdomen, male); D. dorsal, E. ventral. Larva. F. Head capsule, left side ventral, right side dorsal; G. Dorsal head capsule; H. Antenna; I. Antennal apex, detail; J. Mandible; K. Ligula, paraligula; l. Maxilla; M. Submentum; N. Anterior parapod small comb claw; O. Posterior body; P. Posterior parapod comb claw. A, D–F, H–O. N. haplochelus sp. n.; B, C, G, P. N. ctenochelus sp. n. Abbreviations: S5 – S10 – cephalic setae, DP – dorsal pit, VP – ventral pit." figureDoi="http://doi.org/10.5281/zenodo.7996332" httpUri="https://zenodo.org/record/7996332/files/figure.png" pageId="13" pageNumber="25">Fig. 2P</figureCitation>
,
<figureCitation id="03F02A5AFFD33159FDA921E27BB1FA12" box="[616,650,1485,1509]" captionStart="Figure 3" captionStartId="11.[228,291,1986,2008]" captionTargetBox="[237,1350,1444,1959]" captionTargetId="figure-579@11.[227,1361,1431,1973]" captionTargetPageId="11" captionText="Figure 3. Nilotanypus ctenochelus sp. n. A. Wing sheath, tubercle row; B. Larval posterior parapod, comb claw." figureDoi="http://doi.org/10.5281/zenodo.7996334" httpUri="https://zenodo.org/record/7996334/files/figure.png" pageId="13" pageNumber="25">3B</figureCitation>
), amongst otherwise simple claws. Procercus slightly darkened posteriorly, paler anteriorly, length 42–50, width 12–16, bearing 7 anal setae length 300–400. Supra-anal seta strong, 200–250 long. Anal tubules narrow, tapering, hyaline, up to 400 long, often damaged.
</paragraph>
</subSubSection>
</treatment>
</document>
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