209 lines
21 KiB
XML
209 lines
21 KiB
XML
<document id="CED13C0F64BA306095A83AFE883F1FA6" ID-DOI="10.5281/zenodo.213360" ID-GBIF-Dataset="b1a33963-aa99-478f-8b76-50c13ada801c" ID-ISSN="1175-5326" ID-Zenodo-Dep="213360" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460549395632" checkinUser="plazi" docAuthor="Sathiandran, Nithya & Sabu, Thomas K." docDate="2012" docId="0D07755309360F5AFF3705C2FFDAA906" docLanguage="en" docName="zt03526p058.pdf" docOrigin="Zootaxa 3526" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Onthophagus tnai Nithya & Sabu, new species" docType="treatment" docVersion="9" lastPageNumber="56" masterDocId="F13E0D2B09370F59FFA0036DFD7CAA5A" masterDocTitle="New species, new synonym, and redescription of Onthophagus (Coleoptera: Scarabaeidae: Scarabaeinae) from the Western Ghats, India" masterLastPageNumber="58" masterPageNumber="53" pageNumber="54" updateTime="1698318061882" updateUser="plazi">
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<mods:mods id="821A6D75CE2EE3D9E22E2736C9FC3F27" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="D3DCCBAD4BB8370286ADA006E1BEC75B">
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<mods:title id="9ADC00EAFDE040E430D41A6300CB9711">New species, new synonym, and redescription of Onthophagus (Coleoptera: Scarabaeidae: Scarabaeinae) from the Western Ghats, India</mods:title>
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</mods:titleInfo>
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<mods:name id="DEB492CC62E6B56E382B95D54FFFDBCC" type="personal">
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<mods:roleTerm id="FCAEDDA518B5B9E1F9D90C4BAD5F37DB">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="DD4B2FEAA5C89CC2419DB9EAF374E73B">Sathiandran, Nithya</mods:namePart>
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</mods:name>
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<mods:name id="216202AB5D9900F0A7DA9B4B7B3F7E26" type="personal">
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<mods:role id="E8573D9EFEB0E8A02C83EE8A50DBCBA7">
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<mods:roleTerm id="2D3B091CFD8999259786FF430C8839D0">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="11962832335EC6853708383A5DD70589">Sabu, Thomas K.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="F9DB42C3A57BF589934DF92DA76FF863">text</mods:typeOfResource>
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<mods:titleInfo id="8957321179D4F011D2AE7B1DD364EDE2">
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<mods:title id="A3F1370E589071EC04EA4A7A615971AC">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="B540556D62C27CEA1B404A4A4837F3F5">
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<mods:date id="17B45226A8B64810228732FAF9052CD0">2012</mods:date>
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<mods:detail id="FA86B0CE739A0FAB77CF7FA33376F53F" type="volume">
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<mods:number id="4DFB11334BD15C26ADC25BCD4A85BA14">3526</mods:number>
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<mods:start id="90B84BFC696B0D7F3F5A5AED6849D276">53</mods:start>
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<mods:end id="AED630BFC2AC16D4FF64C42A53A829DB">58</mods:end>
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<mods:classification id="D6DBF9D196C30631AA76D38660543A6E">journal article</mods:classification>
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<mods:identifier id="0B17EB22BE920CCE83354C1C3B674B44" type="DOI">10.5281/zenodo.213360</mods:identifier>
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<mods:identifier id="D57E1DED63D632C34768F8378CABA99D" type="GBIF-Dataset">b1a33963-aa99-478f-8b76-50c13ada801c</mods:identifier>
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<mods:identifier id="0CB4603FB9104375A5615282B54F9FB4" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="E859DAA681E14841B7075C592078432A" type="Zenodo-Dep">213360</mods:identifier>
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</mods:mods>
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<treatment id="0D07755309360F5AFF3705C2FFDAA906" ID-DOI="http://doi.org/10.5281/zenodo.6175212" ID-GBIF-Taxon="119529158" ID-Zenodo-Dep="6175212" LSID="urn:lsid:plazi:treatment:0D07755309360F5AFF3705C2FFDAA906" httpUri="http://treatment.plazi.org/id/0D07755309360F5AFF3705C2FFDAA906" lastPageId="3" lastPageNumber="56" pageId="1" pageNumber="54">
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<subSubSection id="CDB497CE09360F58FF3705C2FC6EACB0" pageId="1" pageNumber="54" type="nomenclature">
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<paragraph id="8511C44509360F58FF3705C2FFAFAC93" blockId="1.[151,723,1711,1770]" box="[151,723,1711,1737]" pageId="1" pageNumber="54">
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<heading id="DE59732909360F58FF3705C2FFAFAC93" bold="true" box="[151,723,1711,1737]" fontSize="11" level="1" pageId="1" pageNumber="54" reason="1">
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<emphasis id="B7DA185709360F58FF3705C2FFAFAC93" bold="true" box="[151,723,1711,1737]" pageId="1" pageNumber="54">
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<taxonomicName id="42AEBFC609360F58FF3705C2FF49AC93" ID-CoL="7XPCL" authority="Nithya & Sabu" authorityName="Nithya & Sabu" box="[151,565,1711,1737]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="54" phylum="Arthropoda" rank="species" species="tnai" status="sp. nov.">
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<emphasis id="B7DA185709360F58FF3705C2FC0EAC93" bold="true" box="[151,370,1711,1737]" italics="true" pageId="1" pageNumber="54">Onthophagus tnai</emphasis>
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Nithya & Sabu
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</taxonomicName>
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,
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<taxonomicNameLabel id="ACE9A52C09360F58FDE305C2FFAFAC93" box="[579,723,1711,1737]" pageId="1" pageNumber="54" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8511C44509360F58FF3705BFFC6EACB0" blockId="1.[151,723,1711,1770]" box="[151,274,1746,1770]" pageId="1" pageNumber="54">
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||
(
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<figureCitation id="1D95D8C009360F58FF3F05BFFC75ACB0" box="[159,265,1746,1770]" captionStart="FIGURES 1 – 3" captionStartId="2.[151,264,2006,2029]" captionTargetBox="[184,1422,477,2005]" captionTargetId="figure@2.[169,1433,455,2031]" captionTargetPageId="2" captionText="FIGURES 1 – 3. Onthophagus tnai (holotype): dorsal habitus (1), head (2), aedeagus, lateral view (3). Scale bar = 1 mm." httpUri="https://zenodo.org/record/213361/files/figure.png" pageId="1" pageNumber="54">Figs. 1–3</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="CDB497CE09360F5AFF370477F933AAAD" lastPageId="3" lastPageNumber="56" pageId="1" pageNumber="54" type="description">
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<paragraph id="8511C44509360F5BFF370477FF6DABF6" blockId="1.[151,1436,1818,2022]" lastBlockId="2.[151,1437,151,428]" lastPageId="2" lastPageNumber="55" pageId="1" pageNumber="54">
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<emphasis id="B7DA185709360F58FF370477FC8FAD69" bold="true" box="[151,499,1818,1843]" pageId="1" pageNumber="54">
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Description. Male (
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<typeStatus id="5A157AE709360F58FEDA0477FC9AAD69" box="[378,486,1818,1843]" pageId="1" pageNumber="54" type="holotype">holotype</typeStatus>
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).
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</emphasis>
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Body (
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<figureCitation id="1D95D8C009360F58FDE50477FFF8AD68" box="[581,644,1818,1842]" captionStart="FIGURES 1 – 3" captionStartId="2.[151,264,2006,2029]" captionTargetBox="[184,1422,477,2005]" captionTargetId="figure@2.[169,1433,455,2031]" captionTargetPageId="2" captionText="FIGURES 1 – 3. Onthophagus tnai (holotype): dorsal habitus (1), head (2), aedeagus, lateral view (3). Scale bar = 1 mm." httpUri="https://zenodo.org/record/213361/files/figure.png" pageId="1" pageNumber="54">Fig. 1</figureCitation>
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) shiny black, head and pronotum dark coppery, antennae and tarsi red, body clothed with long yellow setae.
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<emphasis id="B7DA185709360F58FD9C0452FFFFAD0D" bold="true" box="[572,643,1855,1879]" pageId="1" pageNumber="54">Head.</emphasis>
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Broad, ocular lobes gently rounded laterally; clypeus truncate without sinuation; anterior margin reflexed. Head with backwardly produced, elevated lamina with emargination. Clypeus with large punctures laterally; close punctures on the lobes of emargination on head; middle smooth with sparse, fine punctures. Eyes small, moderately elliptic. Frontal carina absent (
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<figureCitation id="1D95D8C009360F58FC1804C7FE83AD98" box="[952,1023,1962,1986]" captionStart="FIGURES 1 – 3" captionStartId="2.[151,264,2006,2029]" captionTargetBox="[184,1422,477,2005]" captionTargetId="figure@2.[169,1433,455,2031]" captionTargetPageId="2" captionText="FIGURES 1 – 3. Onthophagus tnai (holotype): dorsal habitus (1), head (2), aedeagus, lateral view (3). Scale bar = 1 mm." httpUri="https://zenodo.org/record/213361/files/figure.png" pageId="1" pageNumber="54">Fig. 2</figureCitation>
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).
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<emphasis id="B7DA185709360F58FBB504C6F9E9AD99" bold="true" box="[1045,1173,1963,1987]" pageId="1" pageNumber="54">Pronotum.</emphasis>
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Convex; bearing close punctures; with a small, smooth, shiny elongate patch in the middle of the dorsal surface. Front angles produced anteriorly, hind angles obtuse, base strongly rounded. Anterior side of the pronotum smooth with some punctures.
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<emphasis id="B7DA185709350F5BFF3703D1FD96AA8E" bold="true" box="[151,234,188,212]" pageId="2" pageNumber="55">Elytra.</emphasis>
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Black, shiny, finely striate; intervals flat, bearing setiferous punctures; elytron with 7 reddish-yellow patches, 1 at the shoulder, 1 near the middle of the outer margin, 3 at the apical angle, a pair near the base and the suture.
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<emphasis id="B7DA185709350F5BFF4C026EFC43AB46" bold="true" box="[236,319,259,284]" pageId="2" pageNumber="55">Wings.</emphasis>
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<collectingCountry id="FDB984D509350F5BFEE90269FCFDAB46" box="[329,385,260,284]" name="India" pageId="2" pageNumber="55">Hind</collectingCountry>
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wings present.
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<emphasis id="B7DA185709350F5BFD990269FF86AB46" bold="true" box="[569,762,259,284]" pageId="2" pageNumber="55">Ventral surface.</emphasis>
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Pygidium basally bordered; shiny; with moderately close, large punctures. Mesosternum and metasternum with scattered punctures except medially.
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<emphasis id="B7DA185709350F5BFB430245F85EAB1A" bold="true" box="[1251,1314,296,320]" pageId="2" pageNumber="55">Legs.</emphasis>
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Profemur unmodified; protibia elongate, quadridentate externally with serration between teeth; mesofemur and metafemur unmodified; mesotarsi and metatarsi as long as tibia.
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<emphasis id="B7DA185709350F5BFD4D021DFEDFABD2" bold="true" box="[749,931,367,392]" pageId="2" pageNumber="55">Male genetalia.</emphasis>
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Parameres angled relative to basal piece and pointed towards the apex (
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<figureCitation id="1D95D8C009350F5BFE1F02F9FF7FABF6" box="[447,515,404,428]" captionStart="FIGURES 1 – 3" captionStartId="2.[151,264,2006,2029]" captionTargetBox="[184,1422,477,2005]" captionTargetId="figure@2.[169,1433,455,2031]" captionTargetPageId="2" captionText="FIGURES 1 – 3. Onthophagus tnai (holotype): dorsal habitus (1), head (2), aedeagus, lateral view (3). Scale bar = 1 mm." httpUri="https://zenodo.org/record/213361/files/figure.png" pageId="2" pageNumber="55">Fig. 3</figureCitation>
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).
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</paragraph>
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<caption id="D1D194CD09350F5BFF3704BBF820ADB6" box="[151,1372,2006,2029]" httpUri="https://zenodo.org/record/213361/files/figure.png" pageId="2" pageNumber="55" targetBox="[184,1422,477,2005]" targetPageId="2">
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||
<paragraph id="8511C44509350F5BFF3704BBF820ADB6" blockId="2.[151,1372,2006,2029]" box="[151,1372,2006,2029]" pageId="2" pageNumber="55">
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<emphasis id="B7DA185709350F5BFF3704BBFC46ADB6" bold="true" box="[151,314,2006,2029]" pageId="2" pageNumber="55">FIGURES 1–3.</emphasis>
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<taxonomicName id="42AEBFC609350F5BFEE004BAFC8AADB7" box="[320,502,2007,2029]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="55" phylum="Arthropoda" rank="species" species="tnai">
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<emphasis id="B7DA185709350F5BFEE004BAFC8AADB7" box="[320,502,2007,2029]" italics="true" pageId="2" pageNumber="55">Onthophagus tnai</emphasis>
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</taxonomicName>
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(holotype): dorsal habitus (1), head (2), aedeagus, lateral view (3). Scale bar = 1 mm.
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</paragraph>
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</caption>
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<paragraph id="8511C44509340F5AFF6703F5FCDEAAF5" blockId="3.[151,1437,151,860]" box="[199,418,151,176]" pageId="3" pageNumber="56">
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<emphasis id="B7DA185709340F5AFF6703F5FC59AAEA" bold="true" box="[199,293,152,176]" pageId="3" pageNumber="56">Female.</emphasis>
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Unknown.
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</paragraph>
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<paragraph id="8511C44509340F5AFF6703D1F933AAAD" blockId="3.[151,1437,151,860]" pageId="3" pageNumber="56">
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<emphasis id="B7DA185709340F5AFF6703D1FC0AAA8E" bold="true" box="[199,374,188,212]" pageId="3" pageNumber="56">Measurements</emphasis>
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(mm; n = 1:
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<typeStatus id="5A157AE709340F5AFDB303D1FF06AA8E" box="[531,634,188,212]" pageId="3" pageNumber="56" type="holotype">holotype</typeStatus>
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). TL = 7.0, BW = 4.0, PL = 2.0 4, PW = 3.84, EL = 3.3.
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<typeStatus id="5A157AE709340F5AFAAF03D0F800AA8F" box="[1295,1404,189,213]" pageId="3" pageNumber="56" type="paratype">Paratypes</typeStatus>
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(
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<specimenCount id="93A80FCC09340F5AFA2D03D1FDA2AAAD" pageId="3" pageNumber="56" type="male">2 males</specimenCount>
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): TL = 6.75–7.17, BW = 3.9–4.2, PL = 1.95–2.1, PW = 3.3–3.9, EL = 3.3–3.4.
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</paragraph>
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</subSubSection>
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<subSubSection id="CDB497CE09340F5AFF670269FEEDABDD" pageId="3" pageNumber="56" type="materials_examined">
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<paragraph id="8511C44509340F5AFF670269FEA4AB65" blockId="3.[151,1437,151,860]" pageId="3" pageNumber="56">
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<emphasis id="B7DA185709340F5AFF670269FC0BAB46" bold="true" box="[199,375,259,284]" pageId="3" pageNumber="56">
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<typeStatus id="5A157AE709340F5AFF670269FC7EAB46" box="[199,258,260,284]" pageId="3" pageNumber="56">Type</typeStatus>
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material.
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</emphasis>
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<typeStatus id="5A157AE709340F5AFEDF0269FC90AB46" box="[383,492,260,284]" pageId="3" pageNumber="56" type="holotype">Holotype</typeStatus>
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: (male, in ZSI- Ca): “
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<collectingCountry id="FDB984D509340F5AFD460269FE58AB46" box="[742,804,260,284]" name="India" pageId="3" pageNumber="56">India</collectingCountry>
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, Kerala, Palghat, Silent valley National Park, Syrandri,
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<quantity id="425669A009340F5AFF37024AFD92AB65" box="[151,238,295,319]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="56" unit="m" value="1000.0">1000 m</quantity>
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, evergreen forest, dung baited pit fall trap,
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<date id="F110E28509340F5AFD71024AFE47AB65" box="[721,827,295,319]" pageId="3" pageNumber="56" value="2008-05-09">9.v. 2008</date>
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, Vinod, K.V.”
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</paragraph>
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<paragraph id="8511C44509340F5AFF670220FEEDABDD" blockId="3.[151,1437,151,860]" pageId="3" pageNumber="56">
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<typeStatus id="5A157AE709340F5AFF670220FC48AB3F" box="[199,308,333,357]" pageId="3" pageNumber="56" type="paratype">Paratypes</typeStatus>
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||
(2):
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<specimenCount id="93A80FCC09340F5AFECF0221FCC0AB3E" box="[367,444,332,356]" pageId="3" pageNumber="56" type="male">1 male</specimenCount>
|
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in
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<collectionCode id="E3BF5C8009340F5AFE450221FF63AB3E" box="[485,543,332,356]" httpUri="http://grbio.org/cool/nwky-2b63" name="National Pusa Collection" pageId="3" pageNumber="56">NPC</collectionCode>
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; “
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<collectingCountry id="FDB984D509340F5AFD970221FF09AB3E" box="[567,629,332,356]" name="India" pageId="3" pageNumber="56">India</collectingCountry>
|
||
, Kerala, Panathady,
|
||
<quantity id="425669A009340F5AFCC10221FED1AB3E" box="[865,941,332,356]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="3" pageNumber="56" unit="m" value="900.0">900 m</quantity>
|
||
, semi evergreen forest, dung baited pit fall trap,
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<date id="F110E28509340F5AFF700202FC38ABDD" box="[208,324,367,391]" pageId="3" pageNumber="56" value="2007-12-03">3.xii.2007</date>
|
||
, Shiju, T.R.”,
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||
<specimenCount id="93A80FCC09340F5AFE430202FF52ABDD" box="[483,558,367,391]" pageId="3" pageNumber="56" type="male">1 male</specimenCount>
|
||
in SJC; Same data as
|
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<typeStatus id="5A157AE709340F5AFC870202FEF1ABDD" box="[807,909,367,391]" pageId="3" pageNumber="56" type="holotype">holotype</typeStatus>
|
||
.
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||
</paragraph>
|
||
</subSubSection>
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||
<subSubSection id="CDB497CE09340F5AFF6702FEF973ABF6" box="[199,1039,403,428]" pageId="3" pageNumber="56" type="distribution">
|
||
<paragraph id="8511C44509340F5AFF6702FEF973ABF6" blockId="3.[151,1437,151,860]" box="[199,1039,403,428]" pageId="3" pageNumber="56">
|
||
<emphasis id="B7DA185709340F5AFF6702FEFC1CABF6" bold="true" box="[199,352,403,428]" pageId="3" pageNumber="56">Distribution.</emphasis>
|
||
<collectingCountry id="FDB984D509340F5AFEC702F9FCDDABF6" box="[359,417,404,428]" name="India" pageId="3" pageNumber="56">India</collectingCountry>
|
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(South Western Ghats montane rain forests ecoregion).
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</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CDB497CE09340F5AFF6702D5FCE1ABAE" pageId="3" pageNumber="56" type="etymology">
|
||
<paragraph id="8511C44509340F5AFF6702D5FCE1ABAE" blockId="3.[151,1437,151,860]" pageId="3" pageNumber="56">
|
||
<emphasis id="B7DA185709340F5AFF6702D5FC32AB8A" bold="true" box="[199,334,440,464]" pageId="3" pageNumber="56">Etymology.</emphasis>
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This species is named in honor of the doyen of Indian entomology, Prof. T.N. Anathakrishnan (TNA), Chennai,
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<collectingCountry id="FDB984D509340F5AFEFC02B1FCE5ABAE" box="[348,409,476,500]" name="India" pageId="3" pageNumber="56">India</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CDB497CE09340F5AFF67016DFDAFA8B5" pageId="3" pageNumber="56" type="discussion">
|
||
<paragraph id="8511C44509340F5AFF67016DFDAFA8B5" blockId="3.[151,1437,151,860]" pageId="3" pageNumber="56">
|
||
<emphasis id="B7DA185709340F5AFF67016DFC47A842" bold="true" box="[199,315,512,536]" pageId="3" pageNumber="56">Remarks.</emphasis>
|
||
<taxonomicName id="42AEBFC609340F5AFEE8016DFF65A842" box="[328,537,512,536]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="56" phylum="Arthropoda" rank="species" species="tnai">
|
||
<emphasis id="B7DA185709340F5AFEE8016DFF65A842" box="[328,537,512,536]" italics="true" pageId="3" pageNumber="56">Onthophagus tnai</emphasis>
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</taxonomicName>
|
||
has strong similarities to
|
||
<taxonomicName id="42AEBFC609340F5AFCFB016DFE92A842" box="[859,1006,512,536]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="56" phylum="Arthropoda" rank="species" species="kchatriya">
|
||
<emphasis id="B7DA185709340F5AFCFB016DFE92A842" box="[859,1006,512,536]" italics="true" pageId="3" pageNumber="56">O. kchatriya</emphasis>
|
||
</taxonomicName>
|
||
in having black elytra with yellow markings, a pattern of yellow markings on the elytra, and the presence of a backwardly raised lamina on the head. But
|
||
<taxonomicName id="42AEBFC609340F5AFF660125FC68A83A" box="[198,276,584,608]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="56" phylum="Arthropoda" rank="species" species="tnai">
|
||
<emphasis id="B7DA185709340F5AFF660125FC68A83A" box="[198,276,584,608]" italics="true" pageId="3" pageNumber="56">O. tnai</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="42AEBFC609340F5AFE070125FF4FA83A" box="[423,563,584,608]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="56" phylum="Arthropoda" rank="species" species="kchatriya">
|
||
<emphasis id="B7DA185709340F5AFE070125FF4FA83A" box="[423,563,584,608]" italics="true" pageId="3" pageNumber="56">O. kchatriya</emphasis>
|
||
</taxonomicName>
|
||
in the presence of a punctate pronotum and the absence of horns on the head.
|
||
<taxonomicName id="42AEBFC609340F5AFF370101FC19A8DE" box="[151,357,620,644]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="56" phylum="Arthropoda" rank="species" species="tnai">
|
||
<emphasis id="B7DA185709340F5AFF370101FC19A8DE" box="[151,357,620,644]" italics="true" pageId="3" pageNumber="56">Onthophagus tnai</emphasis>
|
||
</taxonomicName>
|
||
is similar to
|
||
<taxonomicName id="42AEBFC609340F5AFDA60100FFCCA8DE" box="[518,688,620,644]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="56" phylum="Arthropoda" rank="species" species="lemniscatus">
|
||
<emphasis id="B7DA185709340F5AFDA60100FFCCA8DE" box="[518,688,620,644]" italics="true" pageId="3" pageNumber="56">O. lemniscatus</emphasis>
|
||
</taxonomicName>
|
||
with a punctate pronotum and broadly truncate front margin of clypeus. It differs from
|
||
<taxonomicName id="42AEBFC609340F5AFE0401FDFF30A8F2" box="[420,588,656,680]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="56" phylum="Arthropoda" rank="species" species="lemniscatus">
|
||
<emphasis id="B7DA185709340F5AFE0401FDFF30A8F2" box="[420,588,656,680]" italics="true" pageId="3" pageNumber="56">O. lemniscatus</emphasis>
|
||
</taxonomicName>
|
||
in the presence of black elytra with yellow markings in contrast to yellow elytra with black markings, and the distribution pattern of black markings on the elytra and lack of horns on the head.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CDB497CE09340F5AFF670196FFDAA906" pageId="3" pageNumber="56" type="description">
|
||
<paragraph id="8511C44509340F5AFF670196FFDAA906" blockId="3.[151,1437,151,860]" pageId="3" pageNumber="56">
|
||
<emphasis id="B7DA185709340F5AFF670196FC47A94E" bold="true" box="[199,315,763,788]" pageId="3" pageNumber="56">Variation.</emphasis>
|
||
The
|
||
<typeStatus id="5A157AE709340F5AFED80190FCA4A94F" box="[376,472,765,789]" pageId="3" pageNumber="56" type="paratype">paratype</typeStatus>
|
||
collected from the same locality as the
|
||
<typeStatus id="5A157AE709340F5AFC350191FE8BA94E" box="[917,1015,764,788]" pageId="3" pageNumber="56" type="holotype">holotype</typeStatus>
|
||
does not vary in any significant way; the other
|
||
<typeStatus id="5A157AE709340F5AFEA2004DFC1EA962" box="[258,354,800,824]" pageId="3" pageNumber="56" type="paratype">paratype</typeStatus>
|
||
has two punctate gibbosities on the anterior side of the pronotum, which contrasts with the lack of punctate gibbosities in other two specimens.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |