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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.656.8914" ID-GBIF-Dataset="51a5b313-2a8b-4bde-8d9d-06bd051fcb42" ID-PMC="PMC5345363" ID-Pensoft-Pub="1313-2970-656-85" ID-PubMed="28331403" ID-ZBK="89234B7B2F01446EB8C077C29CE106AE" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-656-85" ModsDocOrigin="ZooKeys 656" ModsDocTitle="The genus Alaolacon Candèze, a senior synonym of the genus Eumoeus Candèze (Coleoptera, Elateridae, Agrypninae)" checkinTime="1487100179832" checkinUser="pensoft" docAuthor="Arimoto, Koichi &amp; Arimoto, Hisayuki" docDate="2017" docId="00A12B2BEFD6E38933CB83FAD5D17D97" docLanguage="en" docName="ZooKeys 656: 85-110" docOrigin="ZooKeys 656" docSource="http://dx.doi.org/10.3897/zookeys.656.8914" docTitle="Alaolacon candezei Fleutiaux 1928" docType="treatment" docVersion="4" lastPageNumber="91" masterDocId="901EFFE5FFA1FFA53456FFDC5C68FFFC" masterDocTitle="The genus Alaolacon Candeze, a senior synonym of the genus Eumoeus Candeze (Coleoptera, Elateridae, Agrypninae)" masterLastPageNumber="110" masterPageNumber="85" pageNumber="89" updateTime="1668164237870" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>The genus Alaolacon Candeze, a senior synonym of the genus Eumoeus Candeze (Coleoptera, Elateridae, Agrypninae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Arimoto, Koichi</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Arimoto, Hisayuki</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2017</mods:date>
<mods:detail type="volume">
<mods:number>656</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>85</mods:start>
<mods:end>110</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.656.8914</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.656.8914</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-656-85</mods:identifier>
<mods:identifier type="ZBK">89234B7B2F01446EB8C077C29CE106AE</mods:identifier>
<mods:identifier type="ZooBank">89234B7B2F01446EB8C077C29CE106AE</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="127855046" LSID="urn:lsid:plazi:treatment:00A12B2BEFD6E38933CB83FAD5D17D97" httpUri="http://treatment.plazi.org/id/00A12B2BEFD6E38933CB83FAD5D17D97" lastPageId="6" lastPageNumber="91" pageId="4" pageNumber="89">
<subSubSection pageId="4" pageNumber="89" type="nomenclature">
<paragraph pageId="4" pageNumber="89">
<taxonomicName authority="Fleutiaux, 1928" authorityName="Fleutiaux" authorityYear="1928" class="Insecta" family="Elateridae" genus="Alaolacon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Alaolacon candezei" order="Coleoptera" pageId="4" pageNumber="89" phylum="Arthropoda" rank="species" species="candezei">Alaolacon candezei Fleutiaux, 1928</taxonomicName>
Figures 1-3, 4-11, 12-16
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="89" type="reference_group">
<paragraph pageId="4" pageNumber="89">
<taxonomicName class="Insecta" family="Elateridae" genus="Alaolacon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Alaolacon candezei" order="Coleoptera" pageId="4" pageNumber="89" phylum="Arthropoda" rank="species" species="candezei">Alaolacon candezei</taxonomicName>
Fleutiaux, 1928: 177 (original description; type locality: Malaysia, East Malaysia (Sabah), Banggi Island).
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="89" type="type material">
<paragraph pageId="4" pageNumber="89">Type material.</paragraph>
<paragraph pageId="4" pageNumber="89">
Holotype. Female, Banggi Island (located off the northern coast of Borneo), Sabah, Malaysia, Waterstradt leg. [MNHN] (Fig. 3). Label data:
<normalizedToken originalValue="“TYPE”">&quot;TYPE&quot;</normalizedToken>
; [female symbol]; &quot;Banguey/ Borneo/ Waterstradt&quot; &quot;=
<taxonomicName lsidName="cyanipennis" pageId="4" pageNumber="89" rank="species" species="cyanipennis">cyanipennis</taxonomicName>
Cand.?/ Collection FLEUTIAUX&quot;; &quot;
<taxonomicName class="Insecta" family="Elateridae" genus="Alaolacon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Alaolacon" order="Coleoptera" pageId="4" pageNumber="89" phylum="Arthropoda" rank="genus">Alaolacon</taxonomicName>
/
<taxonomicName lsidName="candezei" pageId="4" pageNumber="89" rank="species" species="candezei">candezei</taxonomicName>
/ Fleut. type/ Collection FLEUTIAUX&quot;; &quot;
<taxonomicName class="Insecta" family="Elateridae" genus="Alaolacon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Alaolacon" order="Coleoptera" pageId="4" pageNumber="89" phylum="Arthropoda" rank="genus">Alaolacon</taxonomicName>
/
<taxonomicName lsidName="candezei" pageId="4" pageNumber="89" rank="species" species="candezei">candezei</taxonomicName>
Fleut./ COLLECTION FLEUTIAUX&quot;;
<normalizedToken originalValue="“Muséum">&quot;Museum</normalizedToken>
paris/ Coll./ E. Fleutiaux&quot;.
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="89" type="diagnosis">
<paragraph pageId="4" pageNumber="89">Diagnosis.</paragraph>
<paragraph pageId="4" pageNumber="89">
Body black, elytra blue and with metallic luster, legs red-black; setae white; frontal depression moderate; eye small; female antennomere III subpectinate, 1.2
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide; prothorax almost as long as wide, widest posteriorly; pronotum with anterior angles bisinuate, median longitudinal depression shallow, not reaching anterior margin or base, punctate; anterior angles of hypomeron rounded; prosternal spine inclined weakly behind procoxae; scutellum concave laterally, widest near posterior 2/5; hind wings without wedge cell, with cross vein between veins MP4 and CuA2 located at contact point between veins MP3 and MP4; female sternite VIII with apex concave.
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="89" type="description">
<paragraph pageId="4" pageNumber="89">Measurements.</paragraph>
<paragraph pageId="4" pageNumber="89">BL: 24.0, BW: 8.35, MIE: 2.56, MAE: 3.47, OI: 74, PL: 7.64, PML: 6.67; PW: 7.70, PI: 99, EL: 15.7, EW: 8.35, EI: 188.</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="89" type="redescription">
<paragraph pageId="4" pageNumber="89">Redescription of female.</paragraph>
<paragraph pageId="4" pageNumber="89">Body (Figs 1, 2) shiny; elytra with weak metallic luster. Color. Body black; mouth-parts brown, mandible black, galea and lacinia orange; elytra black-blue; pronotosternal sutures and legs red-black; tarsal claws yellow-brown. Hairs. Body covered with white flatted setae; antennomere I and legs with intermixed brown and white setae; antennomeres II-XI with brown setae. (Most setae of elytra lost.)</paragraph>
<paragraph pageId="4" pageNumber="89">
Head. Frontal depression moderate (Fig. 4). Eyes small. Antennomere II conical; antennomere III longest, subpectinate, 1.2
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide, 3.0
<normalizedToken originalValue="×">x</normalizedToken>
times as long as II; apical half part of antennomere XI thinner than its basal half part (Fig. 5: dotted line). Apical maxillary palpomere 1.6
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide (Fig. 6).
</paragraph>
<paragraph pageId="4" pageNumber="89">
Prothorax almost as long as wide, widest posteriorly; hind angles straight posteriorly. Pronotum with anterior angle bisinuate; median longitudinal depression shallow, not reaching anterior margin or base, punctate. Hypomeron with anterior angles rounded (Fig. 8). Prosternal spine inclined weakly (at 8 degrees) behind procoxae (Fig. 7). Scutellum (Fig. 10) 1.2
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide, concave laterally, widest near posterior 2/5. Hind wings with cross vein between veins MP4 and CuA2 apparent, not completely connected with CuA2, located at contact point between veins MP3 and MP4 (Fig. 11: arrow); wedge cell absent. Elytra widest on basal half; intervals with uniformly small punctures.
</paragraph>
<paragraph pageId="4" pageNumber="89">
Abdomen. Ventrite V 0.59
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide. Tergite VIII (Fig. 12) truncate apically. Sternite VIII (Fig. 13) widest at apical 1/3, apex concave; spiculum ventrale1.4
<normalizedToken originalValue="×">x</normalizedToken>
longer than sternite VIII.
</paragraph>
<paragraph pageId="4" pageNumber="89">Genitalia (Fig. 14). Ovipositor with coxites not sclerotized at apex (Fig. 15). Bursa copulatrix with three short sacs (Fig. 16: arrows); without sclerotized structures.</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="90" type="male">
<paragraph pageId="5" pageNumber="90">
<pageBreakToken pageId="5" pageNumber="90" start="start">Male</pageBreakToken>
.
</paragraph>
<paragraph pageId="5" pageNumber="90">Unknown.</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="90" type="distribution">
<paragraph pageId="5" pageNumber="90">Distribution.</paragraph>
<paragraph pageId="5" pageNumber="90">Malaysia: Sabah: Banggi Island.</paragraph>
</subSubSection>
<subSubSection lastPageId="6" lastPageNumber="91" pageId="5" pageNumber="90" type="remarks">
<paragraph pageId="5" pageNumber="90">Remarks.</paragraph>
<paragraph lastPageId="6" lastPageNumber="91" pageId="5" pageNumber="90">
This species is similar to
<taxonomicName class="Insecta" family="Elateridae" genus="Alaolacon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Alaolacon cyanipennis" order="Coleoptera" pageId="5" pageNumber="90" phylum="Arthropoda" rank="species" species="cyanipennis">Alaolacon cyanipennis</taxonomicName>
<normalizedToken originalValue="Candèze">Candeze</normalizedToken>
, 1865 in large body size (24.0 mm), black body and elytra with metallic luster, but is distinguished by
<pageBreakToken pageId="6" pageNumber="91" start="start">the</pageBreakToken>
following contrasting characters (
<taxonomicName class="Insecta" family="Elateridae" genus="Alaolacon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Alaolacon cyanipennis" order="Coleoptera" pageId="6" pageNumber="91" phylum="Arthropoda" rank="species" species="cyanipennis">Alaolacon cyanipennis</taxonomicName>
in parentheses): female antennomere III pectinate (Fig. 5) (female antennomere III trapezoidal); prothorax widest posteriorly (Fig. 1) (prothorax widest at mid-length except for posterior angles, Fig. 17); scutellum widest near posterior 2/5 (Fig. 10) (scutellum near posterior 1/3); wedge cell of hind wings absent (Fig. 11) (wedge cell of hind wings present, Fig. 18); female sternite VIII with apex concave (Fig. 13) (female sternite VIII with apex rounded).
</paragraph>
<caption pageId="6" pageNumber="91">
<paragraph pageId="6" pageNumber="91">
Figures 17-19.
<taxonomicName class="Insecta" family="Elateridae" genus="Alaolacon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Alaolacon cyanipennis" order="Coleoptera" pageId="6" pageNumber="91" phylum="Arthropoda" rank="species" species="cyanipennis">Alaolacon cyanipennis</taxonomicName>
<normalizedToken originalValue="Candèze">Candeze</normalizedToken>
, 1865, female, lectotype 17 habitus, dorsal view 18 right hind wing 19 data labels and body parts.
</paragraph>
</caption>
<paragraph pageId="6" pageNumber="91">This species are known only from the female holotype. We predict that the males also exhibit blue elytra and with metallic luster, scutellum widest near posterior 2/5, hind wings without wedge cell and with cross vein between veins MP4 and CuA2 located at contact point between veins MP3 and MP4.</paragraph>
</subSubSection>
</treatment>
</document>