treatments-xml/data/FB/B1/12/FBB11244EB675416B51F61EC4CBB7DE4.xml

<document ID-DOI="http://dx.doi.org/10.3897/mycokeys.103.107935" ID-Pensoft-Pub="1314-4049-103-129" ID-Pensoft-UUID="A42A1B46EC1F5C4AA8A83A1514EC444E" ModsDocID="1314-4049-103-129" checkinTime="1711652827384" checkinUser="pensoft" docAuthor="Vadthanarat, Santhiti, Raghoonundon, Bhavesh, Lumyong, Saisamorn &amp; Raspe, Olivier" docDate="2024" docId="FBB11244EB675416B51F61EC4CBB7DE4" docLanguage="en" docName="MycoKeys 103: 129-165" docOrigin="MycoKeys 103" docPubDate="2024-03-28" docSource="http://dx.doi.org/10.3897/mycokeys.103.107935" docTitle="Hemileccinum inferius Vadthanarat, Raghoonundon &amp; Raspe 2024, sp. nov." docType="treatment" docVersion="1" id="A42A1B46EC1F5C4AA8A83A1514EC444E" lastPageNumber="129" masterDocId="A42A1B46EC1F5C4AA8A83A1514EC444E" masterDocTitle="Rostrupomyces, a new genus to accommodate Xerocomus sisongkhramensis, and a new Hemileccinum species (Xerocomoideae, Boletaceae) from Thailand" masterLastPageNumber="165" masterPageNumber="129" pageNumber="129" updateTime="1711652827384" updateUser="pensoft">
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<mods:title id="C7FDC919BDF699A804D33193502EDF6D">Rostrupomyces, a new genus to accommodate Xerocomus sisongkhramensis, and a new Hemileccinum species (Xerocomoideae, Boletaceae) from Thailand</mods:title>
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<mods:namePart id="D1B29B2C406AEF2C0284BC2C7E6DE057">Vadthanarat, Santhiti</mods:namePart>
<mods:nameIdentifier id="C66121A4ECA470E0CCDCE93E890E2C49" type="ORCID">https://orcid.org/0000-0002-9035-0375</mods:nameIdentifier>
<mods:affiliation id="9BD5EBB5EE2D857D1D8AD9C1160F9C4A">School of Science, Mae Fah Luang University, Chiang Rai, 57100, Thailand &amp; Department of Biological Science, Faculty of Science, Ubon Ratchathani University, Ubon Ratchathani, 34190, Thailand</mods:affiliation>
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<mods:namePart id="3E3B189888876242CB2397F324BB3FF6">Raghoonundon, Bhavesh</mods:namePart>
<mods:nameIdentifier id="2A126834D0672D2B2706AC4415640F26" type="ORCID">https://orcid.org/0000-0001-6671-2404</mods:nameIdentifier>
<mods:affiliation id="42FDD2833640B278FC4FC22288FE65C1">School of Science, Mae Fah Luang University, Chiang Rai, 57100, Thailand</mods:affiliation>
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<mods:namePart id="1C8EE1333E58014830C974CE228B2E6C">Lumyong, Saisamorn</mods:namePart>
<mods:affiliation id="CC033BF635CA3E51E75BB15529B6AC9A">Department of Biology, Faculty of Science, Chiang Mai University, Chiang Mai, 50200, Thailand &amp; Research Center of Microbial Diversity and Sustainable Utilization, Faculty of Science, Chiang Mai University, Chiang Mai, 50200, Thailand</mods:affiliation>
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<mods:namePart id="DF4EAB3E897CA8A7FF2F39F8214722BB">Raspe, Olivier</mods:namePart>
<mods:nameIdentifier id="1A34DA227533F43DEC50C28EA6506E86" type="ORCID">https://orcid.org/0000-0002-8426-2133</mods:nameIdentifier>
<mods:affiliation id="3DF5A69F5277C99FCB7459B9FFCF27F6">School of Science, Mae Fah Luang University, Chiang Rai, 57100, Thailand &amp; Academy of Science, The Royal Society of Thailand, Bangkok, Thailand &amp; Meise Botanic Garden, Nieuwelaan 38, 1860 Meise, Belgium</mods:affiliation>
<mods:nameIdentifier id="49D6AAEA5F4D660312733AE76AB18F1C" type="email">Olivier.Ras@mfu.ac.th</mods:nameIdentifier>
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<mods:date id="2E5D49D20B3AEB4E3A9E7263CAF00335">2024</mods:date>
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<subSection id="62EDCDA1A3260476D79CCC350310B1C9" pageId="0" pageNumber="129" type="results">
<treatment id="FBB11244EB675416B51F61EC4CBB7DE4" LSID="urn:lsid:plazi:treatment:FBB11244EB675416B51F61EC4CBB7DE4" httpUri="http://treatment.plazi.org/id/FBB11244EB675416B51F61EC4CBB7DE4" lastPageNumber="129" pageId="0" pageNumber="129">
<subSubSection id="C4896CAB1523782D8EB13218899E0CC0" pageId="0" pageNumber="129" type="nomenclature">
<paragraph id="FEEDCF3E639FD9999787B4C48693DDBD" pageId="0" pageNumber="129">
<taxonomicName id="29F1CF7FFD639214D8B876140EF3D31D" LSID="FBB11244-EB67-5416-B51F-61EC4CBB7DE4" authority="Vadthanarat, Raghoonundon &amp; Raspé" authorityName="Vadthanarat, Raghoonundon &amp; Raspe" authorityYear="2024" class="Agaricomycetes" family="Boletaceae" genus="Hemileccinum" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Hemileccinum inferius" order="Boletales" pageId="0" pageNumber="129" phylum="Basidiomycota" rank="species" species="inferius" status="sp. nov.">
Hemileccinum inferius Vadthanarat, Raghoonundon &amp; 
<normalizedToken id="7ECE6DD06F0FFDD9BE5B232ADBED5D78" originalValue="Raspé">Raspe</normalizedToken>
</taxonomicName>
<taxonomicNameLabel id="94D430DC554FF88AC829874ED1DBF1E3" pageId="0" pageNumber="129">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="041DF9B312BC87B0797729EBD173E545" pageId="0" pageNumber="129" type="description">
<paragraph id="9B4AADA9BA73D056EB61A128D405563F" pageId="0" pageNumber="129">
<figureCitation id="04D5BB0428380F68E0816D49307E6934" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Fresh basidioma of Hemileccinum inferius A, B SV 0282 (holotype). Scale bars: 1 cm (A, B)." figureDoi="10.3897/mycokeys.103.107935.figure6" httpUri="https://binary.pensoft.net/fig/1011683" pageId="0" pageNumber="129">Figs 6</figureCitation>
<figureCitation id="B952A95D09E23ABAD845DB99E1C474F2" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Microscopic features of Hemileccinum inferius A basidiospores B basidia C cheilocystidia D pleurocystidia E pileipellis F stipitipellis showing a cluster of clavate to boardy clavate like cells which moderately scattered on the stipitipellis. Scale bars: 10 µm (A-D); 25 µm (D-E); 50 µm (E-F). All drawings were made from holotype type (SV 0282)." figureDoi="10.3897/mycokeys.103.107935.figure7" httpUri="https://binary.pensoft.net/fig/1011684" pageId="0" pageNumber="129">, 7</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="AA4D0978D2EE7C459DD7DAC7718300D2" pageId="0" pageNumber="129" type="etymology">
<paragraph id="85327A2456DF5589D5DDA99ADCEA4DE0" pageId="0" pageNumber="129">Etymology.</paragraph>
<paragraph id="D1A0CC119D2B3119F69F702CF7AC9288" pageId="0" pageNumber="129">
&quot; 
<taxonomicName id="C2CDBFF474F4E3DC9C10A042EF01F174" authorityName="Vadthanarat, Raghoonundon &amp; Raspe" authorityYear="2024" class="Agaricomycetes" family="Pinaceae" genus="Hemileccinum" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Hemileccinum inferius" order="Boletales" pageId="0" pageNumber="129" phylum="Basidiomycota" rank="species" species="inferius">Hemileccinum inferius</taxonomicName>
&quot; refers to the only lower part of the stipe ornamented with reticulum
</paragraph>
</subSubSection>
<subSubSection id="0BF5B36B33522A9EBB76A261FBBF117D" pageId="0" pageNumber="129" type="holotype">
<paragraph id="06550D30793DD9320E62BF30692C6EA3" pageId="0" pageNumber="129">Holotype.</paragraph>
<paragraph id="383B3E66593B6B8B962E88DCBE97FE4E" pageId="0" pageNumber="129">
Thailand, Chiang Mai Province, Muang District, Doi Suthep-Pui National Park, 
<geoCoordinate id="E2B8E22E39C30DFA53D39A3E4288FB54" degrees="18" direction="north" minutes="47" orientation="latitude" precision="1" seconds="52.8" value="18.798">18°47'52.8&quot;N</geoCoordinate>
, 
<geoCoordinate id="6A6A25629DC8F5A127A86915B8F74EDF" degrees="98" direction="east" minutes="54" orientation="longitude" precision="1" seconds="21.2" value="98.90589">98°54'21.2&quot;E</geoCoordinate>
, elev. 1,170 m, 1 July 2016, 
<emphasis id="49918E63F2E33805F65E5B3F94C56A8C" italics="true" pageId="0" pageNumber="129">Santhiti Vadthanarat</emphasis>
, SV0282 (holotype: CMUB, isotype: BKF, MFUB). ITS sequence accession number PP354892.
</paragraph>
</subSubSection>
<subSubSection id="EC178493AD7867166F058F22554B9F15" pageId="0" pageNumber="129" type="diagnosis">
<paragraph id="E8D768F54D44CDBA645AF6CA277780A0" pageId="0" pageNumber="129">Diagnosis.</paragraph>
<paragraph id="EE79BE0A0EE91F4F5D877DED67A28380" pageId="0" pageNumber="129">
<taxonomicName id="B2FB4E667B91E0FA19E3EC1B0B73DB7E" authorityName="Vadthanarat, Raghoonundon &amp; Raspe" authorityYear="2024" class="Agaricomycetes" family="Pinaceae" genus="Hemileccinum" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Hemileccinum inferius" order="Boletales" pageId="0" pageNumber="129" phylum="Basidiomycota" rank="species" species="inferius">
<emphasis id="68B9C4956B9168C05BF63558AD3C5DCC" italics="true" pageId="0" pageNumber="129">Hemileccinum inferius</emphasis>
</taxonomicName>
can be differentiated from resembling 
<taxonomicName id="F38DC160F6A458E00E90E91E325E9F3F" class="Agaricomycetes" family="Pinaceae" genus="Hemileccinum" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Hemileccinum" order="Boletales" pageId="0" pageNumber="129" phylum="Basidiomycota" rank="genus">
<emphasis id="87E63747ED45E2DF166E91F68C67EB49" italics="true" pageId="0" pageNumber="129">Hemileccinum</emphasis>
</taxonomicName>
species by a grayish red to reddish brown to dark brown, plane to sub-depressed, subrugulose to pitted pileus; and yellow to yellowish white, cylindrical with subbulbous stipe, with surface even on the upper half and subscabrous to delicately reticulate on the lower half, as well as smooth basidiospores even when observed under SEM.
</paragraph>
</subSubSection>
<subSubSection id="1F6F6081009EB1B6C18EFC54E19F36B1" pageId="0" pageNumber="129" type="description">
<paragraph id="1A68C9CCA5D9A7393AD0059FD5ABE84F" pageId="0" pageNumber="129">Description.</paragraph>
<paragraph id="DFE5D54535EFF7CB04D78AF09005B5FB" pageId="0" pageNumber="129">
<emphasis id="3497D54B93C4FBC7DEA73BD70FB38065" bold="true" italics="true" pageId="0" pageNumber="129">Basidiomata</emphasis>
medium-sized. 
<emphasis id="9A16C2E623D156A8AB4A2D953CE9A5C5" bold="true" italics="true" pageId="0" pageNumber="129">Pileus</emphasis>
66-68 mm in diameter, plane to sub-depressed at the centre; 
<emphasis id="A92D9B33BF0E531B492D7EC13DC53D8F" bold="true" italics="true" pageId="0" pageNumber="129">margin</emphasis>
deflexed in age, elastic, slightly exceeding (up to 1 mm); 
<emphasis id="55F78729BE147832E6796E5B6FD92775" bold="true" italics="true" pageId="0" pageNumber="129">surface</emphasis>
subrugulose to pitted especially near the margin, dull, moist to slippery when wet, tomentose, grayish red (8B/C3-4) to reddish brown (8D/E4-6) to dark brown to reddish brown (7-8F4-6), unchanging when bruised; 
<emphasis id="FF6E6FD13BC6A3144F75E63216FEA262" bold="true" italics="true" pageId="0" pageNumber="129">context</emphasis>
8-10 mm thick haft-way to the margin, firm to soft, pale yellow (1A3), slightly brown (7E5) near the pileus surface, light yellow (1A4) above the hymenium in age, unchanging when cut. 
<emphasis id="DDEA2C5B60AAC04043B21B35B45F6C67" bold="true" italics="true" pageId="0" pageNumber="129">Stipe</emphasis>
65-76 
<normalizedToken id="E8CF9C9D84AB1510FA7D8EA69E578941" originalValue="×">x</normalizedToken>
14-18 mm, central, terete, cylindrical with subbulbous base; 
<emphasis id="50159F9A73D79CD09CDC2F70684DF6EF" bold="true" italics="true" pageId="0" pageNumber="129">surface</emphasis>
even on the upper half then subscabrous to delicately reticulate on the lower half, dull, dry to moist, light yellow (2A4-6) to yellowish white to pale yellow (2A2-3) at the base, occasionally with reddish brown to dark brown spots (8D5-8, 8F7) at places, minutely covered with pale yellow to light brown to dark brown (2A3-4 to 7D/E4, 7F8) squamules on the upper half, slightly fibrillose following a reticulate pattern at the middle of the stipe getting less so to the base, unchanging when bruised; 
<emphasis id="8B5165DA3E72D8CE1C4D86AABBED482D" bold="true" italics="true" pageId="0" pageNumber="129">basal mycelium</emphasis>
white (1A1); 
<emphasis id="7496F4288EC34379F447BEA56FB862DC" bold="true" italics="true" pageId="0" pageNumber="129">context</emphasis>
solid, firm, pale yellow (2A3-5) especially in the above half near the stipe surface becoming yellowish white (2A2) to off-white at the base, unchanging when cut. 
<emphasis id="A6BE49F6E7DC2A9F61EE7C062BD44D37" bold="true" italics="true" pageId="0" pageNumber="129">Hymenophore</emphasis>
tubulate, slightly depressed around the stipe, with slightly decurrent tooth, subventricose. 
<emphasis id="EB489621AAD865F5ECEC7CDF2F37103A" bold="true" italics="true" pageId="0" pageNumber="129">Tubes</emphasis>
7-8 mm long half-way to the margin, yellow to grayish yellow (2A7 to 2B7) near the pileus context then olive (2E5) near the pores, separable from the pileus context, unchanging when bruised. 
<emphasis id="AA95705E46A7276E21D0A1DD5AA74826" bold="true" italics="true" pageId="0" pageNumber="129">Pores</emphasis>
0.3-0.8(1.2) mm wide at mid-radius, subangular to angular, even, grayish yellow (2B5), unchanging when touched, irregularly arranged; topography subregular. 
<emphasis id="188D880F3B72D99EDC1DDF18348B1F4E" bold="true" italics="true" pageId="0" pageNumber="129">Odour</emphasis>
mild fungoid. 
<emphasis id="22C1E977FE95951E750E458CC28ECC1D" bold="true" italics="true" pageId="0" pageNumber="129">Taste</emphasis>
mild. 
<emphasis id="64FE3A4E0C2CF5F06C119632561AEB6B" bold="true" italics="true" pageId="0" pageNumber="129">Spore print</emphasis>
olive brown (4E7).
</paragraph>
<paragraph id="A9548EC6B32C59EF5DEE0FBDA8A35011" pageId="0" pageNumber="129">
<emphasis id="770B8E0302361FFD3AA4C69765E96DF4" bold="true" italics="true" pageId="0" pageNumber="129">Macrochemical reactions</emphasis>
: KOH, brownish orange on pileus and hymenophore, pale orange on pileus context and stipe surface, and stipe context; NH4OH, brownish orange with purple aura on pileus, yellowish to brownish orange with purple aura on stipe surface, yellowish to greenish or slightly blue on pileus context and stipe context.
</paragraph>
<caption id="80372C4FB26A49AE122921E5794A5D1F" doi="10.3897/mycokeys.103.107935.figure6" httpUri="https://binary.pensoft.net/fig/1011683" pageId="0" pageNumber="129" start="Figure 6" startId="F6">
<paragraph id="6CCEF97457EB7DFF9F5286AC049027C6" pageId="0" pageNumber="129">
<emphasis id="749A1BA0F754121A59D2BA064E3068E4" bold="true" pageId="0" pageNumber="129">Figure 6.</emphasis>
Fresh basidioma of 
<taxonomicName id="9F8DFC35E3DFC7B76AE61714F2E270A6" authorityName="Vadthanarat, Raghoonundon &amp; Raspe" authorityYear="2024" class="Agaricomycetes" family="Pinaceae" genus="Hemileccinum" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Hemileccinum inferius" order="Boletales" pageId="0" pageNumber="129" phylum="Basidiomycota" rank="species" species="inferius">
<emphasis id="D2CFEC6CEC21BE560354B135D2D3232C" italics="true" pageId="0" pageNumber="129">Hemileccinum inferius</emphasis>
</taxonomicName>
<emphasis id="FFEA82ED6EF08B51CAA610894FAA460D" bold="true" pageId="0" pageNumber="129">A, B</emphasis>
SV0282 (holotype). Scale bars: 1 cm (
<emphasis id="9C37D5A920C46F882E04E2FFE53FAC4C" bold="true" pageId="0" pageNumber="129">A, B</emphasis>
).
</paragraph>
</caption>
<paragraph id="52B523E5E09373206D9C90016D811EA7" pageId="0" pageNumber="129">
<emphasis id="9A22721D55C065273539F33DF800B7BE" bold="true" italics="true" pageId="0" pageNumber="129">Spores</emphasis>
[118/2/2] (10.5-)11.4-12.9-14.6(-15.3) 
<normalizedToken id="716AE2E77ED18E9266B287B07476D566" originalValue="×">x</normalizedToken>
(3.8-)4.2-4.8-5.6(-6.1) 
<normalizedToken id="228648395F64B9BBFC1F31E9929B79BF" originalValue="µm">µm</normalizedToken>
<emphasis id="AD44B3DB28BBF86BC6C7A4A7B570E00E" italics="true" pageId="0" pageNumber="129">Q</emphasis>
= (2.06-)2.4-2.68-3.05(-3.32). From the type (10.8-)11.5-12.7-14.2(-14.5) 
<normalizedToken id="B38F7B3CD9949C45C5661DD4E687B118" originalValue="×">x</normalizedToken>
(4.1-)4.3-4.8-5.5(-6.1) 
<normalizedToken id="74836A6065E5EA201649F29DEECD7200" originalValue="µm">µm</normalizedToken>
, 
<emphasis id="164624E6866A3CF414285276537FCEFA" italics="true" pageId="0" pageNumber="129">Q</emphasis>
= (2.06-)2.33-2.66-3.06(-3.1), 
<emphasis id="C7A857DF2BADB3427D68E24C3CDFE4D7" italics="true" pageId="0" pageNumber="129">N</emphasis>
= 68, narrowly ellipsoid to subcylindrical with a slight suprahilar depression, thin-walled, smooth under light microscope and SEM (Fig. 
<figureCitation id="4D19A78ACB19246E44EDF6FABF97EE1F" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Scanning electron micrographs of basidiospores A-B Rostrupomyces sisongkhramensis (SV 0155) C-D Hemileccinum inferius (SV 0282)." figureDoi="10.3897/mycokeys.103.107935.figure5" httpUri="https://binary.pensoft.net/fig/1011682" pageId="0" pageNumber="129">5C-D</figureCitation>
), yellowish to brownish hyaline in water, yellowish hyaline in KOH, inamyloid. 
<emphasis id="888AE82E29204647CD1AC5E527E0B985" bold="true" italics="true" pageId="0" pageNumber="129">Basidia</emphasis>
4-spored, (23-)24-27-31(-32) 
<normalizedToken id="D4045DEE1AFB6815C8D21E0364D31A41" originalValue="×">x</normalizedToken>
(11-)11-12-14(-14) 
<normalizedToken id="9119AD687C544759E6D39747A5B9C191" originalValue="µm">µm</normalizedToken>
, clavate without basal clamp connection, hyaline to yellowish hyaline in KOH; sterigmata up to 4 
<normalizedToken id="57BF311C4B37CAAE0783C210300D9D48" originalValue="µm">µm</normalizedToken>
long. 
<emphasis id="0681AD46AC403937848D8FB2EFA89D95" bold="true" italics="true" pageId="0" pageNumber="129">Cheilocystidia</emphasis>
(30-)34-54-72(-72) 
<normalizedToken id="82B40B16F34A2A1218485C9BAD4D8738" originalValue="×">x</normalizedToken>
(7-)8-10-14(-14) 
<normalizedToken id="E02EE4C62187962F5EE8CAA807B129E5" originalValue="µm">µm</normalizedToken>
, narrowly fusiform with elongated obtuse apex, frequent, thin-walled, hyaline to yellowish hyaline in KOH. 
<emphasis id="B6A02B7FF104E3E9C6C68470A7BD019A" bold="true" italics="true" pageId="0" pageNumber="129">Pleurocystidia</emphasis>
(34-)34-51-69(-70) 
<normalizedToken id="A2584900718D70F142CAFCE7FDB98CCD" originalValue="×">x</normalizedToken>
(10-)10-11-13(-13) 
<normalizedToken id="17D15441E2924D01D1E8AAF17E6FBB73" originalValue="µm">µm</normalizedToken>
, frequent near the pores, narrowly fusiform with elongated obtuse apex, thin-walled, hyaline to yellowish hyaline in KOH. 
<emphasis id="9581ED466964D8CEF1F2827E0B00F202" bold="true" italics="true" pageId="0" pageNumber="129">Hymenophoral trama</emphasis>
slightly divergent, 62-150 
<normalizedToken id="638D8F4CF2530A1205A5940EDA680A54" originalValue="µm">µm</normalizedToken>
wide composed of cylindrical, 4-12 
<normalizedToken id="2EC16A8ED1DF702C82B1013A7EE90D13" originalValue="µm">µm</normalizedToken>
wide hyphae, with subregular mediostratum 30-100 
<normalizedToken id="3AC77AE1251E0078EE97D13D81C3A961" originalValue="µm">µm</normalizedToken>
wide, hyaline in KOH. 
<emphasis id="F74F9B4344A929F228765F1F1809009F" bold="true" italics="true" pageId="0" pageNumber="129">Pileipellis</emphasis>
a hyphoepithelium, 80-112 
<normalizedToken id="CC2B4A9723564E2AFCBBEDD8BDC5D28F" originalValue="μm">μm</normalizedToken>
thick, the pileipellis composed of ellipsoid to broadly ellipsoid or cylindrical, thin-walled, more or less vertically arranged, occasionally branching or anastomosing, with metablematic, elongated-cylindrical hyphae (2-4 
<normalizedToken id="23394BEDBC2D168C03CD2732A0D461B9" originalValue="µm">µm</normalizedToken>
wide hyphae), branching or anastomosing at places, hyaline to yellowish hyaline in KOH; terminal cells of 2 types: 1) ellipsoid to broadly ellipsoid, 8-15 
<normalizedToken id="99C83C4999DB7B4E4400192A16152EFF" originalValue="×">x</normalizedToken>
12-20 
<normalizedToken id="892211DC7682369D0592EC3027B1083D" originalValue="µm">µm</normalizedToken>
; and 2) clavate to broadly clavate with obtuse apex, 10-20 
<normalizedToken id="CE395F9C3FFC5C9F5E6279BA0D146233" originalValue="×">x</normalizedToken>
4-7 
<normalizedToken id="9D8CF5A8F275425FBCB39A955BC7A914" originalValue="µm">µm</normalizedToken>
. 
<emphasis id="A3BEA360A5CF317E712A5CE2B9D3E5F8" bold="true" italics="true" pageId="0" pageNumber="129">Pileus context</emphasis>
made of moderately interwoven, ellipsoid to broadly ellipsoid, thin-walled hyphae, 10-23 
<normalizedToken id="5B6A667AE7B46E3F4C4D2B86FA31E5E3" originalValue="µm">µm</normalizedToken>
wide, hyaline in KOH. 
<emphasis id="22F7EFAF75FBC6C9A3CB4C2D39BE535A" bold="true" italics="true" pageId="0" pageNumber="129">Stipitipellis</emphasis>
arranged parallel to the surface of the stipe (40-50 
<normalizedToken id="5880039595E40A79EA9CA31544EB0620" originalValue="µm">µm</normalizedToken>
thick), composed of moderately interwoven, cylindrical, thin-walled, 2.5-4 
<normalizedToken id="A5D8D6855E3F2E56A589E96B907173A2" originalValue="µm">µm</normalizedToken>
wide hyphae, anastomosing at places, moderately scattered with groups of rising cells to clusters (50-60 
<normalizedToken id="AEAD155018F0770CB01E8197D24FD2AE" originalValue="µm">µm</normalizedToken>
high) of thin-walled clavate to broadly clavate cells (20-30 
<normalizedToken id="E9D493CC8105D57E4F3EF9B6C62CAC94" originalValue="×">x</normalizedToken>
10-15 
<normalizedToken id="BF83BD6980522A447FF305B8543E8571" originalValue="µm">µm</normalizedToken>
), hyaline to yellowish hyaline in KOH. 
<emphasis id="607C6E41000108DD96A026F277EBAEB1" bold="true" italics="true" pageId="0" pageNumber="129">Caulocystidia</emphasis>
not seen. 
<emphasis id="08FC6AD3C2072790425A387B26E1B11E" bold="true" italics="true" pageId="0" pageNumber="129">Stipe context</emphasis>
composed of parallel, 8-22 
<normalizedToken id="B3BDB55BA09645B6002F9ED858125BE5" originalValue="µm">µm</normalizedToken>
wide hyphae, hyaline in KOH. 
<emphasis id="4819FB682236D414D11B2269878E215B" bold="true" italics="true" pageId="0" pageNumber="129">Clamp connections</emphasis>
not seen in any tissue.
</paragraph>
<caption id="E11AC9154693ED4CDF671D89E3392C60" doi="10.3897/mycokeys.103.107935.figure7" httpUri="https://binary.pensoft.net/fig/1011684" pageId="0" pageNumber="129" start="Figure 7" startId="F7">
<paragraph id="CB2C00C2F37C4214CDBF4FCE629C22D8" pageId="0" pageNumber="129">
<emphasis id="2DF07BC4F76AE250E84DA76B3C15D2A7" bold="true" pageId="0" pageNumber="129">Figure 7.</emphasis>
Microscopic features of 
<taxonomicName id="362E38B1CA1E3FC004A58E16FE5E2B22" authorityName="Vadthanarat, Raghoonundon &amp; Raspe" authorityYear="2024" class="Agaricomycetes" family="Pinaceae" genus="Hemileccinum" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Hemileccinum inferius" order="Boletales" pageId="0" pageNumber="129" phylum="Basidiomycota" rank="species" species="inferius">
<emphasis id="082A1316665587FBA8086AA7D92E644D" italics="true" pageId="0" pageNumber="129">Hemileccinum inferius</emphasis>
</taxonomicName>
<emphasis id="4C2836B04658D96727383D5530FBD747" bold="true" pageId="0" pageNumber="129">A</emphasis>
basidiospores 
<emphasis id="96E94719A744898B1CCC19AE53EE5F55" bold="true" pageId="0" pageNumber="129">B</emphasis>
basidia 
<emphasis id="085787C5BF9E78DE907FE04DE6578DEA" bold="true" pageId="0" pageNumber="129">C</emphasis>
cheilocystidia 
<emphasis id="5B4FB5DB8B34AC4E5A648A2BC93595EC" bold="true" pageId="0" pageNumber="129">D</emphasis>
pleurocystidia 
<emphasis id="46FC6966E5777C7C068EFC47CE27D49F" bold="true" pageId="0" pageNumber="129">E</emphasis>
pileipellis 
<emphasis id="8CF0E0DBFF50B4F21217E69CD48BF19E" bold="true" pageId="0" pageNumber="129">F</emphasis>
stipitipellis showing a cluster of clavate to boardy clavate like cells which moderately scattered on the stipitipellis. Scale bars: 10 
<normalizedToken id="00C15540944425EBF1F1FFF027B2D9BF" originalValue="µm">µm</normalizedToken>
(
<emphasis id="8275F56809ECE59741D82E385839A82B" bold="true" pageId="0" pageNumber="129">A-D</emphasis>
); 25 
<normalizedToken id="1023E4B1DEBFD8D7882C5D40648C8D0E" originalValue="µm">µm</normalizedToken>
(
<emphasis id="ABD31A61862E94822A114908E34E5902" bold="true" pageId="0" pageNumber="129">D-E</emphasis>
); 50 
<normalizedToken id="AC7D8E4A20FA3D020BF2C45CB0BA590E" originalValue="µm">µm</normalizedToken>
(
<emphasis id="6B70746D25C6D29EDDA58004E5845C64" bold="true" pageId="0" pageNumber="129">E-F</emphasis>
). All drawings were made from holotype type (SV0282).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="59E9475F102AD1E1429722D66CFFAC0C" pageId="0" pageNumber="129" type="habitat">
<paragraph id="97A333D3AAE07FF1B8EA562A1DACE53A" pageId="0" pageNumber="129">Habitat and distribution.</paragraph>
<paragraph id="4B64FFF1B9E1D482981C4C318D82D314" pageId="0" pageNumber="129">
Solitary, on loamy soil in hill evergreen forest dominated by 
<taxonomicName id="8CF8F104CA0676C486D8651ADD4CA9FD" family="Fagaceae" kingdom="Fungi" lsidName="" order="Boletales" pageId="0" pageNumber="129" rank="family">Fagaceae</taxonomicName>
scattered with a few 
<taxonomicName id="2C1D89319E927D3FD9D9862EEE9462FA" class="Dicotyledoneae" family="Dipterocarpaceae" genus="Dipterocarpus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Dipterocarpus obtusifolius" order="Guttiferales" pageId="0" pageNumber="129" phylum="Angiospermae" rank="species" species="obtusifolius">
<emphasis id="2E2CFB2F397272FDE426DF269A21FBEA" italics="true" pageId="0" pageNumber="129">Dipterocarpus obtusifolius</emphasis>
</taxonomicName>
, at 985-1,170 m elevation. Currently known from Chiang Mai Province, northern Thailand.
</paragraph>
</subSubSection>
<subSubSection id="6D271AE24FFD554557CBBB9C999B6402" pageId="0" pageNumber="129" type="materials_examined">
<paragraph id="53980506141FAD5A05C0E97B76F76978" pageId="0" pageNumber="129">Additional specimens examined.</paragraph>
<paragraph id="EA772463C4FA3BEA9C5942C1C0377AF2" pageId="0" pageNumber="129">
<materialsCitation id="B0DE89069E6DC99530F9F72E6A19360E" collectingDate="2021-06-06" collectorName="Bhavesh Raghoonundon, BR" country="Thailand" elevation="985" latitude="19.116388" location="Mae Taeng District" longLatPrecision="21" longitude="98.739716" specimenCount="1" stateProvince="Chiang Mai Province">
<collectingCountry id="CE6DFEB7DF494BEA3302A4CB1B3A7ACB" name="Thailand">Thailand</collectingCountry>
, 
<collectingRegion id="BEA9DD73F9E3BFC5CC791EDC73F88787" country="Thailand" name="Chiang Mai">Chiang Mai Province</collectingRegion>
, 
<location id="F4210FCBDF16EA4759C73483EF5FC209" LSID="urn:lsid:plazi:treatment:FBB11244EB675416B51F61EC4CBB7DE4:F4210FCBDF16EA4759C73483EF5FC209" country="Thailand" latitude="19.116388" longLatPrecision="21" longitude="98.739716" name="Mae Taeng District" stateProvince="Chiang Mai Province">Mae Taeng District</location>
, 
<geoCoordinate id="EEF96126DFA7BE6762C5905008F79CE4" degrees="19" direction="north" minutes="06" orientation="latitude" precision="15" seconds="59" value="19.116388">19°06'59&quot;N</geoCoordinate>
, 
<geoCoordinate id="AF2ACE1080906BE2B2EA15587A07E742" degrees="98" direction="east" minutes="44" orientation="longitude" precision="15" seconds="23" value="98.739716">98°44'23&quot;E</geoCoordinate>
, elev. 
<quantity id="3112AFC73E7BEA7FA447BDA92A163E93" metricMagnitude="2" metricUnit="m" metricValue="9.85" unit="m" value="985.0">
<elevation id="6C50198091A5175CFF451884764F5004" metricMagnitude="2" metricUnit="m" metricValue="9.85" unit="m" value="985.0">985 m</elevation>
</quantity>
, 
<collectingDate id="176D22F5CDDA4D59C81097CFFB8CB449" value="2021-06-06">6 June 2021</collectingDate>
, 
<collectorName id="C6779C5B95745CEFD689FEC363BCF372">
<emphasis id="262132F386F406056829F6E6FD0A948E" italics="true" pageId="0" pageNumber="129">Bhavesh Raghoonundon</emphasis>
, BR
</collectorName>
0260 (MFLU)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="0FD8B23139F522956CF9844AFE6A3FDA" pageId="0" pageNumber="129" type="notes">
<paragraph id="6130DDCF2A84E0EC091CD5ACAB8F2E3D" pageId="0" pageNumber="129">Notes.</paragraph>
<paragraph id="A0468F5B7DC6923C80059213EB78B466" pageId="0" pageNumber="129">
<taxonomicName id="E92FFBF7CC0B6C4981E5A546B68EDF7B" authorityName="Vadthanarat, Raghoonundon &amp; Raspe" authorityYear="2024" class="Agaricomycetes" family="Pinaceae" genus="Hemileccinum" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Hemileccinum inferius" order="Boletales" pageId="0" pageNumber="129" phylum="Basidiomycota" rank="species" species="inferius">
<emphasis id="5BC4C668549F0B5530F14BD29BA2D334" italics="true" pageId="0" pageNumber="129">Hemileccinum inferius</emphasis>
</taxonomicName>
is described based on collections from Thailand. The comparison of the new species with the seven known Asian species follows. 
<taxonomicName id="D12DAFC084DF3ED2CBA989517284BA61" class="Agaricomycetes" family="Pinaceae" genus="Hemileccinum" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Hemileccinum albidum" order="Boletales" pageId="0" pageNumber="129" phylum="Basidiomycota" rank="species" species="albidum">
<emphasis id="658426AC51D4F1610979241CD9371481" italics="true" pageId="0" pageNumber="129">Hemileccinum albidum</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="0E86ABBB046E680766B4124CC38244C8" class="Agaricomycetes" family="Boletaceae" kingdom="Fungi" lsidName="H. inferius" order="Boletales" pageId="0" pageNumber="129" phylum="Basidiomycota" rank="species" species="inferius">
<emphasis id="3049B7ED63B32A98E60D80F471976D42" italics="true" pageId="0" pageNumber="129">H. inferius</emphasis>
</taxonomicName>
by gray-brown to chrome yellow to ochraceous or golden brown pileus; longer and slender stipe (up to 160 mm); shorter basidiospores (10-12.5 
<normalizedToken id="58E6C92074AEF78B0FAFA751DC6FEED4" originalValue="×">x</normalizedToken>
4.0-5.5 
<normalizedToken id="F69427BE8DB45A5BD574342E00BE85BE" originalValue="µm">µm</normalizedToken>
); occurrence at higher elevations (1,968-2,490 m; 
<bibRefCitation id="1C180E834A5CF0DE26344C82F8E294CE" DOI="https://doi.org/10.3390/jof7100823" author="Li, MX" journalOrPublisher="Journal of Fungi" pageId="0" pageNumber="129" pagination=": 823" refId="B40" refString="Li, MX, Wu, G, Yang, ZL, 2021. Four new species of Hemileccinum (Xerocomoideae, Boletaceae) from Southwestern China. Journal of Fungi 7 (10): 823, DOI: https://doi.org/10.3390/jof7100823" title="Four new species of Hemileccinum (Xerocomoideae, Boletaceae) from Southwestern China." url="https://doi.org/10.3390/jof7100823" volume="7" year="2021">Li et al. 2021</bibRefCitation>
). 
<taxonomicName id="7BA696116022F67E3BB6FDBA9B8CDA57" class="Agaricomycetes" family="Pinaceae" genus="Hemileccinum" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Hemileccinum brevisporum" order="Boletales" pageId="0" pageNumber="129" phylum="Basidiomycota" rank="species" species="brevisporum">
<emphasis id="6601867CBC2BE4A7924F06D46A75AFB7" italics="true" pageId="0" pageNumber="129">Hemileccinum brevisporum</emphasis>
</taxonomicName>
is similar in pileus color but has shorter basidiospores (9-11 
<normalizedToken id="C44B40060E2AA1D0C4B5C08AC6013C0D" originalValue="×">x</normalizedToken>
4-5 
<normalizedToken id="CE9CA9A79762E53A4AE33EBBC587801D" originalValue="μm">μm</normalizedToken>
); and it occurs under 
<taxonomicName id="0059AFA35FFE96CFCB235B069E925491" family="Fagaceae" kingdom="Fungi" lsidName="" order="Boletales" pageId="0" pageNumber="129" rank="family">Fagaceae</taxonomicName>
and 
<taxonomicName id="1A72E6FAFAF28B1E076508B819267766" family="Pinaceae" kingdom="Fungi" lsidName="" order="Boletales" pageId="0" pageNumber="129" rank="family">Pinaceae</taxonomicName>
, at higher elevations (1,700-2,120 m); 
<bibRefCitation id="8D32897A5C3C1D69F2C20242A8F451A9" DOI="https://doi.org/10.3390/jof7100823" author="Li, MX" journalOrPublisher="Journal of Fungi" pageId="0" pageNumber="129" pagination=": 823" refId="B40" refString="Li, MX, Wu, G, Yang, ZL, 2021. Four new species of Hemileccinum (Xerocomoideae, Boletaceae) from Southwestern China. Journal of Fungi 7 (10): 823, DOI: https://doi.org/10.3390/jof7100823" title="Four new species of Hemileccinum (Xerocomoideae, Boletaceae) from Southwestern China." url="https://doi.org/10.3390/jof7100823" volume="7" year="2021">Li et al. 2021</bibRefCitation>
). 
<taxonomicName id="887E45058084837294EE0574FD852145" class="Agaricomycetes" family="Pinaceae" genus="Hemileccinum" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Hemileccinum duriusculum" order="Boletales" pageId="0" pageNumber="129" phylum="Basidiomycota" rank="species" species="duriusculum">
<emphasis id="28654BA07A50215091B721B756F79539" italics="true" pageId="0" pageNumber="129">Hemileccinum duriusculum</emphasis>
</taxonomicName>
is macromorphologically quite similar, but differs by a strongly venose pileus surface, even when young, the absence of reticulum on the lower half of the stipe, as well as shorter cheilo- and pleurocystidia (
<bibRefCitation id="6950547A646905C70359D2B698FDFE5C" DOI="https://doi.org/10.1007/s13225-023-00529-0" author="Liu, SL" journalOrPublisher="Fungal Diversity" pageId="0" pageNumber="129" pagination="1 - 216" refId="B41" refString="Liu, SL, Wang, XW, Li, GJ, Deng, CY, Rossi, W, Leonardi, M, Liimatainen, K, Kekki, T, Niskanen, T, Smith, ME, Ammirati, J, Bojantchev, D, Abdel-Wahab, MA, Zhang, M, Tian, E, Lu, Y-Z, Zhang, J-Y, Ma, J, Dutta, AK, Acharya, K, Du, T-Y, Xu, J, Kim, JS, Lim, YW, Gerlach, A, Zeng, N-K, Han, Y-X, Razaghi, P, Raza, M, Cai, L, Calabon, MS, Jones, EBG, Saha, R, Kumar, TKA, Krishnapriya, K, Thomas, A, Kaliyaperumal, M, Kezo, K, Gunaseelan, S, Singh, SK, Singh, PN, Lagashetti, AC, Pawar, KS, Jiang, S, Zhang, C, Zhang, H, Qing, Y, Bau, T, Peng, X-C, Wen, T-C, Ramirez, NA, Niveiro, N, Li, M-X, Yang, ZL, Wu, G, Tarafder, E, Tennakoon, DS, Kuo, C-H, da Silva, TM, Souza-Motta, CM, Bezerra, JDP, He, G, Ji, X-H, Suwannarach, N, Kumla, J, Lumyong, S, Wannathes, N, Rana, S, Hyde, KD, Zhou, L-W, 2024. Fungal diversity notes 1717-1817: Taxonomic and phylogenetic contributions on genera and species of fungal taxa. Fungal Diversity 124 (1): 1 - 216, DOI: https://doi.org/10.1007/s13225-023-00529-0" title="Fungal diversity notes 1717 - 1817: Taxonomic and phylogenetic contributions on genera and species of fungal taxa." url="https://doi.org/10.1007/s13225-023-00529-0" volume="124" year="2024">Liu et al. 2024</bibRefCitation>
). 
<taxonomicName id="5442EBA9857FEE7E0E8E3AB608FCE93D" class="Agaricomycetes" family="Pinaceae" genus="Hemileccinum" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Hemileccinum ferrugineipes" order="Boletales" pageId="0" pageNumber="129" phylum="Basidiomycota" rank="species" species="ferrugineipes">
<emphasis id="6667698A85AD8B9B9FD5BD61F374A1B2" italics="true" pageId="0" pageNumber="129">Hemileccinum ferrugineipes</emphasis>
</taxonomicName>
has similar pileus surface and color but can be differentiated by the apparent pale red-brown color on the lower part of the stipe; and shorter basidiospores (11.0-12.5 
<normalizedToken id="1AFD62F4A6E351F4A9C9E8C48F57A32E" originalValue="×">x</normalizedToken>
4-5 
<normalizedToken id="334146583BDACCF605A20B385E2B3EB6" originalValue="μm">μm</normalizedToken>
; 
<bibRefCitation id="FD176AB6EFC69CA9A7445DC78E5E8CE2" DOI="https://doi.org/10.3390/jof7100823" author="Li, MX" journalOrPublisher="Journal of Fungi" pageId="0" pageNumber="129" pagination=": 823" refId="B40" refString="Li, MX, Wu, G, Yang, ZL, 2021. Four new species of Hemileccinum (Xerocomoideae, Boletaceae) from Southwestern China. Journal of Fungi 7 (10): 823, DOI: https://doi.org/10.3390/jof7100823" title="Four new species of Hemileccinum (Xerocomoideae, Boletaceae) from Southwestern China." url="https://doi.org/10.3390/jof7100823" volume="7" year="2021">Li et al. 2021</bibRefCitation>
). 
<taxonomicName id="2F9FA066A6A43982664391B0C4FA478E" class="Agaricomycetes" family="Pinaceae" genus="Hemileccinum" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Hemileccinum indecorum" order="Boletales" pageId="0" pageNumber="129" phylum="Basidiomycota" rank="species" species="indecorum">
<emphasis id="25033A188439314190FDB91BAADE12CE" italics="true" pageId="0" pageNumber="129">Hemileccinum indecorum</emphasis>
</taxonomicName>
is clearly different in having dark red to reddish brown basidiomata with mucilaginous surface densely covered with whitish to dirty white, small conical to subconical to irregularly shaped squamules; incurved margin; and yellowish hymenophore that slowly turns brownish to reddish brown when bruised (
<bibRefCitation id="8B7FF686BE667A0A464FC1A99CFF8CD6" author="Horak, E" journalOrPublisher="Forest Research Institute and Ministry of Natural Resources and Environment, Malaysia" pageId="0" pageNumber="129" refId="B29" refString="Horak, E, 2011. Revision of Malaysian species of Boletales s.l. (Basidiomycota) described by Corner EJH (1972, 1974). Forest Research Institute and Ministry of Natural Resources and Environment, Malaysia" title="Revision of Malaysian species of Boletales s. l. (Basidiomycota) described by Corner EJH (1972, 1974)." year="2011">Horak 2011</bibRefCitation>
; 
<bibRefCitation id="0A960278E187B3C43B109D774799576C" DOI="https://doi.org/10.3852/11-326" author="Zeng, NK" journalOrPublisher="Mycotaxon" pageId="0" pageNumber="129" refId="B76" refString="Zeng, NK, Cai, Q, Yang, ZL, 2012. Corneroboletus, a new genus to accommodate the southeastern Asian Boletus indecorus. Mycologia 104(6): 1420-1432. https://doi.org/10.3852/11-326" title="Corneroboletus, a new genus to accommodate the southeastern Asian Boletus indecorus. Mycologia 104 (6): 1420 - 1432." url="https://doi.org/10.3852/11-326" year="2012">Zeng et al. 2012</bibRefCitation>
). 
<taxonomicName id="1D35A826EC43EF5FDED854B9325DAB75" class="Agaricomycetes" family="Pinaceae" genus="Hemileccinum" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Hemileccinum parvum" order="Boletales" pageId="0" pageNumber="129" phylum="Basidiomycota" rank="species" species="parvum">
<emphasis id="2CECFE4B72373D0A6185D75019EA8B08" italics="true" pageId="0" pageNumber="129">Hemileccinum parvum</emphasis>
</taxonomicName>
has smaller basidiomata (pileus 3.3-3.6 cm diam, stipe 60-97 
<normalizedToken id="3A089CB296D5E31E7C40209DED006599" originalValue="×">x</normalizedToken>
4-9 mm); paler pileus (brownish to yellowish); pale yellow context that slowly turns pale blue when cut (
<bibRefCitation id="84058733FB6B87626AB136B4F57856BD" DOI="https://doi.org/10.3390/jof7100823" author="Li, MX" journalOrPublisher="Journal of Fungi" pageId="0" pageNumber="129" pagination=": 823" refId="B40" refString="Li, MX, Wu, G, Yang, ZL, 2021. Four new species of Hemileccinum (Xerocomoideae, Boletaceae) from Southwestern China. Journal of Fungi 7 (10): 823, DOI: https://doi.org/10.3390/jof7100823" title="Four new species of Hemileccinum (Xerocomoideae, Boletaceae) from Southwestern China." url="https://doi.org/10.3390/jof7100823" volume="7" year="2021">Li et al. 2021</bibRefCitation>
). 
<taxonomicName id="AFBB991C600DA1070441469572278434" class="Agaricomycetes" family="Pinaceae" genus="Hemileccinum" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Hemileccinum rugosum" order="Boletales" pageId="0" pageNumber="129" phylum="Basidiomycota" rank="species" species="rugosum">
<emphasis id="CA72149767DA89A306E05545C719AA8E" italics="true" pageId="0" pageNumber="129">Hemileccinum rugosum</emphasis>
</taxonomicName>
has paler pileus (light orange to reddish orange); very distinctly rugose to wrinkled pileus surface; and shorter basidiospores (9-13 
<normalizedToken id="5C17434822A604A32DC23C0B32C3B2DF" originalValue="×">x</normalizedToken>
4-5 
<normalizedToken id="45C64E2EBABB6458E6AE1803020C27AC" originalValue="µm">µm</normalizedToken>
; 
<bibRefCitation id="39EABD51E2BD38EAA64BC1C8EB5550FA" DOI="https://doi.org/10.1007/s13225-016-0375-8" author="Wu, G" journalOrPublisher="Fungal Diversity" pageId="0" pageNumber="129" pagination="25 - 188" refId="B71" refString="Wu, G, Li, YC, Zhu, XT, Zhao, K, Han, LH, Cui, YY, Li, F, Xu, JP, Yang, ZL, 2016. One hundred noteworthy boletes from China. Fungal Diversity 81 (1): 25 - 188, DOI: https://doi.org/10.1007/s13225-016-0375-8" title="One hundred noteworthy boletes from China." url="https://doi.org/10.1007/s13225-016-0375-8" volume="81" year="2016">Wu et al. 2016</bibRefCitation>
).
</paragraph>
<paragraph id="3E3F3B8BECCA0FA9095D3EAB7879E49C" pageId="0" pageNumber="129">
<taxonomicName id="6685B30BD99B5FC583A6A47979EC5BF8" authorityName="Vadthanarat, Raghoonundon &amp; Raspe" authorityYear="2024" class="Agaricomycetes" family="Pinaceae" genus="Hemileccinum" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Hemileccinum inferius" order="Boletales" pageId="0" pageNumber="129" phylum="Basidiomycota" rank="species" species="inferius">
<emphasis id="63702154B7BBFA143AA8F166A949CE30" italics="true" pageId="0" pageNumber="129">Hemileccinum inferius</emphasis>
</taxonomicName>
is also similar to an American species, 
<taxonomicName id="6475F3EBD6621B55E4550B4F881ECBD9" class="Agaricomycetes" family="Boletaceae" kingdom="Fungi" lsidName="H. floridanum" order="Boletales" pageId="0" pageNumber="129" phylum="Basidiomycota" rank="species" species="floridanum">
<emphasis id="2B75767AB274CEE40F714D4F9E378CC7" italics="true" pageId="0" pageNumber="129">H. floridanum</emphasis>
</taxonomicName>
, which has reddish brown to chestnut brown wrinkled and uneven pileus, whitish to pale yellow stipe, white basal mycelium, yellow hymenophore, and smooth basidiospore under both light microscope and SEM. However, the latter species is different by white context that slowly turns yellow often from the margin toward the center, longer basidiospores (10-17 
<normalizedToken id="3EE28A4B106077BF8A6EFCD0ABF57293" originalValue="×">x</normalizedToken>
4.5-6 
<normalizedToken id="B2C0C449E59B0119748917E4CE60AA9C" originalValue="μm">μm</normalizedToken>
), likely forms association with oak in northern America (
<bibRefCitation id="BC86158674D26A239D76023A461D7B86" DOI="https://doi.org/10.5943/mycosphere/12/1/12" author="Farid, A" journalOrPublisher="Mycosphere" pageId="0" pageNumber="129" pagination="1038 - 1076" refId="B18" refString="Farid, A, Bessette, AE, Bessette, AR, Bolin, JA, Kudzma, LV, Franck, AR, Garey, JR, 2021. Investigations in the boletes (Boletaceae) of southeastern USA: Four novel species and three novel combinations. Mycosphere 12 (1): 1038 - 1076, DOI: https://doi.org/10.5943/mycosphere/12/1/12" title="Investigations in the boletes (Boletaceae) of southeastern USA: Four novel species and three novel combinations." url="https://doi.org/10.5943/mycosphere/12/1/12" volume="12" year="2021">Farid et al. 2021</bibRefCitation>
).
</paragraph>
<paragraph id="7CCED305D8E53F7E691DC50734D40773" pageId="0" pageNumber="129">
Phylogenetically, 
<taxonomicName id="A8D2D8B5E009CC5D62CC769068C7520E" class="Agaricomycetes" family="Boletaceae" kingdom="Fungi" lsidName="H. inferius" order="Boletales" pageId="0" pageNumber="129" phylum="Basidiomycota" rank="species" species="inferius">
<emphasis id="808AB7FA433C6A3A98D7D6F3D876B645" italics="true" pageId="0" pageNumber="129">H. inferius</emphasis>
</taxonomicName>
was most closely related to 
<taxonomicName id="DD4DB5893038FE8E90F6AD1CD23F7BE5" class="Agaricomycetes" family="Boletaceae" kingdom="Fungi" lsidName="H. hortonii" order="Boletales" pageId="0" pageNumber="129" phylum="Basidiomycota" rank="species" species="hortonii">
<emphasis id="CB199FC5D3C73A16E84EE10FFD760C21" italics="true" pageId="0" pageNumber="129">H. hortonii</emphasis>
</taxonomicName>
, 
<taxonomicName id="29EEE6D8DDD4A0FFF5C225767F9C4702" class="Agaricomycetes" family="Boletaceae" kingdom="Fungi" lsidName="H. rugosum" order="Boletales" pageId="0" pageNumber="129" phylum="Basidiomycota" rank="species" species="rugosum">
<emphasis id="6C41627170EF4326531364CF4F0A732F" italics="true" pageId="0" pageNumber="129">H. rugosum</emphasis>
</taxonomicName>
, and an undescribed specimen (voucher HKAS 53421) from China. 
<taxonomicName id="9D200133AD54D4ED73F08D13A4152F81" class="Agaricomycetes" family="Pinaceae" genus="Hemileccinum" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Hemileccinum hortonii" order="Boletales" pageId="0" pageNumber="129" phylum="Basidiomycota" rank="species" species="hortonii">
<emphasis id="5730FABC88AE0B69A4DEAFE84DFD4BD5" italics="true" pageId="0" pageNumber="129">Hemileccinum hortonii</emphasis>
</taxonomicName>
, an American species, can easily be distinguished by its conspicuously pitted pileus, smooth to lightly pruinose stipe that sometimes has delicate reticulation on the upper half, pores that occasionally turn blue on when touched, and slightly longer and narrower basidiospores (12-15 
<normalizedToken id="3553443531ACC0DDC60DCBFB6759C083" originalValue="×">x</normalizedToken>
3.5-4.5; 
<bibRefCitation id="07B27AFF1B7A2810147AB7F198FDF0FC" DOI="https://doi.org/10.1080/00275514.2019.1685351" author="Kuo, M" journalOrPublisher="Mycologia" pageId="0" pageNumber="129" pagination="197 - 211" refId="B35" refString="Kuo, M, Ortiz-Santana, B, 2020. Revision of leccinoid fungi, with emphasis on North American taxa, based on molecular and morphological data. Mycologia 112 (1): 197 - 211, DOI: https://doi.org/10.1080/00275514.2019.1685351" title="Revision of leccinoid fungi, with emphasis on North American taxa, based on molecular and morphological data." url="https://doi.org/10.1080/00275514.2019.1685351" volume="112" year="2020">Kuo and Ortiz-Santana 2020</bibRefCitation>
; 
<bibRefCitation id="FB0AE772D1CB1E2726215A8078D3812B" DOI="https://doi.org/10.5943/mycosphere/12/1/12" author="Farid, A" journalOrPublisher="Mycosphere" pageId="0" pageNumber="129" pagination="1038 - 1076" refId="B18" refString="Farid, A, Bessette, AE, Bessette, AR, Bolin, JA, Kudzma, LV, Franck, AR, Garey, JR, 2021. Investigations in the boletes (Boletaceae) of southeastern USA: Four novel species and three novel combinations. Mycosphere 12 (1): 1038 - 1076, DOI: https://doi.org/10.5943/mycosphere/12/1/12" title="Investigations in the boletes (Boletaceae) of southeastern USA: Four novel species and three novel combinations." url="https://doi.org/10.5943/mycosphere/12/1/12" volume="12" year="2021">Farid et al. 2021</bibRefCitation>
). For morphological comparison with 
<taxonomicName id="12FB888226842BCDB86C51EBF5751350" class="Agaricomycetes" family="Boletaceae" kingdom="Fungi" lsidName="H. rugosum" order="Boletales" pageId="0" pageNumber="129" phylum="Basidiomycota" rank="species" species="rugosum">
<emphasis id="8AC704B66AEEB719D926E091E851F9F7" italics="true" pageId="0" pageNumber="129">H. rugosum</emphasis>
</taxonomicName>
see the above paragraph.
</paragraph>
</subSubSection>
</treatment>
</subSection>
</document>