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<document id="7A9F2D14E840DBDFC6E9E23D7040FD20" ID-DOI="10.1649/0010-065X-73.2.465" ID-ISSN="1938-4394" ID-Zenodo-Dep="5376930" ID-ZooBank="urn:lsid:zoobank.org:pub:07E37C8B-D586-4EB2-A155-9C843584C6B6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630282335579" checkinUser="valdenar" docAuthor="Akhil, S. V. &amp; Thomas, Sabu K." docDate="2019" docId="A91587910B40D219FD74222924471439" docLanguage="en" docName="0010-065X-73.2.465.pdf" docOrigin="The Coleopterists Bulletin 73 (2)" docSource="http://dx.doi.org/10.1649/0010-065x-73.2.465" docStyle="DocumentStyle:F824235CA89CB83023C7D5789122F870.3:TheColeopBull.2017-.journal_article.1cover" docStyleId="F824235CA89CB83023C7D5789122F870" docStyleName="TheColeopBull.2017-.journal_article.1cover" docStyleVersion="3" docTitle="Styphlomerus striatus Akhil and Thomas 2019, new species" docType="treatment" docUuid="CCCD4CF7-BC89-412A-9AE2-64A6F83D3FCF" docUuidSource="ZooBank" docVersion="4" lastPageNumber="469" masterDocId="552CFFE90B44D21CFFBA2713203C1673" masterDocTitle="New Species of Bombardier Beetles of the Genus Styphlomerus Chaudoir, 1875 (Coleoptera: Carabidae) from Southern India, with a Key to the Indomalayan and Palearctic Species" masterLastPageNumber="471" masterPageNumber="465" pageNumber="468" updateTime="1699212806870" updateUser="plazi">
<mods:mods id="D25E1804AC1F3343BC76F2B53E43A31F" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="02C9C9AE3F32B642AEBF8DDBC93ADF7F">
<mods:title id="2831F6A3C7FFCA34EE1F2129DE7784D6">New Species of Bombardier Beetles of the Genus Styphlomerus Chaudoir, 1875 (Coleoptera: Carabidae) from Southern India, with a Key to the Indomalayan and Palearctic Species</mods:title>
</mods:titleInfo>
<mods:name id="013A3701094C6289D0C419005A6C3105" type="personal">
<mods:role id="BB11077482865D74F5A9CF088BC1434B">
<mods:roleTerm id="0C46ACAD657C292A41A7F37416A58BF4">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="779959C17A6DD620ED73878AE4ECB243">Akhil, S. V.</mods:namePart>
</mods:name>
<mods:name id="02F72A207A37E3423A5513201D6B2BAA" type="personal">
<mods:role id="3C202410D5AAEF7184C9674A404021C9">
<mods:roleTerm id="59AC64EBF856E1BA754DB061D0DF2EBA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A46ECC8AD44E081D0103D21A6A2069A8">Thomas, Sabu K.</mods:namePart>
</mods:name>
<mods:typeOfResource id="1A74F98CEBB622D3C5070862B3B2D7B7">text</mods:typeOfResource>
<mods:relatedItem id="6A062C731C20505D087990237391F7BD" type="host">
<mods:titleInfo id="F885B3C5270354E25AB8AB3DE185C954">
<mods:title id="A5FED7A151DDC596544BAB4475B36236">The Coleopterists Bulletin</mods:title>
</mods:titleInfo>
<mods:part id="63C1D77A7FBB372E7A60CEDBA6C4DEFC">
<mods:date id="817270F6D53DACA9A5FFA5B56F393EAE">2019</mods:date>
<mods:detail id="C94B930A4F6229EA99365F9AFCEB82B8" type="pubDate">
<mods:number id="218754E1B49A7BE49DF4E2F3A0C93F31">2019-06-21</mods:number>
</mods:detail>
<mods:detail id="6E390430EEF2FE60235AC7BCF04EA18D" type="volume">
<mods:number id="DC0122E32D5E65994FA25BEA0D58D625">73</mods:number>
</mods:detail>
<mods:detail id="9157786D2173583465F0A69034439F30" type="issue">
<mods:number id="0173EF124E12F454B0B8F7650377E212">2</mods:number>
</mods:detail>
<mods:extent id="C0F9D1EFA2D130D15A79F0696AD1EA24" unit="page">
<mods:start id="C9F924AA8F3269C140031FC7D4932253">465</mods:start>
<mods:end id="91E639FE4FE34DFB508B3050E4C6113C">471</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="3191629877A8D12781356EFE1F2D75A8">
<mods:url id="245889EE9CA1BC83B2E3E483DF4A7B80">http://dx.doi.org/10.1649/0010-065x-73.2.465</mods:url>
</mods:location>
<mods:classification id="D101DB0FA205F68F2A7589C3510023AE">journal article</mods:classification>
<mods:identifier id="240A13944D97A01D64BB2EB383DF2375" type="DOI">10.1649/0010-065X-73.2.465</mods:identifier>
<mods:identifier id="0051FF23B763B39220CB83184621B23A" type="ISSN">1938-4394</mods:identifier>
<mods:identifier id="3063B242A985E95B748251032FDDF6CB" type="Zenodo-Dep">5376930</mods:identifier>
<mods:identifier id="CFDE7F5F350942C5A05A61112A0F0797" type="ZooBank">urn:lsid:zoobank.org:pub:07E37C8B-D586-4EB2-A155-9C843584C6B6</mods:identifier>
</mods:mods>
<treatment id="A91587910B40D219FD74222924471439" LSID="urn:lsid:zoobank.org:act:CCCD4CF7-BC89-412A-9AE2-64A6F83D3FCF" httpUri="http://treatment.plazi.org/id/A91587910B40D219FD74222924471439" lastPageId="5" lastPageNumber="469" pageId="4" pageNumber="468">
<subSubSection id="69A6650C0B40D218FD74222923EA13C9" pageId="4" pageNumber="468" type="nomenclature">
<paragraph id="210336870B40D218FD74222923E71319" blockId="4.[691,1166,1338,1466]" pageId="4" pageNumber="468">
<heading id="7A4B81EB0B40D218FD74222923E71319" centered="true" fontSize="9" level="2" pageId="4" pageNumber="468" reason="2">
<emphasis id="13C8EA950B40D218FD74222923E71319" bold="true" pageId="4" pageNumber="468">
<taxonomicName id="E6BC4D040B40D218FD7422292452133C" ID-CoL="8MCNR" authority="Akhil and Thomas" authorityName="Akhil and Thomas" authorityYear="2019" box="[718,1134,1338,1359]" class="Insecta" family="Carabidae" genus="Styphlomerus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="468" phylum="Arthropoda" rank="species" species="striatus" status="sp. nov.">
<emphasis id="13C8EA950B40D218FD742229239A133C" bold="true" box="[718,934,1338,1359]" italics="true" pageId="4" pageNumber="468">Styphlomerus striatus</emphasis>
Akhil and Thomas
</taxonomicName>
,
<taxonomicNameLabel id="08FB57EE0B40D218FCDC224623E71319" box="[870,987,1365,1386]" pageId="4" pageNumber="468" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="210336870B40D218FD50227C23EA13C9" blockId="4.[691,1166,1338,1466]" pageId="4" pageNumber="468">
Zoobank.org/
<uri id="552D3A850B40D218FCCA227C24B213EC" pageId="4" pageNumber="468">
urn:lsid:zoobank.org:act:
<uuid id="551A0C520B40D218FD09229924B213EC" box="[691,1166,1418,1439]" pageId="4" pageNumber="468">CCCD4CF7-BC89-412A-9AE2-64A6F83D3FCF</uuid>
</uri>
(
<figureCitation id="B9872A020B40D218FCC822B623F213C9" box="[882,974,1444,1466]" captionStart="Fig" captionStartId="3.[197,228,1370,1389]" captionTargetBox="[179,1188,196,1352]" captionTargetId="figure-112@3.[175,1190,194,1354]" captionTargetPageId="3" captionText="Fig. 1. Styphlomerus species. a) S. ruficeps, type in MNHN, b) S. fusciceps, type in NMPC, c) S. fusciceps, variant form, d) S. devagiriensis, holotype, e) S. striatus, holotype, f–g) S. striatus, aedeagus, right and left lateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.5376932" httpUri="https://zenodo.org/record/5376932/files/figure.png" pageId="4" pageNumber="468">Fig. 1e–g</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="69A6650C0B40D218FD7F22CB24B71001" pageId="4" pageNumber="468" type="description">
<paragraph id="210336870B40D218FD7F22CB24B71001" blockId="4.[685,1173,1496,1730]" pageId="4" pageNumber="468">
<emphasis id="13C8EA950B40D218FD7F22CB2384139E" bold="true" box="[709,952,1496,1517]" pageId="4" pageNumber="468">Specimens Examined.</emphasis>
<materialsCitation id="91D43CDA0B40D218FC7F22CB233A1051" collectingDate="2015-05-06" collectorName="S. V. Akhil" country="India" location="Tholpetty" pageId="4" pageNumber="468" specimenCount="1" specimenCount-male="1" stateProvince="Kerala">
<collectingCountry id="59AB76170B40D218FC7F22CB23C2139E" box="[965,1022,1496,1517]" name="India" pageId="4" pageNumber="468">India</collectingCountry>
:
<collectingRegion id="E378F8650B40D218FBB722CB24AC139E" box="[1037,1168,1496,1517]" country="India" name="Kerala" pageId="4" pageNumber="468">Kerala state</collectingRegion>
:
<location id="2463605C0B40D218FD1722E02336107B" LSID="urn:lsid:plazi:treatment:A91587910B40D219FD74222924471439:2463605C0B40D218FD1722E02336107B" box="[685,778,1523,1544]" country="India" name="Tholpetty" pageId="4" pageNumber="468" stateProvince="Kerala">Tholpetty</location>
,
<specimenCount id="37BAFD0E0B40D218FCA822E0236F107B" box="[786,851,1523,1544]" pageId="4" pageNumber="468" type="male">1 male</specimenCount>
, light attracted,
<date id="550210470B40D218FC5C22E02477107B" box="[998,1099,1523,1544]" pageId="4" pageNumber="468" value="2015-05-06">
<collectingDate id="4546E9AF0B40D218FC5C22E02477107B" box="[998,1099,1523,1544]" pageId="4" pageNumber="468" value="2015-05-06">06.V.2015</collectingDate>
</date>
, coll.
<collectorName id="8C4953510B40D218FB3A22E0233A1051" pageId="4" pageNumber="468">S. V. Akhil</collectorName>
</materialsCitation>
;
<materialsCitation id="91D43CDA0B40D218FCAF211E233E102B" collectingDate="2017-09-08" collectorName="S. V. Akhil" country="India" location="Ettimadai" pageId="4" pageNumber="468" specimenCount="5" specimenCount-female="4" specimenCount-male="1" stateProvince="Tamil Nadu">
<collectingCountry id="59AB76170B40D218FCAF211E23751051" box="[789,841,1549,1570]" name="India" pageId="4" pageNumber="468">India</collectingCountry>
:
<collectingRegion id="E378F8650B40D218FCE2211E24301051" box="[856,1036,1549,1570]" country="India" name="Tamil Nadu" pageId="4" pageNumber="468">Tamil Nadu State</collectingRegion>
:
<location id="2463605C0B40D218FBA0211E24441051" LSID="urn:lsid:plazi:treatment:A91587910B40D219FD74222924471439:2463605C0B40D218FBA0211E24441051" box="[1050,1144,1549,1570]" country="India" name="Ettimadai" pageId="4" pageNumber="468" stateProvince="Tamil Nadu">Ettimadai</location>
,
<specimenCount id="37BAFD0E0B40D218FB32211E22E1104E" pageId="4" pageNumber="468" type="male">1 male</specimenCount>
,
<specimenCount id="37BAFD0E0B40D218FD52213B2301104E" box="[744,829,1576,1597]" pageId="4" pageNumber="468" type="female">4 female</specimenCount>
, light attracted,
<date id="550210470B40D218FC6D213B247A104E" box="[983,1094,1576,1597]" pageId="4" pageNumber="468" value="2017-09-08">
<collectingDate id="4546E9AF0B40D218FC6D213B247A104E" box="[983,1094,1576,1597]" pageId="4" pageNumber="468" value="2017-09-08">08.IX.2017</collectingDate>
</date>
, coll.
<collectorName id="8C4953510B40D218FB3A213B233E102B" pageId="4" pageNumber="468">S. V. Akhil</collectorName>
</materialsCitation>
;
<materialsCitation id="91D43CDA0B40D218FCB6215024B71001" collectingDate="2017-10-15" collectorName="S. V. Akhil." country="India" location="Rajapalayam" pageId="4" pageNumber="468" specimenCount="1" specimenCount-male="1" stateProvince="Tamil Nadu">
<collectingCountry id="59AB76170B40D218FCB62150237C102B" box="[780,832,1603,1624]" name="India" pageId="4" pageNumber="468">India</collectingCountry>
:
<collectingRegion id="E378F8650B40D218FCF0215023CA102B" box="[842,1014,1603,1624]" country="India" name="Tamil Nadu" pageId="4" pageNumber="468">Tamil Nadu State</collectingRegion>
:
<location id="2463605C0B40D218FC4521502441102B" LSID="urn:lsid:plazi:treatment:A91587910B40D219FD74222924471439:2463605C0B40D218FC4521502441102B" box="[1023,1149,1603,1624]" country="India" name="Rajapalayam" pageId="4" pageNumber="468" stateProvince="Tamil Nadu">Rajapalayam</location>
,
<specimenCount id="37BAFD0E0B40D218FB32215022E11001" pageId="4" pageNumber="468" type="male">1 male</specimenCount>
, hand picking,
<date id="550210470B40D218FCCE214E23E71001" box="[884,987,1629,1650]" pageId="4" pageNumber="468" value="2017-10-15">
<collectingDate id="4546E9AF0B40D218FCCE214E23E71001" box="[884,987,1629,1650]" pageId="4" pageNumber="468" value="2017-10-15">15.X.2017</collectingDate>
</date>
, coll.
<collectorName id="8C4953510B40D218FBAD214E24B71001" box="[1047,1163,1629,1650]" pageId="4" pageNumber="468">S. V. Akhil.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="69A6650C0B40D219FD7F216B21E817E2" lastPageId="5" lastPageNumber="469" pageId="4" pageNumber="468" type="materials_examined">
<paragraph id="210336870B40D219FD7F216B21E817E2" blockId="4.[685,1173,1496,1730]" lastBlockId="5.[173,661,193,1730]" lastPageId="5" lastPageNumber="469" pageId="4" pageNumber="468">
<emphasis id="13C8EA950B40D218FD7F216B235F10FE" bold="true" box="[709,867,1656,1677]" pageId="4" pageNumber="468">Type Material.</emphasis>
<materialsCitation id="91D43CDA0B40D218FCC8216B241910B1" collectingDate="2017-09-08" collectorName="S. V. Akhil" country="India" county="Male" location="Light" municipality="Ettimadai" pageId="4" pageNumber="468" specimenCount="2" stateProvince="Tamil Nadu" typeStatus="holotype">
<typeStatus id="FE0788250B40D218FCC8216B23F610FE" box="[882,970,1656,1677]" pageId="4" pageNumber="468" type="holotype">Holotype</typeStatus>
labeled “
<typeStatus id="FE0788250B40D218FB8F216B24AC10FE" box="[1077,1168,1656,1677]" pageId="4" pageNumber="468" type="holotype">Holotype</typeStatus>
,
<collectingCounty id="C8624E0B0B40D218FD17218122DC10D4" box="[685,736,1682,1703]" pageId="4" pageNumber="468">Male</collectingCounty>
,
<collectingCountry id="59AB76170B40D218FD4A2181231810D4" box="[752,804,1682,1703]" name="India" pageId="4" pageNumber="468">India</collectingCountry>
:
<collectingRegion id="E378F8650B40D218FC89218123D810D4" box="[819,996,1682,1703]" country="India" name="Tamil Nadu" pageId="4" pageNumber="468">Tamil Nadu state</collectingRegion>
:
<collectingMunicipality id="C167ACFD0B40D218FC492181246D10D4" box="[1011,1105,1682,1703]" pageId="4" pageNumber="468">Ettimadai</collectingMunicipality>
,
<location id="2463605C0B40D218FBDA218124AF10D4" LSID="urn:lsid:plazi:treatment:A91587910B40D219FD74222924471439:2463605C0B40D218FBDA218124AF10D4" box="[1120,1171,1682,1703]" country="India" county="Male" municipality="Ettimadai" name="Light" pageId="4" pageNumber="468" stateProvince="Tamil Nadu">Light</location>
attracted,
<date id="550210470B40D218FCB021BE234510B1" box="[778,889,1709,1730]" pageId="4" pageNumber="468" value="2017-09-08">
<collectingDate id="4546E9AF0B40D218FCB021BE234510B1" box="[778,889,1709,1730]" pageId="4" pageNumber="468" value="2017-09-08">08.IX.2017</collectingDate>
</date>
, coll.
<collectorName id="8C4953510B40D218FC0821BE242710B1" box="[946,1051,1709,1730]" pageId="4" pageNumber="468">S. V. Akhil</collectorName>
”
</materialsCitation>
.
<materialsCitation id="91D43CDA0B40D219FB8B21BE208B1754" collectingDate="2017-09-08" collectorName="S. V. Akhil" country="India" lastPageId="5" lastPageNumber="469" location="Four" pageId="4" pageNumber="468" specimenCount="2" stateProvince="Tamil Nadu" typeStatus="paratype">
<typeStatus id="FE0788250B40D218FB8B21BE24AC10B1" box="[1073,1168,1709,1730]" pageId="4" pageNumber="468" type="paratype">Paratypes</typeStatus>
:
<location id="2463605C0B41D219FF1727D120E016A4" LSID="urn:lsid:plazi:treatment:A91587910B40D219FD74222924471439:2463605C0B41D219FF1727D120E016A4" box="[173,220,194,215]" country="India" name="Four" pageId="5" pageNumber="469" stateProvince="Tamil Nadu">Four</location>
females labeled “
<typeStatus id="FE0788250B41D219FE2227D121CF16A4" box="[408,499,194,215]" pageId="5" pageNumber="469" type="paratype">Paratype</typeStatus>
, Female,
<collectingCountry id="59AB76170B41D219FDE227D122AC16A4" box="[600,656,194,215]" name="India" pageId="5" pageNumber="469">India</collectingCountry>
:
<collectingRegion id="E378F8650B41D219FF1727CF21501682" box="[173,364,220,241]" country="India" name="Tamil Nadu" pageId="5" pageNumber="469">Tamil Nadu state</collectingRegion>
: Ettimadai, Light attracted,
<date id="550210470B41D219FF1727E42127177F" box="[173,283,247,268]" pageId="5" pageNumber="469" value="2017-09-08">
<collectingDate id="4546E9AF0B41D219FF1727E42127177F" box="[173,283,247,268]" pageId="5" pageNumber="469" value="2017-09-08">08.IX.2017</collectingDate>
</date>
, coll.
<collectorName id="8C4953510B41D219FEE627E421F0177F" box="[348,460,247,268]" pageId="5" pageNumber="469">S. V. Akhil</collectorName>
”/ One male labeled “
</materialsCitation>
<materialsCitation id="91D43CDA0B41D219FF0D26022100172F" collectingDate="2015-05-06" collectorName="S. V. Akhil" country="India" county="Male" location="Light" municipality="Tholpetty" pageId="5" pageNumber="469" specimenCount="1" stateProvince="Kerala" typeStatus="paratype">
<typeStatus id="FE0788250B41D219FF0D260221301755" box="[183,268,273,294]" pageId="5" pageNumber="469" type="paratype">Paratype</typeStatus>
,
<collectingCounty id="C8624E0B0B41D219FEA42602216D1755" box="[286,337,273,294]" pageId="5" pageNumber="469">Male</collectingCounty>
,
<collectingCountry id="59AB76170B41D219FED8260221AA1755" box="[354,406,273,294]" name="India" pageId="5" pageNumber="469">India</collectingCountry>
:
<collectingRegion id="E378F8650B41D219FE1D2602221E1755" box="[423,546,273,294]" country="India" name="Kerala" pageId="5" pageNumber="469">Kerala state</collectingRegion>
:
<collectingMunicipality id="C167ACFD0B41D219FD88260222B31755" box="[562,655,273,294]" pageId="5" pageNumber="469">Tholpetty</collectingMunicipality>
,
<location id="2463605C0B41D219FF17263F20DC1732" LSID="urn:lsid:plazi:treatment:A91587910B40D219FD74222924471439:2463605C0B41D219FF17263F20DC1732" box="[173,224,300,321]" country="India" county="Male" municipality="Tholpetty" name="Light" pageId="5" pageNumber="469" stateProvince="Kerala">Light</location>
attracted,
<date id="550210470B41D219FEF3263F21911732" box="[329,429,300,321]" pageId="5" pageNumber="469" value="2015-05-06">
<collectingDate id="4546E9AF0B41D219FEF3263F21911732" box="[329,429,300,321]" pageId="5" pageNumber="469" value="2015-05-06">06.V.2015</collectingDate>
</date>
, coll.
<collectorName id="8C4953510B41D219FE51263F226D1732" box="[491,593,300,321]" pageId="5" pageNumber="469">S.V. Akhil</collectorName>
”/ One male labeled “
</materialsCitation>
<materialsCitation id="91D43CDA0B41D219FE86265521EC17E2" collectingDate="2017-10-15" collectionCode="ZSIC" collectorName="S. V. Akhil. Deposited" country="India" county="Male" location="Hand" municipality="Rajapalayam" pageId="5" pageNumber="469" specimenCount="1" stateProvince="Tamil Nadu" typeStatus="paratype">
<typeStatus id="FE0788250B41D219FE86265521AD1728" box="[316,401,326,347]" pageId="5" pageNumber="469" type="paratype">Paratype</typeStatus>
,
<collectingCounty id="C8624E0B0B41D219FE24265521ED1728" box="[414,465,326,347]" pageId="5" pageNumber="469">Male</collectingCounty>
,
<collectingCountry id="59AB76170B41D219FE642655222E1728" box="[478,530,326,347]" name="India" pageId="5" pageNumber="469">India</collectingCountry>
:
<collectingRegion id="E378F8650B41D219FDA5265520E01705" country="India" name="Tamil Nadu" pageId="5" pageNumber="469">Tamil Nadu state</collectingRegion>
:
<collectingMunicipality id="C167ACFD0B41D219FF5E2672215E1705" box="[228,354,353,374]" pageId="5" pageNumber="469">Rajapalayam</collectingMunicipality>
,
<location id="2463605C0B41D219FED62672219C1705" LSID="urn:lsid:plazi:treatment:A91587910B40D219FD74222924471439:2463605C0B41D219FED62672219C1705" box="[364,416,353,374]" country="India" county="Male" municipality="Rajapalayam" name="Hand" pageId="5" pageNumber="469" stateProvince="Tamil Nadu">Hand</location>
picking,
<date id="550210470B41D219FE432672225C1705" box="[505,608,353,374]" pageId="5" pageNumber="469" value="2017-10-15">
<collectingDate id="4546E9AF0B41D219FE432672225C1705" box="[505,608,353,374]" pageId="5" pageNumber="469" value="2017-10-15">15.X.2017</collectingDate>
</date>
, coll.
<collectorName id="8C4953510B41D219FF17266F214417E2" box="[173,376,380,401]" pageId="5" pageNumber="469">S.V.Akhil. Deposited</collectorName>
in
<collectionCode id="47ADAE420B41D219FE20266F21EC17E2" box="[410,464,380,401]" country="India" httpUri="http://grbio.org/cool/5icv-29es" name="Zoological Survey of India" pageId="5" pageNumber="469">ZSIC</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="69A6650C0B41D219FF7F268522C917D8" pageId="5" pageNumber="469" type="description">
<paragraph id="210336870B41D219FF7F2685218610FF" blockId="5.[173,661,193,1730]" pageId="5" pageNumber="469">
<emphasis id="13C8EA950B41D219FF7F2685210317D8" bold="true" box="[197,319,406,427]" pageId="5" pageNumber="469">Description.</emphasis>
TLA =
<quantity id="E6449B620B41D219FE2E2685223617D8" box="[404,522,406,428]" metricMagnitude="-3" metricUnit="m" metricValue="7.6" metricValueMax="7.7" metricValueMin="7.5" pageId="5" pageNumber="469" unit="mm" value="7.6" valueMax="7.7" valueMin="7.5">7.5–7.7 mm</quantity>
.
<emphasis id="13C8EA950B41D219FDA92685226817D8" bold="true" box="[531,596,406,427]" pageId="5" pageNumber="469">Color:</emphasis>
Head, pronotum, labrum, mandible, scutellum, and region around scutellum reddish brown; scape and legs reddish yellow; pedicel darker reddish yellow, antennomeres 3 and 4 brownish black, other antennomeres dark brown; elytra black; genae, gula, prosternum, ventral prothorax, and mesosternum dark reddish yellow; metasternum, mesepisternum, metepisternum, and mesepimeron brownish yellow; abdomen brownish black.
<emphasis id="13C8EA950B41D219FE18259621E314E9" bold="true" box="[418,479,645,666]" pageId="5" pageNumber="469">Head:</emphasis>
Broader than long, widest between eyes, slightly narrower than pronotum, stout, punctate, region around eyes, sides of frons, and clypeus sparsely pubescent; eyes black, protruding; labral apical angles strongly rounded, form subrectangular, with 6 setae on apical margin; clypeus rectangular, wider than long, apical margin straight, with short setae on lateral margins, midregion glabrous; frontoclypeal suture impressed, distinct; frons with anterolateral setose depressions, midregion glabrous; mandibles sharp, broad; mentum broad, margins pronounced, with short, broad, pointed median tooth, lateral lobes triangular with lateral tooth-like projection, a single seta apicolaterally, remainder of mentum glabrous; palpi pubescent, apical palpomeres pointed, conical, penultimate palpomere with apex wide, base narrow; antennae short, not reaching middle of elytra, antennomeres slender.
<emphasis id="13C8EA950B41D219FE3E237021D3120B" bold="true" box="[388,495,1123,1144]" pageId="5" pageNumber="469">Pronotum:</emphasis>
Punctate, pubescent; anterior margin slightly concave, lateral angles of anterior margin projecting slightly forward, posterior margin straight; anterior 3/4 of disc convex, posterior fourth narrowed, parallel towards base, hind angles quadrate without lateral projection; median furrow weak, shallow, not reaching anterior and posterior margins; lateral ridge wider, margin dark brown.
<emphasis id="13C8EA950B41D219FF41222B2159133E" bold="true" box="[251,357,1336,1357]" pageId="5" pageNumber="469">Scutellum:</emphasis>
Pointed, broad.
<emphasis id="13C8EA950B41D219FDA9222B2265133E" bold="true" box="[531,601,1336,1357]" pageId="5" pageNumber="469">Elytra:</emphasis>
Subparallel, matt, faintly punctate, pubescent with golden brown setae; humerus prominent, rounded; surface flat, apex depressed; striations visible, weak, rounded, intervals narrow, setae along striations and interstices.
<emphasis id="13C8EA950B41D219FF1722AE211913A1" bold="true" box="[173,293,1469,1490]" pageId="5" pageNumber="469">Hind wings:</emphasis>
Well-developed.
<emphasis id="13C8EA950B41D219FE7222AE222D13A1" bold="true" box="[456,529,1469,1490]" pageId="5" pageNumber="469">Venter:</emphasis>
Genae broad, smooth, with few short setae under eyes, remainder glabrous, genae with fine, superficial wrinkles; gula wide, divergent posteriorly, gular sutures brownish black; prothorax ventrally smooth, sparsely pubescent laterally towards neck region; prosternal process smooth, with 4 short setae apically in 2 rows; apical portion of abdomen pointed.
</paragraph>
<paragraph id="210336870B41D219FF7F218122A810B1" blockId="5.[173,661,193,1730]" pageId="5" pageNumber="469">
<emphasis id="13C8EA950B41D219FF7F218121C810D4" bold="true" box="[197,500,1682,1703]" pageId="5" pageNumber="469">
Measurements.
<typeStatus id="FE0788250B41D219FEE22181219210D4" box="[344,430,1682,1703]" pageId="5" pageNumber="469" type="holotype">Holotype</typeStatus>
(male):
</emphasis>
TLA =
<quantity id="E6449B620B41D219FD85218122AC10D4" box="[575,656,1682,1703]" metricMagnitude="-3" metricUnit="m" metricValue="7.55" pageId="5" pageNumber="469" unit="mm" value="7.55">7.55 mm</quantity>
, TLB =
<quantity id="E6449B620B41D219FF4021BE217310B1" box="[250,335,1709,1730]" metricMagnitude="-3" metricUnit="m" metricValue="6.34" pageId="5" pageNumber="469" unit="mm" value="6.34">6.34 mm</quantity>
, PL =
<quantity id="E6449B620B41D219FE2D21BE21D710B1" box="[407,491,1709,1730]" metricMagnitude="-3" metricUnit="m" metricValue="1.05" pageId="5" pageNumber="469" unit="mm" value="1.05">1.05 mm</quantity>
, PW =
<quantity id="E6449B620B41D219FD8121BE22AC10B1" box="[571,656,1709,1730]" metricMagnitude="-3" metricUnit="m" metricValue="1.38" pageId="5" pageNumber="469" unit="mm" value="1.38">1.38 mm</quantity>
,
</paragraph>
<paragraph id="210336870B41D219FD7927D124041755" blockId="5.[707,1194,193,586]" pageId="5" pageNumber="469">
EL =
<quantity id="E6449B620B41D219FD4427D1236D16A4" box="[766,849,194,215]" metricMagnitude="-3" metricUnit="m" metricValue="3.97" pageId="5" pageNumber="469" unit="mm" value="3.97">3.97 mm</quantity>
, TW =
<quantity id="E6449B620B41D219FC2427D123CD16A4" box="[926,1009,194,215]" metricMagnitude="-3" metricUnit="m" metricValue="2.61" pageId="5" pageNumber="469" unit="mm" value="2.61">2.61 mm</quantity>
.
<emphasis id="13C8EA950B41D219FC4027D1249516A4" bold="true" box="[1018,1193,194,215]" pageId="5" pageNumber="469">
<typeStatus id="FE0788250B41D219FC4027D1246C16A4" box="[1018,1104,194,215]" pageId="5" pageNumber="469" type="paratype">Paratype</typeStatus>
(female):
</emphasis>
TLA =
<quantity id="E6449B620B41D219FCA127CF23911682" box="[795,941,220,242]" metricMagnitude="-3" metricUnit="m" metricValue="7.824999999999999" metricValueMax="8.03" metricValueMin="7.62" pageId="5" pageNumber="469" unit="mm" value="7.824999999999999" valueMax="8.03" valueMin="7.62">7.62–8.03 mm</quantity>
, TLB =
<quantity id="E6449B620B41D219FBA927CF24991682" box="[1043,1189,220,242]" metricMagnitude="-3" metricUnit="m" metricValue="6.53" metricValueMax="6.58" metricValueMin="6.48" pageId="5" pageNumber="469" unit="mm" value="6.53" valueMax="6.58" valueMin="6.48">6.48–6.58 mm</quantity>
, PL =
<quantity id="E6449B620B41D219FD4527E423B0177F" box="[767,908,246,268]" metricMagnitude="-3" metricUnit="m" metricValue="1.01" metricValueMax="1.06" metricValueMin="0.96" pageId="5" pageNumber="469" unit="mm" value="1.01" valueMax="1.06" valueMin="0.96">0.96–1.06 mm</quantity>
, PW =
<quantity id="E6449B620B41D219FC6027E42454177F" box="[986,1128,246,268]" metricMagnitude="-3" metricUnit="m" metricValue="1.325" metricValueMax="1.37" metricValueMin="1.28" pageId="5" pageNumber="469" unit="mm" value="1.325" valueMax="1.37" valueMin="1.28">1.28–1.37 mm</quantity>
, EL = 4.00–
<quantity id="E6449B620B41D219FD422602236D1755" box="[760,849,273,294]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="5" pageNumber="469" unit="mm" value="1.1">1.10 mm</quantity>
, TW =
<quantity id="E6449B620B41D219FC1E260224081755" box="[932,1076,273,295]" metricMagnitude="-3" metricUnit="m" metricValue="2.62" metricValueMax="2.74" metricValueMin="2.5" pageId="5" pageNumber="469" unit="mm" value="2.62" valueMax="2.74" valueMin="2.5">2.50–2.74 mm</quantity>
.
</paragraph>
<paragraph id="210336870B41D219FD61263F23191705" blockId="5.[707,1194,193,586]" pageId="5" pageNumber="469">
<emphasis id="13C8EA950B41D219FD61263F23C61732" bold="true" box="[731,1018,300,321]" pageId="5" pageNumber="469">Geographical Distribution.</emphasis>
<materialsCitation id="91D43CDA0B41D219FBB5263F241F1728" country="India" location="Tamil Nadu state" pageId="5" pageNumber="469" specimenCount="1" stateProvince="Tamil Nadu">
<emphasis id="13C8EA950B41D219FBB5263F24611732" bold="true" box="[1039,1117,300,321]" pageId="5" pageNumber="469">
<collectingCountry id="59AB76170B41D219FBB5263F24641732" box="[1039,1112,300,321]" name="India" pageId="5" pageNumber="469">INDIA</collectingCountry>
:
</emphasis>
<collectingRegion id="E378F8650B41D219FBC8263F23121728" country="India" name="Tamil Nadu" pageId="5" pageNumber="469">Tamil Nadu state</collectingRegion>
: Rajapalayam, Ettimadai
</materialsCitation>
;
<collectingRegion id="E378F8650B41D219FB8A2655249A1728" box="[1072,1190,326,347]" country="India" name="Kerala" pageId="5" pageNumber="469">Kerala state</collectingRegion>
: Tholpetty.
</paragraph>
<paragraph id="210336870B41D219FD61266F22C917D8" blockId="5.[707,1194,193,586]" pageId="5" pageNumber="469">
<emphasis id="13C8EA950B41D219FD61266F23A017E2" bold="true" box="[731,924,380,401]" pageId="5" pageNumber="469">Collection Method.</emphasis>
Collected under fluorescent light.
</paragraph>
</subSubSection>
<subSubSection id="69A6650C0B41D219FD6126A224951793" pageId="5" pageNumber="469" type="etymology">
<paragraph id="210336870B41D219FD6126A224951793" blockId="5.[707,1194,193,586]" pageId="5" pageNumber="469">
<emphasis id="13C8EA950B41D219FD6126A2237117B5" bold="true" box="[731,845,433,454]" pageId="5" pageNumber="469">Etymology.</emphasis>
Named for the elytral striations that differentiate it from the closely related
<taxonomicName id="E6BC4D040B41D219FB8726D9249F1793" authorityName="Chaudoir" authorityYear="1876" box="[1085,1187,458,480]" class="Insecta" family="Carabidae" genus="Styphlomerus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="469" phylum="Arthropoda" rank="species" species="ruficeps">
<emphasis id="13C8EA950B41D219FB8726D9249F1793" box="[1085,1187,458,480]" italics="true" pageId="5" pageNumber="469">S. ruficeps</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="69A6650C0B41D219FD6126F524471439" pageId="5" pageNumber="469" type="discussion">
<paragraph id="210336870B41D219FD6126F524471439" blockId="5.[707,1194,193,586]" pageId="5" pageNumber="469">
<emphasis id="13C8EA950B41D219FD6126F523011788" bold="true" box="[731,829,486,507]" pageId="5" pageNumber="469">Remarks.</emphasis>
<taxonomicName id="E6BC4D040B41D219FCF026F624231788" authorityName="Akhil and Thomas" authorityYear="2019" box="[842,1055,485,507]" class="Insecta" family="Carabidae" genus="Styphlomerus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="469" phylum="Arthropoda" rank="species" species="striatus">
<emphasis id="13C8EA950B41D219FCF026F624231788" box="[842,1055,485,507]" italics="true" pageId="5" pageNumber="469">Styphlomerus striatus</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="E6BC4D040B41D219FD79251323101465" authorityName="Chaudoir" authorityYear="1876" box="[707,812,512,534]" class="Insecta" family="Carabidae" genus="Styphlomerus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="469" phylum="Arthropoda" rank="species" species="ruficeps">
<emphasis id="13C8EA950B41D219FD79251323101465" box="[707,812,512,534]" italics="true" pageId="5" pageNumber="469">S. ruficeps</emphasis>
</taxonomicName>
but differs by having weak, shallow striations on the elytra, a pointed scutellum, and the anterior margin of the pronotum concave.
</paragraph>
</subSubSection>
</treatment>
</document>
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