302 lines
29 KiB
XML
302 lines
29 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.259.2970" ID-GBIF-Dataset="37fcf036-b7b9-407b-a090-85e2cea21080" ID-PMC="PMC3591765" ID-Pensoft-Pub="1313-2970-259-1" ID-PubMed="23653501" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1313-2970-259-1" ModsDocOrigin="ZooKeys 259" ModsDocTitle="A taxonomic revision of the Cymindis (Pinacodera) limbata species group (Coleoptera, Carabidae, Lebiini), including description of a new species from Florida, U.S.A." checkinTime="1451247765886" checkinUser="pensoft" docAuthor="Hunting, Wesley M." docDate="2013" docId="A2D6563F6B37D62CEC2683D3BE0CDE81" docLanguage="en" docName="ZooKeys 259: 1-73" docOrigin="ZooKeys 259" docSource="http://dx.doi.org/10.3897/zookeys.259.2970" docTitle="Cymindis (Pinacodera) chevrolati Dejean" docType="treatment" docVersion="4" lastPageNumber="53" masterDocId="FF90904E7A57FFE9CD13EE7858701A20" masterDocTitle="A taxonomic revision of the Cymindis (Pinacodera) limbata species group (Coleoptera, Carabidae, Lebiini), including description of a new species from Florida, U. S. A." masterLastPageNumber="73" masterPageNumber="1" pageNumber="49" updateTime="1668155161763" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A taxonomic revision of the Cymindis (Pinacodera) limbata species group (Coleoptera, Carabidae, Lebiini), including description of a new species from Florida, U. S. A.</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hunting, Wesley M.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2013</mods:date>
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<mods:detail type="volume">
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<mods:number>259</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>73</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.259.2970</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.259.2970</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-259-1</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152039702" LSID="urn:lsid:plazi:treatment:A2D6563F6B37D62CEC2683D3BE0CDE81" httpUri="http://treatment.plazi.org/id/A2D6563F6B37D62CEC2683D3BE0CDE81" lastPageId="52" lastPageNumber="53" pageId="48" pageNumber="49">
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<subSubSection pageId="48" pageNumber="49" type="nomenclature">
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<paragraph pageId="48" pageNumber="49">
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<taxonomicName LSID="http://species-id.net/wiki/Cymindis_chevrolati" authority="Dejean" class="Insecta" family="Carabidae" genus="Cymindis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymindis (Pinacodera) chevrolati" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="chevrolati" subGenus="Pinacodera">Cymindis (Pinacodera) chevrolati Dejean</taxonomicName>
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Figs 30-31, 34A, 35A, 36, 37B-D, 39-41
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</paragraph>
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</subSubSection>
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<subSubSection pageId="48" pageNumber="49" type="reference_group">
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<paragraph pageId="48" pageNumber="49">
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<taxonomicName class="Insecta" family="Carabidae" genus="Cymindis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymindis atrata" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="atrata">Cymindis atrata</taxonomicName>
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<bibRefCitation pageId="48" pageNumber="49">Chevrolat 1835</bibRefCitation>
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, fasc. 7, No. 152 (not
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<bibRefCitation pageId="48" pageNumber="49">Dejean 1831</bibRefCitation>
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: 327).- TYPE MATERIAL in OXUM: four specimens (2 males, 2 females) three of which are in front of the following green box label: "Cymindis/ Chevrolati/ Dj Cat-3, p.9/ atrata Chv Col Mus 1835/ fasc 7- 152/ Mexico/ D.
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<normalizedToken originalValue="Sallé">Salle</normalizedToken>
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" [handwritten]. LECTOTYPE (here designated) male, labeled: [small green square]; "Chevrolat/ Carabidae/ Fr. V. d. Poll/ Pres. 1909, E./ B. Poulton." PARALECTOTYPES, (three, here designated), two of which in front of above box label: male, [two lines of illegible handwriting]; "Chevrolat/ Carabidae/ Fr. V. d. Poll/ Pres. 1909, E./ B. Poulton"; female, "Cymindis nigrita Ch"/
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<normalizedToken originalValue="“Mexico”">"Mexico"</normalizedToken>
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[? handwritten]; "Chevrolat/ Carabidae/ Fr. V. d. Poll/ Pres. 1909, E./ B. Poulton". Paralectotype 3, female, in front of green box label: Cymindis/ nigrita Chd Bull. Mosc./ 1837-VII p. 6/ Mexico D.
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<normalizedToken originalValue="Sallé”">Salle"</normalizedToken>
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[handwritten]. Specimen labeled: "Cymindis/nigrita Chaud./ Mexique"; "Chevrolat/ Carabidae/ Fr. V. d. Poll/ Pres. 1909, E./ B. Poulton". -
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<bibRefCitation pageId="48" pageNumber="49">Dejean 1836</bibRefCitation>
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: 9. -
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<bibRefCitation pageId="48" pageNumber="49">Bates 1883</bibRefCitation>
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: 187, and
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<bibRefCitation pageId="48" pageNumber="49">1891</bibRefCitation>
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: 270.
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</paragraph>
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<paragraph pageId="48" pageNumber="49">
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<taxonomicName class="Insecta" family="Carabidae" genus="Cymindis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymindis chevrolatii" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="chevrolatii">Cymindis chevrolatii</taxonomicName>
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<bibRefCitation pageId="48" pageNumber="49">Dejean 1836</bibRefCitation>
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: 9 [replacement name for
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<taxonomicName class="Insecta" family="Carabidae" genus="Cymindis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymindis atrata" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="atrata">Cymindis atrata</taxonomicName>
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<bibRefCitation pageId="48" pageNumber="49">Chevrolat 1835</bibRefCitation>
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]. -
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<bibRefCitation author="Chaudoir, M de" journalOrPublisher="Berliner Entomologische Zeitschrift" pageId="57" pageNumber="58" pagination="53 - 120" title="Essai monographique sur le genre Cymindis proprement dit." volume="17" year="1873">Chaudoir 1873</bibRefCitation>
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: 54.
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</paragraph>
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<paragraph pageId="48" pageNumber="49">
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<taxonomicName class="Insecta" family="Carabidae" genus="Cymindis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymindis nigrita" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="nigrita">Cymindis nigrita</taxonomicName>
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<bibRefCitation pageId="48" pageNumber="49">Chaudoir 1837</bibRefCitation>
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: 6-8 [unnecessary replacement name for
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<taxonomicName class="Insecta" family="Carabidae" genus="Cymindis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymindis atrata" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="atrata">Cymindis atrata</taxonomicName>
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<bibRefCitation pageId="48" pageNumber="49">Chevrolat 1835</bibRefCitation>
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]. -
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<bibRefCitation author="Chaudoir, M de" journalOrPublisher="Berliner Entomologische Zeitschrift" pageId="57" pageNumber="58" pagination="53 - 120" title="Essai monographique sur le genre Cymindis proprement dit." volume="17" year="1873">Chaudoir 1873</bibRefCitation>
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: 54, and
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<bibRefCitation pageId="48" pageNumber="49">1875</bibRefCitation>
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: 4.
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</paragraph>
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<paragraph pageId="48" pageNumber="49">
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<taxonomicName class="Insecta" family="Carabidae" genus="Cymindis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymindis amblygona" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="amblygona">Cymindis amblygona</taxonomicName>
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<bibRefCitation pageId="48" pageNumber="49">Bates 1878</bibRefCitation>
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: 606. TYPE MATERIAL: not seen. TYPE AREA: Mexico.
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</paragraph>
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<paragraph pageId="48" pageNumber="49">
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<taxonomicName class="Insecta" family="Carabidae" genus="Cymindis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymindis angulifera" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="angulifera">Cymindis angulifera</taxonomicName>
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<bibRefCitation pageId="48" pageNumber="49">Bates 1878</bibRefCitation>
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: 606. TYPE MATERIAL: not seen. TYPE AREA: Mexico.
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</paragraph>
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<paragraph pageId="48" pageNumber="49">
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<taxonomicName class="Insecta" family="Carabidae" genus="Pinacodera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pinacodera atrata" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="atrata">Pinacodera atrata</taxonomicName>
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var. amblygona
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<bibRefCitation pageId="48" pageNumber="49">Bates 1883</bibRefCitation>
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: 187.
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</paragraph>
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<paragraph pageId="48" pageNumber="49">
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<taxonomicName class="Insecta" family="Carabidae" genus="Pinacodera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pinacodera atrata" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="atrata">Pinacodera atrata</taxonomicName>
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var. angulifera
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<bibRefCitation pageId="48" pageNumber="49">Bates 1883</bibRefCitation>
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: 187.
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</paragraph>
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<paragraph pageId="48" pageNumber="49">
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<taxonomicName class="Insecta" family="Carabidae" genus="Pinacodera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pinacodera chevrolati" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="chevrolati">Pinacodera chevrolati</taxonomicName>
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<bibRefCitation pageId="48" pageNumber="49">Csiki 1932</bibRefCitation>
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: 1487. -
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<bibRefCitation pageId="48" pageNumber="49">Blackwelder 1944</bibRefCitation>
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: 62.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="48" pageNumber="49" type="type locality">
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<paragraph pageId="48" pageNumber="49">Type locality.</paragraph>
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<paragraph pageId="48" pageNumber="49">Cruz Blanca, Veracruz, Mexico.</paragraph>
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</subSubSection>
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<subSubSection pageId="48" pageNumber="49" type="specific epithet">
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<paragraph pageId="48" pageNumber="49">Specific epithet.</paragraph>
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<paragraph pageId="48" pageNumber="49">
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The original spelling is
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<normalizedToken originalValue="“chevrolatii”">"chevrolatii"</normalizedToken>
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, but relatively recent catalogues (
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<bibRefCitation pageId="48" pageNumber="49">Csiki 1932</bibRefCitation>
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,
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<bibRefCitation pageId="48" pageNumber="49">Blackwelder 1944</bibRefCitation>
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,
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<bibRefCitation pageId="48" pageNumber="49">Lorenz 2005</bibRefCitation>
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), have omitted the terminal
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<normalizedToken originalValue="“i”">"i"</normalizedToken>
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, thus
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<normalizedToken originalValue="“chevrolati”">"chevrolati"</normalizedToken>
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. Such a modification produces an incorrect subsequent spelling, but I accept the catalogue entries as "prevailing usage", and the latter name as a correct original spelling (
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<bibRefCitation pageId="48" pageNumber="49">International Code of Zoological Nomenclature 1999</bibRefCitation>
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: 43, art. 33.3.1).
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="50" lastPageNumber="51" pageId="49" pageNumber="50" type="notes about types, homonymy, and synonymy">
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<paragraph pageId="49" pageNumber="50">
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<pageBreakToken pageId="49" pageNumber="50" start="start">Notes</pageBreakToken>
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about types, homonymy, and synonymy.
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</paragraph>
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<paragraph pageId="49" pageNumber="50">
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The description of
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<taxonomicName class="Insecta" family="Carabidae" genus="Cymindis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymindis atrata" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="atrata">Cymindis atrata</taxonomicName>
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by
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<bibRefCitation pageId="49" pageNumber="50">Chevrolat (1835</bibRefCitation>
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: fasc. 7, No. 152) was based on a few specimens collected near Cruz Blanca, Veracruz, by
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<normalizedToken originalValue="“Sallé”">"Salle"</normalizedToken>
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.
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</paragraph>
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<paragraph lastPageId="50" lastPageNumber="51" pageId="49" pageNumber="50">
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As implicitly indicated by
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<bibRefCitation pageId="49" pageNumber="50">Dejean (1836</bibRefCitation>
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: 9), the name
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<taxonomicName class="Insecta" family="Carabidae" genus="Cymindis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymindis atrata" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="atrata">Cymindis atrata</taxonomicName>
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Chevrolat was a junior homonym of
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<taxonomicName class="Insecta" family="Carabidae" genus="Cymindis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymindis atrata" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="atrata">Cymindis atrata</taxonomicName>
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Dejean (1831, a species that was subsequently transferred to the genus
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<taxonomicName class="Insecta" family="Carabidae" genus="Inna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Inna" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="genus">Inna</taxonomicName>
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Putzeys (
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<bibRefCitation pageId="49" pageNumber="50">Chaudoir 1872</bibRefCitation>
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) and then to
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<taxonomicName class="Insecta" family="Carabidae" genus="Eucheila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eucheila" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="genus">Eucheila</taxonomicName>
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Dejean (
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<bibRefCitation author="Shpeley, D" journalOrPublisher="Insecta Mundi" pageId="57" pageNumber="58" pagination="1 - 185" title="A taxonomic review of the subtribe Pericalina (Coleoptera: Carabidae: Lebiini) in the Western Hemisphere, with descriptions of new species and notes about classification and zoogeography." volume="14" year="2001">Shpeley and Ball 2001</bibRefCitation>
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). As a replacement name, Dejean proposed
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<taxonomicName class="Insecta" family="Carabidae" genus="Cymindis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymindis chevrolatii" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="chevrolatii">Cymindis chevrolatii</taxonomicName>
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, using as a voucher specimen a male received from Louis Reiche, labeled "chevrolatii mihi/ atrata Chevrolat/ Mexico, d. Reiche " [green paper, handwritten]. This specimen is in the Chaudoir/
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<normalizedToken originalValue="Oberthür">Oberthuer</normalizedToken>
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collection, in front of the box label "nigrita Chaudoir" (MNHP). That specimen might have come originally from the Chevrolat collection to Reiche, and then to Dejean (ultimately to Chaudoir).
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<bibRefCitation pageId="49" pageNumber="50">Chaudoir (1837</bibRefCitation>
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: 8) recorded that he had received specimens from Chevrolat of
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<taxonomicName class="Insecta" family="Carabidae" genus="Cymindis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymindis atrata" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="atrata">Cymindis atrata</taxonomicName>
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, which he renamed
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<taxonomicName class="Insecta" family="Carabidae" genus="Cymindis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymindis nigrita" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="nigrita">Cymindis nigrita</taxonomicName>
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, evidently being unaware that Dejean had provided a new name in 1836. These four specimens (3 males, 1 female, each labeled "Ex Musaeo/ Chaudoir" [red print]),
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<pageBreakToken pageId="50" pageNumber="51" start="start">are</pageBreakToken>
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in front of the box label "nigrita / Chaud./ Mexique/ Cruz Blanca/
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<normalizedToken originalValue="Sallé”">Salle"</normalizedToken>
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(MNHP). As demonstrated by similarity in the male genitalia, the male lectotype is conspecific with the Dejean voucher for
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<taxonomicName class="Insecta" family="Carabidae" genus="Cymindis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymindis chevrolatii" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="chevrolatii">Cymindis chevrolatii</taxonomicName>
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, and for
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<taxonomicName class="Insecta" family="Carabidae" genus="Cymindis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymindis atrata" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="atrata">Cymindis atrata</taxonomicName>
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Chevrolat.
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</paragraph>
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<paragraph pageId="50" pageNumber="51">
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Authentic specimens of
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<taxonomicName class="Insecta" family="Carabidae" genus="Cymindis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymindis amblygona" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="amblygona">Cymindis amblygona</taxonomicName>
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Bates and
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<taxonomicName class="Insecta" family="Carabidae" genus="Cymindis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymindis angulifera" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="angulifera">Cymindis angulifera</taxonomicName>
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Bates were not located in neither The Natural History Museum (London) nor the Museum National
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<normalizedToken originalValue="d’Histoire">d'Histoire</normalizedToken>
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Naturelle (Paris), though they were sought in both of those collections that are known to house the Bates material on which his New World work was based.
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<bibRefCitation pageId="50" pageNumber="51">Bates (1883</bibRefCitation>
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: 187) noted that the features he had originally used to distinguish these two forms were exceedingly variable, even within single populations, citing as an example the series collected at Tehuacan, Mexico. Regarding
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<taxonomicName class="Insecta" family="Carabidae" genus="Cymindis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymindis amblygona" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="amblygona">Cymindis amblygona</taxonomicName>
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, he stated that "even the definition of the form as a variety is impossible". Possibly, then, he removed the type labels from what were the type specimens of these forms. Bates described two additional varieties of "
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<taxonomicName class="Insecta" family="Carabidae" genus="Cymindis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymindis atrata" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="atrata">Cymindis atrata</taxonomicName>
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":
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<taxonomicName class="Insecta" family="Carabidae" genus="Cymindis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymindis atrata subsp. ruficornis" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="subSpecies" species="atrata" subSpecies="ruficornis">Cymindis atrata ruficornis</taxonomicName>
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and
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<taxonomicName class="Insecta" family="Carabidae" genus="Cymindis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymindis atrata subsp. laevior" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="subSpecies" species="atrata" subSpecies="laevior">Cymindis atrata laevior</taxonomicName>
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. These are treated below, as separate species.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="50" pageNumber="51" type="diagnosis">
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<paragraph pageId="50" pageNumber="51">Diagnosis.</paragraph>
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<paragraph pageId="50" pageNumber="51">
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Adults of
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<taxonomicName class="Insecta" family="Carabidae" genus="Cymindis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymindis chevrolati" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="chevrolati">Cymindis chevrolati</taxonomicName>
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(Fig. 30) are distinguishable from those of other species of the limbata species group through genitalic characters: in males a distinct microtrichial patch (mp) on the basal endophallic lobe (bel) of the endophallus (Figs 34A, 36). This patch can be seen in many males through the cleared phallus with endophallus inverted, located near the apex of the phallus. From the right lateral aspect, an everted sac has the microtrichial patch located on the dorsal surface of the basal lobe of the endophallus. Female genitalia differ from other species in the short, stout form of gonocoxite 2 (gc2) (Fig. 35, A2).
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</paragraph>
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<caption pageId="50" pageNumber="51">
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||
<paragraph pageId="50" pageNumber="51">
|
||
Figure 30. Dorsal habitus and color pattern of
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Cymindis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymindis chevrolati" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="chevrolati">Cymindis chevrolati</taxonomicName>
|
||
Dejean (OBL 12.00 mm).
|
||
</paragraph>
|
||
</caption>
|
||
<caption pageId="50" pageNumber="51">
|
||
<paragraph pageId="50" pageNumber="51">
|
||
Figure 31. Pronota, dorsal aspect, of
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Cymindis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymindis chevrolati" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="chevrolati">Cymindis chevrolati</taxonomicName>
|
||
Dejean, showing intrapopulation variation in pronotal punctulation: A, typical, scattered and irregular punctation; B, less common, pronotal disc almost smooth with punctation more visible toward margins.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="51" lastPageNumber="52" pageId="50" pageNumber="51" type="description">
|
||
<paragraph pageId="50" pageNumber="51">Description.</paragraph>
|
||
<paragraph pageId="50" pageNumber="51">
|
||
With character states of subgenus
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Pinacodera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pinacodera" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="genus">Pinacodera</taxonomicName>
|
||
and chevrolati complex restricted as follows: OBL 10.3 - 13.5 mm. Length (n= 10 males, 10 females): head 1.00 - 1.24, pronotum 1.96 - 2.56, elytra 5.41 - 7.08, metepisternum 1.10 - 1.70 mm; width: head 2.04 - 2.60, pronotum 2.48 - 3.28, elytra 3.83 - 5.16, metepisternum 0.66 - 1.02 mm.
|
||
</paragraph>
|
||
<paragraph pageId="51" pageNumber="52">
|
||
<pageBreakToken pageId="51" pageNumber="52" start="start">Body</pageBreakToken>
|
||
proportions. HW/HL 1.88 - 2.31; PWM/PL 1.26 - 1.37; EL/EW 1.25- 1.43; ML/MW 1.66 - 2.00.
|
||
</paragraph>
|
||
<paragraph pageId="51" pageNumber="52">Color (Fig. 30). Dorsum of head black, rarely rufo-piceous in front of eyes; legs piceous to rufo-piceous.</paragraph>
|
||
<paragraph pageId="51" pageNumber="52">Microsculpture. Elytra with mesh pattern isodiametric, microlines shallow in basal half of elytra and shallow to absent from apical half.</paragraph>
|
||
<paragraph pageId="51" pageNumber="52">
|
||
Macrosculpture and pilosity. Dorsal punctures with setae present though short and almost not visible at 50
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
magnification. Ventral surface of head with evenly scattered setigerous punctures (bearing somewhat long setae) from behind eye laterally toward mentum. Pronotum normally with relatively evenly scattered setigerous punctures throughout (Fig. 31A), more densely so toward margins; few specimens with setigerous punctures along margin and few to no punctures on disc (Fig. 31B). Elytral epipleuron glabrous.
|
||
</paragraph>
|
||
<paragraph pageId="51" pageNumber="52">Fixed setae. Pronotum with two fixed setae along each margin. Four to six setae (typically four) along apical margin of sternum VII (Fig 3).</paragraph>
|
||
<paragraph pageId="51" pageNumber="52">Luster. Elytra glossy in basal two thirds, in some specimens slightly less so in apical third.</paragraph>
|
||
<paragraph pageId="51" pageNumber="52">Head (Fig. 37B-D). Mental tooth form varied.</paragraph>
|
||
<paragraph pageId="51" pageNumber="52">Elytra (Fig. 30). Humeri narrowly rounded.</paragraph>
|
||
<paragraph pageId="51" pageNumber="52">Hind wings. Somewhat to markedly reduced (Fig. 39). Length 1.34 - 3.29 mm, mean 2.28 mm.</paragraph>
|
||
<paragraph pageId="51" pageNumber="52">Male genitalia. Phallus (Fig. 34A) length 2.20-2.48 mm. Endophallus with microtrichial patch on basal lobe of everted sac (Fig. 36).</paragraph>
|
||
<paragraph pageId="51" pageNumber="52">Female genitalia. Gonocoxite 2 (gc2) (Fig. 35A) short and stout.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="51" pageNumber="52" type="collection notes and habitat">
|
||
<paragraph pageId="51" pageNumber="52">Collection notes and habitat.</paragraph>
|
||
<paragraph pageId="51" pageNumber="52">
|
||
The known elevational range of
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Cymindis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymindis chevrolati" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="chevrolati">Cymindis chevrolati</taxonomicName>
|
||
extends from sea level to 3400 m, though it is important to note that specimens found at or near sea level were collected from the Tres Marias Islands, the immediately adjacent mainland and one other additional record from El Ebano, San Luis Potosi, that may be mislabeled or misinterpreted by me as Ebano, San Luis Potosi. Typically this species is found further inland and at higher elevation, mostly between 2000-3000 m. Specimens were collected under and around woody debris and stones in forests of oak, pine, fir, juniper and alder, in thorn scrub, and in stands of yucca. As well, the species occupies meadow, desert and grassland habitats.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="51" pageNumber="52" type="geographical distribution">
|
||
<paragraph pageId="51" pageNumber="52">Geographical distribution</paragraph>
|
||
<paragraph pageId="51" pageNumber="52">(Fig. 41) The range of this species is restricted to Mexico, extending in the Sierra Madre Occidental from Chihuahua south to Jalisco, in the Sierra Madre Oriental from central Nuevo Leon south to Hidalgo, and in the Transvolcanic Sierra of central Mexico as far south as central Puebla.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="51" pageNumber="52" type="morphological affinities">
|
||
<paragraph pageId="51" pageNumber="52">Morphological affinities.</paragraph>
|
||
<paragraph pageId="51" pageNumber="52">
|
||
Based on genitalic characteristics and wing length states (Fig. 40), I postulate that
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Cymindis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymindis chevrolati" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="chevrolati">Cymindis chevrolati</taxonomicName>
|
||
is the closest relative of
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Cymindis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymindis laevior" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="laevior">Cymindis laevior</taxonomicName>
|
||
+
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Cymindis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymindis ruficornis" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="ruficornis">Cymindis ruficornis</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="51" pageNumber="52" type="chorological affinities">
|
||
<paragraph pageId="51" pageNumber="52">Chorological affinities.</paragraph>
|
||
<paragraph pageId="51" pageNumber="52">
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Cymindis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymindis chevrolati" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="chevrolati">Cymindis chevrolati</taxonomicName>
|
||
is sympatric in a portion of its range with
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Cymindis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymindis laevior" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="laevior">Cymindis laevior</taxonomicName>
|
||
in the southernmost portion of its range (Fig. 41). It is also sympatric with
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Cymindis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymindis punctigera subsp. punctigera" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="subSpecies" species="punctigera" subSpecies="punctigera">Cymindis punctigera punctigera</taxonomicName>
|
||
in the La Michilia area of southern Durango.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="52" pageNumber="53" type="material examined">
|
||
<paragraph pageId="52" pageNumber="53">
|
||
<pageBreakToken pageId="52" pageNumber="53" start="start">Material</pageBreakToken>
|
||
examined.
|
||
</paragraph>
|
||
<paragraph pageId="52" pageNumber="53">
|
||
I examined 662 specimens: 38 males and 26 females were dissected. For details see University of Alberta Strickland Virtual Entomology Museum Database (
|
||
<bibRefCitation pageId="52" pageNumber="53">University of Alberta 2009</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |