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<document id="BB3806A5B338BA3AE939EF8F46FEE760" ID-CLB-Dataset="25419" ID-DOI="10.11646/zootaxa.4674.1.4" ID-GBIF-Dataset="1a4215e4-efe9-441a-b421-eb004d8e9570" ID-ISSN="1175-5326" ID-Zenodo-Dep="3458436" ID-ZooBank="5F4DF8E8-FB45-451E-966C-0D4557E9A2DA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1569259911767" checkinUser="plazi" docAuthor="Rheims, Cristina A." docDate="2019" docId="A178212F0034FFAFB2C4FA27FBA7F84C" docLanguage="en" docName="zootaxa.4674.1.4.pdf" docOrigin="Zootaxa 4674 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Extraordinarius Rheims 2019, gen. nov." docType="treatment" docUuid="99DB4D62-85BD-4709-8E6A-4C27A58F7E94" docUuidSource="ZooBank" docVersion="9" lastPageNumber="85" masterDocId="5D4159570035FFADB253FFB9FFAEFFF6" masterDocTitle="Extraordinarius gen. nov., a new genus of Sparianthinae spiders (Araneae: Sparassidae) from southeastern Brazil" masterLastPageNumber="99" masterPageNumber="83" pageNumber="84" updateTime="1698756272485" updateUser="ExternalLinkService">
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<mods:title id="6CC9548854D904E4CF6521B87E42E115">Extraordinarius gen. nov., a new genus of Sparianthinae spiders (Araneae: Sparassidae) from southeastern Brazil</mods:title>
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<mods:namePart id="279AFFB1050CF0C45F368F2B64AF3D0A">Rheims, Cristina A.</mods:namePart>
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<emphasis id="1BA54C2B0034FFACB2C4FA27FE7EFA4F" bold="true" box="[151,464,1438,1465]" pageId="1" pageNumber="84">
<taxonomicName id="EED1EBBA0034FFACB2C4FA27FEF5FA4F" ID-CoL="95F6M" authority="Rheims, 2019" authorityName="Rheims" authorityYear="2019" box="[151,347,1438,1465]" class="Arachnida" family="Sparassidae" genus="Extraordinarius" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="84" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="1BA54C2B0034FFACB2C4FA27FEF5FA4F" bold="true" box="[151,347,1438,1465]" italics="true" pageId="1" pageNumber="84">Extraordinarius</emphasis>
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<taxonomicNameLabel id="0096F1500034FFACB331FA26FE7EFA4F" box="[354,464,1439,1465]" pageId="1" pageNumber="84" rank="genus">gen. nov.</taxonomicNameLabel>
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<paragraph id="296E90390034FFACB2C4FA78FC21FA2A" blockId="1.[151,911,1438,1500]" box="[151,911,1473,1500]" pageId="1" pageNumber="84">
http://zoobank.org/
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<subSubSection id="61CBC3B20034FFACB2C4F9B3FBEEF9BE" pageId="1" pageNumber="84" type="etymology">
<paragraph id="296E90390034FFACB2C4F9B3FBEEF9BE" blockId="1.[151,1437,1546,2041]" pageId="1" pageNumber="84">
<emphasis id="1BA54C2B0034FFACB2C4F9B3FEB0F9D2" bold="true" box="[151,286,1546,1572]" pageId="1" pageNumber="84">Etymology.</emphasis>
The generic name is a Latin adjective meaning “remarkable”, “extraordinary” and refers to the welldeveloped, Z-shaped, sclerotized conductor of the male palps. Gender is masculine.
</paragraph>
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<subSubSection id="61CBC3B20034FFACB294F9EBFCA1F99A" box="[199,783,1617,1644]" pageId="1" pageNumber="84" type="type_taxon">
<paragraph id="296E90390034FFACB294F9EBFCA1F99A" blockId="1.[151,1437,1546,2041]" box="[199,783,1617,1644]" pageId="1" pageNumber="84">
<emphasis id="1BA54C2B0034FFACB294F9EBFECCF99A" bold="true" box="[199,354,1617,1644]" pageId="1" pageNumber="84">
<typeStatus id="F66A2E9B0034FFACB294F9EBFEACF99A" box="[199,258,1618,1644]" pageId="1" pageNumber="84">Type</typeStatus>
species.
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<taxonomicName id="EED1EBBA0034FFACB33AF9EBFD01F99A" ID-CoL="95FQN" authority="Rheims, 2019" authorityName="Rheims" authorityYear="2019" box="[361,687,1618,1644]" class="Arachnida" family="Sparassidae" genus="Extraordinarius" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="84" phylum="Arthropoda" rank="species" species="andrematosi" status="sp. nov.">
<emphasis id="1BA54C2B0034FFACB33AF9EBFD01F99A" box="[361,687,1618,1644]" italics="true" pageId="1" pageNumber="84">Extraordinarius andrematosi</emphasis>
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<emphasis id="1BA54C2B0034FFACB0E5F9EBFCA1F99A" bold="true" box="[694,783,1618,1644]" pageId="1" pageNumber="84">
<taxonomicNameLabel id="0096F1500034FFACB0E5F9EBFCA1F99A" box="[694,783,1618,1644]" pageId="1" pageNumber="84" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="61CBC3B20034FFACB294F9CCFD4DF823" pageId="1" pageNumber="84" type="diagnosis">
<paragraph id="296E90390034FFACB294F9CCFD4DF823" blockId="1.[151,1437,1546,2041]" pageId="1" pageNumber="84">
<emphasis id="1BA54C2B0034FFACB294F9CCFEECF966" bold="true" box="[199,322,1653,1680]" pageId="1" pageNumber="84">Diagnosis.</emphasis>
Species of the genus
<taxonomicName id="EED1EBBA0034FFACB06EF9CFFD5EF966" authority="Rheims, 2019" authorityName="Rheims" authorityYear="2019" box="[573,752,1654,1680]" class="Arachnida" family="Sparassidae" genus="Extraordinarius" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="84" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="1BA54C2B0034FFACB06EF9CFFD5EF966" box="[573,752,1654,1680]" italics="true" pageId="1" pageNumber="84">Extraordinarius</emphasis>
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<emphasis id="1BA54C2B0034FFACB0AAF9CFFCCAF966" bold="true" box="[761,868,1654,1680]" pageId="1" pageNumber="84">
<taxonomicNameLabel id="0096F1500034FFACB0AAF9CFFCCAF966" box="[761,868,1654,1680]" pageId="1" pageNumber="84" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
resemble those of the non-Neotropical
<taxonomicName id="EED1EBBA0034FFACB77AF9CFFA3BF966" authorityName="Simon" authorityYear="1898" box="[1321,1429,1654,1680]" class="Arachnida" family="Sparassidae" genus="Pleorotus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="1BA54C2B0034FFACB77AF9CFFA3BF966" box="[1321,1429,1654,1680]" italics="true" pageId="1" pageNumber="84">Pleorotus</emphasis>
</taxonomicName>
,
<taxonomicName id="EED1EBBA0034FFACB2C4F923FE82F942" authorityName="Simon" authorityYear="1897" box="[151,300,1690,1716]" class="Arachnida" family="Sparassidae" genus="Rhacocnemis" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="1BA54C2B0034FFACB2C4F923FE82F942" box="[151,300,1690,1716]" italics="true" pageId="1" pageNumber="84">Rhacocnemis</emphasis>
</taxonomicName>
,
<taxonomicName id="EED1EBBA0034FFACB36EF922FE21F943" box="[317,399,1691,1717]" class="Arachnida" family="Sparassidae" genus="Stasina" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="1BA54C2B0034FFACB36EF922FE21F943" box="[317,399,1691,1717]" italics="true" pageId="1" pageNumber="84">Stasina</emphasis>
</taxonomicName>
,
<taxonomicName id="EED1EBBA0034FFACB3F2F922FE49F943" authorityName="Simon" authorityYear="1898" box="[417,487,1691,1717]" class="Arachnida" family="Sparassidae" genus="Stipax" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="1BA54C2B0034FFACB3F2F922FE49F943" box="[417,487,1691,1717]" italics="true" pageId="1" pageNumber="84">Stipax</emphasis>
</taxonomicName>
,
<taxonomicName id="EED1EBBA0034FFACB3A4F923FD2EF942" authorityName="Karsch" authorityYear="1884" box="[503,640,1690,1716]" class="Arachnida" family="Sparassidae" genus="Thelcticopis" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="1BA54C2B0034FFACB3A4F923FD2EF942" box="[503,640,1690,1716]" italics="true" pageId="1" pageNumber="84">Thelcticopis</emphasis>
</taxonomicName>
and
<taxonomicName id="EED1EBBA0034FFACB0EFF923FCE8F942" authorityName="Hirst" authorityYear="1911" box="[700,838,1690,1716]" class="Arachnida" family="Sparassidae" genus="Thomasettia" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="1BA54C2B0034FFACB0EFF923FCE8F942" box="[700,838,1690,1716]" italics="true" pageId="1" pageNumber="84">Thomasettia</emphasis>
</taxonomicName>
in having a sclerotized conductor in the male palp. They are distinguished from the latter genera by the Z-shape of the conductor, tapering retrolaterally, with tip reaching the retrolateral cymbial margin (
<figureCitation id="B1EA8CBC0034FFACB036F95BFD12F90A" box="[613,700,1761,1788]" captionStart="FIGURES 1418" captionStartId="5.[151,264,1278,1303]" captionTargetBox="[172,1408,209,1220]" captionTargetId="figure@5.[151,1435,196,1229]" captionTargetPageId="5" captionText="FIGURES 1418. E. andrematosi sp. nov. 1416 Male, holotype (IBSP 124323), left palp (14 prolateral, 15 ventral, 16 retrolateral); 1718 Female, paratype (IBSP 124310) (17 epigyne, 18 vulva). Scale lines: 1mm." figureDoi="http://doi.org/10.5281/zenodo.3458442" httpUri="https://zenodo.org/record/3458442/files/figure.png" pageId="1" pageNumber="84">Figs 15</figureCitation>
,
<figureCitation id="B1EA8CBC0034FFACB094F95BFD4BF90A" box="[711,741,1762,1788]" captionStart="FIGURES 2933" captionStartId="8.[151,264,1455,1480]" captionTargetBox="[173,1402,196,1426]" captionTargetId="figure@8.[151,1435,174,1436]" captionTargetPageId="8" captionText="FIGURES 2933. E. brucedickinsoni sp. nov. Male, holotype (IBSP 5390), left palp (29 prolateral, 30 ventral, 31 retrolateral); 3233 Female, paratype (IBSP 60018) (32 epigyne, 33 vulva). Scale lines: 1mm." figureDoi="http://doi.org/10.5281/zenodo.3458448" httpUri="https://zenodo.org/record/3458448/files/figure.png" pageId="1" pageNumber="84">30</figureCitation>
,
<figureCitation id="B1EA8CBC0034FFACB0A3F95BFCA0F90A" box="[752,782,1762,1788]" captionStart="FIGURES 4044" captionStartId="10.[151,264,1970,1995]" captionTargetBox="[161,1425,569,1932]" captionTargetId="figure@10.[151,1435,553,1950]" captionTargetPageId="10" captionText="FIGURES 4044. E. klausmeinei sp. nov., male, holotype (IBSP 60005) 4041 Habitus (40 dorsal, 41 ventral); 4244 Left palp (42 prolateral, 43 ventral, 44 retrolateral). Scale lines: 1011: 2 mm; 1214: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3458452" httpUri="https://zenodo.org/record/3458452/files/figure.png" pageId="1" pageNumber="84">43</figureCitation>
,
<figureCitation id="B1EA8CBC0034FFACB149F958FC94F90A" box="[794,826,1761,1788]" captionStart="FIGURES 5458" captionStartId="13.[151,264,1969,1994]" captionTargetBox="[163,1416,855,1942]" captionTargetId="figure@13.[151,1436,841,1950]" captionTargetPageId="13" captionText="FIGURES 5458. E. rickalleni sp. nov. 5456 Male, holotype (IBSP 121338), left palp (54 prolateral, 55 ventral, 56 retrolateral); 5758 female, paratype (IBSP 121338) (57 epigyne, 58 vulva). Scale lines: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3458458" httpUri="https://zenodo.org/record/3458458/files/figure.png" pageId="1" pageNumber="84">55</figureCitation>
). They are further distinguished from all Sparianthinae genera by the median apophysis arising from tegulum at 5:306:00 oclock position (between 14 oclock in all other genera) and by the presence of a TBE laminar, distally rounded, with hyaline margins (
<figureCitation id="B1EA8CBC0034FFACB6C7F893FB45F8B2" box="[1172,1259,1834,1860]" captionStart="FIGURES 1924" captionStartId="6.[151,264,1215,1240]" captionTargetBox="[158,1428,181,1190]" captionTargetId="figure@6.[151,1435,174,1196]" captionTargetPageId="6" captionText="FIGURES 1924. E. andrematosi sp. nov. 1921 Male, left palp, holotype (IBSP 124323) (19 prolateral, 20 ventral, 21 retrolateral); 2224 Female, paratype (IBSP 124310) (22 epigyne, 23 vulva, 24 schematic course of internal duct system). FD—fertilization duct; LL—lateral lobe; MS—median septum; Sc—scape. Scale lines: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3458444" httpUri="https://zenodo.org/record/3458444/files/figure.png" pageId="1" pageNumber="84">Figs 20</figureCitation>
,
<figureCitation id="B1EA8CBC0034FFACB6A6F890FABDF8B2" box="[1269,1299,1833,1860]" captionStart="FIGURES 3439" captionStartId="9.[151,264,1301,1326]" captionTargetBox="[166,1424,182,1276]" captionTargetId="figure@9.[151,1435,174,1282]" captionTargetPageId="9" captionText="FIGURES 3439. E. brucedickinsoni sp. nov. 3436 Male, left palp, holotype (IBSP 5390) (34 prolateral, 35 ventral, 36 retrolateral); 3739 Female, paratype (IBSP 60018) (37 epigyne, 38 vulva, 39 schematic course of internal duct system). Scale lines: 1mm." figureDoi="http://doi.org/10.5281/zenodo.3458450" httpUri="https://zenodo.org/record/3458450/files/figure.png" pageId="1" pageNumber="84">35</figureCitation>
,
<figureCitation id="B1EA8CBC0034FFACB74DF893FAC8F8B2" box="[1310,1382,1834,1860]" captionStart="FIGURES 4549" captionStartId="11.[151,264,1369,1394]" captionTargetBox="[163,1425,182,1341]" captionTargetId="figure@11.[151,1436,174,1349]" captionTargetPageId="11" captionText="FIGURES 4549. E. klausmeinei sp. nov., male, holotype (IBSP 60005), 4547 left palp (45 prolateral, 46 ventral, 47 retrolateral); 4849 right palp, expanded (48 retrolateral, 49 prolateral). C = conductor; E = embolus; MA = median apophysis; RbP = retrobasal process; RTA = retrolateral tibial apophysis; ST = subtegulum; T = tegulum; TBC = tegular projection at base of conductor; TBE = tegular projection at base of embolus; VTA = ventral tibial apophysis. Scale lines: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3458454" httpUri="https://zenodo.org/record/3458454/files/figure.png" pageId="1" pageNumber="84">46, 49</figureCitation>
,
<figureCitation id="B1EA8CBC0034FFACB723F893FA3EF8B3" box="[1392,1424,1834,1861]" captionStart="FIGURES 5964" captionStartId="14.[151,264,1637,1662]" captionTargetBox="[160,1423,326,1446]" captionTargetId="figure@14.[151,1435,319,1452]" captionTargetPageId="14" captionText="FIGURES 5964. E. rickalleni sp. nov. 5961 Male, holotype (IBSP 121338), left palp (59 prolateral, 60 ventral, 61 retrolateral); 6264 Female, paratype (IBSP 121553) (62 epigyne, 63 vulva, 64 schematic course of internal duct system. Scale lines: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3458460" httpUri="https://zenodo.org/record/3458460/files/figure.png" pageId="1" pageNumber="84">60</figureCitation>
). Females resemble those of
<taxonomicName id="EED1EBBA0034FFACB390F8F7FDE7F89E" authorityName="Rheims &amp; Alayon" authorityYear="2018" box="[451,585,1870,1896]" class="Arachnida" family="Sparassidae" genus="Diminutella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="1BA54C2B0034FFACB390F8F7FDE7F89E" box="[451,585,1870,1896]" italics="true" pageId="1" pageNumber="84">Diminutella</emphasis>
</taxonomicName>
by the vulva with internal ducts running anteriorly as a double helix (
<bibRefCitation id="4D40EDC80034FFACB716F8F7FEE1F87A" author="Rheims, C. A. &amp; Alayon, G." pageId="1" pageNumber="84" pagination="13 - 22" refId="ref7929" refString="Rheims, C. A. &amp; Alayon, G. (2018) A new genus of minute Sparianthinae spiders from the Caribbean (Araneae, Sparassidae). ZooKeys, 742, 13 - 22. https: // doi. org / 10.3897 / zookeys. 742.21819" type="journal article" year="2018">Rheims &amp; Alayón, 2018</bibRefCitation>
, figs 4EF). They are distinguished from the latter genus in having the duct system encapsulated, with an additional convoluted part, not organized in a double helix, close to fertilization ducts, and by lacking a blind ending projection (
<figureCitation id="B1EA8CBC0034FFACB3F8F803FD83F822" box="[427,557,1978,2004]" captionStart="FIGURES 1924" captionStartId="6.[151,264,1215,1240]" captionTargetBox="[158,1428,181,1190]" captionTargetId="figure@6.[151,1435,174,1196]" captionTargetPageId="6" captionText="FIGURES 1924. E. andrematosi sp. nov. 1921 Male, left palp, holotype (IBSP 124323) (19 prolateral, 20 ventral, 21 retrolateral); 2224 Female, paratype (IBSP 124310) (22 epigyne, 23 vulva, 24 schematic course of internal duct system). FD—fertilization duct; LL—lateral lobe; MS—median septum; Sc—scape. Scale lines: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3458444" httpUri="https://zenodo.org/record/3458444/files/figure.png" pageId="1" pageNumber="84">Figs 2324</figureCitation>
,
<figureCitation id="B1EA8CBC0034FFACB06BF803FD2FF822" box="[568,641,1978,2004]" captionStart="FIGURES 3439" captionStartId="9.[151,264,1301,1326]" captionTargetBox="[166,1424,182,1276]" captionTargetId="figure@9.[151,1435,174,1282]" captionTargetPageId="9" captionText="FIGURES 3439. E. brucedickinsoni sp. nov. 3436 Male, left palp, holotype (IBSP 5390) (34 prolateral, 35 ventral, 36 retrolateral); 3739 Female, paratype (IBSP 60018) (37 epigyne, 38 vulva, 39 schematic course of internal duct system). Scale lines: 1mm." figureDoi="http://doi.org/10.5281/zenodo.3458450" httpUri="https://zenodo.org/record/3458450/files/figure.png" pageId="1" pageNumber="84">3839</figureCitation>
,
<figureCitation id="B1EA8CBC0034FFACB0DFF803FD79F823" box="[652,727,1978,2005]" captionStart="FIGURES 5964" captionStartId="14.[151,264,1637,1662]" captionTargetBox="[160,1423,326,1446]" captionTargetId="figure@14.[151,1435,319,1452]" captionTargetPageId="14" captionText="FIGURES 5964. E. rickalleni sp. nov. 5961 Male, holotype (IBSP 121338), left palp (59 prolateral, 60 ventral, 61 retrolateral); 6264 Female, paratype (IBSP 121553) (62 epigyne, 63 vulva, 64 schematic course of internal duct system. Scale lines: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3458460" httpUri="https://zenodo.org/record/3458460/files/figure.png" pageId="1" pageNumber="84">6364</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="61CBC3B20034FFAFB294F864FD40FAF9" lastPageId="2" lastPageNumber="85" pageId="1" pageNumber="84" type="description">
<paragraph id="296E90390034FFAFB294F864FD40FAF9" blockId="1.[151,1437,1546,2041]" lastBlockId="2.[151,1437,152,1979]" lastPageId="2" lastPageNumber="85" pageId="1" pageNumber="84">
<emphasis id="1BA54C2B0034FFACB294F864FEF7F80E" bold="true" box="[199,345,2013,2040]" pageId="1" pageNumber="84">Description.</emphasis>
Total length of males 9.711.8 (n = 6), of females 11.014.3 (n = 6). Prosoma longer than wide; cephalic region higher than thoracic region; fovea slightly conspicuous on posterior third of prosoma. Eight eyes arranged in two rows, the anterior row straight, the posterior straight or slightly procurved; AME larger than ALE and more separated from each other than from laterals; PME smaller than PLE and equidistant (
<figureCitation id="B1EA8CBC0037FFAFB6B5FF59FAC5FF0D" box="[1254,1387,224,251]" captionStart="FIGURES 1013" captionStartId="4.[151,264,1971,1996]" captionTargetBox="[159,1425,388,1941]" captionTargetId="figure@4.[151,1431,373,1951]" captionTargetPageId="4" captionText="FIGURES 1013. E. andrematosi sp. nov., habitus. 1011 Male, holotype (IBSP 124323) (10 dorsal, 11 ventral); 1213 Female, paratype (IBSP 62064) (12 dorsal, 13 ventral). Scale lines: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3458440" httpUri="https://zenodo.org/record/3458440/files/figure.png" pageId="2" pageNumber="85">Figs 10, 12</figureCitation>
,
<figureCitation id="B1EA8CBC0037FFAFB72AFF59FF1BFEE9" captionStart="FIGURES 2528" captionStartId="7.[151,264,1984,2009]" captionTargetBox="[174,1414,382,1950]" captionTargetId="figure@7.[162,1426,373,1965]" captionTargetPageId="7" captionText="FIGURES 2528. E. brucedickinsoni sp. nov., habitus 2526 Male, holotype (IBSP 5390) (25 dorsal, 26 ventral); 2728 Female, holotype (IBSP 60018) (27 dorsal, 28 ventral). Scale lines: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3458446" httpUri="https://zenodo.org/record/3458446/files/figure.png" pageId="2" pageNumber="85">25, 27</figureCitation>
,
<figureCitation id="B1EA8CBC0037FFAFB291FEBCFF4FFEE9" box="[194,225,261,287]" captionStart="FIGURES 4044" captionStartId="10.[151,264,1970,1995]" captionTargetBox="[161,1425,569,1932]" captionTargetId="figure@10.[151,1435,553,1950]" captionTargetPageId="10" captionText="FIGURES 4044. E. klausmeinei sp. nov., male, holotype (IBSP 60005) 4041 Habitus (40 dorsal, 41 ventral); 4244 Left palp (42 prolateral, 43 ventral, 44 retrolateral). Scale lines: 1011: 2 mm; 1214: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3458452" httpUri="https://zenodo.org/record/3458452/files/figure.png" pageId="2" pageNumber="85">40</figureCitation>
,
<figureCitation id="B1EA8CBC0037FFAFB2BEFEBDFE97FEE9" box="[237,313,260,287]" captionStart="FIGURES 5053" captionStartId="12.[151,264,1959,1984]" captionTargetBox="[167,1420,364,1925]" captionTargetId="figure@12.[151,1435,352,1940]" captionTargetPageId="12" captionText="FIGURES 5053. E. rickalleni sp. nov., habitus 5051 Male, holotype (IBSP 121338) (50 dorsal, 51 ventral); female, paratype (IBSP 121338) (52 dorsal, 53 ventral). Scale lines: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3458456" httpUri="https://zenodo.org/record/3458456/files/figure.png" pageId="2" pageNumber="85">50, 52</figureCitation>
). Clypeus low, less than AME diameter. Chelicerae longer than wide, with three promarginal teeth, the basal one smallest, and two small retromarginal teeth; intermarginal denticles absent; one single retromarginal escort setae at the base of fang (
<figureCitation id="B1EA8CBC0037FFAFB3ADFEF5FD91FE91" box="[510,575,332,359]" captionStart="FIGURES 19" captionStartId="3.[151,264,1790,1815]" captionTargetBox="[151,1436,590,1771]" captionTargetId="figure@3.[151,1436,590,1771]" captionTargetPageId="3" captionText="FIGURES 19. E. andrematosi sp. nov. (1, 3, 6, 8 female IBSP 62064, 4 male IBSP 124323); E. rickalleni sp. nov. (2, 5 female IBSP 121533, 7, 9 male IBSP 121338).1 Female, left chelicerae, ventral view (arrow points to retrolateral escort setae); 2 Female, right endite, serrula, ventral view; 3 Female, left palp, claw, prolateral view; 4 Male, left leg I, metatarsus, trilobate membrane, dorsal view; 5 Female, left leg III, tarsus, tricobothria, dorsal view; 6 Female, left leg I, tarsus, tarsal organ, dorsal view; 7 Male, left leg II, tarsal claws, prolateral view; 8 Female, left leg III, tarsal claws, prolateral view; 9 Male, epiandrium, ventral view (arrow points to epiandrous spigots). DP = distal plate; PP = proximal plate." figureDoi="http://doi.org/10.5281/zenodo.3458438" httpUri="https://zenodo.org/record/3458438/files/figure.png" pageId="2" pageNumber="85">Fig. 1</figureCitation>
). Labium slightly longer than wide. Endites convergent with dense scopulae on internal margin. Serrula with a single row of denticles (
<figureCitation id="B1EA8CBC0037FFAFB110FEC9FC22FE7D" box="[835,908,368,395]" captionStart="FIGURES 19" captionStartId="3.[151,264,1790,1815]" captionTargetBox="[151,1436,590,1771]" captionTargetId="figure@3.[151,1436,590,1771]" captionTargetPageId="3" captionText="FIGURES 19. E. andrematosi sp. nov. (1, 3, 6, 8 female IBSP 62064, 4 male IBSP 124323); E. rickalleni sp. nov. (2, 5 female IBSP 121533, 7, 9 male IBSP 121338).1 Female, left chelicerae, ventral view (arrow points to retrolateral escort setae); 2 Female, right endite, serrula, ventral view; 3 Female, left palp, claw, prolateral view; 4 Male, left leg I, metatarsus, trilobate membrane, dorsal view; 5 Female, left leg III, tarsus, tricobothria, dorsal view; 6 Female, left leg I, tarsus, tarsal organ, dorsal view; 7 Male, left leg II, tarsal claws, prolateral view; 8 Female, left leg III, tarsal claws, prolateral view; 9 Male, epiandrium, ventral view (arrow points to epiandrous spigots). DP = distal plate; PP = proximal plate." figureDoi="http://doi.org/10.5281/zenodo.3458438" httpUri="https://zenodo.org/record/3458438/files/figure.png" pageId="2" pageNumber="85">Fig. 2</figureCitation>
). Sternum longer than wide. Female pedipalp with single pectinate claw with 67 short, gently curved teeth (
<figureCitation id="B1EA8CBC0037FFAFB10DFE2DFC0DFE59" box="[862,931,404,431]" captionStart="FIGURES 19" captionStartId="3.[151,264,1790,1815]" captionTargetBox="[151,1436,590,1771]" captionTargetId="figure@3.[151,1436,590,1771]" captionTargetPageId="3" captionText="FIGURES 19. E. andrematosi sp. nov. (1, 3, 6, 8 female IBSP 62064, 4 male IBSP 124323); E. rickalleni sp. nov. (2, 5 female IBSP 121533, 7, 9 male IBSP 121338).1 Female, left chelicerae, ventral view (arrow points to retrolateral escort setae); 2 Female, right endite, serrula, ventral view; 3 Female, left palp, claw, prolateral view; 4 Male, left leg I, metatarsus, trilobate membrane, dorsal view; 5 Female, left leg III, tarsus, tricobothria, dorsal view; 6 Female, left leg I, tarsus, tarsal organ, dorsal view; 7 Male, left leg II, tarsal claws, prolateral view; 8 Female, left leg III, tarsal claws, prolateral view; 9 Male, epiandrium, ventral view (arrow points to epiandrous spigots). DP = distal plate; PP = proximal plate." figureDoi="http://doi.org/10.5281/zenodo.3458438" httpUri="https://zenodo.org/record/3458438/files/figure.png" pageId="2" pageNumber="85">Fig. 3</figureCitation>
). Legs laterigrade, 4123 or 4213. Trochanter smooth. Metatarsi IIV with distal dorsal trilobate membrane with median hook much smaller than lateral projections (
<figureCitation id="B1EA8CBC0037FFAFB28CFE65FE88FE01" box="[223,294,476,503]" captionStart="FIGURES 19" captionStartId="3.[151,264,1790,1815]" captionTargetBox="[151,1436,590,1771]" captionTargetId="figure@3.[151,1436,590,1771]" captionTargetPageId="3" captionText="FIGURES 19. E. andrematosi sp. nov. (1, 3, 6, 8 female IBSP 62064, 4 male IBSP 124323); E. rickalleni sp. nov. (2, 5 female IBSP 121533, 7, 9 male IBSP 121338).1 Female, left chelicerae, ventral view (arrow points to retrolateral escort setae); 2 Female, right endite, serrula, ventral view; 3 Female, left palp, claw, prolateral view; 4 Male, left leg I, metatarsus, trilobate membrane, dorsal view; 5 Female, left leg III, tarsus, tricobothria, dorsal view; 6 Female, left leg I, tarsus, tarsal organ, dorsal view; 7 Male, left leg II, tarsal claws, prolateral view; 8 Female, left leg III, tarsal claws, prolateral view; 9 Male, epiandrium, ventral view (arrow points to epiandrous spigots). DP = distal plate; PP = proximal plate." figureDoi="http://doi.org/10.5281/zenodo.3458438" httpUri="https://zenodo.org/record/3458438/files/figure.png" pageId="2" pageNumber="85">Fig. 4</figureCitation>
). Tarsi and anterior half of metatarsi scopulate. Trichobothria present on dorsal tibiae, metatarsi and tarsi, arranged in several rows that converge to one on metatarsi. Trichobothria with well-differentiated proximal and distal plates; proximal plate with several transverse ridges; distal plate with transverse ridge; trichobothrial setae thickened in a basal bulb (
<figureCitation id="B1EA8CBC0037FFAFB390FDF1FDA7FD95" box="[451,521,584,611]" captionStart="FIGURES 19" captionStartId="3.[151,264,1790,1815]" captionTargetBox="[151,1436,590,1771]" captionTargetId="figure@3.[151,1436,590,1771]" captionTargetPageId="3" captionText="FIGURES 19. E. andrematosi sp. nov. (1, 3, 6, 8 female IBSP 62064, 4 male IBSP 124323); E. rickalleni sp. nov. (2, 5 female IBSP 121533, 7, 9 male IBSP 121338).1 Female, left chelicerae, ventral view (arrow points to retrolateral escort setae); 2 Female, right endite, serrula, ventral view; 3 Female, left palp, claw, prolateral view; 4 Male, left leg I, metatarsus, trilobate membrane, dorsal view; 5 Female, left leg III, tarsus, tricobothria, dorsal view; 6 Female, left leg I, tarsus, tarsal organ, dorsal view; 7 Male, left leg II, tarsal claws, prolateral view; 8 Female, left leg III, tarsal claws, prolateral view; 9 Male, epiandrium, ventral view (arrow points to epiandrous spigots). DP = distal plate; PP = proximal plate." figureDoi="http://doi.org/10.5281/zenodo.3458438" httpUri="https://zenodo.org/record/3458438/files/figure.png" pageId="2" pageNumber="85">Fig. 5</figureCitation>
). Tarsal organ capsulate, with round to oval opening (
<figureCitation id="B1EA8CBC0037FFAFB623FDF1FB18FD95" box="[1136,1206,584,611]" captionStart="FIGURES 19" captionStartId="3.[151,264,1790,1815]" captionTargetBox="[151,1436,590,1771]" captionTargetId="figure@3.[151,1436,590,1771]" captionTargetPageId="3" captionText="FIGURES 19. E. andrematosi sp. nov. (1, 3, 6, 8 female IBSP 62064, 4 male IBSP 124323); E. rickalleni sp. nov. (2, 5 female IBSP 121533, 7, 9 male IBSP 121338).1 Female, left chelicerae, ventral view (arrow points to retrolateral escort setae); 2 Female, right endite, serrula, ventral view; 3 Female, left palp, claw, prolateral view; 4 Male, left leg I, metatarsus, trilobate membrane, dorsal view; 5 Female, left leg III, tarsus, tricobothria, dorsal view; 6 Female, left leg I, tarsus, tarsal organ, dorsal view; 7 Male, left leg II, tarsal claws, prolateral view; 8 Female, left leg III, tarsal claws, prolateral view; 9 Male, epiandrium, ventral view (arrow points to epiandrous spigots). DP = distal plate; PP = proximal plate." figureDoi="http://doi.org/10.5281/zenodo.3458438" httpUri="https://zenodo.org/record/3458438/files/figure.png" pageId="2" pageNumber="85">Fig. 6</figureCitation>
), located distally on tarsi. Tarsi with pair of pectinate claws, with 78 gently curved teeth, closer together on claws of legs IIIIV, and claw tufts with deeply indented setae (
<figureCitation id="B1EA8CBC0037FFAFB018FD29FD1CFD5D" box="[587,690,656,683]" captionStart="FIGURES 19" captionStartId="3.[151,264,1790,1815]" captionTargetBox="[151,1436,590,1771]" captionTargetId="figure@3.[151,1436,590,1771]" captionTargetPageId="3" captionText="FIGURES 19. E. andrematosi sp. nov. (1, 3, 6, 8 female IBSP 62064, 4 male IBSP 124323); E. rickalleni sp. nov. (2, 5 female IBSP 121533, 7, 9 male IBSP 121338).1 Female, left chelicerae, ventral view (arrow points to retrolateral escort setae); 2 Female, right endite, serrula, ventral view; 3 Female, left palp, claw, prolateral view; 4 Male, left leg I, metatarsus, trilobate membrane, dorsal view; 5 Female, left leg III, tarsus, tricobothria, dorsal view; 6 Female, left leg I, tarsus, tarsal organ, dorsal view; 7 Male, left leg II, tarsal claws, prolateral view; 8 Female, left leg III, tarsal claws, prolateral view; 9 Male, epiandrium, ventral view (arrow points to epiandrous spigots). DP = distal plate; PP = proximal plate." figureDoi="http://doi.org/10.5281/zenodo.3458438" httpUri="https://zenodo.org/record/3458438/files/figure.png" pageId="2" pageNumber="85">Figs 78</figureCitation>
). Spination pattern in males: femora IIII: p1-1-1; d0-1-1; r1-1-1; femur IV: p1-1-1; d0-1-1; r0-0-1; patellae IIV: 0; tibiae IIV: p1-0-1; r1-0-1;
<date id="5D6FB6F90037FFAFB1AAFD0CFB81FD39" box="[1017,1071,693,719]" pageId="2" pageNumber="85">v2-2</date>
-2; metatarsi III: p1-1-1; r1-1-0;
<date id="5D6FB6F90037FFAFB2C4FD60FF18FD05" box="[151,182,729,755]" pageId="2" pageNumber="85">v2</date>
-0-0; metatarsi IIIIV: p1-1-1; r1-1-1;
<date id="5D6FB6F90037FFAFB009FD60FD3EFD05" box="[602,656,729,755]" pageId="2" pageNumber="85">v2-2</date>
-0; palp: femur p0-0-1; d0-1-2; patella 0; tibia p1-0; in females: femora III: p1-1-0; d0-1-1; r1-0-0; femur III: p1-1-1; d1-0-0; r1-0-0; femur IV: p1-1-0; d1-1-0; r0-0-1; patellae IIV: 0; tibiae IIII
<date id="5D6FB6F90037FFAFB344FC98FEE3FCCD" box="[279,333,801,827]" pageId="2" pageNumber="85">v2-2</date>
-2; tibia IV: p1-0-1; r1-0-1;
<date id="5D6FB6F90037FFAFB0D2FC98FD35FCCD" box="[641,667,801,827]" pageId="2" pageNumber="85">v1</date>
p-2-2; metatarsi III p1-1-1; r1-1-0;
<date id="5D6FB6F90037FFAFB666FC98FBFDFCCD" box="[1077,1107,801,827]" pageId="2" pageNumber="85">v2</date>
-0-0; metatarsi IIIIV p1-1-1; r1-1-1;
<date id="5D6FB6F90037FFAFB2BDFCFCFE8AFCA9" box="[238,292,837,863]" pageId="2" pageNumber="85">v2-2</date>
-0; palp: femur p0-0-1; d0-1-2; patella p1; r1; tibia p2-1; d1-1; r1-1; tarsus: p2-1; r2-1. Opisthosoma oval, longer than wide, with tegument ring at posterior end, close to base of spinnerets (
<figureCitation id="B1EA8CBC0037FFAFB625FCD1FB63FC75" box="[1142,1229,872,899]" captionStart="FIGURES 1013" captionStartId="4.[151,264,1971,1996]" captionTargetBox="[159,1425,388,1941]" captionTargetId="figure@4.[151,1431,373,1951]" captionTargetPageId="4" captionText="FIGURES 1013. E. andrematosi sp. nov., habitus. 1011 Male, holotype (IBSP 124323) (10 dorsal, 11 ventral); 1213 Female, paratype (IBSP 62064) (12 dorsal, 13 ventral). Scale lines: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3458440" httpUri="https://zenodo.org/record/3458440/files/figure.png" pageId="2" pageNumber="85">Figs 13</figureCitation>
,
<figureCitation id="B1EA8CBC0037FFAFB68AFCD0FB59FC75" box="[1241,1271,873,899]" captionStart="FIGURES 2528" captionStartId="7.[151,264,1984,2009]" captionTargetBox="[174,1414,382,1950]" captionTargetId="figure@7.[162,1426,373,1965]" captionTargetPageId="7" captionText="FIGURES 2528. E. brucedickinsoni sp. nov., habitus 2526 Male, holotype (IBSP 5390) (25 dorsal, 26 ventral); 2728 Female, holotype (IBSP 60018) (27 dorsal, 28 ventral). Scale lines: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3458446" httpUri="https://zenodo.org/record/3458446/files/figure.png" pageId="2" pageNumber="85">27</figureCitation>
,
<figureCitation id="B1EA8CBC0037FFAFB757FCD0FA8DFC75" box="[1284,1315,873,899]" captionStart="FIGURES 4044" captionStartId="10.[151,264,1970,1995]" captionTargetBox="[161,1425,569,1932]" captionTargetId="figure@10.[151,1435,553,1950]" captionTargetPageId="10" captionText="FIGURES 4044. E. klausmeinei sp. nov., male, holotype (IBSP 60005) 4041 Habitus (40 dorsal, 41 ventral); 4244 Left palp (42 prolateral, 43 ventral, 44 retrolateral). Scale lines: 1011: 2 mm; 1214: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3458452" httpUri="https://zenodo.org/record/3458452/files/figure.png" pageId="2" pageNumber="85">40</figureCitation>
,
<figureCitation id="B1EA8CBC0037FFAFB77CFCD1FAE0FC75" box="[1327,1358,872,899]" captionStart="FIGURES 5053" captionStartId="12.[151,264,1959,1984]" captionTargetBox="[167,1420,364,1925]" captionTargetId="figure@12.[151,1435,352,1940]" captionTargetPageId="12" captionText="FIGURES 5053. E. rickalleni sp. nov., habitus 5051 Male, holotype (IBSP 121338) (50 dorsal, 51 ventral); female, paratype (IBSP 121338) (52 dorsal, 53 ventral). Scale lines: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3458456" httpUri="https://zenodo.org/record/3458456/files/figure.png" pageId="2" pageNumber="85">53</figureCitation>
). Male epiandrium with epiandrous spigots arranged in scattered bunches (
<figureCitation id="B1EA8CBC0037FFAFB1D3FC35FC6AFC51" box="[896,964,908,935]" captionStart="FIGURES 19" captionStartId="3.[151,264,1790,1815]" captionTargetBox="[151,1436,590,1771]" captionTargetId="figure@3.[151,1436,590,1771]" captionTargetPageId="3" captionText="FIGURES 19. E. andrematosi sp. nov. (1, 3, 6, 8 female IBSP 62064, 4 male IBSP 124323); E. rickalleni sp. nov. (2, 5 female IBSP 121533, 7, 9 male IBSP 121338).1 Female, left chelicerae, ventral view (arrow points to retrolateral escort setae); 2 Female, right endite, serrula, ventral view; 3 Female, left palp, claw, prolateral view; 4 Male, left leg I, metatarsus, trilobate membrane, dorsal view; 5 Female, left leg III, tarsus, tricobothria, dorsal view; 6 Female, left leg I, tarsus, tarsal organ, dorsal view; 7 Male, left leg II, tarsal claws, prolateral view; 8 Female, left leg III, tarsal claws, prolateral view; 9 Male, epiandrium, ventral view (arrow points to epiandrous spigots). DP = distal plate; PP = proximal plate." figureDoi="http://doi.org/10.5281/zenodo.3458438" httpUri="https://zenodo.org/record/3458438/files/figure.png" pageId="2" pageNumber="85">Fig. 9</figureCitation>
). Six spinnerets: anterior lateral spinnerets contiguous, conical and bi-segmented. Basal segment elongate and cylindrical, distal segment short and truncated. Posterior median spinnerets conical and short. Posterior lateral spinnerets conical and bi-segmented. Basal segment elongate and cylindrical, distal segment short and truncated. Male palp: tibia shorter than one third of cymbium length, with one prolateral spine; VTA slightly displaced retrolaterally; RTA single, arising distally from tibia; cymbium with well-developed retrobasal process, large round alveolus and small, rounded dorsal scopulae; subtegulum ring-shaped; tegulum with large laminar TBE and sclerotized, concave TBC; median apophysis elongate, apically hook-shaped; conductor large, Z-shaped, with tip reaching the retrolateral cymbial margin; embolus filiform, arising from tegulum between 79 oclock positions (
<figureCitation id="B1EA8CBC0037FFAFB0C6FB15FD45FB31" box="[661,747,1196,1223]" captionStart="FIGURES 1924" captionStartId="6.[151,264,1215,1240]" captionTargetBox="[158,1428,181,1190]" captionTargetId="figure@6.[151,1435,174,1196]" captionTargetPageId="6" captionText="FIGURES 1924. E. andrematosi sp. nov. 1921 Male, left palp, holotype (IBSP 124323) (19 prolateral, 20 ventral, 21 retrolateral); 2224 Female, paratype (IBSP 124310) (22 epigyne, 23 vulva, 24 schematic course of internal duct system). FD—fertilization duct; LL—lateral lobe; MS—median septum; Sc—scape. Scale lines: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3458444" httpUri="https://zenodo.org/record/3458444/files/figure.png" pageId="2" pageNumber="85">Figs 20</figureCitation>
,
<figureCitation id="B1EA8CBC0037FFAFB0A6FB15FCBDFB31" box="[757,787,1196,1223]" captionStart="FIGURES 3439" captionStartId="9.[151,264,1301,1326]" captionTargetBox="[166,1424,182,1276]" captionTargetId="figure@9.[151,1435,174,1282]" captionTargetPageId="9" captionText="FIGURES 3439. E. brucedickinsoni sp. nov. 3436 Male, left palp, holotype (IBSP 5390) (34 prolateral, 35 ventral, 36 retrolateral); 3739 Female, paratype (IBSP 60018) (37 epigyne, 38 vulva, 39 schematic course of internal duct system). Scale lines: 1mm." figureDoi="http://doi.org/10.5281/zenodo.3458450" httpUri="https://zenodo.org/record/3458450/files/figure.png" pageId="2" pageNumber="85">35</figureCitation>
,
<figureCitation id="B1EA8CBC0037FFAFB14DFB15FC21FB31" box="[798,911,1196,1223]" captionStart="FIGURES 4549" captionStartId="11.[151,264,1369,1394]" captionTargetBox="[163,1425,182,1341]" captionTargetId="figure@11.[151,1436,174,1349]" captionTargetPageId="11" captionText="FIGURES 4549. E. klausmeinei sp. nov., male, holotype (IBSP 60005), 4547 left palp (45 prolateral, 46 ventral, 47 retrolateral); 4849 right palp, expanded (48 retrolateral, 49 prolateral). C = conductor; E = embolus; MA = median apophysis; RbP = retrobasal process; RTA = retrolateral tibial apophysis; ST = subtegulum; T = tegulum; TBC = tegular projection at base of conductor; TBE = tegular projection at base of embolus; VTA = ventral tibial apophysis. Scale lines: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3458454" httpUri="https://zenodo.org/record/3458454/files/figure.png" pageId="2" pageNumber="85">46, 4849</figureCitation>
,
<figureCitation id="B1EA8CBC0037FFAFB1CAFB15FC17FB31" box="[921,953,1196,1223]" captionStart="FIGURES 5964" captionStartId="14.[151,264,1637,1662]" captionTargetBox="[160,1423,326,1446]" captionTargetId="figure@14.[151,1435,319,1452]" captionTargetPageId="14" captionText="FIGURES 5964. E. rickalleni sp. nov. 5961 Male, holotype (IBSP 121338), left palp (59 prolateral, 60 ventral, 61 retrolateral); 6264 Female, paratype (IBSP 121553) (62 epigyne, 63 vulva, 64 schematic course of internal duct system. Scale lines: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3458460" httpUri="https://zenodo.org/record/3458460/files/figure.png" pageId="2" pageNumber="85">60</figureCitation>
). Female epigyne: divided into lateral lobes and median septum; lateral lobes simple, without projections, partially covering the median septum; median septum bearing short scape (
<figureCitation id="B1EA8CBC0037FFAFB32DFB4DFE78FAF9" box="[382,470,1268,1295]" captionStart="FIGURES 1924" captionStartId="6.[151,264,1215,1240]" captionTargetBox="[158,1428,181,1190]" captionTargetId="figure@6.[151,1435,174,1196]" captionTargetPageId="6" captionText="FIGURES 1924. E. andrematosi sp. nov. 1921 Male, left palp, holotype (IBSP 124323) (19 prolateral, 20 ventral, 21 retrolateral); 2224 Female, paratype (IBSP 124310) (22 epigyne, 23 vulva, 24 schematic course of internal duct system). FD—fertilization duct; LL—lateral lobe; MS—median septum; Sc—scape. Scale lines: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3458444" httpUri="https://zenodo.org/record/3458444/files/figure.png" pageId="2" pageNumber="85">Figs 22</figureCitation>
,
<figureCitation id="B1EA8CBC0037FFAFB3B2FB4CFE50FAF9" box="[481,510,1269,1295]" captionStart="FIGURES 3439" captionStartId="9.[151,264,1301,1326]" captionTargetBox="[166,1424,182,1276]" captionTargetId="figure@9.[151,1435,174,1282]" captionTargetPageId="9" captionText="FIGURES 3439. E. brucedickinsoni sp. nov. 3436 Male, left palp, holotype (IBSP 5390) (34 prolateral, 35 ventral, 36 retrolateral); 3739 Female, paratype (IBSP 60018) (37 epigyne, 38 vulva, 39 schematic course of internal duct system). Scale lines: 1mm." figureDoi="http://doi.org/10.5281/zenodo.3458450" httpUri="https://zenodo.org/record/3458450/files/figure.png" pageId="2" pageNumber="85">37</figureCitation>
) or smooth (
<figureCitation id="B1EA8CBC0037FFAFB0DEFB4DFD4CFAF9" box="[653,738,1268,1295]" captionStart="FIGURES 5964" captionStartId="14.[151,264,1637,1662]" captionTargetBox="[160,1423,326,1446]" captionTargetId="figure@14.[151,1435,319,1452]" captionTargetPageId="14" captionText="FIGURES 5964. E. rickalleni sp. nov. 5961 Male, holotype (IBSP 121338), left palp (59 prolateral, 60 ventral, 61 retrolateral); 6264 Female, paratype (IBSP 121553) (62 epigyne, 63 vulva, 64 schematic course of internal duct system. Scale lines: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3458460" httpUri="https://zenodo.org/record/3458460/files/figure.png" pageId="2" pageNumber="85">Fig. 62</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="61CBC3B20037FFAFB294FAA1FF6AFAA1" pageId="2" pageNumber="85" type="distribution">
<paragraph id="296E90390037FFAFB294FAA1FF6AFAA1" blockId="2.[151,1437,152,1979]" pageId="2" pageNumber="85">
<emphasis id="1BA54C2B0037FFAFB294FAA1FECFFAC5" bold="true" box="[199,353,1304,1331]" pageId="2" pageNumber="85">Distribution.</emphasis>
Known from southeastern
<collectingCountry id="51C6D0A90037FFAFB0CDFAA1FD4BFAC4" box="[670,741,1304,1330]" name="Brazil" pageId="2" pageNumber="85">Brazil</collectingCountry>
, states of
<collectingRegion id="EB155EDB0037FFAFB10DFAA0FC7BFAC4" box="[862,981,1304,1331]" country="Brazil" name="Sao Paulo" pageId="2" pageNumber="85">São Paulo</collectingRegion>
,
<collectingRegion id="EB155EDB0037FFAFB1B0FAA1FB28FAC5" box="[995,1158,1304,1331]" country="Brazil" name="Espirito Santo" pageId="2" pageNumber="85">Espírito Santo</collectingRegion>
and
<collectingRegion id="EB155EDB0037FFAFB690FAA1FAF2FAC4" box="[1219,1372,1304,1330]" country="Brazil" name="Minas Gerais" pageId="2" pageNumber="85">Minas Gerais</collectingRegion>
(
<figureCitation id="B1EA8CBC0037FFAFB73CFAA1FF19FAA1" captionStart="FIGURE 65" captionStartId="15.[151,250,1232,1257]" captionTargetBox="[151,1436,174,1213]" captionTargetId="figure@15.[151,1436,174,1213]" captionTargetPageId="15" captionText="FIGURE 65. Distribution map for species of Extraordinarius gen. nov." figureDoi="http://doi.org/10.5281/zenodo.3458462" httpUri="https://zenodo.org/record/3458462/files/figure.png" pageId="2" pageNumber="85">Fig. 65</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="61CBC3B20037FFAFB294FAD9FBA7F84C" pageId="2" pageNumber="85" type="discussion">
<paragraph id="296E90390037FFAFB294FAD9FE2CFA69" blockId="2.[151,1437,152,1979]" pageId="2" pageNumber="85">
<emphasis id="1BA54C2B0037FFAFB294FAD9FEC9FA8D" bold="true" box="[199,359,1376,1403]" pageId="2" pageNumber="85">Composition.</emphasis>
<taxonomicName id="EED1EBBA0037FFAFB33DFAD9FD1AFA8C" authority="Rheims, 2019" authorityName="Rheims" authorityYear="2019" box="[366,692,1376,1402]" class="Arachnida" family="Sparassidae" genus="Extraordinarius" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="85" phylum="Arthropoda" rank="species" species="andrematosi" status="sp. nov.">
<emphasis id="1BA54C2B0037FFAFB33DFAD9FD1AFA8C" box="[366,692,1376,1402]" italics="true" pageId="2" pageNumber="85">Extraordinarius andrematosi</emphasis>
</taxonomicName>
<emphasis id="1BA54C2B0037FFAFB0E8FAD8FCBBFA8D" bold="true" box="[699,789,1377,1403]" pageId="2" pageNumber="85">
<taxonomicNameLabel id="0096F1500037FFAFB0E8FAD8FCBBFA8D" box="[699,789,1377,1403]" pageId="2" pageNumber="85" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="EED1EBBA0037FFAFB177FAD8FC58FA8C" authority="Rheims, 2019" authorityName="Rheims" authorityYear="2019" box="[804,1014,1376,1402]" class="Arachnida" family="Sparassidae" genus="Extraordinarius" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="85" phylum="Arthropoda" rank="species" species="brucedickinsoni" status="sp. nov.">
<emphasis id="1BA54C2B0037FFAFB177FAD8FC58FA8C" box="[804,1014,1376,1402]" italics="true" pageId="2" pageNumber="85">E. brucedickinsoni</emphasis>
</taxonomicName>
<emphasis id="1BA54C2B0037FFAFB1AEFAD8FBF9FA8D" bold="true" box="[1021,1111,1377,1403]" pageId="2" pageNumber="85">
<taxonomicNameLabel id="0096F1500037FFAFB1AEFAD8FBF9FA8D" box="[1021,1111,1377,1403]" pageId="2" pageNumber="85" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="EED1EBBA0037FFAFB635FAD8FAA5FA8C" authority="Rheims, 2019" authorityName="Rheims" authorityYear="2019" box="[1126,1291,1376,1402]" class="Arachnida" family="Sparassidae" genus="Extraordinarius" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="85" phylum="Arthropoda" rank="species" species="klausmeinei" status="sp. nov.">
<emphasis id="1BA54C2B0037FFAFB635FAD8FAA5FA8C" box="[1126,1291,1376,1402]" italics="true" pageId="2" pageNumber="85">E. klausmeinei</emphasis>
</taxonomicName>
<emphasis id="1BA54C2B0037FFAFB741FAD8FAC2FA8D" bold="true" box="[1298,1388,1377,1403]" pageId="2" pageNumber="85">
<taxonomicNameLabel id="0096F1500037FFAFB741FAD8FAC2FA8D" box="[1298,1388,1377,1403]" pageId="2" pageNumber="85" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="EED1EBBA0037FFAFB2C4FA3CFE8FFA68" authority="Rheims, 2019" authorityName="Rheims" authorityYear="2019" box="[151,289,1412,1438]" class="Arachnida" family="Sparassidae" genus="Extraordinarius" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="85" phylum="Arthropoda" rank="species" species="rickalleni" status="sp. nov.">
<emphasis id="1BA54C2B0037FFAFB2C4FA3CFE8FFA68" box="[151,289,1412,1438]" italics="true" pageId="2" pageNumber="85">E. rickalleni</emphasis>
</taxonomicName>
<emphasis id="1BA54C2B0037FFAFB37AFA3CFE2CFA69" bold="true" box="[297,386,1413,1439]" pageId="2" pageNumber="85">
<taxonomicNameLabel id="0096F1500037FFAFB37AFA3CFE2CFA69" box="[297,386,1413,1439]" pageId="2" pageNumber="85" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="296E90390037FFAFB294FA10FBE4F948" blockId="2.[151,1437,152,1979]" pageId="2" pageNumber="85">
<emphasis id="1BA54C2B0037FFAFB294FA10FD82FA35" bold="true" box="[199,556,1448,1475]" pageId="2" pageNumber="85">Monophyly and relationships.</emphasis>
In the absence of a cladistic analysis, not much can be said about the relationships between
<taxonomicName id="EED1EBBA0037FFAFB312FA75FE5AFA10" authority="Rheims, 2019" authorityName="Rheims" authorityYear="2019" box="[321,500,1484,1510]" class="Arachnida" family="Sparassidae" genus="Extraordinarius" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="85" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="1BA54C2B0037FFAFB312FA75FE5AFA10" box="[321,500,1484,1510]" italics="true" pageId="2" pageNumber="85">Extraordinarius</emphasis>
</taxonomicName>
<emphasis id="1BA54C2B0037FFAFB3ADFA74FDC4FA11" bold="true" box="[510,618,1485,1511]" pageId="2" pageNumber="85">
<taxonomicNameLabel id="0096F1500037FFAFB3ADFA74FDC4FA11" box="[510,618,1485,1511]" pageId="2" pageNumber="85" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
and the remaining Sparianthinae genera. The presence of a sclerotized conductor in the male palp could suggest a closer relationship with the non-Neotropical
<taxonomicName id="EED1EBBA0037FFAFB6D6FA49FB5FF9FC" authorityName="Simon" authorityYear="1898" box="[1157,1265,1520,1546]" class="Arachnida" family="Sparassidae" genus="Pleorotus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="85" phylum="Arthropoda" rank="genus">
<emphasis id="1BA54C2B0037FFAFB6D6FA49FB5FF9FC" box="[1157,1265,1520,1546]" italics="true" pageId="2" pageNumber="85">Pleorotus</emphasis>
</taxonomicName>
,
<taxonomicName id="EED1EBBA0037FFAFB753FA49FA3BF9FC" authorityName="Simon" authorityYear="1897" box="[1280,1429,1520,1546]" class="Arachnida" family="Sparassidae" genus="Rhacocnemis" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="85" phylum="Arthropoda" rank="genus">
<emphasis id="1BA54C2B0037FFAFB753FA49FA3BF9FC" box="[1280,1429,1520,1546]" italics="true" pageId="2" pageNumber="85">Rhacocnemis</emphasis>
</taxonomicName>
,
<taxonomicName id="EED1EBBA0037FFAFB2C4F9ACFF47F9D9" box="[151,233,1557,1583]" class="Arachnida" family="Sparassidae" genus="Stasina" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="85" phylum="Arthropoda" rank="genus">
<emphasis id="1BA54C2B0037FFAFB2C4F9ACFF47F9D9" box="[151,233,1557,1583]" italics="true" pageId="2" pageNumber="85">Stasina</emphasis>
</taxonomicName>
,
<taxonomicName id="EED1EBBA0037FFAFB2A9F9ACFEEEF9D9" authorityName="Simon" authorityYear="1898" box="[250,320,1557,1583]" class="Arachnida" family="Sparassidae" genus="Stipax" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="85" phylum="Arthropoda" rank="genus">
<emphasis id="1BA54C2B0037FFAFB2A9F9ACFEEEF9D9" box="[250,320,1557,1583]" italics="true" pageId="2" pageNumber="85">Stipax</emphasis>
</taxonomicName>
,
<taxonomicName id="EED1EBBA0037FFAFB303F9ADFE77F9D8" authorityName="Karsch" authorityYear="1884" box="[336,473,1556,1582]" class="Arachnida" family="Sparassidae" genus="Thelcticopis" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="85" phylum="Arthropoda" rank="genus">
<emphasis id="1BA54C2B0037FFAFB303F9ADFE77F9D8" box="[336,473,1556,1582]" italics="true" pageId="2" pageNumber="85">Thelcticopis</emphasis>
</taxonomicName>
and
<taxonomicName id="EED1EBBA0037FFAFB047F9ADFD30F9D8" authorityName="Hirst" authorityYear="1911" box="[532,670,1556,1582]" class="Arachnida" family="Sparassidae" genus="Thomasettia" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="85" phylum="Arthropoda" rank="genus">
<emphasis id="1BA54C2B0037FFAFB047F9ADFD30F9D8" box="[532,670,1556,1582]" italics="true" pageId="2" pageNumber="85">Thomasettia</emphasis>
</taxonomicName>
. Nevertheless, cheliceral dentition (two small retromarginal teeth), number of ventral spines on tibiae III (three pairs) and eye arrangement (anterior row straight, posterior straight or gently procurved) are markedly different from the latter genera and suggest it might, in fact, be related to the Neotropical
<taxonomicName id="EED1EBBA0037FFAFB377F938FE0EF96C" authorityName="Rheims &amp; Alayon" authorityYear="2016" box="[292,416,1665,1690]" class="Arachnida" family="Sparassidae" genus="Neostasina" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="85" phylum="Arthropoda" rank="genus">
<emphasis id="1BA54C2B0037FFAFB377F938FE0EF96C" box="[292,416,1665,1690]" italics="true" pageId="2" pageNumber="85">Neostasina</emphasis>
</taxonomicName>
,
<taxonomicName id="EED1EBBA0037FFAFB3E3F939FD2EF96C" authorityName="Simon" authorityYear="1887" box="[432,640,1664,1690]" class="Arachnida" family="Sparassidae" genus="Pseudosparianthis" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="85" phylum="Arthropoda" rank="genus">
<emphasis id="1BA54C2B0037FFAFB3E3F939FD2EF96C" box="[432,640,1664,1690]" italics="true" pageId="2" pageNumber="85">Pseudosparianthis</emphasis>
</taxonomicName>
,
<taxonomicName id="EED1EBBA0037FFAFB0C3F938FCBDF96D" authorityName="Mello-Leitao" authorityYear="1930" box="[656,787,1665,1691]" class="Arachnida" family="Sparassidae" genus="Sampaiosia" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="85" phylum="Arthropoda" rank="genus">
<emphasis id="1BA54C2B0037FFAFB0C3F938FCBDF96D" box="[656,787,1665,1691]" italics="true" pageId="2" pageNumber="85">Sampaiosia</emphasis>
</taxonomicName>
and
<taxonomicName id="EED1EBBA0037FFAFB11EF939FC60F96C" authorityName="Simon" authorityYear="1880" box="[845,974,1664,1690]" class="Arachnida" family="Sparassidae" genus="Sparianthis" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="85" phylum="Arthropoda" rank="genus">
<emphasis id="1BA54C2B0037FFAFB11EF939FC60F96C" box="[845,974,1664,1690]" italics="true" pageId="2" pageNumber="85">Sparianthis</emphasis>
</taxonomicName>
. Thus, until further phylogenetic studies are carried out, the position of the genus within the subfamily remains controversial.
</paragraph>
<paragraph id="296E90390037FFAFB294F971FBA7F84C" blockId="2.[151,1437,152,1979]" pageId="2" pageNumber="85">
The monophyly of the genus, on the other hand, seems to be well substantiated. As pointed out by
<bibRefCitation id="4D40EDC80037FFAFB77AF971FE90F8F1" author="Rheims, C. A. &amp; Alayon, G." pageId="2" pageNumber="85" pagination="301 - 344" refId="ref7868" refString="Rheims, C. A. &amp; Alayon, G. (2016) Neostasina gen. nov., a new genus of huntsman spiders from the Neotropical region (Araneae, Sparassidae, Sparianthinae). Zootaxa, 4079 (3), 301 - 344. https: // doi. org / 10.11646 / zootaxa. 4079.3.1" type="journal article" year="2016">Rheims &amp; Alayón (2016)</bibRefCitation>
,
<taxonomicName id="EED1EBBA0037FFAFB31AF955FE63F8F0" authorityName="Bertkau" authorityYear="1872" box="[329,461,1772,1798]" class="Arachnida" family="Sparassidae" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="85" phylum="Arthropoda" rank="family">Sparassidae</taxonomicName>
species are usually conservative and rarely deviate from the generic
<emphasis id="1BA54C2B0037FFAFB681F955FA80F8F0" box="[1234,1326,1772,1798]" italics="true" pageId="2" pageNumber="85">bauplan</emphasis>
.
<taxonomicName id="EED1EBBA0037FFAFB76FF954FF5AF8DC" authority="Rheims, 2019" authorityName="Rheims" authorityYear="2019" class="Arachnida" family="Sparassidae" genus="Extraordinarius" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="85" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="1BA54C2B0037FFAFB76FF954FF5AF8DC" italics="true" pageId="2" pageNumber="85">Extraordinarius</emphasis>
</taxonomicName>
<emphasis id="1BA54C2B0037FFAFB2AEF8A8FEC9F8DD" bold="true" box="[253,359,1809,1835]" pageId="2" pageNumber="85">
<taxonomicNameLabel id="0096F1500037FFAFB2AEF8A8FEC9F8DD" box="[253,359,1809,1835]" pageId="2" pageNumber="85" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
shows a combination of genitalic characters not found in any other Sparianthinae genera, such as the shape of the sclerotized conductor and tegular apophyses, the position of the median apophysis in the male palps and the general conformation of the internal duct system of the female vulva. I believe that these characters support the hypothesis of this being a distinguished lineage within Sparianthinae and even if new species are to be discovered in the future, they will probably show the same diagnostic features.
</paragraph>
</subSubSection>
</treatment>
</document>