treatments-xml/data/27/73/25/2773254AA33B707AFF31B8B0FAA296A5.xml

<document id="FAB8BF5FA4BFB27C466E42A9EF3115A0" ID-CLB-Dataset="3892" ID-DOI="10.11646/phytotaxa.523.3.6" ID-GBIF-Dataset="b7b21974-e27d-4209-938a-e26cfa2e2c8a" ID-ISSN="1179-3163" ID-Zenodo-Dep="5591458" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1634882310262" checkinUser="plazi" docAuthor="Cascante-Marín, Alfredo &amp; Morales, J. Francisco" docDate="2021" docId="2773254AA33B707AFF31B8B0FAA296A5" docLanguage="en" docName="phytotaxa.523.3.6.pdf" docOrigin="Phytotaxa 523 (3)" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.7:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="7" docTitle="Werauhia talamancana Cascante &amp; J. F. Morales 2021, sp. nov." docType="treatment" docVersion="8" lastPageNumber="257" masterDocId="DB4A5D32A33A7079FFB9B964FFB89040" masterDocTitle="A new species of Werauhia (Bromeliaceae, Tillandsioideae) from the highlands of southern Central America" masterLastPageNumber="258" masterPageNumber="254" pageNumber="255" updateTime="1699464228670" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="1CC7B35A79E648B20ADBB23C54679AC0">A new species of Werauhia (Bromeliaceae, Tillandsioideae) from the highlands of southern Central America</mods:title>
</mods:titleInfo>
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<mods:namePart id="A2D9E3AC6F53F120D60F96AB5F95F61C">Cascante-Marín, Alfredo</mods:namePart>
<mods:nameIdentifier id="99CFAFCEC0D043084D252CD69B4F721E" type="ORCID">0000-0001-6382-9316</mods:nameIdentifier>
<mods:affiliation id="AA9AC3F073BAE17BBA084B60681383C9">Escuela de Biología y Herbario USJ, Universidad de Costa Rica, 11501 - 2060 San Pedro deMontes de Oca, San José, Costa Rica. alfredo. cascante @ ucr. ac. cr; https: // orcid. org / 0000 - 0001 - 6382 - 9316</mods:affiliation>
<mods:nameIdentifier id="7B071705DABFA8C14ACEFC7EF586EC6D" type="email">alfredo.cascante@ucr.ac.cr</mods:nameIdentifier>
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<mods:namePart id="1A155E3DDFE1C6C1D9E95408EF308B6E">Morales, J. Francisco</mods:namePart>
<mods:affiliation id="BC40079342E5CA9628EE6C0B3449FD96">National Herbarium of Trinidad and Tobago, Department of Life Sciences, Faculty of Science and Technology, University of West Indies, St. Augustine Campus, Trinidad and Tobago. Research Associate, Missouri Botanical Garden, 4344 Shaw Blvd., St. Louis, Missouri 63110 USA</mods:affiliation>
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<mods:title id="B289A4952FD0A2E4D73E553C894FBC3E">Phytotaxa</mods:title>
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<mods:date id="3CB1D9A043B6754AAA8E1D40887DB51B">2021</mods:date>
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<treatment id="2773254AA33B707AFF31B8B0FAA296A5" ID-DOI="http://doi.org/10.5281/zenodo.5596929" ID-GBIF-Taxon="189326324" ID-Zenodo-Dep="5596929" LSID="urn:lsid:plazi:treatment:2773254AA33B707AFF31B8B0FAA296A5" httpUri="http://treatment.plazi.org/id/2773254AA33B707AFF31B8B0FAA296A5" lastPageId="3" lastPageNumber="257" pageId="1" pageNumber="255">
<subSubSection id="E7C0C7D7A33B7078FF31B8B0FCCF91AE" box="[136,887,468,495]" pageId="1" pageNumber="255" type="nomenclature">
<paragraph id="AF65945CA33B7078FF31B8B0FCCF91AE" blockId="1.[136,887,468,495]" box="[136,887,468,495]" pageId="1" pageNumber="255">
<heading id="F42D2330A33B7078FF31B8B0FCCF91AE" box="[136,887,468,495]" fontSize="11" level="2" pageId="1" pageNumber="255" reason="3">
<taxonomicName id="68DAEFDFA33B7078FF31B8B0FD0F91AE" ID-CoL="9V7ZS" authority="Cascante &amp; J. F. Morales" authorityName="Cascante &amp; J. F. Morales" authorityYear="2021" box="[136,695,468,495]" class="Liliopsida" family="Bromeliaceae" genus="Werauhia" kingdom="Plantae" order="Poales" pageId="1" pageNumber="255" phylum="Tracheophyta" rank="species" species="talamancana" status="sp. nov.">
<emphasis id="9DAE484EA33B7078FF31B8B0FE2E91AF" bold="true" box="[136,406,468,495]" italics="true" pageId="1" pageNumber="255">Werauhia talamancana</emphasis>
Cascante &amp; J. F. Morales
</taxonomicName>
, 
<emphasis id="9DAE484EA33B7078FD78B8B0FCAF91AE" bold="true" box="[705,791,468,494]" italics="true" pageId="1" pageNumber="255">
<taxonomicNameLabel id="869DF535A33B7078FD78B8B0FCAF91AE" box="[705,791,468,494]" pageId="1" pageNumber="255" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="37E188D9A33B7078FC9FB8B0FCDE91AE" box="[806,870,468,494]" captionStart="FIGURE 1" captionStartId="2.[136,229,1422,1444]" captionTargetBox="[405,1185,280,1363]" captionTargetId="figure-44@2.[367,1219,242,1394]" captionTargetPageId="2" captionText="FIGURE 1. Werauhia talamancana Cascante &amp; J. F. Morales.A. plant habit; B. leaf; C.flower; D. inflorescence; E. cupulate stigmata; F. petal with stamen and basal appendages; G. pistil with antesepalous stamens. Drawings byA. Cascante-Marín." figureDoi="http://doi.org/10.5281/zenodo.5591460" httpUri="https://zenodo.org/record/5591460/files/figure.png" pageId="1" pageNumber="255">Fig. 1</figureCitation>
).
</heading>
</paragraph>
</subSubSection>
<subSubSection id="E7C0C7D7A33B7078FF31BB7BFC969285" pageId="1" pageNumber="255" type="diagnosis">
<paragraph id="AF65945CA33B7078FF31BB7BFC969285" blockId="1.[136,1453,543,818]" pageId="1" pageNumber="255">
<emphasis id="9DAE484EA33B7078FF31BB7BFF4B9275" bold="true" box="[136,243,543,565]" pageId="1" pageNumber="255">Diagnosis:</emphasis>
— 
<taxonomicName id="68DAEFDFA33B7078FEB2BB44FE529275" box="[267,490,544,565]" class="Liliopsida" family="Bromeliaceae" genus="Werauhia" kingdom="Plantae" order="Poales" pageId="1" pageNumber="255" phylum="Tracheophyta" rank="species" species="talamancana">
<emphasis id="9DAE484EA33B7078FEB2BB44FE529275" box="[267,490,544,565]" italics="true" pageId="1" pageNumber="255">Werauhia talamancana</emphasis>
</taxonomicName>
is morphologically similar to 
<taxonomicName id="68DAEFDFA33B7078FCB4BB44FCD19275" authorityName="Grant" authorityYear="1995" baseAuthorityName="Mez &amp; Werckle" baseAuthorityYear="1904" box="[781,873,544,565]" class="Liliopsida" family="Bromeliaceae" genus="Werauhia" kingdom="Plantae" order="Poales" pageId="1" pageNumber="255" phylum="Tracheophyta" rank="species" species="brunei">
<emphasis id="9DAE484EA33B7078FCB4BB44FCD19275" box="[781,873,544,565]" italics="true" pageId="1" pageNumber="255">W. brunei</emphasis>
</taxonomicName>
in its simple spicate inflorescence with smooth and lustrous floral bracts that are distichous (not secund) at anthesis. The new species differs from 
<taxonomicName id="68DAEFDFA33B7078FC4BBB20FBF69219" authorityName="Grant" authorityYear="1995" baseAuthorityName="Mez &amp; Werckle" baseAuthorityYear="1904" box="[1010,1102,580,601]" class="Liliopsida" family="Bromeliaceae" genus="Werauhia" kingdom="Plantae" order="Poales" pageId="1" pageNumber="255" phylum="Tracheophyta" rank="species" species="brunei">
<emphasis id="9DAE484EA33B7078FC4BBB20FBF69219" box="[1010,1102,580,601]" italics="true" pageId="1" pageNumber="255">W. brunei</emphasis>
</taxonomicName>
in having larger flowers with longer petals (5.3–6.0 
<emphasis id="9DAE484EA33B7078FEF7BB0CFED1923D" box="[334,361,616,637]" italics="true" pageId="1" pageNumber="255">vs.</emphasis>
4.0– 
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) and longer stamens (4.0–5.0 
<emphasis id="9DAE484EA33B7078FD4FBB0CFCA9923D" box="[758,785,616,637]" italics="true" pageId="1" pageNumber="255">vs.</emphasis>
<quantity id="682239B9A33B7078FCACBB0CFC3B923D" box="[789,899,616,638]" metricMagnitude="-2" metricUnit="m" metricValue="3.65" metricValueMax="3.7" metricValueMin="3.6" pageId="1" pageNumber="255" unit="cm" value="3.65" valueMax="3.7" valueMin="3.6">3.6–3.7 cm</quantity>
). The lower peduncle bracts in 
<taxonomicName id="68DAEFDFA33B7078FB10BB0CFAFB923D" box="[1193,1347,616,637]" class="Liliopsida" family="Bromeliaceae" genus="Werauhia" kingdom="Plantae" order="Poales" pageId="1" pageNumber="255" phylum="Tracheophyta" rank="species" species="talamancana">
<emphasis id="9DAE484EA33B7078FB10BB0CFAFB923D" box="[1193,1347,616,637]" italics="true" pageId="1" pageNumber="255">W. talamancana</emphasis>
</taxonomicName>
are foliose and longer, compared to the non-foliose and shorter peduncle bracts of 
<taxonomicName id="68DAEFDFA33B7078FCE4BBE8FC0192E1" authorityName="Grant" authorityYear="1995" baseAuthorityName="Mez &amp; Werckle" baseAuthorityYear="1904" box="[861,953,652,673]" class="Liliopsida" family="Bromeliaceae" genus="Werauhia" kingdom="Plantae" order="Poales" pageId="1" pageNumber="255" phylum="Tracheophyta" rank="species" species="brunei">
<emphasis id="9DAE484EA33B7078FCE4BBE8FC0192E1" box="[861,953,652,673]" italics="true" pageId="1" pageNumber="255">W. brunei</emphasis>
</taxonomicName>
. Also, the floral bracts of the new species are usually brown-maroon at anthesis as opposed to light green in 
<taxonomicName id="68DAEFDFA33B7078FD72BBD4FC909285" authorityName="Grant" authorityYear="1995" baseAuthorityName="Mez &amp; Werckle" baseAuthorityYear="1904" box="[715,808,688,709]" class="Liliopsida" family="Bromeliaceae" genus="Werauhia" kingdom="Plantae" order="Poales" pageId="1" pageNumber="255" phylum="Tracheophyta" rank="species" species="brunei">
<emphasis id="9DAE484EA33B7078FD72BBD4FC909285" box="[715,808,688,709]" italics="true" pageId="1" pageNumber="255">W. brunei</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="E7C0C7D7A33B7078FF31BBB0FEAC972A" pageId="1" pageNumber="255" type="materials_examined">
<paragraph id="AF65945CA33B7078FF31BBB0FA939372" blockId="1.[136,1453,543,818]" pageId="1" pageNumber="255">
<emphasis id="9DAE484EA33B7078FF31BBB0FF6192A9" bold="true" box="[136,217,724,745]" pageId="1" pageNumber="255">Type:—</emphasis>
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<collectingCountry id="D7CDD4CCA33B7078FF60BBB0FEDF92A9" box="[217,359,724,745]" name="Costa Rica" pageId="1" pageNumber="255">COSTA RICA</collectingCountry>
. 
<collectingRegion id="6D1E5ABEA33B7078FECABBB0FE7492AA" box="[371,460,724,746]" country="Costa Rica" name="San Jose" pageId="1" pageNumber="255">San José</collectingRegion>
: Jardín, 
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, sector del 
<collectorName id="022FF18AA33B7078FC7FBBB0FB3092A9" box="[966,1160,724,746]" pageId="1" pageNumber="255">Cerro de la Muerte</collectorName>
, carretera interamericana sur. Área no protegida. Fragmento de bosque a aprox. 0.5 Km 
<collectionCode id="C9CB0C99A33B7078FD40BB9CFCBE934D" box="[761,774,760,781]" country="Sweden" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="1" pageNumber="255" type="Herbarium">S</collectionCode>
</materialsCitation>
<materialsCitation id="1FB29E01A33B7078FCB7BB9CFA9F9372" ID-GBIF-Occurrence="3392754307" collectingDate="2021-02-12" collectionCode="USJ, CR" collectorName="de El Empalme. Bosque" country="Costa Rica" elevation="2380" latitude="9.713278" location="Cascante-Marin" longLatPrecision="1" longitude="-83.94953" pageId="1" pageNumber="255" specimenCount="1" stateProvince="San Jose" typeStatus="holotype">
de intersección 
<collectorName id="022FF18AA33B7078FC1EBB9CFB2B934D" box="[935,1171,760,782]" pageId="1" pageNumber="255">de El Empalme. Bosque</collectorName>
muy húmedo montano bajo. 
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, 
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, 
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<elevation id="24F7736FA33B7078FE5EBA78FD8A9371" box="[487,562,796,817]" metricMagnitude="3" metricUnit="m" metricValue="2.38" pageId="1" pageNumber="255" unit="m" value="2380.0">2380 m</elevation>
</quantity>
; 
<date id="DB64B29CA33B7078FD84BA78FD5F9371" box="[573,743,796,818]" pageId="1" pageNumber="255" value="2021-02-12">
<collectingDate id="CB204B74A33B7078FD84BA78FD5F9371" box="[573,743,796,818]" pageId="1" pageNumber="255" value="2021-02-12">12 February 2021</collectingDate>
</date>
(fl.), 
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<location id="AA05C287A33B7078FCA7BA78FC039371" LSID="urn:lsid:plazi:treatment:2773254AA33B707AFF31B8B0FAA296A5:AA05C287A33B7078FCA7BA78FC039371" box="[798,955,796,817]" country="Costa Rica" latitude="9.713278" longLatPrecision="1" longitude="-83.94953" name="Cascante-Marin" pageId="1" pageNumber="255" stateProvince="San Jose">Cascante-Marín</location>
2935
</emphasis>
(
<typeStatus id="70612AFEA33B7078FC47BA78FBE49372" box="[1022,1116,796,818]" pageId="1" pageNumber="255" type="holotype">Holotype</typeStatus>
: 
<collectionCode id="C9CB0C99A33B7078FBDCBA78FB2C9371" box="[1125,1172,796,817]" country="Costa Rica" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15819" lsid="urn:lsid:biocol.org:col:15819" name="Universidad de Costa Rica" pageId="1" pageNumber="255" type="University or college">USJ</collectionCode>
!, Isotype: 
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!)
</materialsCitation>
.
</paragraph>
<paragraph id="AF65945CA33B7078FF31BA04FA37962E" blockId="1.[136,1452,864,2079]" pageId="1" pageNumber="255">
<emphasis id="9DAE484EA33B7078FF31BA04FF69933B" bold="true" box="[136,209,864,891]" italics="true" pageId="1" pageNumber="255">Plants</emphasis>
epiphytic, acaulescent, solitary (forming a single rosette), (40-) 
<quantity id="682239B9A33B7078FC12BA04FBA3933B" box="[939,1051,864,891]" metricMagnitude="-1" metricUnit="m" metricValue="5.75" metricValueMax="6.5" metricValueMin="5.0" pageId="1" pageNumber="255" unit="cm" value="57.5" valueMax="65.0" valueMin="50.0">50–65 cm</quantity>
in diameter, 
<quantity id="682239B9A33B7078FB09BA04FA99933B" box="[1200,1313,864,891]" metricMagnitude="-1" metricUnit="m" metricValue="3.25" metricValueMax="3.5" metricValueMin="3.0" pageId="1" pageNumber="255" unit="cm" value="32.5" valueMax="35.0" valueMin="30.0">30–35 cm</quantity>
tall. 
<emphasis id="9DAE484EA33B7078FAE5BA04FA13933A" bold="true" box="[1372,1451,864,890]" italics="true" pageId="1" pageNumber="255">Leaves</emphasis>
rosulate, (13-) 17–24 per rosette, forming a water compounding tank; 
<emphasis id="9DAE484EA33B7078FC2CBAE0FC5993DF" bold="true" box="[917,993,900,927]" italics="true" pageId="1" pageNumber="255">sheath</emphasis>
elliptic, 
<quantity id="682239B9A33B7078FBFABAE0FB1C93DF" box="[1091,1188,900,927]" metricMagnitude="-1" metricUnit="m" metricValue="1.2" metricValueMax="1.5" metricValueMin="0.9" pageId="1" pageNumber="255" unit="cm" value="12.0" valueMax="15.0" valueMin="9.0">9–15 cm</quantity>
long, 
<quantity id="682239B9A33B7078FB53BAE0FADF93DF" box="[1258,1383,900,927]" metricMagnitude="-2" metricUnit="m" metricValue="6.5" metricValueMax="7.5" metricValueMin="5.5" pageId="1" pageNumber="255" unit="cm" value="6.5" valueMax="7.5" valueMin="5.5">5.5–7.5 cm</quantity>
wide, adaxially whitish-cream, sometimes tinged with magenta, with a dark-brown horizontal band at the base abaxially, lepidote on both surfaces; 
<emphasis id="9DAE484EA33B7078FE79BAA8FE4693A7" bold="true" box="[448,510,972,999]" italics="true" pageId="1" pageNumber="255">blade</emphasis>
ligulate, recurved, with attenuate apex, (31–) 
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long, 
<quantity id="682239B9A33B7078FB58BAA8FADB93A7" box="[1249,1379,972,999]" metricMagnitude="-2" metricUnit="m" metricValue="3.9" metricValueMax="4.3" metricValueMin="3.5" pageId="1" pageNumber="255" unit="cm" value="3.9" valueMax="4.3" valueMin="3.5">3.5–4.3 cm</quantity>
wide, inconspicuously punctulate-lepidote on both sides, concolorous green. 
<emphasis id="9DAE484EA33B7078FC28BA94FB96944B" bold="true" box="[913,1070,1008,1035]" italics="true" pageId="1" pageNumber="255">Inflorescence</emphasis>
terminal, simple; 
<emphasis id="9DAE484EA33B7078FB4BBA94FAE2944B" bold="true" box="[1266,1370,1008,1035]" italics="true" pageId="1" pageNumber="255">peduncle</emphasis>
curved to nearly horizontal, 
<quantity id="682239B9A33B7078FEC1BD70FE51946F" box="[376,489,1044,1071]" metricMagnitude="-1" metricUnit="m" metricValue="2.9" metricValueMax="3.3" metricValueMin="2.5" pageId="1" pageNumber="255" unit="cm" value="29.0" valueMax="33.0" valueMin="25.0">25–33 cm</quantity>
long, 
<quantity id="682239B9A33B7078FD8DBD70FD2B946F" box="[564,659,1044,1071]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="1" pageNumber="255" unit="mm" value="8.5" valueMax="9.0" valueMin="8.0">8–9 mm</quantity>
in diameter, hidden by its bracts; 
<emphasis id="9DAE484EA33B7078FBA7BD70FB6D946F" bold="true" box="[1054,1237,1044,1071]" italics="true" pageId="1" pageNumber="255">peduncle bracts</emphasis>
foliose, imbricate, decreasing in size toward the apex, green, the upper ones tinged with brown-maroon, punctulate-lepidote; its sheath 
<quantity id="682239B9A33B7078FF31BD38FEBE9437" box="[136,262,1116,1143]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" metricValueMax="5.5" metricValueMin="4.5" pageId="1" pageNumber="255" unit="cm" value="5.0" valueMax="5.5" valueMin="4.5">4.5–5.5 cm</quantity>
long (lower ones), 3.5–4.0 cm long (upper ones); its blade narrowly triangular-attenuate, 
<quantity id="682239B9A33B7078FAB8BD38FAC99437" box="[1281,1393,1116,1143]" metricMagnitude="-1" metricUnit="m" metricValue="2.25" metricValueMax="3.0" metricValueMin="1.5" pageId="1" pageNumber="255" unit="cm" value="22.5" valueMax="30.0" valueMin="15.0">15–30 cm</quantity>
long (lower ones), 
<quantity id="682239B9A33B7078FE9EBDE4FE2994DB" box="[295,401,1152,1179]" metricMagnitude="-2" metricUnit="m" metricValue="6.75" metricValueMax="8.5" metricValueMin="5.0" pageId="1" pageNumber="255" unit="cm" value="6.75" valueMax="8.5" valueMin="5.0">5–8.5 cm</quantity>
long (upper ones). 
<emphasis id="9DAE484EA33B7078FDCDBDE4FD0C94DB" bold="true" box="[628,692,1152,1179]" italics="true" pageId="1" pageNumber="255">Spike</emphasis>
narrowly sub-oblong at anthesis, flattened, erecto-patent to nearly horizontal, 
<quantity id="682239B9A33B7078FEB0BDC0FEC194FF" box="[265,377,1188,1215]" metricMagnitude="-1" metricUnit="m" metricValue="1.55" metricValueMax="1.8" metricValueMin="1.3" pageId="1" pageNumber="255" unit="cm" value="15.5" valueMax="18.0" valueMin="13.0">13–18 cm</quantity>
long, 
<quantity id="682239B9A33B7078FE7ABDC0FD9494FF" box="[451,556,1188,1215]" metricMagnitude="-2" metricUnit="m" metricValue="4.35" metricValueMax="4.7" metricValueMin="4.0" pageId="1" pageNumber="255" unit="cm" value="4.35" valueMax="4.7" valueMin="4.0">4–4.7 cm</quantity>
wide, 7–9 flowered, rachis hidden by the bracts; 
<emphasis id="9DAE484EA33B7078FBD9BDC0FB5694FF" bold="true" box="[1120,1262,1188,1215]" italics="true" pageId="1" pageNumber="255">floral bracts</emphasis>
ovate, 
<quantity id="682239B9A33B7078FAFBBDC0FA1394FF" box="[1346,1451,1188,1215]" metricMagnitude="-2" metricUnit="m" metricValue="5.75" metricValueMax="6.0" metricValueMin="5.5" pageId="1" pageNumber="255" unit="cm" value="5.75" valueMax="6.0" valueMin="5.5">5.5–6 cm</quantity>
long, 
<quantity id="682239B9A33B7078FF70BDACFEFF94A3" box="[201,327,1224,1251]" metricMagnitude="-2" metricUnit="m" metricValue="3.8" metricValueMax="4.4" metricValueMin="3.2" pageId="1" pageNumber="255" unit="cm" value="3.8" valueMax="4.4" valueMin="3.2">3.2–4.4 cm</quantity>
wide (when extended and flattened), 3–4 times longer than the internodes, lustrous, slightly nerved (when dry), sparsely punctulate-lepidote, ecarinate, strongly imbricate, not secund with the flowers at anthesis, thin coriaceous, tinged with or completely brown-maroon on the upper side, light-green on the lower side, but turning green on both sides when fruiting. 
<emphasis id="9DAE484EA33B7078FE6ABC50FD88950F" bold="true" box="[467,560,1332,1359]" italics="true" pageId="1" pageNumber="255">Flowers</emphasis>
slightly zygomorphic, nocturnal, downward secund at anthesis; 
<emphasis id="9DAE484EA33B7078FAADBC50FADD950F" bold="true" box="[1300,1381,1332,1359]" italics="true" pageId="1" pageNumber="255">pedicel</emphasis>
stout, white, ca. 
<quantity id="682239B9A33B7078FF46BC3CFEFB9533" box="[255,323,1368,1395]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="1" pageNumber="255" unit="mm" value="9.0">9 mm</quantity>
long, ca. 
<quantity id="682239B9A33B7078FE0FBC3CFE419533" box="[438,505,1368,1395]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="1" pageNumber="255" unit="mm" value="8.0">8 mm</quantity>
in diameter; 
<emphasis id="9DAE484EA33B7078FD2DBC3CFD629533" bold="true" box="[660,730,1368,1395]" italics="true" pageId="1" pageNumber="255">sepals</emphasis>
free, ovate, chartaceous, cymbiform, 
<quantity id="682239B9A33B7078FB35BC3CFAB49533" box="[1164,1292,1368,1395]" metricMagnitude="-2" metricUnit="m" metricValue="3.55" metricValueMax="3.7" metricValueMin="3.4" pageId="1" pageNumber="255" unit="cm" value="3.55" valueMax="3.7" valueMin="3.4">3.4–3.7 cm</quantity>
long, 
<quantity id="682239B9A33B7078FAEEBC3CFF1295D7" metricMagnitude="-2" metricUnit="m" metricValue="1.75" metricValueMax="1.9" metricValueMin="1.6" pageId="1" pageNumber="255" unit="cm" value="1.75" valueMax="1.9" valueMin="1.6">1.6–1.9 cm</quantity>
wide when flattened, ecarinate, whitish-green, lustrous, glabrous; 
<emphasis id="9DAE484EA33B7078FC11BC18FC5395D7" bold="true" box="[936,1003,1404,1431]" italics="true" pageId="1" pageNumber="255">petals</emphasis>
free, narrowly obovate, white, 
<quantity id="682239B9A33B7078FAEEBC18FF1295FB" metricMagnitude="-2" metricUnit="m" metricValue="5.75" metricValueMax="6.2" metricValueMin="5.3" pageId="1" pageNumber="255" unit="cm" value="5.75" valueMax="6.2" valueMin="5.3">5.3–6.2 cm</quantity>
long, 
<quantity id="682239B9A33B7078FF4BBCC4FEE395FB" box="[242,347,1440,1467]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" metricValueMax="2.4" metricValueMin="2.0" pageId="1" pageNumber="255" unit="cm" value="2.2" valueMax="2.4" valueMin="2.0">2–2.4 cm</quantity>
wide when extended and flattened, bearing two appendages at the base; appendages subspatulate, 
<quantity id="682239B9A33B7078FF31BCA0FF47959F" box="[136,255,1476,1503]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" metricValueMax="1.3" metricValueMin="1.1" pageId="1" pageNumber="255" unit="mm" value="12.0" valueMax="13.0" valueMin="11.0">11–13 mm</quantity>
long, ca. 
<quantity id="682239B9A33B7078FED3BCA0FE12959F" box="[362,426,1476,1503]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="255" unit="mm" value="3.0">3 mm</quantity>
wide, basally adnate to the petal, apex dactyloid-dentate; 
<emphasis id="9DAE484EA33B7078FB92BCA0FB30959E" bold="true" box="[1067,1160,1476,1502]" italics="true" pageId="1" pageNumber="255">stamens</emphasis>
subequal, shorter than the petals; 
<emphasis id="9DAE484EA33B7078FF60BC8CFEFC9643" bold="true" box="[217,324,1512,1539]" italics="true" pageId="1" pageNumber="255">filaments</emphasis>
free, linear, flat, 
<quantity id="682239B9A33B7078FDB3BC8CFD309643" box="[522,648,1512,1539]" metricMagnitude="-2" metricUnit="m" metricValue="3.9" metricValueMax="4.2" metricValueMin="3.6" pageId="1" pageNumber="255" unit="cm" value="3.9" valueMax="4.2" valueMin="3.6">3.6–4.2 cm</quantity>
long, white; 
<emphasis id="9DAE484EA33B7078FCA6BC8CFCCE9643" bold="true" box="[799,886,1512,1539]" italics="true" pageId="1" pageNumber="255">anthers</emphasis>
dorsifixed near the base, yellow, oblong, 
<quantity id="682239B9A33B7078FAEBBC8CFA149643" box="[1362,1452,1512,1539]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="1" pageNumber="255" unit="mm" value="8.5" valueMax="9.0" valueMin="8.0">8-9 mm</quantity>
long, arranged at the upper part of the corolla at anthesis; 
<emphasis id="9DAE484EA33B7078FCABBF68FCEA9666" bold="true" box="[786,850,1548,1574]" italics="true" pageId="1" pageNumber="255">ovary</emphasis>
superior, conical, white, ca. 
<quantity id="682239B9A33B7078FB2CBF68FB5D9667" box="[1173,1253,1548,1575]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="255" unit="mm" value="10.0">10 mm</quantity>
long, 
<emphasis id="9DAE484EA33B7078FA95BF68FAE79667" bold="true" box="[1324,1375,1548,1575]" italics="true" pageId="1" pageNumber="255">style</emphasis>
white, 
<quantity id="682239B9A33B7078FF31BF54FF4B960B" box="[136,243,1584,1611]" metricMagnitude="-2" metricUnit="m" metricValue="3.25" metricValueMax="3.5" metricValueMin="3.0" pageId="1" pageNumber="255" unit="cm" value="3.25" valueMax="3.5" valueMin="3.0">3–3.5 cm</quantity>
long; 
<emphasis id="9DAE484EA33B7078FEF8BF54FE35960A" bold="true" box="[321,397,1584,1610]" italics="true" pageId="1" pageNumber="255">stigma</emphasis>
3-lobed, cupulate, ca. 
<quantity id="682239B9A33B7078FD2EBF54FD63960B" box="[663,731,1584,1611]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="255" unit="mm" value="3.0">3 mm</quantity>
in diameter, equaling the anthers. 
<emphasis id="9DAE484EA33B7078FBD7BF54FB73960B" bold="true" box="[1134,1227,1584,1611]" italics="true" pageId="1" pageNumber="255">Capsule</emphasis>
ellipsoid, blackish, lustrous, 
<quantity id="682239B9A33B7078FF56BF30FED4962F" box="[239,364,1620,1647]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" metricValueMax="4.2" metricValueMin="3.8" pageId="1" pageNumber="255" unit="cm" value="4.0" valueMax="4.2" valueMin="3.8">3.8–4.2 cm</quantity>
long, ca. 
<quantity id="682239B9A33B7078FE63BF30FD9E962F" box="[474,550,1620,1647]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="1" pageNumber="255" unit="cm" value="1.2">1.2 cm</quantity>
in diameter, apiculate. 
<emphasis id="9DAE484EA33B7078FC92BF30FCD4962F" bold="true" box="[811,876,1620,1647]" italics="true" pageId="1" pageNumber="255">Seeds</emphasis>
with a plumose white appendage, not measured.
</paragraph>
<paragraph id="AF65945CA33B7078FF04BF1CFEAC972A" blockId="1.[136,1452,864,2079]" pageId="1" pageNumber="255">
<emphasis id="9DAE484EA33B7078FF04BF1CFD8B96D2" bold="true" box="[189,563,1656,1682]" pageId="1" pageNumber="255">Additional specimens examined</emphasis>
<materialsCitation id="1FB29E01A33B7078FD83BF1CFE2996F6" ID-GBIF-Occurrence="3392754303" collectingDate="2017-02-15" collectingDateMax="2018-05-11" collectingDateMin="2017-02-15" collectionCode="USJ" collectorName="Cascante-Marin &amp; et al." country="Costa Rica" county="Cerro Biricuacua" elevation="2600" isTailEnumeration="true" latitude="9.398611" location="COSTA RICA" longLatPrecision="21" longitude="-83.16944" municipality="Ujarras" pageId="1" pageNumber="255" specimenCount="1" stateProvince="Limon" typeStatus="paratype">
<emphasis id="9DAE484EA33B7078FD83BF1CFD7496D2" bold="true" box="[570,716,1656,1682]" pageId="1" pageNumber="255">
(
<typeStatus id="70612AFEA33B7078FDFBBF1CFD0796D2" box="[578,703,1656,1682]" pageId="1" pageNumber="255" type="paratype">paratypes</typeStatus>
):
</emphasis>
— 
<collectingCountry id="D7CDD4CCA33B7078FD51BF1CFC3696D2" box="[744,910,1656,1682]" name="Costa Rica" pageId="1" pageNumber="255">COSTA RICA</collectingCountry>
. Type locality. 
<date id="DB64B29CA33B7078FB8CBF1CFB4396D2" box="[1077,1275,1656,1683]" pageId="1" pageNumber="255" value="2017-02-15">
<collectingDate id="CB204B74A33B7078FB8CBF1CFB4396D2" box="[1077,1275,1656,1683]" pageId="1" pageNumber="255" value="2017-02-15">15 February 2017</collectingDate>
</date>
(fl.), 
<emphasis id="9DAE484EA33B7078FA83BF1CFEFF96F6" italics="true" pageId="1" pageNumber="255">
<collectorName id="022FF18AA33B7078FA83BF1CFEB296F6" pageId="1" pageNumber="255">Cascante-Marín et al.</collectorName>
2652
</emphasis>
(
<collectionCode id="C9CB0C99A33B7078FEECBFF8FE3196F6" box="[341,393,1692,1718]" country="Costa Rica" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15819" lsid="urn:lsid:biocol.org:col:15819" name="Universidad de Costa Rica" pageId="1" pageNumber="255" type="University or college">USJ</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="1FB29E01A33B7078FE22BFF9FBF296F6" ID-GBIF-Occurrence="3392754304" box="[411,1098,1692,1719]" collectingDate="2018-01-31" collectingDateMax="2018-05-11" collectingDateMin="2017-02-15" collectionCode="USJ" collectorName="Cascante-Marin &amp; Trejos" country="Costa Rica" county="Cerro Biricuacua" elevation="2600" isTailEnumeration="true" latitude="9.398611" location="Marin" longLatPrecision="21" longitude="-83.16944" municipality="Ujarras" pageId="1" pageNumber="255" specimenCount="1" stateProvince="Limon">
–, 
<date id="DB64B29CA33B7078FE0FBFF8FDD496F6" box="[438,620,1692,1719]" pageId="1" pageNumber="255" value="2018-01-31">
<collectingDate id="CB204B74A33B7078FE0FBFF8FDD496F6" box="[438,620,1692,1719]" pageId="1" pageNumber="255" value="2018-01-31">31 January 2018</collectingDate>
</date>
(fl.), 
<emphasis id="9DAE484EA33B7078FD10BFF8FBB996F6" box="[681,1025,1692,1718]" italics="true" pageId="1" pageNumber="255">
<collectorName id="022FF18AA33B7078FD10BFF8FCD996F6" box="[681,865,1692,1718]" pageId="1" pageNumber="255">Cascante-Marín</collectorName>
&amp; 
<collectorName id="022FF18AA33B7078FCC6BFF9FC7C96F6" box="[895,964,1693,1718]" pageId="1" pageNumber="255">Trejos</collectorName>
2735
</emphasis>
(
<collectionCode id="C9CB0C99A33B7078FBB7BFF8FBFA96F6" box="[1038,1090,1692,1718]" country="Costa Rica" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15819" lsid="urn:lsid:biocol.org:col:15819" name="Universidad de Costa Rica" pageId="1" pageNumber="255" type="University or college">USJ</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="1FB29E01A33B7078FBEDBFF9FE7B969A" ID-GBIF-Occurrence="3392754305" collectingDate="2018-05-11" collectingDateMax="2018-05-11" collectingDateMin="2017-02-15" collectionCode="USJ" collectorName="Cascante-Marin &amp; Trejos" country="Costa Rica" county="Cerro Biricuacua" elevation="2600" isTailEnumeration="true" latitude="9.398611" location="Marin" longLatPrecision="21" longitude="-83.16944" municipality="Ujarras" pageId="1" pageNumber="255" specimenCount="1" stateProvince="Limon">
–, 
<date id="DB64B29CA33B7078FBD6BFF8FB4696F6" box="[1135,1278,1692,1719]" pageId="1" pageNumber="255" value="2018-05-11">
<collectingDate id="CB204B74A33B7078FBD6BFF8FB4696F6" box="[1135,1278,1692,1719]" pageId="1" pageNumber="255" value="2018-05-11">11 May 2018</collectingDate>
</date>
(fl.), 
<emphasis id="9DAE484EA33B7078FA83BFF8FECF969A" italics="true" pageId="1" pageNumber="255">
<collectorName id="022FF18AA33B7078FA83BFF8FF76969B" pageId="1" pageNumber="255">Cascante-Marín</collectorName>
&amp; 
<collectorName id="022FF18AA33B7078FF4ABFA5FE80969A" box="[243,312,1729,1754]" pageId="1" pageNumber="255">Trejos</collectorName>
2769
</emphasis>
(
<collectionCode id="C9CB0C99A33B7078FE3EBFA4FE03969A" box="[391,443,1728,1754]" country="Costa Rica" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15819" lsid="urn:lsid:biocol.org:col:15819" name="Universidad de Costa Rica" pageId="1" pageNumber="255" type="University or college">USJ</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="1FB29E01A33B7078FE76BFA5FB12969A" ID-GBIF-Occurrence="3392754306" box="[463,1194,1728,1755]" collectingDate="2017-02-15" collectingDateMax="2018-05-11" collectingDateMin="2017-02-15" collectionCode="USJ" collectorName="Cascante-Marin &amp; Trejos" country="Costa Rica" county="Cerro Biricuacua" elevation="2600" isTailEnumeration="true" latitude="9.398611" location="Marin" longLatPrecision="21" longitude="-83.16944" municipality="Ujarras" pageId="1" pageNumber="255" specimenCount="1" stateProvince="Limon">
–, 13 
<collectorName id="022FF18AA33B7078FDA9BFA4FDC2969A" box="[528,634,1728,1754]" pageId="1" pageNumber="255">Setember</collectorName>
2018 (fr.), 
<emphasis id="9DAE484EA33B7078FD42BFA4FBE6969A" box="[763,1118,1728,1754]" italics="true" pageId="1" pageNumber="255">
<collectorName id="022FF18AA33B7078FD42BFA4FC0B969A" box="[763,947,1728,1754]" pageId="1" pageNumber="255">Cascante-Marín</collectorName>
&amp; 
<collectorName id="022FF18AA33B7078FC60BFA5FBA6969A" box="[985,1054,1729,1754]" pageId="1" pageNumber="255">Trejos</collectorName>
2800
</emphasis>
(
<collectionCode id="C9CB0C99A33B7078FBD7BFA4FB1A969A" box="[1134,1186,1728,1754]" country="Costa Rica" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15819" lsid="urn:lsid:biocol.org:col:15819" name="Universidad de Costa Rica" pageId="1" pageNumber="255" type="University or college">USJ</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="1FB29E01A33B7078FB0EBFA4FC009762" ID-GBIF-Occurrence="3392754308" collectingDate="2017-02-15" collectingDateMax="2018-05-11" collectingDateMin="2017-02-15" collectionCode="USJ, P" collectorName="Herrera &amp; Gamboa" country="Costa Rica" county="Cerro Biricuacua" elevation="2600" isTailEnumeration="true" latitude="9.398611" location="P. N. Cordillera de Talamanca, flanco NE y cumbre Cerro Biricuacua, entre Ujarras y San Jose Cabecar" longLatPrecision="21" longitude="-83.16944" municipality="Talamanca" pageId="1" pageNumber="255" specimenCount="1" stateProvince="Limon">
<collectingRegion id="6D1E5ABEA33B7078FB0EBFA4FABD969A" box="[1207,1285,1728,1754]" country="Costa Rica" name="Limon" pageId="1" pageNumber="255">Limón</collectingRegion>
, 
<collectingMunicipality id="4F010E26A33B7078FAA9BFA4FA35969A" box="[1296,1421,1728,1754]" pageId="1" pageNumber="255">Talamanca</collectingMunicipality>
. 
<location id="AA05C287A33B7078FA21BFA5FAAD96BE" LSID="urn:lsid:plazi:treatment:2773254AA33B707AFF31B8B0FAA296A5:AA05C287A33B7078FA21BFA5FAAD96BE" country="Costa Rica" county="Cerro Biricuacua" latitude="9.398611" longLatPrecision="21" longitude="-83.16944" municipality="Talamanca" name="P. N. Cordillera de Talamanca, flanco NE y cumbre Cerro Biricuacua, entre Ujarras y San Jose Cabecar" pageId="1" pageNumber="255" stateProvince="Limon">P. N. Cordillera de Talamanca, flanco NE y cumbre Cerro Biricuacua, entre Ujarrás y San José Cabécar</location>
. 
<geoCoordinate id="CAEEF29BA33B7078FA98BF80FA1F96BF" box="[1313,1447,1764,1791]" degrees="9" direction="north" minutes="23" orientation="latitude" pageId="1" pageNumber="255" precision="15" seconds="55" value="9.398611">9°23’55” N</geoCoordinate>
, 
<geoCoordinate id="CAEEF29BA33B7078FF31BE6CFEA59763" box="[136,285,1800,1827]" degrees="83" direction="west" minutes="10" orientation="longitude" pageId="1" pageNumber="255" precision="15" seconds="10" value="-83.16944">83°10’10” W</geoCoordinate>
, 
<quantity id="682239B9A33B7078FE91BE6CFEC79763" box="[296,383,1800,1827]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="1" pageNumber="255" unit="m" value="2600.0">
<elevation id="24F7736FA33B7078FE91BE6CFEC79763" box="[296,383,1800,1827]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="1" pageNumber="255" unit="m" value="2600.0">2600 m</elevation>
</quantity>
; 
<date id="DB64B29CA33B7078FE32BE6CFDAE9762" box="[395,534,1800,1826]" pageId="1" pageNumber="255" value="1993-04-06">
<collectingDate id="CB204B74A33B7078FE32BE6CFDAE9762" box="[395,534,1800,1826]" pageId="1" pageNumber="255" value="1993-04-06">6April 1993</collectingDate>
</date>
(fl.), 
<emphasis id="9DAE484EA33B7078FDEDBE6DFCC99762" box="[596,881,1800,1826]" italics="true" pageId="1" pageNumber="255">
<collectorName id="022FF18AA33B7078FDEDBE6DFD169762" box="[596,686,1801,1826]" pageId="1" pageNumber="255">Herrera</collectorName>
&amp; 
<collectorName id="022FF18AA33B7078FD6BBE6CFC8A9762" box="[722,818,1800,1826]" pageId="1" pageNumber="255">Gamboa</collectorName>
6218
</emphasis>
(
<collectionCode id="C9CB0C99A33B7078FC38BE6CFC149762" box="[897,940,1800,1826]" country="Costa Rica" lsid="urn:lsid:biocol.org:col:15652" name="Museo Nacional de Costa Rica" pageId="1" pageNumber="255" type="Herbarium">CR</collectionCode>
).
</materialsCitation>
<materialsCitation id="1FB29E01A33B7078FC07BE6DFEA8972A" ID-GBIF-Occurrence="3392754301" collectingDate="2006-08-10" collectingDateMax="2018-05-11" collectingDateMin="2017-02-15" collectionCode="CR" collectorName="Morales &amp; Santamaria" country="Panama" county="Cerro Biricuacua" elevation="2600" isTailEnumeration="true" latitude="9.398611" location="cerro Punta, falda NO, camino desde Finca Dracula hacia el ultimo lodge dentro del Parque" longLatPrecision="21" longitude="-83.16944" municipality="Ujarras" pageId="1" pageNumber="255" specimenCount="1" stateProvince="Chiriqui">
<collectingCountry id="D7CDD4CCA33B7078FC07BE6DFB819764" box="[958,1081,1801,1828]" name="Panama" pageId="1" pageNumber="255">PANAMÁ</collectingCountry>
. 
<collectingRegion id="6D1E5ABEA33B7078FBFABE6CFB1D9762" box="[1091,1189,1800,1826]" country="Panama" name="Chiriqui" pageId="1" pageNumber="255">Chiriquí</collectingRegion>
: 
<location id="AA05C287A33B7078FB16BE6DFC199707" LSID="urn:lsid:plazi:treatment:2773254AA33B707AFF31B8B0FAA296A5:AA05C287A33B7078FB16BE6DFC199707" country="Panama" county="Cerro Biricuacua" latitude="9.398611" longLatPrecision="21" longitude="-83.16944" municipality="Ujarras" name="cerro Punta, falda NO, camino desde Finca Dracula hacia el ultimo lodge dentro del Parque" pageId="1" pageNumber="255" stateProvince="Chiriqui">cerro Punta, falda NO, camino desde Finca Drácula hacia el último lodge dentro del Parque</location>
, 
<date id="DB64B29CA33B7078FC16BE48FBDD9706" box="[943,1125,1836,1862]" pageId="1" pageNumber="255" value="2006-08-10">
<collectingDate id="CB204B74A33B7078FC16BE48FBDD9706" box="[943,1125,1836,1862]" pageId="1" pageNumber="255" value="2006-08-10">10 August 2006</collectingDate>
</date>
(fl), 
<emphasis id="9DAE484EA33B7078FB1ABE48FF76972A" italics="true" pageId="1" pageNumber="255">
<collectorName id="022FF18AA33B7078FB1ABE48FAB89706" box="[1187,1280,1836,1862]" pageId="1" pageNumber="255">Morales</collectorName>
&amp; 
<collectorName id="022FF18AA33B7078FA90BE48FA149706" box="[1321,1452,1836,1862]" pageId="1" pageNumber="255">Santamaría</collectorName>
14625
</emphasis>
(
<collectionCode id="C9CB0C99A33B7078FF64BE34FEB0972A" box="[221,264,1872,1898]" country="Costa Rica" lsid="urn:lsid:biocol.org:col:15652" name="Museo Nacional de Costa Rica" pageId="1" pageNumber="255" type="Herbarium">CR</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="E7C0C7D7A33B707BFF04BE10FC4496E0" lastPageId="2" lastPageNumber="256" pageId="1" pageNumber="255" type="distribution">
<paragraph id="AF65945CA33B707BFF04BE10FD94909C" blockId="1.[136,1452,864,2079]" lastBlockId="2.[136,1451,158,221]" lastPageId="2" lastPageNumber="256" pageId="1" pageNumber="255">
<emphasis id="9DAE484EA33B7078FF04BE10FE5D97CE" bold="true" box="[189,485,1908,1934]" pageId="1" pageNumber="255">Distribution and habitat:</emphasis>
—Highlands of the Talamanca mountain range in 
<collectingCountry id="D7CDD4CCA33B7078FBBDBE10FBC397CE" box="[1028,1147,1908,1934]" name="Costa Rica" pageId="1" pageNumber="255">Costa Rica</collectingCountry>
and 
<collectingCountry id="D7CDD4CCA33B7078FB17BE10FAB297CE" box="[1198,1290,1908,1934]" name="Panama" pageId="1" pageNumber="255">Panama</collectingCountry>
,between 2380 and 
<quantity id="682239B9A33B7078FF0EBEFCFEB497F3" box="[183,268,1944,1971]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="1" pageNumber="255" unit="m" value="2600.0">2600 m</quantity>
asl elevation. At the 
<typeStatus id="70612AFEA33B7078FE4FBEFDFD9E97F3" box="[502,550,1945,1971]" pageId="1" pageNumber="255">type</typeStatus>
locality, plants of this new epiphytic bromeliad are growing in the lower canopy of mixed oak forests in a very humid environment. The vegetation is classified as Very Humid Lower Montane Forest according to Holdridge’s Life Zones System (
<bibRefCitation id="CB4BE9ADA33B7078FD2ABE84FC0897BA" author="Bolanos, R. &amp; Watson, V." box="[659,944,2016,2043]" pageId="1" pageNumber="255" pagination="200" refId="ref2871" refString="Bolanos, R. &amp; Watson, V. (1993) Mapa ecologico de Costa Rica, segun el sistema de clasificacion de Zonas de Vida del mundo de L. R. Holdridge. 1: 200 000. Centro Cientifico Tropical, San Jose, Costa Rica." type="journal article" year="1993">Bolaños &amp; Watson 1993</bibRefCitation>
). Other species in the bromeliad community were the sympatric 
<taxonomicName id="68DAEFDFA33B7078FEDAB161FCC8985E" authority="(Smith 1963: 497) Grant (1995 a: 30)" authorityName="Grant" authorityPageNumber="30" authorityYear="1995" baseAuthorityName="Smith" baseAuthorityPageNumber="497" baseAuthorityYear="1963" box="[355,880,2052,2079]" class="Liliopsida" family="Bromeliaceae" genus="Werauhia" kingdom="Plantae" order="Poales" pageId="1" pageNumber="255" phylum="Tracheophyta" rank="species" species="ampla">
<emphasis id="9DAE484EA33B7078FEDAB161FE73985E" box="[355,459,2052,2078]" italics="true" pageId="1" pageNumber="255">W. ampla</emphasis>
(
<bibRefCitation id="CB4BE9ADA33B7078FE63B160FD2F985E" author="Smith, L. B." box="[474,663,2052,2078]" pageId="1" pageNumber="255" pagination="497 - 506" refId="ref3466" refString="Smith, L. B. (1963) Notes on Bromeliaceae, XIX. Phytologia 8: 497 - 506." type="journal article" year="1963">Smith 1963: 497</bibRefCitation>
) 
<bibRefCitation id="CB4BE9ADA33B7078FD1CB160FCC8985E" author="Grant, J. R." box="[677,880,2052,2079]" pageId="1" pageNumber="255" pagination="16 - 57" refId="ref3061" refString="Grant, J. R. (1995 a) The resurrection of Alcantarea and Werauhia, a new genus. Tropische Und Subtropische Pflanzenwelt 91: 16 - 57." type="journal article" year="1995">Grant (1995a: 30)</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="68DAEFDFA33B7078FCC2B161FA3C985E" authority="(Mez 1896: 9) Grant (1995 a: 44)" authorityName="Grant" authorityPageNumber="44" authorityYear="1995" baseAuthorityName="Mez" baseAuthorityPageNumber="9" baseAuthorityYear="1896" box="[891,1412,2052,2079]" class="Liliopsida" family="Bromeliaceae" genus="Werauhia" kingdom="Plantae" order="Poales" pageId="1" pageNumber="255" phylum="Tracheophyta" rank="species" species="ororiensis">
<emphasis id="9DAE484EA33B7078FCC2B161FBB6985E" box="[891,1038,2053,2078]" italics="true" pageId="1" pageNumber="255">W. ororiensis</emphasis>
(
<bibRefCitation id="CB4BE9ADA33B7078FBA5B160FB13985E" author="Mez, C." box="[1052,1195,2052,2078]" pageId="1" pageNumber="255" pagination="1 - 990" refId="ref3206" refString="Mez, C. (1896) Bromeliaceae. Monographiae Phanerogamarum 9: 1 - 990." type="journal article" year="1896">Mez 1896: 9</bibRefCitation>
) 
<bibRefCitation id="CB4BE9ADA33B7078FB03B160FA3C985E" author="Grant, J. R." box="[1210,1412,2052,2079]" pageId="1" pageNumber="255" pagination="16 - 57" refId="ref3061" refString="Grant, J. R. (1995 a) The resurrection of Alcantarea and Werauhia, a new genus. Tropische Und Subtropische Pflanzenwelt 91: 16 - 57." type="journal article" year="1995">Grant (1995a: 44)</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="68DAEFDFA33B707BFA29B161FDDA90F8" authority="(Mez 1903: 135) Grant (1995 a: 33)" authorityName="Grant" authorityPageNumber="33" authorityYear="1995" baseAuthorityName="Mez" baseAuthorityPageNumber="135" baseAuthorityYear="1903" class="Liliopsida" family="Bromeliaceae" genus="Werauhia" kingdom="Plantae" lastPageId="2" lastPageNumber="256" order="Poales" pageId="1" pageNumber="255" phylum="Tracheophyta" rank="species" species="pittieri">
<emphasis id="9DAE484EA33B707BFA29B161FF6C90F8" italics="true" lastPageId="2" lastPageNumber="256" pageId="1" pageNumber="255">W. pittieri</emphasis>
(
<bibRefCitation id="CB4BE9ADA338707BFF5BB9FAFE3590F9" author="Mez, C." box="[226,397,158,185]" pageId="2" pageNumber="256" pagination="130 - 146" refId="ref3223" refString="Mez, C. (1903) Additamenta monographica. Bulletin de l'Herbier Boissier 3: 130 - 146." type="journal article" year="1903">Mez 1903: 135</bibRefCitation>
) 
<bibRefCitation id="CB4BE9ADA338707BFE25B9FAFDDA90F8" author="Grant, J. R." box="[412,610,158,185]" pageId="2" pageNumber="256" pagination="16 - 57" refId="ref3061" refString="Grant, J. R. (1995 a) The resurrection of Alcantarea and Werauhia, a new genus. Tropische Und Subtropische Pflanzenwelt 91: 16 - 57." type="journal article" year="1995">Grant (1995a: 33)</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="68DAEFDFA338707BFDC9B9FBFB5A90F8" authority="(Mez &amp; Werckle 1904: 878) Grant (1995 a: 47)" authorityName="Grant" authorityPageNumber="47" authorityYear="1995" baseAuthorityName="Mez &amp; Werckle" baseAuthorityPageNumber="878" baseAuthorityYear="1904" box="[624,1250,158,185]" class="Liliopsida" family="Bromeliaceae" genus="Werauhia" kingdom="Plantae" order="Poales" pageId="2" pageNumber="256" phylum="Tracheophyta" rank="species" species="rubra">
<emphasis id="9DAE484EA338707BFDC9B9FBFD6A90F8" box="[624,722,158,184]" italics="true" pageId="2" pageNumber="256">W. rubra</emphasis>
(
<bibRefCitation id="CB4BE9ADA338707BFD59B9FAFBB190F8" author="Mez, C. &amp; Werckle, K." box="[736,1033,158,184]" pageId="2" pageNumber="256" pagination="863 - 878" refId="ref3243" refString="Mez, C. &amp; Werckle, K. (1904) Additamenta monographica. Bulletin de l'Herbier Boissier 9: 863 - 878." type="journal article" year="1904">Mez &amp; Wercklé 1904: 878</bibRefCitation>
) 
<bibRefCitation id="CB4BE9ADA338707BFBA1B9FAFB5A90F8" author="Grant, J. R." box="[1048,1250,158,185]" pageId="2" pageNumber="256" pagination="16 - 57" refId="ref3061" refString="Grant, J. R. (1995 a) The resurrection of Alcantarea and Werauhia, a new genus. Tropische Und Subtropische Pflanzenwelt 91: 16 - 57." type="journal article" year="1995">Grant (1995a: 47)</bibRefCitation>
</taxonomicName>
, and 
<taxonomicName id="68DAEFDFA338707BFAA2B9FBFD91909C" authority="(Smith 1958: 193) Grant (1995 a: 49)" authorityName="Grant" authorityPageNumber="49" authorityYear="1995" baseAuthorityName="Smith" baseAuthorityPageNumber="193" baseAuthorityYear="1958" class="Liliopsida" family="Bromeliaceae" genus="Werauhia" kingdom="Plantae" order="Poales" pageId="2" pageNumber="256" phylum="Tracheophyta" rank="species" species="williamsii">
<emphasis id="9DAE484EA338707BFAA2B9FBFA1390F8" box="[1307,1451,158,184]" italics="true" pageId="2" pageNumber="256">W. williamsii</emphasis>
(
<bibRefCitation id="CB4BE9ADA338707BFF29B9A6FEF5909C" author="Smith, L. B." box="[144,333,194,221]" pageId="2" pageNumber="256" pagination="193 - 196" refId="ref3422" refString="Smith, L. B. (1958) Notes on Bromeliaceae, IX. Phytologia 6: 193 - 196." type="journal article" year="1958">Smith 1958: 193</bibRefCitation>
) 
<bibRefCitation id="CB4BE9ADA338707BFEE5B9A6FD91909C" author="Grant, J. R." box="[348,553,194,221]" pageId="2" pageNumber="256" pagination="16 - 57" refId="ref3061" refString="Grant, J. R. (1995 a) The resurrection of Alcantarea and Werauhia, a new genus. Tropische Und Subtropische Pflanzenwelt 91: 16 - 57." type="journal article" year="1995">Grant (1995a: 49)</bibRefCitation>
</taxonomicName>
.
</paragraph>
<caption id="FBA5C4D4A338707BFF31BCEAFB169588" ID-DOI="http://doi.org/10.5281/zenodo.5591460" ID-Zenodo-Dep="5591460" httpUri="https://zenodo.org/record/5591460/files/figure.png" pageId="2" pageNumber="256" startId="2.[136,229,1422,1444]" targetBox="[405,1185,280,1363]" targetPageId="2">
<paragraph id="AF65945CA338707BFF31BCEAFB169588" blockId="2.[136,1452,1422,1480]" pageId="2" pageNumber="256">
<emphasis id="9DAE484EA338707BFF31BCEAFF4495E3" bold="true" box="[136,252,1422,1444]" pageId="2" pageNumber="256">FIGURE 1.</emphasis>
<taxonomicName id="68DAEFDFA338707BFEBBBCEAFD4E95E3" authority="Cascante &amp; J. F. Morales. A." authorityName="Cascante &amp; J. F. Morales. A." box="[258,758,1422,1444]" class="Liliopsida" family="Bromeliaceae" genus="Werauhia" kingdom="Plantae" order="Poales" pageId="2" pageNumber="256" phylum="Tracheophyta" rank="species" species="talamancana">
<emphasis id="9DAE484EA338707BFEBBBCEAFE5895E3" box="[258,480,1422,1443]" italics="true" pageId="2" pageNumber="256">Werauhia talamancana</emphasis>
Cascante &amp; J. F. Morales.A.
</taxonomicName>
plant habit; B. leaf; C.flower; D. inflorescence; E. cupulate stigmata; F. petal with stamen and basal appendages; G. pistil with antesepalous stamens. Drawings by A. Cascante-Marín.
</paragraph>
</caption>
<paragraph id="AF65945CA338707BFF04BC92FC4496E0" blockId="2.[136,1452,1526,2021]" pageId="2" pageNumber="256">
<emphasis id="9DAE484EA338707BFF04BC92FE639650" bold="true" box="[189,475,1526,1552]" pageId="2" pageNumber="256">Phenology and ecology:</emphasis>
—Flowering from February to May; fruiting from February to December. Anthesis is nocturnal and regularly a single flower opens each night, senescence occurs in the morning of the following day. Floral morphology traits (e.g., white petals and campanulate corolla; 
<figureCitation id="37E188D9A338707BFC22BF5AFC4A9618" box="[923,1010,1598,1624]" captionStart="FIGURE 2" captionStartId="3.[136,229,1346,1368]" captionTargetBox="[167,1424,201,1308]" captionTargetId="figure-20@3.[151,1436,190,1322]" captionTargetPageId="3" captionText="FIGURE 2. A. Reproductive plant of Werauhia talamancana; B. Reproductive plant of W. brunei; C. Inflorescence of W.brunei (left) and W. talamancana (right); D. Flowers in lateral view of W.talamancana (above) and W. brunei (below). Photos byA. Cascante-Marín." figureDoi="http://doi.org/10.5281/zenodo.5591462" httpUri="https://zenodo.org/record/5591462/files/figure.png" pageId="2" pageNumber="256">Fig. 2D</figureCitation>
) and anthesis time suggest a nocturnal pollinator, probably bats, following the chiropterophillous pollination syndrome described by 
<bibRefCitation id="CB4BE9ADA338707BFB13BF06FF6B96E0" author="Faegri, K. &amp; van der Pijl, L." pageId="2" pageNumber="256" refId="ref2980" refString="Faegri, K. &amp; van der Pijl, L. (1979) Principles of Pollination Ecology (3 rd edition). Pergamon Press, Oxford, 256 pp." type="book" year="1979">Faegri and van der Pijl (1979)</bibRefCitation>
and related reports in some congeners (
<bibRefCitation id="CB4BE9ADA338707BFD2CBFE2FC4896E0" author="Aguilar-Rodriguez, P. A. &amp; Kromer, T. &amp; Tschapka, M. &amp; Garcia-Franco, J. G. &amp; Escobedo-Sarti, J. &amp; MacSwiney, M. C." box="[661,1008,1670,1696]" pageId="2" pageNumber="256" pagination="1 - 19" refId="ref2658" refString="Aguilar-Rodriguez, P. A., Kromer, T., Tschapka, M., Garcia-Franco, J. G., Escobedo-Sarti, J. &amp; MacSwiney, M. C. (2019) Bat pollination in Bromeliaceae. Plant Ecology and Diversity 12: 1 - 19." type="journal article" year="2019">
Aguilar-Rodríguez 
<emphasis id="9DAE484EA338707BFCCABFE3FC1496E0" box="[883,940,1670,1696]" italics="true" pageId="2" pageNumber="256">et al.</emphasis>
2019
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="E7C0C7D7A338707BFF04BFCEFBFF96A9" pageId="2" pageNumber="256" type="etymology">
<paragraph id="AF65945CA338707BFF04BFCEFBFF96A9" blockId="2.[136,1452,1526,2021]" pageId="2" pageNumber="256">
<emphasis id="9DAE484EA338707BFF04BFCEFEF09684" bold="true" box="[189,328,1706,1732]" pageId="2" pageNumber="256">Etymology:</emphasis>
—The epithet refers to the mountains of the Talamanca mountain range in southern Central America, the center of diversity of 
<taxonomicName id="68DAEFDFA338707BFE18BFAAFDB596A8" authorityName="Grant" authorityYear="1995" box="[417,525,1742,1768]" class="Liliopsida" family="Bromeliaceae" genus="Werauhia" kingdom="Plantae" order="Poales" pageId="2" pageNumber="256" phylum="Tracheophyta" rank="genus">
<emphasis id="9DAE484EA338707BFE18BFAAFDB596A8" box="[417,525,1742,1768]" italics="true" pageId="2" pageNumber="256">Werauhia</emphasis>
</taxonomicName>
and where nearly two-thirds of thespecies occurs.
</paragraph>
</subSubSection>
<subSubSection id="E7C0C7D7A338707BFF04BF96FDBB97A4" pageId="2" pageNumber="256" type="discussion">
<paragraph id="AF65945CA338707BFF04BF96FDBB97A4" blockId="2.[136,1452,1526,2021]" pageId="2" pageNumber="256">
<emphasis id="9DAE484EA338707BFF04BF96FEDC974C" bold="true" box="[189,356,1778,1804]" pageId="2" pageNumber="256">Comments:—</emphasis>
This new species resembles 
<taxonomicName id="68DAEFDFA338707BFD25BF97FAA8974C" authority="(Mez &amp; Werckle 1904: 865) Grant (1995 a: 31)" authorityName="Grant" authorityPageNumber="31" authorityYear="1995" baseAuthorityName="Mez &amp; Werckle" baseAuthorityPageNumber="865" baseAuthorityYear="1904" box="[668,1296,1778,1805]" class="Liliopsida" family="Bromeliaceae" genus="Werauhia" kingdom="Plantae" order="Poales" pageId="2" pageNumber="256" phylum="Tracheophyta" rank="species" species="brunei">
<emphasis id="9DAE484EA338707BFD25BF97FCBF974C" box="[668,775,1778,1804]" italics="true" pageId="2" pageNumber="256">W. brunei</emphasis>
(
<bibRefCitation id="CB4BE9ADA338707BFCADBF96FB83974D" author="Mez, C. &amp; Werckle, K." box="[788,1083,1778,1805]" pageId="2" pageNumber="256" pagination="863 - 878" refId="ref3243" refString="Mez, C. &amp; Werckle, K. (1904) Additamenta monographica. Bulletin de l'Herbier Boissier 9: 863 - 878." type="journal article" year="1904">Mez &amp; Wercklé 1904: 865</bibRefCitation>
) 
<bibRefCitation id="CB4BE9ADA338707BFBF0BF96FAA8974C" author="Grant, J. R." box="[1097,1296,1778,1805]" pageId="2" pageNumber="256" pagination="16 - 57" refId="ref3061" refString="Grant, J. R. (1995 a) The resurrection of Alcantarea and Werauhia, a new genus. Tropische Und Subtropische Pflanzenwelt 91: 16 - 57." type="journal article" year="1995">Grant (1995a: 31)</bibRefCitation>
</taxonomicName>
in the smooth and lustrous floral bracts that are distichous at anthesis (
<figureCitation id="37E188D9A338707BFD47BE72FCEF9770" box="[766,855,1814,1840]" captionStart="FIGURE 2" captionStartId="3.[136,229,1346,1368]" captionTargetBox="[167,1424,201,1308]" captionTargetId="figure-20@3.[151,1436,190,1322]" captionTargetPageId="3" captionText="FIGURE 2. A. Reproductive plant of Werauhia talamancana; B. Reproductive plant of W. brunei; C. Inflorescence of W.brunei (left) and W. talamancana (right); D. Flowers in lateral view of W.talamancana (above) and W. brunei (below). Photos byA. Cascante-Marín." figureDoi="http://doi.org/10.5281/zenodo.5591462" httpUri="https://zenodo.org/record/5591462/files/figure.png" pageId="2" pageNumber="256">Fig. 2C</figureCitation>
). Also, 
<taxonomicName id="68DAEFDFA338707BFC14BE73FBA19770" authorityName="Grant" authorityYear="1995" baseAuthorityName="Mez &amp; Werckle" baseAuthorityYear="1904" box="[941,1049,1814,1840]" class="Liliopsida" family="Bromeliaceae" genus="Werauhia" kingdom="Plantae" order="Poales" pageId="2" pageNumber="256" phylum="Tracheophyta" rank="species" species="brunei">
<emphasis id="9DAE484EA338707BFC14BE73FBA19770" box="[941,1049,1814,1840]" italics="true" pageId="2" pageNumber="256">W. brunei</emphasis>
</taxonomicName>
inhabits montane forests within the same elevation range as 
<emphasis id="9DAE484EA338707BFE1FBE5FFDDC9714" box="[422,612,1850,1876]" italics="true" pageId="2" pageNumber="256">
<taxonomicName id="68DAEFDFA338707BFE1FBE5FFDD89714" box="[422,608,1850,1876]" class="Liliopsida" family="Bromeliaceae" genus="Werauhia" kingdom="Plantae" order="Poales" pageId="2" pageNumber="256" phylum="Tracheophyta" rank="species" species="talamancana">W. talamancana</taxonomicName>
,
</emphasis>
but this new species differs by its longer petals (5.3–6.0 
<emphasis id="9DAE484EA338707BFAB8BE5FFAA09714" box="[1281,1304,1851,1876]" italics="true" pageId="2" pageNumber="256">vs</emphasis>
.4.0– 
<quantity id="682239B9A338707BFAEDBE5EFA1C9715" box="[1364,1444,1850,1877]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" pageId="2" pageNumber="256" unit="cm" value="4.5">4.5 cm</quantity>
) and stamens (4.0–5.0 
<emphasis id="9DAE484EA338707BFE39BE3BFE2F9738" box="[384,407,1887,1912]" italics="true" pageId="2" pageNumber="256">vs</emphasis>
. 
<quantity id="682239B9A338707BFE1FBE3AFD9E9739" box="[422,550,1886,1913]" metricMagnitude="-2" metricUnit="m" metricValue="3.65" metricValueMax="3.7" metricValueMin="3.6" pageId="2" pageNumber="256" unit="cm" value="3.65" valueMax="3.7" valueMin="3.6">3.6–3.7 cm</quantity>
) (
<figureCitation id="37E188D9A338707BFD87BE3AFD219738" box="[574,665,1886,1912]" captionStart="FIGURE 2" captionStartId="3.[136,229,1346,1368]" captionTargetBox="[167,1424,201,1308]" captionTargetId="figure-20@3.[151,1436,190,1322]" captionTargetPageId="3" captionText="FIGURE 2. A. Reproductive plant of Werauhia talamancana; B. Reproductive plant of W. brunei; C. Inflorescence of W.brunei (left) and W. talamancana (right); D. Flowers in lateral view of W.talamancana (above) and W. brunei (below). Photos byA. Cascante-Marín." figureDoi="http://doi.org/10.5281/zenodo.5591462" httpUri="https://zenodo.org/record/5591462/files/figure.png" pageId="2" pageNumber="256">Fig. 2D</figureCitation>
). The brown-maroon color of the floral bracts in 
<taxonomicName id="68DAEFDFA338707BFB7EBE3BFAC49738" box="[1223,1404,1886,1912]" class="Liliopsida" family="Bromeliaceae" genus="Werauhia" kingdom="Plantae" order="Poales" pageId="2" pageNumber="256" phylum="Tracheophyta" rank="species" species="talamancana">
<emphasis id="9DAE484EA338707BFB7EBE3BFAC49738" box="[1223,1404,1886,1912]" italics="true" pageId="2" pageNumber="256">W. talamancana</emphasis>
</taxonomicName>
is a distinctive trait that distinguishes it from 
<taxonomicName id="68DAEFDFA338707BFDE5BEE7FD7097DC" authorityName="Grant" authorityYear="1995" baseAuthorityName="Mez &amp; Werckle" baseAuthorityYear="1904" box="[604,712,1922,1948]" class="Liliopsida" family="Bromeliaceae" genus="Werauhia" kingdom="Plantae" order="Poales" pageId="2" pageNumber="256" phylum="Tracheophyta" rank="species" species="brunei">
<emphasis id="9DAE484EA338707BFDE5BEE7FD7097DC" box="[604,712,1922,1948]" italics="true" pageId="2" pageNumber="256">W. brunei</emphasis>
</taxonomicName>
(
<figureCitation id="37E188D9A338707BFD60BEE6FC8B97DC" box="[729,819,1922,1948]" captionStart="FIGURE 2" captionStartId="3.[136,229,1346,1368]" captionTargetBox="[167,1424,201,1308]" captionTargetId="figure-20@3.[151,1436,190,1322]" captionTargetPageId="3" captionText="FIGURE 2. A. Reproductive plant of Werauhia talamancana; B. Reproductive plant of W. brunei; C. Inflorescence of W.brunei (left) and W. talamancana (right); D. Flowers in lateral view of W.talamancana (above) and W. brunei (below). Photos byA. Cascante-Marín." figureDoi="http://doi.org/10.5281/zenodo.5591462" httpUri="https://zenodo.org/record/5591462/files/figure.png" pageId="2" pageNumber="256">Fig. 2C</figureCitation>
).This qualitative characteristic is atypical among singlespicate species of 
<emphasis id="9DAE484EA338707BFEEDBEC2FE7F9780" box="[340,455,1958,1984]" italics="true" pageId="2" pageNumber="256">
<taxonomicName id="68DAEFDFA338707BFEEDBEC2FE7B9780" authorityName="Grant" authorityYear="1995" box="[340,451,1958,1984]" class="Liliopsida" family="Bromeliaceae" genus="Werauhia" kingdom="Plantae" order="Poales" pageId="2" pageNumber="256" phylum="Tracheophyta" rank="genus">Werauhia</taxonomicName>
,
</emphasis>
which duringanthesis usually exhibit floral bracts with green tones and sometimes with brownish bract tips due to drying.
</paragraph>
</subSubSection>
<caption id="FBA5C4D4A339707AFF31BC26FACD953C" ID-DOI="http://doi.org/10.5281/zenodo.5591462" ID-Zenodo-Dep="5591462" httpUri="https://zenodo.org/record/5591462/files/figure.png" pageId="3" pageNumber="257" startId="3.[136,229,1346,1368]" targetBox="[167,1424,201,1308]" targetPageId="3">
<paragraph id="AF65945CA339707AFF31BC26FACD953C" blockId="3.[136,1452,1346,1404]" pageId="3" pageNumber="257">
<emphasis id="9DAE484EA339707AFF31BC26FF459517" bold="true" box="[136,253,1346,1368]" pageId="3" pageNumber="257">FIGURE 2.</emphasis>
A. Reproductive plant of 
<taxonomicName id="68DAEFDFA339707AFE4CBC26FD6B9517" box="[501,723,1346,1367]" class="Liliopsida" family="Bromeliaceae" genus="Werauhia" kingdom="Plantae" order="Poales" pageId="3" pageNumber="257" phylum="Tracheophyta" rank="species" species="talamancana">
<emphasis id="9DAE484EA339707AFE4CBC26FD6B9517" box="[501,723,1346,1367]" italics="true" pageId="3" pageNumber="257">Werauhia talamancana</emphasis>
</taxonomicName>
; B. Reproductive plant of 
<taxonomicName id="68DAEFDFA339707AFC68BC27FB959517" authorityName="Grant" authorityYear="1995" baseAuthorityName="Mez &amp; Werckle" baseAuthorityYear="1904" box="[977,1069,1346,1368]" class="Liliopsida" family="Bromeliaceae" genus="Werauhia" kingdom="Plantae" order="Poales" pageId="3" pageNumber="257" phylum="Tracheophyta" rank="species" species="brunei">
<emphasis id="9DAE484EA339707AFC68BC27FB959517" box="[977,1069,1346,1368]" italics="true" pageId="3" pageNumber="257">W. brunei</emphasis>
</taxonomicName>
; C. Inflorescence of 
<taxonomicName id="68DAEFDFA339707AFB4DBC27FAF79517" authorityName="Grant" authorityYear="1995" baseAuthorityName="Mez &amp; Werckle" baseAuthorityYear="1904" box="[1268,1359,1346,1368]" class="Liliopsida" family="Bromeliaceae" genus="Werauhia" kingdom="Plantae" order="Poales" pageId="3" pageNumber="257" phylum="Tracheophyta" rank="species" species="brunei">
<emphasis id="9DAE484EA339707AFB4DBC27FAF79517" box="[1268,1359,1346,1368]" italics="true" pageId="3" pageNumber="257">W. brunei</emphasis>
</taxonomicName>
(left) and 
<taxonomicName id="68DAEFDFA339707AFF31BC03FE9A953B" box="[136,290,1382,1404]" class="Liliopsida" family="Bromeliaceae" genus="Werauhia" kingdom="Plantae" order="Poales" pageId="3" pageNumber="257" phylum="Tracheophyta" rank="species" species="talamancana">
<emphasis id="9DAE484EA339707AFF31BC03FE9A953B" box="[136,290,1382,1404]" italics="true" pageId="3" pageNumber="257">W. talamancana</emphasis>
</taxonomicName>
(right); D. Flowers in lateral view of 
<taxonomicName id="68DAEFDFA339707AFD30BC03FCA6953B" box="[649,798,1382,1404]" class="Liliopsida" family="Bromeliaceae" genus="Werauhia" kingdom="Plantae" order="Poales" pageId="3" pageNumber="257" phylum="Tracheophyta" rank="species" species="talamancana">
<emphasis id="9DAE484EA339707AFD30BC03FCA6953B" box="[649,798,1382,1404]" italics="true" pageId="3" pageNumber="257">W.talamancana</emphasis>
</taxonomicName>
(above) and 
<taxonomicName id="68DAEFDFA339707AFC25BC03FC40953B" authorityName="Grant" authorityYear="1995" baseAuthorityName="Mez &amp; Werckle" baseAuthorityYear="1904" box="[924,1016,1382,1404]" class="Liliopsida" family="Bromeliaceae" genus="Werauhia" kingdom="Plantae" order="Poales" pageId="3" pageNumber="257" phylum="Tracheophyta" rank="species" species="brunei">
<emphasis id="9DAE484EA339707AFC25BC03FC40953B" box="[924,1016,1382,1404]" italics="true" pageId="3" pageNumber="257">W. brunei</emphasis>
</taxonomicName>
(below). Photos by A. Cascante-Marín.
</paragraph>
</caption>
<subSubSection id="E7C0C7D7A339707AFF04BCCEFAA296A5" pageId="3" pageNumber="257" type="reference_group">
<paragraph id="AF65945CA339707AFF04BCCEFAA296A5" blockId="3.[136,1452,1450,1765]" pageId="3" pageNumber="257">
<treatmentCitationGroup id="8FCAB372A339707AFF04BCCEFAA296A5" pageId="3" pageNumber="257">
<taxonomicName id="68DAEFDFA339707AFF04BCCEFE069584" ID-CoL="9V7ZS" authorityName="Cascante &amp; J. F. Morales" authorityYear="2021" box="[189,446,1450,1476]" class="Liliopsida" family="Bromeliaceae" genus="Werauhia" kingdom="Plantae" order="Poales" pageId="3" pageNumber="257" phylum="Tracheophyta" rank="species" species="talamancana">
<emphasis id="9DAE484EA339707AFF04BCCEFE069584" box="[189,446,1450,1476]" italics="true" pageId="3" pageNumber="257">Werauhia talamancana</emphasis>
</taxonomicName>
falls into 
<bibRefCitation id="CB4BE9ADA339707AFD9ABCCFFD689585" author="Grant, J. R." box="[547,720,1450,1477]" pageId="3" pageNumber="257" pagination="16 - 57" refId="ref3061" refString="Grant, J. R. (1995 a) The resurrection of Alcantarea and Werauhia, a new genus. Tropische Und Subtropische Pflanzenwelt 91: 16 - 57." type="journal article" year="1995">Grant’s (1995a)</bibRefCitation>
section 
<taxonomicName id="68DAEFDFA339707AFC9CBCCEFC299584" ID-CoL="8WM3G" authorityName="Grant" authorityYear="1995" box="[805,913,1450,1476]" class="Liliopsida" family="Bromeliaceae" genus="Werauhia" kingdom="Plantae" order="Poales" pageId="3" pageNumber="257" phylum="Tracheophyta" rank="genus">
<emphasis id="9DAE484EA339707AFC9CBCCEFC299584" box="[805,913,1450,1476]" italics="true" pageId="3" pageNumber="257">Werauhia</emphasis>
</taxonomicName>
,which includes plants with simple inflorescence. Following the taxonomic key provided by 
<bibRefCitation id="CB4BE9ADA339707AFDD2BCAAFC9095A9" author="Morales, J. F." box="[619,808,1486,1513]" pageId="3" pageNumber="257" pagination="297 - 375" refId="ref3301" refString="Morales, J. F. (2003 a) Bromeliaceae. Manual de Plantas de Costa Rica. Volumen 2. Gimnospermas y Monocotiledoneas (Agavaceae- Musaceae). Monographs in Systematic Botany from the Missouri Botanical Garden 92: 297 - 375." type="journal article" year="2003">Morales (2003a)</bibRefCitation>
, the new species belongs to a species group that exhibits relatively large floral bracts (3.0– 
<quantity id="682239B9A339707AFDBEBC96FDEF964D" box="[519,599,1522,1549]" metricMagnitude="-2" metricUnit="m" metricValue="6.3" pageId="3" pageNumber="257" unit="cm" value="6.3">6.3 cm</quantity>
) with smooth and lustrous or rugose surfaces. This species group includes among others 
<taxonomicName id="68DAEFDFA339707AFE9FBF73FE369670" ID-CoL="7FW2J" authorityName="Grant" authorityYear="1995" baseAuthorityName="Smith" baseAuthorityYear="1963" box="[294,398,1558,1584]" class="Liliopsida" family="Bromeliaceae" genus="Werauhia" kingdom="Plantae" order="Poales" pageId="3" pageNumber="257" phylum="Tracheophyta" rank="species" species="ampla">
<emphasis id="9DAE484EA339707AFE9FBF73FE369670" box="[294,398,1558,1584]" italics="true" pageId="3" pageNumber="257">W. ampla</emphasis>
</taxonomicName>
, 
<treatmentCitation id="2E7BB24DA339707AFE25BF73FC0C9671" author="Grant, J. R." box="[412,948,1558,1585]" page="174" pageId="3" pageNumber="257" year="1995">
<taxonomicName id="68DAEFDFA339707AFE25BF73FC0C9671" ID-CoL="7G82K" authority="(Smith 1960: 174) Grant (1995 a: 30)" authorityName="Grant" authorityPageNumber="30" authorityYear="1995" baseAuthorityName="Smith" baseAuthorityPageNumber="174" baseAuthorityYear="1960" box="[412,948,1558,1585]" class="Liliopsida" family="Bromeliaceae" genus="Werauhia" kingdom="Plantae" order="Poales" pageId="3" pageNumber="257" phylum="Tracheophyta" rank="species" species="bicolor">
<emphasis id="9DAE484EA339707AFE25BF73FDB79670" box="[412,527,1558,1584]" italics="true" pageId="3" pageNumber="257">W. bicolor</emphasis>
(
<bibRefCitation id="CB4BE9ADA339707AFDA4BF72FD629671" author="Smith, L. B." box="[541,730,1558,1585]" pageId="3" pageNumber="257" pagination="169 - 179" refId="ref3444" refString="Smith, L. B. (1960) Notes on Bromeliaceae, XIV. Phytologia 7: 169 - 179." type="journal article" year="1960">Smith 1960: 174</bibRefCitation>
) 
<bibRefCitation id="CB4BE9ADA339707AFD50BF73FC0C9671" author="Grant, J. R." box="[745,948,1558,1585]" pageId="3" pageNumber="257" pagination="16 - 57" refId="ref3061" refString="Grant, J. R. (1995 a) The resurrection of Alcantarea and Werauhia, a new genus. Tropische Und Subtropische Pflanzenwelt 91: 16 - 57." type="journal article" year="1995">Grant (1995a: 30)</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, 
<treatmentCitation id="2E7BB24DA339707AFC06BF73FF069615" author="Grant, J. R." page="326" pageId="3" pageNumber="257" year="1995">
<taxonomicName id="68DAEFDFA339707AFC06BF73FF069615" ID-CoL="7FW2M" authority="(Smith 1974: 326) Grant (1995 b: 121)" authorityName="Grant" authorityPageNumber="121" authorityYear="1995" baseAuthorityName="Smith" baseAuthorityPageNumber="326" baseAuthorityYear="1974" class="Liliopsida" family="Bromeliaceae" genus="Werauhia" kingdom="Plantae" order="Poales" pageId="3" pageNumber="257" phylum="Tracheophyta" rank="species" species="burgeri">
<emphasis id="9DAE484EA339707AFC06BF73FB8C9670" box="[959,1076,1558,1584]" italics="true" pageId="3" pageNumber="257">W. burgeri</emphasis>
(
<bibRefCitation id="CB4BE9ADA339707AFBFBBF72FAB89671" author="Smith, L. B." box="[1090,1280,1558,1585]" pageId="3" pageNumber="257" pagination="319 - 333" refId="ref3488" refString="Smith, L. B. (1974) Notes on Bromeliaceae, XXXVI. Phytologia 28: 319 - 333." type="journal article" year="1974">Smith 1974: 326</bibRefCitation>
) 
<bibRefCitation id="CB4BE9ADA339707AFAB7BF73FF069615" author="Grant, J. R." pageId="3" pageNumber="257" pagination="119 - 123" refId="ref3092" refString="Grant, J. R. (1995 b) Addendum to &quot; the resurrection of Alcantarea and Werauhia, a new genus &quot; (Bromeliaceae: Tillandsioideae). Phytologia 78: 119 - 123." type="journal article" year="1995">Grant (1995b: 121)</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, 
<treatmentCitation id="2E7BB24DA339707AFF74BF5FFCE49615" author="Grant, J. R." box="[205,860,1594,1621]" page="31" pageId="3" pageNumber="257" year="1863">
<taxonomicName id="68DAEFDFA339707AFF74BF5FFCE49615" ID-CoL="5BXBD" authority="(Wendland 1863: 31) Grant (1995 a: 31)" authorityName="Grant" authorityPageNumber="31" authorityYear="1995" baseAuthorityName="Wendland" baseAuthorityPageNumber="31" baseAuthorityYear="1863" box="[205,860,1594,1621]" class="Liliopsida" family="Bromeliaceae" genus="Werauhia" kingdom="Plantae" order="Poales" pageId="3" pageNumber="257" phylum="Tracheophyta" rank="species" species="gladioliflora">
<emphasis id="9DAE484EA339707AFF74BF5FFE399614" box="[205,385,1594,1620]" italics="true" pageId="3" pageNumber="257">W. gladioliflora</emphasis>
(
<bibRefCitation id="CB4BE9ADA339707AFE2ABF5EFDC09615" author="Wendland, H. A." box="[403,632,1594,1621]" pageId="3" pageNumber="257" pagination="29 - 34" refId="ref3567" refString="Wendland, H. A. (1863) Beschreiburgen einiger neuer Pflanzen aus dem Kgl. Berggarten zu Herrenhausen bei Hannover. Hamburger Garten-Blumenzeitung 19: 29 - 34." type="journal article" year="1863">Wendland 1863: 31</bibRefCitation>
) 
<bibRefCitation id="CB4BE9ADA339707AFD32BF5FFCE49615" author="Grant, J. R." box="[651,860,1594,1621]" pageId="3" pageNumber="257" pagination="16 - 57" refId="ref3061" refString="Grant, J. R. (1995 a) The resurrection of Alcantarea and Werauhia, a new genus. Tropische Und Subtropische Pflanzenwelt 91: 16 - 57." type="journal article" year="1995">Grant (1995a: 31)</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, 
<treatmentCitation id="2E7BB24DA339707AFCD5BF5FFEA59639" author="Mez, C. &amp; Werckle, K." page="867" pageId="3" pageNumber="257" year="1904">
<taxonomicName id="68DAEFDFA339707AFCD5BF5FFEA59639" ID-CoL="5BXBZ" authority="(Mez &amp; Werckle 1904: 867) Grant (1995 b: 121)" authorityName="Grant" authorityPageNumber="121" authorityYear="1995" baseAuthorityName="Mez &amp; Werckle" baseAuthorityPageNumber="867" baseAuthorityYear="1904" class="Liliopsida" family="Bromeliaceae" genus="Werauhia" kingdom="Plantae" order="Poales" pageId="3" pageNumber="257" phylum="Tracheophyta" rank="species" species="macrantha">
<emphasis id="9DAE484EA339707AFCD5BF5FFBB49614" box="[876,1036,1594,1620]" italics="true" pageId="3" pageNumber="257">W. macrantha</emphasis>
(
<bibRefCitation id="CB4BE9ADA339707AFBA7BF5FFAE19615" author="Mez, C. &amp; Werckle, K." box="[1054,1369,1594,1621]" pageId="3" pageNumber="257" pagination="863 - 878" refId="ref3243" refString="Mez, C. &amp; Werckle, K. (1904) Additamenta monographica. Bulletin de l'Herbier Boissier 9: 863 - 878." type="journal article" year="1904">Mez &amp; Wercklé 1904: 867</bibRefCitation>
) 
<bibRefCitation id="CB4BE9ADA339707AFAD5BF5FFEA59639" author="Grant, J. R." pageId="3" pageNumber="257" pagination="119 - 123" refId="ref3092" refString="Grant, J. R. (1995 b) Addendum to &quot; the resurrection of Alcantarea and Werauhia, a new genus &quot; (Bromeliaceae: Tillandsioideae). Phytologia 78: 119 - 123." type="journal article" year="1995">Grant (1995b: 121)</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, 
<treatmentCitation id="2E7BB24DA339707AFE93BF3BFB949639" author="Mez, C. &amp; Werckle, K." box="[298,1068,1630,1657]" page="865" pageId="3" pageNumber="257" year="1904">
<taxonomicName id="68DAEFDFA339707AFE93BF3BFB949639" ID-CoL="5BXC2" authority="(Mez &amp; Werckle 1904: 865) Morales (2003 b: 65)" authorityName="Morales" authorityPageNumber="65" authorityYear="2003" baseAuthorityName="Mez &amp; Werckle" baseAuthorityPageNumber="865" baseAuthorityYear="1904" box="[298,1068,1630,1657]" class="Liliopsida" family="Bromeliaceae" genus="Werauhia" kingdom="Plantae" order="Poales" pageId="3" pageNumber="257" phylum="Tracheophyta" rank="species" species="macrochlamys">
<emphasis id="9DAE484EA339707AFE93BF3BFE489638" box="[298,496,1630,1656]" italics="true" pageId="3" pageNumber="257">W. macrochlamys</emphasis>
(
<bibRefCitation id="CB4BE9ADA339707AFDB9BF3BFC889639" author="Mez, C. &amp; Werckle, K." box="[512,816,1630,1657]" pageId="3" pageNumber="257" pagination="863 - 878" refId="ref3243" refString="Mez, C. &amp; Werckle, K. (1904) Additamenta monographica. Bulletin de l'Herbier Boissier 9: 863 - 878." type="journal article" year="1904">Mez &amp; Wercklé 1904: 865</bibRefCitation>
) 
<bibRefCitation id="CB4BE9ADA339707AFCF8BF3AFB949639" author="Morales, J. F." box="[833,1068,1630,1657]" pageId="3" pageNumber="257" pagination="65" refId="ref3346" refString="Morales, J. F. (2003 b) New combinations in Werauhia (Bromeliaceae) from Costa Rica. Lundiana 4: 65." type="journal article" year="2003">Morales (2003b: 65)</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, 
<treatmentCitation id="2E7BB24DA339707AFB81BF3BFED796DD" author="Morales, J. F." page="403" pageId="3" pageNumber="257" year="2003">
<taxonomicName id="68DAEFDFA339707AFB81BF3BFED796DD" ID-CoL="5BXCF" authority="(Morales 1999: 403) Morales (2003 c: 15)" authorityName="Morales" authorityPageNumber="15" authorityYear="2003" baseAuthorityName="Morales" baseAuthorityPageNumber="403" baseAuthorityYear="1999" class="Liliopsida" family="Bromeliaceae" genus="Werauhia" kingdom="Plantae" order="Poales" pageId="3" pageNumber="257" phylum="Tracheophyta" rank="species" species="osaensis">
<emphasis id="9DAE484EA339707AFB81BF3BFB039638" box="[1080,1211,1631,1656]" italics="true" pageId="3" pageNumber="257">W. osaensis</emphasis>
(
<bibRefCitation id="CB4BE9ADA339707AFB73BF3AFA1B9639" author="Morales, J. F." box="[1226,1443,1630,1657]" pageId="3" pageNumber="257" pagination="401 - 406" refId="ref3268" refString="Morales, J. F. (1999) Seis nuevas especies de Vriesea sect. Xiphion (Bromeliaceae: Tillandsioideae) para Costa Rica. Novon 9: 401 - 406." type="journal article" year="1999">Morales 1999: 403</bibRefCitation>
) 
<bibRefCitation id="CB4BE9ADA339707AFF31BFE6FED796DD" author="Morales, J. F." box="[136,367,1666,1693]" pageId="3" pageNumber="257" pagination="109 - 111" refId="ref3372" refString="Morales, J. F. (2003 c) Nuevas combinaciones y un nuevo nombre en las Bromeliaceae de Costa Rica. Polibotanica 15: 109 - 111." type="journal article" year="2003">Morales (2003c: 15)</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, 
<treatmentCitation id="2E7BB24DA339707AFEC2BFE7FC4496DD" author="Morales, J. F." box="[379,1020,1666,1693]" page="405" pageId="3" pageNumber="257" year="1999">
<taxonomicName id="68DAEFDFA339707AFEC2BFE7FC4496DD" ID-CoL="5BXD5" authority="(Morales 1999: 405) Morales (2003 c: 110)" authorityName="Morales" authorityPageNumber="110" authorityYear="2003" baseAuthorityName="Morales" baseAuthorityPageNumber="405" baseAuthorityYear="1999" box="[379,1020,1666,1693]" class="Liliopsida" family="Bromeliaceae" genus="Werauhia" kingdom="Plantae" order="Poales" pageId="3" pageNumber="257" phylum="Tracheophyta" rank="species" species="tiquirensis">
<emphasis id="9DAE484EA339707AFEC2BFE7FDAB96DC" box="[379,531,1667,1692]" italics="true" pageId="3" pageNumber="257">W. tiquirensis</emphasis>
(
<bibRefCitation id="CB4BE9ADA339707AFD9BBFE6FD4196DD" author="Morales, J. F." box="[546,761,1666,1693]" pageId="3" pageNumber="257" pagination="401 - 406" refId="ref3268" refString="Morales, J. F. (1999) Seis nuevas especies de Vriesea sect. Xiphion (Bromeliaceae: Tillandsioideae) para Costa Rica. Novon 9: 401 - 406." type="journal article" year="1999">Morales 1999: 405</bibRefCitation>
) 
<bibRefCitation id="CB4BE9ADA339707AFCB1BFE6FC4496DD" author="Morales, J. F." box="[776,1020,1666,1693]" pageId="3" pageNumber="257" pagination="109 - 111" refId="ref3372" refString="Morales, J. F. (2003 c) Nuevas combinaciones y un nuevo nombre en las Bromeliaceae de Costa Rica. Polibotanica 15: 109 - 111." type="journal article" year="2003">Morales (2003c: 110)</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, and 
<treatmentCitation id="2E7BB24DA339707AFB8EBFE7FEEB9681" author="Grant, J. R." page="166" pageId="3" pageNumber="257" year="1995">
<taxonomicName id="68DAEFDFA339707AFB8EBFE7FEEB9681" ID-CoL="5BXD6" authority="(Smith 1938: 166) Grant (1995 a: 35)" authorityName="Grant" authorityPageNumber="35" authorityYear="1995" baseAuthorityName="Smith" baseAuthorityPageNumber="166" baseAuthorityYear="1938" class="Liliopsida" family="Bromeliaceae" genus="Werauhia" kingdom="Plantae" order="Poales" pageId="3" pageNumber="257" phylum="Tracheophyta" rank="species" species="tonduziana">
<emphasis id="9DAE484EA339707AFB8EBFE7FB6E96DC" box="[1079,1238,1666,1692]" italics="true" pageId="3" pageNumber="257">W. tonduziana</emphasis>
(
<bibRefCitation id="CB4BE9ADA339707AFB5CBFE6FA1B96DD" author="Smith, L. B." box="[1253,1443,1666,1693]" pageId="3" pageNumber="257" pagination="61 - 228" refId="ref3402" refString="Smith, L. B. (1938) Bromeliaceae. North American Flora 19: 61 - 228." type="journal article" year="1938">Smith 1938: 166</bibRefCitation>
) 
<bibRefCitation id="CB4BE9ADA339707AFF31BFC3FEEB9681" author="Grant, J. R." box="[136,339,1702,1729]" pageId="3" pageNumber="257" pagination="16 - 57" refId="ref3061" refString="Grant, J. R. (1995 a) The resurrection of Alcantarea and Werauhia, a new genus. Tropische Und Subtropische Pflanzenwelt 91: 16 - 57." type="journal article" year="1995">Grant (1995a: 35)</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, that are very similar in appearance and difficult to keep apart from another (
<bibRefCitation id="CB4BE9ADA339707AFB15BFC2FAF49681" author="Morales, J. F." box="[1196,1356,1702,1729]" pageId="3" pageNumber="257" pagination="401 - 406" refId="ref3268" refString="Morales, J. F. (1999) Seis nuevas especies de Vriesea sect. Xiphion (Bromeliaceae: Tillandsioideae) para Costa Rica. Novon 9: 401 - 406." type="journal article" year="1999">Morales 1999</bibRefCitation>
, 
<bibRefCitation id="CB4BE9ADA339707AFAEEBFC3FA189681" author="Morales, J. F." box="[1367,1440,1703,1729]" pageId="3" pageNumber="257" pagination="297 - 375" refId="ref3301" refString="Morales, J. F. (2003 a) Bromeliaceae. Manual de Plantas de Costa Rica. Volumen 2. Gimnospermas y Monocotiledoneas (Agavaceae- Musaceae). Monographs in Systematic Botany from the Missouri Botanical Garden 92: 297 - 375." type="journal article" year="2003">2003a</bibRefCitation>
). Molecular and floral morphology studies are necessary to elucidate the relationships between these taxa.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
</treatment>
</document>