428 lines
60 KiB
XML
428 lines
60 KiB
XML
<document id="84EEA78974B002CE2685D0E472F4B86D" ID-CLB-Dataset="28395" ID-DOI="10.11646/zootaxa.4565.2.4" ID-GBIF-Dataset="f0b05826-808f-41ea-94be-a06da11e1b2b" ID-ISSN="1175-5326" ID-Zenodo-Dep="2589876" ID-ZooBank="9C32EE7B-2D00-4E83-8C55-2E84758F8F80" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1552284609444" checkinUser="plazi" docAuthor="Zhibin, Gan & Xinzheng, Li" docDate="2019" docId="271587955016FFF0FF2422B9FD2E386B" docLanguage="en" docName="zootaxa.4565.2.4.pdf" docOrigin="Zootaxa 4565 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Trianguloscalpellum regium" docType="treatment" docVersion="13" lastPageNumber="205" masterDocId="DB2CFFED5017FFF4FFB32619FF8C3E75" masterDocTitle="Report on four deep-water barnacles (Cirripedia, Thoracica) from the north west Pacific, with remarks on Trianguloscalpellum regium (Wyville-Thomson, 1873)" masterLastPageNumber="212" masterPageNumber="201" pageNumber="202" updateTime="1698701496248" updateUser="ExternalLinkService">
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<mods:title id="F7668D47E1B5C5DEB60ABE220BA97BCE">Report on four deep-water barnacles (Cirripedia, Thoracica) from the north west Pacific, with remarks on Trianguloscalpellum regium (Wyville-Thomson, 1873)</mods:title>
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<mods:namePart id="D4E15D26DD5398CD226FF69240B6264A">Zhibin, Gan</mods:namePart>
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<mods:namePart id="190A6AECC930B117B7DCD1A0B1221253">Xinzheng, Li</mods:namePart>
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<mods:title id="1A1349658233303EC2FF3DFA243AF445">Zootaxa</mods:title>
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<mods:date id="F2F142F778A86E9673E71A6ECBD86857">2019</mods:date>
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<treatment id="271587955016FFF0FF2422B9FD2E386B" ID-DOI="http://doi.org/10.5281/zenodo.3513801" ID-GBIF-Taxon="154366903" ID-Zenodo-Dep="3513801" LSID="urn:lsid:plazi:treatment:271587955016FFF0FF2422B9FD2E386B" httpUri="http://treatment.plazi.org/id/271587955016FFF0FF2422B9FD2E386B" lastPageId="4" lastPageNumber="205" pageId="1" pageNumber="202">
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<subSubSection id="E7A665085016FFF5FF2422B9FE903AAE" pageId="1" pageNumber="202" type="nomenclature">
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<paragraph id="AF0336835016FFF5FF2422B9FCBD3ACF" blockId="1.[151,817,1184,1243]" box="[151,817,1184,1210]" pageId="1" pageNumber="202">
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<heading id="F44B81EF5016FFF5FF2422B9FCBD3ACF" bold="true" box="[151,817,1184,1210]" fontSize="11" level="1" pageId="1" pageNumber="202" reason="1">
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<taxonomicName id="68BC4D005016FFF5FF2422B9FCBD3ACF" authority="(Wyville-Thomson, 1873)" baseAuthorityName="Wyville-Thomson" baseAuthorityYear="1873" box="[151,817,1184,1210]" class="Maxillopoda" family="Scalpellidae" genus="Trianguloscalpellum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pedunculata" pageId="1" pageNumber="202" phylum="Arthropoda" rank="species" species="regium">
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<emphasis id="9DC8EA915016FFF5FF2422B9FCBD3ACF" bold="true" box="[151,817,1184,1210]" pageId="1" pageNumber="202">
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<emphasis id="9DC8EA915016FFF5FF2422B9FE633ACF" bold="true" box="[151,495,1184,1210]" italics="true" pageId="1" pageNumber="202">Trianguloscalpellum regium</emphasis>
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(
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<bibRefCitation id="CB2D4B725016FFF5FE4C22B9FCA53ACF" author="Wyville-Thomson, C." box="[511,809,1184,1210]" pageId="1" pageNumber="202" pagination="347 - 349" refId="ref5756" refString="Wyville-Thomson, C. (1873) Notes from the " Challenger ". Nature, 8, 347 - 349. https: // doi. org / 10.1038 / 008347 a 0" type="journal article" year="1873">Wyville-Thomson, 1873</bibRefCitation>
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)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="AF0336835016FFF5FF2422DBFE903AAE" blockId="1.[151,817,1184,1243]" box="[151,284,1218,1243]" pageId="1" pageNumber="202">Figures 1–3</paragraph>
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</subSubSection>
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<subSubSection id="E7A665085016FFF5FF242313FE253B09" pageId="1" pageNumber="202" type="reference_group">
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<paragraph id="AF0336835016FFF5FF242313FAE63B55" blockId="1.[151,1437,1290,1404]" box="[151,1386,1290,1312]" pageId="1" pageNumber="202">
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<treatmentCitationGroup id="8FAC11AD5016FFF5FF242313FAE63B55" box="[151,1386,1290,1312]" pageId="1" pageNumber="202">
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<taxonomicName id="68BC4D005016FFF5FF242313FD093B55" authority="Wyville-Thomson, 1873: 347" authorityName="Wyville-Thomson" authorityPageNumber="347" authorityYear="1873" box="[151,645,1290,1312]" class="Maxillopoda" family="Scalpellidae" genus="Scalpellum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pedunculata" pageId="1" pageNumber="202" phylum="Arthropoda" rank="species" species="regium">
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<emphasis id="9DC8EA915016FFF5FF242313FED83B55" box="[151,340,1290,1312]" italics="true" pageId="1" pageNumber="202">Scalpellum regium</emphasis>
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<treatmentCitation id="2E1D10925016FFF5FEE92313FD093B55" author="Wyville-Thomson, C." box="[346,645,1290,1312]" page="347" pageId="1" pageNumber="202" year="1873">
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<bibRefCitation id="CB2D4B725016FFF5FEE92313FDD93B55" author="Wyville-Thomson, C." box="[346,597,1290,1312]" pageId="1" pageNumber="202" pagination="347 - 349" refId="ref5756" refString="Wyville-Thomson, C. (1873) Notes from the " Challenger ". Nature, 8, 347 - 349. https: // doi. org / 10.1038 / 008347 a 0" type="journal article" year="1873">Wyville-Thomson, 1873</bibRefCitation>
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: 347
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</treatmentCitation>
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</taxonomicName>
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–348.—
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<treatmentCitation id="2E1D10925016FFF5FD662313FCF03B55" author="Hoek, P. P. C." box="[725,892,1290,1312]" page="106" pageId="1" pageNumber="202" year="1883">
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<bibRefCitation id="CB2D4B725016FFF5FD662313FCC63B55" author="Hoek, P. P. C." box="[725,842,1290,1312]" pageId="1" pageNumber="202" pagination="1 - 169" refId="ref4892" refString="Hoek, P. P. C. (1883) Report on the Cirripedia collected by H. M. S. " Challenger " during the years 1873 - 1876. Report on the Scientific Results of the Foyage H. M. S. " Challenger ", Zoology, part 25, 8, 1 - 169." type="journal article" year="1883">Hoek 1883</bibRefCitation>
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: 106
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</treatmentCitation>
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, pl. IV figs 3–5.—
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<treatmentCitation id="2E1D10925016FFF5FB852313FB683B55" author="Pilsbry, H. A." box="[1078,1252,1290,1312]" page="28" pageId="1" pageNumber="202" year="1907">
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<bibRefCitation id="CB2D4B725016FFF5FB852313FB323B55" author="Pilsbry, H. A." box="[1078,1214,1290,1312]" pageId="1" pageNumber="202" pagination="1 - 122" refId="ref5429" refString="Pilsbry, H. A. (1907) The Barnacles (Cirripedia) contained in the collections of the U. S. National Museum. Bulletin of the United States National Museum, 60, 1 - 122. https: // doi. org / 10.5479 / si. 03629236.60.1" type="journal article" year="1907">Pilsbry 1907</bibRefCitation>
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: 28
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</treatmentCitation>
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, pl. III fig. 5.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="AF0336835016FFF5FF242330FE253B09" blockId="1.[151,1437,1290,1404]" pageId="1" pageNumber="202">
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<treatmentCitationGroup id="8FAC11AD5016FFF5FF242330FE253B09" pageId="1" pageNumber="202">
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<taxonomicName id="68BC4D005016FFF5FF242330FE3A3B4A" baseAuthorityName="Wyville-Thomson" baseAuthorityYear="1873" box="[151,438,1321,1343]" class="Maxillopoda" family="Scalpellidae" genus="Trianguloscalpellum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pedunculata" pageId="1" pageNumber="202" phylum="Arthropoda" rank="species" species="regium">
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<emphasis id="9DC8EA915016FFF5FF242330FE3A3B4A" box="[151,438,1321,1343]" italics="true" pageId="1" pageNumber="202">Trianguloscalpellum regium</emphasis>
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</taxonomicName>
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.—
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<treatmentCitation id="2E1D10925016FFF5FE602333FD093B4A" author="Young, P. S." box="[467,645,1321,1343]" page="28" pageId="1" pageNumber="202" year="1998">
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<bibRefCitation id="CB2D4B725016FFF5FE602333FDED3B4A" author="Young, P. S." box="[467,609,1321,1343]" pageId="1" pageNumber="202" pagination="31 - 92" refId="ref5846" refString="Young, P. S. (1998 a) Cirripedia (Crustacea) from the " Campagne Biacores " in the Azores region, including a generic revision of Verrucidae. Zoosystema, 20, 31 - 92." type="journal article" year="1998">Young 1998a</bibRefCitation>
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: 28
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</treatmentCitation>
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–30, figs 20a–h, 22.—1998b: 111, fig. 2c.—1999: 617, fig. 5B.—2001a: 464, fig. 7.—2001b: 741.—
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<treatmentCitation id="2E1D10925016FFF5FE1D2351FD0F3B2B" ID-DOI="http://doi.org/10.5281/zenodo.3506916" ID-Zenodo-Dep="3506916" author="Chan, B. K. & Prabowo, R. E. & Lee, K. S." box="[430,643,1352,1374]" httpUri="http://treatment.plazi.org/id/C1306911FFCA0F6EC988F95BFAB9FCD3" page="13" pageId="1" pageNumber="202" year="2010">
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<bibRefCitation id="CB2D4B725016FFF5FE1D2351FDD23B2B" author="Chan, B. K. & Prabowo, R. E. & Lee, K. S." box="[430,606,1352,1374]" pageId="1" pageNumber="202" pagination="1 - 47" refId="ref4418" refString="Chan, B. K., Prabowo, R. E. & Lee, K. S. (2010) North West Pacific deep-sea barnacles (Cirripedia, Thoracica) collected by the TAIWAN expeditions, with descriptions of two new species. Zootaxa, 2405, 1 - 47. https: // doi. org / 10.11646 / zootaxa. 2405.1.1" type="journal article" year="2010">
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Chan
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<emphasis id="9DC8EA915016FFF5FE5F2350FDAD3B2B" box="[492,545,1352,1374]" italics="true" pageId="1" pageNumber="202">et al.</emphasis>
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2010
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</bibRefCitation>
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||
: 13
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||
</treatmentCitation>
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, figs 1G, 10.—
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||
<treatmentCitation id="2E1D10925016FFF5FCA92351FC623B2B" author="Poltarukha, O. P." box="[794,1006,1352,1374]" page="37" pageId="1" pageNumber="202" year="2012">
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<bibRefCitation id="CB2D4B725016FFF5FCA92351FC463B2B" author="Poltarukha, O. P." box="[794,970,1352,1374]" pageId="1" pageNumber="202" pagination="29 - 42" refId="ref5482" refString="Poltarukha, O. P. (2012) On the Deep-sea barnacle (Cirripedia, Thoracica) fauna of the Western Part of Indian Ocean. Bulletin of Moscow Society of Naturalists, 117, 29 - 42. [in Russian]" type="journal article" year="2012">Poltarukha 2012</bibRefCitation>
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: 37
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</treatmentCitation>
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–38, fig. 11.—
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<treatmentCitation id="2E1D10925016FFF5FBCC2351FF6D3B09" ID-DOI="http://doi.org/10.5281/zenodo.5114821" ID-Zenodo-Dep="5114821" author="Shalaeva, K. & Boxshall, G." httpUri="http://treatment.plazi.org/id/03B9EA727F1BFF88FF7DF94B3C16FE95" page="45" pageId="1" pageNumber="202" year="2014">
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<bibRefCitation id="CB2D4B725016FFF5FBCC2351FA153B2B" author="Shalaeva, K. & Boxshall, G." box="[1151,1433,1352,1374]" pageId="1" pageNumber="202" pagination="1 - 63" refId="ref5689" refString="Shalaeva, K. & Boxshall, G. (2014) An illustrated catalogue of the scalpellid barnacles (Crustacea: Cirripedia: Scalpellidae) collected during the HMS " Challenger " expedition and deposited in the Natural History Museum, London. Zootaxa, 3804 (1), 1 - 63. https: // doi. org / 10.11646 / zootaxa. 3804.1.1" type="journal article" year="2014">Shalaeva & Boxshall 2014</bibRefCitation>
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: 45
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</treatmentCitation>
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–49, figs 33A, 33B.
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</treatmentCitationGroup>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="E7A665085016FFF5FF2423B0FCCB387F" pageId="1" pageNumber="202" type="materials_examined">
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<paragraph id="AF0336835016FFF5FF2423B0FCCB387F" blockId="1.[151,1437,1449,2014]" pageId="1" pageNumber="202">
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<emphasis id="9DC8EA915016FFF5FF2423B0FE083BB7" bold="true" box="[151,388,1449,1474]" pageId="1" pageNumber="202">Material examined.</emphasis>
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JL-Dive114-S01-01,
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<specimenCount id="B9BAFD0A5016FFF5FDCC23B3FC8E3BB7" box="[639,770,1449,1474]" pageId="1" pageNumber="202" type="generic">1 specimen</specimenCount>
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(CL
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<quantity id="68449B665016FFF5FCF623B3FC223BB4" box="[837,942,1450,1474]" metricMagnitude="-2" metricUnit="m" metricValue="3.9899999999999998" pageId="1" pageNumber="202" unit="mm" value="39.9">39.9 mm</quantity>
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, CW
|
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<quantity id="68449B665016FFF5FC4523B3FBEC3BB4" box="[1014,1120,1450,1474]" metricMagnitude="-2" metricUnit="m" metricValue="2.18" pageId="1" pageNumber="202" unit="mm" value="21.8">21.8 mm</quantity>
|
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, PL
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||
<quantity id="68449B665016FFF5FB2823B3FA8A3BB7" box="[1179,1286,1450,1474]" metricMagnitude="-2" metricUnit="m" metricValue="2.03" pageId="1" pageNumber="202" unit="mm" value="20.3">20.3 mm</quantity>
|
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), Station JL- Dive114 (
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<geoCoordinate id="CA8850445016FFF5FEBD23D4FE283B93" box="[270,420,1485,1510]" direction="north" orientation="latitude" pageId="1" pageNumber="202" precision="1" value="10.861517">10°51.691’N</geoCoordinate>
|
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,
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<geoCoordinate id="CA8850445016FFF5FE0423D4FDDB3B93" box="[439,599,1485,1510]" direction="east" orientation="longitude" pageId="1" pageNumber="202" precision="1" value="141.95183">141°57.110’E</geoCoordinate>
|
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), Mariana Trench, depth
|
||
<quantity id="68449B665016FFF5FC2123D4FC743B90" box="[914,1016,1485,1510]" metricMagnitude="3" metricUnit="m" metricValue="5.462" pageId="1" pageNumber="202" unit="m" value="5462.0">5,462 m</quantity>
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||
, collected by manned submersible
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||
<emphasis id="9DC8EA915016FFF5FF2423E8FF75387F" box="[151,249,1521,1546]" italics="true" pageId="1" pageNumber="202">Jiaolong</emphasis>
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||
from onboard R/V
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||
<emphasis id="9DC8EA915016FFF5FE6623E8FD17387F" box="[469,667,1521,1546]" italics="true" pageId="1" pageNumber="202">Xiangyanghong 9</emphasis>
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||
, 0
|
||
<date id="DB0210435016FFF5FD0823EBFCCF387F" box="[699,835,1521,1546]" pageId="1" pageNumber="202" value="2016-06-01">1 June 2016</date>
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||
.
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||
</paragraph>
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||
</subSubSection>
|
||
<subSubSection id="E7A665085016FFF5FF74200CFBF638CB" pageId="1" pageNumber="202" type="diagnosis">
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||
<paragraph id="AF0336835016FFF5FF74200CFBF638CB" blockId="1.[151,1437,1449,2014]" pageId="1" pageNumber="202">
|
||
<emphasis id="9DC8EA915016FFF5FF74200CFECD385B" bold="true" box="[199,321,1557,1582]" pageId="1" pageNumber="202">Diagnosis.</emphasis>
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||
Capitulum leaf-like, with 14 closely packed plates, all plates with umbo apical; tergum triangular; carina arched, with narrow laterals, tectum flat or slightly raised medially with longitudinal bordering ribs and medial V-shaped ribs; inframedian latus triangular; carinolatus as high as wide or wider than high, with two apicobasal ridges, both plates basally contacting below carina; carinolatus significantly longer than inframedian latus; peduncle slightly longer than half length of capitulum, with compactly concentric scales.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="E7A665085016FFF6FF7420D0FE9A3985" lastPageId="2" lastPageNumber="203" pageId="1" pageNumber="202" type="description">
|
||
<paragraph id="AF0336835016FFF5FF7420D0FE55395F" blockId="1.[151,1437,1449,2014]" pageId="1" pageNumber="202">
|
||
<emphasis id="9DC8EA915016FFF5FF7420D0FED53897" bold="true" box="[199,345,1737,1762]" pageId="1" pageNumber="202">Description.</emphasis>
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Capitulum (Fig. 1A–D) entirely white, coated by very thin cuticle with short setae; 14 plates closely packed, growth lines conspicuous. Peduncle slightly longer than half length of capitulum, covered by compactly concentric scales.
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</paragraph>
|
||
<paragraph id="AF0336835016FFF6FF74212FFD633FFD" blockId="1.[151,1437,1449,2014]" lastBlockId="2.[151,1437,151,392]" lastPageId="2" lastPageNumber="203" pageId="1" pageNumber="202">Tergum large, occludent and scutal margins straight, carinal margin arched; apex somewhat curved towards carinal side; surface with distinct medial apico-basal ridge, very low ridge from apex to basilateral margin; umbo apical. Scutum smaller than tergum, quadrangular, with prominent apico-basal ridge; occludent margin very straight, basal margin slightly concave; apex bluntly pointed; umbo apical. Carina arciform, exceeding apex of scutum in height; lateral margin narrow; tectum flat, with longitudinal bordering ribs and medial V-shaped ribs. Upper latus quadrangular; tergal margin nearly straight, scutal margin concave; umbo apical, apex pointed. Rostrum (Fig. 1E) very small, strip-like in exposed part. Rostro-latus trapeziform, scutal margin longer than basal margin; surface with apico-basal ridge; basal margin slightly longer than that of inframedian latus. Inframedian latus triangular, height slightly longer than width; umbo apical, apex pointed. Carino-latus as high as wide; upper latus margin with distinct upper flange; surface with two prominent apico-basal ridges; basal margin obviously longer than that of inframedian latus; umbo apical, apex curved; both plates basally contacting below carina, junction length ~0.27 length of carino-latus (Fig. 1F).</paragraph>
|
||
<paragraph id="AF0336835015FFF6FF2423F6FC643851" blockId="2.[151,1435,1519,1572]" pageId="2" pageNumber="203">
|
||
<emphasis id="9DC8EA915015FFF6FF2423F6FE943871" bold="true" box="[151,280,1519,1541]" pageId="2" pageNumber="203">FIGURE. 1</emphasis>
|
||
<taxonomicName id="68BC4D005015FFF6FE9023F6FCED3870" authority="(Wyville-Thomson, 1873)" baseAuthorityName="Wyville-Thomson" baseAuthorityYear="1873" box="[291,865,1519,1542]" class="Maxillopoda" family="Scalpellidae" genus="Trianguloscalpellum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pedunculata" pageId="2" pageNumber="203" phylum="Arthropoda" rank="species" species="regium">
|
||
<emphasis id="9DC8EA915015FFF6FE9023F6FDCA3873" box="[291,582,1519,1542]" italics="true" pageId="2" pageNumber="203">Trianguloscalpellum regium</emphasis>
|
||
(
|
||
<bibRefCitation id="CB2D4B725015FFF6FDEB23F6FCD63870" author="Wyville-Thomson, C." box="[600,858,1519,1541]" pageId="2" pageNumber="203" pagination="347 - 349" refId="ref5756" refString="Wyville-Thomson, C. (1873) Notes from the " Challenger ". Nature, 8, 347 - 349. https: // doi. org / 10.1038 / 008347 a 0" type="journal article" year="1873">Wyville-Thomson, 1873</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. A, B, lateral views of specimen; C, carinal view of specimen; D, rostral view of specimen; E, Rostrum; F, carinal view of carino-latera.
|
||
</paragraph>
|
||
<paragraph id="AF0336835015FFF6FF742052FBDC3949" blockId="2.[151,1436,1611,2032]" pageId="2" pageNumber="203">Soft body without filamentary processes and penis (Fig. 2A). Cirrus I (Fig. 2B) separated from, and shorter than, remaining cirri; inner ramus broad, 11 segmented, medial segments broader than others; outer ramus long, 16 segmented. Cirrus II (Fig. 2C) much longer than cirrus I, rami of nearly equal length, inner ramus 28 segmented, outer ramus 30 segmented. Cirrus VI (Fig. 2D) longest, both rami nearly equal length, inner ramus 37 segmented, outer ramus 41 segmented, intermediate segments of rami with 3–4 (mostly 4) pairs of long setae on ventral margin (2–6 short seta among each pair of long setae) and 1 pair of long setae on dorsal margin (sometimes with 1–2 short seta). Caudal appendage (Fig. 2E) long, 5 segmented, reaching to end of protopodite.</paragraph>
|
||
<paragraph id="AF0336835015FFF6FF63215EFE9A3985" blockId="2.[151,1436,1611,2032]" pageId="2" pageNumber="203">Labrum swollen (Fig. 3A), cutting edge with flat ridge; mandibular palp narrow (Fig. 3B), with long setae along apical margin. Mandible with 3 large teeth excluding inferior angle (Fig. 3C), apex of inferior angle rough, without setae, inferior margin with long spinules (Fig. 3D). Maxillule slightly curved (Fig. 3E), cutting edge nearly straight, with many long, spinous setae. Maxilla with dense long setae along margins (Fig. 3F), anterior margin with notch.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="E7A665085014FFF7FF74268EFE5B3B50" pageId="3" pageNumber="204" type="distribution">
|
||
<paragraph id="AF0336835014FFF7FF74268EFD5D3F10" blockId="3.[151,1436,151,357]" pageId="3" pageNumber="204">
|
||
<emphasis id="9DC8EA915014FFF7FF74268EFEEC3EC5" bold="true" box="[199,352,151,176]" pageId="3" pageNumber="204">Distribution.</emphasis>
|
||
<taxonomicName id="68BC4D005014FFF7FEDB268EFC593EC5" authority="(Wyville-Thomson, 1873)" baseAuthorityName="Wyville-Thomson" baseAuthorityYear="1873" box="[360,981,151,176]" class="Maxillopoda" family="Scalpellidae" genus="Trianguloscalpellum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pedunculata" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="regium">
|
||
<emphasis id="9DC8EA915014FFF7FEDB268EFD2B3EC5" box="[360,679,151,176]" italics="true" pageId="3" pageNumber="204">Trianguloscalpellum regium</emphasis>
|
||
(
|
||
<bibRefCitation id="CB2D4B725014FFF7FD04268EFC403EC5" author="Wyville-Thomson, C." box="[695,972,151,176]" pageId="3" pageNumber="204" pagination="347 - 349" refId="ref5756" refString="Wyville-Thomson, C. (1873) Notes from the " Challenger ". Nature, 8, 347 - 349. https: // doi. org / 10.1038 / 008347 a 0" type="journal article" year="1873">Wyville-Thomson, 1873</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
is a cosmopolitan species distributed in the Atlantic, Indian and Pacific Oceans (
|
||
<bibRefCitation id="CB2D4B725014FFF7FDD226A5FD763EA1" author="Zevina, G. B." box="[609,762,188,213]" pageId="3" pageNumber="204" refId="ref6098" refString="Zevina, G. B. (1981) Cirripede crustaceans of the suborder Lepadomorpha (Cirripedia, Thoracica) of the World Ocean. Part 1. Family Scalpellidae. Opredeliteli po Faune SSSR, Leningrad, 398 pp. [in Russian]" type="book" year="1981">Zevina, 1981</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="CB2D4B725014FFF7FCB426A4FBB13EA1" author="Foster, B. A. & Buckeridge, J. S." box="[775,1085,188,213]" pageId="3" pageNumber="204" pagination="345 - 382" refId="ref4728" refString="Foster, B. A. & Buckeridge, J. S. (1994) Barnacles (Cirripedia, Thoracica) of seas off Reunion Island and the East Indies. Bulletin du Museum national d'histoire naturelle. Section A, Zoologie, biologie et ecologie animales, 16, 345 - 382." type="journal article" year="1994">Foster & Buckeridge, 1994</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="CB2D4B725014FFF7FBF926A5FA9C3EA1" author="Chan, B. K. & Prabowo, R. E. & Lee, K. S." box="[1098,1296,188,213]" pageId="3" pageNumber="204" pagination="1 - 47" refId="ref4418" refString="Chan, B. K., Prabowo, R. E. & Lee, K. S. (2010) North West Pacific deep-sea barnacles (Cirripedia, Thoracica) collected by the TAIWAN expeditions, with descriptions of two new species. Zootaxa, 2405, 1 - 47. https: // doi. org / 10.11646 / zootaxa. 2405.1.1" type="journal article" year="2010">
|
||
Chan
|
||
<emphasis id="9DC8EA915014FFF7FB3F26A4FB323EA0" box="[1164,1214,188,213]" italics="true" pageId="3" pageNumber="204">et al</emphasis>
|
||
., 2010
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="CB2D4B725014FFF7FAAF26A5FF5F3E8D" author="Poltarukha, O. P." pageId="3" pageNumber="204" pagination="29 - 42" refId="ref5482" refString="Poltarukha, O. P. (2012) On the Deep-sea barnacle (Cirripedia, Thoracica) fauna of the Western Part of Indian Ocean. Bulletin of Moscow Society of Naturalists, 117, 29 - 42. [in Russian]" type="journal article" year="2012">Poltarukha, 2012</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="CB2D4B725014FFF7FF6C26F9FE5E3E8D" author="Jones, D. S. & Hosie, A. M." box="[223,466,223,248]" pageId="3" pageNumber="204" pagination="241 - 311" refId="ref5117" refString="Jones, D. S. & Hosie, A. M. (2016) A checklist of the barnacles (Cirripedia: Thoracica) of Singapore and neighbouring waters. Raffles Bulletin of Zoology, 34, 241 - 311." type="journal article" year="2016">Jones & Hosie, 2016</bibRefCitation>
|
||
). However,
|
||
<bibRefCitation id="CB2D4B725014FFF7FDE926F9FD733E8D" author="Young, P. S." box="[602,767,223,248]" pageId="3" pageNumber="204" pagination="1 - 54" refId="ref5886" refString="Young, P. S. (1998 b) The Cirripedia (Crustacea) collected by the " Fisheries Steamer Meteor " in the eastern Atlantic. Arquivos do Museu Nacional Rio de Janeiro, 58, 1 - 54." type="journal article" year="1998">Young (1998b</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="CB2D4B725014FFF7FCBF26C6FCDF3E8D" author="Young, P. S." box="[780,851,223,248]" pageId="3" pageNumber="204" pagination="705 - 756" refId="ref6022" refString="Young, P. S. (2001 b) Deep-sea Cirripedia Thoracica (Crustacea) from the northeastern Atlantic collected by French expeditions. Zoosystema, 23, 705 - 756." type="journal article" year="2001">2001b</bibRefCitation>
|
||
) considered it to be distributed only in the Atlantic Ocean. As noted by
|
||
<bibRefCitation id="CB2D4B725014FFF7FECA271CFDA83F68" author="Young, P. S." box="[377,548,260,285]" pageId="3" pageNumber="204" pagination="1 - 54" refId="ref5886" refString="Young, P. S. (1998 b) The Cirripedia (Crustacea) collected by the " Fisheries Steamer Meteor " in the eastern Atlantic. Arquivos do Museu Nacional Rio de Janeiro, 58, 1 - 54." type="journal article" year="1998">Young (1998b)</bibRefCitation>
|
||
, it is difficult to determine the geographic distribution patterns of
|
||
<taxonomicName id="68BC4D005014FFF7FAB7271CFAE03F69" baseAuthorityName="Wyville-Thomson" baseAuthorityYear="1873" box="[1284,1388,261,284]" class="Maxillopoda" family="Scalpellidae" genus="Trianguloscalpellum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pedunculata" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="regium">
|
||
<emphasis id="9DC8EA915014FFF7FAB7271CFAE03F69" box="[1284,1388,261,284]" italics="true" pageId="3" pageNumber="204">T. regium</emphasis>
|
||
</taxonomicName>
|
||
due to taxonomic confusion, and the present specimen does exhibit some features that differ to those reported by
|
||
<bibRefCitation id="CB2D4B725014FFF7FAD2273EFEAD3F11" author="Chan, B. K. & Prabowo, R. E. & Lee, K. S." pageId="3" pageNumber="204" pagination="1 - 47" refId="ref4418" refString="Chan, B. K., Prabowo, R. E. & Lee, K. S. (2010) North West Pacific deep-sea barnacles (Cirripedia, Thoracica) collected by the TAIWAN expeditions, with descriptions of two new species. Zootaxa, 2405, 1 - 47. https: // doi. org / 10.11646 / zootaxa. 2405.1.1" type="journal article" year="2010">
|
||
Chan
|
||
<emphasis id="9DC8EA915014FFF7FF242754FF443F10" box="[151,200,332,357]" italics="true" pageId="3" pageNumber="204">et al</emphasis>
|
||
. (2010)
|
||
</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="CB2D4B725014FFF7FEE42755FDAE3F11" author="Poltarukha, O. P." box="[343,546,332,357]" pageId="3" pageNumber="204" pagination="29 - 42" refId="ref5482" refString="Poltarukha, O. P. (2012) On the Deep-sea barnacle (Cirripedia, Thoracica) fauna of the Western Part of Indian Ocean. Bulletin of Moscow Society of Naturalists, 117, 29 - 42. [in Russian]" type="journal article" year="2012">Poltarukha (2012)</bibRefCitation>
|
||
(see Remarks).
|
||
</paragraph>
|
||
<paragraph id="AF0336835014FFF7FF2422E9FE5B3B50" blockId="3.[151,1435,1264,1317]" pageId="3" pageNumber="204">
|
||
<emphasis id="9DC8EA915014FFF7FF2422E9FE983B70" bold="true" box="[151,276,1264,1286]" pageId="3" pageNumber="204">FIGURE. 2</emphasis>
|
||
<taxonomicName id="68BC4D005014FFF7FEAF22E9FCDE3B73" authority="(Wyville-Thomson, 1873)" baseAuthorityName="Wyville-Thomson" baseAuthorityYear="1873" box="[284,850,1264,1286]" class="Maxillopoda" family="Scalpellidae" genus="Trianguloscalpellum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pedunculata" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="regium">
|
||
<emphasis id="9DC8EA915014FFF7FEAF22E9FDB13B73" box="[284,573,1264,1286]" italics="true" pageId="3" pageNumber="204">Trianguloscalpellum regium</emphasis>
|
||
(
|
||
<bibRefCitation id="CB2D4B725014FFF7FDF822E9FCC63B73" author="Wyville-Thomson, C." box="[587,842,1264,1286]" pageId="3" pageNumber="204" pagination="347 - 349" refId="ref5756" refString="Wyville-Thomson, C. (1873) Notes from the " Challenger ". Nature, 8, 347 - 349. https: // doi. org / 10.1038 / 008347 a 0" type="journal article" year="1873">Wyville-Thomson, 1873</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. A, lateral view of soft body; B, cirrus I; C, cirrus II; D, cirrus VI; E, caudal appendage.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="E7A665085014FFF0FF742348FD2E386B" lastPageId="4" lastPageNumber="205" pageId="3" pageNumber="204" type="discussion">
|
||
<paragraph id="AF0336835014FFF0FF742348FC363C6D" blockId="3.[151,1437,1361,2035]" lastBlockId="4.[151,1436,151,536]" lastPageId="4" lastPageNumber="205" pageId="3" pageNumber="204">
|
||
<emphasis id="9DC8EA915014FFF7FF742348FEB73B1F" bold="true" box="[199,315,1361,1386]" pageId="3" pageNumber="204">Remarks.</emphasis>
|
||
Based on cladistic analysis,
|
||
<bibRefCitation id="CB2D4B725014FFF7FD292348FCAA3B1F" author="Gale, A. S." box="[666,806,1361,1386]" pageId="3" pageNumber="204" pagination="266 - 304" refId="ref4783" refString="Gale, A. S. (2016) Phylogeny of the deep-sea cirripede family Scalpellidae (Crustacea, Thoracica) based on shell capitular plate morphology. Zoological Journal of the Linnean Society, 176, 266 - 304. https: // doi. org / 10.1111 / zoj. 12321" type="journal article" year="2016">Gale (2016)</bibRefCitation>
|
||
proposed a new classification system for the family
|
||
<taxonomicName id="68BC4D005014FFF7FF24236CFE353BFB" authority="Pilsbry, 1907" authorityName="Pilsbry" authorityYear="1907" box="[151,441,1397,1422]" class="Maxillopoda" family="Scalpellidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pedunculata" pageId="3" pageNumber="204" phylum="Arthropoda" rank="family">
|
||
Scalpellidae
|
||
<bibRefCitation id="CB2D4B725014FFF7FE95236CFE353BFB" author="Pilsbry, H. A." box="[294,441,1397,1422]" pageId="3" pageNumber="204" pagination="1 - 122" refId="ref5429" refString="Pilsbry, H. A. (1907) The Barnacles (Cirripedia) contained in the collections of the U. S. National Museum. Bulletin of the United States National Museum, 60, 1 - 122. https: // doi. org / 10.5479 / si. 03629236.60.1" type="journal article" year="1907">Pilsbry, 1907</bibRefCitation>
|
||
</taxonomicName>
|
||
and established a new genus,
|
||
<emphasis id="9DC8EA915014FFF7FCBA236CFC483BFB" box="[777,964,1397,1422]" italics="true" pageId="3" pageNumber="204">Regioscalpellum</emphasis>
|
||
, to embrace
|
||
<taxonomicName id="68BC4D005014FFF7FBE0236CFF5E3BC7" authority="Thomson, 1873" authorityName="Wyville-Thomson" authorityYear="1873" class="Maxillopoda" family="Scalpellidae" genus="Scalpellum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pedunculata" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="regium">
|
||
<emphasis id="9DC8EA915014FFF7FBE0236CFAA83BFB" box="[1107,1316,1397,1422]" italics="true" pageId="3" pageNumber="204">Scalpellum regium</emphasis>
|
||
<bibRefCitation id="CB2D4B725014FFF7FA98236CFF5E3BC7" author="Wyville-Thomson, C." pageId="3" pageNumber="204" pagination="347 - 349" refId="ref5756" refString="Wyville-Thomson, C. (1873) Notes from the " Challenger ". Nature, 8, 347 - 349. https: // doi. org / 10.1038 / 008347 a 0" type="journal article" year="1873">Thomson, 1873</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="68BC4D005014FFF7FF512383FE7C3BC7" authority="Hoek, 1883" authorityName="Hoek" authorityYear="1883" box="[226,496,1433,1458]" class="Maxillopoda" family="Scalpellidae" genus="Scalpellum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pedunculata" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="darwini">
|
||
<emphasis id="9DC8EA915014FFF7FF512383FED63BC7" box="[226,346,1433,1458]" italics="true" pageId="3" pageNumber="204">S. darwini</emphasis>
|
||
<bibRefCitation id="CB2D4B725014FFF7FED62380FE7C3BC7" author="Hoek, P. P. C." box="[357,496,1433,1458]" pageId="3" pageNumber="204" pagination="1 - 169" refId="ref4892" refString="Hoek, P. P. C. (1883) Report on the Cirripedia collected by H. M. S. " Challenger " during the years 1873 - 1876. Report on the Scientific Results of the Foyage H. M. S. " Challenger ", Zoology, part 25, 8, 1 - 169." type="journal article" year="1883">Hoek, 1883</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="68BC4D005014FFF7FDB32383FD7F3BC7" authority="Hoek, 1883" authorityName="Hoek" authorityYear="1883" box="[512,755,1433,1458]" class="Maxillopoda" family="Poecilasmatidae" genus="Glyptelasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pedunculata" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="gigas">
|
||
<emphasis id="9DC8EA915014FFF7FDB32383FDD13BC7" box="[512,605,1434,1458]" italics="true" pageId="3" pageNumber="204">S. gigas</emphasis>
|
||
<bibRefCitation id="CB2D4B725014FFF7FDDA2380FD7F3BC7" author="Hoek, P. P. C." box="[617,755,1433,1458]" pageId="3" pageNumber="204" pagination="1 - 169" refId="ref4892" refString="Hoek, P. P. C. (1883) Report on the Cirripedia collected by H. M. S. " Challenger " during the years 1873 - 1876. Report on the Scientific Results of the Foyage H. M. S. " Challenger ", Zoology, part 25, 8, 1 - 169." type="journal article" year="1883">Hoek, 1883</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="68BC4D005014FFF7FCB72383FBC63BC7" authority="Hoek, 1883" authorityName="Hoek" authorityYear="1883" box="[772,1098,1433,1458]" class="Maxillopoda" family="Scalpellidae" genus="Scalpellum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pedunculata" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="moluccanum">
|
||
<emphasis id="9DC8EA915014FFF7FCB72383FC3F3BC7" box="[772,947,1433,1458]" italics="true" pageId="3" pageNumber="204">S. moluccanum</emphasis>
|
||
<bibRefCitation id="CB2D4B725014FFF7FC0C2380FBC63BC7" author="Hoek, P. P. C." box="[959,1098,1433,1458]" pageId="3" pageNumber="204" pagination="1 - 169" refId="ref4892" refString="Hoek, P. P. C. (1883) Report on the Cirripedia collected by H. M. S. " Challenger " during the years 1873 - 1876. Report on the Scientific Results of the Foyage H. M. S. " Challenger ", Zoology, part 25, 8, 1 - 169." type="journal article" year="1883">Hoek, 1883</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="68BC4D005014FFF7FBE92380FAB63BC7" box="[1114,1338,1433,1458]" class="Maxillopoda" family="Pollicipedidae" genus="Pollicipes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pedunculata" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="maximus">
|
||
<emphasis id="9DC8EA915014FFF7FBE92380FAB63BC7" box="[1114,1338,1433,1458]" italics="true" pageId="3" pageNumber="204">Pollicipes maximus</emphasis>
|
||
</taxonomicName>
|
||
J. de C Sowerby, 1829, and
|
||
<taxonomicName id="68BC4D005014FFF7FE3223A4FCDB3BA3" authority="Withers, 1953" authorityName="Withers" authorityYear="1953" box="[385,855,1469,1494]" class="Maxillopoda" family="Scalpellidae" genus="Arcoscalpellum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pedunculata" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="sanchezae">
|
||
<emphasis id="9DC8EA915014FFF7FE3223A4FD203BA3" box="[385,684,1469,1494]" italics="true" pageId="3" pageNumber="204">Arcoscalpellum sanchezae</emphasis>
|
||
Withers, 1953
|
||
</taxonomicName>
|
||
; however, the taxonomic status of some remaining species assigned to
|
||
<taxonomicName id="68BC4D005014FFF7FEC623F8FDD13B8F" authorityName="Zevina" authorityYear="1978" box="[373,605,1505,1530]" class="Maxillopoda" family="Scalpellidae" genus="Trianguloscalpellum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pedunculata" pageId="3" pageNumber="204" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="9DC8EA915014FFF7FEC623F8FDD13B8F" box="[373,605,1505,1530]" italics="true" pageId="3" pageNumber="204">Trianguloscalpellum</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="CB2D4B725014FFF7FD3B23F8FCAE3B8F" author="Zevina, G. B." box="[648,802,1505,1530]" pageId="3" pageNumber="204" pagination="1343 - 1352" refId="ref6055" refString="Zevina, G. B. (1978) A new system of the family Scalpellidae Pilsbry (Cirripedia, Thoracica). 2. Subfamilies Arcoscalpellinae and Meroscalpellinae. Zoologicheskii Zhurnal, 57, 1343 - 1352. [in Russian]" type="journal article" year="1978">Zevina (1978</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="CB2D4B725014FFF7FC9D23FBFCEB3B8F" author="Zevina, G. B." box="[814,871,1506,1530]" pageId="3" pageNumber="204" refId="ref6098" refString="Zevina, G. B. (1981) Cirripede crustaceans of the suborder Lepadomorpha (Cirripedia, Thoracica) of the World Ocean. Part 1. Family Scalpellidae. Opredeliteli po Faune SSSR, Leningrad, 398 pp. [in Russian]" type="book" year="1981">1981</bibRefCitation>
|
||
) is unclear. The present study only focuses on the morphological characters of
|
||
<taxonomicName id="68BC4D005014FFF7FE6E201CFBC7386B" authority="(Wyville-Thomson, 1873)" baseAuthorityName="Wyville-Thomson" baseAuthorityYear="1873" box="[477,1099,1541,1566]" class="Maxillopoda" family="Scalpellidae" genus="Trianguloscalpellum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pedunculata" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="regium">
|
||
<emphasis id="9DC8EA915014FFF7FE6E201CFC90386B" box="[477,796,1541,1566]" italics="true" pageId="3" pageNumber="204">Trianguloscalpellum regium</emphasis>
|
||
(
|
||
<bibRefCitation id="CB2D4B725014FFF7FC9F201CFBCF386B" author="Wyville-Thomson, C." box="[812,1091,1541,1566]" pageId="3" pageNumber="204" pagination="347 - 349" refId="ref5756" refString="Wyville-Thomson, C. (1873) Notes from the " Challenger ". Nature, 8, 347 - 349. https: // doi. org / 10.1038 / 008347 a 0" type="journal article" year="1873">Wyville-Thomson, 1873</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and we use the classification system of
|
||
<bibRefCitation id="CB2D4B725014FFF7FEBD2030FE3E3837" author="Zevina, G. B." box="[270,434,1577,1602]" pageId="3" pageNumber="204" pagination="1343 - 1352" refId="ref6055" refString="Zevina, G. B. (1978) A new system of the family Scalpellidae Pilsbry (Cirripedia, Thoracica). 2. Subfamilies Arcoscalpellinae and Meroscalpellinae. Zoologicheskii Zhurnal, 57, 1343 - 1352. [in Russian]" type="journal article" year="1978">Zevina (1978)</bibRefCitation>
|
||
, which is convenient for comparison between
|
||
<taxonomicName id="68BC4D005014FFF7FC782032FBBB3837" baseAuthorityName="Wyville-Thomson" baseAuthorityYear="1873" box="[971,1079,1579,1602]" class="Maxillopoda" family="Scalpellidae" genus="Trianguloscalpellum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pedunculata" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="regium">
|
||
<emphasis id="9DC8EA915014FFF7FC782032FBBB3837" box="[971,1079,1579,1602]" italics="true" pageId="3" pageNumber="204">T. regium</emphasis>
|
||
</taxonomicName>
|
||
and its closely related species. The plates of the present specimen agree well with the detailed re-description of
|
||
<bibRefCitation id="CB2D4B725014FFF7FB952056FB5C3813" author="Young, P. S." box="[1062,1232,1613,1639]" pageId="3" pageNumber="204" pagination="1 - 54" refId="ref5886" refString="Young, P. S. (1998 b) The Cirripedia (Crustacea) collected by the " Fisheries Steamer Meteor " in the eastern Atlantic. Arquivos do Museu Nacional Rio de Janeiro, 58, 1 - 54." type="journal article" year="1998">Young (1998b)</bibRefCitation>
|
||
based on the
|
||
<typeStatus id="700788215014FFF7FADF2056FA103812" box="[1388,1436,1615,1639]" pageId="3" pageNumber="204">type</typeStatus>
|
||
specimen of
|
||
<emphasis id="9DC8EA915014FFF7FE962068FDE438FF" box="[293,616,1649,1674]" italics="true" pageId="3" pageNumber="204">
|
||
<taxonomicName id="68BC4D005014FFF7FE962068FDE838FF" baseAuthorityName="Wyville-Thomson" baseAuthorityYear="1873" box="[293,612,1649,1674]" class="Maxillopoda" family="Scalpellidae" genus="Trianguloscalpellum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pedunculata" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="regium">Trianguloscalpellum regium</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and with the photographs in
|
||
<bibRefCitation id="CB2D4B725014FFF7FC1F2068FB6538FF" author="Shalaeva, K. & Boxshall, G." box="[940,1257,1649,1674]" pageId="3" pageNumber="204" pagination="1 - 63" refId="ref5689" refString="Shalaeva, K. & Boxshall, G. (2014) An illustrated catalogue of the scalpellid barnacles (Crustacea: Cirripedia: Scalpellidae) collected during the HMS " Challenger " expedition and deposited in the Natural History Museum, London. Zootaxa, 3804 (1), 1 - 63. https: // doi. org / 10.11646 / zootaxa. 3804.1.1" type="journal article" year="2014">Shalaeva & Boxshall (2014)</bibRefCitation>
|
||
also taken from the
|
||
<typeStatus id="700788215014FFF7FF72208EFF7D38DA" box="[193,241,1687,1711]" pageId="3" pageNumber="204">type</typeStatus>
|
||
specimen. The present specimen exhibits all the morphological differences between
|
||
<taxonomicName id="68BC4D005014FFF7FB01208EFAA138DB" baseAuthorityName="Hoek" baseAuthorityYear="1883" box="[1202,1325,1685,1710]" class="Maxillopoda" family="Scalpellidae" genus="Trianguloscalpellum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pedunculata" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="darwinii">
|
||
<emphasis id="9DC8EA915014FFF7FB01208EFAA138DB" box="[1202,1325,1685,1710]" italics="true" pageId="3" pageNumber="204">T. darwinii</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="68BC4D005014FFF7FA8F208EFA1838DB" authorityName="Hoek" authorityYear="1883" box="[1340,1428,1687,1710]" class="Maxillopoda" family="Poecilasmatidae" genus="Glyptelasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pedunculata" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="gigas">
|
||
<emphasis id="9DC8EA915014FFF7FA8F208EFA1838DB" box="[1340,1428,1687,1710]" italics="true" pageId="3" pageNumber="204">T. gigas</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="68BC4D005014FFF7FF2420A2FEB238A7" box="[151,318,1721,1746]" class="Maxillopoda" family="Scalpellidae" genus="Trianguloscalpellum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pedunculata" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="mollucanum">
|
||
<emphasis id="9DC8EA915014FFF7FF2420A2FEB238A7" box="[151,318,1721,1746]" italics="true" pageId="3" pageNumber="204">T. mollucanum</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="68BC4D005014FFF7FEC420A0FE4438A7" baseAuthorityName="Hoek" baseAuthorityYear="1883" box="[375,456,1721,1746]" class="Maxillopoda" family="Scalpellidae" genus="Trianguloscalpellum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pedunculata" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="ovale">
|
||
<emphasis id="9DC8EA915014FFF7FEC420A0FE4438A7" box="[375,456,1721,1746]" italics="true" pageId="3" pageNumber="204">T.ovale</emphasis>
|
||
</taxonomicName>
|
||
proposed by
|
||
<bibRefCitation id="CB2D4B725014FFF7FDD720A3FC9D38A7" author="Young, P. S." box="[612,785,1721,1746]" pageId="3" pageNumber="204" pagination="1 - 54" refId="ref5886" refString="Young, P. S. (1998 b) The Cirripedia (Crustacea) collected by the " Fisheries Steamer Meteor " in the eastern Atlantic. Arquivos do Museu Nacional Rio de Janeiro, 58, 1 - 54." type="journal article" year="1998">Young (1998b)</bibRefCitation>
|
||
. For example, it differs from
|
||
<taxonomicName id="68BC4D005014FFF7FBD220A2FB3738A7" authorityName="Hoek" authorityYear="1883" box="[1121,1211,1723,1746]" class="Maxillopoda" family="Poecilasmatidae" genus="Glyptelasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pedunculata" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="gigas">
|
||
<emphasis id="9DC8EA915014FFF7FBD220A2FB3738A7" box="[1121,1211,1723,1746]" italics="true" pageId="3" pageNumber="204">T. gigas</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="68BC4D005014FFF7FB4720A2FAC238A7" baseAuthorityName="Hoek" baseAuthorityYear="1883" box="[1268,1358,1721,1746]" class="Maxillopoda" family="Scalpellidae" genus="Trianguloscalpellum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pedunculata" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="ovale">
|
||
<emphasis id="9DC8EA915014FFF7FB4720A2FAC238A7" box="[1268,1358,1721,1746]" italics="true" pageId="3" pageNumber="204">T. ovale</emphasis>
|
||
</taxonomicName>
|
||
by the carinolatus (only as high as wide vs. significantly higher than wide), from
|
||
<taxonomicName id="68BC4D005014FFF7FC5D20C6FBE73883" baseAuthorityName="Hoek" baseAuthorityYear="1883" box="[1006,1131,1757,1782]" class="Maxillopoda" family="Scalpellidae" genus="Trianguloscalpellum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pedunculata" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="darwinii">
|
||
<emphasis id="9DC8EA915014FFF7FC5D20C6FBE73883" box="[1006,1131,1757,1782]" italics="true" pageId="3" pageNumber="204">T. darwinii</emphasis>
|
||
</taxonomicName>
|
||
by the shape of the carina (tectum flatter and lateral margin narrower vs. tectum pronounced and lateral margin broad), and is immediately distinguishable from
|
||
<taxonomicName id="68BC4D005014FFF7FE34213EFDA2394B" box="[391,558,1829,1854]" class="Maxillopoda" family="Scalpellidae" genus="Trianguloscalpellum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pedunculata" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="mollucanum">
|
||
<emphasis id="9DC8EA915014FFF7FE34213EFDA2394B" box="[391,558,1829,1854]" italics="true" pageId="3" pageNumber="204">T. mollucanum</emphasis>
|
||
</taxonomicName>
|
||
by features of the carinolatus and inframedian latus (carinolatus significantly wider than inframedian latus vs. carinolatus as wide as inframedian latus).
|
||
<bibRefCitation id="CB2D4B725014FFF7FBB82153FB683917" author="Chan, B. K. & Prabowo, R. E. & Lee, K. S." box="[1035,1252,1866,1891]" pageId="3" pageNumber="204" pagination="1 - 47" refId="ref4418" refString="Chan, B. K., Prabowo, R. E. & Lee, K. S. (2010) North West Pacific deep-sea barnacles (Cirripedia, Thoracica) collected by the TAIWAN expeditions, with descriptions of two new species. Zootaxa, 2405, 1 - 47. https: // doi. org / 10.11646 / zootaxa. 2405.1.1" type="journal article" year="2010">
|
||
Chan
|
||
<emphasis id="9DC8EA915014FFF7FBE22152FB0A3916" box="[1105,1158,1866,1891]" italics="true" pageId="3" pageNumber="204">et al</emphasis>
|
||
. (2010)
|
||
</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="CB2D4B725014FFF7FA902153FF6D39F3" author="Poltarukha, O. P." pageId="3" pageNumber="204" pagination="29 - 42" refId="ref5482" refString="Poltarukha, O. P. (2012) On the Deep-sea barnacle (Cirripedia, Thoracica) fauna of the Western Part of Indian Ocean. Bulletin of Moscow Society of Naturalists, 117, 29 - 42. [in Russian]" type="journal article" year="2012">Poltarukha (2012)</bibRefCitation>
|
||
reported
|
||
<taxonomicName id="68BC4D005014FFF7FEE02176FE3339F3" baseAuthorityName="Wyville-Thomson" baseAuthorityYear="1873" box="[339,447,1903,1926]" class="Maxillopoda" family="Scalpellidae" genus="Trianguloscalpellum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pedunculata" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="regium">
|
||
<emphasis id="9DC8EA915014FFF7FEE02176FE3339F3" box="[339,447,1903,1926]" italics="true" pageId="3" pageNumber="204">T. regium</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="D7AB76135014FFF7FDB92174FDD039F3" box="[522,604,1901,1926]" name="Taiwan" pageId="3" pageNumber="204">Taiwan</collectingCountry>
|
||
and the Indian Ocean, respectively; however, their specimens show the carinolatus higher than wide, a morphological character presented by
|
||
<taxonomicName id="68BC4D005014FFF7FC1F218AFB8A39DF" authorityName="Hoek" authorityYear="1883" box="[940,1030,1939,1962]" class="Maxillopoda" family="Poecilasmatidae" genus="Glyptelasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pedunculata" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="gigas">
|
||
<emphasis id="9DC8EA915014FFF7FC1F218AFB8A39DF" box="[940,1030,1939,1962]" italics="true" pageId="3" pageNumber="204">T. gigas</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="68BC4D005014FFF7FB8D218AFB1A39DE" baseAuthorityName="Hoek" baseAuthorityYear="1883" box="[1086,1174,1938,1963]" class="Maxillopoda" family="Scalpellidae" genus="Trianguloscalpellum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pedunculata" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="ovale">
|
||
<emphasis id="9DC8EA915014FFF7FB8D218AFB1A39DE" box="[1086,1174,1938,1963]" italics="true" pageId="3" pageNumber="204">T. ovale</emphasis>
|
||
</taxonomicName>
|
||
. The present specimen also differs from their specimens in the appendages of the soft body. For example, Cirrus I has more segments than that of
|
||
<bibRefCitation id="CB2D4B725014FFF7FF5621C3FE3F3987" author="Chan, B. K. & Prabowo, R. E. & Lee, K. S." box="[229,435,2010,2035]" pageId="3" pageNumber="204" pagination="1 - 47" refId="ref4418" refString="Chan, B. K., Prabowo, R. E. & Lee, K. S. (2010) North West Pacific deep-sea barnacles (Cirripedia, Thoracica) collected by the TAIWAN expeditions, with descriptions of two new species. Zootaxa, 2405, 1 - 47. https: // doi. org / 10.11646 / zootaxa. 2405.1.1" type="journal article" year="2010">
|
||
Chan
|
||
<emphasis id="9DC8EA915014FFF7FE9421C2FED43986" box="[295,344,2010,2035]" italics="true" pageId="3" pageNumber="204">et al</emphasis>
|
||
. (2010)
|
||
</bibRefCitation>
|
||
; the caudal appendage of the present specimen is longer (5-segmented) and reaches to the end of the second segment of the protopodite, which is short (4-segmented) and falls short of the end of the first segment of the protopodite in that of
|
||
<bibRefCitation id="CB2D4B725013FFF0FDE626A5FCA23EA1" author="Chan, B. K. & Prabowo, R. E. & Lee, K. S." box="[597,814,188,213]" pageId="4" pageNumber="205" pagination="1 - 47" refId="ref4418" refString="Chan, B. K., Prabowo, R. E. & Lee, K. S. (2010) North West Pacific deep-sea barnacles (Cirripedia, Thoracica) collected by the TAIWAN expeditions, with descriptions of two new species. Zootaxa, 2405, 1 - 47. https: // doi. org / 10.11646 / zootaxa. 2405.1.1" type="journal article" year="2010">
|
||
Chan
|
||
<emphasis id="9DC8EA915013FFF0FD2826A4FD5D3EA0" box="[667,721,188,213]" italics="true" pageId="4" pageNumber="205">et al</emphasis>
|
||
. (2010)
|
||
</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="CB2D4B725013FFF0FCDF26A5FBB33EA1" author="Poltarukha, O. P." box="[876,1087,188,213]" pageId="4" pageNumber="205" pagination="29 - 42" refId="ref5482" refString="Poltarukha, O. P. (2012) On the Deep-sea barnacle (Cirripedia, Thoracica) fauna of the Western Part of Indian Ocean. Bulletin of Moscow Society of Naturalists, 117, 29 - 42. [in Russian]" type="journal article" year="2012">Poltarukha (2012)</bibRefCitation>
|
||
; and the inferior angle of the mandible of the present specimen has five small spines that are absent in that of
|
||
<bibRefCitation id="CB2D4B725013FFF0FB9826C6FB713E8D" author="Chan, B. K. & Prabowo, R. E. & Lee, K. S." box="[1067,1277,223,248]" pageId="4" pageNumber="205" pagination="1 - 47" refId="ref4418" refString="Chan, B. K., Prabowo, R. E. & Lee, K. S. (2010) North West Pacific deep-sea barnacles (Cirripedia, Thoracica) collected by the TAIWAN expeditions, with descriptions of two new species. Zootaxa, 2405, 1 - 47. https: // doi. org / 10.11646 / zootaxa. 2405.1.1" type="journal article" year="2010">
|
||
Chan
|
||
<emphasis id="9DC8EA915013FFF0FBDD26F8FB2C3E8D" box="[1134,1184,223,248]" italics="true" pageId="4" pageNumber="205">et al</emphasis>
|
||
. (2010)
|
||
</bibRefCitation>
|
||
. Furthermore, the similarities of the nucleotide sequences of the genes COI from the present specimen is less than 90% in comparison to that provided by
|
||
<bibRefCitation id="CB2D4B725013FFF0FE49273EFD3C3F35" author="Lin, H. C. & Hoeg, J. T. & Yusa, Y. & Chan, B. K." box="[506,688,295,320]" pageId="4" pageNumber="205" pagination="1 - 11" refId="ref5273" refString="Lin, H. C., Hoeg, J. T., Yusa, Y. & Chan, B. K. (2015) The origins and evolution of dwarf males and habitat use in thoracican barnacles. Molecular phylogenetics and evolution, 91, 1 - 11. https: // doi. org / 10.1016 / j. ympev. 2015.04.026" type="journal article" year="2015">
|
||
Lin
|
||
<emphasis id="9DC8EA915013FFF0FD942730FDDB3F35" box="[551,599,295,320]" italics="true" pageId="4" pageNumber="205">et al</emphasis>
|
||
. (2015)
|
||
</bibRefCitation>
|
||
(genbank number:
|
||
<accessionNumber id="B0EFAB605013FFF0FC392731FB8B3F35" box="[906,1031,296,320]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KF484203" pageId="4" pageNumber="205">KF484203</accessionNumber>
|
||
). The taxonomic status of
|
||
<emphasis id="9DC8EA915013FFF0FA9F2730FE213F11" italics="true" pageId="4" pageNumber="205">
|
||
<taxonomicName id="68BC4D005013FFF0FA9F2730FA143F35" baseAuthorityName="Wyville-Thomson" baseAuthorityYear="1873" box="[1324,1432,297,320]" class="Maxillopoda" family="Scalpellidae" genus="Trianguloscalpellum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pedunculata" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="regium">T. regium</taxonomicName>
|
||
,
|
||
<taxonomicName id="68BC4D005013FFF0FF242754FECE3F10" box="[151,322,332,357]" class="Maxillopoda" family="Scalpellidae" genus="Trianguloscalpellum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pedunculata" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="mollucanum">T. mollucanum</taxonomicName>
|
||
,
|
||
<taxonomicName id="68BC4D005013FFF0FEE22754FE213F11" authorityName="Hoek" authorityYear="1883" box="[337,429,333,356]" class="Maxillopoda" family="Poecilasmatidae" genus="Glyptelasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pedunculata" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="gigas">T. gigas</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="68BC4D005013FFF0FE5A2754FDC93F10" baseAuthorityName="Hoek" baseAuthorityYear="1883" box="[489,581,332,357]" class="Maxillopoda" family="Scalpellidae" genus="Trianguloscalpellum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pedunculata" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="ovale">
|
||
<emphasis id="9DC8EA915013FFF0FE5A2754FDC93F10" box="[489,581,332,357]" italics="true" pageId="4" pageNumber="205">T. ovale</emphasis>
|
||
</taxonomicName>
|
||
has always been disputed in the previous literature (
|
||
<bibRefCitation id="CB2D4B725013FFF0FB1F2755FAC43F11" author="Zevina, G. B." box="[1196,1352,332,357]" pageId="4" pageNumber="205" refId="ref6098" refString="Zevina, G. B. (1981) Cirripede crustaceans of the suborder Lepadomorpha (Cirripedia, Thoracica) of the World Ocean. Part 1. Family Scalpellidae. Opredeliteli po Faune SSSR, Leningrad, 398 pp. [in Russian]" type="book" year="1981">Zevina, 1981</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="CB2D4B725013FFF0FAE42754FF5F3FFD" author="Jones, D. S." pageId="4" pageNumber="205" pagination="154 - 178" refId="ref5034" refString="Jones, D. S. (1992) Scalpellid barnacles (Cirripedia: Thoracica) from the northeastern and central eastern Australian continental shelf and slope. Memoirs of the Queensland Museum, 32, 154 - 178." type="journal article" year="1992">Jones, 1992</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="CB2D4B725013FFF0FF6E2769FEF73FFD" author="Young, P. S." box="[221,379,367,392]" pageId="4" pageNumber="205" pagination="1 - 54" refId="ref5886" refString="Young, P. S. (1998 b) The Cirripedia (Crustacea) collected by the " Fisheries Steamer Meteor " in the eastern Atlantic. Arquivos do Museu Nacional Rio de Janeiro, 58, 1 - 54." type="journal article" year="1998">Young, 1998b</bibRefCitation>
|
||
) and some authors consider that
|
||
<taxonomicName id="68BC4D005013FFF0FD5B2768FCDD3FFD" baseAuthorityName="Wyville-Thomson" baseAuthorityYear="1873" box="[744,849,369,392]" class="Maxillopoda" family="Scalpellidae" genus="Trianguloscalpellum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pedunculata" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="regium">
|
||
<emphasis id="9DC8EA915013FFF0FD5B2768FCDD3FFD" box="[744,849,369,392]" italics="true" pageId="4" pageNumber="205">T. regium</emphasis>
|
||
</taxonomicName>
|
||
represents a complex of cryptic species (
|
||
<bibRefCitation id="CB2D4B725013FFF0FAAA2776FEC93FD9" author="Shalaeva, K. & Boxshall, G." pageId="4" pageNumber="205" pagination="1 - 63" refId="ref5689" refString="Shalaeva, K. & Boxshall, G. (2014) An illustrated catalogue of the scalpellid barnacles (Crustacea: Cirripedia: Scalpellidae) collected during the HMS " Challenger " expedition and deposited in the Natural History Museum, London. Zootaxa, 3804 (1), 1 - 63. https: // doi. org / 10.11646 / zootaxa. 3804.1.1" type="journal article" year="2014">Shalaeva & Boxshall, 2014</bibRefCitation>
|
||
). Therefore, more detailed morphological characteristics, such as the features of the soft body of the
|
||
<typeStatus id="700788215013FFF0FF2427A1FF4B3FA5" box="[151,199,440,464]" pageId="4" pageNumber="205">type</typeStatus>
|
||
specimens, are imperative, as well as molecular data. The nucleotide sequences of 16S rRNA and mt COI gene from the present specimen were submitted to tGenbank (genbank number:
|
||
<accessionNumber id="B0EFAB605013FFF0FBE127C5FB5F3F80" box="[1106,1235,476,501]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MH938354" pageId="4" pageNumber="205">MH938354</accessionNumber>
|
||
and
|
||
<accessionNumber id="B0EFAB605013FFF0FAA727C5FA143F80" box="[1300,1432,476,501]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MH973609" pageId="4" pageNumber="205">MH973609</accessionNumber>
|
||
, respectively), and we hope they are useful for future taxonomic studies.
|
||
</paragraph>
|
||
<paragraph id="AF0336835013FFF0FF2423F0FD2E386B" blockId="4.[151,1435,1513,1566]" pageId="4" pageNumber="205">
|
||
<emphasis id="9DC8EA915013FFF0FF2423F0FE9B3B8B" bold="true" box="[151,279,1513,1535]" pageId="4" pageNumber="205">FIGURE. 3</emphasis>
|
||
<taxonomicName id="68BC4D005013FFF0FE9123F0FCD33B8A" authority="(Wyville-Thomson, 1873)" baseAuthorityName="Wyville-Thomson" baseAuthorityYear="1873" box="[290,863,1513,1535]" class="Maxillopoda" family="Scalpellidae" genus="Trianguloscalpellum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pedunculata" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="regium">
|
||
<emphasis id="9DC8EA915013FFF0FE9123F0FDC93B8A" box="[290,581,1513,1535]" italics="true" pageId="4" pageNumber="205">Trianguloscalpellum regium</emphasis>
|
||
(
|
||
<bibRefCitation id="CB2D4B725013FFF0FDE523F0FCDB3B8A" author="Wyville-Thomson, C." box="[598,855,1513,1535]" pageId="4" pageNumber="205" pagination="347 - 349" refId="ref5756" refString="Wyville-Thomson, C. (1873) Notes from the " Challenger ". Nature, 8, 347 - 349. https: // doi. org / 10.1038 / 008347 a 0" type="journal article" year="1873">Wyville-Thomson, 1873</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. A, mouthparts; B, mandibular palp; C, mandible; D, inferior angle of mandible; E, maxillule; F, maxilla.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |