treatments-xml/data/03/FB/C3/03FBC31AFFDFFF83FEB1D5F2D05A4492.xml
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<emphasis id="B926AE1EFFDFFF83FEB1D5F2D49C4542" bold="true" box="[293,478,749,773]" italics="true" pageId="1" pageNumber="228">Scymnomorphus</emphasis>
<bibRefCitation id="EFC30FFDFFDFFF83FE70D5F1D72B4542" author="Weise" box="[484,617,750,773]" firstAuthor="Weise" journalOrPublisher="Deutsche Entomologische Zeitschrift" pageId="1" pageNumber="228" pagination="289 - 304" part="1897" refId="ref7966" refString="WEISE J. 1897: Coccinellen aus Ostafrika (Usambara). Deutsche Entomologische Zeitschrift 1897: 289 - 304." title="Coccinellen aus Ostafrika (Usambara)" type="journal article" year="1897">Weise, 1897</bibRefCitation>
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rotundatus
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<bibRefCitation id="EFC30FFDFFDFFF83FE8DD431D4C74506" author="Weise" box="[281,389,814,833]" firstAuthor="Weise" journalOrPublisher="Deutsche Entomologische Zeitschrift" pageId="1" pageNumber="228" pagination="289 - 304" part="1897" refId="ref7966" refString="WEISE J. 1897: Coccinellen aus Ostafrika (Usambara). Deutsche Entomologische Zeitschrift 1897: 289 - 304." title="Coccinellen aus Ostafrika (Usambara)" type="journal article" year="1897">Weise, 1897</bibRefCitation>
(Africa) (designated by
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<treatmentCitation id="0AF3541DFFDFFF83FF21D456D4AC451B" box="[181,494,841,860]" pageId="1" pageNumber="228">ŚLIPIŃSKI &amp; TOMASZEWSKA (2005:380</treatmentCitation>
)
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;
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<bibRefCitation id="EFC30FFDFFDFFF83FE68D456D7DF451B" author="KOVAR" bookContentInfo="935 pp." box="[508,669,841,860]" editor="LOBL I. &amp; SMETANA A." firstAuthor="KOVAR" journalOrPublisher="Apollo Books, Stenstrup" pageId="1" pageNumber="228" pagination="71 - 73" refId="ref7033" refString="KOVAR I. 2007: New nomenclatorial and taxonomic acts, and comments. Coccinellidae. Pp. 71 - 73. In: LOBL I. &amp; SMETANA A. (eds.): Catalogue of Palaearctic Coleoptera. Vol. 4. Elateroidea - Derodontoidea - Bostrichoidea- Lymexyloidea - Cleroidea - Cucujoidea. Apollo Books, Stenstrup, 935 pp." title="New nomenclatorial and taxonomic acts, and comments. Coccinellidae. Pp" type="book chapter" volumeTitle="Catalogue of Palaearctic Coleoptera. Vol. 4. Elateroidea - Derodontoidea - Bostrichoidea- Lymexyloidea - Cleroidea - Cucujoidea" year="2007">KOVÁŘ (2007: 572)</bibRefCitation>
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;
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<bibRefCitation id="EFC30FFDFFDFFF83FD31D456D4C04530" author="ESCALONA &amp; SLIPINSKI" firstAuthor="ESCALONA" journalOrPublisher="Systematic Entomology" pageId="1" pageNumber="228" pagination="125 - 171" part="37" refId="ref6785" refString="ESCALONA H. &amp; SLIPINSKI A. 2012: Generic revision and phylogeny of Microweiseinae (Coleoptera: Coccinellidae). Systematic Entomology 37: 125 - 171." title="Generic revision and phylogeny of Microweiseinae (Coleoptera: Coccinellidae)" type="book" year="2012">ESCALONA &amp; ŚLIPIŃSKI (2012: 152)</bibRefCitation>
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<treatmentCitationGroup id="AB425522FFDFFF83FF03D460D7264580" pageId="1" pageNumber="228">
<emphasis id="B926AE1EFFDFFF83FF03D460D44D45D5" box="[151,271,895,914]" italics="true" pageId="1" pageNumber="228">Scotoscymnus</emphasis>
<treatmentCitation id="0AF3541DFFDFFF83FE83D461D4F245D6" author="WEISE J." box="[279,432,894,913]" page="458" pageId="1" pageNumber="228" year="1901">
<bibRefCitation id="EFC30FFDFFDFFF83FE83D461D4F245D6" author="Weise" box="[279,432,894,913]" firstAuthor="Weise" journalOrPublisher="Deutsche Entomologische Zeitschrift" pageId="1" pageNumber="228" pagination="446 - 459" part="1900" refId="ref7987" refString="WEISE J. 1901: Beschreibungen Africanischer Chrysomeliden nebst synonymischen Bemerkungen. Deutsche Entomologische Zeitschrift 1900: 446 - 459." title="Beschreibungen Africanischer Chrysomeliden nebst synonymischen Bemerkungen" type="journal article" year="1901">Weise, 1901: 458</bibRefCitation>
</treatmentCitation>
(unnecessary replacement name, see
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<bibRefCitation id="EFC30FFDFFDFFF83FF21D486D4FC45EB" author="SLIPINSKI &amp; TOMASZEWSKA" box="[181,446,921,940]" firstAuthor="SLIPINSKI" journalOrPublisher="Australian Journal of Entomology" pageId="1" pageNumber="228" pagination="369 - 384" part="44" refId="ref7717" refString="SLIPINSKI A. &amp; TOMASZEWSKA W. 2005: Revision of the Australian Coccinellidae (Coleoptera). Part 3. Tribe Sukunahikonini. Australian Journal of Entomology 44: 369 - 384." title="Revision of the Australian Coccinellidae (Coleoptera). Part 3. Tribe Sukunahikonini" type="journal article" year="2005">ŚLIPIŃSKI &amp; TOMASZEWSKA 2005</bibRefCitation>
</treatmentCitation>
).
<bibRefCitation id="EFC30FFDFFDFFF83FE5FD486D73A45EB" author="GORDON" box="[459,632,921,940]" firstAuthor="GORDON" journalOrPublisher="Coleopterist Bulletin" pageId="1" pageNumber="228" pagination="185 - 228" part="31" refId="ref6913" refString="GORDON R. D. 1977: Classification and phylogeny of the New World Sticholotidinae (Coccinellidae). Coleopterist Bulletin 31: 185 - 228." title="Classification and phylogeny of the New World Sticholotidinae (Coccinellidae)" type="journal article" year="1977">
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)
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;
<bibRefCitation id="EFC30FFDFFDFFF83FDEBD486D59D4580" author="FURSCH" firstAuthor="FURSCH" pageId="1" pageNumber="228" pagination="279 - 295" refId="ref6814" refString="FURSCH H. 1985: Die Afrikanischen Sukunahikonini und Microweiseini mit Diskussion uber alle Gattungen (Col. Cocc.). Deutsche Entomologische Zeitschrift, Neue Folge 32: 279 - 295." type="journal article" year="1985">
<treatmentCitation id="0AF3541DFFDFFF83FDEBD486D59A4580" pageId="1" pageNumber="228">FÜRSCH (1985: 283</treatmentCitation>
)
</bibRefCitation>
;
<bibRefCitation id="EFC30FFDFFDFFF83FF7CD4ABD4E64580" author="MIYATAKE" box="[232,420,948,967]" firstAuthor="MIYATAKE" journalOrPublisher="Memoirs of the College of Agriculture, Ehime University" pageId="1" pageNumber="228" pagination="223 - 292" part="38" refId="ref7385" refString="MIYATAKE M. 1994: Revisional studies on Asian genera of the subfamily Sticholotidinae (Coleoptera: Coccinellidae). Memoirs of the College of Agriculture, Ehime University 38: 223 - 292." title="Revisional studies on Asian genera of the subfamily Sticholotidinae (Coleoptera: Coccinellidae)" type="journal article" year="1994">
<treatmentCitation id="0AF3541DFFDFFF83FF7CD4ABD4DC4580" box="[232,414,948,967]" pageId="1" pageNumber="228">MIYATAKE (1994: 235</treatmentCitation>
)
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;
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<emphasis id="B926AE1EFFDFFF83FF03D4D1D45245A6" box="[151,272,974,993]" italics="true" pageId="1" pageNumber="228">Sukunahikona</emphasis>
<treatmentCitation id="0AF3541DFFDFFF83FE83D4D1D4F445A6" author="KAMIYA H." box="[279,438,974,993]" page="22" pageId="1" pageNumber="228" year="1960">
<bibRefCitation id="EFC30FFDFFDFFF83FE83D4D1D4F445A6" author="Kamiya" box="[279,438,974,993]" firstAuthor="Kamiya" journalOrPublisher="Kontyu" pageId="1" pageNumber="228" pagination="22 - 26" part="28" refId="ref7012" refString="KAMIYA H. 1960: A new tribe of Coccinellidae (Coleoptera). Kontyu 28: 22 - 26." title="A new tribe of Coccinellidae (Coleoptera)" type="journal article" year="1960">Kamiya, 1960: 22</bibRefCitation>
</treatmentCitation>
.
<typeStatus id="54E9CCAEFFDFFF83FE2BD4D1D4AB45A6" box="[447,489,974,993]" pageId="1" pageNumber="228">Type</typeStatus>
species:
<emphasis id="B926AE1EFFDFFF83FDAED4D1D58B45BB" italics="true" pageId="1" pageNumber="228">Sukunahikona japonica</emphasis>
<bibRefCitation id="EFC30FFDFFDFFF83FF46D4F6D40C45BB" author="Kamiya" box="[210,334,1001,1020]" firstAuthor="Kamiya" journalOrPublisher="Kontyu" pageId="1" pageNumber="228" pagination="22 - 26" part="28" refId="ref7012" refString="KAMIYA H. 1960: A new tribe of Coccinellidae (Coleoptera). Kontyu 28: 22 - 26." title="A new tribe of Coccinellidae (Coleoptera)" type="journal article" year="1960">Kamiya, 1960</bibRefCitation>
(original designation). Synonymized by
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<treatmentCitationGroup id="AB425522FFDFFF83FF03D301D7704220" pageId="1" pageNumber="228">
<emphasis id="B926AE1EFFDFFF83FF03D301D4504276" box="[151,274,1054,1073]" italics="true" pageId="1" pageNumber="228">Hikonasukuna</emphasis>
<treatmentCitation id="0AF3541DFFDFFF83FE8DD301D4DE4276" author="SASAJI H." box="[281,412,1054,1073]" page="4" pageId="1" pageNumber="228" year="1967">
<bibRefCitation id="EFC30FFDFFDFFF83FE8DD301D4DE4276" author="Sasaji" box="[281,412,1054,1073]" firstAuthor="Sasaji" journalOrPublisher="Etizenia, Occasional Publications of the Biological Laboratory, Fukui University" pageId="1" pageNumber="228" pagination="1 - 28" part="25" refId="ref7638" refString="SASAJI H. 1967: A revision of the Formosan Coccinellidae (I) the subfamily Sticholotidinae, with an establishment of a new tribe (Coleoptera). Etizenia, Occasional Publications of the Biological Laboratory, Fukui University 25: 1 - 28." title="A revision of the Formosan Coccinellidae (I) the subfamily Sticholotidinae, with an establishment of a new tribe (Coleoptera)" type="journal article" year="1967">Sasaji, 1967: 4</bibRefCitation>
</treatmentCitation>
.
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<emphasis id="B926AE1EFFDFFF83FDB4D301D7D94276" box="[544,667,1054,1073]" italics="true" pageId="1" pageNumber="228">Hikonasukuna</emphasis>
<taxonomicName id="4C52098FFFDFFF83FD36D301D45C420B" authority="Sasaji, 1967" authorityName="Sasaji" authorityYear="1967" class="Insecta" family="Coccinellidae" genus="Scymnomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="228" phylum="Arthropoda" rank="species" species="monticola">
<emphasis id="B926AE1EFFDFFF83FD36D301D7B54276" box="[674,759,1054,1073]" italics="true" pageId="1" pageNumber="228">monticola</emphasis>
<bibRefCitation id="EFC30FFDFFDFFF83FF21D326D45C420B" author="Sasaji" box="[181,286,1081,1100]" firstAuthor="Sasaji" journalOrPublisher="Etizenia, Occasional Publications of the Biological Laboratory, Fukui University" pageId="1" pageNumber="228" pagination="1 - 28" part="25" refId="ref7638" refString="SASAJI H. 1967: A revision of the Formosan Coccinellidae (I) the subfamily Sticholotidinae, with an establishment of a new tribe (Coleoptera). Etizenia, Occasional Publications of the Biological Laboratory, Fukui University 25: 1 - 28." title="A revision of the Formosan Coccinellidae (I) the subfamily Sticholotidinae, with an establishment of a new tribe (Coleoptera)" type="journal article" year="1967">Sasaji, 1967</bibRefCitation>
</taxonomicName>
(by original designation).
<treatmentCitation id="0AF3541DFFDFFF83FE6BD326D7E5420B" author="FURSCH" box="[511,679,1081,1100]" page="287" pageId="1" pageNumber="228" year="1985">
<bibRefCitation id="EFC30FFDFFDFFF83FE6BD326D7E5420B" author="FURSCH" box="[511,679,1081,1100]" firstAuthor="FURSCH" pageId="1" pageNumber="228" pagination="279 - 295" refId="ref6814" refString="FURSCH H. 1985: Die Afrikanischen Sukunahikonini und Microweiseini mit Diskussion uber alle Gattungen (Col. Cocc.). Deutsche Entomologische Zeitschrift, Neue Folge 32: 279 - 295." type="journal article" year="1985">FÜRSCH (1985: 287)</bibRefCitation>
</treatmentCitation>
. Synonymized by
<treatmentCitation id="0AF3541DFFDFFF83FE9ED34BD76D4220" author="ESCALONA H. &amp; SLIPINSKI A." box="[266,559,1108,1127]" page="152" pageId="1" pageNumber="228" year="2012">
<bibRefCitation id="EFC30FFDFFDFFF83FE9ED34BD76D4220" author="ESCALONA &amp; SLIPINSKI" box="[266,559,1108,1127]" firstAuthor="ESCALONA" journalOrPublisher="Systematic Entomology" pageId="1" pageNumber="228" pagination="125 - 171" part="37" refId="ref6785" refString="ESCALONA H. &amp; SLIPINSKI A. 2012: Generic revision and phylogeny of Microweiseinae (Coleoptera: Coccinellidae). Systematic Entomology 37: 125 - 171." title="Generic revision and phylogeny of Microweiseinae (Coleoptera: Coccinellidae)" type="book" year="2012">ESCALONA &amp; ŚLIPIŃSKI (2012: 152)</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BED720CFFDFFF83FF03D371D42D42F0" blockId="1.[151,761,749,1207]" pageId="1" pageNumber="228">
<treatmentCitationGroup id="AB425522FFDFFF83FF03D371D42E42F0" pageId="1" pageNumber="228">
<emphasis id="B926AE1EFFDFFF83FF03D371D59942C6" box="[151,219,1134,1153]" italics="true" pageId="1" pageNumber="228">Orculus</emphasis>
<treatmentCitation id="0AF3541DFFDFFF83FF4BD371D43542C6" author="SICARD A." box="[223,375,1134,1153]" page="233" pageId="1" pageNumber="228" year="1931">
<bibRefCitation id="EFC30FFDFFDFFF83FF4BD371D43542C6" author="Sicard" box="[223,375,1134,1153]" firstAuthor="Sicard" journalOrPublisher="Annals and Magazine of Natural History" pageId="1" pageNumber="228" pagination="228 - 234" part="8" refId="ref7684" refString="SICARD A. 1931: Descriptions d'especes nouvelles appartenanta la famille des Coccinellides (Col.). Annals and Magazine of Natural History, Series 10 8: 228 - 234." title="Descriptions d'especes nouvelles appartenanta la famille des Coccinellides (Col.)" type="journal article" year="1931">Sicard, 1931: 233</bibRefCitation>
</treatmentCitation>
.
<typeStatus id="54E9CCAEFFDFFF83FEE9D371D4E542C6" box="[381,423,1134,1153]" pageId="1" pageNumber="228">Type</typeStatus>
species
<emphasis id="B926AE1EFFDFFF83FE79D371D7CB42C5" box="[493,649,1134,1154]" italics="true" pageId="1" pageNumber="228">Orculus castaneus</emphasis>
<bibRefCitation id="EFC30FFDFFDFFF83FD19D371D7BA42C6" author="Sicard" box="[653,760,1134,1153]" firstAuthor="Sicard" journalOrPublisher="Annals and Magazine of Natural History" pageId="1" pageNumber="228" pagination="228 - 234" part="8" refId="ref7684" refString="SICARD A. 1931: Descriptions d'especes nouvelles appartenanta la famille des Coccinellides (Col.). Annals and Magazine of Natural History, Series 10 8: 228 - 234." title="Descriptions d'especes nouvelles appartenanta la famille des Coccinellides (Col.)" type="journal article" year="1931">Sicard, 1931</bibRefCitation>
(by monotypy).
<treatmentCitation id="0AF3541DFFDFFF83FED5D396D4AF42DB" author="FURSCH" box="[321,493,1161,1180]" page="289" pageId="1" pageNumber="228" year="1985">
<bibRefCitation id="EFC30FFDFFDFFF83FED5D396D4AF42DB" author="FURSCH" box="[321,493,1161,1180]" firstAuthor="FURSCH" pageId="1" pageNumber="228" pagination="279 - 295" refId="ref6814" refString="FURSCH H. 1985: Die Afrikanischen Sukunahikonini und Microweiseini mit Diskussion uber alle Gattungen (Col. Cocc.). Deutsche Entomologische Zeitschrift, Neue Folge 32: 279 - 295." type="journal article" year="1985">FÜRSCH (1985: 289)</bibRefCitation>
</treatmentCitation>
. Synonymized by
<treatmentCitation id="0AF3541DFFDFFF83FD19D396D42E42F0" author="ESCALONA H. &amp; SLIPINSKI A." page="152" pageId="1" pageNumber="228" year="2012">
<bibRefCitation id="EFC30FFDFFDFFF83FD19D396D42E42F0" author="ESCALONA &amp; SLIPINSKI" firstAuthor="ESCALONA" journalOrPublisher="Systematic Entomology" pageId="1" pageNumber="228" pagination="125 - 171" part="37" refId="ref6785" refString="ESCALONA H. &amp; SLIPINSKI A. 2012: Generic revision and phylogeny of Microweiseinae (Coleoptera: Coccinellidae). Systematic Entomology 37: 125 - 171." title="Generic revision and phylogeny of Microweiseinae (Coleoptera: Coccinellidae)" type="book" year="2012">ESCALONA &amp; ŚLIPIŃSKI (2012: 152)</bibRefCitation>
</treatmentCitation>
</treatmentCitationGroup>
.
</paragraph>
</subSubSection>
<subSubSection id="C3482187FFDFFF83FF03D3D0D15D4432" pageId="1" pageNumber="228" type="diagnosis">
<paragraph id="8BED720CFFDFFF83FF03D3D0D4FF4001" blockId="1.[151,760,1231,2086]" pageId="1" pageNumber="228">
<emphasis id="B926AE1EFFDFFF83FF03D3D0D4CC42A1" bold="true" box="[151,398,1231,1254]" pageId="1" pageNumber="228">Differential diagnosis.</emphasis>
<taxonomicName id="4C52098FFFDFFF83FE06D3D0D70342A1" authorityName="Weise" authorityYear="1897" box="[402,577,1231,1254]" class="Insecta" family="Coccinellidae" genus="Scymnomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="234" phylum="Arthropoda" rank="genus">
<emphasis id="B926AE1EFFDFFF83FE06D3D0D70342A1" box="[402,577,1231,1254]" italics="true" pageId="1" pageNumber="228">Scymnomorphus</emphasis>
</taxonomicName>
is similar to
<emphasis id="B926AE1EFFDFFF83FD50D3D0D5914341" italics="true" pageId="1" pageNumber="228">Pharellus</emphasis>
Sicard, 1928 and
<emphasis id="B926AE1EFFDFFF83FE1AD3F0D7524341" box="[398,528,1263,1286]" italics="true" pageId="1" pageNumber="228">Paraphellus</emphasis>
Chazeau, 1981. From
<emphasis id="B926AE1EFFDFFF83FF03D210D45A4361" box="[151,280,1295,1318]" italics="true" pageId="1" pageNumber="228">Paraphellus</emphasis>
it can be easily separated by club consisting of two or three antennomeres, in contrast to one antennomere in
<emphasis id="B926AE1EFFDFFF83FF26D250D4764321" box="[178,308,1359,1382]" italics="true" pageId="1" pageNumber="228">Paraphellus</emphasis>
. From
<emphasis id="B926AE1EFFDFFF83FE15D250D4AA4321" box="[385,488,1359,1382]" italics="true" pageId="1" pageNumber="228">Pharellus</emphasis>
it differs in the following characters: maxillary cardo broad, visible externally (very narrow, partially visible in
<emphasis id="B926AE1EFFDFFF83FE53D290D76C43E1" box="[455,558,1423,1446]" italics="true" pageId="1" pageNumber="228">Pharellus</emphasis>
), submentum distinctly narrower than mentum (submentum and mentum equally broad in
<emphasis id="B926AE1EFFDFFF83FEC6D2D0D4FB43A1" box="[338,441,1487,1510]" italics="true" pageId="1" pageNumber="228">Pharellus</emphasis>
), mentum deeply emarginate apically (mentum truncate apically in
<emphasis id="B926AE1EFFDFFF83FDD6D2F0D7E84041" box="[578,682,1519,1542]" italics="true" pageId="1" pageNumber="228">Pharellus</emphasis>
), male genitalia with parameres bearing setae (parameres reduced, without setae in
<emphasis id="B926AE1EFFDFFF83FED3D130D4EC4001" box="[327,430,1583,1606]" italics="true" pageId="1" pageNumber="228">Pharellus</emphasis>
).
</paragraph>
<paragraph id="8BED720CFFDFFF83FF03D150D1674452" blockId="1.[151,760,1231,2086]" lastBlockId="1.[828,1437,190,725]" pageId="1" pageNumber="228">
<emphasis id="B926AE1EFFDFFF83FF03D150D44C4021" bold="true" box="[151,270,1615,1638]" pageId="1" pageNumber="228">Diagnosis.</emphasis>
Body minute, round to oval, convex, dorsum pubescent (
<figureCitation id="13696E89FFDFFF83FE83D170D4D440C1" box="[279,406,1647,1670]" captionStart="Fig" captionStartId="2.[151,187,2036,2055]" captionTargetBox="[193,1417,192,2003]" captionTargetId="figure@2.[159,1428,180,2014]" captionTargetPageId="2" captionText="Fig.1. Habitus of Scymnomorphus species from Mt. Wilhelm, New Guinea. AF S. bimaculatus sp. nov.; GI S. kausi sp. nov.; JL S. papuensis sp. nov. A, D, G, J dorsal view; B, E, H, K lateral view; C, F, I, L frontal view." figureDoi="http://doi.org/10.5281/zenodo.3699220" httpUri="https://zenodo.org/record/3699220/files/figure.png" pageId="1" pageNumber="228">Figs 1AL</figureCitation>
). Head transverse with clypeus and frons usually produced, sometimes rostrate, clypeus bordered laterally (
<figureCitation id="13696E89FFDFFF83FE18D1B0D7414081" box="[396,515,1711,1734]" captionStart="Fig" captionStartId="6.[153,184,2010,2029]" captionTargetBox="[153,1438,457,1984]" captionTargetId="figure@6.[153,1438,457,1984]" captionTargetPageId="6" captionText="Fig. 4. Morphology of Scymnomorphus species. AC, EG S. bimaculatus sp. nov.: A habitus, dorsal; B magnification of the elytral punctures; C mouthparts, ventral; E magnification of the metaventral punctures; F meso- and metaventrite; G head, anterior view. D, HJ S. papuensis sp. nov.: D habitus, ventral; H magnification of the metaventral punctures; I meso- and metaventrite; J head, anterior view." figureDoi="http://doi.org/10.5281/zenodo.3699226" httpUri="https://zenodo.org/record/3699226/files/figure.png" pageId="1" pageNumber="228">Figs 4G, J</figureCitation>
). Eyes small, convex, coarsely faceted. Antenna consisting of 10 antennomeres, two (sometimes three) terminal antennomeres forming distinct club (
<figureCitation id="13696E89FFDFFF83FEBED010D4DB4161" box="[298,409,1807,1830]" captionStart="Fig" captionStartId="4.[151,182,1972,1991]" captionTargetBox="[167,1427,196,1883]" captionTargetId="figure@4.[151,1436,186,1908]" captionTargetPageId="4" captionText="Fig. 2. Morphology of Scymnomorphus species. AH S. bimaculatus sp. nov.: A abdomen, female; B antenna; C ovipositor; D spermatheca; E tegmen; inner view; F tegmen, lateral view; G penis, lateral view; H abdomen, male. IP S. kausi sp. nov.: I abdomen, female; J antenna; K ovipositor; L spermatheca; M tegmen, inner view; N tegmen, lateral view; O penis, lateral view; P abdomen, male (arrows indicate glandular opening pores)." figureDoi="http://doi.org/10.5281/zenodo.3699222" httpUri="https://zenodo.org/record/3699222/files/figure.png" pageId="1" pageNumber="228">Figs 2B, J</figureCitation>
,
<figureCitation id="13696E89FFDFFF83FE37D010D48A4161" box="[419,456,1807,1830]" captionStart="Fig" captionStartId="5.[151,182,2036,2055]" captionTargetBox="[227,1358,1374,2009]" captionTargetId="figure@5.[218,1370,1368,2012]" captionTargetPageId="5" captionText="Fig. 3. Morphology of Scymnomorphus species. AC S. papuensis sp. nov.: A antenna; B female genitalia; C abdomen, female. D S. kausi sp. nov., head and mouthparts, ventral (lp labial palp, m mentum, pm prementum, sm submentum)." figureDoi="http://doi.org/10.5281/zenodo.3699224" httpUri="https://zenodo.org/record/3699224/files/figure.png" pageId="1" pageNumber="228">3A</figureCitation>
). Maxillary cardo broad and clearly visible externally (
<figureCitation id="13696E89FFDFFF83FE20D030D7484101" box="[436,522,1839,1862]" captionStart="Fig" captionStartId="6.[153,184,2010,2029]" captionTargetBox="[153,1438,457,1984]" captionTargetId="figure@6.[153,1438,457,1984]" captionTargetPageId="6" captionText="Fig. 4. Morphology of Scymnomorphus species. AC, EG S. bimaculatus sp. nov.: A habitus, dorsal; B magnification of the elytral punctures; C mouthparts, ventral; E magnification of the metaventral punctures; F meso- and metaventrite; G head, anterior view. D, HJ S. papuensis sp. nov.: D habitus, ventral; H magnification of the metaventral punctures; I meso- and metaventrite; J head, anterior view." figureDoi="http://doi.org/10.5281/zenodo.3699226" httpUri="https://zenodo.org/record/3699226/files/figure.png" pageId="1" pageNumber="228">Fig. 4C</figureCitation>
). Terminal palpomere elongate-conical (
<figureCitation id="13696E89FFDFFF83FECFD050D4F74121" box="[347,437,1871,1894]" captionStart="Fig" captionStartId="5.[151,182,2036,2055]" captionTargetBox="[227,1358,1374,2009]" captionTargetId="figure@5.[218,1370,1368,2012]" captionTargetPageId="5" captionText="Fig. 3. Morphology of Scymnomorphus species. AC S. papuensis sp. nov.: A antenna; B female genitalia; C abdomen, female. D S. kausi sp. nov., head and mouthparts, ventral (lp labial palp, m mentum, pm prementum, sm submentum)." figureDoi="http://doi.org/10.5281/zenodo.3699224" httpUri="https://zenodo.org/record/3699224/files/figure.png" pageId="1" pageNumber="228">Figs 3D</figureCitation>
,
<figureCitation id="13696E89FFDFFF83FE56D050D4BC4121" box="[450,510,1871,1894]" captionStart="Fig" captionStartId="6.[153,184,2010,2029]" captionTargetBox="[153,1438,457,1984]" captionTargetId="figure@6.[153,1438,457,1984]" captionTargetPageId="6" captionText="Fig. 4. Morphology of Scymnomorphus species. AC, EG S. bimaculatus sp. nov.: A habitus, dorsal; B magnification of the elytral punctures; C mouthparts, ventral; E magnification of the metaventral punctures; F meso- and metaventrite; G head, anterior view. D, HJ S. papuensis sp. nov.: D habitus, ventral; H magnification of the metaventral punctures; I meso- and metaventrite; J head, anterior view." figureDoi="http://doi.org/10.5281/zenodo.3699226" httpUri="https://zenodo.org/record/3699226/files/figure.png" pageId="1" pageNumber="228">4G, J</figureCitation>
). Submentum distinctly narrower than mentum, mentum deeply emarginate apically (
<figureCitation id="13696E89FFDFFF83FF6BD090D41F41E1" box="[255,349,1935,1958]" captionStart="Fig" captionStartId="5.[151,182,2036,2055]" captionTargetBox="[227,1358,1374,2009]" captionTargetId="figure@5.[218,1370,1368,2012]" captionTargetPageId="5" captionText="Fig. 3. Morphology of Scymnomorphus species. AC S. papuensis sp. nov.: A antenna; B female genitalia; C abdomen, female. D S. kausi sp. nov., head and mouthparts, ventral (lp labial palp, m mentum, pm prementum, sm submentum)." figureDoi="http://doi.org/10.5281/zenodo.3699224" httpUri="https://zenodo.org/record/3699224/files/figure.png" pageId="1" pageNumber="228">Figs 3D</figureCitation>
,
<figureCitation id="13696E89FFDFFF83FEF8D090D4CD41E1" box="[364,399,1935,1958]" captionStart="Fig" captionStartId="6.[153,184,2010,2029]" captionTargetBox="[153,1438,457,1984]" captionTargetId="figure@6.[153,1438,457,1984]" captionTargetPageId="6" captionText="Fig. 4. Morphology of Scymnomorphus species. AC, EG S. bimaculatus sp. nov.: A habitus, dorsal; B magnification of the elytral punctures; C mouthparts, ventral; E magnification of the metaventral punctures; F meso- and metaventrite; G head, anterior view. D, HJ S. papuensis sp. nov.: D habitus, ventral; H magnification of the metaventral punctures; I meso- and metaventrite; J head, anterior view." figureDoi="http://doi.org/10.5281/zenodo.3699226" httpUri="https://zenodo.org/record/3699226/files/figure.png" pageId="1" pageNumber="228">4C</figureCitation>
). Pronotum with line separating anterior corners from pronotal disc. Prosternum strongly reduced, prosternal process reduced to narrow carina (
<figureCitation id="13696E89FFDFFF83FF0BD0F0D5B54E41" box="[159,247,2031,2054]" captionStart="Fig" captionStartId="6.[153,184,2010,2029]" captionTargetBox="[153,1438,457,1984]" captionTargetId="figure@6.[153,1438,457,1984]" captionTargetPageId="6" captionText="Fig. 4. Morphology of Scymnomorphus species. AC, EG S. bimaculatus sp. nov.: A habitus, dorsal; B magnification of the elytral punctures; C mouthparts, ventral; E magnification of the metaventral punctures; F meso- and metaventrite; G head, anterior view. D, HJ S. papuensis sp. nov.: D habitus, ventral; H magnification of the metaventral punctures; I meso- and metaventrite; J head, anterior view." figureDoi="http://doi.org/10.5281/zenodo.3699226" httpUri="https://zenodo.org/record/3699226/files/figure.png" pageId="1" pageNumber="228">Fig. 4F</figureCitation>
). Mesoventrite short and broad, fused in mid part with metaventrite (
<figureCitation id="13696E89FFDFFF83FE02DF10D7424E61" box="[406,512,2063,2086]" captionStart="Fig" captionStartId="6.[153,184,2010,2029]" captionTargetBox="[153,1438,457,1984]" captionTargetId="figure@6.[153,1438,457,1984]" captionTargetPageId="6" captionText="Fig. 4. Morphology of Scymnomorphus species. AC, EG S. bimaculatus sp. nov.: A habitus, dorsal; B magnification of the elytral punctures; C mouthparts, ventral; E magnification of the metaventral punctures; F meso- and metaventrite; G head, anterior view. D, HJ S. papuensis sp. nov.: D habitus, ventral; H magnification of the metaventral punctures; I meso- and metaventrite; J head, anterior view." figureDoi="http://doi.org/10.5281/zenodo.3699226" httpUri="https://zenodo.org/record/3699226/files/figure.png" pageId="1" pageNumber="228">Figs 4F, I</figureCitation>
,
<figureCitation id="13696E89FFDFFF83FD9FDF10D76C4E61" box="[523,558,2063,2086]" captionStart="Fig" captionStartId="7.[151,182,1201,1220]" captionTargetBox="[151,1436,190,1186]" captionTargetId="figure@7.[151,1436,190,1190]" captionTargetPageId="7" captionText="Fig. 5. Morphology of Scymnomorphus species. AD, F S. kausi sp. nov.: A habitus, dorsal; B ventral view; C magnification of metaventral punctures; D glandular openings on abdominal ventrite 5 in male; F head, anterior view. E S. bimaculatus sp. nov., glandular opening on abdominal ventrite 5 in male." figureDoi="http://doi.org/10.5281/zenodo.3699228" httpUri="https://zenodo.org/record/3699228/files/figure.png" pageId="1" pageNumber="228">5B</figureCitation>
). Elytra punctured (
<figureCitation id="13696E89FFDFFF83FCD7D7A1D6DD4692" box="[835,927,190,213]" captionStart="Fig" captionStartId="6.[153,184,2010,2029]" captionTargetBox="[153,1438,457,1984]" captionTargetId="figure@6.[153,1438,457,1984]" captionTargetPageId="6" captionText="Fig. 4. Morphology of Scymnomorphus species. AC, EG S. bimaculatus sp. nov.: A habitus, dorsal; B magnification of the elytral punctures; C mouthparts, ventral; E magnification of the metaventral punctures; F meso- and metaventrite; G head, anterior view. D, HJ S. papuensis sp. nov.: D habitus, ventral; H magnification of the metaventral punctures; I meso- and metaventrite; J head, anterior view." figureDoi="http://doi.org/10.5281/zenodo.3699226" httpUri="https://zenodo.org/record/3699226/files/figure.png" pageId="1" pageNumber="228">Figs 4A</figureCitation>
,
<figureCitation id="13696E89FFDFFF83FC38D7A1D6934692" box="[940,977,190,213]" captionStart="Fig" captionStartId="7.[151,182,1201,1220]" captionTargetBox="[151,1436,190,1186]" captionTargetId="figure@7.[151,1436,190,1190]" captionTargetPageId="7" captionText="Fig. 5. Morphology of Scymnomorphus species. AD, F S. kausi sp. nov.: A habitus, dorsal; B ventral view; C magnification of metaventral punctures; D glandular openings on abdominal ventrite 5 in male; F head, anterior view. E S. bimaculatus sp. nov., glandular opening on abdominal ventrite 5 in male." figureDoi="http://doi.org/10.5281/zenodo.3699228" httpUri="https://zenodo.org/record/3699228/files/figure.png" pageId="1" pageNumber="228">5A</figureCitation>
), epipleuron incomplete apically with carina not reaching elytral base (
<figureCitation id="13696E89FFDFFF83FB1DD7C1D1A346B2" box="[1161,1249,222,245]" captionStart="Fig" captionStartId="6.[153,184,2010,2029]" captionTargetBox="[153,1438,457,1984]" captionTargetId="figure@6.[153,1438,457,1984]" captionTargetPageId="6" captionText="Fig. 4. Morphology of Scymnomorphus species. AC, EG S. bimaculatus sp. nov.: A habitus, dorsal; B magnification of the elytral punctures; C mouthparts, ventral; E magnification of the metaventral punctures; F meso- and metaventrite; G head, anterior view. D, HJ S. papuensis sp. nov.: D habitus, ventral; H magnification of the metaventral punctures; I meso- and metaventrite; J head, anterior view." figureDoi="http://doi.org/10.5281/zenodo.3699226" httpUri="https://zenodo.org/record/3699226/files/figure.png" pageId="1" pageNumber="228">Fig. 4D</figureCitation>
), associated with pits and pores (
<figureCitation id="13696E89FFDFFF83FC70D7E1D1124752" box="[996,1104,254,277]" captionStart="Fig" captionStartId="6.[153,184,2010,2029]" captionTargetBox="[153,1438,457,1984]" captionTargetId="figure@6.[153,1438,457,1984]" captionTargetPageId="6" captionText="Fig. 4. Morphology of Scymnomorphus species. AC, EG S. bimaculatus sp. nov.: A habitus, dorsal; B magnification of the elytral punctures; C mouthparts, ventral; E magnification of the metaventral punctures; F meso- and metaventrite; G head, anterior view. D, HJ S. papuensis sp. nov.: D habitus, ventral; H magnification of the metaventral punctures; I meso- and metaventrite; J head, anterior view." figureDoi="http://doi.org/10.5281/zenodo.3699226" httpUri="https://zenodo.org/record/3699226/files/figure.png" pageId="1" pageNumber="228">Figs 4F, I</figureCitation>
). Metaventrite without visible discrimen, usually densely punctured with postcoxal lines descending laterally, associated with pits and pores (
<figureCitation id="13696E89FFDFFF83FCD7D641D6E64732" box="[835,932,350,373]" captionStart="Fig" captionStartId="6.[153,184,2010,2029]" captionTargetBox="[153,1438,457,1984]" captionTargetId="figure@6.[153,1438,457,1984]" captionTargetPageId="6" captionText="Fig. 4. Morphology of Scymnomorphus species. AC, EG S. bimaculatus sp. nov.: A habitus, dorsal; B magnification of the elytral punctures; C mouthparts, ventral; E magnification of the metaventral punctures; F meso- and metaventrite; G head, anterior view. D, HJ S. papuensis sp. nov.: D habitus, ventral; H magnification of the metaventral punctures; I meso- and metaventrite; J head, anterior view." figureDoi="http://doi.org/10.5281/zenodo.3699226" httpUri="https://zenodo.org/record/3699226/files/figure.png" pageId="1" pageNumber="228">Figs 4F,I</figureCitation>
,
<figureCitation id="13696E89FFDFFF83FC3AD641D6924732" box="[942,976,350,373]" captionStart="Fig" captionStartId="7.[151,182,1201,1220]" captionTargetBox="[151,1436,190,1186]" captionTargetId="figure@7.[151,1436,190,1190]" captionTargetPageId="7" captionText="Fig. 5. Morphology of Scymnomorphus species. AD, F S. kausi sp. nov.: A habitus, dorsal; B ventral view; C magnification of metaventral punctures; D glandular openings on abdominal ventrite 5 in male; F head, anterior view. E S. bimaculatus sp. nov., glandular opening on abdominal ventrite 5 in male." figureDoi="http://doi.org/10.5281/zenodo.3699228" httpUri="https://zenodo.org/record/3699228/files/figure.png" pageId="1" pageNumber="228">5B</figureCitation>
). Abdomen with six ventrites in both sexes. Abdominal postcoxal lines incomplete or reaching lateral margin, divided into two lines, usually associated with pits and pores (
<figureCitation id="13696E89FFDFFF83FC74D6A1D1C84792" box="[992,1162,446,469]" captionStart="Fig" captionStartId="4.[151,182,1972,1991]" captionTargetBox="[167,1427,196,1883]" captionTargetId="figure@4.[151,1436,186,1908]" captionTargetPageId="4" captionText="Fig. 2. Morphology of Scymnomorphus species. AH S. bimaculatus sp. nov.: A abdomen, female; B antenna; C ovipositor; D spermatheca; E tegmen; inner view; F tegmen, lateral view; G penis, lateral view; H abdomen, male. IP S. kausi sp. nov.: I abdomen, female; J antenna; K ovipositor; L spermatheca; M tegmen, inner view; N tegmen, lateral view; O penis, lateral view; P abdomen, male (arrows indicate glandular opening pores)." figureDoi="http://doi.org/10.5281/zenodo.3699222" httpUri="https://zenodo.org/record/3699222/files/figure.png" pageId="1" pageNumber="228">Figs 2A, H, I, P</figureCitation>
,
<figureCitation id="13696E89FFDFFF83FB01D6A1D1F54792" box="[1173,1207,446,469]" captionStart="Fig" captionStartId="5.[151,182,2036,2055]" captionTargetBox="[227,1358,1374,2009]" captionTargetId="figure@5.[218,1370,1368,2012]" captionTargetPageId="5" captionText="Fig. 3. Morphology of Scymnomorphus species. AC S. papuensis sp. nov.: A antenna; B female genitalia; C abdomen, female. D S. kausi sp. nov., head and mouthparts, ventral (lp labial palp, m mentum, pm prementum, sm submentum)." figureDoi="http://doi.org/10.5281/zenodo.3699224" httpUri="https://zenodo.org/record/3699224/files/figure.png" pageId="1" pageNumber="228">3C</figureCitation>
). Tarsal formula 4-4- 4. Male genitalia asymmetrical, with parameres reduced, always bearing setae.
</paragraph>
<paragraph id="8BED720CFFDFFF83FCCED501D15D4432" blockId="1.[828,1437,190,725]" pageId="1" pageNumber="228">
For detailed description of the genus, see
<bibRefCitation id="EFC30FFDFFDFFF83FAB6D501D1634412" author="SLIPINSKI &amp; TOMASZEWSKA" firstAuthor="SLIPINSKI" journalOrPublisher="Australian Journal of Entomology" pageId="1" pageNumber="228" pagination="369 - 384" part="44" refId="ref7717" refString="SLIPINSKI A. &amp; TOMASZEWSKA W. 2005: Revision of the Australian Coccinellidae (Coleoptera). Part 3. Tribe Sukunahikonini. Australian Journal of Entomology 44: 369 - 384." title="Revision of the Australian Coccinellidae (Coleoptera). Part 3. Tribe Sukunahikonini" type="journal article" year="2005">ŚLIPIŃSKI &amp; TOMASZEWSKA (2005)</bibRefCitation>
,
<bibRefCitation id="EFC30FFDFFDFFF83FBBAD521D0284412" author="ESCALONA &amp; SLIPINSKI" box="[1070,1386,574,597]" firstAuthor="ESCALONA" journalOrPublisher="Systematic Entomology" pageId="1" pageNumber="228" pagination="125 - 171" part="37" refId="ref6785" refString="ESCALONA H. &amp; SLIPINSKI A. 2012: Generic revision and phylogeny of Microweiseinae (Coleoptera: Coccinellidae). Systematic Entomology 37: 125 - 171." title="Generic revision and phylogeny of Microweiseinae (Coleoptera: Coccinellidae)" type="book" year="2012">ESCALONA &amp; ŚLIPIŃSKI (2012)</bibRefCitation>
, and
<bibRefCitation id="EFC30FFDFFDFFF83FCA8D541D1594432" author="WANG &amp; REN" box="[828,1051,606,629]" firstAuthor="WANG" journalOrPublisher="Pan-Pacific Entomologist" pageId="1" pageNumber="228" pagination="287 - 293" part="87" refId="ref7926" refString="WANG X. - M. &amp; REN S. - X. 2012: A new genus Pangia belonging to the tribe Serangiini from China (Coleoptera: Coccinellidae). Pan-Pacific Entomologist 87: 287 - 293." title="A new genus Pangia belonging to the tribe Serangiini from China (Coleoptera: Coccinellidae)" type="journal article" year="2012">WANG &amp; REN (2012)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3482187FFDFFF83FCA8D561D05A4492" pageId="1" pageNumber="228" type="distribution">
<paragraph id="8BED720CFFDFFF83FCA8D561D05A4492" blockId="1.[828,1437,190,725]" pageId="1" pageNumber="228">
<emphasis id="B926AE1EFFDFFF83FCA8D561D68F44D2" bold="true" box="[828,973,638,661]" pageId="1" pageNumber="228">Distribution.</emphasis>
Pantropical:Africa,
<collectingCountry id="F345329CFFDFFF83FB36D561D04444D2" box="[1186,1286,638,661]" name="Australia" pageId="1" pageNumber="228">Australia</collectingCountry>
, Central America,
<collectingCountry id="F345329CFFDFFF83FCE7D581D6F744F2" box="[883,949,670,693]" name="China" pageId="1" pageNumber="228">China</collectingCountry>
,
<collectingCountry id="F345329CFFDFFF83FC56D581D10744F2" box="[962,1093,670,693]" name="Madagascar" pageId="1" pageNumber="228">Madagascar</collectingCountry>
, Oriental Region, New
<collectingCountry id="F345329CFFDFFF83FADDD581D0DB44F2" box="[1353,1433,670,693]" name="Guinea-Bissau" pageId="1" pageNumber="228">Guinea</collectingCountry>
,
<collectingCountry id="F345329CFFDFFF83FCA8D5A1D6974492" box="[828,981,702,725]" name="Vanuatu" pageId="1" pageNumber="228">New Hebrides</collectingCountry>
(
<bibRefCitation id="EFC30FFDFFDFFF83FC77D5A1D04E4492" author="ESCALONA &amp; SLIPINSKI" box="[995,1292,702,725]" firstAuthor="ESCALONA" journalOrPublisher="Systematic Entomology" pageId="1" pageNumber="228" pagination="125 - 171" part="37" refId="ref6785" refString="ESCALONA H. &amp; SLIPINSKI A. 2012: Generic revision and phylogeny of Microweiseinae (Coleoptera: Coccinellidae). Systematic Entomology 37: 125 - 171." title="Generic revision and phylogeny of Microweiseinae (Coleoptera: Coccinellidae)" type="book" year="2012">ESCALONA &amp; ŚLIPIŃSKI 2012</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>