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<document id="529806ECD9B87A3E52956DC7B0F30490" ID-DOI="10.11646/zootaxa.3857.2.5" ID-GBIF-Dataset="4ad4cc3b-5758-4f51-beab-68b53c9d25fd" ID-ISSN="1175-5326" ID-Zenodo-Dep="229668" ID-ZooBank="32D449D9-0E04-4848-A3F9-88FB0FC24025" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460897827241" checkinUser="plazi" docAuthor="García-Romera, Carlos &amp; Barrientos, Jose Antonio" docDate="2014" docId="03FB5521FFDA3D03FF09FB8CEA16669E" docLanguage="en" docName="zt03857p260.pdf" docOrigin="Zootaxa 3857 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Megaselia longianalis Garcia-Romera, sp. nov." docType="treatment" docVersion="7" lastPageNumber="253" masterDocId="FFC22D59FFD23D0AFF9EFFCEEB116008" masterDocTitle="Nine new species of Phoridae (Insecta: Diptera) from Spain" masterLastPageNumber="260" masterPageNumber="244" pageNumber="252" updateTime="1698360198051" updateUser="plazi">
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<mods:title id="BEFD14393D269463CDA82115F0DA6447">Nine new species of Phoridae (Insecta: Diptera) from Spain</mods:title>
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<mods:namePart id="B9D5AA586E2379E010ACF76ADDD7426F">García-Romera, Carlos</mods:namePart>
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<mods:namePart id="2BC19285CB5BDF34E1A64C0EA47DD949">Barrientos, Jose Antonio</mods:namePart>
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<mods:title id="ECD257AA354B6CBAB1833765CAA93B2D">Zootaxa</mods:title>
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<mods:date id="12A677A1EF8137D17FB1305A66E8A6E6">2014</mods:date>
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<treatment id="03FB5521FFDA3D03FF09FB8CEA16669E" ID-DOI="http://doi.org/10.5281/zenodo.6136533" ID-GBIF-Taxon="119587649" ID-Zenodo-Dep="6136533" LSID="urn:lsid:plazi:treatment:03FB5521FFDA3D03FF09FB8CEA16669E" httpUri="http://treatment.plazi.org/id/03FB5521FFDA3D03FF09FB8CEA16669E" lastPageId="9" lastPageNumber="253" pageId="8" pageNumber="252">
<subSubSection id="C348B7BCFFDA3D02FF09FB8CEA376477" pageId="8" pageNumber="252" type="nomenclature">
<paragraph id="8BEDE437FFDA3D02FF09FB8CE9A96453" blockId="8.[151,696,1090,1151]" box="[151,696,1090,1115]" pageId="8" pageNumber="252">
<heading id="D0A5535BFFDA3D02FF09FB8CE9A96453" bold="true" box="[151,696,1090,1115]" fontSize="10" level="2" pageId="8" pageNumber="252" reason="2">
<emphasis id="B9263825FFDA3D02FF09FB8CE9A96453" bold="true" box="[151,696,1090,1115]" pageId="8" pageNumber="252">
<taxonomicName id="4C529FB4FFDA3D02FF09FB8CE9496453" ID-CoL="8CS7B" authority="Garcia-Romera" authorityName="Garcia-Romera" box="[151,600,1090,1115]" class="Insecta" family="Phoridae" genus="Megaselia" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="252" phylum="Arthropoda" rank="species" species="longianalis" status="sp. nov.">
<emphasis id="B9263825FFDA3D02FF09FB8CEA846453" bold="true" box="[151,405,1090,1115]" italics="true" pageId="8" pageNumber="252">Megaselia longianalis</emphasis>
García-Romera
</taxonomicName>
<taxonomicNameLabel id="A215855EFFDA3D02FDC1FB8CE9A96453" box="[607,696,1090,1115]" pageId="8" pageNumber="252" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BEDE437FFDA3D02FF09FBA8EA376477" blockId="8.[151,696,1090,1151]" box="[151,294,1126,1151]" pageId="8" pageNumber="252">
(
<figureCitation id="1369F8B2FFDA3D02FF01FBA8EA0E6477" box="[159,287,1126,1151]" captionStart="FIGURES 12 13" captionStartId="9.[151,264,810,833]" captionTargetBox="[160,1436,190,796]" captionTargetId="figure@9.[156,1441,183,802]" captionTargetPageId="9" captionText="FIGURES 12 13. Megaselia longianalis García-Romera sp. nov. male. 12, left face of hypopygium. 13, right face of hypopygium. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/229674/files/figure.png" pageId="8" pageNumber="252">Figs 1213</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C348B7BCFFDA3D02FF09FB60EE2864CF" box="[151,1337,1198,1223]" pageId="8" pageNumber="252" type="etymology">
<paragraph id="8BEDE437FFDA3D02FF09FB60EE2864CF" blockId="8.[151,1437,1198,2016]" box="[151,1337,1198,1223]" pageId="8" pageNumber="252">
<emphasis id="B9263825FFDA3D02FF09FB60EA0F64CF" bold="true" box="[151,286,1198,1223]" pageId="8" pageNumber="252">Etymology.</emphasis>
The species is named after its clearly longer anal tube in relation to dorsal face of epandrium.
</paragraph>
</subSubSection>
<subSubSection id="C348B7BCFFDA3D02FF59FB1CE8D765CB" pageId="8" pageNumber="252" type="diagnosis">
<paragraph id="8BEDE437FFDA3D02FF59FB1CE8D765CB" blockId="8.[151,1437,1198,2016]" pageId="8" pageNumber="252">
<emphasis id="B9263825FFDA3D02FF59FB1CEA5064E3" bold="true" box="[199,321,1234,1259]" pageId="8" pageNumber="252">Diagnosis.</emphasis>
Palps yellow with longest bristle clearly longer than maximum width of palp; labella enlarged, but without short pale spinules below; mesopleuron with hairs, bristles absent; scutellum with an anterior pair of small hairs and a posterior pair of bristles; left side of epandrium with four long bristles stronger than hairs on cerci and proctiger; anal tube yellow and clearly longer than length of dorsal face of epandrium; posterior lobes of hypandrium vestigial; all fore tarsal segments yellowish, with only metatarsus a little enlarged and without complete longitudinal rows of hairs reduced to short blunt spines; knob of haltere brown; costal index less than 0.44; first costal section less twice longer than second and third together.
</paragraph>
</subSubSection>
<subSubSection id="C348B7BCFFDA3D03FF59FA00EE84660E" lastPageId="9" lastPageNumber="253" pageId="8" pageNumber="252" type="description">
<paragraph id="8BEDE437FFDA3D02FF59FA00E8CE670F" blockId="8.[151,1437,1198,2016]" pageId="8" pageNumber="252">
<emphasis id="B9263825FFDA3D02FF59FA00EABD65EF" bold="true" box="[199,428,1486,1511]" pageId="8" pageNumber="252">Description (male).</emphasis>
Head. Frons light brown, about as broad as long, with 6061 hairs. Upper supra-antennal bristles a little stronger than lower pair (missing in
<typeStatus id="54E95A95FFDA3D02FD42FA3CE82F6603" box="[732,830,1522,1547]" pageId="8" pageNumber="252" type="holotype">holotype</typeStatus>
and
<typeStatus id="54E95A95FFDA3D02FCE9FA3DE8CD6603" box="[887,988,1523,1547]" pageId="8" pageNumber="252" type="paratype">paratype</typeStatus>
), and they are as closer together as preocellars, which are lower on frons than mediolaterals and clearly closer together than either is from a mediolateral bristles. Upper supra-antennal bristles at about the same level on frons as antials, which are slightly lower on frons than anterolaterales, and clearly closer to these than to upper supra-antennal bristles. Gena with three long bristles and four shorter ones. Antennal postpedicel subglobose, brown, diameter
<quantity id="4CAA49D2FFDA3D02FC50F94DEF236692" box="[974,1074,1667,1691]" metricMagnitude="-4" metricUnit="m" metricValue="1.2" pageId="8" pageNumber="252" unit="mm" value="0.12">0.12 mm</quantity>
and without SPS vesicles. Palps yellow,
<quantity id="4CAA49D2FFDA3D02FF6AF969EA4D66B7" box="[244,348,1703,1727]" metricMagnitude="-4" metricUnit="m" metricValue="1.7" pageId="8" pageNumber="252" unit="mm" value="0.17">0.17 mm</quantity>
long and
<quantity id="4CAA49D2FFDA3D02FE47F968E95366B7" box="[473,578,1702,1727]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="252" unit="mm" value="0.05">0.05 mm</quantity>
maximum width, with 78 bristles, the longest bristle being longer than greatest width of palp. Labella enlarged, dusky yellow, without short spinules below, with only a few long hairs. Labrum yellow brown, about 0.80.9 × as wide as an antennal postpedicel.
</paragraph>
<paragraph id="8BEDE437FFDA3D02FF59F8DCE821677A" blockId="8.[151,1437,1198,2016]" pageId="8" pageNumber="252">Thorax. Light brown. Mesopleuron with 10 hairs, bristles absent. Three notopleural bristles and no cleft in front of these. Scutellum with an anterior pair of small hairs and a posterior pair of bristles. Propleura with four hairs on posterodorsal edge and four on posteroventral one.</paragraph>
<paragraph id="8BEDE437FFDA3D02FF59F8B0EF1467E8" blockId="8.[151,1437,1198,2016]" pageId="8" pageNumber="252">Abdomen. Tergites brown, with scattered short hairs, especially on posterior half, with those of the rear margin only a little longer, apart from those of T6, which are clearly longer than the rest and so long as those on cerci. Venter yellow, with hairs on segments 36, longer in the latest three segments.</paragraph>
<caption id="DF2DB4BFFFDB3D03FF09FCE4EAEB6357" httpUri="https://zenodo.org/record/229674/files/figure.png" pageId="9" pageNumber="253" targetBox="[160,1436,190,796]" targetPageId="9">
<paragraph id="8BEDE437FFDB3D03FF09FCE4EAEB6357" blockId="9.[151,1436,810,863]" pageId="9" pageNumber="253">
<emphasis id="B9263825FFDB3D03FF09FCE4EA486348" bold="true" box="[151,345,810,833]" pageId="9" pageNumber="253">FIGURES 1213.</emphasis>
<taxonomicName id="4C529FB4FFDB3D03FEFBFCE5E9E56349" authority="Garcia-Romera" authorityName="Garcia-Romera" box="[357,756,811,833]" class="Insecta" family="Phoridae" genus="Megaselia" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="253" phylum="Arthropoda" rank="species" species="longianalis" status="sp. nov.">
<emphasis id="B9263825FFDB3D03FEFBFCE5E95D6349" box="[357,588,811,833]" italics="true" pageId="9" pageNumber="253">Megaselia longianalis</emphasis>
García-Romera
</taxonomicName>
<emphasis id="B9263825FFDB3D03FC9EFCE5E8466348" bold="true" box="[768,855,811,832]" pageId="9" pageNumber="253">
<taxonomicNameLabel id="A215855EFFDB3D03FC9EFCE5E8466348" box="[768,855,811,832]" pageId="9" pageNumber="253" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
male.
<emphasis id="B9263825FFDB3D03FC38FCE5E8AE6348" bold="true" box="[934,959,811,832]" pageId="9" pageNumber="253">12</emphasis>
, left face of hypopygium.
<emphasis id="B9263825FFDB3D03FB77FCE5EE136348" bold="true" box="[1257,1282,811,832]" pageId="9" pageNumber="253">13</emphasis>
, right face of hypopygium. Scale bars = 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BEDE437FFDB3D03FF59FC48E9CF63EF" blockId="9.[151,1437,902,1686]" pageId="9" pageNumber="253">
<emphasis id="B9263825FFDB3D03FF59FC48EA726397" bold="true" box="[199,355,902,927]" pageId="9" pageNumber="253">Hypopygium</emphasis>
(
<figureCitation id="1369F8B2FFDB3D03FEEFFC48EAE16397" box="[369,496,902,927]" captionStart="FIGURES 12 13" captionStartId="9.[151,264,810,833]" captionTargetBox="[160,1436,190,796]" captionTargetId="figure@9.[156,1441,183,802]" captionTargetPageId="9" captionText="FIGURES 12 13. Megaselia longianalis García-Romera sp. nov. male. 12, left face of hypopygium. 13, right face of hypopygium. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/229674/files/figure.png" pageId="9" pageNumber="253">Figs 1213</figureCitation>
). Left side of epandrium brown, about as high as long, with four long bristles longer and stronger than hairs on cerci and proctiger; anal tube yellow, nearly twice as long as dorsal face of epandrium. Hypandrium yellow brown, without posterior lobes.
</paragraph>
<paragraph id="8BEDE437FFDB3D03FF59FC3AEA2864B7" blockId="9.[151,1437,902,1686]" pageId="9" pageNumber="253">
Legs. Mainly brown, clearing from hind to front legs, with the fore tarse and tibia yellowish. Front legs with a posterodorsal hair palisade on tarsal segments 14 and metatarsus only slightly enlarged. Length ratios of fore tarsal segments 3.2:1.31.6:1.11.2:1:1, with fifth segment as wide as fourth one. Mid legs with a dorsal hair palisade on tibia extends about 2/3 of its length. Longest hairs below basal half of hind femur longer (1.3 ×) than those of anteroventral row of outer third.
<collectingCountry id="F345A4A7FFDB3D03FDFCFB4DE98B6494" box="[610,666,1155,1180]" name="India" pageId="9" pageNumber="253">Hind</collectingCountry>
tibia with 14 differentiated posterodorsal hairs and spinules of apical combs simple.
</paragraph>
<paragraph id="8BEDE437FFDB3D03FF59FB05EE4D6526" blockId="9.[151,1437,902,1686]" pageId="9" pageNumber="253">
Wing. Length
<quantity id="4CAA49D2FFDB3D03FEECFB05EACF64EB" box="[370,478,1227,1251]" metricMagnitude="-3" metricUnit="m" metricValue="1.84" pageId="9" pageNumber="253" unit="mm" value="1.84">1.84 mm</quantity>
. Membrane lightly tinged grey and veins light brown; with 23 axillary bristles. Subcosta (Sc) not reaching R1. With 01 hair at base of vein Rs. Costal index 0.43; costal ratios 3.84:1.7:1. Costal cilia (section 3)
<quantity id="4CAA49D2FFDB3D03FED2FAD8EAA16526" box="[332,432,1302,1326]" metricMagnitude="-4" metricUnit="m" metricValue="1.2" pageId="9" pageNumber="253" unit="mm" value="0.12">0.12 mm</quantity>
long, a little shorter than longest axillary bristle (
<quantity id="4CAA49D2FFDB3D03FC47FAD8EF556526" box="[985,1092,1302,1326]" metricMagnitude="-4" metricUnit="m" metricValue="1.4" pageId="9" pageNumber="253" unit="mm" value="0.14">0.14 mm</quantity>
). Knob of haltere brown.
</paragraph>
<paragraph id="8BEDE437FFDB3D03FF59FAF7EE84660E" blockId="9.[151,1437,902,1686]" pageId="9" pageNumber="253">
<emphasis id="B9263825FFDB3D03FF59FAF7EA4C655A" bold="true" box="[199,349,1337,1362]" pageId="9" pageNumber="253">Recognition.</emphasis>
In the keys to the males of the British species of
<taxonomicName id="4C529FB4FFDB3D03FC08FAF7EFAA655A" authority="Disney, 1989" authorityName="Disney" authorityYear="1989" box="[918,1211,1337,1362]" class="Insecta" family="Phoridae" genus="Megaselia" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="253" phylum="Arthropoda" rank="genus">
<emphasis id="B9263825FFDB3D03FC08FAF7EF1A655A" box="[918,1035,1337,1362]" italics="true" pageId="9" pageNumber="253">Megaselia</emphasis>
(
<bibRefCitation id="EFC399C6FFDB3D03FB85FAF7EFA2655A" author="Disney" box="[1051,1203,1337,1362]" pageId="9" pageNumber="253" refString="Disney, R. H. L. (1989) Scuttle flies. Diptera, Phoridae, genus Megaselia. Handbooks for the Identification of British Insects, 10 (8), 1 - 155." type="journal article" year="1989">Disney, 1989</bibRefCitation>
)
</taxonomicName>
this species runs to couplet 91, after return of 126, where neither option applies; it differs from
<taxonomicName id="4C529FB4FFDB3D03FB96FA91EFE5657E" authority="Disney" authorityName="Disney" box="[1032,1268,1373,1398]" class="Insecta" family="Phoridae" genus="Megaselia" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="253" phylum="Arthropoda" rank="species" species="horsfieldi">
<emphasis id="B9263825FFDB3D03FB96FA91EF8D657E" box="[1032,1180,1373,1398]" italics="true" pageId="9" pageNumber="253">M. horsfieldi</emphasis>
Disney
</taxonomicName>
by having the ventral hair palisade of hind metatarsus without hairs with recurved tips, and from
<taxonomicName id="4C529FB4FFDB3D03FBF5FA4DEE646592" authority="Schmitz" authorityName="Schmitz" box="[1131,1397,1409,1434]" class="Insecta" family="Phoridae" genus="Megaselia" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="253" phylum="Arthropoda" rank="species" species="protarsalis">
<emphasis id="B9263825FFDB3D03FBF5FA4DEE1F6592" box="[1131,1294,1409,1434]" italics="true" pageId="9" pageNumber="253">M. protarsalis</emphasis>
Schmitz
</taxonomicName>
by having all fore tarsal segments the same colour, and from both species in the hypopygium, especially in shape and hairs of epandrium. In the keys of Palaearctic
<taxonomicName id="4C529FB4FFDB3D03FD33FA07E80065EA" box="[685,785,1481,1506]" class="Insecta" family="Phoridae" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="253" phylum="Arthropoda" rank="family">Phoridae</taxonomicName>
(
<bibRefCitation id="EFC399C6FFDB3D03FCBFFA07EF1565EA" author="Schmitz" box="[801,1028,1481,1506]" pageId="9" pageNumber="253" refString="Schmitz, H., Beyer, H. y. &amp; Delage, A. (1938 - 1981) Phoridae. In: Lindner, E. (Ed.), Die Fliegen der palaearktischen Region, 4 (33), pp. 1 - 712. [Stuttgart]" type="journal article" year="1938">
Schmitz
<emphasis id="B9263825FFDB3D03FC16FA05E8D365EA" box="[904,962,1481,1506]" italics="true" pageId="9" pageNumber="253">et al,</emphasis>
<date id="FFECC2F7FFDB3D03FC55FA04EF1565EA" box="[971,1028,1482,1506]" pageId="9" pageNumber="253" value="1938" valueMax="1981">1938</date>
</bibRefCitation>
81) this species runs to couplet 36 (Abteilung V, page 640) to
<taxonomicName id="4C529FB4FFDB3D03FE57FA21E9DD660E" authority="Colyer" authorityName="Colyer" box="[457,716,1517,1542]" class="Insecta" family="Phoridae" genus="Megaselia" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="253" phylum="Arthropoda" rank="species" species="criniticauda">
<emphasis id="B9263825FFDB3D03FE57FA21E966660E" box="[457,631,1517,1542]" italics="true" pageId="9" pageNumber="253">M. criniticauda</emphasis>
Colyer
</taxonomicName>
, which clearly differs from the new species in the hypopygium.
</paragraph>
</subSubSection>
<subSubSection id="C348B7BCFFDB3D03FF59F9DFEA16669E" pageId="9" pageNumber="253" type="materials_examined">
<paragraph id="8BEDE437FFDB3D03FF59F9DFEF606646" blockId="9.[151,1437,902,1686]" pageId="9" pageNumber="253">
<emphasis id="B9263825FFDB3D03FF59F9DFE93B6622" bold="true" box="[199,554,1553,1578]" pageId="9" pageNumber="253">
Material examined.
<typeStatus id="54E95A95FFDB3D03FE23F9DFE93B6622" box="[445,554,1553,1578]" pageId="9" pageNumber="253" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="F345A4A7FFDB3D03FDCDF9DFE9876622" box="[595,662,1553,1578]" name="Spain" pageId="9" pageNumber="253">Spain</collectingCountry>
, Barcelona: Montseny, Montseny Natural Park, beech forest, UTM 31TDG530274, 1130 m a.s.l.,
<date id="FFECC2F7FFDB3D03FE75F9F8E9896646" box="[491,664,1590,1614]" pageId="9" pageNumber="253" value="1990-07-01" valueMax="1990-07-14" valueMin="1990-07-01">0114/07/1990</date>
. Window trap. J.A. Barrientos leg (
<collectionCode id="ED437CF2FFDB3D03FBBAF9F8EF746646" box="[1060,1125,1590,1614]" httpUri="http://grbio.org/cool/0md6-ynwr" name="Universidad Autonoma de Barcelona" pageId="9" pageNumber="253">UAB</collectionCode>
).
</paragraph>
<paragraph id="8BEDE437FFDB3D03FF59F997EA16669E" blockId="9.[151,1437,902,1686]" pageId="9" pageNumber="253">
<emphasis id="B9263825FFDB3D03FF59F997EA25667A" bold="true" box="[199,308,1625,1650]" pageId="9" pageNumber="253">
<typeStatus id="54E95A95FFDB3D03FF59F997EA25667A" box="[199,308,1625,1650]" pageId="9" pageNumber="253" type="paratype">Paratype</typeStatus>
</emphasis>
♂, same locality and habitat as
<typeStatus id="54E95A95FFDB3D03FD35F997E801667A" box="[683,784,1625,1650]" pageId="9" pageNumber="253" type="holotype">holotype</typeStatus>
, but
<date id="FFECC2F7FFDB3D03FCD6F994E8E3667A" box="[840,1010,1626,1650]" pageId="9" pageNumber="253" value="1990-07-14" valueMax="1990-07-28" valueMin="1990-07-14">1428/07/1990</date>
and window trap. J.A. Barrientos leg (CCG-R).
</paragraph>
</subSubSection>
</treatment>
</document>