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<mods:title id="CC27A1818956E778BD8C6BE6242A5457">Redescription of Paraleptophlebia falcula Traver 1934 with notes on status and composition of Paraleptophlebia Lestage 1917 and Neoleptophlebia Kluge 1997 (Ephemeroptera: Leptophlebiidae)</mods:title>
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<treatment id="03E08C6BFFB5FFD874E6FDC632FE2A29" ID-DOI="http://doi.org/10.5281/zenodo.6075462" ID-GBIF-Taxon="119375080" ID-Zenodo-Dep="6075462" LSID="urn:lsid:plazi:treatment:03E08C6BFFB5FFD874E6FDC632FE2A29" httpUri="http://treatment.plazi.org/id/03E08C6BFFB5FFD874E6FDC632FE2A29" lastPageNumber="372" pageId="3" pageNumber="372">
<subSubSection id="C3536EF6FFB5FFD874E6FDC6314F2CDF" box="[151,614,619,646]" pageId="3" pageNumber="372" type="nomenclature">
<paragraph id="8BF63D7DFFB5FFD874E6FDC6314F2CDF" blockId="3.[151,614,619,646]" box="[151,614,619,646]" pageId="3" pageNumber="372">
<heading id="D0BE8A11FFB5FFD874E6FDC6314F2CDF" bold="true" box="[151,614,619,646]" fontSize="11" level="1" pageId="3" pageNumber="372" reason="1">
<emphasis id="B93DE16FFFB5FFD874E6FDC6314F2CDF" bold="true" box="[151,614,619,646]" pageId="3" pageNumber="372">
Genus
<taxonomicName id="4C4946FEFFB5FFD8749FFDC1314F2CDF" ID-CoL="6F8B" ID-ENA="202397" authority="Lestage 1917" authorityName="Lestage" authorityYear="1917" box="[238,614,619,646]" class="Insecta" family="Leptophlebiidae" genus="Paraleptophlebia" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="genus">
<emphasis id="B93DE16FFFB5FFD8749FFDC132942CDC" bold="true" box="[238,445,619,645]" italics="true" pageId="3" pageNumber="372">Paraleptophlebia</emphasis>
<bibRefCitation id="EFD8408CFFB5FFD875B5FDC6314F2CDF" author="Lestage" box="[452,614,620,646]" pageId="3" pageNumber="372" refString="Lestage, J. - A. (1917) Contribution a l'etude des larves des Ephemeres palearctiques. Annales de Biologie Lacustre, 8 (3 - 4), 213 - 459." type="journal article" year="1917">Lestage 1917</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3536EF6FFB5FFD874E6FD1930E32CBE" pageId="3" pageNumber="372" type="materials_examined">
<paragraph id="8BF63D7DFFB5FFD874E6FD19314D2C91" blockId="3.[151,970,690,743]" box="[151,612,690,712]" pageId="3" pageNumber="372">
<typeStatus id="54F283DFFFB5FFD874E6FD1933E02C91" box="[151,201,691,712]" pageId="3" pageNumber="372">Type</typeStatus>
species:
<taxonomicName id="4C4946FEFFB5FFD87557FD1831482C91" authority="Retzius 1783" authorityName="Retzius" authorityYear="1783" box="[294,609,690,712]" class="Insecta" family="Ephemeridae" genus="Ephemera" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="species" species="cincta">
<emphasis id="B93DE16FFFB5FFD87557FD1832FB2C91" box="[294,466,690,712]" italics="true" pageId="3" pageNumber="372">Ephemera cincta</emphasis>
<bibRefCitation id="EFD8408CFFB5FFD875A9FD1831482C91" author="Retzius" box="[472,609,690,712]" pageId="3" pageNumber="372" refString="Retzius, A. J. (1783) Caroli de Geer. Genera et species insectorum e generosissimi auctoris scriptis. Apud Siegfried Lebrecht Crusium, Lipsiae, 220 pp." type="book" year="1783">Retzius 1783</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="8BF63D7DFFB5FFD874E6FD7830E32CBE" blockId="3.[151,970,690,743]" box="[151,970,721,743]" pageId="3" pageNumber="372">
In circumscription corresponds to subgenus
<taxonomicName id="4C4946FEFFB5FFD87627FD7B30EF2CBE" authority="Kluge 1997" authorityName="Kluge" authorityYear="1997" box="[598,966,721,743]" class="Insecta" family="Leptophlebiidae" genus="Paraleptophlebia" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="genus">
<emphasis id="B93DE16FFFB5FFD87627FD7B302E2CBE" box="[598,775,721,743]" italics="true" pageId="3" pageNumber="372">Paraleptophlebia</emphasis>
sensu
<bibRefCitation id="EFD8408CFFB5FFD8773CFD7B30EF2CBE" author="Kluge" box="[845,966,721,743]" pageId="3" pageNumber="372" refString="Kluge, N. J. (1997) New subgenera of Holarctic mayflies (Ephemeroptera: Heptageniidae, Leptophlebiidae, Ephemerellidae). Zoosystematica Rossica, 5 (2), 233 - 235. [1996]" type="journal article" year="1997">Kluge 1997</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3536EF6FFB5FFD874E6FCBE30F02DE4" pageId="3" pageNumber="372" type="diagnosis">
<paragraph id="8BF63D7DFFB5FFD874E6FCBE37082D09" blockId="3.[151,1436,788,1136]" pageId="3" pageNumber="372">
<emphasis id="B93DE16FFFB5FFD874E6FCBE32B92D74" bold="true" box="[151,400,788,813]" pageId="3" pageNumber="372">
Diagnosis.
<emphasis id="B93DE16FFFB5FFD87569FCBE324A2D72" bold="true" box="[280,355,788,811]" italics="true" pageId="3" pageNumber="372">Larva.</emphasis>
(1)
</emphasis>
Third segment of maxillary palp widest in middle part (
<figureCitation id="137221F8FFB5FFD87072FCBE376C2D75" box="[1027,1093,788,813]" captionStart="FIGURES 1 13" captionStartId="5.[151,264,1920,1943]" captionTargetBox="[191,1398,196,1901]" captionTargetId="figure@5.[187,1402,180,1912]" captionTargetPageId="5" captionText="FIGURES 1 13. Paraleptophlebia falcula, larvae. 1, labrum, dorsal view; 2, 3, left and right mandibles, dorsal view; 4, 5, incisors and kinetodonts of left and right mandibles; 6, maxillary palp, ventral view; 7, labium, dorsal view; 8, segments 2 and 3 of labial palp; 9, hypopharynx and superlinguae, dorsal view; 10, femur of foreleg; 11, foreclaw, dorsal view; 12, posterior margin of abdominal tergum IV; 13, abdominal sternum IX, female." httpUri="https://zenodo.org/record/266228/files/figure.png" pageId="3" pageNumber="372">Fig. 8</figureCitation>
) (the same in
<taxonomicName id="4C4946FEFFB5FFD87095FCBE36512D75" box="[1252,1400,788,812]" class="Insecta" family="Leptophlebiidae" genus="Leptophlebia" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="genus">
<emphasis id="B93DE16FFFB5FFD87095FCBE36512D75" box="[1252,1400,788,812]" italics="true" pageId="3" pageNumber="372">Leptophlebia</emphasis>
</taxonomicName>
; in contrast to
<taxonomicName id="4C4946FEFFB5FFD87562FC9232E52D09" box="[275,460,824,848]" class="Insecta" family="Leptophlebiidae" genus="Neoleptophlebia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="genus">
<emphasis id="B93DE16FFFB5FFD87562FC9232E52D09" box="[275,460,824,848]" italics="true" pageId="3" pageNumber="372">Neoleptophlebia</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4946FEFFB5FFD875ABFC9231B22D09" box="[474,667,824,848]" class="Insecta" family="Leptophlebiidae" genus="Habrophlebiodes" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="genus">
<emphasis id="B93DE16FFFB5FFD875ABFC9231B22D09" box="[474,667,824,848]" italics="true" pageId="3" pageNumber="372">Habrophlebiodes</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4946FEFFB5FFD876DBFC9230502D09" box="[682,889,824,848]" class="Insecta" family="Leptophlebiidae" genus="Dipterophlebiodes" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="genus">
<emphasis id="B93DE16FFFB5FFD876DBFC9230502D09" box="[682,889,824,848]" italics="true" pageId="3" pageNumber="372">Dipterophlebiodes</emphasis>
</taxonomicName>
and
<emphasis id="B93DE16FFFB5FFD877DEFC9237082D09" box="[943,1057,824,848]" italics="true" pageId="3" pageNumber="372">
<taxonomicName id="4C4946FEFFB5FFD877DEFC92373D2D09" box="[943,1044,824,848]" class="Insecta" family="Leptophlebiidae" genus="Gilliesia" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="genus">Gilliesia</taxonomicName>
).
</emphasis>
</paragraph>
<paragraph id="8BF63D7DFFB5FFD874B6FCF630F02DE4" blockId="3.[151,1436,788,1136]" pageId="3" pageNumber="372">
<emphasis id="B93DE16FFFB5FFD874B6FCF633C12D2C" bold="true" box="[199,232,860,885]" pageId="3" pageNumber="372">(2)</emphasis>
Tergalii V-shape, i.e. with two slender lobes separated nearly up to base (
<figureCitation id="137221F8FFB5FFD8702FFCF637CA2D2D" box="[1118,1251,860,885]" captionStart-0="FIGURES 14 15" captionStart-1="FIGURES 16 22" captionStartId-0="6.[151,264,1121,1144]" captionStartId-1="6.[151,264,1988,2011]" captionTargetBox-0="[266,1320,193,1100]" captionTargetBox-1="[199,1380,1197,1952]" captionTargetId-0="figure@6.[266,1321,193,1100]" captionTargetId-1="figure@6.[193,1390,1193,1972]" captionTargetPageId-0="6" captionTargetPageId-1="6" captionText-0="FIGURES 14 15. Paraleptophlebia falcula, larvae, habitus. 14, dorsal view; 15, ventral view." captionText-1="FIGURES 16 22. Paraleptophlebia falcula, larvae, tergalii of I, II, III, IV, V, VI and VII pairs." httpUri-0="https://zenodo.org/record/266229/files/figure.png" httpUri-1="https://zenodo.org/record/266230/files/figure.png" pageId="3" pageNumber="372">Figs 1422</figureCitation>
) (in contrast to
<taxonomicName id="4C4946FEFFB5FFD874E6FC2A32022DC1" box="[151,299,896,920]" class="Insecta" family="Leptophlebiidae" genus="Leptophlebia" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="genus">
<emphasis id="B93DE16FFFB5FFD874E6FC2A32022DC1" box="[151,299,896,920]" italics="true" pageId="3" pageNumber="372">Leptophlebia</emphasis>
</taxonomicName>
, whose tergalii IIVII have lobes widened; in contrast to
<taxonomicName id="4C4946FEFFB5FFD87075FC2A37942DC1" box="[1028,1213,896,920]" class="Insecta" family="Leptophlebiidae" genus="Neoleptophlebia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="genus">
<emphasis id="B93DE16FFFB5FFD87075FC2A37942DC1" box="[1028,1213,896,920]" italics="true" pageId="3" pageNumber="372">Neoleptophlebia</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4946FEFFB5FFD870A5FC2A36BC2DC1" box="[1236,1429,896,920]" class="Insecta" family="Leptophlebiidae" genus="Habrophlebiodes" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="genus">
<emphasis id="B93DE16FFFB5FFD870A5FC2A36BC2DC1" box="[1236,1429,896,920]" italics="true" pageId="3" pageNumber="372">Habrophlebiodes</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4946FEFFB5FFD874E6FC0E324F2DE5" box="[151,358,932,956]" class="Insecta" family="Leptophlebiidae" genus="Dipterophlebiodes" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="genus">
<emphasis id="B93DE16FFFB5FFD874E6FC0E324F2DE5" box="[151,358,932,956]" italics="true" pageId="3" pageNumber="372">Dipterophlebiodes</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4946FEFFB5FFD875EDFC0E32D42DE5" box="[412,509,932,956]" class="Insecta" family="Leptophlebiidae" genus="Gilliesia" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="genus">
<emphasis id="B93DE16FFFB5FFD875EDFC0E32D42DE5" box="[412,509,932,956]" italics="true" pageId="3" pageNumber="372">Gilliesia</emphasis>
</taxonomicName>
, whose tergalii have lobes less separated).
</paragraph>
</subSubSection>
<subSubSection id="C3536EF6FFB5FFD874B6FC6D30C12A14" pageId="3" pageNumber="372" type="description">
<paragraph id="8BF63D7DFFB5FFD874B6FC6D36B12A5D" blockId="3.[151,1436,788,1136]" pageId="3" pageNumber="372">
<emphasis id="B93DE16FFFB5FFD874B6FC6D32562DB9" bold="true" box="[199,383,967,992]" pageId="3" pageNumber="372">
<emphasis id="B93DE16FFFB5FFD874B6FC6D32782DB9" bold="true" box="[199,337,967,992]" italics="true" pageId="3" pageNumber="372">Male imago</emphasis>
. (3)
</emphasis>
<collectingCountry id="F35E7DEDFFB5FFD875F7FC6D32972DB9" box="[390,446,967,992]" name="India" pageId="3" pageNumber="372">Hind</collectingCountry>
wing fully developed, without costal projection, with Sc terminating near apex (
<figureCitation id="137221F8FFB5FFD87133FC6D36BD2DB9" box="[1346,1428,967,992]" captionStart="FIGURES 23 25" captionStartId="7.[151,264,1805,1828]" captionTargetBox="[151,1436,193,1783]" captionTargetId="figure@7.[151,1436,193,1785]" captionTargetPageId="7" captionText="FIGURES 23 25. Paraleptophlebia falcula, male imago. 23, lateral view; 24, dorsal view; 25, ventral view." httpUri="https://zenodo.org/record/266231/files/figure.png" pageId="3" pageNumber="372">Fig. 23</figureCitation>
) (the same in
<taxonomicName id="4C4946FEFFB5FFD87556FC4632922A5D" box="[295,443,1004,1028]" class="Insecta" family="Leptophlebiidae" genus="Leptophlebia" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="genus">
<emphasis id="B93DE16FFFB5FFD87556FC4632922A5D" box="[295,443,1004,1028]" italics="true" pageId="3" pageNumber="372">Leptophlebia</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4946FEFFB5FFD87580FC4631832A5D" box="[497,682,1004,1028]" class="Insecta" family="Leptophlebiidae" genus="Neoleptophlebia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="genus">
<emphasis id="B93DE16FFFB5FFD87580FC4631832A5D" box="[497,682,1004,1028]" italics="true" pageId="3" pageNumber="372">Neoleptophlebia</emphasis>
</taxonomicName>
; in contrast to
<taxonomicName id="4C4946FEFFB5FFD87723FC46373A2A5D" box="[850,1043,1004,1028]" class="Insecta" family="Leptophlebiidae" genus="Habrophlebiodes" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="genus">
<emphasis id="B93DE16FFFB5FFD87723FC46373A2A5D" box="[850,1043,1004,1028]" italics="true" pageId="3" pageNumber="372">Habrophlebiodes</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4946FEFFB5FFD87050FC4637D92A5D" box="[1057,1264,1004,1028]" class="Insecta" family="Leptophlebiidae" genus="Dipterophlebiodes" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="genus">
<emphasis id="B93DE16FFFB5FFD87050FC4637D92A5D" box="[1057,1264,1004,1028]" italics="true" pageId="3" pageNumber="372">Dipterophlebiodes</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4946FEFFB5FFD87156FC4636AE2A5D" box="[1319,1415,1004,1028]" class="Insecta" family="Leptophlebiidae" genus="Gilliesia" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="genus">
<emphasis id="B93DE16FFFB5FFD87156FC4636AE2A5D" box="[1319,1415,1004,1028]" italics="true" pageId="3" pageNumber="372">Gilliesia</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8BF63D7DFFB5FFD874B6FBA530C12A14" blockId="3.[151,1436,788,1136]" pageId="3" pageNumber="372">
<emphasis id="B93DE16FFFB5FFD874B6FBA533C12A71" bold="true" box="[199,232,1039,1064]" pageId="3" pageNumber="372">(4)</emphasis>
Penis deeply divided, with a pair of ventral appendages arising from apex of each penis lobe and directed proximally (
<figureCitation id="137221F8FFB5FFD87550FB9E32882A14" box="[289,417,1076,1101]" captionStart="FIGURES 26 30" captionStartId="8.[151,264,1748,1771]" captionTargetBox="[172,1411,195,1716]" captionTargetId="figure@8.[170,1417,193,1728]" captionTargetPageId="8" captionText="FIGURES 26 30. Paraleptophlebia falcula, genitals of male imago. 26 28, specimens from Transbaikal Kray; 29, 30, specimen from Komi Republic. 26, 29, ventral view; 27, penis, ventral view; 28, penis, dorsal view, 30, lateral view." httpUri="https://zenodo.org/record/266232/files/figure.png" pageId="3" pageNumber="372">Figs 2630</figureCitation>
) (similar to
<taxonomicName id="4C4946FEFFB5FFD8765BFB9E31972A15" box="[554,702,1076,1100]" class="Insecta" family="Leptophlebiidae" genus="Leptophlebia" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="genus">
<emphasis id="B93DE16FFFB5FFD8765BFB9E31972A15" box="[554,702,1076,1100]" italics="true" pageId="3" pageNumber="372">Leptophlebia</emphasis>
</taxonomicName>
, possibly synapomorphy).
</paragraph>
</subSubSection>
<subSubSection id="C3536EF6FFB5FFD874B6FBFD32FE2A29" box="[199,471,1111,1136]" pageId="3" pageNumber="372" type="distribution">
<paragraph id="8BF63D7DFFB5FFD874B6FBFD32FE2A29" blockId="3.[151,1436,788,1136]" box="[199,471,1111,1136]" pageId="3" pageNumber="372">
<emphasis id="B93DE16FFFB5FFD874B6FBFD32492A29" bold="true" box="[199,352,1111,1136]" pageId="3" pageNumber="372">Distribution.</emphasis>
Holarctic.
</paragraph>
</subSubSection>
</treatment>
</document>