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<document id="7F95606DDEB84B756838187880B51F4A" ID-DOI="10.5281/zenodo.3708186" ID-GBIF-Dataset="f503e20e-e3fb-41ac-bf17-bdaf221bc7c7" ID-ISSN="1942-1354" ID-Zenodo-Dep="3708186" ID-ZooBank="55CCB217-771C-499D-9110-36F143C375C5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1584033498187" checkinUser="felipe" docAuthor="Edmonds, W. D." docDate="2018" docId="03E087E4FFFDFF9CFF249C9FFDA9F979" docLanguage="en" docName="InsectaMundi.0642.1-30.pdf.imf" docOrigin="Insecta Mundi 642" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Copris arizonensis Schaeffer 1906" docType="treatment" docVersion="5" lastPageNumber="13" masterDocId="FFD9FF9CFFF3FF92FFE49E74FFB1FF91" masterDocTitle="The dung beetle fauna of the Big Bend region of Texas (Coleoptera: Scarabaeidae: Scarabaeinae)" masterLastPageNumber="30" masterPageNumber="1" pageNumber="13" updateTime="1698727366089" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
<mods:mods id="170099E45439127CBEF48E6F112EBE3B" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="56F35973CBF19ED2B5A7A0B0EE5AEC48">
<mods:title id="333F1808914A3B8AA7B4327C2FF82421">The dung beetle fauna of the Big Bend region of Texas (Coleoptera: Scarabaeidae: Scarabaeinae)</mods:title>
</mods:titleInfo>
<mods:name id="5F5FB473D14C93F1DE08453D111A00B8" type="personal">
<mods:role id="C594AA9C807B52B3A899D6AC4B00C4D8">
<mods:roleTerm id="E549091EEB3B621C39D7070C8D839407">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="ABE88A40F11202019FE218859EDCEA9C">Edmonds, W. D.</mods:namePart>
</mods:name>
<mods:typeOfResource id="D68F2F66F8210C58268236DD4DEB0B0A">text</mods:typeOfResource>
<mods:relatedItem id="8E45C0561BAF2DAE0438DD19EB5C06B7" type="host">
<mods:titleInfo id="BBA19A01F5A48AEC73EA1CBE6F2DC0F9">
<mods:title id="820C4A9DA8454A1543F100BAC996CAFA">Insecta Mundi</mods:title>
</mods:titleInfo>
<mods:part id="6E07934E7A97B7FE729CC4371F3F3FAA">
<mods:date id="251C89A36F315125A7739BD3DA46D6FB">2018</mods:date>
<mods:detail id="D16BBD9FBA73C82237B35E0F8A994010" type="pubDate">
<mods:number id="15F37B22E25A405DEBE16DE1897F7219">2018-07-27</mods:number>
</mods:detail>
<mods:detail id="140E77E19F6A57FDEB77E8C035760C04" type="issue">
<mods:number id="EDBC813F4B7048480AEB3F9864B20B42">642</mods:number>
</mods:detail>
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<mods:start id="2157B7AC4C361683E4D94D214008777C">1</mods:start>
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</mods:extent>
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<mods:classification id="6E4E17B04DE7E33BC1CD0988A8152837">journal article</mods:classification>
<mods:identifier id="534EEA1CB7192B3723CFDAD272D30170" type="DOI">10.5281/zenodo.3708186</mods:identifier>
<mods:identifier id="B8C7EC13CCAA2B3BDF2FCC435CD6D25C" type="GBIF-Dataset">f503e20e-e3fb-41ac-bf17-bdaf221bc7c7</mods:identifier>
<mods:identifier id="CAF3E36F614662859F63BA376EB7087C" type="ISSN">1942-1354</mods:identifier>
<mods:identifier id="7F6555EBAC1A3F7C7E5B39E6D3093F28" type="Zenodo-Dep">3708186</mods:identifier>
<mods:identifier id="37FD49AC9AE5CAE4F54A85F958CE8E0F" type="ZooBank">55CCB217-771C-499D-9110-36F143C375C5</mods:identifier>
</mods:mods>
<treatment id="03E087E4FFFDFF9CFF249C9FFDA9F979" ID-DOI="http://doi.org/10.5281/zenodo.3717146" ID-GBIF-Taxon="162485165" ID-Zenodo-Dep="3717146" LSID="urn:lsid:plazi:treatment:03E087E4FFFDFF9CFF249C9FFDA9F979" httpUri="http://treatment.plazi.org/id/03E087E4FFFDFF9CFF249C9FFDA9F979" lastPageNumber="13" pageId="14" pageNumber="13">
<subSubSection id="C3536579FFFDFF9CFF249C9FFDF8FC92" box="[192,585,747,771]" pageId="14" pageNumber="13" type="nomenclature">
<paragraph id="8BF636F2FFFDFF9CFF249C9FFDF8FC92" blockId="14.[192,585,747,804]" box="[192,585,747,771]" pageId="14" pageNumber="13">
<heading id="D0BE819EFFFDFF9CFF249C9FFDF8FC92" bold="true" box="[192,585,747,771]" fontSize="10" level="2" pageId="14" pageNumber="13" reason="3">
<taxonomicName id="4C494D71FFFDFF9CFF249C9FFDF8FC92" ID-CoL="7ZMF9" authority="Schaeffer" authorityName="Schaeffer" authorityYear="1906" box="[192,585,747,771]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="arizonensis">
<emphasis id="B93DEAE0FFFDFF9CFF249C9FFDF8FC92" bold="true" box="[192,585,747,771]" pageId="14" pageNumber="13">
<emphasis id="B93DEAE0FFFDFF9CFF249C9FFE0FFC92" bold="true" box="[192,446,747,771]" italics="true" pageId="14" pageNumber="13">Copris arizonensis</emphasis>
Schaeffer
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3536579FFFDFF9CFF249D78FEF0FCB5" box="[192,321,780,804]" pageId="14" pageNumber="13" type="description">
<paragraph id="8BF636F2FFFDFF9CFF249D78FEF0FCB5" blockId="14.[192,585,747,804]" box="[192,321,780,804]" pageId="14" pageNumber="13">
<figureCitation id="13722A77FFFDFF9CFF249D78FEF0FCB5" box="[192,321,780,804]" captionStart="Figures 60–69" captionStartId="30.[192,286,1790,1812]" captionTargetBox="[253,1390,282,1714]" captionTargetId="figure@30.[227,1407,249,1724]" captionTargetPageId="30" captionText="Figures 60–69. Euoniticellus and Copris. 60) E. intermedius, dorsal view, female. 61) Same, lateral view of forebody. 62) Same, dorsal view, male. 63) Same, lateral view of forebody. 64) C. arizonensis, dorsal view of forebody, large male (southwestern Arizona). 65) Same, dorsal view, small male (Big Bend). 66) Same, lateral view of forebody. 67) Same, lateral view of forebody, female. 68) Same, oblique dorsal view. 69) Same, dorsal view of head and prothoracic margin." figureDoi="http://doi.org/10.5281/zenodo.3708204" httpUri="https://zenodo.org/record/3708204/files/figure.png" pageId="14" pageNumber="13">Fig. 64–69</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3536579FFFDFF9CFF249D48FD92FBA4" pageId="14" pageNumber="13" type="diagnosis">
<paragraph id="8BF636F2FFFDFF9CFF249D48FD92FBA4" blockId="14.[192,1440,828,1077]" pageId="14" pageNumber="13">
<emphasis id="B93DEAE0FFFDFF9CFF249D48FEFEFCC5" bold="true" box="[192,335,828,852]" pageId="14" pageNumber="13">Diagnosis.</emphasis>
Black, surface lustrous, sides of body more-or-less parallel. Length
<quantity id="4CB19B17FFFDFF9CFB769D48FAA6FCC5" box="[1170,1303,828,852]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" metricValueMax="2.2" metricValueMin="1.3" pageId="14" pageNumber="13" unit="mm" value="17.5" valueMax="22.0" valueMin="13.0">13–22 mm</quantity>
. Male (
<figureCitation id="13722A77FFFDFF9CFA949D48FEBBFCE5" captionStart="Figures 60–69" captionStartId="30.[192,286,1790,1812]" captionTargetBox="[253,1390,282,1714]" captionTargetId="figure@30.[227,1407,249,1724]" captionTargetPageId="30" captionText="Figures 60–69. Euoniticellus and Copris. 60) E. intermedius, dorsal view, female. 61) Same, lateral view of forebody. 62) Same, dorsal view, male. 63) Same, lateral view of forebody. 64) C. arizonensis, dorsal view of forebody, large male (southwestern Arizona). 65) Same, dorsal view, small male (Big Bend). 66) Same, lateral view of forebody. 67) Same, lateral view of forebody, female. 68) Same, oblique dorsal view. 69) Same, dorsal view of head and prothoracic margin." figureDoi="http://doi.org/10.5281/zenodo.3708204" httpUri="https://zenodo.org/record/3708204/files/figure.png" pageId="14" pageNumber="13">Fig. 64–66</figureCitation>
) – Head of large individuals bearing posteriorly curved horn, tip of which approaches bifurcated median process of pronotum; flanks of pronotum each with large, acute, forward-directed, blade-like process separated from apically bifurcated median process by broad, declivitous concavity (in smaller individuals horn and processes progressively attenuated,
<figureCitation id="13722A77FFFDFF9CFC709DC9FC5FFC45" box="[916,1006,956,981]" captionStart="Figures 60–69" captionStartId="30.[192,286,1790,1812]" captionTargetBox="[253,1390,282,1714]" captionTargetId="figure@30.[227,1407,249,1724]" captionTargetPageId="30" captionText="Figures 60–69. Euoniticellus and Copris. 60) E. intermedius, dorsal view, female. 61) Same, lateral view of forebody. 62) Same, dorsal view, male. 63) Same, lateral view of forebody. 64) C. arizonensis, dorsal view of forebody, large male (southwestern Arizona). 65) Same, dorsal view, small male (Big Bend). 66) Same, lateral view of forebody. 67) Same, lateral view of forebody, female. 68) Same, oblique dorsal view. 69) Same, dorsal view of head and prothoracic margin." figureDoi="http://doi.org/10.5281/zenodo.3708204" httpUri="https://zenodo.org/record/3708204/files/figure.png" pageId="14" pageNumber="13">Fig. 65</figureCitation>
). Female
<figureCitation id="13722A77FFFDFF9CFB889DC9FB41FC44" box="[1132,1264,957,981]" captionStart="Figures 60–69" captionStartId="30.[192,286,1790,1812]" captionTargetBox="[253,1390,282,1714]" captionTargetId="figure@30.[227,1407,249,1724]" captionTargetPageId="30" captionText="Figures 60–69. Euoniticellus and Copris. 60) E. intermedius, dorsal view, female. 61) Same, lateral view of forebody. 62) Same, dorsal view, male. 63) Same, lateral view of forebody. 64) C. arizonensis, dorsal view of forebody, large male (southwestern Arizona). 65) Same, dorsal view, small male (Big Bend). 66) Same, lateral view of forebody. 67) Same, lateral view of forebody, female. 68) Same, oblique dorsal view. 69) Same, dorsal view of head and prothoracic margin." figureDoi="http://doi.org/10.5281/zenodo.3708204" httpUri="https://zenodo.org/record/3708204/files/figure.png" pageId="14" pageNumber="13">Fig. 67–69</figureCitation>
) – Head with short, erect horn, apex widened, scoop-like; pronotum convex except for pair of blunt tubercles, one on each side near anterior angle and separated by thick transverse ridge.
<bibRefCitation id="EFD84B03FFFDFF9CFBF99D89FB5DFB84" author="Matthews, E. G." box="[1053,1260,1021,1045]" pageId="14" pageNumber="13" pagination="1 - 139" refId="ref18649" refString="Matthews, E. G. 1961. A revision of the genus Copris Muller of the western hemisphere. Entomologica Americana 41: 1 - 139." type="journal article" year="1961">Matthews (1961)</bibRefCitation>
provides a full re-description of this species.
</paragraph>
</subSubSection>
<subSubSection id="C3536579FFFDFF9CFF249A39FB98FA97" pageId="14" pageNumber="13" type="materials_examined">
<paragraph id="8BF636F2FFFDFF9CFF249A39FC06FBF4" blockId="14.[192,1440,1101,1286]" box="[192,951,1101,1126]" pageId="14" pageNumber="13">
<emphasis id="B93DEAE0FFFDFF9CFF249A39FDA2FBF4" bold="true" box="[192,531,1101,1125]" pageId="14" pageNumber="13">Big Bend collection sites</emphasis>
(altitudinal range:
<quantity id="4CB19B17FFFDFF9CFCE79A39FC1BFBF4" box="[771,938,1101,1125]" metricMagnitude="3" metricUnit="m" metricValue="1.5875" metricValueMax="1.8" metricValueMin="1.375" pageId="14" pageNumber="13" unit="m" value="1587.5" valueMax="1800.0" valueMin="1375.0">1375–1800 m</quantity>
).
</paragraph>
<paragraph id="8BF636F2FFFDFF9CFF0C9A1AFD12FB57" blockId="14.[192,1440,1101,1286]" pageId="14" pageNumber="13">
<materialsCitation id="3B213CAFFFFDFF9CFF0C9A1AFADEFB14" ID-GBIF-Occurrence="2575871655" box="[232,1391,1132,1158]" county="Brewster Co." latitude="30.367779" location="Sul Ross State University" longLatPrecision="20" longitude="-103.64777" municipality="Alpine" pageId="14" pageNumber="13" specimenCount="1">
<collectingCounty id="62974E7EFFFDFF9CFF0C9A1AFE3DFB17" box="[232,396,1134,1158]" pageId="14" pageNumber="13">Brewster Co.</collectingCounty>
:
<emphasis id="B93DEAE0FFFDFF9CFE7C9A19FE0AFB14" bold="true" box="[408,443,1133,1157]" pageId="14" pageNumber="13">[1]</emphasis>
<collectingMunicipality id="6B92AC88FFFDFF9CFE279A19FDA7FB14" box="[451,534,1133,1157]" pageId="14" pageNumber="13">Alpine</collectingMunicipality>
,
<location id="8E966029FFFDFF9CFDC79A19FCD3FB14" LSID="urn:lsid:plazi:treatment:03E087E4FFFDFF9CFF249C9FFDA9F979:8E966029FFFDFF9CFDC79A19FCD3FB14" box="[547,866,1133,1158]" county="Brewster Co." latitude="30.367779" longLatPrecision="20" longitude="-103.64777" municipality="Alpine" name="Sul Ross State University" pageId="14" pageNumber="13">Sul Ross State University</location>
campus,
<geoCoordinate id="EE7D5035FFFDFF9CFC339A1AFBEEFB14" box="[983,1119,1132,1158]" degrees="30" direction="north" minutes="22" orientation="latitude" pageId="14" pageNumber="13" precision="15" seconds="04" value="30.367779">30°22′04″N</geoCoordinate>
<geoCoordinate id="EE7D5035FFFDFF9CFB839A1AFAB7FB14" box="[1127,1286,1132,1158]" degrees="103" direction="west" minutes="38" orientation="longitude" pageId="14" pageNumber="13" precision="15" seconds="52" value="-103.64777">103°38′52″W</geoCoordinate>
,
<quantity id="4CB19B17FFFDFF9CFAF69A19FADEFB14" box="[1298,1391,1133,1157]" metricMagnitude="3" metricUnit="m" metricValue="1.375" pageId="14" pageNumber="13" unit="m" value="1375.0">1375 m</quantity>
</materialsCitation>
;
<materialsCitation id="3B213CAFFFFDFF9CFA989A19FBDBFB37" ID-GBIF-Occurrence="2575871674" elevation="1660" latitude="29.267778" location="Big Bend National Park" longLatPrecision="20" longitude="-103.29417" municipality="Basin area" pageId="14" pageNumber="13" specimenCount="1">
<emphasis id="B93DEAE0FFFDFF9CFA989A19FA11FB14" bold="true" box="[1404,1440,1133,1157]" pageId="14" pageNumber="13">[2]</emphasis>
*
<location id="8E966029FFFDFF9CFF289AFAFE43FB37" LSID="urn:lsid:plazi:treatment:03E087E4FFFDFF9CFF249C9FFDA9F979:8E966029FFFDFF9CFF289AFAFE43FB37" box="[204,498,1166,1190]" latitude="29.267778" longLatPrecision="20" longitude="-103.29417" municipality="Basin area" name="Big Bend National Park" pageId="14" pageNumber="13">Big Bend National Park</location>
,
<collectingMunicipality id="6B92AC88FFFDFF9CFE189AFAFD30FB34" box="[508,641,1166,1190]" pageId="14" pageNumber="13">Basin area</collectingMunicipality>
,
<geoCoordinate id="EE7D5035FFFDFF9CFD689AF9FCA5FB34" box="[652,788,1164,1189]" degrees="29" direction="north" minutes="16" orientation="latitude" pageId="14" pageNumber="13" precision="15" seconds="04" value="29.267778">29°16′04″N</geoCoordinate>
<geoCoordinate id="EE7D5035FFFDFF9CFCFE9AFAFC09FB34" box="[794,952,1164,1190]" degrees="103" direction="west" minutes="17" orientation="longitude" pageId="14" pageNumber="13" precision="15" seconds="39" value="-103.29417">103°17′39″W</geoCoordinate>
,
<quantity id="4CB19B17FFFDFF9CFC269AF9FBAAFB34" box="[962,1051,1165,1189]" metricMagnitude="3" metricUnit="m" metricValue="1.66" pageId="14" pageNumber="13" unit="m" value="1660.0">
<elevation id="0064D1C1FFFDFF9CFC269AF9FBAAFB34" box="[962,1051,1165,1189]" metricMagnitude="3" metricUnit="m" metricValue="1.66" pageId="14" pageNumber="13" unit="m" value="1660.0">1660 m</elevation>
</quantity>
(May)
</materialsCitation>
;
<materialsCitation id="3B213CAFFFFDFF9CFB919AF9FD2EFB57" ID-GBIF-Occurrence="2575871697" elevation="1445" latitude="30.378056" location="Alpine" longLatPrecision="20" longitude="-103.75166" municipality="Sunny Glen" pageId="14" pageNumber="13" specimenCount="1">
<emphasis id="B93DEAE0FFFDFF9CFB919AF9FB28FB34" bold="true" box="[1141,1177,1165,1189]" pageId="14" pageNumber="13">[3]</emphasis>
*
<location id="8E966029FFFDFF9CFB4E9AFAFB4EFB37" LSID="urn:lsid:plazi:treatment:03E087E4FFFDFF9CFF249C9FFDA9F979:8E966029FFFDFF9CFB4E9AFAFB4EFB37" box="[1194,1279,1166,1190]" latitude="30.378056" longLatPrecision="20" longitude="-103.75166" municipality="Sunny Glen" name="Alpine" pageId="14" pageNumber="13">Alpine</location>
,
<collectingMunicipality id="6B92AC88FFFDFF9CFAED9AFAFA2AFB37" box="[1289,1435,1166,1190]" pageId="14" pageNumber="13">Sunny Glen</collectingMunicipality>
,
<geoCoordinate id="EE7D5035FFFDFF9CFF249ADAFEF8FB54" box="[192,329,1196,1222]" degrees="30" direction="north" minutes="22" orientation="latitude" pageId="14" pageNumber="13" precision="15" seconds="41" value="30.378056">30°22′41″N</geoCoordinate>
<geoCoordinate id="EE7D5035FFFDFF9CFEB49ADAFE5FFB54" box="[336,494,1196,1222]" degrees="103" direction="west" minutes="45" orientation="longitude" pageId="14" pageNumber="13" precision="15" seconds="06" value="-103.75166">103°45′06″W</geoCoordinate>
,
<quantity id="4CB19B17FFFDFF9CFE1E9ADAFDE5FB54" box="[506,596,1198,1222]" metricMagnitude="3" metricUnit="m" metricValue="1.4449999999999998" pageId="14" pageNumber="13" unit="m" value="1445.0">
<elevation id="0064D1C1FFFDFF9CFE1E9ADAFDE5FB54" box="[506,596,1198,1222]" metricMagnitude="3" metricUnit="m" metricValue="1.4449999999999998" pageId="14" pageNumber="13" unit="m" value="1445.0">1445 m</elevation>
</quantity>
(Jun)
</materialsCitation>
.
</paragraph>
<paragraph id="8BF636F2FFFDFF9CFF0C9ABAFB98FA97" blockId="14.[192,1440,1101,1286]" pageId="14" pageNumber="13">
<materialsCitation id="3B213CAFFFFDFF9CFF0C9ABAFADAFB77" ID-GBIF-Occurrence="2575871686" box="[232,1387,1228,1254]" county="Jeff Davis Co." latitude="30.595278" location="Davis Mountains State Park" longLatPrecision="20" longitude="-103.93472" municipality="Davis Mountains State Park" pageId="14" pageNumber="13" specimenCount="1">
<collectingCounty id="62974E7EFFFDFF9CFF0C9ABAFE28FB77" box="[232,409,1230,1254]" pageId="14" pageNumber="13">Jeff Davis Co.</collectingCounty>
:
<emphasis id="B93DEAE0FFFDFF9CFE429ABAFE7BFB77" bold="true" box="[422,458,1230,1254]" pageId="14" pageNumber="13">[1]</emphasis>
*
<collectingMunicipality id="6B92AC88FFFDFF9CFE3B9ABAFCF6FB77" box="[479,839,1230,1254]" pageId="14" pageNumber="13">Davis Mountains State Park</collectingMunicipality>
,
<geoCoordinate id="EE7D5035FFFDFF9CFCB19ABAFC6CFB74" box="[853,989,1228,1254]" degrees="30" direction="north" minutes="35" orientation="latitude" pageId="14" pageNumber="13" precision="15" seconds="43" value="30.595278">30°35′43″N</geoCoordinate>
<geoCoordinate id="EE7D5035FFFDFF9CFC029ABAFBC7FB74" box="[998,1142,1228,1254]" degrees="103" direction="west" minutes="56" orientation="longitude" pageId="14" pageNumber="13" precision="15" seconds="5" value="-103.93472">103°56′5″W</geoCoordinate>
,
<quantity id="4CB19B17FFFDFF9CFB679ABAFB51FB74" box="[1155,1248,1230,1254]" metricMagnitude="3" metricUnit="m" metricValue="1.54" pageId="14" pageNumber="13" unit="m" value="1540.0">1540 m</quantity>
(Jun, Aug)
</materialsCitation>
;
<materialsCitation id="3B213CAFFFFDFF9CFA989ABAFB94FA97" ID-GBIF-Occurrence="2575871682" county="Mountains Resort" elevation="1800" latitude="30.625" location="Davis Mountains Resort" longLatPrecision="20" longitude="-104.09167" municipality="Davis Mountains" pageId="14" pageNumber="13" specimenCount="1">
<emphasis id="B93DEAE0FFFDFF9CFA989ABAFF7CFA97" bold="true" pageId="14" pageNumber="13">[2] *</emphasis>
<location id="8E966029FFFDFF9CFF299A9AFE4BFA97" LSID="urn:lsid:plazi:treatment:03E087E4FFFDFF9CFF249C9FFDA9F979:8E966029FFFDFF9CFF299A9AFE4BFA97" box="[205,506,1262,1286]" county="Mountains Resort" latitude="30.625" longLatPrecision="20" longitude="-104.09167" municipality="Davis Mountains" name="Davis Mountains Resort" pageId="14" pageNumber="13">
<collectingMunicipality id="6B92AC88FFFDFF9CFF299A9AFE11FA97" box="[205,416,1262,1286]" pageId="14" pageNumber="13">
Davis
<collectingCounty id="62974E7EFFFDFF9CFEFF9A9AFE11FA97" box="[283,416,1262,1286]" pageId="14" pageNumber="13">Mountains</collectingCounty>
</collectingMunicipality>
Resort
</location>
,
<geoCoordinate id="EE7D5035FFFDFF9CFDE19A9AFD3FFA97" box="[517,654,1261,1286]" degrees="30" direction="north" minutes="37" orientation="latitude" pageId="14" pageNumber="13" precision="15" seconds="30" value="30.625">30°37′30″N</geoCoordinate>
<geoCoordinate id="EE7D5035FFFDFF9CFD719A9AFC85FA97" box="[661,820,1261,1286]" degrees="104" direction="west" minutes="05" orientation="longitude" pageId="14" pageNumber="13" precision="15" seconds="30" value="-104.09167">104°05′30″W</geoCoordinate>
,
<quantity id="4CB19B17FFFDFF9CFCDB9A9AFC2BFA97" box="[831,922,1262,1286]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="14" pageNumber="13" unit="m" value="1800.0">
<elevation id="0064D1C1FFFDFF9CFCDB9A9AFC2BFA97" box="[831,922,1262,1286]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="14" pageNumber="13" unit="m" value="1800.0">1800 m</elevation>
</quantity>
(Jun–Aug)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3536579FFFDFF9CFF249B6AFB50FAA7" box="[192,1249,1310,1334]" pageId="14" pageNumber="13" type="nomenclature">
<paragraph id="8BF636F2FFFDFF9CFF249B6AFB50FAA7" blockId="14.[192,1249,1310,1334]" box="[192,1249,1310,1334]" pageId="14" pageNumber="13">
<emphasis id="B93DEAE0FFFDFF9CFF249B6AFE57FAA7" bold="true" box="[192,486,1310,1334]" pageId="14" pageNumber="13">Collection method(s).</emphasis>
a) UV light trap; b) *direct capture beneath nests of
<taxonomicName id="4C494D71FFFDFF9CFB8A9B6BFB6BFAA7" baseAuthorityName="McCleve and Kohlmann" baseAuthorityYear="2005" box="[1134,1242,1311,1334]" class="Mammalia" family="Muridae" genus="Neotoma" kingdom="Animalia" order="Rodentia" pageId="14" pageNumber="13" phylum="Chordata" rank="genus">
<emphasis id="B93DEAE0FFFDFF9CFB8A9B6BFB6BFAA7" box="[1134,1242,1311,1334]" italics="true" pageId="14" pageNumber="13">Neotoma</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3536579FFFDFF9CFF249B3AFCB2FA26" pageId="14" pageNumber="13" type="biology_ecology">
<paragraph id="8BF636F2FFFDFF9CFF249B3AFD9FFAF7" blockId="14.[192,558,1358,1382]" box="[192,558,1358,1382]" pageId="14" pageNumber="13">
<emphasis id="B93DEAE0FFFDFF9CFF249B3AFE12FAF7" bold="true" box="[192,419,1358,1382]" pageId="14" pageNumber="13">Surface activity.</emphasis>
Nocturnal.
</paragraph>
<paragraph id="8BF636F2FFFDFF9CFF249B0AFCB2FA26" blockId="14.[192,1440,1406,1463]" pageId="14" pageNumber="13">
<emphasis id="B93DEAE0FFFDFF9CFF249B0AFE80FA07" bold="true" box="[192,305,1406,1430]" pageId="14" pageNumber="13">Habitat.</emphasis>
Montane, juniper-pinyon woodlands and arroyos in association with the wood rat,
<taxonomicName id="4C494D71FFFDFF9CFAD09B0BFE97FA27" authorityName="Hartley" authorityYear="1894" class="Mammalia" family="Muridae" genus="Neotoma" kingdom="Animalia" order="Rodentia" pageId="14" pageNumber="13" phylum="Chordata" rank="species" species="albigula">
<emphasis id="B93DEAE0FFFDFF9CFAD09B0BFE97FA27" italics="true" pageId="14" pageNumber="13">Neotoma albigula</emphasis>
</taxonomicName>
(and possibly also
<taxonomicName id="4C494D71FFFDFF9CFDEA9BEBFD47FA26" authority="Baird" authorityName="Baird" authorityYear="1855" box="[526,758,1439,1463]" class="Mammalia" family="Muridae" genus="Neotoma" kingdom="Animalia" order="Rodentia" pageId="14" pageNumber="13" phylum="Chordata" rank="species" species="mexicana">
<emphasis id="B93DEAE0FFFDFF9CFDEA9BEBFD17FA27" box="[526,678,1439,1462]" italics="true" pageId="14" pageNumber="13">N. mexicana</emphasis>
Baird
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C3536579FFFDFF9CFF249BBAFDA9F979" pageId="14" pageNumber="13" type="discussion">
<paragraph id="8BF636F2FFFDFF9CFF249BBAFDA9F979" blockId="14.[192,1440,1486,1768]" pageId="14" pageNumber="13">
<emphasis id="B93DEAE0FFFDFF9CFF249BBAFEE8FA77" bold="true" box="[192,345,1486,1510]" pageId="14" pageNumber="13">Comments.</emphasis>
This is by far the rarest dung beetle in the Big Bend. All examined specimens from the region were collected at lights.
<taxonomicName id="4C494D71FFFDFF9CFDD79B9BFCA2F997" authorityName="Schaeffer" authorityYear="1906" box="[563,787,1519,1542]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="arizonensis">
<emphasis id="B93DEAE0FFFDFF9CFDD79B9BFCA2F997" box="[563,787,1519,1542]" italics="true" pageId="14" pageNumber="13">Copris arizonensis</emphasis>
</taxonomicName>
is also known from various locations in the mountains of southeastern Arizona and adjacent areas of southwestern New
<collectingCountry id="F35E7662FFFDFF9CFC1B987BFBE4F9B6" box="[1023,1109,1551,1575]" name="Mexico" pageId="14" pageNumber="13">Mexico</collectingCountry>
and northern
<collectingRegion id="498DF810FFFDFF9CFAEB987BFA2DF9B6" box="[1295,1436,1551,1575]" country="Mexico" name="Chihuahua" pageId="14" pageNumber="13">Chihuahua</collectingRegion>
,
<collectingCountry id="F35E7662FFFDFF9CFF24985BFEA7F9D6" box="[192,278,1583,1607]" name="Mexico" pageId="14" pageNumber="13">Mexico</collectingCountry>
[
<bibRefCitation id="EFD84B03FFFDFF9CFEC2985BFE58F9D6" author="Matthews, E. G." box="[294,489,1583,1607]" pageId="14" pageNumber="13" pagination="1 - 139" refId="ref18649" refString="Matthews, E. G. 1961. A revision of the genus Copris Muller of the western hemisphere. Entomologica Americana 41: 1 - 139." type="journal article" year="1961">Matthews 1961</bibRefCitation>
;
<bibRefCitation id="EFD84B03FFFDFF9CFE12985BFCD6F9D6" author="McCleve, S. &amp; B. Kohlmann" box="[502,871,1583,1607]" pageId="14" pageNumber="13" pagination="17 - 28" refId="ref18677" refString="McCleve, S., and B. Kohlmann. 2005. New species and new records of Copris (Coleoptera: Scarabaeidae: Scarabaeinae) from Mexico and the United States. Zootaxa 1096: 17 - 28." type="journal article" year="2005">McCleve and Kohlmann 2005</bibRefCitation>
;
<bibRefCitation id="EFD84B03FFFDFF9CFC909844FBABF9D6" author="Warner, W. B." box="[884,1050,1583,1607]" pageId="14" pageNumber="13" pagination="232 - 240" refId="ref19485" refString="Warner, W. B. 1990. Two new North American Copris Muller, with notes on other species. Pan-Pacific Entomologist 66: 232 - 240." type="journal article" year="1990">Warner 1990</bibRefCitation>
].)
<figureCitation id="13722A77FFFDFF9CFBD3985BFB1FF9D6" box="[1079,1198,1583,1607]" captionStart="Figures 60–69" captionStartId="30.[192,286,1790,1812]" captionTargetBox="[253,1390,282,1714]" captionTargetId="figure@30.[227,1407,249,1724]" captionTargetPageId="30" captionText="Figures 60–69. Euoniticellus and Copris. 60) E. intermedius, dorsal view, female. 61) Same, lateral view of forebody. 62) Same, dorsal view, male. 63) Same, lateral view of forebody. 64) C. arizonensis, dorsal view of forebody, large male (southwestern Arizona). 65) Same, dorsal view, small male (Big Bend). 66) Same, lateral view of forebody. 67) Same, lateral view of forebody, female. 68) Same, oblique dorsal view. 69) Same, dorsal view of head and prothoracic margin." figureDoi="http://doi.org/10.5281/zenodo.3708204" httpUri="https://zenodo.org/record/3708204/files/figure.png" pageId="14" pageNumber="13">Figure 64</figureCitation>
depicts a specimen from southeastern Arizona (Dragoon Mts., Cochise Co.) exhibiting maximum expression of male armament, which I have not observed in Big Bend specimens (
<figureCitation id="13722A77FFFDFF9CFC60981BFC6DF916" box="[900,988,1647,1671]" captionStart="Figures 60–69" captionStartId="30.[192,286,1790,1812]" captionTargetBox="[253,1390,282,1714]" captionTargetId="figure@30.[227,1407,249,1724]" captionTargetPageId="30" captionText="Figures 60–69. Euoniticellus and Copris. 60) E. intermedius, dorsal view, female. 61) Same, lateral view of forebody. 62) Same, dorsal view, male. 63) Same, lateral view of forebody. 64) C. arizonensis, dorsal view of forebody, large male (southwestern Arizona). 65) Same, dorsal view, small male (Big Bend). 66) Same, lateral view of forebody. 67) Same, lateral view of forebody, female. 68) Same, oblique dorsal view. 69) Same, dorsal view of head and prothoracic margin." figureDoi="http://doi.org/10.5281/zenodo.3708204" httpUri="https://zenodo.org/record/3708204/files/figure.png" pageId="14" pageNumber="13">Fig. 65</figureCitation>
). The occurrence of
<taxonomicName id="4C494D71FFFDFF9CFB379804FACEF916" authorityName="Schaeffer" authorityYear="1906" box="[1235,1407,1648,1671]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="arizonensis">
<emphasis id="B93DEAE0FFFDFF9CFB379804FACEF916" box="[1235,1407,1648,1671]" italics="true" pageId="14" pageNumber="13">C. arizonensis</emphasis>
</taxonomicName>
in scattered higher mountainous areas of northern
<collectingRegion id="498DF810FFFDFF9CFCF798E4FC2CF939" box="[787,925,1680,1704]" country="Mexico" name="Chihuahua" pageId="14" pageNumber="13">Chihuahua</collectingRegion>
and the desert southwest is doubtless the result of shrinking and vertical isolation of suitable, high-elevation habitat during post-Pleistocene desertification of the region.
</paragraph>
</subSubSection>
</treatment>
</document>
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