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<document id="261275BBB17329DA36008AA2C9D5756D" ID-DOI="10.5281/zenodo.3708186" ID-GBIF-Dataset="f503e20e-e3fb-41ac-bf17-bdaf221bc7c7" ID-ISSN="1942-1354" ID-Zenodo-Dep="3708186" ID-ZooBank="55CCB217-771C-499D-9110-36F143C375C5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1584033498187" checkinUser="felipe" docAuthor="Edmonds, W. D." docDate="2018" docId="03E087E4FFF9FF99FF249B68FCDDFCA7" docLanguage="en" docName="InsectaMundi.0642.1-30.pdf.imf" docOrigin="Insecta Mundi 642" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Onthophagus velutinus Horn 1875" docType="treatment" docVersion="5" lastPageNumber="10" masterDocId="FFD9FF9CFFF3FF92FFE49E74FFB1FF91" masterDocTitle="The dung beetle fauna of the Big Bend region of Texas (Coleoptera: Scarabaeidae: Scarabaeinae)" masterLastPageNumber="30" masterPageNumber="1" pageNumber="9" updateTime="1698727366089" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
<mods:mods id="D2B3CF091BB8B25E2CFF260F97049608" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="00AF5619FEAB443DCC5984264AADE658">
<mods:title id="5E31EB1680815C4EEE5D9BE3216C196E">The dung beetle fauna of the Big Bend region of Texas (Coleoptera: Scarabaeidae: Scarabaeinae)</mods:title>
</mods:titleInfo>
<mods:name id="EF6D6505D3A3CFD6B5BFEB813BFA8E4D" type="personal">
<mods:role id="21F6D180367E58C218862AF96F1D0433">
<mods:roleTerm id="F3C2CB694CAB778E6E6E5DD7CA07246F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1D5C14C527EC18B23EEBD08ED737FCDB">Edmonds, W. D.</mods:namePart>
</mods:name>
<mods:typeOfResource id="F1DE420FECA777D841F913DA45391647">text</mods:typeOfResource>
<mods:relatedItem id="92A4EFB6FAF10F4BE65D16EC35E22A74" type="host">
<mods:titleInfo id="27E4AC9BEA53FE48B70C1768EBDE9D5E">
<mods:title id="0A8D7EBE73C5352DA8CD3C71EFAAC5F9">Insecta Mundi</mods:title>
</mods:titleInfo>
<mods:part id="2F1FA96A9AA0E2B2B06158470E59C2A6">
<mods:date id="AC632698027711A400B67A41C5B30DFA">2018</mods:date>
<mods:detail id="EC281755651F22C9F7EEB10ECA9948A6" type="pubDate">
<mods:number id="EF7C49628948D2F0114A9BA157B8B58D">2018-07-27</mods:number>
</mods:detail>
<mods:detail id="12FAD25AEF84C94A293E2D29DBECB567" type="issue">
<mods:number id="EB29AB16A8923E8F004056EE70EC2777">642</mods:number>
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<mods:start id="CA2426691D4A8281B66B193B7C9F36A9">1</mods:start>
<mods:end id="072BC578401B990ECFF4E56E92F04ED3">30</mods:end>
</mods:extent>
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<mods:classification id="26649307FF17309E1BF13592BFC314F2">journal article</mods:classification>
<mods:identifier id="A9BA1D94570B0E73EA9246E4B0E5A338" type="DOI">10.5281/zenodo.3708186</mods:identifier>
<mods:identifier id="5271FE41EAF2ECE72546A8DE3440A8FE" type="GBIF-Dataset">f503e20e-e3fb-41ac-bf17-bdaf221bc7c7</mods:identifier>
<mods:identifier id="44508C57F7F15169788CFA529261300C" type="ISSN">1942-1354</mods:identifier>
<mods:identifier id="D524970B676B97E045D0E370231B9C97" type="Zenodo-Dep">3708186</mods:identifier>
<mods:identifier id="2DBDB464E186A20B6B0AA7BBA8184FAC" type="ZooBank">55CCB217-771C-499D-9110-36F143C375C5</mods:identifier>
</mods:mods>
<treatment id="03E087E4FFF9FF99FF249B68FCDDFCA7" ID-DOI="http://doi.org/10.5281/zenodo.3717136" ID-GBIF-Taxon="162485172" ID-Zenodo-Dep="3717136" LSID="urn:lsid:plazi:treatment:03E087E4FFF9FF99FF249B68FCDDFCA7" httpUri="http://treatment.plazi.org/id/03E087E4FFF9FF99FF249B68FCDDFCA7" lastPageId="11" lastPageNumber="10" pageId="10" pageNumber="9">
<subSubSection id="C3536579FFF9FF98FF249B68FDFFFAA4" box="[192,590,1308,1333]" pageId="10" pageNumber="9" type="nomenclature">
<paragraph id="8BF636F2FFF9FF98FF249B68FDFFFAA4" blockId="10.[192,590,1308,1366]" box="[192,590,1308,1333]" pageId="10" pageNumber="9">
<heading id="D0BE819EFFF9FF98FF249B68FDFFFAA4" bold="true" box="[192,590,1308,1333]" fontSize="10" level="2" pageId="10" pageNumber="9" reason="3">
<taxonomicName id="4C494D71FFF9FF98FF249B68FDFFFAA4" ID-CoL="7W5HS" authority="Horn" authorityName="Horn" authorityYear="1875" box="[192,590,1308,1333]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="velutinus">
<emphasis id="B93DEAE0FFF9FF98FF249B68FDFFFAA4" bold="true" box="[192,590,1308,1333]" pageId="10" pageNumber="9">
<emphasis id="B93DEAE0FFF9FF98FF249B68FE4EFAA5" bold="true" box="[192,511,1308,1332]" italics="true" pageId="10" pageNumber="9">Onthophagus velutinus</emphasis>
Horn
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3536579FFF9FF98FF249B4AFEF0FAC7" box="[192,321,1342,1366]" pageId="10" pageNumber="9" type="description">
<paragraph id="8BF636F2FFF9FF98FF249B4AFEF0FAC7" blockId="10.[192,590,1308,1366]" box="[192,321,1342,1366]" pageId="10" pageNumber="9">Fig, 44–48</paragraph>
</subSubSection>
<subSubSection id="C3536579FFF9FF98FF249B1AFBD6F942" pageId="10" pageNumber="9" type="diagnosis">
<paragraph id="8BF636F2FFF9FF98FF249B1AFBD6F942" blockId="10.[192,1440,1390,1747]" pageId="10" pageNumber="9">
<emphasis id="B93DEAE0FFF9FF98FF249B1AFEFEFA17" bold="true" box="[192,335,1390,1414]" pageId="10" pageNumber="9">Diagnosis.</emphasis>
Dorsum black to brownish black, often with greenish reflections. Length
<quantity id="4CB19B17FFF9FF98FB2D9B1BFAF4FA17" box="[1225,1349,1391,1415]" metricMagnitude="-3" metricUnit="m" metricValue="5.25" metricValueMax="6.5" metricValueMin="4.0" pageId="10" pageNumber="9" unit="mm" value="5.25" valueMax="6.5" valueMin="4.0">4–6.5 mm</quantity>
. Prono- tum evenly but not densely covered by small shining granules, each with associated short seta, on field of extremely fine shagreening (
<figureCitation id="13722A77FFF9FF98FDDA9BC5FD27FA58" box="[574,662,1457,1481]" captionStart="Figures 36–48" captionStartId="28.[192,287,1790,1812]" captionTargetBox="[255,1384,219,1755]" captionTargetId="figure@28.[223,1408,207,1757]" captionTargetPageId="28" captionText="Figures 36–48. Onthophagus. 36) O. brevifrons, dorsal view, male. 37) Same, oblique frontal view. 38) Same, dorsal view of forebody, female. 39) Same, prolegs (A, female; B, male. 40) Same, pronotal sculpturing. 41) O. browni, dorsal view, male. 42) Same, oblique frontal view. 43) Same, oblique frontal view, female. 44) O. velutinus, dorsal view, male. 45) Same, oblique frontal view. 46) Same, dorsal view of female forebody. 47) Same, oblique frontal view. 48) Same, pronotal sculpturing." figureDoi="http://doi.org/10.5281/zenodo.3708200" httpUri="https://zenodo.org/record/3708200/files/figure.png" pageId="10" pageNumber="9">Fig. 48</figureCitation>
); elytral interstriae with two longitudinal rows of small, setose granules on field of fine shagreening; pygidium evenly punctate, punctures setose. Front legs similar in both sexes. Male (
<figureCitation id="13722A77FFF9FF98FE7C9B80FDA8F99A" box="[408,537,1523,1548]" captionStart="Figures 36–48" captionStartId="28.[192,287,1790,1812]" captionTargetBox="[255,1384,219,1755]" captionTargetId="figure@28.[223,1408,207,1757]" captionTargetPageId="28" captionText="Figures 36–48. Onthophagus. 36) O. brevifrons, dorsal view, male. 37) Same, oblique frontal view. 38) Same, dorsal view of forebody, female. 39) Same, prolegs (A, female; B, male. 40) Same, pronotal sculpturing. 41) O. browni, dorsal view, male. 42) Same, oblique frontal view. 43) Same, oblique frontal view, female. 44) O. velutinus, dorsal view, male. 45) Same, oblique frontal view. 46) Same, dorsal view of female forebody. 47) Same, oblique frontal view. 48) Same, pronotal sculpturing." figureDoi="http://doi.org/10.5281/zenodo.3708200" httpUri="https://zenodo.org/record/3708200/files/figure.png" pageId="10" pageNumber="9">Fig. 44–45</figureCitation>
) – Clypeus weakly upturned medially. Clypeal carina weak, frontal carina replaced by pair of slender, nearly parallel horns (reduced in small individuals). Pronotum convex except for shallowly concave, vertical anterior surface; concavity partially divided by weak, median, vertical carina extending partway down from upper margin. Female (
<figureCitation id="13722A77FFF9FF98FC409823FB94F9FE" box="[932,1061,1623,1647]" captionStart="Figures 36–48" captionStartId="28.[192,287,1790,1812]" captionTargetBox="[255,1384,219,1755]" captionTargetId="figure@28.[223,1408,207,1757]" captionTargetPageId="28" captionText="Figures 36–48. Onthophagus. 36) O. brevifrons, dorsal view, male. 37) Same, oblique frontal view. 38) Same, dorsal view of forebody, female. 39) Same, prolegs (A, female; B, male. 40) Same, pronotal sculpturing. 41) O. browni, dorsal view, male. 42) Same, oblique frontal view. 43) Same, oblique frontal view, female. 44) O. velutinus, dorsal view, male. 45) Same, oblique frontal view. 46) Same, dorsal view of female forebody. 47) Same, oblique frontal view. 48) Same, pronotal sculpturing." figureDoi="http://doi.org/10.5281/zenodo.3708200" httpUri="https://zenodo.org/record/3708200/files/figure.png" pageId="10" pageNumber="9">Fig. 46–47</figureCitation>
) – Clypeus not upturned. Head with two transverse carinae, clypeal carina weak, frontal carina stronger, ends often distinctly raised higher than middle. Pronotum evenly convex, rarely with slightest hint of anteromedian process. A formal description of this species appears in
<bibRefCitation id="EFD84B03FFF9FF98FD0598CFFBD5F942" author="Howden, H. F. &amp; O. L. Cartwright" box="[737,1124,1723,1747]" pageId="10" pageNumber="9" pagination="1 - 135" refId="ref18230" refString="Howden, H. F., and O. L. Cartwright. 1963. Scarab beetles of the genus Onthophagus Latreille north of Mexico (Coleoptera: Scarabaeidae). Proceedings of the United States National Museum 114: 1 - 135." type="journal article" year="1963">Howden and Cartwright (1963)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3536579FFF9FF99FF24989FFCCBFDB4" lastPageId="11" lastPageNumber="10" pageId="10" pageNumber="9" type="materials_examined">
<paragraph id="8BF636F2FFF9FF98FF24989FFC19F895" blockId="10.[192,1440,1771,1929]" box="[192,936,1771,1796]" pageId="10" pageNumber="9">
<emphasis id="B93DEAE0FFF9FF98FF24989FFDA2F892" bold="true" box="[192,531,1771,1796]" pageId="10" pageNumber="9">Big Bend collection sites</emphasis>
(altitudinal range:
<quantity id="4CB19B17FFF9FF98FCE79898FC2AF895" box="[771,923,1772,1796]" metricMagnitude="3" metricUnit="m" metricValue="1.0474999999999999" metricValueMax="1.54" metricValueMin="0.5549999999999999" pageId="10" pageNumber="9" unit="m" value="1047.5" valueMax="1540.0" valueMin="555.0">555–1540 m</quantity>
).
</paragraph>
<paragraph id="8BF636F2FFF9FF98FF0C9979FCD3F818" blockId="10.[192,1440,1771,1929]" pageId="10" pageNumber="9">
<materialsCitation id="3B213CAFFFF9FF98FF0C9979FEBBF8D7" ID-GBIF-Occurrence="2575871582" county="Brewster Co." latitude="29.324446" location="Panther Junction" longLatPrecision="20" longitude="-103.20583" municipality="Big Bend National Park" pageId="10" pageNumber="9" specimenCount="1">
<collectingCounty id="62974E7EFFF9FF98FF0C9979FE3FF8B4" box="[232,398,1805,1829]" pageId="10" pageNumber="9">Brewster Co.</collectingCounty>
:
<emphasis id="B93DEAE0FFF9FF98FE789979FE71F8B4" bold="true" box="[412,448,1805,1829]" pageId="10" pageNumber="9">[1]</emphasis>
*
<collectingMunicipality id="6B92AC88FFF9FF98FE329979FCBBF8B4" box="[470,778,1805,1829]" pageId="10" pageNumber="9">Big Bend National Park</collectingMunicipality>
,
<location id="8E966029FFF9FF98FCFC9979FC44F8B4" LSID="urn:lsid:plazi:treatment:03E087E4FFF9FF99FF249B68FCDDFCA7:8E966029FFF9FF98FCFC9979FC44F8B4" box="[792,1013,1805,1829]" county="Brewster Co." latitude="29.324446" longLatPrecision="20" longitude="-103.20583" municipality="Big Bend National Park" name="Panther Junction" pageId="10" pageNumber="9">Panther Junction</location>
,
<geoCoordinate id="EE7D5035FFF9FF98FBE79979FB3DF8B4" box="[1027,1164,1804,1829]" degrees="29" direction="north" minutes="19" orientation="latitude" pageId="10" pageNumber="9" precision="15" seconds="28" value="29.324446">29°19′28″N</geoCoordinate>
<geoCoordinate id="EE7D5035FFF9FF98FB729979FA85F8B5" box="[1174,1332,1804,1829]" degrees="103" direction="west" minutes="12" orientation="longitude" pageId="10" pageNumber="9" precision="15" seconds="21" value="-103.20583">103°12′21″W</geoCoordinate>
,
<quantity id="4CB19B17FFF9FF98FAA79979FA11F8B5" box="[1347,1440,1805,1829]" metricMagnitude="3" metricUnit="m" metricValue="1.17" pageId="10" pageNumber="9" unit="m" value="1170.0">1170 m</quantity>
(May)
</materialsCitation>
;
<materialsCitation id="3B213CAFFFF9FF98FEF2995AFA2AF8D7" ID-GBIF-Occurrence="2575871585" box="[278,1435,1837,1863]" elevation="555" latitude="29.183054" location="Big Bend National Park" longLatPrecision="20" longitude="-102.96027" municipality="Boquillas Ranger Station" pageId="10" pageNumber="9" specimenCount="1">
<emphasis id="B93DEAE0FFF9FF98FEF2995AFE8BF8D7" bold="true" box="[278,314,1838,1862]" pageId="10" pageNumber="9">[2]</emphasis>
*
<location id="8E966029FFF9FF98FEA9995AFDC6F8D7" LSID="urn:lsid:plazi:treatment:03E087E4FFF9FF99FF249B68FCDDFCA7:8E966029FFF9FF98FEA9995AFDC6F8D7" box="[333,631,1838,1862]" latitude="29.183054" longLatPrecision="20" longitude="-102.96027" municipality="Boquillas Ranger Station" name="Big Bend National Park" pageId="10" pageNumber="9">Big Bend National Park</location>
,
<collectingMunicipality id="6B92AC88FFF9FF98FD67995AFC08F8D7" box="[643,953,1838,1863]" pageId="10" pageNumber="9">Boquillas Ranger Station</collectingMunicipality>
,
<geoCoordinate id="EE7D5035FFF9FF98FC22995AFBFFF8D7" box="[966,1102,1837,1862]" degrees="29" direction="north" minutes="10" orientation="latitude" pageId="10" pageNumber="9" precision="15" seconds="59" value="29.183054">29°10′59″N</geoCoordinate>
<geoCoordinate id="EE7D5035FFF9FF98FBB1995AFB45F8D7" box="[1109,1268,1837,1862]" degrees="102" direction="west" minutes="57" orientation="longitude" pageId="10" pageNumber="9" precision="15" seconds="37" value="-102.96027">102°57′37″W</geoCoordinate>
,
<quantity id="4CB19B17FFF9FF98FB1B995AFAFBF8D7" box="[1279,1354,1838,1862]" metricMagnitude="2" metricUnit="m" metricValue="5.55" pageId="10" pageNumber="9" unit="m" value="555.0">
<elevation id="0064D1C1FFF9FF98FB1B995AFAFBF8D7" box="[1279,1354,1838,1862]" metricMagnitude="2" metricUnit="m" metricValue="5.55" pageId="10" pageNumber="9" unit="m" value="555.0">555 m</elevation>
</quantity>
(May)
</materialsCitation>
;
<materialsCitation id="3B213CAFFFF9FF98FF24993BFB7DF8F6" ID-GBIF-Occurrence="2575871576" box="[192,1228,1870,1895]" elevation="1325" latitude="29.281944" location="Big Bend National Park" longLatPrecision="20" longitude="-103.336945" municipality="Oak Spring" pageId="10" pageNumber="9" specimenCount="1">
<emphasis id="B93DEAE0FFF9FF98FF24993BFF55F8F6" bold="true" box="[192,228,1871,1895]" pageId="10" pageNumber="9">[3]</emphasis>
*
<location id="8E966029FFF9FF98FF1D993BFD9CF8F6" LSID="urn:lsid:plazi:treatment:03E087E4FFF9FF99FF249B68FCDDFCA7:8E966029FFF9FF98FF1D993BFD9CF8F6" box="[249,557,1871,1895]" latitude="29.281944" longLatPrecision="20" longitude="-103.336945" municipality="Oak Spring" name="Big Bend National Park" pageId="10" pageNumber="9">Big Bend National Park</location>
,
<collectingMunicipality id="6B92AC88FFF9FF98FDDF993BFD7CF8F6" box="[571,717,1871,1895]" pageId="10" pageNumber="9">Oak Spring</collectingMunicipality>
,
<geoCoordinate id="EE7D5035FFF9FF98FD38993BFCD5F8F6" box="[732,868,1870,1895]" degrees="29" direction="north" minutes="16" orientation="latitude" pageId="10" pageNumber="9" precision="15" seconds="55" value="29.281944">29°16′55″N</geoCoordinate>
<geoCoordinate id="EE7D5035FFF9FF98FC8B993BFBBCF8F6" box="[879,1037,1870,1895]" degrees="103" direction="west" minutes="20" orientation="longitude" pageId="10" pageNumber="9" precision="15" seconds="13" value="-103.336945">103°20′13″W</geoCoordinate>
,
<quantity id="4CB19B17FFF9FF98FBF8993BFBC8F8F6" box="[1052,1145,1871,1895]" metricMagnitude="3" metricUnit="m" metricValue="1.325" pageId="10" pageNumber="9" unit="m" value="1325.0">
<elevation id="0064D1C1FFF9FF98FBF8993BFBC8F8F6" box="[1052,1145,1871,1895]" metricMagnitude="3" metricUnit="m" metricValue="1.325" pageId="10" pageNumber="9" unit="m" value="1325.0">1325 m</elevation>
</quantity>
(May)
</materialsCitation>
;
<materialsCitation id="3B213CAFFFF9FF98FB3F993BFCEFF818" ID-GBIF-Occurrence="2575871581" collectionCode="RV" elevation="770" latitude="29.644444" location="Stillwell" longLatPrecision="20" longitude="-103.07972" municipality="Park" pageId="10" pageNumber="9" specimenCode="FM 2627" specimenCount="1">
<emphasis id="B93DEAE0FFF9FF98FB3F993BFB4EF8F6" bold="true" box="[1243,1279,1871,1895]" pageId="10" pageNumber="9">[4]</emphasis>
<location id="8E966029FFF9FF98FAED993BFAC1F8F6" LSID="urn:lsid:plazi:treatment:03E087E4FFF9FF99FF249B68FCDDFCA7:8E966029FFF9FF98FAED993BFAC1F8F6" box="[1289,1392,1871,1895]" latitude="29.644444" longLatPrecision="20" longitude="-103.07972" municipality="Park" name="Stillwell" pageId="10" pageNumber="9">Stillwell</location>
<collectionCode id="ED58AE37FFF9FF98FA9E9924FA11F8F6" box="[1402,1440,1872,1895]" country="Italy" name="Collection of Leptospira Strains" pageId="10" pageNumber="9">RV</collectionCode>
<collectingMunicipality id="6B92AC88FFF9FF98FF249904FF4EF819" box="[192,255,1904,1928]" pageId="10" pageNumber="9">Park</collectingMunicipality>
,
<specimenCode id="DBEF9E89FFF9FF98FEEF9905FECAF819" box="[267,379,1904,1928]" collectionCode="FM" country="China" name="Department of Nature, Fujian Province Museum" pageId="10" pageNumber="9">FM 2627</specimenCode>
,
<geoCoordinate id="EE7D5035FFF9FF98FE629905FDBEF819" box="[390,527,1903,1929]" degrees="29" direction="north" minutes="38" orientation="latitude" pageId="10" pageNumber="9" precision="15" seconds="40" value="29.644444">29°38′40″N</geoCoordinate>
<geoCoordinate id="EE7D5035FFF9FF98FDF29905FD0DF819" box="[534,700,1903,1929]" degrees="103" direction="west" minutes="04" orientation="longitude" pageId="10" pageNumber="9" precision="15" seconds="47" value="-103.07972">103° 04′47″W</geoCoordinate>
,
<quantity id="4CB19B17FFF9FF98FD2C9905FCA2F819" box="[712,787,1905,1929]" metricMagnitude="2" metricUnit="m" metricValue="7.7" pageId="10" pageNumber="9" unit="m" value="770.0">
<elevation id="0064D1C1FFF9FF98FD2C9905FCA2F819" box="[712,787,1905,1929]" metricMagnitude="2" metricUnit="m" metricValue="7.7" pageId="10" pageNumber="9" unit="m" value="770.0">770 m</elevation>
</quantity>
(Jun)
</materialsCitation>
.
</paragraph>
<paragraph id="8BF636F2FFF8FF99FF0C9EA9FE90FEA4" blockId="11.[192,1440,221,470]" pageId="11" pageNumber="10">
<materialsCitation id="3B213CAFFFF8FF99FF0C9EA9FD51FE84" ID-GBIF-Occurrence="2575871579" county="Jeff Davis Co." latitude="30.535" location="Chihuahuan Desert Research Institute" longLatPrecision="20" longitude="-103.84361" municipality="Fort Davis" pageId="11" pageNumber="10" specimenCount="1">
<collectingCounty id="62974E7EFFF8FF99FF0C9EA9FE20FF64" box="[232,401,221,245]" pageId="11" pageNumber="10">Jeff Davis Co.</collectingCounty>
:
<emphasis id="B93DEAE0FFF8FF99FE7F9EA9FE0FFF64" bold="true" box="[411,446,221,245]" pageId="11" pageNumber="10">[1]</emphasis>
~
<locationDeviation id="9A442FD0FFF8FF99FE309EA9FD89FF64" bearing="135.0" box="[468,568,221,245]" distance="8000.0" pageId="11" pageNumber="10">
<quantity id="4CB19B17FFF8FF99FE309EA9FDBEFF64" box="[468,527,221,245]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="11" pageNumber="10" unit="km" value="8.0">8 km</quantity>
SE
</locationDeviation>
<collectingMunicipality id="6B92AC88FFF8FF99FDDA9EAAFD0FFF64" box="[574,702,221,245]" pageId="11" pageNumber="10">Fort Davis</collectingMunicipality>
(via TX 118),
<location id="8E966029FFF8FF99FC869EA9FA89FF64" LSID="urn:lsid:plazi:treatment:03E087E4FFF9FF99FF249B68FCDDFCA7:8E966029FFF8FF99FC869EA9FA89FF64" box="[866,1336,221,245]" county="Jeff Davis Co." latitude="30.535" longLatPrecision="20" longitude="-103.84361" municipality="Fort Davis" name="Chihuahuan Desert Research Institute" pageId="11" pageNumber="10">Chihuahuan Desert Research Institute</location>
(
<location id="8E966029FFF8FF99FAA29EA9FF4BFE84" LSID="urn:lsid:plazi:treatment:03E087E4FFF9FF99FF249B68FCDDFCA7:8E966029FFF8FF99FAA29EA9FF4BFE84" county="Jeff Davis Co." latitude="30.535" longLatPrecision="20" longitude="-103.84361" municipality="Fort Davis" name="Quarry Unit" pageId="11" pageNumber="10">Quarry Unit</location>
),
<geoCoordinate id="EE7D5035FFF8FF99FEE99E89FE25FE84" box="[269,404,252,277]" degrees="30" direction="north" minutes="32" orientation="latitude" pageId="11" pageNumber="10" precision="15" seconds="06" value="30.535">30°32′06″N</geoCoordinate>
<geoCoordinate id="EE7D5035FFF8FF99FE7D9E89FD87FE84" box="[409,566,252,277]" degrees="103" direction="west" minutes="50" orientation="longitude" pageId="11" pageNumber="10" precision="15" seconds="37" value="-103.84361">103°50′37″W</geoCoordinate>
,
<quantity id="4CB19B17FFF8FF99FDDB9E89FD24FE84" box="[575,661,253,277]" metricMagnitude="3" metricUnit="m" metricValue="1.48" pageId="11" pageNumber="10" unit="m" value="1480.0">1480 m</quantity>
(Aug)
</materialsCitation>
;
<materialsCitation id="3B213CAFFFF8FF99FD0E9E89FEACFEA4" ID-GBIF-Occurrence="2575871575" elevation="1540" latitude="30.595278" location="Davis Mountains State Park" longLatPrecision="20" longitude="-103.93472" pageId="11" pageNumber="10" specimenCount="1">
<emphasis id="B93DEAE0FFF8FF99FD0E9E89FCBCFE84" bold="true" box="[746,781,253,277]" pageId="11" pageNumber="10">[2]</emphasis>
<location id="8E966029FFF8FF99FCF79E89FBD8FE84" LSID="urn:lsid:plazi:treatment:03E087E4FFF9FF99FF249B68FCDDFCA7:8E966029FFF8FF99FCF79E89FBD8FE84" box="[787,1129,253,277]" latitude="30.595278" longLatPrecision="20" longitude="-103.93472" name="Davis Mountains State Park" pageId="11" pageNumber="10">Davis Mountains State Park</location>
,
<geoCoordinate id="EE7D5035FFF8FF99FB979E89FB4BFE84" box="[1139,1274,252,277]" degrees="30" direction="north" minutes="35" orientation="latitude" pageId="11" pageNumber="10" precision="15" seconds="43" value="30.595278">30°35′43″N</geoCoordinate>
<geoCoordinate id="EE7D5035FFF8FF99FB1B9E89FA2DFE84" box="[1279,1436,252,277]" degrees="103" direction="west" minutes="56" orientation="longitude" pageId="11" pageNumber="10" precision="15" seconds="05" value="-103.93472">103°56′05″W</geoCoordinate>
,
<quantity id="4CB19B17FFF8FF99FF249F69FEACFEA4" box="[192,285,285,309]" metricMagnitude="3" metricUnit="m" metricValue="1.54" pageId="11" pageNumber="10" unit="m" value="1540.0">
<elevation id="0064D1C1FFF8FF99FF249F69FEACFEA4" box="[192,285,285,309]" metricMagnitude="3" metricUnit="m" metricValue="1.54" pageId="11" pageNumber="10" unit="m" value="1540.0">1540 m</elevation>
</quantity>
</materialsCitation>
.
</paragraph>
<paragraph id="8BF636F2FFF8FF99FF0C9F49FEDDFE47" blockId="11.[192,1440,221,470]" pageId="11" pageNumber="10">
<materialsCitation id="3B213CAFFFF8FF99FF0C9F49FEDAFEE4" ID-GBIF-Occurrence="2575871630" country="Netherlands" county="Presidio Co." elevation="1410" latitude="30.547222" location="Presidio Co." longLatPrecision="20" longitude="-104.66111" pageId="11" pageNumber="10" specimenCount="1">
<collectingCounty id="62974E7EFFF8FF99FF0C9F49FECFFEC4" box="[232,382,317,341]" pageId="11" pageNumber="10">Presidio Co.</collectingCounty>
:
<emphasis id="B93DEAE0FFF8FF99FE6C9F49FE1DFEC4" bold="true" box="[392,428,317,341]" pageId="11" pageNumber="10">[1]</emphasis>
C.E. Miller Ranch (~
<locationDeviation id="9A442FD0FFF8FF99FD489F49FCA9FEC4" bearing="270.0" box="[684,792,317,341]" distance="16000.0" pageId="11" pageNumber="10">
<quantity id="4CB19B17FFF8FF99FD489F49FD46FEC4" box="[684,759,317,341]" metricMagnitude="4" metricUnit="m" metricValue="1.6" pageId="11" pageNumber="10" unit="km" value="16.0">16 km</quantity>
W
</locationDeviation>
Valentine),
<geoCoordinate id="EE7D5035FFF8FF99FC489F49FB84FEC4" box="[940,1077,316,341]" degrees="30" direction="north" minutes="32" orientation="latitude" pageId="11" pageNumber="10" precision="15" seconds="50" value="30.547222">30°32′50″N</geoCoordinate>
<geoCoordinate id="EE7D5035FFF8FF99FBDF9F49FB66FEC4" box="[1083,1239,316,341]" degrees="104" direction="west" minutes="39" orientation="longitude" pageId="11" pageNumber="10" precision="15" seconds="40" value="-104.66111">104°39′40″W</geoCoordinate>
(Camp Holland)
<quantity id="4CB19B17FFF8FF99FF249F29FEADFEE4" box="[192,284,349,373]" metricMagnitude="3" metricUnit="m" metricValue="1.41" pageId="11" pageNumber="10" unit="m" value="1410.0">
<elevation id="0064D1C1FFF8FF99FF249F29FEADFEE4" box="[192,284,349,373]" metricMagnitude="3" metricUnit="m" metricValue="1.41" pageId="11" pageNumber="10" unit="m" value="1410.0">1410 m</elevation>
</quantity>
(Aug)
</materialsCitation>
;
<materialsCitation id="3B213CAFFFF8FF99FE9E9F29FE24FE04" ID-GBIF-Occurrence="2575871574" elevation="1350" location="Valentine" longitude="-104.64555" municipality="Miller Ranch" pageId="11" pageNumber="10" specimenCount="1">
<emphasis id="B93DEAE0FFF8FF99FE9E9F29FE2CFEE4" bold="true" box="[378,413,349,373]" pageId="11" pageNumber="10">[2]</emphasis>
~16 kmW
<location id="8E966029FFF8FF99FDCD9F29FD10FEE4" LSID="urn:lsid:plazi:treatment:03E087E4FFF9FF99FF249B68FCDDFCA7:8E966029FFF8FF99FDCD9F29FD10FEE4" box="[553,673,349,373]" longitude="-104.64555" municipality="Miller Ranch" name="Valentine" pageId="11" pageNumber="10">Valentine</location>
(
<collectingMunicipality id="6B92AC88FFF8FF99FD569F29FCE9FEE4" box="[690,856,349,373]" pageId="11" pageNumber="10">Miller Ranch</collectingMunicipality>
, near headquarters),
<geoCoordinate id="EE7D5035FFF8FF99FB829F29FB45FEE4" box="[1126,1268,348,373]" pageId="11" pageNumber="10">30°33°30″N</geoCoordinate>
<geoCoordinate id="EE7D5035FFF8FF99FB199F29FA2DFEE4" box="[1277,1436,348,373]" degrees="104" direction="west" minutes="38" orientation="longitude" pageId="11" pageNumber="10" precision="15" seconds="44" value="-104.64555">104°38′44″W</geoCoordinate>
,
<quantity id="4CB19B17FFF8FF99FF249F09FEA2FE04" box="[192,275,381,405]" metricMagnitude="3" metricUnit="m" metricValue="1.35" pageId="11" pageNumber="10" unit="m" value="1350.0">
<elevation id="0064D1C1FFF8FF99FF249F09FEA2FE04" box="[192,275,381,405]" metricMagnitude="3" metricUnit="m" metricValue="1.35" pageId="11" pageNumber="10" unit="m" value="1350.0">1350m</elevation>
</quantity>
(Jul–Aug)
</materialsCitation>
;
<materialsCitation id="3B213CAFFFF8FF99FE449F09FAD0FE07" ID-GBIF-Occurrence="2575871670" box="[416,1377,380,406]" elevation="1530" latitude="30.365" location="Ranch" longLatPrecision="20" longitude="-104.12" pageId="11" pageNumber="10" specimenCount="1">
<emphasis id="B93DEAE0FFF8FF99FE449F09FE75FE04" bold="true" box="[416,452,381,405]" pageId="11" pageNumber="10">[3]</emphasis>
~
<locationDeviation id="9A442FD0FFF8FF99FE3F9F0AFD3BFE04" box="[475,650,381,405]" location="Marfa" pageId="11" pageNumber="10">
<quantity id="4CB19B17FFF8FF99FE3F9F0AFDA6FE04" box="[475,535,381,405]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="11" pageNumber="10" unit="km" value="7.0">7 km</quantity>
W Marfa
</locationDeviation>
(Hip-O
<location id="8E966029FFF8FF99FD0F9F09FC8DFE04" LSID="urn:lsid:plazi:treatment:03E087E4FFF9FF99FF249B68FCDDFCA7:8E966029FFF8FF99FD0F9F09FC8DFE04" box="[747,828,381,405]" latitude="30.365" longLatPrecision="20" longitude="-104.12" name="Ranch" pageId="11" pageNumber="10">Ranch</location>
),
<geoCoordinate id="EE7D5035FFF8FF99FCB49F0AFC69FE04" box="[848,984,380,406]" degrees="30" direction="north" minutes="21" orientation="latitude" pageId="11" pageNumber="10" precision="15" seconds="54" value="30.365">30°21′54″N</geoCoordinate>
<geoCoordinate id="EE7D5035FFF8FF99FC3B9F0AFBC4FE04" box="[991,1141,380,406]" degrees="104" direction="west" minutes="7" orientation="longitude" pageId="11" pageNumber="10" precision="15" seconds="12" value="-104.12">104° 7′12″W</geoCoordinate>
,
<quantity id="4CB19B17FFF8FF99FB649F09FB6BFE04" box="[1152,1242,381,405]" metricMagnitude="3" metricUnit="m" metricValue="1.53" pageId="11" pageNumber="10" unit="m" value="1530.0">
<elevation id="0064D1C1FFF8FF99FB649F09FB6BFE04" box="[1152,1242,381,405]" metricMagnitude="3" metricUnit="m" metricValue="1.53" pageId="11" pageNumber="10" unit="m" value="1530.0">1530 m</elevation>
</quantity>
(Aug–Sep)
</materialsCitation>
;–
<materialsCitation id="3B213CAFFFF8FF99FA989F09FED9FE47" ID-GBIF-Occurrence="2575871583" collectionCode="SSW" elevation="1475" latitude="30.021666" location="Pinto Canyon Ranch" longLatPrecision="20" longitude="-104.46166" municipality="Marfa" pageId="11" pageNumber="10" specimenCode="FM 2810" specimenCount="1">
<emphasis id="B93DEAE0FFF8FF99FA989F09FA11FE04" bold="true" box="[1404,1440,381,405]" pageId="11" pageNumber="10">[4]</emphasis>
<location id="8E966029FFF8FF99FF249FE9FE0CFE24" LSID="urn:lsid:plazi:treatment:03E087E4FFF9FF99FF249B68FCDDFCA7:8E966029FFF8FF99FF249FE9FE0CFE24" box="[192,445,413,438]" latitude="30.021666" longLatPrecision="20" longitude="-104.46166" municipality="Marfa" name="Pinto Canyon Ranch" pageId="11" pageNumber="10">Pinto Canyon Ranch</location>
(~
<quantity id="4CB19B17FFF8FF99FE389FE9FD98FE24" box="[476,553,413,437]" metricMagnitude="4" metricUnit="m" metricValue="5.8" pageId="11" pageNumber="10" unit="km" value="58.0">58 km</quantity>
<collectionCode id="ED58AE37FFF8FF99FDD59FEAFDDCFE27" box="[561,621,414,438]" pageId="11" pageNumber="10">SSW</collectionCode>
<collectingMunicipality id="6B92AC88FFF8FF99FD909FE9FD0EFE24" box="[628,703,413,437]" pageId="11" pageNumber="10">Marfa</collectingMunicipality>
on
<specimenCode id="DBEF9E89FFF8FF99FD089FEAFCECFE27" box="[748,861,414,438]" collectionCode="FM" country="China" name="Department of Nature, Fujian Province Museum" pageId="11" pageNumber="10">FM 2810</specimenCode>
),
<geoCoordinate id="EE7D5035FFF8FF99FC959FEAFC4BFE24" box="[881,1018,412,438]" degrees="30" direction="north" minutes="01" orientation="latitude" pageId="11" pageNumber="10" precision="15" seconds="18" value="30.021666">30°01′18″N</geoCoordinate>
<geoCoordinate id="EE7D5035FFF8FF99FBE59FEAFB2CFE24" box="[1025,1181,412,438]" degrees="104" direction="west" minutes="27" orientation="longitude" pageId="11" pageNumber="10" precision="15" seconds="42" value="-104.46166">104°27′42″W</geoCoordinate>
(headquarters area),
<quantity id="4CB19B17FFF8FF99FF249FC9FEABFE44" box="[192,282,445,469]" metricMagnitude="3" metricUnit="m" metricValue="1.475" pageId="11" pageNumber="10" unit="m" value="1475.0">
<elevation id="0064D1C1FFF8FF99FF249FC9FEABFE44" box="[192,282,445,469]" metricMagnitude="3" metricUnit="m" metricValue="1.475" pageId="11" pageNumber="10" unit="m" value="1475.0">1475 m</elevation>
</quantity>
(Aug)
</materialsCitation>
.
</paragraph>
<paragraph id="8BF636F2FFF8FF99FF249F99FCCBFDB4" blockId="11.[192,1440,493,550]" pageId="11" pageNumber="10">
<emphasis id="B93DEAE0FFF8FF99FF249F99FE59FD94" bold="true" box="[192,488,493,517]" pageId="11" pageNumber="10">Collection method(s).</emphasis>
a) baited pitfall trap (human, laboratory rat feces); (b) incandescent and UV light trap; (c)*direct capture in nest of
<emphasis id="B93DEAE0FFF8FF99FD7D9C7AFCCBFDB4" box="[665,890,525,549]" italics="true" pageId="11" pageNumber="10">
<taxonomicName id="4C494D71FFF8FF99FD7D9C7AFCC6FDB4" authorityName="Hartley" authorityYear="1894" box="[665,887,525,549]" class="Mammalia" family="Muridae" genus="Neotoma" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="10" phylum="Chordata" rank="species" species="albigula">Neotoma albigula</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C3536579FFF8FF99FF249C49FD9FFDC7" box="[192,558,573,598]" pageId="11" pageNumber="10" type="diagnosis">
<paragraph id="8BF636F2FFF8FF99FF249C49FD9FFDC7" blockId="11.[192,558,573,598]" box="[192,558,573,598]" pageId="11" pageNumber="10">
<emphasis id="B93DEAE0FFF8FF99FF249C49FE12FDC4" bold="true" box="[192,419,573,597]" pageId="11" pageNumber="10">Surface activity.</emphasis>
Nocturnal.
</paragraph>
</subSubSection>
<subSubSection id="C3536579FFF8FF99FF249C19FE3AFD34" pageId="11" pageNumber="10" type="biology_ecology">
<paragraph id="8BF636F2FFF8FF99FF249C19FE3AFD34" blockId="11.[192,1440,621,678]" pageId="11" pageNumber="10">
<emphasis id="B93DEAE0FFF8FF99FF249C19FE80FD14" bold="true" box="[192,305,621,645]" pageId="11" pageNumber="10">Habitat.</emphasis>
In association with wood rats (
<taxonomicName id="4C494D71FFF8FF99FD219C1AFC80FD14" baseAuthorityName="McCleve and Kohlmann" baseAuthorityYear="2005" box="[709,817,622,645]" class="Mammalia" family="Muridae" genus="Neotoma" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="10" phylum="Chordata" rank="genus">
<emphasis id="B93DEAE0FFF8FF99FD219C1AFC80FD14" box="[709,817,622,645]" italics="true" pageId="11" pageNumber="10">Neotoma</emphasis>
</taxonomicName>
) in all zones, but more common in montane and grassland zones.
</paragraph>
</subSubSection>
<subSubSection id="C3536579FFF8FF99FF249CCAFCDDFCA7" pageId="11" pageNumber="10" type="discussion">
<paragraph id="8BF636F2FFF8FF99FF249CCAFCDDFCA7" blockId="11.[192,1440,702,822]" pageId="11" pageNumber="10">
<emphasis id="B93DEAE0FFF8FF99FF249CCAFEE8FD47" bold="true" box="[192,345,702,726]" pageId="11" pageNumber="10">Comments.</emphasis>
<taxonomicName id="4C494D71FFF8FF99FE849CCAFDCAFD47" authorityName="Horn" authorityYear="1875" box="[352,635,702,726]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="velutinus">
<emphasis id="B93DEAE0FFF8FF99FE849CCAFDCAFD47" box="[352,635,702,726]" italics="true" pageId="11" pageNumber="10">Onthophagus velutinus</emphasis>
</taxonomicName>
is a common species in southern
<collectingRegion id="498DF810FFF8FF99FBEF9CCAFBDAFD47" box="[1035,1131,702,726]" country="United States of America" name="Arizona" pageId="11" pageNumber="10">Arizona</collectingRegion>
and is recorded also from
<collectingRegion id="498DF810FFF8FF99FF249CAAFE80FD67" box="[192,305,734,758]" country="United States of America" name="Colorado" pageId="11" pageNumber="10">Colorado</collectingRegion>
. It is a close relative of
<taxonomicName id="4C494D71FFF8FF99FDAA9CAAFCDAFD67" authority="Howden" authorityName="Howden" authorityYear="1973" box="[590,875,734,758]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="cartwrighti">
<emphasis id="B93DEAE0FFF8FF99FDAA9CAAFD4DFD67" box="[590,764,734,758]" italics="true" pageId="11" pageNumber="10">O. cartwrighti</emphasis>
Howden
</taxonomicName>
, which occurs in southern
<collectingRegion id="498DF810FFF8FF99FB4B9CAAFA9AFD67" box="[1199,1323,734,758]" country="United States of America" name="California" pageId="11" pageNumber="10">California</collectingRegion>
and
<collectingRegion id="498DF810FFF8FF99FA839CAAFE8EFC87" country="Mexico" name="Baja California" pageId="11" pageNumber="10">Baja California</collectingRegion>
, of
<taxonomicName id="4C494D71FFF8FF99FE8D9C8AFCA2FC87" authority="Howden and Cartwright" authorityName="Howden and Cartwright" box="[361,787,766,790]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="arnetti">
<emphasis id="B93DEAE0FFF8FF99FE8D9C8AFECFFC84" box="[361,382,766,789]" italics="true" pageId="11" pageNumber="10">O</emphasis>
.
<emphasis id="B93DEAE0FFF8FF99FE699C8BFE6EFC87" box="[397,479,767,790]" italics="true" pageId="11" pageNumber="10">arnetti</emphasis>
Howden and Cartwright
</taxonomicName>
(southern
<collectingRegion id="498DF810FFF8FF99FC7E9C8AFC4FFC87" box="[922,1022,766,790]" country="United States of America" name="Arizona" pageId="11" pageNumber="10">Arizona</collectingRegion>
), as well as of
<taxonomicName id="4C494D71FFF8FF99FB559C8AFEB9FCA7" authority="(Howden 1973)" baseAuthorityName="Howden" baseAuthorityYear="1973" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="browni">
<emphasis id="B93DEAE0FFF8FF99FB559C8AFB77FC84" box="[1201,1222,766,789]" italics="true" pageId="11" pageNumber="10">O</emphasis>
.
<emphasis id="B93DEAE0FFF8FF99FB319C8AFA9BFC87" box="[1237,1322,766,790]" italics="true" pageId="11" pageNumber="10">browni</emphasis>
(
<bibRefCitation id="EFD84B03FFF8FF99FADE9C8AFF4EFCA7" author="Howden, H. F." pageId="11" pageNumber="10" pagination="329 - 337" refId="ref18192" refString="Howden, H. F. 1973. Four new species of Onthophagus from Mexico and the United States (Coleoptera: Scarabaeidae). Proceedings of the Entomological Society of Washington 75: 329 - 337." type="journal article" year="1973">Howden 1973</bibRefCitation>
)
</taxonomicName>
. (See also comments below concerning
<taxonomicName id="4C494D71FFF8FF99FD079D6AFCEDFCA7" baseAuthorityName="Howden" baseAuthorityYear="1973" box="[739,860,798,822]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="browni">
<emphasis id="B93DEAE0FFF8FF99FD079D6AFD49FCA4" box="[739,760,798,821]" italics="true" pageId="11" pageNumber="10">O</emphasis>
.
<emphasis id="B93DEAE0FFF8FF99FCE39D6AFCEDFCA7" box="[775,860,798,822]" italics="true" pageId="11" pageNumber="10">browni</emphasis>
</taxonomicName>
.)
</paragraph>
</subSubSection>
</treatment>
</document>
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