treatments-xml/data/03/E0/87/03E087C3697D7A29DDA82B920477FEF2.xml
2024-06-21 12:22:17 +02:00

206 lines
24 KiB
XML

<document id="E8732AF3AEBF2795BDDF60BBBFE6C3E2" ID-CLB-Dataset="55724" ID-DOI="10.5281/zenodo.4507307" ID-GBIF-Dataset="72a1f456-f29c-471e-baf8-6c026cc900a1" ID-ISSN="0253-116X" ID-RefBankID="1643FFB2ED42FFC2FFE4FFE28E27FFAF" ID-Zenodo-Dep="4507307" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612537541968" checkinUser="felipe" docAuthor="Assing, V." docDate="2013" docId="03E087C3697D7A29DDA82B920477FEF2" docLanguage="en" docName="LinzerbiolBeitr.45.1.0171-0190.pdf" docOrigin="Linzer biologische Beiträge 45 (1)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Rugilus (Rugilus) atronitidus Assing 2013, nov.sp." docType="treatment" docVersion="10" lastPageNumber="181" masterDocId="FFD9FFBB69747A23DD0B2B5B0548FFA4" masterDocTitle="A revision of Palaearctic and Oriental Rugilus. III. Five new species from the Palaearctic region and additional records (Coleoptera: Staphylinidae: Paederinae)" masterLastPageNumber="190" masterPageNumber="171" pageNumber="180" updateTime="1698867118016" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="992BDDBFFE138A6B041391EBC0813DD4" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A5660B38ED1ECB262DA72BBA3578E569">
<mods:title id="B6E86E3128620530E7B60DB8568B2F5F">A revision of Palaearctic and Oriental Rugilus. III. Five new species from the Palaearctic region and additional records (Coleoptera: Staphylinidae: Paederinae)</mods:title>
</mods:titleInfo>
<mods:name id="552775416CDCAB1A1F7790F6F4713758" type="personal">
<mods:role id="165DA4560E5C552C09ED764586E81882">
<mods:roleTerm id="71182E88994A66839BC180969F941ACF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0A5B747733F0C3D2C3DDA9D54D684EF5">Assing, V.</mods:namePart>
</mods:name>
<mods:typeOfResource id="13A399B80CA36719A6D28AF2F99A87A3">text</mods:typeOfResource>
<mods:relatedItem id="AB447207B367F3E22C1041E8AC483A99" type="host">
<mods:titleInfo id="39009D4DB6E48FD9696EDAE87465B8A1">
<mods:title id="6FCFC7B43DC62BCDDCAD041CE38B5790">Linzer biologische Beiträge</mods:title>
</mods:titleInfo>
<mods:part id="F52CA94F9EEF196FEDD49D842BC39DC7">
<mods:date id="AD738F172BFB4260E89A88F0726C3AE2">2013</mods:date>
<mods:detail id="540EED3C7B0A955F7643F81C58C69D8C" type="pubDate">
<mods:number id="558DB90E7DE046ACA498596A8FDA42A2">2013-07-31</mods:number>
</mods:detail>
<mods:detail id="D8DC0F099DAB50F1B1EBEDC58AAE6E25" type="volume">
<mods:number id="843AF7B1E41A3C3B294C04A20712DA97">45</mods:number>
</mods:detail>
<mods:detail id="D1A7F455F39990E95556D532326A0FD1" type="issue">
<mods:number id="B5E05AF231C18484B345EE32A5477E68">1</mods:number>
</mods:detail>
<mods:extent id="F4286EB410685EF9DB403FF44DAE312A" unit="page">
<mods:start id="B87290E1D2E0CEA045F0D688A562E252">171</mods:start>
<mods:end id="F8B4DC26863FE1FF72EA74203E968677">190</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="AD61FA9024CFA64A367E5E1DC6D0EB58">journal article</mods:classification>
<mods:identifier id="286E0CAD05FB79FF60DCD9581CE8DD03" type="CLB-Dataset">55724</mods:identifier>
<mods:identifier id="B147069CCD57FB2083C50494FC6C6000" type="DOI">10.5281/zenodo.4507307</mods:identifier>
<mods:identifier id="13A8FE0A049D18EC571BFA370DDCF37B" type="GBIF-Dataset">72a1f456-f29c-471e-baf8-6c026cc900a1</mods:identifier>
<mods:identifier id="FE3F825F6DB13923201264B0E0757A92" type="ISSN">0253-116X</mods:identifier>
<mods:identifier id="E57A3E9FE315912789D7FB236F88604B" type="RefBankID">1643FFB2ED42FFC2FFE4FFE28E27FFAF</mods:identifier>
<mods:identifier id="9DFCD2BE4E56CB8691EAAE435A9EC5D5" type="Zenodo-Dep">4507307</mods:identifier>
<mods:identifier id="098D086ED55AD2D08586A782E4D30F05" type="ZooBank">8683FCEB-A495-4147-A79E-996E04C23526</mods:identifier>
</mods:mods>
<treatment id="03E087C3697D7A29DDA82B920477FEF2" ID-DOI="http://doi.org/10.5281/zenodo.6521065" ID-GBIF-Taxon="195191770" ID-Zenodo-Dep="6521065" LSID="urn:lsid:plazi:treatment:03E087C3697D7A29DDA82B920477FEF2" httpUri="http://treatment.plazi.org/id/03E087C3697D7A29DDA82B920477FEF2" lastPageId="10" lastPageNumber="181" pageId="9" pageNumber="180">
<subSubSection id="C353655E697D7A2ADDA82B920760FF7B" box="[163,552,201,223]" pageId="9" pageNumber="180" type="nomenclature">
<paragraph id="8BF636D5697D7A2ADDA82B920760FF7B" blockId="9.[163,640,201,223]" box="[163,552,201,223]" pageId="9" pageNumber="180">
<heading id="D0BE81B9697D7A2ADDA82B920760FF7B" box="[163,552,201,223]" fontSize="8" level="2" pageId="9" pageNumber="180" reason="2">
<emphasis id="B93DEAC7697D7A2ADDA82B920760FF7B" bold="true" box="[163,552,201,223]" pageId="9" pageNumber="180">
<taxonomicName id="4C494D56697D7A2ADDA82B92049EFF7B" ID-CoL="4TNVK" authority="Assing, 2013" authorityName="Assing" authorityYear="2013" box="[163,470,201,223]" class="Insecta" family="Staphylinidae" genus="Rugilus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="180" phylum="Arthropoda" rank="species" species="atronitidus" status="sp. nov." subGenus="Rugilus">
<emphasis id="B93DEAC7697D7A2ADDA82B9205BCFF7B" bold="true" box="[163,244,201,223]" italics="true" pageId="9" pageNumber="180">Rugilus</emphasis>
(
<emphasis id="B93DEAC7697D7A2ADC082B92041DFF7B" bold="true" box="[259,341,201,223]" italics="true" pageId="9" pageNumber="180">Rugilus</emphasis>
)
<emphasis id="B93DEAC7697D7A2ADC6F2B92049EFF7B" bold="true" box="[356,470,201,223]" italics="true" pageId="9" pageNumber="180">atronitidus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20E57BC697D7A2ADCD72B920760FF7B" box="[476,552,201,223]" pageId="9" pageNumber="180" rank="species">nov.sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C353655E697D7A2ADF252B9207C8FF7B" box="[558,640,201,223]" pageId="9" pageNumber="180" type="description">
<paragraph id="8BF636D5697D7A2ADF252B9207C8FF7B" blockId="9.[163,640,201,223]" box="[558,640,201,223]" pageId="9" pageNumber="180">
<heading id="D0BE81B9697D7A2ADF252B9207C8FF7B" box="[558,640,201,223]" fontSize="8" level="2" pageId="9" pageNumber="180" reason="2">
(
<figureCitation id="13722A50697D7A2ADF3E2B92073EFF7B" box="[565,630,201,223]" captionStart="Map 1" captionStartId="7.[163,208,898,918]" captionTargetBox="[163,1069,196,879]" captionTargetId="figure-4@7.[163,1069,459,879]" captionTargetPageId="7" captionText="Map 1: Distributions of Rugilus gansuensis ROUGEMONT (triangles), R. confluens ASSING (diamonds), R. meilixuensis ASSING (circles), and R. atronitidus nov.sp. (star), based on examined records. Filled symbols: male-based records; open symbols: female-based records." figureDoi="http://doi.org/10.5281/zenodo.4507327" httpUri="https://zenodo.org/record/4507327/files/figure.png" pageId="9" pageNumber="180">Map 1</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C353655E697D7A2ADDA82BAB0707FE95" pageId="9" pageNumber="180" type="materials_examined">
<paragraph id="8BF636D5697D7A2ADDA82BAB0707FE95" blockId="9.[163,1070,238,305]" pageId="9" pageNumber="180">
Type material:
<materialsCitation id="3B213C88697D7A2ADCAF2BAB0707FE95" ID-GBIF-Occurrence="3762949381" collectingDate="2010-05-27" collectionCode="CAS" collectorName="V. Grebennikov" country="China" determinerName="V. Assing" elevation="3000" latitude="27.983612" location="Gaoligong Shan" longitude="98.54914" municipality="Gaoligong Shan" pageId="9" pageNumber="180" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="holotype">
<typeStatus id="54F28877697D7A2ADCAF2BAB04B0FEA0" box="[420,504,240,260]" pageId="9" pageNumber="180" type="holotype">Holotype</typeStatus>
<specimenCount id="9D4FFD5C697D7A2ADF0E2BB50750FEA7" box="[517,536,238,259]" pageId="9" pageNumber="180" type="male"></specimenCount>
: &quot;P. R.
<collectingCountry id="F35E7645697D7A2ADF7E2BAB07E6FEA0" box="[629,686,240,260]" name="China" pageId="9" pageNumber="180">China</collectingCountry>
,
<collectingRegion id="498DF837697D7A2ADFB52BAB0640FEA0" box="[702,776,240,260]" country="China" name="Yunnan" pageId="9" pageNumber="180">Yunnan</collectingRegion>
, E slope N
<collectingMunicipality id="6B92ACAF697D7A2ADE852BAB0163FEA0" box="[910,1067,240,260]" pageId="9" pageNumber="180">Gaoligong Shan</collectingMunicipality>
,
<geoCoordinate id="EE7D5012697D7A2ADDA82A5C0456FEBF" box="[163,286,263,283]" degrees="27" direction="north" minutes="59" orientation="latitude" pageId="9" pageNumber="180" precision="1" seconds="01.0" value="27.983612">N27°59'01.0''</geoCoordinate>
<geoCoordinate id="EE7D5012697D7A2ADC282A5C04E1FEBF" box="[291,425,263,283]" degrees="098" direction="east" minutes="32" orientation="longitude" pageId="9" pageNumber="180" precision="1" seconds="56.9" value="98.54914">E098°32'56.9''</geoCoordinate>
,
<date id="FFF71015697D7A2ADCB82A5C0758FEBF" box="[435,528,263,283]" pageId="9" pageNumber="180" value="2010-05-27">
<collectingDate id="EFB3E9FD697D7A2ADCB82A5C0758FEBF" box="[435,528,263,283]" pageId="9" pageNumber="180" value="2010-05-27">27.v.2010</collectingDate>
</date>
, 3018 m, sifting22,
<collectorName id="26BC5303697D7A2ADFCD2A5C0611FEBF" box="[710,857,263,283]" pageId="9" pageNumber="180">V. Grebennikov</collectorName>
/ Holotypus ♂
<originalDetermination id="A8578E29697D7A2ADEE32A5E0477FE95" pageId="9" pageNumber="180">
<taxonomicName id="4C494D56697D7A2ADEE32A5E044EFE95" authority="Assing, 2013" authorityName="Assing" authorityYear="2013" class="Insecta" family="Staphylinidae" genus="Rugilus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="180" phylum="Arthropoda" rank="species" species="atronitidus" status="sp. nov.">
<emphasis id="B93DEAC7697D7A2ADEE32A5E044EFE95" italics="true" pageId="9" pageNumber="180">Rugilus atronitidus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20E57BC697D7A2ADC072A460477FE95" box="[268,319,285,305]" pageId="9" pageNumber="180" rank="species">sp. n.</taxonomicNameLabel>
</originalDetermination>
, det.
<determinerName id="7E2ADA11697D7A2ADC7B2A460482FE95" box="[368,458,285,305]" pageId="9" pageNumber="180">V. Assing</determinerName>
2013&quot; (
<collectionCode id="ED58AE10697D7A2ADF1F2A46070DFE95" box="[532,581,285,305]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="9" pageNumber="180" type="Museum">CAS</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C353655E697D7A2ADDA82A1A06ADFED1" pageId="9" pageNumber="180" type="etymology">
<paragraph id="8BF636D5697D7A2ADDA82A1A06ADFED1" blockId="9.[163,1070,321,373]" pageId="9" pageNumber="180">
Etymology: The specific epithet (Latin, adjective) refers to the glossy elytra, one of the characters distinguishing this species from
<taxonomicName id="4C494D56697D7A2ADF932A3B0679FED1" box="[664,817,352,373]" class="Insecta" family="Staphylinidae" genus="Rugilus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="180" phylum="Arthropoda" rank="species" species="meilixuensis">
<emphasis id="B93DEAC7697D7A2ADF932A3B0679FED1" box="[664,817,352,373]" italics="true" pageId="9" pageNumber="180">R. meilixuensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C494D56697D7A2ADE692A3B0697FED1" box="[866,991,352,373]" class="Insecta" family="Staphylinidae" genus="Rugilus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="180" phylum="Arthropoda" rank="species" species="confluens">
<emphasis id="B93DEAC7697D7A2ADE692A3B0697FED1" box="[866,991,352,373]" italics="true" pageId="9" pageNumber="180">R. confluens</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C353655E697D7A2ADDA82ADE07B6FB7E" pageId="9" pageNumber="180" type="description">
<paragraph id="8BF636D5697D7A2ADDA82ADE071EFE57" blockId="9.[163,1070,389,499]" pageId="9" pageNumber="180">
Description: Body length
<quantity id="4CB19B30697D7A2ADF0C2ADE0711FE3F" box="[519,601,389,411]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="9" pageNumber="180" unit="mm" value="5.2">5.2 mm</quantity>
; length of forebody 3.0 mm. For illustrations of the habitus and the forebody see figures
<quantity id="4CB19B30697D7A2ADF712AF907F2FE1C" box="[634,698,418,440]" metricMagnitude="-2" metricUnit="m" metricValue="6.35" metricValueMax="7.62" metricValueMin="5.08" pageId="9" pageNumber="180" unit="in" value="2.5" valueMax="3.0" valueMin="2.0">2-3 in</quantity>
<bibRefCitation id="EFD84B24697D7A2ADFCE2AF90627FE1C" author="ASSING V" box="[709,879,418,440]" pageId="9" pageNumber="180" pagination="137 - 149" refId="ref11108" refString="ASSING V. (2012 b): A revision of Palaearctic and Oriental Rugilus LEACH, 1819. II. Three new species from China and additional records (Coleoptera: Staphylinidae: Paederinae). - Koleopterologische Rundschau 82: 137 - 149." type="journal article" year="2012">ASSING (2012b)</bibRefCitation>
. Coloration: body black; legs brown with somewhat darker femora; antennae pale-brown, with antennomeres I and II somewhat infuscate.
</paragraph>
<paragraph id="8BF636D5697D7A2ADDA82959049EFCA6" blockId="9.[163,1072,514,770]" pageId="9" pageNumber="180">Head 1.05 times as broad as long, broadest across eyes; margins behind eyes weakly convex, posterior angles obsolete; dorsal surface somewhat uneven, median dorsal portion transversely elevated, anterior and posterior to this elevation irregularly, transversely impressed; punctation coarse and dense, largely confluent in median dorsal portion; interstices mostly reduced to narrow ridges, but surface with some shine. Eyes large and bulging, approximately 0.7 times as long as distance from posterior margin of eyes to posterior constriction, and with distinct dense and dark pubescence. Anterior margin of labrum with two pronounced, basally fused, and strongly projecting teeth on either side of median incision.</paragraph>
<paragraph id="8BF636D5697D7A2ADDA82849041DFCC6" blockId="9.[163,1070,786,866]" pageId="9" pageNumber="180">Pronotum 1.15 times as long as broad and approximately 0.7 times as wide as head; punctation dense, coarse, and granulose; midline with short, indistinct, oblong glossy patch posteriorly.</paragraph>
<paragraph id="8BF636D5697D7A2ADDA8282A04BEFC7B" blockId="9.[163,1070,881,991]" pageId="9" pageNumber="180">Elytra long and large, approximately 1.12 times as long and 1.5 times as broad as pronotum, and distinctly glossy; punctation dense, moderately fine; interstices without microsculpture. Hind wings fully developed. Metatarsomere I approximately as long as the combined length of II and III.</paragraph>
<paragraph id="8BF636D5697D7A2ADDA828B40685FBF8" blockId="9.[163,1070,1007,1116]" pageId="9" pageNumber="180">Abdomen distinctly narrower than elytra; tergites III-VI with shallow impressions anteriorly, these impressions with coarse punctation; punctation of remaining tergal surfaces fine and moderately dense; tergite VII with very shallow, tergites III-VI without appreciable microsculpture; posterior margin of tergite VII with palisade fringe.</paragraph>
<paragraph id="8BF636D5697D7A2ADDA82F3707B6FB7E" blockId="9.[163,1070,1132,1242]" pageId="9" pageNumber="180">
♂: sternite VII (
<bibRefCitation id="EFD84B24697D7A2ADC402F3704ACFB26" author="ASSING V" box="[331,484,1132,1154]" pageId="9" pageNumber="180" pagination="137 - 149" refId="ref11108" refString="ASSING V. (2012 b): A revision of Palaearctic and Oriental Rugilus LEACH, 1819. II. Three new species from China and additional records (Coleoptera: Staphylinidae: Paederinae). - Koleopterologische Rundschau 82: 137 - 149." type="journal article" year="2012">ASSING 2012b</bibRefCitation>
: figure 5) distinctly transverse and with broadly concave posterior margin; sternite VIII (
<bibRefCitation id="EFD84B24697D7A2ADCE12FD207CCFB3B" author="ASSING V" box="[490,644,1161,1183]" pageId="9" pageNumber="180" pagination="137 - 149" refId="ref11108" refString="ASSING V. (2012 b): A revision of Palaearctic and Oriental Rugilus LEACH, 1819. II. Three new species from China and additional records (Coleoptera: Staphylinidae: Paederinae). - Koleopterologische Rundschau 82: 137 - 149." type="journal article" year="2012">ASSING 2012b</bibRefCitation>
: figure 4) with moderately deep, broadly V-shaped posterior excision; aedeagus (
<bibRefCitation id="EFD84B24697D7A2ADF342FFD0790FB18" author="ASSING V" box="[575,728,1190,1212]" pageId="9" pageNumber="180" pagination="137 - 149" refId="ref11108" refString="ASSING V. (2012 b): A revision of Palaearctic and Oriental Rugilus LEACH, 1819. II. Three new species from China and additional records (Coleoptera: Staphylinidae: Paederinae). - Koleopterologische Rundschau 82: 137 - 149." type="journal article" year="2012">ASSING 2012b</bibRefCitation>
: figure 6)
<quantity id="4CB19B30697D7A2ADE4C2FFD06EAFB18" box="[839,930,1190,1212]" metricMagnitude="-4" metricUnit="m" metricValue="6.1" pageId="9" pageNumber="180" unit="mm" value="0.61">0.61 mm</quantity>
long; ventral process of similar shape as in
<taxonomicName id="4C494D56697D7A2ADCD92F9F07B2FB7E" authority="ASSING 2012" authorityName="ASSING" authorityYear="2012" box="[466,762,1220,1242]" class="Insecta" family="Staphylinidae" genus="Rugilus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="180" phylum="Arthropoda" rank="species" species="meilixuensis">
<emphasis id="B93DEAC7697D7A2ADCD92F9F0723FB7D" box="[466,619,1220,1241]" italics="true" pageId="9" pageNumber="180">R. meilixuensis</emphasis>
ASSING 2012
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C353655E697D7A2ADDA82FB20781FAD0" pageId="9" pageNumber="180" type="discussion">
<paragraph id="8BF636D5697D7A2ADDA82FB20781FAD0" blockId="9.[163,1071,1257,1396]" pageId="9" pageNumber="180">
Comment: The
<typeStatus id="54F28877697D7A2ADC812FB204ABFB5B" box="[394,483,1257,1279]" pageId="9" pageNumber="180" type="holotype">holotype</typeStatus>
had previously been identified and illustrated as
<taxonomicName id="4C494D56697D7A2AD9132FB10468FABF" class="Insecta" family="Staphylinidae" genus="Rugilus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="180" phylum="Arthropoda" rank="species" species="meilixuensis">
<emphasis id="B93DEAC7697D7A2AD9132FB10468FABF" italics="true" pageId="9" pageNumber="180">R. meilixuensis</emphasis>
</taxonomicName>
, primarily based on the similar shape of the aedeagus (
<bibRefCitation id="EFD84B24697D7A2ADE8D2E5D016BFAB8" author="ASSING V" box="[902,1059,1286,1308]" pageId="9" pageNumber="180" pagination="137 - 149" refId="ref11108" refString="ASSING V. (2012 b): A revision of Palaearctic and Oriental Rugilus LEACH, 1819. II. Three new species from China and additional records (Coleoptera: Staphylinidae: Paederinae). - Koleopterologische Rundschau 82: 137 - 149." type="journal article" year="2012">ASSING 2012b</bibRefCitation>
). However, a re-examination of this specimen and of material of
<taxonomicName id="4C494D56697D7A2ADE412E7F06AFFA9D" box="[842,999,1316,1337]" class="Insecta" family="Staphylinidae" genus="Rugilus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="180" phylum="Arthropoda" rank="species" species="meilixuensis">
<emphasis id="B93DEAC7697D7A2ADE412E7F06AFFA9D" box="[842,999,1316,1337]" italics="true" pageId="9" pageNumber="180">R. meilixuensis</emphasis>
</taxonomicName>
in the course of a comparative study of the variation of
<taxonomicName id="4C494D56697D7A2ADFB32E190671FAF2" box="[696,825,1345,1367]" class="Insecta" family="Staphylinidae" genus="Rugilus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="180" phylum="Arthropoda" rank="species" species="confluens">
<emphasis id="B93DEAC7697D7A2ADFB32E190671FAF2" box="[696,825,1345,1367]" italics="true" pageId="9" pageNumber="180">R. confluens</emphasis>
</taxonomicName>
revealed that the male from the Gaoligong
<collectingRegion id="498DF837697D7A2ADC7A2E0504ECFAD0" box="[369,420,1374,1396]" country="Myanmar" name="Shan" pageId="9" pageNumber="180">Shan</collectingRegion>
represents a distinct species.
</paragraph>
</subSubSection>
<subSubSection id="C353655E697D7A2ADDA82ED8044FF988" pageId="9" pageNumber="180" type="diagnosis">
<paragraph id="8BF636D5697D7A2ADDA82ED8044FF988" blockId="9.[163,1070,1411,1580]" pageId="9" pageNumber="180">
Comparative notes
<taxonomicName id="4C494D56697D7A2ADF112ED807A6FA3C" authorityName="Assing" authorityYear="2013" box="[538,750,1411,1432]" class="Insecta" family="Staphylinidae" genus="Rugilus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="180" phylum="Arthropoda" rank="species" species="atronitidus">
<emphasis id="B93DEAC7697D7A2ADF112ED807A6FA3C" box="[538,750,1411,1432]" italics="true" pageId="9" pageNumber="180">Rugilus atronitidus</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="4C494D56697D7A2AD9132EDF0468FA12" class="Insecta" family="Staphylinidae" genus="Rugilus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="180" phylum="Arthropoda" rank="species" species="meilixuensis">
<emphasis id="B93DEAC7697D7A2AD9132EDF0468FA12" italics="true" pageId="9" pageNumber="180">R. meilixuensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C494D56697D7A2ADC542EFA04ABFA12" box="[351,483,1441,1462]" class="Insecta" family="Staphylinidae" genus="Rugilus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="180" phylum="Arthropoda" rank="species" species="confluens">
<emphasis id="B93DEAC7697D7A2ADC542EFA04ABFA12" box="[351,483,1441,1462]" italics="true" pageId="9" pageNumber="180">R. confluens</emphasis>
</taxonomicName>
by the completely black coloration of the body, the distinctly more shiny, less densely and more finely punctate elytra, the more glossy head and pronotum, a shorter head, the denser and more distinct pubescence of the eyes, the more uneven dorsal surface of the head, and by the slightly different shape of the aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="C353655E697E7A29DDA82B900477FEF2" pageId="10" pageNumber="181" type="distribution">
<paragraph id="8BF636D5697E7A29DDA82B900477FEF2" blockId="10.[163,1070,203,343]" pageId="10" pageNumber="181">
Distribution and natural history:The type locality is situated in the northern Gaoligong Shan in the northwest of
<collectingRegion id="498DF837697E7A29DFBE2BB20622FF5B" box="[693,874,233,255]" country="China" name="Yunnan" pageId="10" pageNumber="181">Yunnan province</collectingRegion>
,
<collectingCountry id="F35E7645697E7A29DE722BB206F0FF5B" box="[889,952,233,255]" name="China" pageId="10" pageNumber="181">China</collectingCountry>
, where the
<typeStatus id="54F28877697E7A29DDA82A5D05B4FEB8" box="[163,252,262,284]" pageId="10" pageNumber="181" type="holotype">holotype</typeStatus>
was sifted at an altitude of approximately
<quantity id="4CB19B30697E7A29DFD62A5D067AFEB8" box="[733,818,262,284]" metricMagnitude="3" metricUnit="m" metricValue="3.02" pageId="10" pageNumber="181" unit="m" value="3020.0">3020 m</quantity>
. The pronounced wing dimorphism in closely related species suggests that
<taxonomicName id="4C494D56697E7A29DFE62A7F06C9FE9C" authorityName="Assing" authorityYear="2013" box="[749,897,291,313]" class="Insecta" family="Staphylinidae" genus="Rugilus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="181" phylum="Arthropoda" rank="species" species="atronitidus">
<emphasis id="B93DEAC7697E7A29DFE62A7F06C9FE9C" box="[749,897,291,313]" italics="true" pageId="10" pageNumber="181">R. atronitidus</emphasis>
</taxonomicName>
may be wingdimorphic, too.
</paragraph>
</subSubSection>
</treatment>
</document>