treatments-xml/data/03/E0/87/03E087C3697B7A32DDA82F8F0627FEB8.xml
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<mods:title id="E7516DE115290F228522050863332730">A revision of Palaearctic and Oriental Rugilus. III. Five new species from the Palaearctic region and additional records (Coleoptera: Staphylinidae: Paederinae)</mods:title>
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<mods:namePart id="B74C358F6430A2DB8B66459F85F97DA6">Assing, V.</mods:namePart>
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<treatment id="03E087C3697B7A32DDA82F8F0627FEB8" ID-DOI="http://doi.org/10.5281/zenodo.4507560" ID-GBIF-Taxon="195191772" ID-Zenodo-Dep="4507560" LSID="urn:lsid:plazi:treatment:03E087C3697B7A32DDA82F8F0627FEB8" httpUri="http://treatment.plazi.org/id/03E087C3697B7A32DDA82F8F0627FEB8" lastPageId="17" lastPageNumber="188" pageId="15" pageNumber="186">
<subSubSection id="C353655E697B7A2CDDA82F8F0483FB4E" box="[163,459,1236,1258]" pageId="15" pageNumber="186" type="nomenclature">
<paragraph id="8BF636D5697B7A2CDDA82F8F0483FB4E" blockId="15.[163,669,1236,1258]" box="[163,459,1236,1258]" pageId="15" pageNumber="186">
<heading id="D0BE81B9697B7A2CDDA82F8F0483FB4E" box="[163,459,1236,1258]" fontSize="8" level="2" pageId="15" pageNumber="186" reason="2">
<taxonomicName id="4C494D56697B7A2CDDA82F8F0483FB4E" ID-CoL="4TP39" authority="ASSING 2012" authorityName="ASSING" authorityYear="2012" box="[163,459,1236,1258]" class="Insecta" family="Staphylinidae" genus="Rugilus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="186" phylum="Arthropoda" rank="species" species="lucens">
<emphasis id="B93DEAC7697B7A2CDDA82F8F0483FB4E" bold="true" box="[163,459,1236,1258]" pageId="15" pageNumber="186">
<emphasis id="B93DEAC7697B7A2CDDA82F8F0476FB4E" bold="true" box="[163,318,1236,1258]" italics="true" pageId="15" pageNumber="186">Rugilus lucens</emphasis>
ASSING 2012
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C353655E697B7A2CDCDA2F8F07D5FB4E" box="[465,669,1236,1258]" pageId="15" pageNumber="186" type="description">
<paragraph id="8BF636D5697B7A2CDCDA2F8F07D5FB4E" blockId="15.[163,669,1236,1258]" box="[465,669,1236,1258]" pageId="15" pageNumber="186">
<heading id="D0BE81B9697B7A2CDCDA2F8F07D5FB4E" box="[465,669,1236,1258]" fontSize="8" level="2" pageId="15" pageNumber="186" reason="2">
(
<figureCitation id="13722A50697B7A2CDCD32F8F0700FB4E" box="[472,584,1236,1258]" captionStart="Figs 23-30" captionStartId="16.[163,203,1425,1445]" captionTargetBox="[168,1061,201,1403]" captionTargetId="figure-92@16.[163,1070,197,1407]" captionTargetPageId="16" captionText="Figs 23-30: Rugilus lucens ASSING (23-25) and R. saudicus nov.sp. (26-30): (23, 27) male sternite VIII; (24, 28) aedeagus in lateral view; (25, 29) aedeagus in ventral view; (26) forebody; (30) apical portion of aedeagus in lateral view. Scale bars: 26: 1.0 mm; 23, 27: 0.2 mm; 24-25, 28-30: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4507339" httpUri="https://zenodo.org/record/4507339/files/figure.png" pageId="15" pageNumber="186">Figs 23-25</figureCitation>
,
<figureCitation id="13722A50697B7A2CDF582F8E07DEFB4E" box="[595,662,1236,1258]" captionStart="Map 4" captionStartId="15.[163,208,1152,1172]" captionTargetBox="[163,1070,395,1132]" captionTargetId="figure-83@15.[163,1070,756,1132]" captionTargetPageId="15" captionText="Map 4: Distributions of Rugilus pygmaeus (KRAATZ) (triangles) and R. lucens ASSING (circles), based on examined records." figureDoi="http://doi.org/10.5281/zenodo.6518966" httpUri="https://zenodo.org/record/6518966/files/figure.png" pageId="15" pageNumber="186">Map 4</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C353655E697B7A2CDDA82E58070BFAE1" pageId="15" pageNumber="186" type="materials_examined">
<paragraph id="8BF636D5697B7A2CDDA82E58070BFAE1" blockId="15.[163,1072,1283,1349]" pageId="15" pageNumber="186">
Material examined:
<materialsCitation id="3B213C88697B7A2CDCFB2E58070BFAE1" ID-GBIF-Occurrence="3762949372" latitude="15.233334" longitude="106.53333" pageId="15" pageNumber="186">
<collectingCountry id="F35E7645697B7A2CDCFB2E580756FAB3" box="[496,542,1283,1303]" name="Laos" pageId="15" pageNumber="186">Laos</collectingCountry>
:
<specimenCount id="9D4FFD5C697B7A2CDF262E580722FAB3" box="[557,618,1283,1303]" pageId="15" pageNumber="186" type="generic">6 exs.</specimenCount>
,
<collectingRegion id="498DF837697B7A2CDF722E580672FAB3" box="[633,826,1283,1303]" country="Laos" name="Champasak" pageId="15" pageNumber="186">Champasak province</collectingRegion>
, Bolaven plateau, Muang Paxong, Ban Thongvay,
<geoCoordinate id="EE7D5012697B7A2CDCBC2E410749FA8A" box="[439,513,1306,1326]" degrees="15" direction="north" minutes="14" orientation="latitude" pageId="15" pageNumber="186" precision="925" value="15.233334">15°14'N</geoCoordinate>
,
<geoCoordinate id="EE7D5012697B7A2CDF042E410729FA8A" box="[527,609,1306,1326]" degrees="106" direction="east" minutes="32" orientation="longitude" pageId="15" pageNumber="186" precision="925" value="106.53333">106°32'E</geoCoordinate>
,
<quantity id="4CB19B30697B7A2CDF652E4107A2FA8A" box="[622,746,1306,1326]" metricMagnitude="3" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="15" pageNumber="186" unit="m" value="1100.0" valueMax="1200.0" valueMin="1000.0">1000-1200 m</quantity>
, disturbed primary rain forest, leg. Solodovnikov &amp; Pedersen (
<collectionCode id="ED58AE10697B7A2CDCB42E6A074AFAE1" box="[447,514,1329,1349]" pageId="15" pageNumber="186">ZMUC</collectionCode>
,
<collectionCode id="ED58AE10697B7A2CDF012E6A0770FAE1" box="[522,568,1329,1349]" pageId="15" pageNumber="186">cAss</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C353655E697B7A32DDA82E0F0627FEB8" lastPageId="17" lastPageNumber="188" pageId="15" pageNumber="186" type="discussion">
<paragraph id="8BF636D5697B7A2CDDA82E0F045AF9BE" blockId="15.[163,1070,1364,1562]" pageId="15" pageNumber="186">
Comment: The original description of this conspicuous species is based on two females from
<collectingCountry id="F35E7645697B7A2CDC3A2E2A0425FA23" box="[305,365,1393,1415]" name="Nepal" pageId="15" pageNumber="186">Nepal</collectingCountry>
(
<bibRefCitation id="EFD84B24697B7A2CDC762E2A075CFA23" author="ASSING V" box="[381,532,1393,1415]" pageId="15" pageNumber="186" pagination="115 - 190" refId="ref11068" refString="ASSING V. (2012 a): The Rugilus species of the Palaearctic and Oriental regions (Coleoptera: Staphylinidae: Paederinae). - Stuttgarter Beitrage zur Naturkunde A, Neue Serie 5: 115 - 190." type="journal article" year="2012">ASSING 2012a</bibRefCitation>
). The above first record from
<collectingCountry id="F35E7645697B7A2CDE402E290634FA23" box="[843,892,1394,1415]" name="Laos" pageId="15" pageNumber="186">Laos</collectingCountry>
suggests that the species is widespread in the East Palaearctic and Oriental regions (
<figureCitation id="13722A50697B7A2CDE962ED506A3FA00" box="[925,1003,1422,1444]" captionStart="Map 4" captionStartId="15.[163,208,1152,1172]" captionTargetBox="[163,1070,395,1132]" captionTargetId="figure-83@15.[163,1070,756,1132]" captionTargetPageId="15" captionText="Map 4: Distributions of Rugilus pygmaeus (KRAATZ) (triangles) and R. lucens ASSING (circles), based on examined records." figureDoi="http://doi.org/10.5281/zenodo.6518966" httpUri="https://zenodo.org/record/6518966/files/figure.png" pageId="15" pageNumber="186">Map 4</figureCitation>
). The subgeneric affiliations are still unclarified, but its hypothesized close relationship to
<taxonomicName id="4C494D56697B7A2CD9132EF605B3FA7A" authorityName="Assing" authorityYear="2013" class="Insecta" family="Staphylinidae" genus="Rugilus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="186" phylum="Arthropoda" rank="species" species="saudicus">
<emphasis id="B93DEAC7697B7A2CD9132EF605B3FA7A" italics="true" pageId="15" pageNumber="186">R. saudicus</emphasis>
</taxonomicName>
(see the following section) suggests that both species may have their closest relatives in the Afrotropical region. The previously unknown male sexual characters are as follows:
</paragraph>
<caption id="DF36665D69647A33DDA82ECA05A7FA4E" ID-DOI="http://doi.org/10.5281/zenodo.4507339" ID-Zenodo-Dep="4507339" httpUri="https://zenodo.org/record/4507339/files/figure.png" pageId="16" pageNumber="187" startId="16.[163,203,1425,1445]" targetBox="[168,1061,201,1403]" targetPageId="16">
<paragraph id="8BF636D569647A33DDA82ECA05A7FA4E" blockId="16.[163,1071,1424,1514]" pageId="16" pageNumber="187">
<emphasis id="B93DEAC769647A33DDA82ECA044FFA01" bold="true" box="[163,263,1425,1445]" pageId="16" pageNumber="187">Figs 23-30</emphasis>
:
<taxonomicName id="4C494D5669647A33DC182ECB04AEFA00" authority="ASSING" authorityName="ASSING" authorityYear="2012" box="[275,486,1424,1445]" class="Insecta" family="Staphylinidae" genus="Rugilus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="187" phylum="Arthropoda" rank="species" species="lucens">
<emphasis id="B93DEAC769647A33DC182ECB04D1FA01" box="[275,409,1424,1445]" italics="true" pageId="16" pageNumber="187">Rugilus lucens</emphasis>
ASSING
</taxonomicName>
(
<emphasis id="B93DEAC769647A33DCFF2ECA0762FA01" bold="true" box="[500,554,1425,1445]" pageId="16" pageNumber="187">23-25</emphasis>
) and
<taxonomicName id="4C494D5669647A33DF6B2ECA0782FA01" authority="Assing, 2013" authorityName="Assing" authorityYear="2013" box="[608,714,1424,1446]" class="Insecta" family="Staphylinidae" genus="Rugilus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="187" phylum="Arthropoda" rank="species" species="saudicus" status="sp. nov.">
<emphasis id="B93DEAC769647A33DF6B2ECA0782FA01" box="[608,714,1424,1446]" italics="true" pageId="16" pageNumber="187">R. saudicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20E57BC69647A33DFC42ECA065AFA01" box="[719,786,1425,1445]" pageId="16" pageNumber="187" rank="species">nov.sp.</taxonomicNameLabel>
(
<emphasis id="B93DEAC769647A33DE142ECA061DFA01" bold="true" box="[799,853,1425,1445]" pageId="16" pageNumber="187">26-30</emphasis>
): (
<emphasis id="B93DEAC769647A33DE7B2ECA06E2FA01" bold="true" box="[880,938,1425,1445]" pageId="16" pageNumber="187">23, 27</emphasis>
) male sternite VIII; (
<emphasis id="B93DEAC769647A33DDEA2EF30456FA18" bold="true" box="[225,286,1448,1468]" pageId="16" pageNumber="187">24, 28</emphasis>
) aedeagus in lateral view; (
<emphasis id="B93DEAC769647A33DF272EF30722FA18" bold="true" box="[556,618,1448,1468]" pageId="16" pageNumber="187">25, 29</emphasis>
) aedeagus in ventral view; (
<emphasis id="B93DEAC769647A33DE752EF306DDFA18" bold="true" box="[894,917,1448,1468]" pageId="16" pageNumber="187">26</emphasis>
) forebody; (
<emphasis id="B93DEAC769647A33D91B2EF3016FFA18" bold="true" box="[1040,1063,1448,1468]" pageId="16" pageNumber="187">30</emphasis>
) apical portion of aedeagus in lateral view. Scale bars: 26: 1.0 mm; 23, 27: 0.2 mm; 24-25, 28-30: 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BF636D569657A32DDA82B970627FEB8" blockId="17.[163,1071,203,284]" pageId="17" pageNumber="188">
♂: sternite VII transverse and with broadly concave posterior margin; sternite VIII (
<figureCitation id="13722A5069657A32D90E2B9005F5FF5B" captionStart="Figs 23-30" captionStartId="16.[163,203,1425,1445]" captionTargetBox="[168,1061,201,1403]" captionTargetId="figure-92@16.[163,1070,197,1407]" captionTargetPageId="16" captionText="Figs 23-30: Rugilus lucens ASSING (23-25) and R. saudicus nov.sp. (26-30): (23, 27) male sternite VIII; (24, 28) aedeagus in lateral view; (25, 29) aedeagus in ventral view; (26) forebody; (30) apical portion of aedeagus in lateral view. Scale bars: 26: 1.0 mm; 23, 27: 0.2 mm; 24-25, 28-30: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4507339" httpUri="https://zenodo.org/record/4507339/files/figure.png" pageId="17" pageNumber="188">Fig. 23</figureCitation>
) with dense pubescence, posteriorly with broad and shallow excision; aedeagus (
<figureCitation id="13722A5069657A32D90A2BB20597FEB8" captionStart="Figs 23-30" captionStartId="16.[163,203,1425,1445]" captionTargetBox="[168,1061,201,1403]" captionTargetId="figure-92@16.[163,1070,197,1407]" captionTargetPageId="16" captionText="Figs 23-30: Rugilus lucens ASSING (23-25) and R. saudicus nov.sp. (26-30): (23, 27) male sternite VIII; (24, 28) aedeagus in lateral view; (25, 29) aedeagus in ventral view; (26) forebody; (30) apical portion of aedeagus in lateral view. Scale bars: 26: 1.0 mm; 23, 27: 0.2 mm; 24-25, 28-30: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4507339" httpUri="https://zenodo.org/record/4507339/files/figure.png" pageId="17" pageNumber="188">Figs 24-25</figureCitation>
) approximately
<quantity id="4CB19B3069657A32DC8E2A5D049AFEB8" box="[389,466,262,284]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="17" pageNumber="188" unit="mm" value="0.5">0.5 mm</quantity>
long, with pair of long apical structures.
</paragraph>
</subSubSection>
</treatment>
</document>