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<document id="4A26373ACA092CD229780179A17C417D" ID-CLB-Dataset="42318" ID-DOI="10.11646/zootaxa.3878.4.1" ID-GBIF-Dataset="e778cf07-b718-454a-98d1-1a9edbe071aa" ID-ISSN="1175-5326" ID-Zenodo-Dep="252701" ID-ZooBank="B6AF841A-2D56-4F76-847F-44E881DF38B5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460947067534" checkinUser="plazi" docAuthor="Watters, G. Thomas" docDate="2014" docId="03E087C1FFAB0273F681FF55FE602A55" docLanguage="en" docName="zt03878p350.pdf" docOrigin="Zootaxa 3878 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Halotudora gaigei Bequaert &amp; Clench 1931" docType="treatment" docVersion="8" lastPageNumber="310" masterDocId="FFD9FFB9FFA3027AF616FFCDFFF62C2E" masterDocTitle="A revision of the Annulariidae of Central America (Gastropoda: Littorinoidea)" masterLastPageNumber="350" masterPageNumber="301" pageNumber="309" updateTime="1698584952024" updateUser="ExternalLinkService">
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<mods:title id="36FC723F3E9589E34A8E71DC65FE7EEF">A revision of the Annulariidae of Central America (Gastropoda: Littorinoidea)</mods:title>
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<mods:namePart id="E3A10581F8BEA7F50A2BB53D933D1DC9">Watters, G. Thomas</mods:namePart>
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<mods:typeOfResource id="35535C281865FFD3B605A756E468DC61">text</mods:typeOfResource>
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<mods:date id="49831348728841FD9E746CE8EE8B0DCF">2014</mods:date>
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<mods:classification id="CA85B740578BAB831EA72EFB01CB4D59">journal article</mods:classification>
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<mods:identifier id="D0408C929490D7EE0C9613279831E5CF" type="DOI">10.11646/zootaxa.3878.4.1</mods:identifier>
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<treatment id="03E087C1FFAB0273F681FF55FE602A55" ID-DOI="http://doi.org/10.5281/zenodo.6134480" ID-GBIF-Taxon="119591814" ID-Zenodo-Dep="6134480" LSID="urn:lsid:plazi:treatment:03E087C1FFAB0273F681FF55FE602A55" httpUri="http://treatment.plazi.org/id/03E087C1FFAB0273F681FF55FE602A55" lastPageId="9" lastPageNumber="310" pageId="8" pageNumber="309">
<subSubSection id="C353655CFFAB0272F681FF55FE772CFC" pageId="8" pageNumber="309" type="nomenclature">
<paragraph id="8BF636D7FFAB0272F681FF55FD3F2C9C" blockId="8.[151,713,152,211]" box="[151,713,152,178]" pageId="8" pageNumber="309">
<heading id="D0BE81BBFFAB0272F681FF55FD3F2C9C" bold="true" box="[151,713,152,178]" fontSize="11" level="1" pageId="8" pageNumber="309" reason="1">
<taxonomicName id="4C494D54FFAB0272F681FF55FD3F2C9C" authority="Bequaert &amp; Clench, 1931" authorityName="Bequaert &amp; Clench" authorityYear="1931" box="[151,713,152,178]" class="Gastropoda" family="Annulariidae" genus="Halotudora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="8" pageNumber="309" phylum="Mollusca" rank="species" species="gaigei">
<emphasis id="B93DEAC5FFAB0272F681FF55FD3F2C9C" bold="true" box="[151,713,152,178]" pageId="8" pageNumber="309">
<emphasis id="B93DEAC5FFAB0272F681FF55FE852C9C" bold="true" box="[151,371,152,178]" italics="true" pageId="8" pageNumber="309">Halotudora gaigei</emphasis>
(
<bibRefCitation id="EFD84B26FFAB0272F794FF55FD342C9C" author="Bequaert" box="[386,706,152,178]" pageId="8" pageNumber="309" refString="Bequaert, J. &amp; Clench, W. J. (1931) Three new terrestrial snails from Yucatan. Occasional Papers of the Boston Society of Natural History, 5, 423 - 426." type="journal article" year="1931">Bequaert &amp; Clench, 1931</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BF636D7FFAB0272F681FF77FE772CFC" blockId="8.[151,713,152,211]" box="[151,385,186,211]" pageId="8" pageNumber="309">
<figureCitation id="13722A52FFAB0272F681FF77FEB62CFD" box="[151,320,186,211]" captionStart="FIGURE 2 A – K" captionStartId="9.[151,250,977,999]" captionTargetBox="[151,1436,193,955]" captionTargetId="figure@9.[151,1436,193,956]" captionTargetPageId="9" captionText="FIGURE 2 A – K. Halotudora gaigei (Bequaert &amp; Clench, 1931). A. UMMZ 49190, holotype, 14.5 mm. B – C. UF 19088, 15.2 mm. D. UF 155822, 13.4 mm. E. UF 133597, 16.6 mm. F. UF 19092, 15.7 mm. G. UF 19152, 13.5 mm. H. GTW 13724 c, 15.6 mm. I. GTW 13725 b, 14.0 mm [operculum glued in aperture backward by collector]. J. GTW 13725 b, 13.2 mm. K. GTW 13725 a, 16.1 mm." httpUri="https://zenodo.org/record/252703/files/figure.png" pageId="8" pageNumber="309">Figures 2 A–K</figureCitation>
,
<figureCitation id="13722A52FFAB0272F75DFF76FE772CFC" box="[331,385,187,211]" captionStart="FIGURE 12 A – D" captionStartId="43.[151,250,1358,1380]" captionTargetBox="[151,1436,195,1336]" captionTargetId="figure@43.[151,1436,195,1336]" captionTargetPageId="43" captionText="FIGURE 12 A – D. Distribution maps. A. Choanopomops largillierti (Pfeiffer, 1846) (black); Parachondria rubicundus (Morelet, 1849) (red); Parachondria cordovanus Pfeiffer, 1857 (blue); Paridoxipoma enigmaticum new species (white). B. Halotudora sumichrasti (Crosse &amp; Fischer, 1874) (blue); Halotudora gaigei (Bequaert &amp; Clench, 1931) (black); Diplopoma osberti (Tristram, 1861) (red); Halotudora gruneri (Pfeiffer, 1846) (white). C. Gouldipoma chiapasense (Crosse &amp; Fischer, 1877) (red); Diplopoma rigidulum (Morelet, 1851) (black); Tudorisca andrewsae (Ancey, 1886) (blue); Gouldipoma coltrorum new species (white); Gouldipoma thomasi (Solem, 1961) (purple). D. Gouldipoma trochlearis (Pfeiffer, 1852) (red); Halotudora kuesteri (Pfeiffer, 1852) (black); Gouldipoma terecostatum (Thompson, 1966) (blue); Gouldipoma chrysostiria new species (purple); Gouldipoma callipeplum (Solem, 1961) (white)." httpUri="https://zenodo.org/record/252713/files/figure.png" pageId="8" pageNumber="309">12 B</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C353655CFFAB0272F681FECFFE922DA9" pageId="8" pageNumber="309" type="materials_examined">
<paragraph id="8BF636D7FFAB0272F681FECFFAC22D35" blockId="8.[151,1437,258,1975]" box="[151,1332,258,283]" pageId="8" pageNumber="309">
<emphasis id="B93DEAC5FFAB0272F681FECFFEB32D35" bold="true" box="[151,325,258,283]" pageId="8" pageNumber="309">
<typeStatus id="54F28875FFAB0272F681FECFFF242D35" box="[151,210,258,283]" pageId="8" pageNumber="309">Type</typeStatus>
material.
</emphasis>
<collectionCode id="ED58AE12FFAB0272F75AFECEFE552D34" box="[332,419,259,282]" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="8" pageNumber="309">UMMZ</collectionCode>
49190,
<typeStatus id="54F28875FFAB0272F7EBFECFFDA92D35" box="[509,607,258,283]" pageId="8" pageNumber="309" type="holotype">holotype</typeStatus>
(fig. 2 A);
<collectionCode id="ED58AE12FFAB0272F4C8FECEFCEC2D35" LSID="urn:lsid:biocol.org:col:33791" box="[734,794,259,283]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="8" pageNumber="309">MCZ</collectionCode>
79390 (1),
<typeStatus id="54F28875FFAB0272F588FECEFBF42D35" box="[926,1026,259,283]" pageId="8" pageNumber="309" type="paratype">paratype</typeStatus>
;
<collectionCode id="ED58AE12FFAB0272F21BFECEFBBF2D35" LSID="urn:lsid:biocol.org:col:33791" box="[1037,1097,259,283]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="8" pageNumber="309">MCZ</collectionCode>
79783 (1),
<typeStatus id="54F28875FFAB0272F2DAFECEFAC62D35" box="[1228,1328,259,283]" pageId="8" pageNumber="309" type="paratype">paratype</typeStatus>
.
</paragraph>
<paragraph id="8BF636D7FFAB0272F6D1FEEBFD892D11" blockId="8.[151,1437,258,1975]" box="[199,639,294,319]" pageId="8" pageNumber="309">
<emphasis id="B93DEAC5FFAB0272F6D1FEEBFE8C2D11" bold="true" box="[199,378,294,319]" pageId="8" pageNumber="309">
<typeStatus id="54F28875FFAB0272F6D1FEEBFEF42D11" box="[199,258,294,319]" pageId="8" pageNumber="309">Type</typeStatus>
locality. “
</emphasis>
Chichin Itzá, Yucatan.”
</paragraph>
<paragraph id="8BF636D7FFAB0272F6D1FE87FC102D4D" blockId="8.[151,1437,258,1975]" box="[199,998,330,355]" pageId="8" pageNumber="309">
<emphasis id="B93DEAC5FFAB0272F6D1FE87FE902D4D" bold="true" box="[199,358,330,355]" pageId="8" pageNumber="309">
<typeStatus id="54F28875FFAB0272F6D1FE87FEF62D4D" box="[199,256,330,355]" pageId="8" pageNumber="309">Type</typeStatus>
figured.
</emphasis>
<bibRefCitation id="EFD84B26FFAB0272F77BFE86FD792D4D" author="Bequaert" box="[365,655,330,355]" pageId="8" pageNumber="309" refString="Bequaert, J. &amp; Clench, W. J. (1933) The non-marine Mollusca of Yucatan, In: Shattuck, G. C. (Ed.), The peninsula of Yucatan. Medical, biological, meteorological and sociological studies. Carnegie Institute of Washington Publication, pp. 525 - 545. [(431) 576 pp] [plates 68]" type="book" year="1933">Bequaert &amp; Clench, 1933</bibRefCitation>
: text fig. 26, pl. 68, figs. 8–11.
</paragraph>
<paragraph id="8BF636D7FFAB0272F6D1FEA3FE922DA9" blockId="8.[151,1437,258,1975]" box="[199,356,366,391]" pageId="8" pageNumber="309">
<emphasis id="B93DEAC5FFAB0272F6D1FEA3FE922DA9" bold="true" box="[199,356,366,391]" pageId="8" pageNumber="309">Chresonymy.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C353655CFFAB0272F6D1FE5FFC142EAD" pageId="8" pageNumber="309" type="reference_group">
<paragraph id="8BF636D7FFAB0272F6D1FE5FFBFC2DDD" blockId="8.[151,1437,258,1975]" pageId="8" pageNumber="309">
<treatmentCitationGroup id="AB5911F9FFAB0272F6D1FE5FFBFC2DDD" pageId="8" pageNumber="309">
<taxonomicName id="4C494D54FFAB0272F6D1FE5FFCF82D85" authority="Bequaert &amp; Clench, 1931: 425" authorityName="Bequaert &amp; Clench" authorityPageNumber="425" authorityYear="1931" box="[199,782,402,427]" class="Gastropoda" family="Pomatiidae" genus="Choanopoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="8" pageNumber="309" phylum="Mollusca" rank="species" species="gaigei">
<emphasis id="B93DEAC5FFAB0272F6D1FE5FFE5D2D85" box="[199,427,402,427]" italics="true" pageId="8" pageNumber="309">Choanopoma gaigei</emphasis>
<treatmentCitation id="0AE810C6FFAB0272F7A5FE5EFCF82D85" author="Bequaert" box="[435,782,402,427]" page="425" pageId="8" pageNumber="309" year="1931">
<bibRefCitation id="EFD84B26FFAB0272F7A5FE5EFD212D85" author="Bequaert" box="[435,727,402,427]" pageId="8" pageNumber="309" refString="Bequaert, J. &amp; Clench, W. J. (1931) Three new terrestrial snails from Yucatan. Occasional Papers of the Boston Society of Natural History, 5, 423 - 426." type="journal article" year="1931">Bequaert &amp; Clench, 1931</bibRefCitation>
: 425
</treatmentCitation>
</taxonomicName>
–426;
<treatmentCitation id="0AE810C6FFAB0272F542FE5EFB592D85" author="Bequaert" box="[852,1199,402,427]" page="540" pageId="8" pageNumber="309" year="1933">
<bibRefCitation id="EFD84B26FFAB0272F542FE5EFB8F2D85" author="Bequaert" box="[852,1145,402,427]" pageId="8" pageNumber="309" refString="Bequaert, J. &amp; Clench, W. J. (1933) The non-marine Mollusca of Yucatan, In: Shattuck, G. C. (Ed.), The peninsula of Yucatan. Medical, biological, meteorological and sociological studies. Carnegie Institute of Washington Publication, pp. 525 - 545. [(431) 576 pp] [plates 68]" type="book" year="1933">Bequaert &amp; Clench, 1933</bibRefCitation>
: 540
</treatmentCitation>
–542, text fig. 26, pl. 68, figs. 8–11; Goodrich &amp; van der
<treatmentCitation id="0AE810C6FFAB0272F430FE7BFD1D2DE1" author="Schalie" box="[550,747,438,463]" page="12" pageId="8" pageNumber="309" year="1937">
<bibRefCitation id="EFD84B26FFAB0272F430FE7BFD342DE1" author="Schalie" box="[550,706,438,463]" pageId="8" pageNumber="309" refString="Goodrich, C. &amp; van der Schalie, H. (1937) Mollusca of Peten and North Alta Vera Paz, Guatemala. University of Michigan Museum of Zoology Miscellaneous Publications, 34, 1 - 50. [1 plate, 1 map]" type="book chapter" year="1937">Schalie, 1937</bibRefCitation>
: 12
</treatmentCitation>
, 15, 22, 32;
<treatmentCitation id="0AE810C6FFAB0272F560FE7AFBEC2DE0" author="Harry" box="[886,1050,438,463]" page="7" pageId="8" pageNumber="309" year="1950">
<bibRefCitation id="EFD84B26FFAB0272F560FE7AFBF62DE1" author="Harry" box="[886,1024,438,463]" pageId="8" pageNumber="309" refString="Harry, H. W. (1950) Studies on the nonmarine Mollusca of Yucatan. Occasional Papers of the Museum of Zoology, University of Michigan, 524, 1 - 34." type="journal article" year="1950">Harry, 1950</bibRefCitation>
: 7
</treatmentCitation>
, 26–28;
<treatmentCitation id="0AE810C6FFAB0272F26DFE7AFF322DDD" author="Branson" page="104" pageId="8" pageNumber="309" year="1963">
<bibRefCitation id="EFD84B26FFAB0272F26DFE7AFA6E2DE1" author="Branson" box="[1147,1432,438,463]" pageId="8" pageNumber="309" refString="Branson, B. A. &amp; McCoy, C. J. (1963) Gastropoda of the 1961 University of Colorado Museum Expedition in Mexico. Nautilus, 76, 101 - 108." type="journal article" year="1963">Branson &amp; McCoy, 1963</bibRefCitation>
: 104
</treatmentCitation>
;
<treatmentCitation id="0AE810C6FFAB0272F6C6FE17FE4C2DDD" author="Thompson" box="[208,442,474,499]" page="27" pageId="8" pageNumber="309" year="1966">
<bibRefCitation id="EFD84B26FFAB0272F6C6FE17FE672DDD" author="Thompson" box="[208,401,474,499]" pageId="8" pageNumber="309" refString="Thompson, F. G. (1966) A new pomatiasid from Chiapas, Mexico. Nautilus, 80, 24 - 28." type="journal article" year="1966">Thompson, 1966</bibRefCitation>
: 27
</treatmentCitation>
, 28;
<treatmentCitation id="0AE810C6FFAB0272F7E7FE17FD1D2DDD" author="Thompson" box="[497,747,474,499]" page="225" pageId="8" pageNumber="309" year="1967">
<bibRefCitation id="EFD84B26FFAB0272F7E7FE17FD452DDD" author="Thompson" box="[497,691,474,499]" pageId="8" pageNumber="309" refString="Thompson, F. G. (1967) The land and freshwater snails of Campeche. Bulletin of the Florida State Museum, Biological Sciences, 2 (4), 221 - 256. [6 figures]" type="book chapter" year="1967">Thompson, 1967</bibRefCitation>
: 225
</treatmentCitation>
, 227;
<treatmentCitation id="0AE810C6FFAB0272F538FE16FBF02DDD" author="Watters" box="[814,1030,474,499]" page="269" pageId="8" pageNumber="309" year="2006">
<bibRefCitation id="EFD84B26FFAB0272F538FE16FC3B2DDD" author="Watters" box="[814,973,474,499]" pageId="8" pageNumber="309" refString="Watters. G. T. (2006) The Caribbean landsnail family Annulariidae. A revision of the higher taxa and catalog of the species. Backhuys Publ., Leiden, 584 pp." type="book" year="2006">Watters, 2006</bibRefCitation>
: 269
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BF636D7FFAB0272F6D1FE33FB842E15" blockId="8.[151,1437,258,1975]" pageId="8" pageNumber="309">
<treatmentCitationGroup id="AB5911F9FFAB0272F6D1FE33FB842E15" pageId="8" pageNumber="309">
<taxonomicName id="4C494D54FFAB0272F6D1FE33FC562E39" authority="Bequaert &amp; Clench, 1931" authorityName="Bequaert &amp; Clench" authorityYear="1931" box="[199,928,510,535]" class="Gastropoda" family="Pomatiidae" genus="Choanopoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="8" pageNumber="309" phylum="Mollusca" rank="species" species="gaigei" subGenus="Choanopomops">
<emphasis id="B93DEAC5FFAB0272F6D1FE33FEA82E39" box="[199,350,510,535]" italics="true" pageId="8" pageNumber="309">Choanopoma</emphasis>
(
<emphasis id="B93DEAC5FFAB0272F778FE33FDE82E39" box="[366,542,510,535]" italics="true" pageId="8" pageNumber="309">Choanopomops</emphasis>
)
<emphasis id="B93DEAC5FFAB0272F439FE32FD822E38" box="[559,628,511,534]" italics="true" pageId="8" pageNumber="309">gaigei</emphasis>
<bibRefCitation id="EFD84B26FFAB0272F46BFE32FC562E39" author="Bequaert" box="[637,928,510,535]" pageId="8" pageNumber="309" refString="Bequaert, J. &amp; Clench, W. J. (1931) Three new terrestrial snails from Yucatan. Occasional Papers of the Boston Society of Natural History, 5, 423 - 426." type="journal article" year="1931">Bequaert &amp; Clench, 1931</bibRefCitation>
</taxonomicName>
.
<treatmentCitation id="0AE810C6FFAB0272F5BDFE32FB0C2E39" author="Bequaert" box="[939,1274,510,535]" page="72" pageId="8" pageNumber="309" year="1936">
<bibRefCitation id="EFD84B26FFAB0272F5BDFE32FB392E39" author="Bequaert" box="[939,1231,510,535]" pageId="8" pageNumber="309" refString="Bequaert, J. &amp; Clench, W. J. (1936) A second contribution to the molluscan fauna of Yucatan, In: Pearse, A. S., Creaser, E. P. &amp; Hall, F. G. (Eds.), The cenotes of Yucatan, a zoological and hydrographic survey. Carnegie Institute of Washington Publication, pp. 61 - 75. [(457), 304 pp. 2 plates]" type="book chapter" year="1936">Bequaert &amp; Clench, 1936</bibRefCitation>
: 72
</treatmentCitation>
;
<treatmentCitation id="0AE810C6FFAB0272F310FE33FF352E15" author="Solem" page="197" pageId="8" pageNumber="309" year="1961">
<bibRefCitation id="EFD84B26FFAB0272F310FE33FA6E2E39" author="Solem" box="[1286,1432,510,535]" pageId="8" pageNumber="309" refString="Solem, A. (1961) A preliminary review of the pomatiasid land snails of Central America (Mollusca, Prosobranchia). Archiv fur Molluskenkunde, 90 (4 / 6), 191 - 213." type="journal article" year="1961">Solem, 1961</bibRefCitation>
: 197
</treatmentCitation>
, 203, pl. 10, fig. 5, pl. 12, fig. 24e, map 1;
<treatmentCitation id="0AE810C6FFAB0272F4B5FDEFFC7B2E15" author="Thompson" box="[675,909,546,571]" page="46" pageId="8" pageNumber="309" year="2011">
<bibRefCitation id="EFD84B26FFAB0272F4B5FDEFFC922E15" author="Thompson" box="[675,868,546,571]" pageId="8" pageNumber="309" refString="Thompson, F. G. (2011) An annotated checklist and bibliography of the land and freshwater snails of Mexico and Central America. Bulletin of the Florida Museum of Natural History, 50 (1), 1 - 299." type="journal article" year="2011">Thompson, 2011</bibRefCitation>
: 46
</treatmentCitation>
, 280, 284, 291, 296.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BF636D7FFAB0272F6D1FD8BFC0F2E71" blockId="8.[151,1437,258,1975]" box="[199,1017,582,607]" pageId="8" pageNumber="309">
<treatmentCitationGroup id="AB5911F9FFAB0272F6D1FD8BFC0F2E71" box="[199,1017,582,607]" pageId="8" pageNumber="309">
<taxonomicName id="4C494D54FFAB0272F6D1FD8BFE552E70" box="[199,419,582,607]" class="Gastropoda" family="Pomatiidae" genus="Choanopoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="8" pageNumber="309" phylum="Mollusca" rank="species" species="gagei">
<emphasis id="B93DEAC5FFAB0272F6D1FD8BFE552E70" box="[199,419,582,607]" italics="true" pageId="8" pageNumber="309">Choanopoma gagei</emphasis>
</taxonomicName>
[
<emphasis id="B93DEAC5FFAB0272F7A5FD8AFE242E70" box="[435,466,583,606]" italics="true" pageId="8" pageNumber="309">sic</emphasis>
]
<bibRefCitation id="EFD84B26FFAB0272F7F5FD8AFCF22E71" author="Bequaert" box="[483,772,582,607]" pageId="8" pageNumber="309" refString="Bequaert, J. &amp; Clench, W. J. (1931) Three new terrestrial snails from Yucatan. Occasional Papers of the Boston Society of Natural History, 5, 423 - 426." type="journal article" year="1931">Bequaert &amp; Clench, 1931</bibRefCitation>
.
<treatmentCitation id="0AE810C6FFAB0272F519FD8BFC022E71" author="Richards" box="[783,1012,582,607]" page="257" pageId="8" pageNumber="309" year="1937">
<bibRefCitation id="EFD84B26FFAB0272F519FD8BFC4A2E71" author="Richards" box="[783,956,582,607]" pageId="8" pageNumber="309" refString="Richards, H. G. (1937) Land and freshwater mollusks from the island of Cozumel, Mexico, and their bearing on the geological history of the region. Proceedings of the American Philosophical Society, 77 (3), 249 - 262. [plates. 1 - 4]" type="journal article" year="1937">Richards, 1937</bibRefCitation>
: 257
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BF636D7FFAB0272F6D1FDA7FC142EAD" blockId="8.[151,1437,258,1975]" box="[199,994,618,643]" pageId="8" pageNumber="309">
<treatmentCitationGroup id="AB5911F9FFAB0272F6D1FDA7FC142EAD" box="[199,994,618,643]" pageId="8" pageNumber="309">
<taxonomicName id="4C494D54FFAB0272F6D1FDA7FD242EAD" authority="Bequaert &amp; Clench, 1931" authorityName="Bequaert &amp; Clench" authorityYear="1931" box="[199,722,618,643]" class="Gastropoda" family="Annulariidae" genus="Halotudora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="8" pageNumber="309" phylum="Mollusca" rank="species" species="gaigei">
<emphasis id="B93DEAC5FFAB0272F6D1FDA7FE602EAD" box="[199,406,618,643]" italics="true" pageId="8" pageNumber="309">Halotudora gaigei</emphasis>
(
<bibRefCitation id="EFD84B26FFAB0272F7B3FDA6FD3F2EAD" author="Bequaert" box="[421,713,618,643]" pageId="8" pageNumber="309" refString="Bequaert, J. &amp; Clench, W. J. (1931) Three new terrestrial snails from Yucatan. Occasional Papers of the Boston Society of Natural History, 5, 423 - 426." type="journal article" year="1931">Bequaert &amp; Clench, 1931</bibRefCitation>
)
</taxonomicName>
.
<treatmentCitation id="0AE810C6FFAB0272F4CAFDA6FC532EAD" author="Watters" box="[732,933,618,643]" page="73" pageId="8" pageNumber="309" year="2006">
<bibRefCitation id="EFD84B26FFAB0272F4CAFDA6FC8D2EAD" author="Watters" box="[732,891,618,643]" pageId="8" pageNumber="309" refString="Watters. G. T. (2006) The Caribbean landsnail family Annulariidae. A revision of the higher taxa and catalog of the species. Backhuys Publ., Leiden, 584 pp." type="book" year="2006">Watters, 2006</bibRefCitation>
: 73
</treatmentCitation>
, 269.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C353655CFFAB0272F6D1FD43FE842F19" pageId="8" pageNumber="309" type="distribution">
<paragraph id="8BF636D7FFAB0272F6D1FD43FB8B2EC1" blockId="8.[151,1437,258,1975]" pageId="8" pageNumber="309">
<emphasis id="B93DEAC5FFAB0272F6D1FD43FE022E89" bold="true" box="[199,500,654,679]" pageId="8" pageNumber="309">Distribution and habitat.</emphasis>
Yucatán Peninsula of Campeche, Quintana Roo, and Yucatán states in
<collectingCountry id="F35E7647FFAB0272F337FD43FA8F2E89" box="[1313,1401,654,679]" name="Mexico" pageId="8" pageNumber="309">México</collectingCountry>
, in
<collectingCountry id="F35E7647FFAB0272F681FD7FFF172EE5" box="[151,225,690,715]" name="Belize" pageId="8" pageNumber="309">Belize</collectingCountry>
, and into Izabal Department of
<collectingCountry id="F35E7647FFAB0272F44DFD7FFD2E2EE5" box="[603,728,690,715]" name="Guatemala" pageId="8" pageNumber="309">Guatemala</collectingCountry>
. Snails occur on lowland limestone outcrops and in cenotes. Often encountered at the Mayan ruins of Chichén-Itzá, Chetumal, Kohunlich, and others.
</paragraph>
<paragraph id="8BF636D7FFAB0272F6D1FD37FE842F19" blockId="8.[151,1437,258,1975]" pageId="8" pageNumber="309">
<emphasis id="B93DEAC5FFAB0272F6D1FD37FE992F3D" bold="true" box="[199,367,762,787]" pageId="8" pageNumber="309">Conservation.</emphasis>
A significant portion of the range occurs in reserves, parks, and archeological sites in both
<collectingCountry id="F35E7647FFAB0272F681FCD3FF292F19" box="[151,223,798,823]" name="Belize" pageId="8" pageNumber="309">Belize</collectingCountry>
and
<collectingCountry id="F35E7647FFAB0272F703FCD3FE992F19" box="[277,367,798,823]" name="Mexico" pageId="8" pageNumber="309">México</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C353655CFFAB0272F6D1FC8FFAF228B1" pageId="8" pageNumber="309" type="materials_examined">
<paragraph id="8BF636D7FFAB0272F6D1FC8FFAF228B1" blockId="8.[151,1437,258,1975]" pageId="8" pageNumber="309">
<emphasis id="B93DEAC5FFAB0272F6D1FC8FFBC22F75" bold="true" box="[199,1076,834,859]" pageId="8" pageNumber="309">
Other material (specimens examined: 201).
<collectingCountry id="F35E7647FFAB0272F4E0FC8FFCB92F75" box="[758,847,834,859]" name="Mexico" pageId="8" pageNumber="309">México</collectingCountry>
. Campeche State:
</emphasis>
UF 19093 (4),
<quantity id="4CB19B32FFAB0272F317FC8EFAA02F75" box="[1281,1366,834,859]" metricMagnitude="3" metricUnit="m" metricValue="9.2" pageId="8" pageNumber="309" unit="km" value="9.2">9.2 km</quantity>
E of Campeche; UF 19088 (17), UF 213712 (10),
<quantity id="4CB19B32FFAB0272F483FCABFD152F51" box="[661,739,870,895]" metricMagnitude="3" metricUnit="m" metricValue="6.4" pageId="8" pageNumber="309" unit="km" value="6.4">6.4 km</quantity>
E of Xpujil; UF 155822 (4), Becán; UF 19080 (17),
<quantity id="4CB19B32FFAB0272F321FCABFA702F51" box="[1335,1414,870,895]" metricMagnitude="3" metricUnit="m" metricValue="5.8" pageId="8" pageNumber="309" unit="km" value="5.8">5.8 km</quantity>
S of Hopelchén; UF 19087 (11),
<quantity id="4CB19B32FFAB0272F7E4FC46FDB92F8D" box="[498,591,906,931]" metricMagnitude="4" metricUnit="m" metricValue="3.09" pageId="8" pageNumber="309" unit="km" value="30.9">30.9 km</quantity>
E of Silvituc; UF 19091 (10),
<quantity id="4CB19B32FFAB0272F5BEFC46FC012F8D" box="[936,1015,906,931]" metricMagnitude="3" metricUnit="m" metricValue="8.2" pageId="8" pageNumber="309" unit="km" value="8.2">8.2 km</quantity>
W of Tikimul; UF 19090 (3),
<quantity id="4CB19B32FFAB0272F358FC47FA6A2F8D" box="[1358,1436,906,931]" metricMagnitude="3" metricUnit="m" metricValue="5.5" pageId="8" pageNumber="309" unit="km" value="5.5">5.5 km</quantity>
S of Cayal; UF 19098 (2),
<quantity id="4CB19B32FFAB0272F7DFFC63FDEF2FE9" box="[457,537,942,967]" metricMagnitude="3" metricUnit="m" metricValue="3.5" pageId="8" pageNumber="309" unit="km" value="3.5">3.5 km</quantity>
S of San Pablo Pixtún.
<emphasis id="B93DEAC5FFAB0272F53CFC63FBD62FE9" bold="true" box="[810,1056,942,967]" pageId="8" pageNumber="309">Quintana Roo State:</emphasis>
GTW 13725a (1), Chetumal; UF 133597 (11), 30.0 km S of Felipe Carrillo Puerto; GTW 13725c (1), UF 155826 (5), Kohunlich; UF 19084 (1),
<quantity id="4CB19B32FFAB0272F36FFC1EFF4D2821" metricMagnitude="3" metricUnit="m" metricValue="9.7" pageId="8" pageNumber="309" unit="km" value="9.7">9.7 km</quantity>
N of Dzibanche; UF 19092 (5),
<quantity id="4CB19B32FFAB0272F435FC3AFD872821" box="[547,625,1014,1039]" metricMagnitude="3" metricUnit="m" metricValue="3.7" pageId="8" pageNumber="309" unit="km" value="3.7">3.7 km</quantity>
SSE of Polyuc; UF 19089 (3),
<quantity id="4CB19B32FFAB0272F5DDFC3AFBD12821" box="[971,1063,1014,1039]" metricMagnitude="4" metricUnit="m" metricValue="1.04" pageId="8" pageNumber="309" unit="km" value="10.4">10.4 km</quantity>
NNW of Polyuc.
<emphasis id="B93DEAC5FFAB0272F2F8FC3BFA6A2821" bold="true" box="[1262,1436,1014,1039]" pageId="8" pageNumber="309">Yucatán State:</emphasis>
GTW 13725b (2), UF 275359 (3), Chichén-Itzá; UF 19085 (11),
<quantity id="4CB19B32FFAB0272F561FBD6FC30281D" box="[887,966,1050,1075]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="8" pageNumber="309" unit="km" value="1.3">1.3 km</quantity>
NE of Becanchén.
<emphasis id="B93DEAC5FFAB0272F2B2FBD7FF0C2879" bold="true" pageId="8" pageNumber="309">
<collectingCountry id="F35E7647FFAB0272F2B2FBD7FB18281D" box="[1188,1262,1050,1075]" name="Belize" pageId="8" pageNumber="309">Belize</collectingCountry>
. Orange Walk District:
</emphasis>
UF 207140 (30), UF 207164 (33), 1.0 km E of Neuendorf,
<quantity id="4CB19B32FFAB0272F585FBF2FC2B2878" box="[915,989,1087,1111]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="309" unit="m" value="100.0">100 m</quantity>
; UF 207181 (1), UF 207182 (8),
<quantity id="4CB19B32FFAB0272F359FBF3FA6A2879" box="[1359,1436,1086,1111]" metricMagnitude="3" metricUnit="m" metricValue="5.4" pageId="8" pageNumber="309" unit="km" value="5.4">5.4 km</quantity>
W of Neuendorf.
<emphasis id="B93DEAC5FFAB0272F74BFBAFFDF02855" bold="true" box="[349,518,1122,1147]" pageId="8" pageNumber="309">Cayo District:</emphasis>
GTW 13742c (1), Río Frío Cave.
<emphasis id="B93DEAC5FFAB0272F59AFBAFFB7D2855" bold="true" box="[908,1163,1122,1147]" pageId="8" pageNumber="309">Stann Creek District:</emphasis>
UF 207591 (1), mogote
<quantity id="4CB19B32FFAB0272F681FB4AFF1328B1" box="[151,229,1158,1183]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="8" pageNumber="309" unit="km" value="2.2">2.2 km</quantity>
S of Trio Village,
<specimenCount id="9D4FFD5EFFAB0272F7A3FB4BFE0028B0" box="[437,502,1158,1183]" pageId="8" pageNumber="309" type="generic">
<quantity id="4CB19B32FFAB0272F7A3FB4BFE0728B0" box="[437,497,1158,1183]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="309" unit="m" value="50.0">50 m</quantity>
.
</specimenCount>
<emphasis id="B93DEAC5FFAB0272F7EAFB4BFC8C28B1" bold="true" box="[508,890,1158,1183]" pageId="8" pageNumber="309">
<collectingCountry id="F35E7647FFAB0272F7EAFB4BFD7428B1" box="[508,642,1158,1183]" name="Guatemala" pageId="8" pageNumber="309">Guatemala</collectingCountry>
. Izabal Department:
</emphasis>
UF 19152 (6),
<quantity id="4CB19B32FFAB0272F231FB4AFB9728B1" box="[1063,1121,1158,1183]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="8" pageNumber="309" unit="km" value="4.0">4 km</quantity>
N of Morales.
</paragraph>
</subSubSection>
<subSubSection id="C353655CFFAB0272F6D1FB67FB8B2AB9" pageId="8" pageNumber="309" type="description">
<paragraph id="8BF636D7FFAB0272F6D1FB67FB8B2AB9" blockId="8.[151,1437,258,1975]" pageId="8" pageNumber="309">
<emphasis id="B93DEAC5FFAB0272F6D1FB67FEAF28ED" bold="true" box="[199,345,1194,1219]" pageId="8" pageNumber="309">Description.</emphasis>
Shell conical, moderately high-spired, thin but solid. Smallest adult specimen seen
<quantity id="4CB19B32FFAB0272F30EFB67FA8828EC" box="[1304,1406,1194,1219]" metricMagnitude="-2" metricUnit="m" metricValue="1.06" pageId="8" pageNumber="309" unit="mm" value="10.6">10.6 mm</quantity>
in length, largest
<quantity id="4CB19B32FFAB0272F757FB02FE5C28C8" box="[321,426,1231,1255]" metricMagnitude="-2" metricUnit="m" metricValue="1.8199999999999998" pageId="8" pageNumber="309" unit="mm" value="18.2">18.2 mm</quantity>
, average
<quantity id="4CB19B32FFAB0272F40FFB03FD7628C8" box="[537,640,1230,1255]" metricMagnitude="-2" metricUnit="m" metricValue="1.46" pageId="8" pageNumber="309" unit="mm" value="14.6">14.6 mm</quantity>
(non-decollate). Protoconch usually retained in adult, 1.5 prominent, rounded, pale whorls. Teleoconch of 4.5–5 rounded whorls. Umbilicus wide, partially occluded by outer lip. Spiral sculpture of numerous faint, low cords, separated by about twice their width, much stronger in the umbilicus. Axial sculpture of numerous fine lines that render the shell minutely decussate; this sculpture is more lamellate and widely separated on earlier whorls. Suture strongly indented, narrowly channeled in some specimens. Tufts absent but groups of axial lamella may be somewhat wider at the suture rendering the suture finely serrate. Aperture nearly circular. Inner lip smooth, narrowly exserted. Outer lip strongly lamellate, widely to moderately reflected perpendicular to whorl, evenly expanded, usually auriculate posteriorly, narrowly solute or (less commonly) scarcely attached to previous whorl. Base color dingy white to yellowish-tan, first teleoconch whorl darker in some specimens. Rarely unicolor, usually with spiral, more or less continuous brown bands, occasionally broken into spots, bands lacking in umbilicus. Outer lip white; the bands do not extend to the front of the peristome. Inside of aperture brownish-orange or white with the outer pattern showing through. Operculum paucispiral, with a thick reflected plate composed of numerous recurved lamellae. Radula and anatomy unknown.
</paragraph>
</subSubSection>
<subSubSection id="C353655CFFAB0272F6D1F96FFBFD2B09" pageId="8" pageNumber="309" type="materials_examined">
<paragraph id="8BF636D7FFAB0272F6D1F96FFB7D2A95" blockId="8.[151,1437,258,1975]" box="[199,1163,1698,1723]" pageId="8" pageNumber="309">
<emphasis id="B93DEAC5FFAB0272F6D1F96FFE2B2A95" bold="true" box="[199,477,1698,1723]" pageId="8" pageNumber="309">Variation in specimens.</emphasis>
The color pattern varies from all white to strongly patterned.
</paragraph>
<paragraph id="8BF636D7FFAB0272F6D1F90BFBFD2B09" blockId="8.[151,1437,258,1975]" pageId="8" pageNumber="309">
<emphasis id="B93DEAC5FFAB0272F6D1F90BFDB42AF1" bold="true" box="[199,578,1734,1759]" pageId="8" pageNumber="309">Comparison with other species.</emphasis>
This species differs from
<taxonomicName id="4C494D54FFAB0272F578F90AFC1D2AF1" box="[878,1003,1734,1759]" class="Gastropoda" family="Annulariidae" genus="Halotudora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="8" pageNumber="309" phylum="Mollusca" rank="species" species="kuesteri">
<emphasis id="B93DEAC5FFAB0272F578F90AFC1D2AF1" box="[878,1003,1734,1759]" italics="true" pageId="8" pageNumber="309">H. kuesteri</emphasis>
</taxonomicName>
and
<taxonomicName id="4C494D54FFAB0272F232F90AFB6A2AF0" box="[1060,1180,1735,1758]" class="Gastropoda" family="Assimineidae" genus="Cyclostoma" kingdom="Animalia" order="Littorinimorpha" pageId="8" pageNumber="309" phylum="Mollusca" rank="species" species="gruneri">
<emphasis id="B93DEAC5FFAB0272F232F90AFB6A2AF0" box="[1060,1180,1735,1758]" italics="true" pageId="8" pageNumber="309">H. gruneri</emphasis>
</taxonomicName>
in the retention of the protoconch (usually) and in the narrowly solute outer lip. It lacks the fused tufts of
<taxonomicName id="4C494D54FFAB0272F286F926FADD2B2D" box="[1168,1323,1770,1795]" class="Gastropoda" family="Assimineidae" genus="Cyclostoma" kingdom="Animalia" order="Littorinimorpha" pageId="8" pageNumber="309" phylum="Mollusca" rank="species" species="largillierti">
<emphasis id="B93DEAC5FFAB0272F286F926FADD2B2D" box="[1168,1323,1770,1795]" italics="true" pageId="8" pageNumber="309">C. largillierti</emphasis>
</taxonomicName>
. From
<taxonomicName id="4C494D54FFAB0272F391F921FEEF2B09" class="Gastropoda" family="Pomatiidae" genus="Parachondria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="8" pageNumber="309" phylum="Mollusca" rank="species" species="cordovanus">
<emphasis id="B93DEAC5FFAB0272F391F921FEEF2B09" italics="true" pageId="8" pageNumber="309">P. cordovanus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C494D54FFAB0272F759F8C2FE1E2B09" box="[335,488,1806,1831]" class="Gastropoda" family="Pomatiidae" genus="Parachondria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="8" pageNumber="309" phylum="Mollusca" rank="species" species="rubicundus">
<emphasis id="B93DEAC5FFAB0272F759F8C2FE1E2B09" box="[335,488,1806,1831]" italics="true" pageId="8" pageNumber="309">P. rubicundus</emphasis>
</taxonomicName>
it differs in the heavily calcified opercular plate.
</paragraph>
</subSubSection>
<subSubSection id="C353655CFFAB0273F6D1F8FFFCC82A1D" lastPageId="9" lastPageNumber="310" pageId="8" pageNumber="309" type="discussion">
<paragraph id="8BF636D7FFAB0272F6D1F8FFFD4E2B99" blockId="8.[151,1437,258,1975]" pageId="8" pageNumber="309">
<emphasis id="B93DEAC5FFAB0272F6D1F8FFFECD2B65" bold="true" box="[199,315,1842,1867]" pageId="8" pageNumber="309">Remarks.</emphasis>
It is peculiar that this widespread and distinctive species, commonly occurring at oft-visited Mayan ruins, should have remained undescribed until 1931, particularly since it is one of the most widely distributed of all the
<taxonomicName id="4C494D54FFAB0272F6D6F8B7FEA42BBD" box="[192,338,1914,1939]" class="Gastropoda" family="Annulariidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="8" pageNumber="309" phylum="Mollusca" rank="family">Annulariidae</taxonomicName>
(the Venezuelan
<taxonomicName id="4C494D54FFAB0272F402F8B7FC242BBD" authority="Pfeiffer, 1846" authorityName="Pfeiffer" authorityYear="1846" box="[532,978,1914,1939]" class="Gastropoda" family="Annulariidae" genus="Gouldipoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="8" pageNumber="309" phylum="Mollusca" rank="species" species="plicatulum">
<emphasis id="B93DEAC5FFAB0272F402F8B7FCE82BBD" box="[532,798,1914,1939]" italics="true" pageId="8" pageNumber="309">Gouldipoma plicatulum</emphasis>
(
<bibRefCitation id="EFD84B26FFAB0272F53BF8B7FC3C2BBD" author="Pfeiffer" box="[813,970,1914,1939]" pageId="8" pageNumber="309" refString="Pfeiffer, L. (1846) Revision der Gattung Cyclostoma. Zeitschrift fur Malakozoologie fur Marz, 3, 32 - 48. [1846]" type="journal article" year="1846">Pfeiffer, 1846</bibRefCitation>
)
</taxonomicName>
appears to hold the distinction of having the broadest zoogeographic range of the family).
</paragraph>
<caption id="DF36665FFFAA0273F681FC1CFEB9286D" httpUri="https://zenodo.org/record/252703/files/figure.png" pageId="9" pageNumber="310" targetBox="[151,1436,193,955]" targetPageId="9">
<paragraph id="8BF636D7FFAA0273F681FC1CFEB9286D" blockId="9.[151,1436,977,1091]" pageId="9" pageNumber="310">
<emphasis id="B93DEAC5FFAA0273F681FC1CFEBA2FC9" bold="true" box="[151,332,977,999]" pageId="9" pageNumber="310">FIGURE 2 A–K.</emphasis>
<taxonomicName id="4C494D54FFAA0273F744FC1CFCDD2FC9" authority="Bequaert &amp; Clench, 1931" authorityName="Bequaert &amp; Clench" authorityYear="1931" box="[338,811,977,999]" class="Gastropoda" family="Annulariidae" genus="Halotudora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="9" pageNumber="310" phylum="Mollusca" rank="species" species="gaigei">
<emphasis id="B93DEAC5FFAA0273F744FC1CFDF82FC9" box="[338,526,977,999]" italics="true" pageId="9" pageNumber="310">Halotudora gaigei</emphasis>
(Bequaert &amp; Clench, 1931)
</taxonomicName>
. A. UMMZ 49190, holotype, 14.5 mm. B–C. UF 19088, 15.2 mm. D. UF 155822, 13.4 mm. E. UF 133597, 16.6 mm. F. UF 19092, 15.7 mm. G. UF 19152, 13.5 mm. H. GTW 13724c, 15.6 mm. I. GTW 13725b, 14.0 mm [operculum glued in aperture backward by collector]. J. GTW 13725b, 13.2 mm. K. GTW 13725a, 16.1 mm.
</paragraph>
</caption>
<paragraph id="8BF636D7FFAA0273F6D1FBA7FCC82A1D" blockId="9.[151,1437,1130,1659]" pageId="9" pageNumber="310">
<emphasis id="B93DEAC5FFAA0273F6D1FBA7FE3428AD" bold="true" box="[199,450,1130,1155]" pageId="9" pageNumber="310">Original description.</emphasis>
“Medium in size, acuminate, elongate-conic, narrowly umbilicate. Color pale brown, with spiral bands of mahogany red, the bands usually irregularly broken, more strongly developed on the last two whorls. Whorls 6, convex, regularly increasing in size, and often all preserved in adult specimens; sometimes only the embryonic whorls lost. Spire acute, but the apex truncate, the first whorl being depressed. Aperture entire and nearly circular. Peristome duplex: inner rim sharp and circular; outer rim reflected, somewhat wavy, and forming a slight rounded expansion near the parietal angle. Sutures deep, not channeled, strongly but finely and regularly beaded. Sculpture of both axial and spiral threads. Embryonic whorls smooth; third and fourth whorls with strong, axial riblets, regularly spaced. On the remaining whorls the spiral sculpture gradually approaches and eventually equals the axial riblets, which on the last whorl are more crowded. Under the lens the last whorl is finely reticulate. The fine sutural beading is caused by the insertion and slight thickening of the axial riblets. Operculum calcified, paucispiral, circular, composed of three whorls; occasionally the edge irregularly crenulate; the last whorl rapidly increasing in size, being equal in width to the other two whorls together; the lamella only slightly elevated and not extending over the spiral suture; nucleus strongly eccentric.”
</paragraph>
</subSubSection>
<subSubSection id="C353655CFFAA0273F6D1F9F3FE602A55" pageId="9" pageNumber="310" type="etymology">
<paragraph id="8BF636D7FFAA0273F6D1F9F3FE602A55" blockId="9.[151,1437,1130,1659]" pageId="9" pageNumber="310">
<emphasis id="B93DEAC5FFAA0273F6D1F9F3FEB82A79" bold="true" box="[199,334,1598,1623]" pageId="9" pageNumber="310">Etymology.</emphasis>
Frederick McMahon Gaige (
<date id="FFF71017FFAA0273F4A7F9F3FCC22A79" box="[689,820,1598,1623]" pageId="9" pageNumber="310" value="1890" valueMax="1976">1890–1976</date>
), entomologist at the University of Michigan, who collected the
<typeStatus id="54F28875FFAA0273F73AF9AFFE642A55" box="[300,402,1634,1659]" pageId="9" pageNumber="310" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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