1251 lines
167 KiB
XML
1251 lines
167 KiB
XML
<document id="824ABC95208E9CFF7541906BA40FE73E" ID-CLB-Dataset="24691" ID-DOI="10.11646/zootaxa.4706.2.2" ID-GBIF-Dataset="e6914caf-bb5b-41f6-bf49-eaa87ef1fb1f" ID-ISSN="1175-5326" ID-Zenodo-Dep="3567336" ID-ZooBank="41805144-D639-454C-98F3-6C3A013EC8CE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1575876155473" checkinUser="plazi" docAuthor="Schliewen, Ulrich K., Kovačić, Marcelo, Cerwenka, Alexander F., Svensen, Rudolf & Ordines, Francesc" docDate="2019" docId="03E087B3FF88FFDAA8E4F940FBC9FE0F" docLanguage="en" docName="zootaxa.4706.2.2.pdf" docOrigin="Zootaxa 4706 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lebetus patzneri Schliewen & Kovačić & Cerwenka & Svensen & Ordines 2019, sp. nov." docType="treatment" docVersion="8" lastPageNumber="243" masterDocId="FFD9FFCBFF8DFFD4A873FFD4FF9CFFDC" masterDocTitle="Lebetus patzneri (Teleostei: Gobiidae), a new goby species from the Balearic Islands, western Mediterranean, with first records of Lebetus guilleti (Le Danois, 1913) from this area and Norway, and with notes on its biology" masterLastPageNumber="254" masterPageNumber="231" pageNumber="236" updateTime="1698773588651" updateUser="ExternalLinkService">
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<mods:title id="C60EE9BEA59BC74DCCCA3F15B044D544">Lebetus patzneri (Teleostei: Gobiidae), a new goby species from the Balearic Islands, western Mediterranean, with first records of Lebetus guilleti (Le Danois, 1913) from this area and Norway, and with notes on its biology</mods:title>
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<mods:namePart id="2F063FB04C7CE9F2F6B282EDD3F09658">Schliewen, Ulrich K.</mods:namePart>
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<mods:namePart id="347DEF5B524BD3FCDFC83E0349AAC37B">Kovačić, Marcelo</mods:namePart>
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<mods:namePart id="51D9D2ACCB75E459FDA1DE16F5E83000">Cerwenka, Alexander F.</mods:namePart>
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<mods:namePart id="4D534034368D622B52B8881216791578">Svensen, Rudolf</mods:namePart>
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<mods:namePart id="18D1A3F539B95EEA42E92C032C97B5EA">Ordines, Francesc</mods:namePart>
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<mods:title id="27BEE9365D1851832CF07A05B3ED471E">Zootaxa</mods:title>
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<mods:date id="FFD1603A69B22FE38BDA8AB30F474D21">2019</mods:date>
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<mods:number id="B9E83556422FEA7C48D9383826A1A6D8">2019-12-09</mods:number>
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<mods:number id="D8874FD9A16F3127A0D5946BFB6E7F69">4706</mods:number>
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<subSubSection id="C353652EFF88FFD1A8E4F940FCD5F972" box="[151,841,1683,1711]" pageId="5" pageNumber="236" type="nomenclature">
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<paragraph id="8BF636A5FF88FFD1A8E4F940FCD5F972" blockId="5.[151,841,1683,1746]" box="[151,841,1683,1711]" pageId="5" pageNumber="236">
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<heading id="D0BE81C9FF88FFD1A8E4F940FCD5F972" bold="true" box="[151,841,1683,1711]" fontSize="11" level="1" pageId="5" pageNumber="236" reason="1">
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<emphasis id="B93DEAB7FF88FFD1A8E4F940FCD5F972" bold="true" box="[151,841,1683,1711]" pageId="5" pageNumber="236">
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<taxonomicName id="4C494D26FF88FFD1A8E4F940FEC3F972" authority="Schliewen & Kovačić & Cerwenka & Svensen & Ordines, 2019" authorityName="Schliewen & Kovačić & Cerwenka & Svensen & Ordines" authorityYear="2019" box="[151,351,1684,1711]" class="Actinopterygii" family="Gobiidae" genus="Lebetus" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="236" phylum="Chordata" rank="species" species="patzneri" status="sp. nov.">
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<emphasis id="B93DEAB7FF88FFD1A8E4F940FEC3F972" bold="true" box="[151,351,1684,1711]" italics="true" pageId="5" pageNumber="236">Lebetus patzneri</emphasis>
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<taxonomicNameLabel id="A20E57CCFF88FFD1A915F940FE58F972" box="[358,452,1684,1710]" pageId="5" pageNumber="236" rank="species">sp. nov.</taxonomicNameLabel>
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Schliewen, Kovačić & Ordines
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="C353652EFF88FFD1A8E4F96CFE8EF90E" box="[151,274,1720,1746]" pageId="5" pageNumber="236" type="description">
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<paragraph id="8BF636A5FF88FFD1A8E4F96CFE8EF90E" blockId="5.[151,841,1683,1746]" box="[151,274,1720,1746]" pageId="5" pageNumber="236">
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(
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<figureCitation id="13722A20FF88FFD1A8ECF96CFE97F90E" box="[159,267,1720,1746]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStart-4="FIGURE 6" captionStart-5="FIGURE 7" captionStart-6="FIGURE 8" captionStart-7="FIGURE 9" captionStartId-0="6.[151,250,1935,1960]" captionStartId-1="7.[151,250,1726,1751]" captionStartId-2="8.[151,250,1427,1452]" captionStartId-3="9.[151,250,1975,2000]" captionStartId-4="11.[151,250,1257,1282]" captionStartId-5="12.[151,250,922,947]" captionStartId-6="13.[151,250,1801,1826]" captionStartId-7="14.[151,250,1193,1218]" captionTargetBox-0="[151,1436,733,1915]" captionTargetBox-1="[177,1410,917,1703]" captionTargetBox-2="[151,1436,519,1407]" captionTargetBox-3="[170,1425,953,1948]" captionTargetBox-4="[160,1436,607,1226]" captionTargetBox-5="[151,1435,337,902]" captionTargetBox-6="[179,1412,181,1770]" captionTargetBox-7="[156,1429,482,1167]" captionTargetId-0="figure@6.[151,1436,733,1915]" captionTargetId-1="figure@7.[170,1418,913,1707]" captionTargetId-2="figure@8.[151,1436,517,1407]" captionTargetId-3="figure@9.[162,1425,949,1956]" captionTargetId-4="figure@11.[151,1436,589,1238]" captionTargetId-5="figure@12.[151,1435,337,902]" captionTargetId-6="figure@13.[159,1429,166,1782]" captionTargetId-7="figure@14.[151,1437,481,1173]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="8" captionTargetPageId-3="9" captionTargetPageId-4="11" captionTargetPageId-5="12" captionTargetPageId-6="13" captionTargetPageId-7="14" captionText-0="FIGURE 2. Lebetus patzneri sp. nov., aspects of the freshly collected holotype, ZSM 47486, female, 16.6 mm SL, from off SW Mallorca Island, 72 m depth, preserved. a: left lateral view; b: right lateral view; c: ventral view; c: dorsal view; d: ventral view left lateral view. Photographs by F. Ordinas." captionText-1="FIGURE 3. Lebetus patzneri sp. nov., aspects of the preserved holotype, reversibly stained with Cyanine Blue, ZSM 47486, female, 16.6 mm SL, from off SW Mallorca Island, 72 m depth, preserved. a: left lateral view b: dorsal view; c: ventral view. Photographs by M. Kovačić." captionText-2="FIGURE 4. Lebetus patzneri sp. nov., SEM aspects of the paratype ZSM 46865, female, 16.8 mm SL; from off SW Mallorca Island, 60m depth. a: right lateral view; b: ventral view of pelvic fins and urogenital papilla. SEM photo by A. Cerwenka." captionText-3="FIGURE 5. Lebetus patzneri sp. nov., MicroCT reconstruction of the paratype ZSM 46865, female, 16.8 mm SL; from off SW Mallorca Island, 60m depth. a: left lateral view; b: dorsal view; c: ventral view. MicroCT reconstruction by A. Cerwenka." captionText-4="FIGURE 6. Lebetus patzneri sp. nov., head lateral line system of the holotype ZSM 47486, female, 16.6 mm SL, from off SW Mallorca Island, 72 m depth. Terminology in text. Drawing by M. Kovačić." captionText-5="FIGURE 7. Lebetus patzneri sp. nov., right saccular otolith of the paratype ZSM 46865, female, 16.8 mm SL; from off SW Mallorca, 60m depth. a: in medial/inner view; b: left otolith in anterior view. Orientation of otolith with respect to fish symmetry is indicated as follows: a, anterior; d, dorsal; l, lateral; m, medial; p, posterior; v, ventral. MicroCT reconstruction by A. Cerwenka." captionText-6="FIGURE 8. Lebetus patzneri sp. nov., aspects of the paratype and two unpreserved specimens in life coloration shortly after collection. a: left lateral view of paratype ZSM 46865, female, 16.8 mm SL; from off SW Mallorca Island, 60m depth (based on montage of two separate photographs (slight artifacts at montage suture at about D1 origin due to slightly different position of the specimen between the two photographs); b: left dorsolateral view of head and anterior flank view of paratype ZSM 46865; c: left lateral view of the second known specimen (unpreserved), unknown size, Menorca channel, 67m depth (based on montage of two separate photographs; stitching artifacts visible before D1 origin); d: left slightly dorsolateral view of third known specimen (unpreserved) of unknown size, Menorca channel, 67m depth. Photographs by F. Ordines." captionText-7="FIGURE 9. Maximum likelihood phylogenetic tree topology depicting the phylogenetic position and small intraspecific genetic distance of COI-barcode haplotypes (520 bp) of the holotype of Lebetus patzneri sp. nov. and five Lebetus guilleti from the northeastern Atlantic (Norway, two specimens), the Western Mediterranean (Baleares, one specimen) and from the northern Adriatic (Croatia, two specimens). All sand goby genera are included, except for Hyrcanogobius and Pseudaphya. Numbers on branches are bootstrap support values (%) for the Maximum Likelihood analysis (values below 50% not shown). Tip labels are composed of the genus and species name followed by ZSM collection number (if newly sequenced in this study) or of the GenBank accession number. Inserted photos depict the second known specimen of L. patzneri sp. nov. (as Fig. 8c; upper photograph) and the first L. guilleti specimen photographed and collected in 15m depth in Norway (ZSM-PIS-GO 1227; lower photograph). Photographs above and below by F. Ordines and R. Svensen, respectively." figureDoi-0="http://doi.org/10.5281/zenodo.3567340" figureDoi-1="http://doi.org/10.5281/zenodo.3567342" figureDoi-2="http://doi.org/10.5281/zenodo.3567344" figureDoi-3="http://doi.org/10.5281/zenodo.3567346" figureDoi-4="http://doi.org/10.5281/zenodo.3567348" figureDoi-5="http://doi.org/10.5281/zenodo.3567350" figureDoi-6="http://doi.org/10.5281/zenodo.3567352" figureDoi-7="http://doi.org/10.5281/zenodo.3567354" httpUri-0="https://zenodo.org/record/3567340/files/figure.png" httpUri-1="https://zenodo.org/record/3567342/files/figure.png" httpUri-2="https://zenodo.org/record/3567344/files/figure.png" httpUri-3="https://zenodo.org/record/3567346/files/figure.png" httpUri-4="https://zenodo.org/record/3567348/files/figure.png" httpUri-5="https://zenodo.org/record/3567350/files/figure.png" httpUri-6="https://zenodo.org/record/3567352/files/figure.png" httpUri-7="https://zenodo.org/record/3567354/files/figure.png" pageId="5" pageNumber="236">Figs. 2–9</figureCitation>
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)
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</paragraph>
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<subSubSection id="C353652EFF88FFD2A8E4F92BFB11FE9F" lastPageId="6" lastPageNumber="237" pageId="5" pageNumber="236" type="materials_examined">
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<paragraph id="8BF636A5FF88FFD1A8E4F92BFC2FF85A" blockId="5.[151,1436,1791,2035]" pageId="5" pageNumber="236">
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<materialsCitation id="3B213CF8FF88FFD1A8E4F92BFC2FF85A" ID-GBIF-Occurrence="2465031520" collectingDate="2019-04-27" collectionCode="ZSM" collectorName="F. Ordines & Preserved" country="Spain" elevation="72" latitude="39.859" location="Channel" longLatPrecision="55" longitude="3.5413" pageId="5" pageNumber="236" specimenCount="1" stateProvince="Illes Baleares" typeStatus="holotype">
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<emphasis id="B93DEAB7FF88FFD1A8E4F92BFF63F8C6" bold="true" box="[151,255,1791,1818]" italics="true" pageId="5" pageNumber="236">
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<typeStatus id="54F28807FF88FFD1A8E4F92BFF63F8C6" box="[151,255,1791,1818]" pageId="5" pageNumber="236" type="holotype">Holotype</typeStatus>
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</emphasis>
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.
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</emphasis>
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<collectionCode id="ED58AE60FF88FFD1A97DF8D4FED4F8C6" box="[270,328,1792,1818]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="5" pageNumber="236" type="Museum">ZSM</collectionCode>
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47486, female,
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SL (
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TL,
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<collectingCountry id="F35E7635FF88FFD1AB4FF8D4FCE3F8C6" box="[828,895,1792,1818]" name="Spain" pageId="5" pageNumber="236">Spain</collectingCountry>
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,
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<collectingRegion id="498DF847FF88FFD1ABF8F8D4FBDEF8C6" box="[907,1090,1792,1818]" country="Spain" name="Illes Baleares" pageId="5" pageNumber="236">Balearic Islands</collectingRegion>
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, Menorca
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(
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,
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<geoCoordinate id="EE7D5062FF88FFD1A8E4F8F7FE98F8E2" box="[151,260,1827,1854]" degrees="3.5413" direction="east" orientation="longitude" pageId="5" pageNumber="236" precision="5" value="3.5413">3.5413°E</geoCoordinate>
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), rodolith beds,
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<quantity id="4CB19B40FF88FFD1A9B2F8F0FE60F8E3" box="[449,508,1828,1855]" metricMagnitude="1" metricUnit="m" metricValue="7.2" pageId="5" pageNumber="236" unit="m" value="72.0">72 m</quantity>
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depth
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</elevation>
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, coll.
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<collectorName id="26BC5373FF88FFD1AAFCF8F1FC9AF8E2" box="[655,774,1828,1855]" pageId="5" pageNumber="236">F. Ordines</collectorName>
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during INTEMARES-CANAL0419 survey on
|
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<date id="FFF71065FF88FFD1AD55F8F0FF53F8BE" pageId="5" pageNumber="236" value="2019-04-27">
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<collectingDate id="EFB3E98DFF88FFD1AD55F8F0FF53F8BE" pageId="5" pageNumber="236" value="2019-04-27">April 27th 2019</collectingDate>
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</date>
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||
with haul #16 (
|
||
<figureCitation id="13722A20FF88FFD1A9F6F89CFE6EF8BE" box="[389,498,1864,1890]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="2.[151,250,1666,1691]" captionStartId-1="6.[151,250,1935,1960]" captionStartId-2="7.[151,250,1726,1751]" captionTargetBox-0="[151,1435,737,1642]" captionTargetBox-1="[151,1436,733,1915]" captionTargetBox-2="[177,1410,917,1703]" captionTargetId-0="figure@2.[151,1435,733,1647]" captionTargetId-1="figure@6.[151,1436,733,1915]" captionTargetId-2="figure@7.[170,1418,913,1707]" captionTargetPageId-0="2" captionTargetPageId-1="6" captionTargetPageId-2="7" captionText-0="FIGURE 1. Map showing the three sampling stations where all the individuals of Lebetus patzneri sp. nov. (stars) and L. guilleti newly recorded individuals (circles) from the Balearic Islands were collected. The holotype and the paratype of L. patzneri sp. nov. were collected from the sampling station in the Menorca Channel and off southeast Mallorca, respectively. The holotype, paratype and the two unpreserved specimens off south Menorca were collected at 72, 60 and 67 m, respectively. The 50, 100, 500 and 800 m isobaths are shown. Map drawn by F. Ordinas." captionText-1="FIGURE 2. Lebetus patzneri sp. nov., aspects of the freshly collected holotype, ZSM 47486, female, 16.6 mm SL, from off SW Mallorca Island, 72 m depth, preserved. a: left lateral view; b: right lateral view; c: ventral view; c: dorsal view; d: ventral view left lateral view. Photographs by F. Ordinas." captionText-2="FIGURE 3. Lebetus patzneri sp. nov., aspects of the preserved holotype, reversibly stained with Cyanine Blue, ZSM 47486, female, 16.6 mm SL, from off SW Mallorca Island, 72 m depth, preserved. a: left lateral view b: dorsal view; c: ventral view. Photographs by M. Kovačić." figureDoi-0="http://doi.org/10.5281/zenodo.3567338" figureDoi-1="http://doi.org/10.5281/zenodo.3567340" figureDoi-2="http://doi.org/10.5281/zenodo.3567342" httpUri-0="https://zenodo.org/record/3567338/files/figure.png" httpUri-1="https://zenodo.org/record/3567340/files/figure.png" httpUri-2="https://zenodo.org/record/3567342/files/figure.png" pageId="5" pageNumber="236">Figs. 1–3</figureCitation>
|
||
;
|
||
<figureCitation id="13722A20FF88FFD1AA73F89CFD8CF8BF" box="[512,528,1864,1891]" captionStart="FIGURE 6" captionStartId="11.[151,250,1257,1282]" captionTargetBox="[160,1436,607,1226]" captionTargetId="figure@11.[151,1436,589,1238]" captionTargetPageId="11" captionText="FIGURE 6. Lebetus patzneri sp. nov., head lateral line system of the holotype ZSM 47486, female, 16.6 mm SL, from off SW Mallorca Island, 72 m depth. Terminology in text. Drawing by M. Kovačić." figureDoi="http://doi.org/10.5281/zenodo.3567348" httpUri="https://zenodo.org/record/3567348/files/figure.png" pageId="5" pageNumber="236">6</figureCitation>
|
||
).
|
||
<collectorName id="26BC5373FF88FFD1AA54F89CFD09F8BE" box="[551,661,1864,1890]" pageId="5" pageNumber="236">Preserved</collectorName>
|
||
in 4% formol solution after capture, later transferred to 75% EtOH, right pectoral fin removed as DNA-tissue voucher ZSM-PIS-GO 2124.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BF636A5FF88FFD2A8B4F844FC5CFF04" blockId="5.[151,1436,1791,2035]" lastBlockId="6.[151,1437,152,719]" lastPageId="6" lastPageNumber="237" pageId="5" pageNumber="236">
|
||
<materialsCitation id="3B213CF8FF88FFD2A8B4F844FC20FF04" ID-GBIF-Occurrence="2465031521" collectingDate="2014-09-08" collectionCode="ZSM" collectorName="F. Ordines & Preserved" country="Spain" elevation="60" lastPageId="6" lastPageNumber="237" latitude="39.2372" location="Mallorca" longLatPrecision="6" longitude="2.9777" pageId="5" pageNumber="236" specimenCount="1" stateProvince="Illes Baleares" typeStatus="paratype">
|
||
<emphasis id="B93DEAB7FF88FFD1A8B4F844FEA9F876" bold="true" box="[199,309,1936,1962]" pageId="5" pageNumber="236">
|
||
<emphasis id="B93DEAB7FF88FFD1A8B4F844FEB2F876" bold="true" box="[199,302,1936,1962]" italics="true" pageId="5" pageNumber="236">
|
||
<typeStatus id="54F28807FF88FFD1A8B4F844FEB2F876" box="[199,302,1936,1962]" pageId="5" pageNumber="236" type="paratype">Paratype</typeStatus>
|
||
</emphasis>
|
||
.
|
||
</emphasis>
|
||
<collectionCode id="ED58AE60FF88FFD1A94DF844FEE4F876" box="[318,376,1936,1962]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="5" pageNumber="236" type="Museum">ZSM</collectionCode>
|
||
46865, female,
|
||
<quantity id="4CB19B40FF88FFD1AA47F844FD07F877" box="[564,667,1936,1963]" metricMagnitude="-2" metricUnit="m" metricValue="1.68" pageId="5" pageNumber="236" unit="mm" value="16.8">16.8 mm</quantity>
|
||
SL (caudal fin damaged),
|
||
<collectingCountry id="F35E7635FF88FFD1ABA3F844FB8FF876" box="[976,1043,1936,1962]" name="Spain" pageId="5" pageNumber="236">Spain</collectingCountry>
|
||
,
|
||
<collectingRegion id="498DF847FF88FFD1AC53F844FB45F876" box="[1056,1241,1936,1962]" country="Spain" name="Illes Baleares" pageId="5" pageNumber="236">Balearic Islands</collectingRegion>
|
||
, off the southern coast of
|
||
<location id="8E96607EFF88FFD1A88AF860FEC2F812" LSID="urn:lsid:plazi:treatment:03E087B3FF88FFDAA8E4F940FBC9FE0F:8E96607EFF88FFD1A88AF860FEC2F812" box="[249,350,1972,1998]" country="Spain" latitude="39.2372" longLatPrecision="6" longitude="2.9777" name="Mallorca" pageId="5" pageNumber="236" stateProvince="Illes Baleares">Mallorca</location>
|
||
(
|
||
<geoCoordinate id="EE7D5062FF88FFD1A91DF860FE71F812" box="[366,493,1972,1998]" degrees="39.2372" direction="north" orientation="latitude" pageId="5" pageNumber="236" precision="5" value="39.2372">39.2372°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE7D5062FF88FFD1A98AF860FDFBF812" box="[505,615,1972,1998]" degrees="2.9777" direction="east" orientation="longitude" pageId="5" pageNumber="236" precision="5" value="2.9777">2.9777°E</geoCoordinate>
|
||
), rodolith beds,
|
||
<elevation id="0064D196FF88FFD1AB6CF860FC3EF812" box="[799,930,1972,1999]" metricMagnitude="1" metricUnit="m" metricValue="6.0" pageId="5" pageNumber="236" unit="m" value="60.0">
|
||
<quantity id="4CB19B40FF88FFD1AB6CF860FCC5F813" box="[799,857,1972,1999]" metricMagnitude="1" metricUnit="m" metricValue="6.0" pageId="5" pageNumber="236" unit="m" value="60.0">60 m</quantity>
|
||
depth
|
||
</elevation>
|
||
, coll.
|
||
<collectorName id="26BC5373FF88FFD1AB9BF861FBC2F812" box="[1000,1118,1972,1999]" pageId="5" pageNumber="236">F. Ordines</collectorName>
|
||
during DRAGONSAL0914 survey on
|
||
<date id="FFF71065FF88FFD1A979F80CFE7DF82E" box="[266,481,2007,2034]" pageId="5" pageNumber="236" value="2014-09-08">
|
||
<collectingDate id="EFB3E98DFF88FFD1A979F80CFE7DF82E" box="[266,481,2007,2034]" pageId="5" pageNumber="236" value="2014-09-08">
|
||
September 8
|
||
<superScript id="7C3C9BEDFF88FFD1A9E5F803FE3FF839" attach="left" box="[406,419,2007,2021]" fontSize="6" pageId="5" pageNumber="236">th</superScript>
|
||
2014
|
||
</collectingDate>
|
||
</date>
|
||
with haul #25 (
|
||
<figureCitation id="13722A20FF88FFD1AAFEF80CFD40F82E" box="[653,732,2008,2034]" captionStart="FIGURE 4" captionStartId="8.[151,250,1427,1452]" captionTargetBox="[151,1436,519,1407]" captionTargetId="figure@8.[151,1436,517,1407]" captionTargetPageId="8" captionText="FIGURE 4. Lebetus patzneri sp. nov., SEM aspects of the paratype ZSM 46865, female, 16.8 mm SL; from off SW Mallorca Island, 60m depth. a: right lateral view; b: ventral view of pelvic fins and urogenital papilla. SEM photo by A. Cerwenka." figureDoi="http://doi.org/10.5281/zenodo.3567344" httpUri="https://zenodo.org/record/3567344/files/figure.png" pageId="5" pageNumber="236">Figs. 4</figureCitation>
|
||
,
|
||
<figureCitation id="13722A20FF88FFD1AA94F803FD6BF82E" box="[743,759,2007,2034]" captionStart="FIGURE 5" captionStartId="9.[151,250,1975,2000]" captionTargetBox="[170,1425,953,1948]" captionTargetId="figure@9.[162,1425,949,1956]" captionTargetPageId="9" captionText="FIGURE 5. Lebetus patzneri sp. nov., MicroCT reconstruction of the paratype ZSM 46865, female, 16.8 mm SL; from off SW Mallorca Island, 60m depth. a: left lateral view; b: dorsal view; c: ventral view. MicroCT reconstruction by A. Cerwenka." figureDoi="http://doi.org/10.5281/zenodo.3567346" httpUri="https://zenodo.org/record/3567346/files/figure.png" pageId="5" pageNumber="236">5</figureCitation>
|
||
,
|
||
<figureCitation id="13722A20FF88FFD1AB71F80DFC8EF82F" box="[770,786,2009,2035]" captionStart="FIGURE 7" captionStartId="12.[151,250,922,947]" captionTargetBox="[151,1435,337,902]" captionTargetId="figure@12.[151,1435,337,902]" captionTargetPageId="12" captionText="FIGURE 7. Lebetus patzneri sp. nov., right saccular otolith of the paratype ZSM 46865, female, 16.8 mm SL; from off SW Mallorca, 60m depth. a: in medial/inner view; b: left otolith in anterior view. Orientation of otolith with respect to fish symmetry is indicated as follows: a, anterior; d, dorsal; l, lateral; m, medial; p, posterior; v, ventral. MicroCT reconstruction by A. Cerwenka." figureDoi="http://doi.org/10.5281/zenodo.3567350" httpUri="https://zenodo.org/record/3567350/files/figure.png" pageId="5" pageNumber="236">7</figureCitation>
|
||
,
|
||
<figureCitation id="13722A20FF88FFD1AB6EF80CFCCCF82F" box="[797,848,2008,2035]" captionStart="FIGURE 8" captionStartId="13.[151,250,1801,1826]" captionTargetBox="[179,1412,181,1770]" captionTargetId="figure@13.[159,1429,166,1782]" captionTargetPageId="13" captionText="FIGURE 8. Lebetus patzneri sp. nov., aspects of the paratype and two unpreserved specimens in life coloration shortly after collection. a: left lateral view of paratype ZSM 46865, female, 16.8 mm SL; from off SW Mallorca Island, 60m depth (based on montage of two separate photographs (slight artifacts at montage suture at about D1 origin due to slightly different position of the specimen between the two photographs); b: left dorsolateral view of head and anterior flank view of paratype ZSM 46865; c: left lateral view of the second known specimen (unpreserved), unknown size, Menorca channel, 67m depth (based on montage of two separate photographs; stitching artifacts visible before D1 origin); d: left slightly dorsolateral view of third known specimen (unpreserved) of unknown size, Menorca channel, 67m depth. Photographs by F. Ordines." figureDoi="http://doi.org/10.5281/zenodo.3567352" httpUri="https://zenodo.org/record/3567352/files/figure.png" pageId="5" pageNumber="236">8a,b</figureCitation>
|
||
).
|
||
<collectorName id="26BC5373FF88FFD1AB10F80CFC4DF82E" box="[867,977,2008,2034]" pageId="5" pageNumber="236">Preserved</collectorName>
|
||
in 4% formol solution after capture, later transferred to 75% EtOH, and now stored as a critical-point-dried and gold-sputtered SEM specimen mounted on SEM mount (details of preparation for SEM see Material and Methods)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BF636A5FF8BFFD2A8B4FF35FB11FE9F" blockId="6.[151,1437,152,719]" pageId="6" pageNumber="237">
|
||
<emphasis id="B93DEAB7FF8BFFD2A8B4FF35FE31FF27" bold="true" box="[199,429,224,251]" pageId="6" pageNumber="237">Non-type material.</emphasis>
|
||
Photographs of two specimens (
|
||
<figureCitation id="13722A20FF8BFFD2AB5DFF35FC3EFF20" box="[814,930,225,252]" captionStart="FIGURE 8" captionStartId="13.[151,250,1801,1826]" captionTargetBox="[179,1412,181,1770]" captionTargetId="figure@13.[159,1429,166,1782]" captionTargetPageId="13" captionText="FIGURE 8. Lebetus patzneri sp. nov., aspects of the paratype and two unpreserved specimens in life coloration shortly after collection. a: left lateral view of paratype ZSM 46865, female, 16.8 mm SL; from off SW Mallorca Island, 60m depth (based on montage of two separate photographs (slight artifacts at montage suture at about D1 origin due to slightly different position of the specimen between the two photographs); b: left dorsolateral view of head and anterior flank view of paratype ZSM 46865; c: left lateral view of the second known specimen (unpreserved), unknown size, Menorca channel, 67m depth (based on montage of two separate photographs; stitching artifacts visible before D1 origin); d: left slightly dorsolateral view of third known specimen (unpreserved) of unknown size, Menorca channel, 67m depth. Photographs by F. Ordines." figureDoi="http://doi.org/10.5281/zenodo.3567352" httpUri="https://zenodo.org/record/3567352/files/figure.png" pageId="6" pageNumber="237">Figs. 8c,d</figureCitation>
|
||
) of unknown size and sex,
|
||
<collectingCountry id="F35E7635FF8BFFD2AC9EFF35FAACFF27" box="[1261,1328,225,251]" name="Spain" pageId="6" pageNumber="237">Spain</collectingCountry>
|
||
,
|
||
<collectingRegion id="498DF847FF8BFFD2AD4CFF35FF75FEC3" country="Spain" name="Illes Baleares" pageId="6" pageNumber="237">Balearic Islands</collectingRegion>
|
||
,
|
||
<collectingCountry id="F35E7635FF8BFFD2A884FED1FEC7FEC3" box="[247,347,261,287]" name="Spain" pageId="6" pageNumber="237">Menorca</collectingCountry>
|
||
channel (
|
||
<geoCoordinate id="EE7D5062FF8BFFD2A9BFFED1FDD6FEC3" box="[460,586,261,287]" degrees="39.8628" direction="north" orientation="latitude" pageId="6" pageNumber="237" precision="5" value="39.8628">39.8628°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE7D5062FF8BFFD2AA2BFED1FD59FEC3" box="[600,709,261,287]" degrees="4.0747" direction="east" orientation="longitude" pageId="6" pageNumber="237" precision="5" value="4.0747">4.0747°E</geoCoordinate>
|
||
), red algae beds,
|
||
<quantity id="4CB19B40FF8BFFD2ABFCFED1FC5DFEC3" box="[911,961,261,287]" metricMagnitude="1" metricUnit="m" metricValue="6.7" pageId="6" pageNumber="237" unit="m" value="67.0">67m</quantity>
|
||
depth, coll. F. Ordines during the INDE- MARES-CANAL0411 survey on
|
||
<date id="FFF71065FF8BFFD2AA67FEFDFD77FE9F" box="[532,747,296,323]" pageId="6" pageNumber="237" value="2011-09-07">
|
||
September 7
|
||
<superScript id="7C3C9BEDFF8BFFD2AAD2FEFCFD32FEEA" attach="left" box="[673,686,296,310]" fontSize="6" pageId="6" pageNumber="237">th</superScript>
|
||
2011
|
||
</date>
|
||
with beam trawl #26. Not preserved.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C353652EFF8BFFD2A8B4FE98FEF0FD13" pageId="6" pageNumber="237" type="diagnosis">
|
||
<paragraph id="8BF636A5FF8BFFD2A8B4FE98FEF0FD13" blockId="6.[151,1437,152,719]" pageId="6" pageNumber="237">
|
||
<emphasis id="B93DEAB7FF8BFFD2A8B4FE98FEA6FEBB" bold="true" box="[199,314,332,359]" pageId="6" pageNumber="237">Diagnosis</emphasis>
|
||
.
|
||
<taxonomicName id="4C494D26FF8BFFD2A93AFE9AFE5FFEBB" authority="Schliewen & Kovačić & Cerwenka & Svensen & Ordines, 2019" authorityName="Schliewen & Kovačić & Cerwenka & Svensen & Ordines" authorityYear="2019" box="[329,451,334,359]" class="Actinopterygii" family="Gobiidae" genus="Lebetus" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="237" phylum="Chordata" rank="species" species="patzneri" status="sp. nov.">
|
||
<emphasis id="B93DEAB7FF8BFFD2A93AFE9AFE5FFEBB" box="[329,451,334,359]" italics="true" pageId="6" pageNumber="237">L. patzneri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B93DEAB7FF8BFFD2A9B8FE99FDB9FEBB" bold="true" box="[459,549,333,359]" pageId="6" pageNumber="237">
|
||
<taxonomicNameLabel id="A20E57CCFF8BFFD2A9B8FE99FDB9FEBB" box="[459,549,333,359]" pageId="6" pageNumber="237" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is distinguished from the one or both other
|
||
<taxonomicName id="4C494D26FF8BFFD2AC66FE99FBF0FEBB" authorityName="Whinter" authorityYear="1877" box="[1045,1132,333,359]" class="Actinopterygii" family="Gobiidae" genus="Lebetus" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="237" phylum="Chordata" rank="genus">
|
||
<emphasis id="B93DEAB7FF8BFFD2AC66FE99FBF0FEBB" box="[1045,1132,333,359]" italics="true" pageId="6" pageNumber="237">Lebetus</emphasis>
|
||
</taxonomicName>
|
||
species by each of following characters: (1) P 15–16 vs.
|
||
<quantity id="4CB19B40FF8BFFD2A985FEA5FDC6FE57" box="[502,602,369,395]" metricMagnitude="-1" metricUnit="m" metricValue="4.826" metricValueMax="5.334" metricValueMin="4.318" pageId="6" pageNumber="237" unit="in" value="19.0" valueMax="21.0" valueMin="17.0">17–21 in</quantity>
|
||
<emphasis id="B93DEAB7FF8BFFD2AA12FEA6FC96FE57" box="[609,778,369,395]" italics="true" pageId="6" pageNumber="237">
|
||
<taxonomicName id="4C494D26FF8BFFD2AA12FEA6FC9EFE57" baseAuthorityName="Collett" baseAuthorityYear="1874" box="[609,770,369,395]" class="Actinopterygii" family="Gobiidae" genus="Lebetus" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="237" phylum="Chordata" rank="species" species="scorpioides">L. scorpioides</taxonomicName>
|
||
;
|
||
</emphasis>
|
||
(2) D2 segmented fin rays 7–8 vs.
|
||
<quantity id="4CB19B40FF8BFFD2ACEBFEA5FB72FE57" box="[1176,1262,369,395]" metricMagnitude="-1" metricUnit="m" metricValue="2.413" metricValueMax="2.54" metricValueMin="2.286" pageId="6" pageNumber="237" unit="in" value="9.5" valueMax="10.0" valueMin="9.0">9–10 in</quantity>
|
||
<taxonomicName id="4C494D26FF8BFFD2AC86FEA6FA08FE57" baseAuthorityName="Collett" baseAuthorityYear="1874" box="[1269,1428,369,395]" class="Actinopterygii" family="Gobiidae" genus="Lebetus" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="237" phylum="Chordata" rank="species" species="scorpioides">
|
||
<emphasis id="B93DEAB7FF8BFFD2AC86FEA6FA08FE57" box="[1269,1428,369,395]" italics="true" pageId="6" pageNumber="237">L. scorpioides</emphasis>
|
||
</taxonomicName>
|
||
; (3) snout slightly larger than eye diameter vs. snout clearly shorter than eye diameter in
|
||
<emphasis id="B93DEAB7FF8BFFD2AC0AFE42FB73FE73" box="[1145,1263,405,431]" italics="true" pageId="6" pageNumber="237">
|
||
<taxonomicName id="4C494D26FF8BFFD2AC0AFE42FB74FE73" baseAuthorityName="Le Danois" baseAuthorityYear="1913" box="[1145,1256,405,431]" class="Actinopterygii" family="Gobiidae" genus="Lebetus" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="237" phylum="Chordata" rank="species" species="guilleti">L. guilleti</taxonomicName>
|
||
;
|
||
</emphasis>
|
||
(4) eyes small, 23.3–25.5% of head length vs. eyes large, 29.4–35.7% of head length in
|
||
<taxonomicName id="4C494D26FF8BFFD2ABAFFE6EFBE0FE0F" baseAuthorityName="Collett" baseAuthorityYear="1874" box="[988,1148,441,467]" class="Actinopterygii" family="Gobiidae" genus="Lebetus" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="237" phylum="Chordata" rank="species" species="scorpioides">
|
||
<emphasis id="B93DEAB7FF8BFFD2ABAFFE6EFBE0FE0F" box="[988,1148,441,467]" italics="true" pageId="6" pageNumber="237">L. scorpioides</emphasis>
|
||
</taxonomicName>
|
||
and 28.8–35.5% of head length in
|
||
<taxonomicName id="4C494D26FF8BFFD2A977FE0AFEEFFE2B" baseAuthorityName="Le Danois" baseAuthorityYear="1913" box="[260,371,477,503]" class="Actinopterygii" family="Gobiidae" genus="Lebetus" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="237" phylum="Chordata" rank="species" species="guilleti">
|
||
<emphasis id="B93DEAB7FF8BFFD2A977FE0AFEEFFE2B" box="[260,371,477,503]" italics="true" pageId="6" pageNumber="237">L. guilleti</emphasis>
|
||
</taxonomicName>
|
||
; (5) body height low, body depth at pelvic fin 14.9–16.4% of SL vs. 16.6–21.1% of SL in
|
||
<taxonomicName id="4C494D26FF8BFFD2ADF5FE0AFF7AFDC7" baseAuthorityName="Le Danois" baseAuthorityYear="1913" class="Actinopterygii" family="Gobiidae" genus="Lebetus" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="237" phylum="Chordata" rank="species" species="guilleti">
|
||
<emphasis id="B93DEAB7FF8BFFD2ADF5FE0AFF7AFDC7" italics="true" pageId="6" pageNumber="237">L. guilleti</emphasis>
|
||
</taxonomicName>
|
||
; (6) lips thick, giving mouth a broad, almost rectangular shape in dorsal and lateral view vs. lips of normal size, mouth of oval shape in dorsal and lateral view in both
|
||
<taxonomicName id="4C494D26FF8BFFD2AB58FDF2FC55FDE3" baseAuthorityName="Collett" baseAuthorityYear="1874" box="[811,969,549,575]" class="Actinopterygii" family="Gobiidae" genus="Lebetus" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="237" phylum="Chordata" rank="species" species="scorpioides">
|
||
<emphasis id="B93DEAB7FF8BFFD2AB58FDF2FC55FDE3" box="[811,969,549,575]" italics="true" pageId="6" pageNumber="237">L. scorpioides</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C494D26FF8BFFD2AB8CFDF2FBF7FDE3" baseAuthorityName="Le Danois" baseAuthorityYear="1913" box="[1023,1131,549,575]" class="Actinopterygii" family="Gobiidae" genus="Lebetus" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="237" phylum="Chordata" rank="species" species="guilleti">
|
||
<emphasis id="B93DEAB7FF8BFFD2AB8CFDF2FBF7FDE3" box="[1023,1131,549,575]" italics="true" pageId="6" pageNumber="237">L. guilleti</emphasis>
|
||
</taxonomicName>
|
||
; (7) minute black “pepper” spots on predorsal area, dorsum, flanks and caudal peduncle, visible in fresh and preserved specimens vs. no similar spots in
|
||
<taxonomicName id="4C494D26FF8BFFD2A880FDBAFEC3FD5B" baseAuthorityName="Le Danois" baseAuthorityYear="1913" box="[243,351,621,647]" class="Actinopterygii" family="Gobiidae" genus="Lebetus" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="237" phylum="Chordata" rank="species" species="guilleti">
|
||
<emphasis id="B93DEAB7FF8BFFD2A880FDBAFEC3FD5B" box="[243,351,621,647]" italics="true" pageId="6" pageNumber="237">L. guilleti</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C494D26FF8BFFD2A9E7FDBAFDAEFD5B" baseAuthorityName="Collett" baseAuthorityYear="1874" box="[404,562,621,647]" class="Actinopterygii" family="Gobiidae" genus="Lebetus" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="237" phylum="Chordata" rank="species" species="scorpioides">
|
||
<emphasis id="B93DEAB7FF8BFFD2A9E7FDBAFDAEFD5B" box="[404,562,621,647]" italics="true" pageId="6" pageNumber="237">L. scorpioides</emphasis>
|
||
</taxonomicName>
|
||
; (8) the most of the caudal peduncle similarly colored as the lateral area below D2, reddish or brown vs. sharp dark-pale boundary between caudal peduncle and the rest of the body in
|
||
<taxonomicName id="4C494D26FF8BFFD2AD5CFD46FA00FD77" baseAuthorityName="Le Danois" baseAuthorityYear="1913" box="[1327,1436,657,683]" class="Actinopterygii" family="Gobiidae" genus="Lebetus" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="237" phylum="Chordata" rank="species" species="guilleti">
|
||
<emphasis id="B93DEAB7FF8BFFD2AD5CFD46FA00FD77" box="[1327,1436,657,683]" italics="true" pageId="6" pageNumber="237">L. guilleti</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C494D26FF8BFFD2A8B4FD62FEF9FD13" baseAuthorityName="Collett" baseAuthorityYear="1874" box="[199,357,693,719]" class="Actinopterygii" family="Gobiidae" genus="Lebetus" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="237" phylum="Chordata" rank="species" species="scorpioides">
|
||
<emphasis id="B93DEAB7FF8BFFD2A8B4FD62FEF9FD13" box="[199,357,693,719]" italics="true" pageId="6" pageNumber="237">L. scorpioides</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF36662DFF8BFFD2A8E4F85BFDCAF82C" ID-DOI="http://doi.org/10.5281/zenodo.3567340" ID-Zenodo-Dep="3567340" httpUri="https://zenodo.org/record/3567340/files/figure.png" pageId="6" pageNumber="237" startId="6.[151,250,1935,1960]" targetBox="[151,1436,733,1915]" targetPageId="6">
|
||
<paragraph id="8BF636A5FF8BFFD2A8E4F85BFDCAF82C" blockId="6.[151,1437,1935,2032]" pageId="6" pageNumber="237">
|
||
<emphasis id="B93DEAB7FF8BFFD2A8E4F85BFE8EF874" bold="true" box="[151,274,1935,1960]" pageId="6" pageNumber="237">FIGURE 2.</emphasis>
|
||
<taxonomicName id="4C494D26FF8BFFD2A96BF85BFE23F87B" authority="Schliewen & Kovačić & Cerwenka & Svensen & Ordines, 2019" authorityName="Schliewen & Kovačić & Cerwenka & Svensen & Ordines" authorityYear="2019" box="[280,447,1935,1959]" class="Actinopterygii" family="Gobiidae" genus="Lebetus" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="237" phylum="Chordata" rank="species" species="patzneri" status="sp. nov.">
|
||
<emphasis id="B93DEAB7FF8BFFD2A96BF85BFE23F87B" box="[280,447,1935,1959]" italics="true" pageId="6" pageNumber="237">Lebetus patzneri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B93DEAB7FF8BFFD2A9B6F85BFD88F87B" bold="true" box="[453,532,1935,1959]" pageId="6" pageNumber="237">
|
||
<taxonomicNameLabel id="A20E57CCFF8BFFD2A9B6F85BFD88F87B" box="[453,532,1935,1959]" pageId="6" pageNumber="237" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, aspects of the freshly collected holotype, ZSM 47486, female, 16.6 mm SL, from off SW Mallorca Island, 72 m depth, preserved.
|
||
<emphasis id="B93DEAB7FF8BFFD2AA5CF867FDA0F817" bold="true" box="[559,572,1971,1995]" pageId="6" pageNumber="237">a</emphasis>
|
||
: left lateral view;
|
||
<emphasis id="B93DEAB7FF8BFFD2AA80F867FC9DF817" bold="true" box="[755,769,1971,1995]" pageId="6" pageNumber="237">b</emphasis>
|
||
: right lateral view;
|
||
<emphasis id="B93DEAB7FF8BFFD2ABB4F867FC4EF817" bold="true" box="[967,978,1971,1995]" pageId="6" pageNumber="237">c</emphasis>
|
||
: ventral view;
|
||
<emphasis id="B93DEAB7FF8BFFD2AC19F867FBE9F817" bold="true" box="[1130,1141,1971,1995]" pageId="6" pageNumber="237">c</emphasis>
|
||
: dorsal view;
|
||
<emphasis id="B93DEAB7FF8BFFD2AD77F867FA8EF817" bold="true" box="[1284,1298,1971,1995]" pageId="6" pageNumber="237">d</emphasis>
|
||
: ventral view left lateral view. Photographs by F. Ordinas.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C353652EFF8AFFD8A8B4FF4CFAF0FB0F" lastPageId="12" lastPageNumber="243" pageId="7" pageNumber="238" type="description">
|
||
<paragraph id="8BF636A5FF8AFFD3A8B4FF4CFE91FC5F" blockId="7.[151,1437,152,899]" pageId="7" pageNumber="238">
|
||
<emphasis id="B93DEAB7FF8AFFD3A8B4FF4CFECDFF6F" bold="true" box="[199,337,152,179]" pageId="7" pageNumber="238">Description</emphasis>
|
||
(all morphometric values and meristics in the text are presented as
|
||
<typeStatus id="54F28807FF8AFFD3AC14FF4DFB55FF6F" box="[1127,1225,153,179]" pageId="7" pageNumber="238" type="holotype">holotype</typeStatus>
|
||
first and
|
||
<typeStatus id="54F28807FF8AFFD3AD4FFF4DFA00FF6F" box="[1340,1436,153,179]" pageId="7" pageNumber="238" type="paratype">paratype</typeStatus>
|
||
in parentheses).
|
||
<emphasis id="B93DEAB7FF8AFFD3A922FF69FDA3FF0B" box="[337,575,189,215]" italics="true" pageId="7" pageNumber="238">General Morphology</emphasis>
|
||
(
|
||
<figureCitation id="13722A20FF8AFFD3AA23FF69FD22FF0B" box="[592,702,189,215]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStartId-0="6.[151,250,1935,1960]" captionStartId-1="7.[151,250,1726,1751]" captionStartId-2="8.[151,250,1427,1452]" captionTargetBox-0="[151,1436,733,1915]" captionTargetBox-1="[177,1410,917,1703]" captionTargetBox-2="[151,1436,519,1407]" captionTargetId-0="figure@6.[151,1436,733,1915]" captionTargetId-1="figure@7.[170,1418,913,1707]" captionTargetId-2="figure@8.[151,1436,517,1407]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="8" captionText-0="FIGURE 2. Lebetus patzneri sp. nov., aspects of the freshly collected holotype, ZSM 47486, female, 16.6 mm SL, from off SW Mallorca Island, 72 m depth, preserved. a: left lateral view; b: right lateral view; c: ventral view; c: dorsal view; d: ventral view left lateral view. Photographs by F. Ordinas." captionText-1="FIGURE 3. Lebetus patzneri sp. nov., aspects of the preserved holotype, reversibly stained with Cyanine Blue, ZSM 47486, female, 16.6 mm SL, from off SW Mallorca Island, 72 m depth, preserved. a: left lateral view b: dorsal view; c: ventral view. Photographs by M. Kovačić." captionText-2="FIGURE 4. Lebetus patzneri sp. nov., SEM aspects of the paratype ZSM 46865, female, 16.8 mm SL; from off SW Mallorca Island, 60m depth. a: right lateral view; b: ventral view of pelvic fins and urogenital papilla. SEM photo by A. Cerwenka." figureDoi-0="http://doi.org/10.5281/zenodo.3567340" figureDoi-1="http://doi.org/10.5281/zenodo.3567342" figureDoi-2="http://doi.org/10.5281/zenodo.3567344" httpUri-0="https://zenodo.org/record/3567340/files/figure.png" httpUri-1="https://zenodo.org/record/3567342/files/figure.png" httpUri-2="https://zenodo.org/record/3567344/files/figure.png" pageId="7" pageNumber="238">Figs. 2–4</figureCitation>
|
||
;
|
||
<tableCitation id="C6CB031EFF8AFFD3AAB9FF69FCBDFF0B" box="[714,801,189,215]" captionStart="TABLE 2" captionStartId="10.[151,239,154,178]" captionTargetBox="[166,1345,225,1833]" captionText="TABLE 2. Morphometric characters of Lebetus patzneri sp. nov. holotype (ZSM 47486; female) and paratype (ZSM 46865; female); absolute values and % values in brackets. An asterik (*) indicates a damaged character." httpUri="http://table.plazi.org/id/DF36662DFF87FFDEA8E4FF4EFB74FF0A" pageId="7" pageNumber="238" tableUuid="DF36662DFF87FFDEA8E4FF4EFB74FF0A">Table 2</tableCitation>
|
||
). Body proportions and meristic counts of
|
||
<typeStatus id="54F28807FF8AFFD3AD7AFF69FAF7FF0B" box="[1289,1387,189,215]" pageId="7" pageNumber="238" type="holotype">holotype</typeStatus>
|
||
and
|
||
<typeStatus id="54F28807FF8AFFD3A8E4FF35FF6BFF27" box="[151,247,225,251]" pageId="7" pageNumber="238" type="paratype">paratype</typeStatus>
|
||
are given in
|
||
<tableCitation id="C6CB031EFF8AFFD3A9FEFF35FE78FF27" box="[397,484,225,251]" captionStart="TABLE 2" captionStartId="10.[151,239,154,178]" captionTargetBox="[166,1345,225,1833]" captionText="TABLE 2. Morphometric characters of Lebetus patzneri sp. nov. holotype (ZSM 47486; female) and paratype (ZSM 46865; female); absolute values and % values in brackets. An asterik (*) indicates a damaged character." httpUri="http://table.plazi.org/id/DF36662DFF87FFDEA8E4FF4EFB74FF0A" pageId="7" pageNumber="238" tableUuid="DF36662DFF87FFDEA8E4FF4EFB74FF0A">Table 2</tableCitation>
|
||
. Body elongate, subcylindrical and slightly laterally compressed towards caudal fin. Head fairly long 32.2 (32.5) % SL, slightly depressed, with a nearly horizontal predorsal profile. Snout moderately long and pointed, about as long as eye 105.2 (106.3) % in eye length, dorsal profile of snout gently sloping. Eyes dorsolaterally extending in fresh specimens above dorsal profile, large 25.5 (23.3) % of head length, interorbital space narrow, about one-fourth of eye length. Anterior and posterior nostrils both short, not reaching upper lip, tubular, erect, without process from rim; anterior nostrils slightly longer than posterior ones. Mouth oblique from lateral view, posterior angle of jaws below anterior third of eye. Lips ending about equally anteriorly, thick, giving mouth a broad, almost rectangular shape in dorsal and lateral view. Minimum teeth counts of inner rows were made for
|
||
<typeStatus id="54F28807FF8AFFD3A8CDFDD5FE82FDC7" box="[190,286,513,539]" pageId="7" pageNumber="238" type="paratype">paratype</typeStatus>
|
||
only, because for the
|
||
<typeStatus id="54F28807FF8AFFD3AA63FDD5FDEEFDC7" box="[528,626,513,539]" pageId="7" pageNumber="238" type="holotype">holotype</typeStatus>
|
||
no high-resolution MicroCT image was available; however roughly estimated numbers for
|
||
<typeStatus id="54F28807FF8AFFD3A902FDF1FE4FFDE3" box="[369,467,549,575]" pageId="7" pageNumber="238" type="holotype">holotype</typeStatus>
|
||
are similar to the
|
||
<typeStatus id="54F28807FF8AFFD3AAEFFDF1FC9DFDE3" box="[668,769,549,575]" pageId="7" pageNumber="238" type="paratype">paratype</typeStatus>
|
||
: lower jaw with three rows of unicuspid teeth at symphysis, the outer row with more than twenty (left: 22 (25), right; 26 (21)) upright teeth, the second (intermediate) row with very small and irregularly set teeth along the anterior jaw part, and the third row with teeth as large as in the first row, but more slender, the first reaching back to the jaw angle, the third row reaching back almost as far as the first row. Upper jaw with two rows, the outer row with slender and backwards curved teeth (more than 20 on both sides in
|
||
<typeStatus id="54F28807FF8AFFD3A8C6FD0DFE86FD2F" box="[181,282,729,755]" pageId="7" pageNumber="238" type="holotype">holotype</typeStatus>
|
||
, at least 17 on each side in
|
||
<typeStatus id="54F28807FF8AFFD3AA22FD0DFD2AFD2F" box="[593,694,729,755]" pageId="7" pageNumber="238" type="paratype">paratype</typeStatus>
|
||
), and an interior row of numerous small to minute and irregularly set teeth, reaching farther back to the jaw angle than the anterior row. Chin without mental flap (fold) or barbels. Cranial roof covered with dorsal axial musculature. Pectoral girdle without flaps on anterior edge. Genital papilla short, broad and with terminal villose processes on the posterior and lateral parts, but without on the anterior part of the rim.
|
||
</paragraph>
|
||
<caption id="DF36662DFF8AFFD3A8E4F96AFE27F8C3" ID-DOI="http://doi.org/10.5281/zenodo.3567342" ID-Zenodo-Dep="3567342" httpUri="https://zenodo.org/record/3567342/files/figure.png" pageId="7" pageNumber="238" startId="7.[151,250,1726,1751]" targetBox="[177,1410,917,1703]" targetPageId="7">
|
||
<paragraph id="8BF636A5FF8AFFD3A8E4F96AFE27F8C3" blockId="7.[151,1437,1726,1823]" pageId="7" pageNumber="238">
|
||
<emphasis id="B93DEAB7FF8AFFD3A8E4F96AFE88F90B" bold="true" box="[151,276,1726,1751]" pageId="7" pageNumber="238">FIGURE 3.</emphasis>
|
||
<taxonomicName id="4C494D26FF8AFFD3A96FF96BFE59F90B" authority="Schliewen & Kovačić & Cerwenka & Svensen & Ordines, 2019" authorityName="Schliewen & Kovačić & Cerwenka & Svensen & Ordines" authorityYear="2019" box="[284,453,1727,1751]" class="Actinopterygii" family="Gobiidae" genus="Lebetus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="238" phylum="Chordata" rank="species" species="patzneri" status="sp. nov.">
|
||
<emphasis id="B93DEAB7FF8AFFD3A96FF96BFE59F90B" box="[284,453,1727,1751]" italics="true" pageId="7" pageNumber="238">Lebetus patzneri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B93DEAB7FF8AFFD3A9BEF96BFD82F90B" bold="true" box="[461,542,1727,1751]" pageId="7" pageNumber="238">
|
||
<taxonomicNameLabel id="A20E57CCFF8AFFD3A9BEF96BFD82F90B" box="[461,542,1727,1751]" pageId="7" pageNumber="238" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, aspects of the preserved holotype, reversibly stained with Cyanine Blue, ZSM 47486, female, 16.6 mm SL, from off SW Mallorca Island, 72 m depth, preserved.
|
||
<emphasis id="B93DEAB7FF8AFFD3ABEEF937FC36F927" bold="true" box="[925,938,1763,1787]" pageId="7" pageNumber="238">a</emphasis>
|
||
: left lateral view
|
||
<emphasis id="B93DEAB7FF8AFFD3AC2CF937FBF1F927" bold="true" box="[1119,1133,1763,1787]" pageId="7" pageNumber="238">b</emphasis>
|
||
: dorsal view;
|
||
<emphasis id="B93DEAB7FF8AFFD3AD73F937FA97F927" bold="true" box="[1280,1291,1763,1787]" pageId="7" pageNumber="238">c</emphasis>
|
||
: ventral view. Photographs by M. Kovačić.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF636A5FF8AFFDCA8B4F89AFD65FE9F" blockId="7.[151,1436,1869,2040]" lastBlockId="8.[151,1437,153,504]" lastPageId="8" lastPageNumber="239" pageId="7" pageNumber="238">
|
||
<emphasis id="B93DEAB7FF8AFFD3A8B4F89AFF65F8BB" box="[199,249,1870,1895]" italics="true" pageId="7" pageNumber="238">Fins</emphasis>
|
||
(
|
||
<figureCitation id="13722A20FF8AFFD3A97AF899FED2F8B4" box="[265,334,1869,1896]" captionStart="FIGURE 5" captionStartId="9.[151,250,1975,2000]" captionTargetBox="[170,1425,953,1948]" captionTargetId="figure@9.[162,1425,949,1956]" captionTargetPageId="9" captionText="FIGURE 5. Lebetus patzneri sp. nov., MicroCT reconstruction of the paratype ZSM 46865, female, 16.8 mm SL; from off SW Mallorca Island, 60m depth. a: left lateral view; b: dorsal view; c: ventral view. MicroCT reconstruction by A. Cerwenka." figureDoi="http://doi.org/10.5281/zenodo.3567346" httpUri="https://zenodo.org/record/3567346/files/figure.png" pageId="7" pageNumber="238">Fig. 5</figureCitation>
|
||
). D
|
||
<date id="FFF71065FF8AFFD3A90FF89AFE37F8B4" box="[380,427,1870,1896]" pageId="7" pageNumber="238">1 VI</date>
|
||
(VII); D
|
||
<date id="FFF71065FF8AFFD3AA7CF89AFDB1F8B4" box="[527,557,1870,1896]" pageId="7" pageNumber="238">2 I</date>
|
||
/7 (
|
||
<date id="FFF71065FF8AFFD3AA20F89AFDE8F8B4" box="[595,628,1870,1896]" bridgedPair="/" pageId="7" pageNumber="238">I/8</date>
|
||
) (last bifid); A
|
||
<date id="FFF71065FF8AFFD3AB51F899FCD8F8BB" box="[802,836,1869,1895]" bridgedPair="/" pageId="7" pageNumber="238">I/6</date>
|
||
; C 9 branched rays, 13 segmented rays (caudal fin of
|
||
<typeStatus id="54F28807FF8AFFD3A8E4F8A6FF6BF850" box="[151,247,1906,1932]" pageId="7" pageNumber="238" type="paratype">paratype</typeStatus>
|
||
damaged, therefore identification as branched and segmented tentative); P 16 (16 left and 15 right), V
|
||
<date id="FFF71065FF8AFFD3AD14F8A5FA1BF850" box="[1383,1415,1905,1932]" bridgedPair="/" pageId="7" pageNumber="238">I/5</date>
|
||
+
|
||
<date id="FFF71065FF8AFFD3A8E4F841FF21F86C" box="[151,189,1941,1968]" bridgedPair="/" pageId="7" pageNumber="238">5/I.</date>
|
||
Fin-bases and lengths in proportion to standard length are given in
|
||
<tableCitation id="C6CB031EFF8AFFD3ABD5F841FB81F86C" box="[934,1053,1941,1968]" captionStart="TABLE 2" captionStartId="10.[151,239,154,178]" captionTargetBox="[166,1345,225,1833]" captionText="TABLE 2. Morphometric characters of Lebetus patzneri sp. nov. holotype (ZSM 47486; female) and paratype (ZSM 46865; female); absolute values and % values in brackets. An asterik (*) indicates a damaged character." httpUri="http://table.plazi.org/id/DF36662DFF87FFDEA8E4FF4EFB74FF0A" pageId="7" pageNumber="238" tableUuid="DF36662DFF87FFDEA8E4FF4EFB74FF0A">Table 2. D</tableCitation>
|
||
<date id="FFF71065FF8AFFD3AC6EF842FB0CF86C" box="[1053,1168,1941,1968]" pageId="7" pageNumber="238">
|
||
<tableCitation id="C6CB031EFF8AFFD3AC6EF842FBBBF86C" box="[1053,1063,1942,1968]" captionStart="TABLE 1" captionStartId="4.[151,239,982,1006]" captionTargetBox="[166,1387,1090,1962]" captionText="TABLE 1. Adjustment of the Lebetus lateral line system description to the terminology for gobies with a more complex head lateral line system (Sanzo 1911, Miller 1986) compared to the same in Kovačić et al. (2017, 2018) for the reduced head lateral line systems of related Buenia species and to Miller’s (1963) terminology for Lebetus." httpUri="http://table.plazi.org/id/DF36662DFF89FFD0A8E4FC02FB2CFBEA" pageId="7" pageNumber="238" tableUuid="DF36662DFF89FFD0A8E4FC02FB2CFBEA">1</tableCitation>
|
||
II and III
|
||
</date>
|
||
the longest spine of D1, their tips just reaching D
|
||
<date id="FFF71065FF8AFFD3A9D4F86EFE58F808" box="[423,452,1978,2004]" pageId="7" pageNumber="238">2 I</date>
|
||
anterior origin. Fin membrane of D
|
||
<date id="FFF71065FF8AFFD3AB21F86EFC1CF808" box="[850,896,1978,2004]" pageId="7" pageNumber="238">1 VI</date>
|
||
not connected with base of D
|
||
<date id="FFF71065FF8AFFD3ACB4F86EFB76F808" box="[1223,1258,1978,2004]" pageId="7" pageNumber="238">2 I.</date>
|
||
D2 commences at vertical of urogenital papilla, with its last ray over vertical of last A ray. A commences below third (
|
||
<typeStatus id="54F28807FF8AFFD3AD62F809FAEAF824" box="[1297,1398,2013,2040]" pageId="7" pageNumber="238" type="holotype">holotype</typeStatus>
|
||
) or second (
|
||
<typeStatus id="54F28807FF85FFDCA887FF4DFEC4FF6F" box="[244,344,153,179]" pageId="8" pageNumber="239" type="paratype">paratype</typeStatus>
|
||
) segmented ray of D2. C rounded, shorter than head length. Uppermost P rays within membrane, all P rays unbranched, ending at vertical of D2 spine (
|
||
<typeStatus id="54F28807FF85FFDCAA9BFF69FCCCFF04" box="[744,848,189,216]" pageId="8" pageNumber="239" type="holotype">holotype</typeStatus>
|
||
), or just before it (
|
||
<typeStatus id="54F28807FF85FFDCAC6AFF69FBE2FF0B" box="[1049,1150,189,215]" pageId="8" pageNumber="239" type="paratype">paratype</typeStatus>
|
||
, but potential damage has to be taken into account). V without an anterior membrane; V fin membranes strongly damaged, i.e. it is not possible to infer shape and completeness of pectoral disc. V longest ray either not reaching genital papilla (
|
||
<typeStatus id="54F28807FF85FFDCACA9FED1FADDFEC3" box="[1242,1345,261,287]" pageId="8" pageNumber="239" type="holotype">holotype</typeStatus>
|
||
; longest ray damaged?) or reaching slightly beyond (
|
||
<typeStatus id="54F28807FF85FFDCAAF5FEFDFD71FE9F" box="[646,749,297,323]" pageId="8" pageNumber="239" type="paratype">paratype</typeStatus>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BF636A5FF85FFDCA8B4FE99FA06FE2B" blockId="8.[151,1437,153,504]" pageId="8" pageNumber="239">
|
||
<emphasis id="B93DEAB7FF85FFDCA8B4FE99FED1FEBB" box="[199,333,333,359]" italics="true" pageId="8" pageNumber="239">Squamation</emphasis>
|
||
(
|
||
<figureCitation id="13722A20FF85FFDCA928FE99FE35FEBB" box="[347,425,333,359]" captionStart="FIGURE 3" captionStartId="7.[151,250,1726,1751]" captionTargetBox="[177,1410,917,1703]" captionTargetId="figure@7.[170,1418,913,1707]" captionTargetPageId="7" captionText="FIGURE 3. Lebetus patzneri sp. nov., aspects of the preserved holotype, reversibly stained with Cyanine Blue, ZSM 47486, female, 16.6 mm SL, from off SW Mallorca Island, 72 m depth, preserved. a: left lateral view b: dorsal view; c: ventral view. Photographs by M. Kovačić." figureDoi="http://doi.org/10.5281/zenodo.3567342" httpUri="https://zenodo.org/record/3567342/files/figure.png" pageId="8" pageNumber="239">Figs. 3</figureCitation>
|
||
,
|
||
<figureCitation id="13722A20FF85FFDCA9C6FE99FE4FFEBB" box="[437,467,333,359]" captionStart="FIGURE 4" captionStartId="8.[151,250,1427,1452]" captionTargetBox="[151,1436,519,1407]" captionTargetId="figure@8.[151,1436,517,1407]" captionTargetPageId="8" captionText="FIGURE 4. Lebetus patzneri sp. nov., SEM aspects of the paratype ZSM 46865, female, 16.8 mm SL; from off SW Mallorca Island, 60m depth. a: right lateral view; b: ventral view of pelvic fins and urogenital papilla. SEM photo by A. Cerwenka." figureDoi="http://doi.org/10.5281/zenodo.3567344" httpUri="https://zenodo.org/record/3567344/files/figure.png" pageId="8" pageNumber="239">4a</figureCitation>
|
||
): Body scaled. Since both specimens lost scales, scale counts are based on scale pouches visible on the stained
|
||
<typeStatus id="54F28807FF85FFDCA9C2FEA5FD8FFE57" box="[433,531,369,395]" pageId="8" pageNumber="239" type="holotype">holotype</typeStatus>
|
||
and on the REM image of the
|
||
<typeStatus id="54F28807FF85FFDCAB04FEA5FC4BFE57" box="[887,983,369,395]" pageId="8" pageNumber="239" type="paratype">paratype</typeStatus>
|
||
and scale types could not be identified. Head including opercle and cheek naked. Predorsal area, including nape, and the area below D1 downwards to the horizontal level of upper edge of pectoral fin base naked. Breast and belly naked. Scales in lateral series 25 on both sides plus one on caudal fin origin (23 plus one on caudal fin count on the right side). Scales in transverse series 7.
|
||
</paragraph>
|
||
<caption id="DF36662DFF85FFDCA8E4FA47FAF6FA0C" ID-DOI="http://doi.org/10.5281/zenodo.3567344" ID-Zenodo-Dep="3567344" httpUri="https://zenodo.org/record/3567344/files/figure.png" pageId="8" pageNumber="239" startId="8.[151,250,1427,1452]" targetBox="[151,1436,519,1407]" targetPageId="8">
|
||
<paragraph id="8BF636A5FF85FFDCA8E4FA47FAF6FA0C" blockId="8.[151,1436,1427,1488]" pageId="8" pageNumber="239">
|
||
<emphasis id="B93DEAB7FF85FFDCA8E4FA47FE8FFA70" bold="true" box="[151,275,1427,1452]" pageId="8" pageNumber="239">FIGURE 4.</emphasis>
|
||
<taxonomicName id="4C494D26FF85FFDCA969FA40FE5FFA77" authority="Schliewen & Kovačić & Cerwenka & Svensen & Ordines, 2019" authorityName="Schliewen & Kovačić & Cerwenka & Svensen & Ordines" authorityYear="2019" box="[282,451,1428,1452]" class="Actinopterygii" family="Gobiidae" genus="Lebetus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="239" phylum="Chordata" rank="species" species="patzneri" status="sp. nov.">
|
||
<emphasis id="B93DEAB7FF85FFDCA969FA40FE5FFA77" box="[282,451,1428,1452]" italics="true" pageId="8" pageNumber="239">Lebetus patzneri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B93DEAB7FF85FFDCA9B9FA40FD87FA70" bold="true" box="[458,539,1428,1452]" pageId="8" pageNumber="239">
|
||
<taxonomicNameLabel id="A20E57CCFF85FFDCA9B9FA40FD87FA70" box="[458,539,1428,1452]" pageId="8" pageNumber="239" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, SEM aspects of the paratype ZSM 46865, female, 16.8 mm SL; from off SW Mallorca Island, 60m depth.
|
||
<emphasis id="B93DEAB7FF85FFDCA929FA6CFEFBFA0C" bold="true" box="[346,359,1464,1488]" pageId="8" pageNumber="239">a</emphasis>
|
||
: right lateral view;
|
||
<emphasis id="B93DEAB7FF85FFDCAA5CFA6CFDA1FA0C" bold="true" box="[559,573,1464,1488]" pageId="8" pageNumber="239">b</emphasis>
|
||
: ventral view of pelvic fins and urogenital papilla. SEM photo by A. Cerwenka.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF636A5FF85FFDCA8B4FA2AFC9FF881" blockId="8.[151,1437,1533,2029]" pageId="8" pageNumber="239">
|
||
<emphasis id="B93DEAB7FF85FFDCA8B4FA2AFEA5F9C4" box="[199,313,1534,1560]" italics="true" pageId="8" pageNumber="239">Osteology</emphasis>
|
||
. Postcranial skeleton and pterygiophore insertion pattern (
|
||
<figureCitation id="13722A20FF85FFDCABA1FA2AFB84F9C4" box="[978,1048,1533,1560]" captionStart="FIGURE 5" captionStartId="9.[151,250,1975,2000]" captionTargetBox="[170,1425,953,1948]" captionTargetId="figure@9.[162,1425,949,1956]" captionTargetPageId="9" captionText="FIGURE 5. Lebetus patzneri sp. nov., MicroCT reconstruction of the paratype ZSM 46865, female, 16.8 mm SL; from off SW Mallorca Island, 60m depth. a: left lateral view; b: dorsal view; c: ventral view. MicroCT reconstruction by A. Cerwenka." figureDoi="http://doi.org/10.5281/zenodo.3567346" httpUri="https://zenodo.org/record/3567346/files/figure.png" pageId="8" pageNumber="239">Fig. 5</figureCitation>
|
||
). 11 (12) precaudal and 15 caudal vertebrae including urostyle; total count 26 (27). In lateral view, first three vertebral centra short, becoming progressively longer in vertebrae 4 and 5, and approximately of equal lengths further posteriorly. First three vertebrae substantially, fourth vertebrae slightly shorter than posterior ones. Neural spines and haemal spines of last four vertebrae before urostyl increasingly enlarged posteriorly, especially last two haemal spine. D1 pterygiophore formula 3-
|
||
<date id="FFF71065FF85FFDCA8D9F966FE9DF910" box="[170,257,1714,1740]" pageId="8" pageNumber="239" value="00-11-22">221100</date>
|
||
(3-2220100); one prehaemal A pterygiophore. Four branched caudal fin rays inserting at hypurals 1 + 2 (fused), five at hypurals 3 + 4 (fused with each other and with urostyl); 1 branched and 1 unbranched ray in hypural 5; one branched ray in parhypural; each one unbranched caudal ray beyond hypural 5 and parhypural, respectively; total number of fin rays inserting in hypuralia and parhypural 12. Hypural 5 and parhypural small splint-like bones. One epural. Five dorsal and six ventral procurrent rays.
|
||
</paragraph>
|
||
<paragraph id="8BF636A5FF85FFDDA8B4F8B2FD32FC7B" blockId="8.[151,1437,1533,2029]" lastBlockId="9.[151,1437,152,935]" lastPageId="9" lastPageNumber="240" pageId="8" pageNumber="239">
|
||
<emphasis id="B93DEAB7FF85FFDCA8B4F8B2FE01F85C" box="[199,413,1894,1920]" italics="true" pageId="8" pageNumber="239">Lateral line system</emphasis>
|
||
(
|
||
<figureCitation id="13722A20FF85FFDCA9D8F8B2FE6CF85D" box="[427,496,1894,1921]" captionStart="FIGURE 6" captionStartId="11.[151,250,1257,1282]" captionTargetBox="[160,1436,607,1226]" captionTargetId="figure@11.[151,1436,589,1238]" captionTargetPageId="11" captionText="FIGURE 6. Lebetus patzneri sp. nov., head lateral line system of the holotype ZSM 47486, female, 16.6 mm SL, from off SW Mallorca Island, 72 m depth. Terminology in text. Drawing by M. Kovačić." figureDoi="http://doi.org/10.5281/zenodo.3567348" httpUri="https://zenodo.org/record/3567348/files/figure.png" pageId="8" pageNumber="239">Fig. 6</figureCitation>
|
||
). No head canals present. Suborbital sensory papillae of the head lateral line system were identified on the
|
||
<typeStatus id="54F28807FF85FFDCA9E7F85EFE6AF878" box="[404,502,1930,1956]" pageId="8" pageNumber="239" type="holotype">holotype</typeStatus>
|
||
and confirmed in part on the
|
||
<typeStatus id="54F28807FF85FFDCAB37F85FFC38F879" box="[836,932,1931,1957]" pageId="8" pageNumber="239" type="paratype">paratype</typeStatus>
|
||
where many rows are lost due to the heavily damaged surface. The sensory papillae of the head lateral line system are large and reduced in number in all rows, many represented by a single papilla. Rows with count of sensory papillae in parentheses: (1)
|
||
<emphasis id="B93DEAB7FF85FFDCACAEF806FAD2F830" box="[1245,1358,2002,2028]" italics="true" pageId="8" pageNumber="239">preorbital</emphasis>
|
||
: snout with two rows in median preorbital series visible, row
|
||
<taxonomicName id="4C494D26FF84FFDDAB7AFF4EFCDDFF6F" box="[777,833,153,179]" pageId="9" pageNumber="240" rank="subSpecies" subSpecies="not">
|
||
<emphasis id="B93DEAB7FF84FFDDAB7AFF4EFC88FF6F" box="[777,788,154,179]" italics="true" pageId="9" pageNumber="240">r</emphasis>
|
||
not
|
||
</taxonomicName>
|
||
visible, superior row
|
||
<emphasis id="B93DEAB7FF84FFDDAC4FFF4EFBD3FF7A" box="[1084,1103,152,179]" italics="true" pageId="9" pageNumber="240">
|
||
s
|
||
<superScript id="7C3C9BEDFF84FFDDAC34FF4CFBD3FF7A" attach="left" box="[1095,1103,152,166]" fontSize="6" pageId="9" pageNumber="240">1</superScript>
|
||
</emphasis>
|
||
(1) close to posterior nostril, inferior row
|
||
<emphasis id="B93DEAB7FF84FFDDA95AFF6AFEA0FF16" box="[297,316,188,215]" italics="true" pageId="9" pageNumber="240">
|
||
s
|
||
<superScript id="7C3C9BEDFF84FFDDA947FF68FEA0FF16" attach="left" box="[308,316,188,202]" fontSize="6" pageId="9" pageNumber="240">3</superScript>
|
||
</emphasis>
|
||
(1) above upper lip visible only in
|
||
<typeStatus id="54F28807FF84FFDDAAA9FF69FCDCFF0B" box="[730,832,189,215]" pageId="9" pageNumber="240" type="holotype">holotype</typeStatus>
|
||
. Lateral series
|
||
<emphasis id="B93DEAB7FF84FFDDAB9DFF6AFC66FF0B" box="[1006,1018,190,215]" italics="true" pageId="9" pageNumber="240">c</emphasis>
|
||
: superior
|
||
<emphasis id="B93DEAB7FF84FFDDAC02FF6AFB1AFF16" box="[1137,1158,188,215]" italics="true" pageId="9" pageNumber="240">
|
||
c
|
||
<superScript id="7C3C9BEDFF84FFDDAC0DFF68FB1AFF16" attach="left" box="[1150,1158,188,202]" fontSize="6" pageId="9" pageNumber="240">2</superScript>
|
||
</emphasis>
|
||
as single papilla visible only in
|
||
<typeStatus id="54F28807FF84FFDDA883FF35FEC9FF27" box="[240,341,225,251]" pageId="9" pageNumber="240" type="holotype">holotype</typeStatus>
|
||
, superior
|
||
<emphasis id="B93DEAB7FF84FFDDA9B4FF36FE47FF32" box="[455,475,224,251]" italics="true" pageId="9" pageNumber="240">
|
||
c
|
||
<superScript id="7C3C9BEDFF84FFDDA9A0FF34FE47FF32" attach="left" box="[467,475,224,238]" fontSize="6" pageId="9" pageNumber="240">1</superScript>
|
||
</emphasis>
|
||
(2) behind and below anterior nostril, upper inferior
|
||
<emphasis id="B93DEAB7FF84FFDDAC4DFF36FBCEFEDC" box="[1086,1106,226,256]" italics="true" pageId="9" pageNumber="240">
|
||
c
|
||
<subScript id="17CD34E0FF84FFDDAC39FF26FBCEFEDC" attach="left" box="[1098,1106,242,256]" fontSize="6" pageId="9" pageNumber="240">2</subScript>
|
||
</emphasis>
|
||
not visible, lower inferior
|
||
<emphasis id="B93DEAB7FF84FFDDADFBFF36FA00FEDC" box="[1416,1436,226,256]" italics="true" pageId="9" pageNumber="240">
|
||
c
|
||
<subScript id="17CD34E0FF84FFDDADE7FF26FA00FEDC" attach="left" box="[1428,1436,242,256]" fontSize="6" pageId="9" pageNumber="240">1</subScript>
|
||
</emphasis>
|
||
(1) above upper lip. (2)
|
||
<emphasis id="B93DEAB7FF84FFDDA9D7FED1FD8BFEC3" box="[420,535,261,287]" italics="true" pageId="9" pageNumber="240">suborbital</emphasis>
|
||
: rows
|
||
<emphasis id="B93DEAB7FF84FFDDAA10FED2FDEDFEC3" box="[611,625,262,287]" italics="true" pageId="9" pageNumber="240">a</emphasis>
|
||
and
|
||
<emphasis id="B93DEAB7FF84FFDDAADBFED2FD28FEC3" box="[680,692,262,287]" italics="true" pageId="9" pageNumber="240">c</emphasis>
|
||
, including
|
||
<emphasis id="B93DEAB7FF84FFDDAB47FED2FCD2FEC3" box="[820,846,262,287]" italics="true" pageId="9" pageNumber="240">cp</emphasis>
|
||
, without transverse proliferation; row
|
||
<emphasis id="B93DEAB7FF84FFDDAD73FED2FA92FEC3" box="[1280,1294,262,287]" italics="true" pageId="9" pageNumber="240">a</emphasis>
|
||
longitudinal (3) below rear part of eye; row
|
||
<emphasis id="B93DEAB7FF84FFDDA98BFEFEFD98FE9F" box="[504,516,298,323]" italics="true" pageId="9" pageNumber="240">c</emphasis>
|
||
(4) longitudinal below frontal and middle part of eye anteriorly with two papillae superimposed each other between anterior part of eye and posterior part of upper lip, the row
|
||
<emphasis id="B93DEAB7FF84FFDDACACFE9AFB77FEBB" box="[1247,1259,334,359]" italics="true" pageId="9" pageNumber="240">c</emphasis>
|
||
visible only in
|
||
<typeStatus id="54F28807FF84FFDDA8E4FEA5FF61FE57" box="[151,253,369,395]" pageId="9" pageNumber="240" type="holotype">holotype</typeStatus>
|
||
; two
|
||
<emphasis id="B93DEAB7FF84FFDDA948FEA6FEC9FE57" box="[315,341,370,395]" italics="true" pageId="9" pageNumber="240">cp</emphasis>
|
||
larger papillae bellow
|
||
<emphasis id="B93DEAB7FF84FFDDAA2FFEA6FDF6FE57" box="[604,618,370,395]" italics="true" pageId="9" pageNumber="240">a</emphasis>
|
||
row anterior part, lower
|
||
<emphasis id="B93DEAB7FF84FFDDABFBFEA5FC30FE57" box="[904,940,369,395]" italics="true" pageId="9" pageNumber="240">cp’</emphasis>
|
||
anterior to upper
|
||
<emphasis id="B93DEAB7FF84FFDDAC0AFEA6FB0FFE57" box="[1145,1171,370,395]" italics="true" pageId="9" pageNumber="240">cp</emphasis>
|
||
. Longitudinal row
|
||
<emphasis id="B93DEAB7FF84FFDDAD1CFEA5FAE1FE57" box="[1391,1405,369,395]" italics="true" pageId="9" pageNumber="240">b</emphasis>
|
||
as two papillae below rear border of eye, only one papilla visible in
|
||
<typeStatus id="54F28807FF84FFDDAB08FE41FC43FE73" box="[891,991,405,431]" pageId="9" pageNumber="240" type="paratype">paratype</typeStatus>
|
||
. Longitudinal row
|
||
<emphasis id="B93DEAB7FF84FFDDACC5FE41FB58FE73" box="[1206,1220,405,431]" italics="true" pageId="9" pageNumber="240">d</emphasis>
|
||
with single papilla above and behind mouth corner. (3)
|
||
<emphasis id="B93DEAB7FF84FFDDAA59FE6DFCA1FE0F" box="[554,829,441,467]" italics="true" pageId="9" pageNumber="240">preoperculo-mandibular</emphasis>
|
||
: internal row
|
||
<emphasis id="B93DEAB7FF84FFDDABABFE6EFC7CFE0F" box="[984,992,442,467]" italics="true" pageId="9" pageNumber="240">i</emphasis>
|
||
divided into anterior (
|
||
<emphasis id="B93DEAB7FF84FFDDACAAFE6EFB7DFE0F" box="[1241,1249,442,467]" italics="true" pageId="9" pageNumber="240">i</emphasis>
|
||
:
|
||
<quantity id="4CB19B40FF84FFDDAC9CFE6DFA85FE0F" box="[1263,1305,441,467]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="9" pageNumber="240" unit="in" value="6.0">6 in</quantity>
|
||
<typeStatus id="54F28807FF84FFDDAD6CFE6DFA18FE0F" box="[1311,1412,441,467]" pageId="9" pageNumber="240" type="holotype">holotype</typeStatus>
|
||
,
|
||
<quantity id="4CB19B40FF84FFDDADFDFE6CFF31FE2B" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="9" pageNumber="240" unit="in" value="5.0">5 in</quantity>
|
||
<typeStatus id="54F28807FF84FFDDA8C7FE09FE85FE2B" box="[180,281,477,503]" pageId="9" pageNumber="240" type="paratype">paratype</typeStatus>
|
||
), and posterior sections, posterior section visible only in
|
||
<typeStatus id="54F28807FF84FFDDABE6FE09FC6BFE2B" box="[917,1015,477,503]" pageId="9" pageNumber="240" type="holotype">holotype</typeStatus>
|
||
(
|
||
<emphasis id="B93DEAB7FF84FFDDAC74FE0AFB93FE2B" box="[1031,1039,478,503]" italics="true" pageId="9" pageNumber="240">i</emphasis>
|
||
: 2 + 2); external row
|
||
<emphasis id="B93DEAB7FF84FFDDAD71FE0AFA92FE2B" box="[1282,1294,478,503]" italics="true" pageId="9" pageNumber="240">e</emphasis>
|
||
divided into anterior and posterior section (2 + 1), anterior visible only in
|
||
<typeStatus id="54F28807FF84FFDDAB32FDD5FC3BFDC7" box="[833,935,513,539]" pageId="9" pageNumber="240" type="holotype">holotype</typeStatus>
|
||
; mental row
|
||
<emphasis id="B93DEAB7FF84FFDDAC44FDD5FBA3FDC7" box="[1079,1087,513,539]" italics="true" pageId="9" pageNumber="240">f</emphasis>
|
||
(1) present; two larger papillae marked
|
||
<emphasis id="B93DEAB7FF84FFDDA885FDF2FE8CFDE3" box="[246,272,550,575]" italics="true" pageId="9" pageNumber="240">pc</emphasis>
|
||
present on the place of the absent PC. (4)
|
||
<emphasis id="B93DEAB7FF84FFDDAB63FDF1FC2CFDE3" box="[784,944,549,575]" italics="true" pageId="9" pageNumber="240">oculoscapular</emphasis>
|
||
: longitudinal row marked
|
||
<emphasis id="B93DEAB7FF84FFDDAC99FDF2FA8EFDE3" box="[1258,1298,550,575]" italics="true" pageId="9" pageNumber="240">aoc</emphasis>
|
||
(2) starting behind from posterior edge of orbit on the place of the lost AOC; anterior row
|
||
<date id="FFF71065FF84FFDDAC55FD9EFBA7FD8A" box="[1062,1083,584,611]" pageId="9" pageNumber="240">
|
||
<emphasis id="B93DEAB7FF84FFDDAC55FD9EFBA7FD8A" box="[1062,1083,584,611]" italics="true" pageId="9" pageNumber="240">
|
||
x
|
||
<superScript id="7C3C9BEDFF84FFDDAC40FD9CFBA7FD8A" attach="left" box="[1075,1083,584,598]" fontSize="6" pageId="9" pageNumber="240">1</superScript>
|
||
</emphasis>
|
||
</date>
|
||
as single visible papilla above preopercle visible also in
|
||
<typeStatus id="54F28807FF84FFDDA9C5FDB9FD85FD5B" box="[438,537,621,647]" pageId="9" pageNumber="240" type="paratype">paratype</typeStatus>
|
||
, all other oculoscapular rows recorded only on the
|
||
<typeStatus id="54F28807FF84FFDDAC20FDB9FB25FD5B" box="[1107,1209,621,647]" pageId="9" pageNumber="240" type="holotype">holotype</typeStatus>
|
||
: posterior row
|
||
<date id="FFF71065FF84FFDDAD13FDBAFAE9FDA6" box="[1376,1397,620,647]" pageId="9" pageNumber="240">
|
||
<emphasis id="B93DEAB7FF84FFDDAD13FDBAFAE9FDA6" box="[1376,1397,620,647]" italics="true" pageId="9" pageNumber="240">
|
||
x
|
||
<superScript id="7C3C9BEDFF84FFDDAD1EFDB8FAE9FDA6" attach="left" box="[1389,1397,620,634]" fontSize="6" pageId="9" pageNumber="240">2</superScript>
|
||
</emphasis>
|
||
</date>
|
||
(1) above posterior opercle, row
|
||
<emphasis id="B93DEAB7FF84FFDDA990FD46FE6DFD77" box="[483,497,658,683]" italics="true" pageId="9" pageNumber="240">u</emphasis>
|
||
(2) on the place of the lost POC, axillary rows
|
||
<emphasis id="B93DEAB7FF84FFDDAC62FD46FBADFD42" box="[1041,1073,656,683]" italics="true" pageId="9" pageNumber="240">
|
||
as
|
||
<superScript id="7C3C9BEDFF84FFDDAC5AFD44FBADFD42" attach="left" box="[1065,1073,656,670]" fontSize="6" pageId="9" pageNumber="240">1</superScript>
|
||
</emphasis>
|
||
(2) and
|
||
<emphasis id="B93DEAB7FF84FFDDACE3FD46FB2DFD42" box="[1168,1201,656,683]" italics="true" pageId="9" pageNumber="240">
|
||
as
|
||
<superScript id="7C3C9BEDFF84FFDDACDAFD44FB2DFD42" attach="left" box="[1193,1201,656,670]" fontSize="6" pageId="9" pageNumber="240">2</superScript>
|
||
</emphasis>
|
||
(1) present, no other oculoscapular papillae are visible; (5)
|
||
<emphasis id="B93DEAB7FF84FFDDAA3AFD61FD2AFD13" box="[585,694,693,719]" italics="true" pageId="9" pageNumber="240">opercular</emphasis>
|
||
: two larger papillae marked
|
||
<emphasis id="B93DEAB7FF84FFDDAB8DFD62FB84FD13" box="[1022,1048,694,719]" italics="true" pageId="9" pageNumber="240">pc</emphasis>
|
||
present on the place of the absent PC are visible, transverse row
|
||
<emphasis id="B93DEAB7FF84FFDDA99DFD0EFD98FD2F" box="[494,516,730,755]" italics="true" pageId="9" pageNumber="240">ot</emphasis>
|
||
(3) present with lower part; superior longitudinal row
|
||
<emphasis id="B93DEAB7FF84FFDDAC19FD0EFB1FFD2F" box="[1130,1155,730,755]" italics="true" pageId="9" pageNumber="240">os</emphasis>
|
||
and inferior longitudinal row
|
||
<emphasis id="B93DEAB7FF84FFDDA8B9FD2AFF7CFCCB" box="[202,224,766,791]" italics="true" pageId="9" pageNumber="240">oi</emphasis>
|
||
both as single papilla in
|
||
<typeStatus id="54F28807FF84FFDDA989FD29FDC0FCCB" box="[506,604,765,791]" pageId="9" pageNumber="240" type="holotype">holotype</typeStatus>
|
||
while the surface of the opercular area was destroyed on both sides in the
|
||
<typeStatus id="54F28807FF84FFDDA8E4FCF5FF66FCE7" box="[151,250,801,827]" pageId="9" pageNumber="240" type="paratype">paratype</typeStatus>
|
||
, so no papillae are visible; (6)
|
||
<emphasis id="B93DEAB7FF84FFDDAA27FCF6FD63FCE7" box="[596,767,801,827]" italics="true" pageId="9" pageNumber="240">anterior dorsal</emphasis>
|
||
: rows
|
||
<emphasis id="B93DEAB7FF84FFDDAB38FCF6FCC5FCE7" box="[843,857,802,827]" italics="true" pageId="9" pageNumber="240">n</emphasis>
|
||
(1) at posterior edge of orbit, another single visible papilla is assigned to row
|
||
<emphasis id="B93DEAB7FF84FFDDA9C5FC92FE58FC83" box="[438,452,838,863]" italics="true" pageId="9" pageNumber="240">g</emphasis>
|
||
since row
|
||
<emphasis id="B93DEAB7FF84FFDDAA48FC92FDD3FC83" box="[571,591,838,863]" italics="true" pageId="9" pageNumber="240">m</emphasis>
|
||
is usually lower and first lost in species with reduced lateral line system and row
|
||
<emphasis id="B93DEAB7FF84FFDDA8B9FCBEFF44FC5F" box="[202,216,874,899]" italics="true" pageId="9" pageNumber="240">o</emphasis>
|
||
more anteriorly placed, so no rows
|
||
<emphasis id="B93DEAB7FF84FFDDAA1FFCBEFDE6FC5F" box="[620,634,874,899]" italics="true" pageId="9" pageNumber="240">o</emphasis>
|
||
and
|
||
<emphasis id="B93DEAB7FF84FFDDAAC2FCBEFD59FC5F" box="[689,709,874,899]" italics="true" pageId="9" pageNumber="240">m</emphasis>
|
||
, row
|
||
<emphasis id="B93DEAB7FF84FFDDAB75FCBDFC88FC5F" box="[774,788,873,899]" italics="true" pageId="9" pageNumber="240">h</emphasis>
|
||
(1) in front of the first dorsal fin visible only in
|
||
<typeStatus id="54F28807FF84FFDDAD40FCBDFA05FC5F" box="[1331,1433,873,899]" pageId="9" pageNumber="240" type="holotype">holotype</typeStatus>
|
||
. (7)
|
||
<emphasis id="B93DEAB7FF84FFDDA8CCFC59FEA3FC7B" box="[191,319,909,935]" italics="true" pageId="9" pageNumber="240">interorbital</emphasis>
|
||
: interorbital papillae not visible.
|
||
</paragraph>
|
||
<caption id="DF36662DFF84FFDDA8E4F863FAF5F828" ID-DOI="http://doi.org/10.5281/zenodo.3567346" ID-Zenodo-Dep="3567346" httpUri="https://zenodo.org/record/3567346/files/figure.png" pageId="9" pageNumber="240" startId="9.[151,250,1975,2000]" targetBox="[170,1425,953,1948]" targetPageId="9">
|
||
<paragraph id="8BF636A5FF84FFDDA8E4F863FAF5F828" blockId="9.[151,1437,1975,2036]" pageId="9" pageNumber="240">
|
||
<emphasis id="B93DEAB7FF84FFDDA8E4F863FE8EF80C" bold="true" box="[151,274,1975,2000]" pageId="9" pageNumber="240">FIGURE 5.</emphasis>
|
||
<taxonomicName id="4C494D26FF84FFDDA96AF86CFE5CF80C" authority="Schliewen & Kovačić & Cerwenka & Svensen & Ordines, 2019" authorityName="Schliewen & Kovačić & Cerwenka & Svensen & Ordines" authorityYear="2019" box="[281,448,1976,2000]" class="Actinopterygii" family="Gobiidae" genus="Lebetus" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="240" phylum="Chordata" rank="species" species="patzneri" status="sp. nov.">
|
||
<emphasis id="B93DEAB7FF84FFDDA96AF86CFE5CF80C" box="[281,448,1976,2000]" italics="true" pageId="9" pageNumber="240">Lebetus patzneri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B93DEAB7FF84FFDDA9B5F86CFD8BF80C" bold="true" box="[454,535,1976,2000]" pageId="9" pageNumber="240">
|
||
<taxonomicNameLabel id="A20E57CCFF84FFDDA9B5F86CFD8BF80C" box="[454,535,1976,2000]" pageId="9" pageNumber="240" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, MicroCT reconstruction of the paratype ZSM 46865, female, 16.8 mm SL; from off SW Mallorca Island, 60m depth.
|
||
<emphasis id="B93DEAB7FF84FFDDA9CFF808FE55F828" bold="true" box="[444,457,2012,2036]" pageId="9" pageNumber="240">a</emphasis>
|
||
: left lateral view;
|
||
<emphasis id="B93DEAB7FF84FFDDAAF1F808FD0CF828" bold="true" box="[642,656,2012,2036]" pageId="9" pageNumber="240">b</emphasis>
|
||
: dorsal view;
|
||
<emphasis id="B93DEAB7FF84FFDDAB52F808FCB0F828" bold="true" box="[801,812,2012,2036]" pageId="9" pageNumber="240">c</emphasis>
|
||
: ventral view. MicroCT reconstruction by A. Cerwenka.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF36662DFF87FFDEA8E4FF4EFB74FF0A" ID-Table-UUID="DF36662DFF87FFDEA8E4FF4EFB74FF0A" httpUri="http://table.plazi.org/id/DF36662DFF87FFDEA8E4FF4EFB74FF0A" pageId="10" pageNumber="241" startId="10.[151,239,154,178]" targetBox="[166,1345,225,1833]" targetIsTable="true" targetPageId="10">
|
||
<paragraph id="8BF636A5FF87FFDEA8E4FF4EFA00FF6E" blockId="10.[151,1436,154,214]" box="[151,1436,154,178]" pageId="10" pageNumber="241">
|
||
<emphasis id="B93DEAB7FF87FFDEA8E4FF4EFE90FF6E" bold="true" box="[151,268,154,178]" pageId="10" pageNumber="241">TABLE 2.</emphasis>
|
||
Morphometric characters of
|
||
<taxonomicName id="4C494D26FF87FFDEAA3DFF4EFC9DFF6D" authority="Schliewen & Kovačić & Cerwenka & Svensen & Ordines, 2019" authorityName="Schliewen & Kovačić & Cerwenka & Svensen & Ordines" authorityYear="2019" box="[590,769,154,178]" class="Actinopterygii" family="Gobiidae" genus="Lebetus" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="241" phylum="Chordata" rank="species" species="patzneri" status="sp. nov.">
|
||
<emphasis id="B93DEAB7FF87FFDEAA3DFF4EFC9DFF6D" box="[590,769,154,178]" italics="true" pageId="10" pageNumber="241">Lebetus patzneri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B93DEAB7FF87FFDEAB79FF4EFCFEFF6E" bold="true" box="[778,866,154,178]" pageId="10" pageNumber="241">
|
||
<taxonomicNameLabel id="A20E57CCFF87FFDEAB79FF4EFCFEFF6E" box="[778,866,154,178]" pageId="10" pageNumber="241" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
holotype (ZSM 47486; female) and paratype (ZSM
|
||
</paragraph>
|
||
<paragraph id="8BF636A5FF87FFDEA8E4FF6AFB74FF0A" blockId="10.[151,1436,154,214]" box="[151,1256,190,214]" pageId="10" pageNumber="241">
|
||
<tableNote id="76AF372BFF87FFDEA8E4FF6AFB74FF0A" box="[151,1256,190,214]" pageId="10" pageNumber="241" targetBox="[166,1345,225,1833]" targetPageId="10">46865; female); absolute values and % values in brackets. An asterik (*) indicates a damaged character.</tableNote>
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF636A5FF87FFDEABE9FF35FADDF8F5" pageId="10" pageNumber="241">
|
||
<table id="F949C405FF87002BA8D5FF35FADDF8F5" box="[166,1345,225,1833]" gridcols="4" gridrows="42" pageId="10" pageNumber="241">
|
||
<tr id="357934E7FF87002BA8D5FF35FADDFF27" box="[166,1345,225,251]" gridrow="0" pageId="10" pageNumber="241" rowspan-0="1" rowspan-1="1">
|
||
<th id="76A85D9BFF87002BABE9FF35FBDFFF27" box="[922,1091,225,251]" gridcol="2" gridrow="0" pageId="10" pageNumber="241">Holotype</th>
|
||
<th id="76A85D9BFF87002BACB6FF35FADDFF27" box="[1221,1345,225,251]" gridcol="3" gridrow="0" pageId="10" pageNumber="241">Paratype</th>
|
||
</tr>
|
||
<tr id="357934E7FF87002BA8D5FED3FADDFEFC" box="[166,1345,263,288]" gridrow="1" pageId="10" pageNumber="241" rowspan-0="1">
|
||
<td id="76A85D9BFF87002BA9DBFED3FC89FEFC" box="[424,789,263,288]" gridcol="1" gridrow="1" pageId="10" pageNumber="241">Standard length (Sl) in mm</td>
|
||
<td id="76A85D9BFF87002BABE9FED3FBDFFEFC" box="[922,1091,263,288]" gridcol="2" gridrow="1" pageId="10" pageNumber="241">16.56</td>
|
||
<td id="76A85D9BFF87002BACB6FED3FADDFEFC" box="[1221,1345,263,288]" gridcol="3" gridrow="1" pageId="10" pageNumber="241">16.79</td>
|
||
</tr>
|
||
<tr id="357934E7FF87002BA8D5FEF9FADDFE9A" box="[166,1345,301,326]" gridrow="2" pageId="10" pageNumber="241">
|
||
<th id="76A85D9BFF87002BA8D5FEF9FEF6FE9A" box="[166,362,301,326]" gridcol="0" gridrow="2" pageId="10" pageNumber="241">% Standard length</th>
|
||
<td id="76A85D9BFF87002BA9DBFEF9FC89FE9A" box="[424,789,301,326]" gridcol="1" gridrow="2" pageId="10" pageNumber="241">Anal fin base</td>
|
||
<td id="76A85D9BFF87002BABE9FEF9FBDFFE9A" box="[922,1091,301,326]" gridcol="2" gridrow="2" pageId="10" pageNumber="241">2.48 (14.9)</td>
|
||
<td id="76A85D9BFF87002BACB6FEF9FADDFE9A" box="[1221,1345,301,326]" gridcol="3" gridrow="2" pageId="10" pageNumber="241">2.70 (16.1)</td>
|
||
</tr>
|
||
<tr id="357934E7FF87002BA8D5FE87FADDFEB0" box="[166,1345,339,364]" gridrow="3" pageId="10" pageNumber="241" rowspan-0="1">
|
||
<td id="76A85D9BFF87002BA9DBFE87FC89FEB0" box="[424,789,339,364]" gridcol="1" gridrow="3" pageId="10" pageNumber="241">Anal fin spine length</td>
|
||
<td id="76A85D9BFF87002BABE9FE87FBDFFEB0" box="[922,1091,339,364]" gridcol="2" gridrow="3" pageId="10" pageNumber="241">1.40 (8.4)</td>
|
||
<td id="76A85D9BFF87002BACB6FE87FADDFEB0" box="[1221,1345,339,364]" gridcol="3" gridrow="3" pageId="10" pageNumber="241">1.36 (8.1)</td>
|
||
</tr>
|
||
<tr id="357934E7FF87002BA8D5FEADFADDFE4E" box="[166,1345,377,402]" gridrow="4" pageId="10" pageNumber="241" rowspan-0="1">
|
||
<td id="76A85D9BFF87002BA9DBFEADFC89FE4E" box="[424,789,377,402]" gridcol="1" gridrow="4" pageId="10" pageNumber="241">Body depth at anal fin origin</td>
|
||
<td id="76A85D9BFF87002BABE9FEADFBDFFE4E" box="[922,1091,377,402]" gridcol="2" gridrow="4" pageId="10" pageNumber="241">2.68 (16.1)</td>
|
||
<td id="76A85D9BFF87002BACB6FEADFADDFE4E" box="[1221,1345,377,402]" gridcol="3" gridrow="4" pageId="10" pageNumber="241">2.39 (14.2)</td>
|
||
</tr>
|
||
<tr id="357934E7FF87002BA8D5FE4AFADDFE6B" box="[166,1345,414,439]" gridrow="5" pageId="10" pageNumber="241" rowspan-0="1">
|
||
<td id="76A85D9BFF87002BA9DBFE4AFC89FE6B" box="[424,789,414,439]" gridcol="1" gridrow="5" pageId="10" pageNumber="241">Body width at anal fin origin</td>
|
||
<td id="76A85D9BFF87002BABE9FE4AFBDFFE6B" box="[922,1091,414,439]" gridcol="2" gridrow="5" pageId="10" pageNumber="241">1.65 (9.9)</td>
|
||
<td id="76A85D9BFF87002BACB6FE4AFADDFE6B" box="[1221,1345,414,439]" gridcol="3" gridrow="5" pageId="10" pageNumber="241">1.30 (7.7)</td>
|
||
</tr>
|
||
<tr id="357934E7FF87002BA8D5FE10FADDFE01" box="[166,1345,452,477]" gridrow="6" pageId="10" pageNumber="241" rowspan-0="1">
|
||
<td id="76A85D9BFF87002BA9DBFE10FC89FE01" box="[424,789,452,477]" gridcol="1" gridrow="6" pageId="10" pageNumber="241">Body depth at pelvic fin origin</td>
|
||
<td id="76A85D9BFF87002BABE9FE10FBDFFE01" box="[922,1091,452,477]" gridcol="2" gridrow="6" pageId="10" pageNumber="241">2.73 (16.4)</td>
|
||
<td id="76A85D9BFF87002BACB6FE10FADDFE01" box="[1221,1345,452,477]" gridcol="3" gridrow="6" pageId="10" pageNumber="241">2.50 (14.9)</td>
|
||
</tr>
|
||
<tr id="357934E7FF87002BA8D5FE3EFADDFDDF" box="[166,1345,490,515]" gridrow="7" pageId="10" pageNumber="241" rowspan-0="1">
|
||
<td id="76A85D9BFF87002BA9DBFE3EFC89FDDF" box="[424,789,490,515]" gridcol="1" gridrow="7" pageId="10" pageNumber="241">Body width at pelvic fin origin</td>
|
||
<td id="76A85D9BFF87002BABE9FE3EFBDFFDDF" box="[922,1091,490,515]" gridcol="2" gridrow="7" pageId="10" pageNumber="241">2.91 (17.6)</td>
|
||
<td id="76A85D9BFF87002BACB6FE3EFADDFDDF" box="[1221,1345,490,515]" gridcol="3" gridrow="7" pageId="10" pageNumber="241">2.59 (15.4)</td>
|
||
</tr>
|
||
<tr id="357934E7FF87002BA8D5FDC4FADDFDF5" box="[166,1345,528,553]" gridrow="8" pageId="10" pageNumber="241" rowspan-0="1">
|
||
<td id="76A85D9BFF87002BA9DBFDC4FC89FDF5" box="[424,789,528,553]" gridcol="1" gridrow="8" pageId="10" pageNumber="241">Caudal fin length (longest ray)</td>
|
||
<td id="76A85D9BFF87002BABE9FDC4FBDFFDF5" box="[922,1091,528,553]" gridcol="2" gridrow="8" pageId="10" pageNumber="241">4.22 (25.4)</td>
|
||
<td id="76A85D9BFF87002BACB6FDC4FADDFDF5" box="[1221,1345,528,553]" gridcol="3" gridrow="8" pageId="10" pageNumber="241">*</td>
|
||
</tr>
|
||
<tr id="357934E7FF87002BA8D5FDE2FADDFD93" box="[166,1345,566,591]" gridrow="9" pageId="10" pageNumber="241" rowspan-0="1">
|
||
<td id="76A85D9BFF87002BA9DBFDE2FC89FD93" box="[424,789,566,591]" gridcol="1" gridrow="9" pageId="10" pageNumber="241">Caudal peduncle depth</td>
|
||
<td id="76A85D9BFF87002BABE9FDE2FBDFFD93" box="[922,1091,566,591]" gridcol="2" gridrow="9" pageId="10" pageNumber="241">1.61 (9.7)</td>
|
||
<td id="76A85D9BFF87002BACB6FDE2FADDFD93" box="[1221,1345,566,591]" gridcol="3" gridrow="9" pageId="10" pageNumber="241">1.38 (8.2)</td>
|
||
</tr>
|
||
<tr id="357934E7FF87002BA8D5FD8FFADDFDA8" box="[166,1345,603,628]" gridrow="10" pageId="10" pageNumber="241" rowspan-0="1">
|
||
<td id="76A85D9BFF87002BA9DBFD8FFC89FDA8" box="[424,789,603,628]" gridcol="1" gridrow="10" pageId="10" pageNumber="241">Caudal peduncle length</td>
|
||
<td id="76A85D9BFF87002BABE9FD8FFBDFFDA8" box="[922,1091,603,628]" gridcol="2" gridrow="10" pageId="10" pageNumber="241">3.79 (22.8)</td>
|
||
<td id="76A85D9BFF87002BACB6FD8FFADDFDA8" box="[1221,1345,603,628]" gridcol="3" gridrow="10" pageId="10" pageNumber="241">3.91 (23.3)</td>
|
||
</tr>
|
||
<tr id="357934E7FF87002BA8D5FD55FADDFD46" box="[166,1345,641,666]" gridrow="11" pageId="10" pageNumber="241" rowspan-0="1">
|
||
<td id="76A85D9BFF87002BA9DBFD55FC89FD46" box="[424,789,641,666]" gridcol="1" gridrow="11" pageId="10" pageNumber="241">Eye diameter (horizontal)</td>
|
||
<td id="76A85D9BFF87002BABE9FD55FBDFFD46" box="[922,1091,641,666]" gridcol="2" gridrow="11" pageId="10" pageNumber="241">1.36 (8.2)</td>
|
||
<td id="76A85D9BFF87002BACB6FD55FADDFD46" box="[1221,1345,641,666]" gridcol="3" gridrow="11" pageId="10" pageNumber="241">1.27 (7.6)</td>
|
||
</tr>
|
||
<tr id="357934E7FF87002BA8D5FD73FADDFD1C" box="[166,1345,679,704]" gridrow="12" pageId="10" pageNumber="241" rowspan-0="1">
|
||
<td id="76A85D9BFF87002BA9DBFD73FC89FD1C" box="[424,789,679,704]" gridcol="1" gridrow="12" pageId="10" pageNumber="241">Eye diameter (vertical)</td>
|
||
<td id="76A85D9BFF87002BABE9FD73FBDFFD1C" box="[922,1091,679,704]" gridcol="2" gridrow="12" pageId="10" pageNumber="241">1.06 (6.4)</td>
|
||
<td id="76A85D9BFF87002BACB6FD73FADDFD1C" box="[1221,1345,679,704]" gridcol="3" gridrow="12" pageId="10" pageNumber="241">0.88 (5.2)</td>
|
||
</tr>
|
||
<tr id="357934E7FF87002BA8D5FD19FADDFD3A" box="[166,1345,717,742]" gridrow="13" pageId="10" pageNumber="241" rowspan-0="1">
|
||
<td id="76A85D9BFF87002BA9DBFD19FC89FD3A" box="[424,789,717,742]" gridcol="1" gridrow="13" pageId="10" pageNumber="241">First dorsal fin base</td>
|
||
<td id="76A85D9BFF87002BABE9FD19FBDFFD3A" box="[922,1091,717,742]" gridcol="2" gridrow="13" pageId="10" pageNumber="241">1.72 (10.4)</td>
|
||
<td id="76A85D9BFF87002BACB6FD19FADDFD3A" box="[1221,1345,717,742]" gridcol="3" gridrow="13" pageId="10" pageNumber="241">1.90 (11.3)</td>
|
||
</tr>
|
||
<tr id="357934E7FF87002BA8D5FD27FADDFCD0" box="[166,1345,755,780]" gridrow="14" pageId="10" pageNumber="241" rowspan-0="1">
|
||
<td id="76A85D9BFF87002BA9DBFD27FC89FCD0" box="[424,789,755,780]" gridcol="1" gridrow="14" pageId="10" pageNumber="241">Head length</td>
|
||
<td id="76A85D9BFF87002BABE9FD27FBDFFCD0" box="[922,1091,755,780]" gridcol="2" gridrow="14" pageId="10" pageNumber="241">5.34 (32.2)</td>
|
||
<td id="76A85D9BFF87002BACB6FD27FADDFCD0" box="[1221,1345,755,780]" gridcol="3" gridrow="14" pageId="10" pageNumber="241">5.45 (32.5)</td>
|
||
</tr>
|
||
<tr id="357934E7FF87002BA8D5FCCCFADDFCED" box="[166,1345,792,817]" gridrow="15" pageId="10" pageNumber="241" rowspan-0="1">
|
||
<td id="76A85D9BFF87002BA9DBFCCCFC89FCED" box="[424,789,792,817]" gridcol="1" gridrow="15" pageId="10" pageNumber="241">Head width</td>
|
||
<td id="76A85D9BFF87002BABE9FCCCFBDFFCED" box="[922,1091,792,817]" gridcol="2" gridrow="15" pageId="10" pageNumber="241">3.22 (19.4)</td>
|
||
<td id="76A85D9BFF87002BACB6FCCCFADDFCED" box="[1221,1345,792,817]" gridcol="3" gridrow="15" pageId="10" pageNumber="241">3.30 (19.7)</td>
|
||
</tr>
|
||
<tr id="357934E7FF87002BA8D5FCEAFADDFC8B" box="[166,1345,830,855]" gridrow="16" pageId="10" pageNumber="241" rowspan-0="1">
|
||
<td id="76A85D9BFF87002BA9DBFCEAFC89FC8B" box="[424,789,830,855]" gridcol="1" gridrow="16" pageId="10" pageNumber="241">Pectoral fin length</td>
|
||
<td id="76A85D9BFF87002BABE9FCEAFBDFFC8B" box="[922,1091,830,855]" gridcol="2" gridrow="16" pageId="10" pageNumber="241">4.87 (29.3)</td>
|
||
<td id="76A85D9BFF87002BACB6FCEAFADDFC8B" box="[1221,1345,830,855]" gridcol="3" gridrow="16" pageId="10" pageNumber="241">4.50 (26.8)</td>
|
||
</tr>
|
||
<tr id="357934E7FF87002BA8D5FCB0FADDFCA1" box="[166,1345,868,893]" gridrow="17" pageId="10" pageNumber="241" rowspan-0="1">
|
||
<td id="76A85D9BFF87002BA9DBFCB0FC89FCA1" box="[424,789,868,893]" gridcol="1" gridrow="17" pageId="10" pageNumber="241">Pelvic fin length</td>
|
||
<td id="76A85D9BFF87002BABE9FCB0FBDFFCA1" box="[922,1091,868,893]" gridcol="2" gridrow="17" pageId="10" pageNumber="241">3.50 (21.1)</td>
|
||
<td id="76A85D9BFF87002BACB6FCB0FADDFCA1" box="[1221,1345,868,893]" gridcol="3" gridrow="17" pageId="10" pageNumber="241">3.91 (23.3)</td>
|
||
</tr>
|
||
<tr id="357934E7FF87002BA8D5FC5EFADDFC7F" box="[166,1345,906,931]" gridrow="18" pageId="10" pageNumber="241" rowspan-0="1">
|
||
<td id="76A85D9BFF87002BA9DBFC5EFC89FC7F" box="[424,789,906,931]" gridcol="1" gridrow="18" pageId="10" pageNumber="241">Pelvic to anus</td>
|
||
<td id="76A85D9BFF87002BABE9FC5EFBDFFC7F" box="[922,1091,906,931]" gridcol="2" gridrow="18" pageId="10" pageNumber="241">4.45 (26.8)</td>
|
||
<td id="76A85D9BFF87002BACB6FC5EFADDFC7F" box="[1221,1345,906,931]" gridcol="3" gridrow="18" pageId="10" pageNumber="241">4.80 (28.6)</td>
|
||
</tr>
|
||
<tr id="357934E7FF87002BA8D5FC7BFADDFC14" box="[166,1345,943,968]" gridrow="19" pageId="10" pageNumber="241" rowspan-0="1">
|
||
<td id="76A85D9BFF87002BA9DBFC7BFC89FC14" box="[424,789,943,968]" gridcol="1" gridrow="19" pageId="10" pageNumber="241">Second dorsal fin base</td>
|
||
<td id="76A85D9BFF87002BABE9FC7BFBDFFC14" box="[922,1091,943,968]" gridcol="2" gridrow="19" pageId="10" pageNumber="241">3.38 (20.4)</td>
|
||
<td id="76A85D9BFF87002BACB6FC7BFADDFC14" box="[1221,1345,943,968]" gridcol="3" gridrow="19" pageId="10" pageNumber="241">3.28 (19.5)</td>
|
||
</tr>
|
||
<tr id="357934E7FF87002BA8D5FC01FADDFC32" box="[166,1345,981,1006]" gridrow="20" pageId="10" pageNumber="241" rowspan-0="1">
|
||
<td id="76A85D9BFF87002BA9DBFC01FC89FC32" box="[424,789,981,1006]" gridcol="1" gridrow="20" pageId="10" pageNumber="241">Second dorsal fin spine length</td>
|
||
<td id="76A85D9BFF87002BABE9FC01FBDFFC32" box="[922,1091,981,1006]" gridcol="2" gridrow="20" pageId="10" pageNumber="241">1.32 (8.0)</td>
|
||
<td id="76A85D9BFF87002BACB6FC01FADDFC32" box="[1221,1345,981,1006]" gridcol="3" gridrow="20" pageId="10" pageNumber="241">*</td>
|
||
</tr>
|
||
<tr id="357934E7FF87002BA8D5FC2FFADDFBC8" box="[166,1345,1019,1044]" gridrow="21" pageId="10" pageNumber="241" rowspan-0="1">
|
||
<td id="76A85D9BFF87002BA9DBFC2FFC89FBC8" box="[424,789,1019,1044]" gridcol="1" gridrow="21" pageId="10" pageNumber="241">Snout to origin of first dorsal fin</td>
|
||
<td id="76A85D9BFF87002BABE9FC2FFBDFFBC8" box="[922,1091,1019,1044]" gridcol="2" gridrow="21" pageId="10" pageNumber="241">6.53 (39.4)</td>
|
||
<td id="76A85D9BFF87002BACB6FC2FFADDFBC8" box="[1221,1345,1019,1044]" gridcol="3" gridrow="21" pageId="10" pageNumber="241">6.23 (37.1)</td>
|
||
</tr>
|
||
<tr id="357934E7FF87002BA8D5FBF5FADDFBE6" box="[166,1345,1057,1082]" gridrow="22" pageId="10" pageNumber="241" rowspan-0="1">
|
||
<td id="76A85D9BFF87002BA9DBFBF5FC89FBE6" box="[424,789,1057,1082]" gridcol="1" gridrow="22" pageId="10" pageNumber="241">Snout to origin of second dorsal fin</td>
|
||
<td id="76A85D9BFF87002BABE9FBF5FBDFFBE6" box="[922,1091,1057,1082]" gridcol="2" gridrow="22" pageId="10" pageNumber="241">9.59 (57.9)</td>
|
||
<td id="76A85D9BFF87002BACB6FBF5FADDFBE6" box="[1221,1345,1057,1082]" gridcol="3" gridrow="22" pageId="10" pageNumber="241">9.69 (57.7)</td>
|
||
</tr>
|
||
<tr id="357934E7FF87002BA8D5FB93FADDFBBC" box="[166,1345,1095,1120]" gridrow="23" pageId="10" pageNumber="241" rowspan-0="1">
|
||
<td id="76A85D9BFF87002BA9DBFB93FC89FBBC" box="[424,789,1095,1120]" gridcol="1" gridrow="23" pageId="10" pageNumber="241">Snout to vertical of anal fin origin</td>
|
||
<td id="76A85D9BFF87002BABE9FB93FBDFFBBC" box="[922,1091,1095,1120]" gridcol="2" gridrow="23" pageId="10" pageNumber="241">10.4 (62.8)</td>
|
||
<td id="76A85D9BFF87002BACB6FB93FADDFBBC" box="[1221,1345,1095,1120]" gridcol="3" gridrow="23" pageId="10" pageNumber="241">10.32 (61.5)</td>
|
||
</tr>
|
||
<tr id="357934E7FF87002BA8D5FBB8FADDFB59" box="[166,1345,1132,1157]" gridrow="24" pageId="10" pageNumber="241" rowspan-0="1">
|
||
<td id="76A85D9BFF87002BA9DBFBB8FC89FB59" box="[424,789,1132,1157]" gridcol="1" gridrow="24" pageId="10" pageNumber="241">Snout to vertical of pelvic fin origin</td>
|
||
<td id="76A85D9BFF87002BABE9FBB8FBDFFB59" box="[922,1091,1132,1157]" gridcol="2" gridrow="24" pageId="10" pageNumber="241">5.75 (34.7)</td>
|
||
<td id="76A85D9BFF87002BACB6FBB8FADDFB59" box="[1221,1345,1132,1157]" gridcol="3" gridrow="24" pageId="10" pageNumber="241">5.80 (34.5)</td>
|
||
</tr>
|
||
<tr id="357934E7FF87002BA8D5FB46FADDFB77" box="[166,1345,1170,1195]" gridrow="25" pageId="10" pageNumber="241" rowspan-0="1">
|
||
<td id="76A85D9BFF87002BA9DBFB46FC89FB77" box="[424,789,1170,1195]" gridcol="1" gridrow="25" pageId="10" pageNumber="241">1st spine length of 1st dorsal fin</td>
|
||
<td id="76A85D9BFF87002BABE9FB46FBDFFB77" box="[922,1091,1170,1195]" gridcol="2" gridrow="25" pageId="10" pageNumber="241">2.41 (14.5)</td>
|
||
<td id="76A85D9BFF87002BACB6FB46FADDFB77" box="[1221,1345,1170,1195]" gridcol="3" gridrow="25" pageId="10" pageNumber="241">*</td>
|
||
</tr>
|
||
<tr id="357934E7FF87002BA8D5FB6CFADDFB0D" box="[166,1345,1208,1233]" gridrow="26" pageId="10" pageNumber="241" rowspan-0="1">
|
||
<td id="76A85D9BFF87002BA9DBFB6CFC89FB0D" box="[424,789,1208,1233]" gridcol="1" gridrow="26" pageId="10" pageNumber="241">2nd spine length of 1st dorsal fin</td>
|
||
<td id="76A85D9BFF87002BABE9FB6CFBDFFB0D" box="[922,1091,1208,1233]" gridcol="2" gridrow="26" pageId="10" pageNumber="241">2.93 (17.7)</td>
|
||
<td id="76A85D9BFF87002BACB6FB6CFADDFB0D" box="[1221,1345,1208,1233]" gridcol="3" gridrow="26" pageId="10" pageNumber="241">*</td>
|
||
</tr>
|
||
<tr id="357934E7FF87002BA8D5FB0AFADDFB2B" box="[166,1345,1246,1271]" gridrow="27" pageId="10" pageNumber="241" rowspan-0="1">
|
||
<td id="76A85D9BFF87002BA9DBFB0AFC89FB2B" box="[424,789,1246,1271]" gridcol="1" gridrow="27" pageId="10" pageNumber="241">3rd spine length of 1st dorsal fin</td>
|
||
<td id="76A85D9BFF87002BABE9FB0AFBDFFB2B" box="[922,1091,1246,1271]" gridcol="2" gridrow="27" pageId="10" pageNumber="241">2.64 15.9)</td>
|
||
<td id="76A85D9BFF87002BACB6FB0AFADDFB2B" box="[1221,1345,1246,1271]" gridcol="3" gridrow="27" pageId="10" pageNumber="241">*</td>
|
||
</tr>
|
||
<tr id="357934E7FF87002BA8D5FAD0FADDFAC1" box="[166,1345,1284,1309]" gridrow="28" pageId="10" pageNumber="241" rowspan-0="1">
|
||
<td id="76A85D9BFF87002BA9DBFAD0FC89FAC1" box="[424,789,1284,1309]" gridcol="1" gridrow="28" pageId="10" pageNumber="241">4th spine length of first dorsal fin</td>
|
||
<td id="76A85D9BFF87002BABE9FAD0FBDFFAC1" box="[922,1091,1284,1309]" gridcol="2" gridrow="28" pageId="10" pageNumber="241">2.17 (13.1)</td>
|
||
<td id="76A85D9BFF87002BACB6FAD0FADDFAC1" box="[1221,1345,1284,1309]" gridcol="3" gridrow="28" pageId="10" pageNumber="241">*</td>
|
||
</tr>
|
||
<tr id="357934E7FF87002BA8D5FAFDFADDFA9E" box="[166,1345,1321,1346]" gridrow="29" pageId="10" pageNumber="241" rowspan-0="1">
|
||
<td id="76A85D9BFF87002BA9DBFAFDFC89FA9E" box="[424,789,1321,1346]" gridcol="1" gridrow="29" pageId="10" pageNumber="241">5th spine length of first dorsal fin</td>
|
||
<td id="76A85D9BFF87002BABE9FAFDFBDFFA9E" box="[922,1091,1321,1346]" gridcol="2" gridrow="29" pageId="10" pageNumber="241">2.08 12.5)</td>
|
||
<td id="76A85D9BFF87002BACB6FAFDFADDFA9E" box="[1221,1345,1321,1346]" gridcol="3" gridrow="29" pageId="10" pageNumber="241">*</td>
|
||
</tr>
|
||
<tr id="357934E7FF87002BA8D5FA9BFADDFAB4" box="[166,1345,1359,1384]" gridrow="30" pageId="10" pageNumber="241" rowspan-0="1">
|
||
<td id="76A85D9BFF87002BA9DBFA9BFC89FAB4" box="[424,789,1359,1384]" gridcol="1" gridrow="30" pageId="10" pageNumber="241">6th spine length of first dorsal fin</td>
|
||
<td id="76A85D9BFF87002BABE9FA9BFBDFFAB4" box="[922,1091,1359,1384]" gridcol="2" gridrow="30" pageId="10" pageNumber="241">0.92 (5.5)</td>
|
||
<td id="76A85D9BFF87002BACB6FA9BFADDFAB4" box="[1221,1345,1359,1384]" gridcol="3" gridrow="30" pageId="10" pageNumber="241">*</td>
|
||
</tr>
|
||
<tr id="357934E7FF87002BA8D5FAA1FADDFA52" box="[166,1345,1397,1422]" gridrow="31" pageId="10" pageNumber="241" rowspan-0="1">
|
||
<td id="76A85D9BFF87002BA9DBFAA1FC89FA52" box="[424,789,1397,1422]" gridcol="1" gridrow="31" pageId="10" pageNumber="241">1st ray length of second dorsal fin</td>
|
||
<td id="76A85D9BFF87002BABE9FAA1FBDFFA52" box="[922,1091,1397,1422]" gridcol="2" gridrow="31" pageId="10" pageNumber="241">1.88 (11.3)</td>
|
||
<td id="76A85D9BFF87002BACB6FAA1FADDFA52" box="[1221,1345,1397,1422]" gridcol="3" gridrow="31" pageId="10" pageNumber="241">1.83 (10.9)</td>
|
||
</tr>
|
||
<tr id="357934E7FF87002BA8D5FA4FFADDFA68" box="[166,1345,1435,1460]" gridrow="32" pageId="10" pageNumber="241">
|
||
<th id="76A85D9BFF87002BA8D5FA4FFEF6FA68" box="[166,362,1435,1460]" gridcol="0" gridrow="32" pageId="10" pageNumber="241">% Caudal peduncle</th>
|
||
<td id="76A85D9BFF87002BA9DBFA4FFC89FA68" box="[424,789,1435,1460]" gridcol="1" gridrow="32" pageId="10" pageNumber="241">Caudal Peduncle depth</td>
|
||
<td id="76A85D9BFF87002BABE9FA4FFBDFFA68" box="[922,1091,1435,1460]" gridcol="2" gridrow="32" pageId="10" pageNumber="241">1.61 (42.5)</td>
|
||
<td id="76A85D9BFF87002BACB6FA4FFADDFA68" box="[1221,1345,1435,1460]" gridcol="3" gridrow="32" pageId="10" pageNumber="241">1.38 (35.3)</td>
|
||
</tr>
|
||
<tr id="357934E7FF87002BA8D5FA6BFADDFA25" box="[166,1345,1471,1529]" gridrow="33" pageId="10" pageNumber="241">
|
||
<th id="76A85D9BFF87002BA8D5FA6BFEF6FA25" box="[166,362,1471,1529]" gridcol="0" gridrow="33" pageId="10" pageNumber="241">length % Head length</th>
|
||
<td id="76A85D9BFF87002BA9DBFA6BFC89FA25" box="[424,789,1471,1529]" gridcol="1" gridrow="33" pageId="10" pageNumber="241">Cheek depth</td>
|
||
<td id="76A85D9BFF87002BABE9FA6BFBDFFA25" box="[922,1091,1471,1529]" gridcol="2" gridrow="33" pageId="10" pageNumber="241">0.97 (18.2)</td>
|
||
<td id="76A85D9BFF87002BACB6FA6BFADDFA25" box="[1221,1345,1471,1529]" gridcol="3" gridrow="33" pageId="10" pageNumber="241">1.06 (19.5)</td>
|
||
</tr>
|
||
<tr id="357934E7FF87002BA8D5F9D1FADDF9C2" box="[166,1345,1541,1566]" gridrow="34" pageId="10" pageNumber="241" rowspan-0="1">
|
||
<td id="76A85D9BFF87002BA9DBF9D1FC89F9C2" box="[424,789,1541,1566]" gridcol="1" gridrow="34" pageId="10" pageNumber="241">Eye diameter</td>
|
||
<td id="76A85D9BFF87002BABE9F9D1FBDFF9C2" box="[922,1091,1541,1566]" gridcol="2" gridrow="34" pageId="10" pageNumber="241">1.36 (25.5)</td>
|
||
<td id="76A85D9BFF87002BACB6F9D1FADDF9C2" box="[1221,1345,1541,1566]" gridcol="3" gridrow="34" pageId="10" pageNumber="241">1.27 (23.3)</td>
|
||
</tr>
|
||
<tr id="357934E7FF87002BA8D5F9FFFADDF998" box="[166,1345,1579,1604]" gridrow="35" pageId="10" pageNumber="241" rowspan-0="1">
|
||
<td id="76A85D9BFF87002BA9DBF9FFFC89F998" box="[424,789,1579,1604]" gridcol="1" gridrow="35" pageId="10" pageNumber="241">Head depth</td>
|
||
<td id="76A85D9BFF87002BABE9F9FFFBDFF998" box="[922,1091,1579,1604]" gridcol="2" gridrow="35" pageId="10" pageNumber="241">2.55 (47.8)</td>
|
||
<td id="76A85D9BFF87002BACB6F9FFFADDF998" box="[1221,1345,1579,1604]" gridcol="3" gridrow="35" pageId="10" pageNumber="241">2.10 (38.5)</td>
|
||
</tr>
|
||
<tr id="357934E7FF87002BA8D5F985FADDF9B6" box="[166,1345,1617,1642]" gridrow="36" pageId="10" pageNumber="241" rowspan-0="1">
|
||
<td id="76A85D9BFF87002BA9DBF985FC89F9B6" box="[424,789,1617,1642]" gridcol="1" gridrow="36" pageId="10" pageNumber="241">Head width</td>
|
||
<td id="76A85D9BFF87002BABE9F985FBDFF9B6" box="[922,1091,1617,1642]" gridcol="2" gridrow="36" pageId="10" pageNumber="241">(expanded head)</td>
|
||
<td id="76A85D9BFF87002BACB6F985FADDF9B6" box="[1221,1345,1617,1642]" gridcol="3" gridrow="36" pageId="10" pageNumber="241">3.62 (66.4)</td>
|
||
</tr>
|
||
<tr id="357934E7FF87002BA8D5F9A3FADDF94C" box="[166,1345,1655,1680]" gridrow="37" pageId="10" pageNumber="241" rowspan-0="1">
|
||
<td id="76A85D9BFF87002BA9DBF9A3FC89F94C" box="[424,789,1655,1680]" gridcol="1" gridrow="37" pageId="10" pageNumber="241">Postorbital length</td>
|
||
<td id="76A85D9BFF87002BABE9F9A3FBDFF94C" box="[922,1091,1655,1680]" gridcol="2" gridrow="37" pageId="10" pageNumber="241">2.39 (44.8)</td>
|
||
<td id="76A85D9BFF87002BACB6F9A3FADDF94C" box="[1221,1345,1655,1680]" gridcol="3" gridrow="37" pageId="10" pageNumber="241">2.50 (45.9)</td>
|
||
</tr>
|
||
<tr id="357934E7FF87002BA8D5F948FADDF969" box="[166,1345,1692,1717]" gridrow="38" pageId="10" pageNumber="241" rowspan-0="1">
|
||
<td id="76A85D9BFF87002BA9DBF948FC89F969" box="[424,789,1692,1717]" gridcol="1" gridrow="38" pageId="10" pageNumber="241">Snout length</td>
|
||
<td id="76A85D9BFF87002BABE9F948FBDFF969" box="[922,1091,1692,1717]" gridcol="2" gridrow="38" pageId="10" pageNumber="241">1.43 (26.8)</td>
|
||
<td id="76A85D9BFF87002BACB6F948FADDF969" box="[1221,1345,1692,1717]" gridcol="3" gridrow="38" pageId="10" pageNumber="241">1.35 (24.8)</td>
|
||
</tr>
|
||
<tr id="357934E7FF87002BA8D5F916FADDF907" box="[166,1345,1730,1755]" gridrow="39" pageId="10" pageNumber="241">
|
||
<th id="76A85D9BFF87002BA8D5F916FEF6F907" box="[166,362,1730,1755]" gridcol="0" gridrow="39" pageId="10" pageNumber="241">% Eye length</th>
|
||
<td id="76A85D9BFF87002BA9DBF916FC89F907" box="[424,789,1730,1755]" gridcol="1" gridrow="39" pageId="10" pageNumber="241">Interorbital width</td>
|
||
<td id="76A85D9BFF87002BABE9F916FBDFF907" box="[922,1091,1730,1755]" gridcol="2" gridrow="39" pageId="10" pageNumber="241">0.24 (17.7)</td>
|
||
<td id="76A85D9BFF87002BACB6F916FADDF907" box="[1221,1345,1730,1755]" gridcol="3" gridrow="39" pageId="10" pageNumber="241">0.33 (26.0)</td>
|
||
</tr>
|
||
<tr id="357934E7FF87002BA8D5F93EFADDF8DF" box="[166,1345,1770,1795]" gridrow="40" pageId="10" pageNumber="241" rowspan-0="1">
|
||
<td id="76A85D9BFF87002BA9DBF93EFC89F8DF" box="[424,789,1770,1795]" gridcol="1" gridrow="40" pageId="10" pageNumber="241">Snout length</td>
|
||
<td id="76A85D9BFF87002BABE9F93EFBDFF8DF" box="[922,1091,1770,1795]" gridcol="2" gridrow="40" pageId="10" pageNumber="241">1.43 (105.2)</td>
|
||
<td id="76A85D9BFF87002BACB6F93EFADDF8DF" box="[1221,1345,1770,1795]" gridcol="3" gridrow="40" pageId="10" pageNumber="241">1.35 (106.3)</td>
|
||
</tr>
|
||
<tr id="357934E7FF87002BA8D5F8C4FADDF8F5" box="[166,1345,1808,1833]" gridrow="41" pageId="10" pageNumber="241">
|
||
<th id="76A85D9BFF87002BA8D5F8C4FEF6F8F5" box="[166,362,1808,1833]" gridcol="0" gridrow="41" pageId="10" pageNumber="241">% Pelvic to anus</th>
|
||
<td id="76A85D9BFF87002BA9DBF8C4FC89F8F5" box="[424,789,1808,1833]" gridcol="1" gridrow="41" pageId="10" pageNumber="241">Pelvic fin length</td>
|
||
<td id="76A85D9BFF87002BABE9F8C4FBDFF8F5" box="[922,1091,1808,1833]" gridcol="2" gridrow="41" pageId="10" pageNumber="241">3.5 (78.7)</td>
|
||
<td id="76A85D9BFF87002BACB6F8C4FADDF8F5" box="[1221,1345,1808,1833]" gridcol="3" gridrow="41" pageId="10" pageNumber="241">5.91 (122.6)</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="8BF636A5FF87FFDFA8B4F88DFCB2FDE3" blockId="10.[151,1437,1880,2016]" lastBlockId="11.[151,1437,153,575]" lastPageId="11" lastPageNumber="242" pageId="10" pageNumber="241">
|
||
<emphasis id="B93DEAB7FF87FFDEA8B4F88DFEBDF8AF" box="[199,289,1881,1907]" italics="true" pageId="10" pageNumber="241">Otoliths</emphasis>
|
||
(
|
||
<figureCitation id="13722A20FF87FFDEA941F88DFEE4F8A8" box="[306,376,1881,1908]" captionStart="FIGURE 7" captionStartId="12.[151,250,922,947]" captionTargetBox="[151,1435,337,902]" captionTargetId="figure@12.[151,1435,337,902]" captionTargetPageId="12" captionText="FIGURE 7. Lebetus patzneri sp. nov., right saccular otolith of the paratype ZSM 46865, female, 16.8 mm SL; from off SW Mallorca, 60m depth. a: in medial/inner view; b: left otolith in anterior view. Orientation of otolith with respect to fish symmetry is indicated as follows: a, anterior; d, dorsal; l, lateral; m, medial; p, posterior; v, ventral. MicroCT reconstruction by A. Cerwenka." figureDoi="http://doi.org/10.5281/zenodo.3567350" httpUri="https://zenodo.org/record/3567350/files/figure.png" pageId="10" pageNumber="241">Fig. 7</figureCitation>
|
||
). The Micro-CT of the
|
||
<typeStatus id="54F28807FF87FFDEAAF4F88EFD7BF8A8" box="[647,743,1882,1908]" pageId="10" pageNumber="241" type="paratype">paratype</typeStatus>
|
||
ZSM 46865 shows a right otolith in medial/inner view (
|
||
<figureCitation id="13722A20FF87FFDEAD1CF88DFF2EF844" captionStart="FIGURE 7" captionStartId="12.[151,250,922,947]" captionTargetBox="[151,1435,337,902]" captionTargetId="figure@12.[151,1435,337,902]" captionTargetPageId="12" captionText="FIGURE 7. Lebetus patzneri sp. nov., right saccular otolith of the paratype ZSM 46865, female, 16.8 mm SL; from off SW Mallorca, 60m depth. a: in medial/inner view; b: left otolith in anterior view. Orientation of otolith with respect to fish symmetry is indicated as follows: a, anterior; d, dorsal; l, lateral; m, medial; p, posterior; v, ventral. MicroCT reconstruction by A. Cerwenka." figureDoi="http://doi.org/10.5281/zenodo.3567350" httpUri="https://zenodo.org/record/3567350/files/figure.png" pageId="10" pageNumber="241">Fig. 7a</figureCitation>
|
||
) and a left otolith seen from the anterior margin (
|
||
<figureCitation id="13722A20FF87FFDEAAA8F8A9FCB3F844" box="[731,815,1917,1944]" captionStart="FIGURE 7" captionStartId="12.[151,250,922,947]" captionTargetBox="[151,1435,337,902]" captionTargetId="figure@12.[151,1435,337,902]" captionTargetPageId="12" captionText="FIGURE 7. Lebetus patzneri sp. nov., right saccular otolith of the paratype ZSM 46865, female, 16.8 mm SL; from off SW Mallorca, 60m depth. a: in medial/inner view; b: left otolith in anterior view. Orientation of otolith with respect to fish symmetry is indicated as follows: a, anterior; d, dorsal; l, lateral; m, medial; p, posterior; v, ventral. MicroCT reconstruction by A. Cerwenka." figureDoi="http://doi.org/10.5281/zenodo.3567350" httpUri="https://zenodo.org/record/3567350/files/figure.png" pageId="10" pageNumber="241">Fig. 7b</figureCitation>
|
||
). A small utricular otolith is visible in the ventral sector of each saccular otolith. The otolith shown in
|
||
<figureCitation id="13722A20FF87FFDEAAE5F875FD7FF860" box="[662,739,1953,1980]" captionStart="FIGURE 7" captionStartId="12.[151,250,922,947]" captionTargetBox="[151,1435,337,902]" captionTargetId="figure@12.[151,1435,337,902]" captionTargetPageId="12" captionText="FIGURE 7. Lebetus patzneri sp. nov., right saccular otolith of the paratype ZSM 46865, female, 16.8 mm SL; from off SW Mallorca, 60m depth. a: in medial/inner view; b: left otolith in anterior view. Orientation of otolith with respect to fish symmetry is indicated as follows: a, anterior; d, dorsal; l, lateral; m, medial; p, posterior; v, ventral. MicroCT reconstruction by A. Cerwenka." figureDoi="http://doi.org/10.5281/zenodo.3567350" httpUri="https://zenodo.org/record/3567350/files/figure.png" pageId="10" pageNumber="241">Fig. 7a</figureCitation>
|
||
displays smooth margins and an almost quadratic outline. The dorsal margin ascends slightly toward the posterior margin, at the highest tip it declines and forms a slightly bulged posterodorsal projection. The posterior margin is slightly concave in the middle, its transition to the ventral margin is skewed. The ventral margin is straight, only the passage to the anterior margin is slightly curved. The anterior margin is straight and displays a small predorsal angle. A clear ventral line extends with some distance from the ventral rim and extends up to the tip and end of the sulcus, respectively. The sulcus is relatively small, its length is 30% of otolith length, its height equals 10% of the otolith length. The orientation of the sulcus is horizontal. It appears that the cauda is wider than the ostium, but this would have to be checked with an SEM image. The otolith shown in
|
||
<figureCitation id="13722A20FF86FFDFA976FEA5FEC9FE57" box="[261,341,369,395]" captionStart="FIGURE 7" captionStartId="12.[151,250,922,947]" captionTargetBox="[151,1435,337,902]" captionTargetId="figure@12.[151,1435,337,902]" captionTargetPageId="12" captionText="FIGURE 7. Lebetus patzneri sp. nov., right saccular otolith of the paratype ZSM 46865, female, 16.8 mm SL; from off SW Mallorca, 60m depth. a: in medial/inner view; b: left otolith in anterior view. Orientation of otolith with respect to fish symmetry is indicated as follows: a, anterior; d, dorsal; l, lateral; m, medial; p, posterior; v, ventral. MicroCT reconstruction by A. Cerwenka." figureDoi="http://doi.org/10.5281/zenodo.3567350" httpUri="https://zenodo.org/record/3567350/files/figure.png" pageId="11" pageNumber="242">Fig. 7b</figureCitation>
|
||
reveals a bent, thickened lateral face and a relatively plane medial face; the total thickness of the otolith is approx. 20% of otolith length. The general outline of this otolith conforms well to that of the otoliths of
|
||
<taxonomicName id="4C494D26FF86FFDFA8E4FE6DFEB5FE0F" box="[151,297,441,467]" class="Actinopterygii" family="Oxudercidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="242" phylum="Chordata" rank="family">Oxudercidae</taxonomicName>
|
||
, and in particular to that of the otoliths of species of the
|
||
<taxonomicName id="4C494D26FF86FFDFABE9FE6DFBD6FE0F" box="[922,1098,441,467]" class="Actinopterygii" family="Gobiidae" genus="Pomatoschistus" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="242" phylum="Chordata" rank="genus">
|
||
<emphasis id="B93DEAB7FF86FFDFABE9FE6DFBD6FE0F" box="[922,1098,441,467]" italics="true" pageId="11" pageNumber="242">Pomatoschistus</emphasis>
|
||
</taxonomicName>
|
||
-lineage (see
|
||
<bibRefCitation id="EFD84B54FF86FFDFACADFE6DFA04FE0F" author="Gierl, C. & Liebl, D. & Sanda, R. & Vukic, J. & Esmaeili, H. R. & Reichenbacher, B." box="[1246,1432,441,467]" pageId="11" pageNumber="242" pagination="349 - 363" refId="ref16348" refString="Gierl, C., Liebl, D., Sanda, R., Vukic, J., Esmaeili, H. R. & Reichenbacher, B. (2018) What can goby otolith morphology tell us? Cybium, 42, 349 - 363. https: // doi. org / 10.26028 / cybium / 2018 - 424 - 006" type="journal article" year="2018">
|
||
Gierl
|
||
<emphasis id="B93DEAB7FF86FFDFAD6DFE6EFACAFE0F" box="[1310,1366,441,467]" italics="true" pageId="11" pageNumber="242">et al.</emphasis>
|
||
2018
|
||
</bibRefCitation>
|
||
:
|
||
<figureCitation id="13722A20FF86FFDFA8E4FE09FF7AFE2B" box="[151,230,477,503]" captionStart="FIGURE 2" captionStartId="6.[151,250,1935,1960]" captionTargetBox="[151,1436,733,1915]" captionTargetId="figure@6.[151,1436,733,1915]" captionTargetPageId="6" captionText="FIGURE 2. Lebetus patzneri sp. nov., aspects of the freshly collected holotype, ZSM 47486, female, 16.6 mm SL, from off SW Mallorca Island, 72 m depth, preserved. a: left lateral view; b: right lateral view; c: ventral view; c: dorsal view; d: ventral view left lateral view. Photographs by F. Ordinas." figureDoi="http://doi.org/10.5281/zenodo.3567340" httpUri="https://zenodo.org/record/3567340/files/figure.png" pageId="11" pageNumber="242">Figs. 2</figureCitation>
|
||
,
|
||
<figureCitation id="13722A20FF86FFDFA880FE09FE9FFE2B" box="[243,259,477,503]" captionStart="FIGURE 3" captionStartId="7.[151,250,1726,1751]" captionTargetBox="[177,1410,917,1703]" captionTargetId="figure@7.[170,1418,913,1707]" captionTargetPageId="7" captionText="FIGURE 3. Lebetus patzneri sp. nov., aspects of the preserved holotype, reversibly stained with Cyanine Blue, ZSM 47486, female, 16.6 mm SL, from off SW Mallorca Island, 72 m depth, preserved. a: left lateral view b: dorsal view; c: ventral view. Photographs by M. Kovačić." figureDoi="http://doi.org/10.5281/zenodo.3567342" httpUri="https://zenodo.org/record/3567342/files/figure.png" pageId="11" pageNumber="242">3</figureCitation>
|
||
). The quadratic otolith shape of
|
||
<taxonomicName id="4C494D26FF86FFDFAA06FE0AFD6CFE2B" authority="Schliewen & Kovačić & Cerwenka & Svensen & Ordines, 2019" authorityName="Schliewen & Kovačić & Cerwenka & Svensen & Ordines" authorityYear="2019" box="[629,752,477,503]" class="Actinopterygii" family="Gobiidae" genus="Lebetus" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="242" phylum="Chordata" rank="species" species="patzneri" status="sp. nov.">
|
||
<emphasis id="B93DEAB7FF86FFDFAA06FE0AFD19FE2B" box="[629,645,478,503]" italics="true" pageId="11" pageNumber="242">L</emphasis>
|
||
.
|
||
<emphasis id="B93DEAB7FF86FFDFAAE7FE0AFD6CFE2B" box="[660,752,478,503]" italics="true" pageId="11" pageNumber="242">patzneri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B93DEAB7FF86FFDFAA8BFE09FCCEFE2B" bold="true" box="[760,850,477,503]" pageId="11" pageNumber="242">
|
||
<taxonomicNameLabel id="A20E57CCFF86FFDFAA8BFE09FCCEFE2B" box="[760,850,477,503]" pageId="11" pageNumber="242" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
resembles that of
|
||
<taxonomicName id="4C494D26FF86FFDFAC50FE09FAF8FE2B" baseAuthorityName="Risso" baseAuthorityYear="1810" box="[1059,1380,477,503]" class="Actinopterygii" family="Gobiidae" genus="Pomatoschistus" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="242" phylum="Chordata" rank="species" species="marmoratus">
|
||
<emphasis id="B93DEAB7FF86FFDFAC50FE09FAF8FE2B" box="[1059,1380,477,503]" italics="true" pageId="11" pageNumber="242">Pomatoschistus marmoratus</emphasis>
|
||
</taxonomicName>
|
||
(imaged in
|
||
<bibRefCitation id="EFD84B54FF86FFDFA881FDD5FE5EFDC7" author="Gierl, C. & Liebl, D. & Sanda, R. & Vukic, J. & Esmaeili, H. R. & Reichenbacher, B." box="[242,450,513,539]" pageId="11" pageNumber="242" pagination="349 - 363" refId="ref16348" refString="Gierl, C., Liebl, D., Sanda, R., Vukic, J., Esmaeili, H. R. & Reichenbacher, B. (2018) What can goby otolith morphology tell us? Cybium, 42, 349 - 363. https: // doi. org / 10.26028 / cybium / 2018 - 424 - 006" type="journal article" year="2018">
|
||
Gierl
|
||
<emphasis id="B93DEAB7FF86FFDFA947FDD6FEF1FDC7" box="[308,365,513,539]" italics="true" pageId="11" pageNumber="242">et al.</emphasis>
|
||
(2018)
|
||
</bibRefCitation>
|
||
,
|
||
<figureCitation id="13722A20FF86FFDFA9BCFDD5FD80FDC0" box="[463,540,513,540]" captionStart="FIGURE 3" captionStartId="7.[151,250,1726,1751]" captionTargetBox="[177,1410,917,1703]" captionTargetId="figure@7.[170,1418,913,1707]" captionTargetPageId="7" captionText="FIGURE 3. Lebetus patzneri sp. nov., aspects of the preserved holotype, reversibly stained with Cyanine Blue, ZSM 47486, female, 16.6 mm SL, from off SW Mallorca Island, 72 m depth, preserved. a: left lateral view b: dorsal view; c: ventral view. Photographs by M. Kovačić." figureDoi="http://doi.org/10.5281/zenodo.3567342" httpUri="https://zenodo.org/record/3567342/files/figure.png" pageId="11" pageNumber="242">Figs. 3</figureCitation>
|
||
d–f). However, a horizontal position of the sulcus as seen in the present otolith does not occur in otoliths of the species of
|
||
<taxonomicName id="4C494D26FF86FFDFAA04FDF1FCBBFDE3" box="[631,807,549,575]" class="Actinopterygii" family="Gobiidae" genus="Pomatoschistus" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="242" phylum="Chordata" rank="genus">
|
||
<emphasis id="B93DEAB7FF86FFDFAA04FDF1FCBBFDE3" box="[631,807,549,575]" italics="true" pageId="11" pageNumber="242">Pomatoschistus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF36662DFF86FFDFA8E4FB3DFC09FAFA" ID-DOI="http://doi.org/10.5281/zenodo.3567348" ID-Zenodo-Dep="3567348" httpUri="https://zenodo.org/record/3567348/files/figure.png" pageId="11" pageNumber="242" startId="11.[151,250,1257,1282]" targetBox="[160,1436,607,1226]" targetPageId="11">
|
||
<paragraph id="8BF636A5FF86FFDFA8E4FB3DFC09FAFA" blockId="11.[151,1437,1257,1318]" pageId="11" pageNumber="242">
|
||
<emphasis id="B93DEAB7FF86FFDFA8E4FB3DFE8FFADE" bold="true" box="[151,275,1257,1282]" pageId="11" pageNumber="242">FIGURE 6.</emphasis>
|
||
<taxonomicName id="4C494D26FF86FFDFA96AFB3EFE5DFADE" authority="Schliewen & Kovačić & Cerwenka & Svensen & Ordines, 2019" authorityName="Schliewen & Kovačić & Cerwenka & Svensen & Ordines" authorityYear="2019" box="[281,449,1258,1282]" class="Actinopterygii" family="Gobiidae" genus="Lebetus" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="242" phylum="Chordata" rank="species" species="patzneri" status="sp. nov.">
|
||
<emphasis id="B93DEAB7FF86FFDFA96AFB3EFE5DFADE" box="[281,449,1258,1282]" italics="true" pageId="11" pageNumber="242">Lebetus patzneri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B93DEAB7FF86FFDFA9BBFB3EFD84FADE" bold="true" box="[456,536,1258,1282]" pageId="11" pageNumber="242">
|
||
<taxonomicNameLabel id="A20E57CCFF86FFDFA9BBFB3EFD84FADE" box="[456,536,1258,1282]" pageId="11" pageNumber="242" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, head lateral line system of the holotype ZSM 47486, female, 16.6 mm SL, from off SW Mallorca Island, 72 m depth. Terminology in text. Drawing by M. Kovačić.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF636A5FF86FFD8A8B4FA81FBDFFE9F" blockId="11.[151,1437,1364,2039]" lastBlockId="12.[151,1436,153,323]" lastPageId="12" lastPageNumber="243" pageId="11" pageNumber="242">
|
||
<emphasis id="B93DEAB7FF86FFDFA8B4FA81FEDEFAB3" box="[199,322,1365,1391]" italics="true" pageId="11" pageNumber="242">Coloration</emphasis>
|
||
. Color of the freshly collected females,
|
||
<typeStatus id="54F28807FF86FFDFAB75FA80FCF4FAB2" box="[774,872,1364,1390]" pageId="11" pageNumber="242" type="holotype">holotype</typeStatus>
|
||
(
|
||
<figureCitation id="13722A20FF86FFDFAB04FA80FC59FAB3" box="[887,965,1364,1391]" captionStart="FIGURE 2" captionStartId="6.[151,250,1935,1960]" captionTargetBox="[151,1436,733,1915]" captionTargetId="figure@6.[151,1436,733,1915]" captionTargetPageId="6" captionText="FIGURE 2. Lebetus patzneri sp. nov., aspects of the freshly collected holotype, ZSM 47486, female, 16.6 mm SL, from off SW Mallorca Island, 72 m depth, preserved. a: left lateral view; b: right lateral view; c: ventral view; c: dorsal view; d: ventral view left lateral view. Photographs by F. Ordinas." figureDoi="http://doi.org/10.5281/zenodo.3567340" httpUri="https://zenodo.org/record/3567340/files/figure.png" pageId="11" pageNumber="242">Figs. 2</figureCitation>
|
||
.a–d) and
|
||
<typeStatus id="54F28807FF86FFDFAC47FA81FB08FAB3" box="[1076,1172,1365,1391]" pageId="11" pageNumber="242" type="paratype">paratype</typeStatus>
|
||
(
|
||
<figureCitation id="13722A20FF86FFDFACD7FA80FA88FAB3" box="[1188,1300,1364,1391]" captionStart="FIGURE 8" captionStartId="13.[151,250,1801,1826]" captionTargetBox="[179,1412,181,1770]" captionTargetId="figure@13.[159,1429,166,1782]" captionTargetPageId="13" captionText="FIGURE 8. Lebetus patzneri sp. nov., aspects of the paratype and two unpreserved specimens in life coloration shortly after collection. a: left lateral view of paratype ZSM 46865, female, 16.8 mm SL; from off SW Mallorca Island, 60m depth (based on montage of two separate photographs (slight artifacts at montage suture at about D1 origin due to slightly different position of the specimen between the two photographs); b: left dorsolateral view of head and anterior flank view of paratype ZSM 46865; c: left lateral view of the second known specimen (unpreserved), unknown size, Menorca channel, 67m depth (based on montage of two separate photographs; stitching artifacts visible before D1 origin); d: left slightly dorsolateral view of third known specimen (unpreserved) of unknown size, Menorca channel, 67m depth. Photographs by F. Ordines." figureDoi="http://doi.org/10.5281/zenodo.3567352" httpUri="https://zenodo.org/record/3567352/files/figure.png" pageId="11" pageNumber="242">Figs. 8a,b</figureCitation>
|
||
) and photographs of two additional specimens (
|
||
<figureCitation id="13722A20FF86FFDFAA41FAACFD38FA4F" box="[562,676,1400,1427]" captionStart="FIGURE 8" captionStartId="13.[151,250,1801,1826]" captionTargetBox="[179,1412,181,1770]" captionTargetId="figure@13.[159,1429,166,1782]" captionTargetPageId="13" captionText="FIGURE 8. Lebetus patzneri sp. nov., aspects of the paratype and two unpreserved specimens in life coloration shortly after collection. a: left lateral view of paratype ZSM 46865, female, 16.8 mm SL; from off SW Mallorca Island, 60m depth (based on montage of two separate photographs (slight artifacts at montage suture at about D1 origin due to slightly different position of the specimen between the two photographs); b: left dorsolateral view of head and anterior flank view of paratype ZSM 46865; c: left lateral view of the second known specimen (unpreserved), unknown size, Menorca channel, 67m depth (based on montage of two separate photographs; stitching artifacts visible before D1 origin); d: left slightly dorsolateral view of third known specimen (unpreserved) of unknown size, Menorca channel, 67m depth. Photographs by F. Ordines." figureDoi="http://doi.org/10.5281/zenodo.3567352" httpUri="https://zenodo.org/record/3567352/files/figure.png" pageId="11" pageNumber="242">Figs. 8c,d</figureCitation>
|
||
).
|
||
<typeStatus id="54F28807FF86FFDFAACBFAACFCBCFA4E" box="[696,800,1400,1426]" pageId="11" pageNumber="242" type="holotype">Holotype</typeStatus>
|
||
and
|
||
<typeStatus id="54F28807FF86FFDFAB25FAADFC27FA4F" box="[854,955,1401,1427]" pageId="11" pageNumber="242" type="paratype">paratype</typeStatus>
|
||
: ground color of body, head, snout and lips orange-red except for white areas of lower head, lower postopercular region belly and ventral caudal peduncle; back with four whitish-beige saddle marks, one before D1 origin, one below D1, one at D2 origin and one forming a broad vertical band on caudal peduncle, the latter vermiculated in the
|
||
<typeStatus id="54F28807FF86FFDFAB11FA30FC54FA22" box="[866,968,1508,1534]" pageId="11" pageNumber="242" type="holotype">holotype</typeStatus>
|
||
; additional very small whitish blotches in the predorsal area and along posterior lower flanks. Chin, suborbital region and lower parts of preopercle porcelain white with two well defined red bands almost meeting at a moderately acute angle below eye diverging radially downwards; ventral part of preopercle fringed with a narrow red margin in the
|
||
<typeStatus id="54F28807FF86FFDFAB8CF985FBFEF9B7" box="[1023,1122,1617,1643]" pageId="11" pageNumber="242" type="paratype">paratype</typeStatus>
|
||
, but reduced to a red narrow spot in the
|
||
<typeStatus id="54F28807FF86FFDFA962F9A0FEEBF952" box="[273,375,1652,1678]" pageId="11" pageNumber="242" type="holotype">holotype</typeStatus>
|
||
. Iris orange-red, partially blackish or with a golden hue; interior iris margin golden. Dorsal area and flanks with many minute blackish spots, fewer in the
|
||
<typeStatus id="54F28807FF86FFDFAB43F94DFC0CF96F" box="[816,912,1689,1715]" pageId="11" pageNumber="242" type="paratype">paratype</typeStatus>
|
||
on predorsal region than in the
|
||
<typeStatus id="54F28807FF86FFDFAD71F94CFAFBF96E" box="[1282,1383,1688,1714]" pageId="11" pageNumber="242" type="holotype">holotype</typeStatus>
|
||
, and generally more densely aggregated on caudal peduncle and anal fin base, and forming in the
|
||
<typeStatus id="54F28807FF86FFDFACDCF969FA93F90B" box="[1199,1295,1725,1751]" pageId="11" pageNumber="242" type="paratype">paratype</typeStatus>
|
||
two vertical narrow bands at vertical of D1 anterior and posterior origins. Head, preopercle, opercle, pectoral and anterior two thirds of ventral area almost without minute small black spots. D1 black (folded down on photos), in
|
||
<typeStatus id="54F28807FF86FFDFAD70F8D1FAFFF8C3" box="[1283,1379,1797,1823]" pageId="11" pageNumber="242" type="paratype">paratype</typeStatus>
|
||
with two narrow white bands indicated at distal margin; D2 (folded down on photos) red with white bands indicated. P base white with scattered orange-red spots, P transparent with few thin reddish bands indicated in
|
||
<typeStatus id="54F28807FF86FFDFAC91F899FADAF8BB" box="[1250,1350,1869,1895]" pageId="11" pageNumber="242" type="paratype">paratype</typeStatus>
|
||
. C with a porcelain white basal blotch, sharply delineated anteriorly, otherwise transparent orange and with a whitish band at about middle part (visible only in
|
||
<typeStatus id="54F28807FF86FFDFAA46F840FD0BF872" box="[565,663,1940,1966]" pageId="11" pageNumber="242" type="holotype">holotype</typeStatus>
|
||
and one of the additional only photographed specimens). The coloration of the two additional specimens is similar, matching the
|
||
<typeStatus id="54F28807FF86FFDFAB23F86CFC2EF80E" box="[848,946,1976,2002]" pageId="11" pageNumber="242" type="holotype">holotype</typeStatus>
|
||
coloration, D1 of the one unpreserved only photographed specimen (
|
||
<figureCitation id="13722A20FF86FFDFA9DCF808FE61F82B" box="[431,509,2012,2039]" captionStart="FIGURE 8" captionStartId="13.[151,250,1801,1826]" captionTargetBox="[179,1412,181,1770]" captionTargetId="figure@13.[159,1429,166,1782]" captionTargetPageId="13" captionText="FIGURE 8. Lebetus patzneri sp. nov., aspects of the paratype and two unpreserved specimens in life coloration shortly after collection. a: left lateral view of paratype ZSM 46865, female, 16.8 mm SL; from off SW Mallorca Island, 60m depth (based on montage of two separate photographs (slight artifacts at montage suture at about D1 origin due to slightly different position of the specimen between the two photographs); b: left dorsolateral view of head and anterior flank view of paratype ZSM 46865; c: left lateral view of the second known specimen (unpreserved), unknown size, Menorca channel, 67m depth (based on montage of two separate photographs; stitching artifacts visible before D1 origin); d: left slightly dorsolateral view of third known specimen (unpreserved) of unknown size, Menorca channel, 67m depth. Photographs by F. Ordines." figureDoi="http://doi.org/10.5281/zenodo.3567352" httpUri="https://zenodo.org/record/3567352/files/figure.png" pageId="11" pageNumber="242">Fig. 8c</figureCitation>
|
||
) is partially unfolded and exhibits an overall black coloration and small pinkish-red elements along the base of two spines as well as a white blotch at the upper posterior area and around the membrane between the last D1 spine and the dorsum; furthermore, whitish-transparent V are visible in the
|
||
<typeStatus id="54F28807FF81FFD8ACA3FF69FAAEFF0B" box="[1232,1330,189,215]" pageId="12" pageNumber="243" type="holotype">holotype</typeStatus>
|
||
(
|
||
<figureCitation id="13722A20FF81FFD8AD31FF69FA08FF04" box="[1346,1428,189,216]" captionStart="FIGURE 2" captionStartId="6.[151,250,1935,1960]" captionTargetBox="[151,1436,733,1915]" captionTargetId="figure@6.[151,1436,733,1915]" captionTargetPageId="6" captionText="FIGURE 2. Lebetus patzneri sp. nov., aspects of the freshly collected holotype, ZSM 47486, female, 16.6 mm SL, from off SW Mallorca Island, 72 m depth, preserved. a: left lateral view; b: right lateral view; c: ventral view; c: dorsal view; d: ventral view left lateral view. Photographs by F. Ordinas." figureDoi="http://doi.org/10.5281/zenodo.3567340" httpUri="https://zenodo.org/record/3567340/files/figure.png" pageId="12" pageNumber="243">Fig. 2d</figureCitation>
|
||
) and second unpreserved and only photographed specimen (
|
||
<figureCitation id="13722A20FF81FFD8AB5FFF35FC1CFF20" box="[812,896,225,252]" captionStart="FIGURE 8" captionStartId="13.[151,250,1801,1826]" captionTargetBox="[179,1412,181,1770]" captionTargetId="figure@13.[159,1429,166,1782]" captionTargetPageId="13" captionText="FIGURE 8. Lebetus patzneri sp. nov., aspects of the paratype and two unpreserved specimens in life coloration shortly after collection. a: left lateral view of paratype ZSM 46865, female, 16.8 mm SL; from off SW Mallorca Island, 60m depth (based on montage of two separate photographs (slight artifacts at montage suture at about D1 origin due to slightly different position of the specimen between the two photographs); b: left dorsolateral view of head and anterior flank view of paratype ZSM 46865; c: left lateral view of the second known specimen (unpreserved), unknown size, Menorca channel, 67m depth (based on montage of two separate photographs; stitching artifacts visible before D1 origin); d: left slightly dorsolateral view of third known specimen (unpreserved) of unknown size, Menorca channel, 67m depth. Photographs by F. Ordines." figureDoi="http://doi.org/10.5281/zenodo.3567352" httpUri="https://zenodo.org/record/3567352/files/figure.png" pageId="12" pageNumber="243">Fig. 8d</figureCitation>
|
||
). In the second photographed only specimen the C base has an inverted brownish triangular bar, darker at the caudal and ventral margins, more reddish on the dorsal side, and the posterior base of the same vertical boundary as in the other specimens.
|
||
</paragraph>
|
||
<caption id="DF36662DFF81FFD8A8E4FC4EFF7DFBC3" ID-DOI="http://doi.org/10.5281/zenodo.3567350" ID-Zenodo-Dep="3567350" httpUri="https://zenodo.org/record/3567350/files/figure.png" pageId="12" pageNumber="243" startId="12.[151,250,922,947]" targetBox="[151,1435,337,902]" targetPageId="12">
|
||
<paragraph id="8BF636A5FF81FFD8A8E4FC4EFF7DFBC3" blockId="12.[151,1437,922,1055]" pageId="12" pageNumber="243">
|
||
<emphasis id="B93DEAB7FF81FFD8A8E4FC4EFE88FC6E" bold="true" box="[151,276,922,947]" pageId="12" pageNumber="243">FIGURE 7.</emphasis>
|
||
<taxonomicName id="4C494D26FF81FFD8A96FFC4EFE5AFC6E" authority="Schliewen & Kovačić & Cerwenka & Svensen & Ordines, 2019" authorityName="Schliewen & Kovačić & Cerwenka & Svensen & Ordines" authorityYear="2019" box="[284,454,922,946]" class="Actinopterygii" family="Gobiidae" genus="Lebetus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="243" phylum="Chordata" rank="species" species="patzneri" status="sp. nov.">
|
||
<emphasis id="B93DEAB7FF81FFD8A96FFC4EFE5AFC6E" box="[284,454,922,946]" italics="true" pageId="12" pageNumber="243">Lebetus patzneri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B93DEAB7FF81FFD8A9BDFC4EFDBCFC6E" bold="true" box="[462,544,922,946]" pageId="12" pageNumber="243">
|
||
<taxonomicNameLabel id="A20E57CCFF81FFD8A9BDFC4EFDBCFC6E" box="[462,544,922,946]" pageId="12" pageNumber="243" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, right saccular otolith of the paratype ZSM 46865, female, 16.8 mm SL; from off SW Mallorca, 60m depth.
|
||
<emphasis id="B93DEAB7FF81FFD8A907FC6AFE1DFC0A" bold="true" box="[372,385,958,982]" pageId="12" pageNumber="243">a</emphasis>
|
||
: in medial/inner view;
|
||
<emphasis id="B93DEAB7FF81FFD8AA1AFC6AFDEBFC0A" bold="true" box="[617,631,958,982]" pageId="12" pageNumber="243">b</emphasis>
|
||
: left otolith in anterior view. Orientation of otolith with respect to fish symmetry is indicated as follows: a, anterior; d, dorsal; l, lateral; m, medial; p, posterior; v, ventral. MicroCT reconstruction by A. Cerwenka.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF636A5FF81FFD8A8B4FB99FAF0FB0F" blockId="12.[151,1437,1101,2028]" pageId="12" pageNumber="243">
|
||
<emphasis id="B93DEAB7FF81FFD8A8B4FB99FD0CFBBB" box="[199,656,1101,1127]" italics="true" pageId="12" pageNumber="243">Color of preserved specimens in alcohol</emphasis>
|
||
. Body of
|
||
<typeStatus id="54F28807FF81FFD8AB70FB99FCF9FBBB" box="[771,869,1101,1127]" pageId="12" pageNumber="243" type="holotype">holotype</typeStatus>
|
||
yellowish white, with orange-red coloration completely lost. The minute small black spots on the dorsal area and flanks with still clearly visible. D1 black and other fins transparent. The
|
||
<typeStatus id="54F28807FF81FFD8A9F9FB41FE76FB73" box="[394,490,1173,1199]" pageId="12" pageNumber="243" type="paratype">paratype</typeStatus>
|
||
was completely discolored (whitish) before preparation of the specimen for REM, except for the distinct and scattered and minute black spots on predorsal dorsal and posterior flanks. D1 black.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C353652EFF81FFDAA8B4FB09FBC9FE0F" lastPageId="14" lastPageNumber="245" pageId="12" pageNumber="243" type="etymology">
|
||
<paragraph id="8BF636A5FF81FFD8A8B4FB09FCA4FAE3" blockId="12.[151,1437,1101,2028]" pageId="12" pageNumber="243">
|
||
<emphasis id="B93DEAB7FF81FFD8A8B4FB09FED4FB2B" bold="true" box="[199,328,1245,1271]" pageId="12" pageNumber="243">Etymology</emphasis>
|
||
. The species is named
|
||
<taxonomicName id="4C494D26FF81FFD8AA24FB0AFD2FFB2B" authorityName="Schliewen & Kovačić & Cerwenka & Svensen & Ordines" authorityYear="2019" box="[599,691,1246,1271]" class="Actinopterygii" family="Gobiidae" genus="Lebetus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="243" phylum="Chordata" rank="species" species="patzneri">
|
||
<emphasis id="B93DEAB7FF81FFD8AA24FB0AFD2FFB2B" box="[599,691,1246,1271]" italics="true" pageId="12" pageNumber="243">patzneri</emphasis>
|
||
</taxonomicName>
|
||
, to honor Prof. Robert (“Bobby”) Patzner, Professor Emeritus at the University of
|
||
<collectingRegion id="498DF847FF81FFD8A92CFAD5FE5EFAC7" box="[351,450,1281,1307]" country="Austria" name="Salzburg" pageId="12" pageNumber="243">Salzburg</collectingRegion>
|
||
(
|
||
<collectingCountry id="F35E7635FF81FFD8A9A1FAD5FDB6FAC0" box="[466,554,1281,1308]" name="Austria" pageId="12" pageNumber="243">Austria</collectingCountry>
|
||
), in recognition for his dedication to the study of gobies in the Mediterranean including the Balearic Islands, see e.g.
|
||
<bibRefCitation id="EFD84B54FF81FFD8AA3EFAF1FCA8FAE3" author="Patzner, R. A. & Van Tassell, J. L. & Kovacic, M. & Kapoora, B. G." box="[589,820,1317,1343]" pageId="12" pageNumber="243" refId="ref17751" refString="Patzner, R. A., Van Tassell, J. L., Kovacic, M. & Kapoora, B. G. (2011) The Biology of Gobies. Science Publishers CRC Press, Boca Raton, Florida, 702 pp. https: // doi. org / 10.1201 / b 11397" type="book" year="2011">
|
||
Patzner
|
||
<emphasis id="B93DEAB7FF81FFD8AADBFAF2FD7CFAE3" box="[680,736,1317,1343]" italics="true" pageId="12" pageNumber="243">et al.</emphasis>
|
||
(2011)
|
||
</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BF636A5FF81FFD8A8B4FA9CFD7BF95F" blockId="12.[151,1437,1101,2028]" pageId="12" pageNumber="243">
|
||
<emphasis id="B93DEAB7FF81FFD8A8B4FA9CFDF9FABF" bold="true" box="[199,613,1352,1379]" pageId="12" pageNumber="243">Genetics and phylogenetic position</emphasis>
|
||
(
|
||
<figureCitation id="13722A20FF81FFD8AA07FA9DFD25FABF" box="[628,697,1353,1379]" captionStart="FIGURE 9" captionStartId="14.[151,250,1193,1218]" captionTargetBox="[156,1429,482,1167]" captionTargetId="figure@14.[151,1437,481,1173]" captionTargetPageId="14" captionText="FIGURE 9. Maximum likelihood phylogenetic tree topology depicting the phylogenetic position and small intraspecific genetic distance of COI-barcode haplotypes (520 bp) of the holotype of Lebetus patzneri sp. nov. and five Lebetus guilleti from the northeastern Atlantic (Norway, two specimens), the Western Mediterranean (Baleares, one specimen) and from the northern Adriatic (Croatia, two specimens). All sand goby genera are included, except for Hyrcanogobius and Pseudaphya. Numbers on branches are bootstrap support values (%) for the Maximum Likelihood analysis (values below 50% not shown). Tip labels are composed of the genus and species name followed by ZSM collection number (if newly sequenced in this study) or of the GenBank accession number. Inserted photos depict the second known specimen of L. patzneri sp. nov. (as Fig. 8c; upper photograph) and the first L. guilleti specimen photographed and collected in 15m depth in Norway (ZSM-PIS-GO 1227; lower photograph). Photographs above and below by F. Ordines and R. Svensen, respectively." figureDoi="http://doi.org/10.5281/zenodo.3567354" httpUri="https://zenodo.org/record/3567354/files/figure.png" pageId="12" pageNumber="243">Fig. 9</figureCitation>
|
||
). Partial cytochrome oxidase subunit I (COI) sequences (520 base pairs) of
|
||
<taxonomicName id="4C494D26FF81FFD8A88FFAB9FE2AFA5B" authority="Schliewen & Kovačić & Cerwenka & Svensen & Ordines, 2019" authorityName="Schliewen & Kovačić & Cerwenka & Svensen & Ordines" authorityYear="2019" box="[252,438,1389,1415]" class="Actinopterygii" family="Gobiidae" genus="Lebetus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="243" phylum="Chordata" rank="species" species="patzneri" status="sp. nov.">
|
||
<emphasis id="B93DEAB7FF81FFD8A88FFAB9FE2AFA5B" box="[252,438,1389,1415]" italics="true" pageId="12" pageNumber="243">Lebetus patzneri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B93DEAB7FF81FFD8A9CEFAB9FD8BFA5B" bold="true" box="[445,535,1389,1415]" pageId="12" pageNumber="243">
|
||
<taxonomicNameLabel id="A20E57CCFF81FFD8A9CEFAB9FD8BFA5B" box="[445,535,1389,1415]" pageId="12" pageNumber="243" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
compared with five
|
||
<taxonomicName id="4C494D26FF81FFD8AA8DFABAFCF7FA5B" baseAuthorityName="Le Danois" baseAuthorityYear="1913" box="[766,875,1389,1415]" class="Actinopterygii" family="Gobiidae" genus="Lebetus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="243" phylum="Chordata" rank="species" species="guilleti">
|
||
<emphasis id="B93DEAB7FF81FFD8AA8DFABAFCF7FA5B" box="[766,875,1389,1415]" italics="true" pageId="12" pageNumber="243">L. guilleti</emphasis>
|
||
</taxonomicName>
|
||
and one
|
||
<taxonomicName id="4C494D26FF81FFD8ABA2FABAFBECFA5B" baseAuthorityName="Collett" baseAuthorityYear="1874" box="[977,1136,1389,1415]" class="Actinopterygii" family="Gobiidae" genus="Lebetus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="243" phylum="Chordata" rank="species" species="scorpioides">
|
||
<emphasis id="B93DEAB7FF81FFD8ABA2FABAFBECFA5B" box="[977,1136,1389,1415]" italics="true" pageId="12" pageNumber="243">L. scorpioides</emphasis>
|
||
</taxonomicName>
|
||
sequences, as well as with those of members of all other sand goby and DLBS genera provide good support for the phylogenetic position as the sisterclade of the
|
||
<taxonomicName id="4C494D26FF81FFD8A9F6FA62FDB8FA13" baseAuthorityName="Collett" baseAuthorityYear="1874" box="[389,548,1461,1487]" class="Actinopterygii" family="Gobiidae" genus="Lebetus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="243" phylum="Chordata" rank="species" species="scorpioides">
|
||
<emphasis id="B93DEAB7FF81FFD8A9F6FA62FDB8FA13" box="[389,548,1461,1487]" italics="true" pageId="12" pageNumber="243">L. scorpioides</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="F35E7635FF81FFD8AA1BFA61FD5EFA13" box="[616,706,1461,1487]" name="Norway" pageId="12" pageNumber="243">Norway</collectingCountry>
|
||
(BS 87), as well as for its species-level distinctiveness. K2P-distance between the single
|
||
<taxonomicName id="4C494D26FF81FFD8A9C4FA0EFDAEFA2F" authorityName="Schliewen & Kovačić & Cerwenka & Svensen & Ordines" authorityYear="2019" box="[439,562,1498,1523]" class="Actinopterygii" family="Gobiidae" genus="Lebetus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="243" phylum="Chordata" rank="species" species="patzneri">
|
||
<emphasis id="B93DEAB7FF81FFD8A9C4FA0EFDAEFA2F" box="[439,562,1498,1523]" italics="true" pageId="12" pageNumber="243">L. patzneri</emphasis>
|
||
</taxonomicName>
|
||
and the single
|
||
<taxonomicName id="4C494D26FF81FFD8AA92FA0EFC1DFA2F" baseAuthorityName="Collett" baseAuthorityYear="1874" box="[737,897,1497,1523]" class="Actinopterygii" family="Gobiidae" genus="Lebetus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="243" phylum="Chordata" rank="species" species="scorpioides">
|
||
<emphasis id="B93DEAB7FF81FFD8AA92FA0EFC1DFA2F" box="[737,897,1497,1523]" italics="true" pageId="12" pageNumber="243">L. scorpioides</emphasis>
|
||
</taxonomicName>
|
||
COI-haplotype is 0.150 (±0.018), and between
|
||
<taxonomicName id="4C494D26FF81FFD8A8E4FA2AFE8FF9CB" authorityName="Schliewen & Kovačić & Cerwenka & Svensen & Ordines" authorityYear="2019" box="[151,275,1534,1559]" class="Actinopterygii" family="Gobiidae" genus="Lebetus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="243" phylum="Chordata" rank="species" species="patzneri">
|
||
<emphasis id="B93DEAB7FF81FFD8A8E4FA2AFE8FF9CB" box="[151,275,1534,1559]" italics="true" pageId="12" pageNumber="243">L. patzneri</emphasis>
|
||
</taxonomicName>
|
||
and each of the five
|
||
<taxonomicName id="4C494D26FF81FFD8AA7AFA2AFDEBF9CB" baseAuthorityName="Le Danois" baseAuthorityYear="1913" box="[521,631,1533,1559]" class="Actinopterygii" family="Gobiidae" genus="Lebetus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="243" phylum="Chordata" rank="species" species="guilleti">
|
||
<emphasis id="B93DEAB7FF81FFD8AA7AFA2AFDEBF9CB" box="[521,631,1533,1559]" italics="true" pageId="12" pageNumber="243">L. guilleti</emphasis>
|
||
</taxonomicName>
|
||
haplotypes it ranges from 0.195 (±
|
||
<geoCoordinate id="EE7D5062FF81FFD8AC78FA29FBFAF9CB" box="[1035,1126,1533,1559]" degrees="0.019" direction="south" orientation="latitude" pageId="12" pageNumber="243" precision="55" value="-0.019">0.019 S</geoCoordinate>
|
||
.E.) to 0.196 (±
|
||
<geoCoordinate id="EE7D5062FF81FFD8AD60FA29FAF2F9CB" box="[1299,1390,1533,1559]" degrees="0.019" direction="south" orientation="latitude" pageId="12" pageNumber="243" precision="55" value="-0.019">0.019 S</geoCoordinate>
|
||
.E.). Thus, pairwise K2P-distances between the three congeneric
|
||
<taxonomicName id="4C494D26FF81FFD8AB4BF9F5FC13F9E7" authorityName="Whinter" authorityYear="1877" box="[824,911,1569,1595]" class="Actinopterygii" family="Gobiidae" genus="Lebetus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="243" phylum="Chordata" rank="genus">
|
||
<emphasis id="B93DEAB7FF81FFD8AB4BF9F5FC13F9E7" box="[824,911,1569,1595]" italics="true" pageId="12" pageNumber="243">Lebetus</emphasis>
|
||
</taxonomicName>
|
||
taxa range well above distances between other congeneric sand goby and DLBS-taxa, e.g.
|
||
<taxonomicName id="4C494D26FF81FFD8AAF3F991FCACF983" box="[640,816,1605,1631]" class="Actinopterygii" family="Gobiidae" genus="Pomatoschistus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="243" phylum="Chordata" rank="genus">
|
||
<emphasis id="B93DEAB7FF81FFD8AAF3F991FCACF983" box="[640,816,1605,1631]" italics="true" pageId="12" pageNumber="243">Pomatoschistus</emphasis>
|
||
</taxonomicName>
|
||
species (
|
||
<bibRefCitation id="EFD84B54FF81FFD8ABE5F991FB41F983" author="Knebelsberger, T. & Thiel, R." box="[918,1245,1605,1631]" pageId="12" pageNumber="243" pagination="831 - 845" refId="ref17052" refString="Knebelsberger, T. & Thiel, R. (2014) Identification of gobies (Teleostei: Perciformes: Gobiidae) from the North and Baltic Seas combining morphological analysis and DNA barcoding. Zoological Journal of the Linnean Society, 172, 831 - 845. https: // doi. org / 10.1111 / zoj. 12189." type="journal article" year="2014">Knebelsberger & Thiel 2014</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFD84B54FF81FFD8AC94F990FF4DF95F" author="Ozturk, D. S. & Engin, S." pageId="12" pageNumber="243" pagination="142 - 150" refId="ref17693" refString="Ozturk, D. S. & Engin, S. (2019) Taxonomic status of the Mediterranean-endemic goby Pomatoschistus adriaticus Miller, 1973 inferred with both morphological and genetic data. Zoology of the Middle East, 65 (4), 142 - 150. https: // doi. org / 10.1080 / 09397140.2019.1580932" type="journal article" year="2019">Öztürk & Engin 2019</bibRefCitation>
|
||
) or
|
||
<taxonomicName id="4C494D26FF81FFD8A88DF9BEFED1F95F" authorityName="Iljin" authorityYear="1930" box="[254,333,1642,1667]" class="Actinopterygii" family="Gobiidae" genus="Buenia" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="243" phylum="Chordata" rank="genus">
|
||
<emphasis id="B93DEAB7FF81FFD8A88DF9BEFED1F95F" box="[254,333,1642,1667]" italics="true" pageId="12" pageNumber="243">Buenia</emphasis>
|
||
</taxonomicName>
|
||
species (
|
||
<bibRefCitation id="EFD84B54FF81FFD8A9C7F9BDFD08F95F" author="Kovacic, M. & Ordines, F. & Schliewen, U. K." box="[436,660,1641,1667]" pageId="12" pageNumber="243" pagination="447 - 460" refId="ref16846" refString="Kovacic, M., Ordines, F. & Schliewen, U. K. (2017) A new species of Buenia (Teleostei: Gobiidae) from the western Mediterranean Sea, with the description of this genus. Zootaxa, 4250 (5), 447 - 460. https: // doi. org / 10.11646 / zootaxa. 4250.5.3" type="journal article" year="2017">
|
||
Kovačić
|
||
<emphasis id="B93DEAB7FF81FFD8AA69F9BEFDCEF95F" box="[538,594,1641,1667]" italics="true" pageId="12" pageNumber="243">et al.</emphasis>
|
||
2017
|
||
</bibRefCitation>
|
||
, 2018).
|
||
</paragraph>
|
||
<paragraph id="8BF636A5FF81FFDAA8B4F958FBC9FE0F" blockId="12.[151,1437,1101,2028]" lastBlockId="14.[151,1437,153,467]" lastPageId="14" lastPageNumber="245" pageId="12" pageNumber="243">
|
||
<emphasis id="B93DEAB7FF81FFD8A8B4F958FD2CF97B" bold="true" box="[199,688,1676,1703]" pageId="12" pageNumber="243">Ecological and geographical distribution</emphasis>
|
||
.
|
||
<taxonomicName id="4C494D26FF81FFD8AACCF959FCE7F97B" authority="Schliewen & Kovačić & Cerwenka & Svensen & Ordines, 2019" authorityName="Schliewen & Kovačić & Cerwenka & Svensen & Ordines" authorityYear="2019" box="[703,891,1677,1703]" class="Actinopterygii" family="Gobiidae" genus="Lebetus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="243" phylum="Chordata" rank="species" species="patzneri" status="sp. nov.">
|
||
<emphasis id="B93DEAB7FF81FFD8AACCF959FCE7F97B" box="[703,891,1677,1703]" italics="true" pageId="12" pageNumber="243">Lebetus patzneri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B93DEAB7FF81FFD8ABF0F959FC42F97B" bold="true" box="[899,990,1677,1703]" pageId="12" pageNumber="243">
|
||
<taxonomicNameLabel id="A20E57CCFF81FFD8ABF0F959FC42F97B" box="[899,990,1677,1703]" pageId="12" pageNumber="243" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
has only been caught so far from sampling stations on red algae beds in south
|
||
<collectingCountry id="F35E7635FF81FFD8AA1EF965FD4FF917" box="[621,723,1713,1739]" name="Spain" pageId="12" pageNumber="243">Menorca</collectingCountry>
|
||
, the
|
||
<collectingCountry id="F35E7635FF81FFD8AB78F965FCF3F917" box="[779,879,1713,1739]" name="Spain" pageId="12" pageNumber="243">Menorca</collectingCountry>
|
||
Channel and off south Mallorca, at depths ranging between 60 and
|
||
<specimenCount id="9D4FFD2CFF81FFD8A90FF901FE21F933" box="[380,445,1749,1775]" pageId="12" pageNumber="243" type="generic">
|
||
<quantity id="4CB19B40FF81FFD8A90FF901FE25F933" box="[380,441,1749,1775]" metricMagnitude="1" metricUnit="m" metricValue="7.2" pageId="12" pageNumber="243" unit="m" value="72.0">72 m</quantity>
|
||
.
|
||
</specimenCount>
|
||
The first two specimens caught, not preserved, were collected from a
|
||
<taxonomicName id="4C494D26FF81FFD8ACABF901FAF0F933" authorityName="Decaisne" authorityYear="1841" box="[1240,1388,1749,1775]" class="Florideophyceae" family="Peyssonneliaceae" genus="Peyssonnelia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Gigartinales" pageId="12" pageNumber="243" phylum="Rhodophyta" rank="genus">
|
||
<emphasis id="B93DEAB7FF81FFD8ACABF901FAF0F933" box="[1240,1388,1749,1775]" italics="true" pageId="12" pageNumber="243">Peyssonnelia</emphasis>
|
||
</taxonomicName>
|
||
bed off the southern coast of
|
||
<collectingCountry id="F35E7635FF81FFD8A9C0F92DFD8BF8CF" box="[435,535,1785,1811]" name="Spain" pageId="12" pageNumber="243">Menorca</collectingCountry>
|
||
(
|
||
<figureCitation id="13722A20FF81FFD8AA55F92DFDF5F8CF" box="[550,617,1785,1811]" captionStart="FIGURE 1" captionStartId="2.[151,250,1666,1691]" captionTargetBox="[151,1435,737,1642]" captionTargetId="figure@2.[151,1435,733,1647]" captionTargetPageId="2" captionText="FIGURE 1. Map showing the three sampling stations where all the individuals of Lebetus patzneri sp. nov. (stars) and L. guilleti newly recorded individuals (circles) from the Balearic Islands were collected. The holotype and the paratype of L. patzneri sp. nov. were collected from the sampling station in the Menorca Channel and off southeast Mallorca, respectively. The holotype, paratype and the two unpreserved specimens off south Menorca were collected at 72, 60 and 67 m, respectively. The 50, 100, 500 and 800 m isobaths are shown. Map drawn by F. Ordinas." figureDoi="http://doi.org/10.5281/zenodo.3567338" httpUri="https://zenodo.org/record/3567338/files/figure.png" pageId="12" pageNumber="243">Fig. 1</figureCitation>
|
||
). In that sample
|
||
<quantity id="4CB19B40FF81FFD8AB58F92DFCF7F8CF" box="[811,875,1785,1811]" metricMagnitude="1" metricUnit="kg" metricValue="8.7" pageId="12" pageNumber="243" unit="kg" value="87.0">87 kg</quantity>
|
||
of algae were collected from a swept area of
|
||
<quantity id="4CB19B40FF81FFD8AD01F92CFF31F8EB" metricMagnitude="2" metricUnit="m" metricValue="5.55" pageId="12" pageNumber="243" unit="m" value="555.0">555 m</quantity>
|
||
<superScript id="7C3C9BEDFF81FFD8A8DEF8C8FF29F8F6" attach="left" box="[173,181,1820,1834]" fontSize="6" pageId="12" pageNumber="243">2</superScript>
|
||
, with dominance of red algae species belonging to the
|
||
<taxonomicName id="4C494D26FF81FFD8AB43F8C9FC68F8EB" authorityName="Denizot" authorityYear="1968" box="[816,1012,1821,1847]" class="Florideophyceae" family="Peyssonneliaceae" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cryptonemiales" pageId="12" pageNumber="243" phylum="Rhodophyta" rank="family">Peyssonneliaceae</taxonomicName>
|
||
family (
|
||
<quantity id="4CB19B40FF81FFD8AC24F8C9FB00F8E4" box="[1111,1180,1821,1848]" metricMagnitude="1" metricUnit="kg" metricValue="7.9" pageId="12" pageNumber="243" unit="kg" value="79.0">79 kg</quantity>
|
||
), mainly
|
||
<taxonomicName id="4C494D26FF81FFD8AD7BF8C9FE99F887" authorityName="Pilger" authorityYear="1911" class="Florideophyceae" family="Peyssonneliaceae" genus="Peyssonnelia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Gigartinales" pageId="12" pageNumber="243" phylum="Rhodophyta" rank="species" species="inamoena">
|
||
<emphasis id="B93DEAB7FF81FFD8AD7BF8C9FE99F887" italics="true" pageId="12" pageNumber="243">Peyssonnelia inamoena</emphasis>
|
||
</taxonomicName>
|
||
but also with some presence of rhodolith forming species such as the coralline algae
|
||
<taxonomicName id="4C494D26FF81FFD8ACBFF895FF58F8A3" authorityName="Foslie" authorityYear="1909" class="Florideophyceae" family="Corallinaceae" genus="Lithothamnion" higherTaxonomySource="GBIF" kingdom="Plantae" order="Corallinales" pageId="12" pageNumber="243" phylum="Rhodophyta" rank="species" species="valens">
|
||
<emphasis id="B93DEAB7FF81FFD8ACBFF895FF58F8A3" italics="true" pageId="12" pageNumber="243">Lithothamnion valens</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C494D26FF81FFD8A88DF8B1FE64F8A3" authorityName="Kutzing" authorityYear="1841" box="[254,504,1893,1919]" class="Florideophyceae" family="Corallinaceae" genus="Spongites" higherTaxonomySource="GBIF" kingdom="Plantae" order="Corallinales" pageId="12" pageNumber="243" phylum="Rhodophyta" rank="species" species="fruticulosus">
|
||
<emphasis id="B93DEAB7FF81FFD8A88DF8B1FE64F8A3" box="[254,504,1893,1919]" italics="true" pageId="12" pageNumber="243">Spongites fruticulosus</emphasis>
|
||
</taxonomicName>
|
||
(3 and
|
||
<quantity id="4CB19B40FF81FFD8AA22F8B1FD00F8A3" box="[593,668,1893,1919]" metricMagnitude="0" metricUnit="kg" metricValue="1.2" pageId="12" pageNumber="243" unit="kg" value="1.2">1.2 kg</quantity>
|
||
, respectively). The
|
||
<typeStatus id="54F28807FF81FFD8AB0EF8B1FC41F8A3" box="[893,989,1893,1919]" pageId="12" pageNumber="243" type="paratype">paratype</typeStatus>
|
||
was collected from a rhodolith bed off the south coast of Mallorca (
|
||
<figureCitation id="13722A20FF81FFD8A993F85DFDBFF87F" box="[480,547,1929,1955]" captionStart="FIGURE 1" captionStartId="2.[151,250,1666,1691]" captionTargetBox="[151,1435,737,1642]" captionTargetId="figure@2.[151,1435,733,1647]" captionTargetPageId="2" captionText="FIGURE 1. Map showing the three sampling stations where all the individuals of Lebetus patzneri sp. nov. (stars) and L. guilleti newly recorded individuals (circles) from the Balearic Islands were collected. The holotype and the paratype of L. patzneri sp. nov. were collected from the sampling station in the Menorca Channel and off southeast Mallorca, respectively. The holotype, paratype and the two unpreserved specimens off south Menorca were collected at 72, 60 and 67 m, respectively. The 50, 100, 500 and 800 m isobaths are shown. Map drawn by F. Ordinas." figureDoi="http://doi.org/10.5281/zenodo.3567338" httpUri="https://zenodo.org/record/3567338/files/figure.png" pageId="12" pageNumber="243">Fig. 1</figureCitation>
|
||
). In that sample
|
||
<quantity id="4CB19B40FF81FFD8AA94F85DFCAAF87F" box="[743,822,1929,1955]" metricMagnitude="2" metricUnit="kg" metricValue="1.66" pageId="12" pageNumber="243" unit="kg" value="166.0">166 kg</quantity>
|
||
of algae were collected from a swept area of
|
||
<quantity id="4CB19B40FF81FFD8AD36F85DFA11F87F" box="[1349,1421,1929,1955]" metricMagnitude="2" metricUnit="m" metricValue="2.73" pageId="12" pageNumber="243" unit="m" value="273.0">273 m</quantity>
|
||
<superScript id="7C3C9BEDFF81FFD8ADFEF85CFA09F84A" attach="left" box="[1421,1429,1928,1942]" fontSize="6" pageId="12" pageNumber="243">2</superScript>
|
||
, with dominance of coralline red algae species, mainly
|
||
<taxonomicName id="4C494D26FF81FFD8AB75F879FBA5F81B" box="[774,1081,1965,1991]" class="Florideophyceae" family="Corallinaceae" genus="Lithothamnion" higherTaxonomySource="GBIF" kingdom="Plantae" order="Corallinales" pageId="12" pageNumber="243" phylum="Rhodophyta" rank="species" species="corallioides">
|
||
<emphasis id="B93DEAB7FF81FFD8AB75F879FBA5F81B" box="[774,1081,1965,1991]" italics="true" pageId="12" pageNumber="243">Lithothamnion corallioides</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C494D26FF81FFD8AC3AF879FAF7F81B" authorityName="W.H.Adey & D.L.McKibbin" authorityYear="1970" baseAuthorityName="Pallas" baseAuthorityYear="1766" box="[1097,1387,1965,1991]" class="Florideophyceae" family="Corallinaceae" genus="Phymatolithon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Corallinales" pageId="12" pageNumber="243" phylum="Rhodophyta" rank="species" species="calcareum">
|
||
<emphasis id="B93DEAB7FF81FFD8AC3AF879FAF7F81B" box="[1097,1387,1965,1991]" italics="true" pageId="12" pageNumber="243">Phymatolithon calcareum</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C494D26FF81FFD8A8E4F805FE08F837" authorityName="Foslie" authorityYear="1901" baseAuthorityName="Lamarck" box="[151,404,2001,2027]" family="Corallinaceae" genus="Lithophyllum" higherTaxonomySource="CoL" kingdom="Plantae" order="Cryptonemiales" pageId="12" pageNumber="243" phylum="Rhodophyta" rank="species" species="racemus">
|
||
<emphasis id="B93DEAB7FF81FFD8A8E4F805FE08F837" box="[151,404,2001,2027]" italics="true" pageId="12" pageNumber="243">Lithophyllum racemus</emphasis>
|
||
</taxonomicName>
|
||
(39, 10.6 and
|
||
<quantity id="4CB19B40FF81FFD8AA48F805FD19F837" box="[571,645,2001,2027]" metricMagnitude="0" metricUnit="kg" metricValue="4.1" pageId="12" pageNumber="243" unit="kg" value="4.1">4.1 kg</quantity>
|
||
, respectively), but also with brown algae, mainly
|
||
<taxonomicName id="4C494D26FF81FFD8ACCEF806FA00F837" box="[1213,1436,2001,2027]" class="Insecta" family="Tettigoniidae" genus="Dictyota" higherTaxonomySource="CoL" kingdom="Animalia" order="Orthoptera" pageId="12" pageNumber="243" phylum="Arthropoda" rank="species" species="dichotoma">
|
||
<emphasis id="B93DEAB7FF81FFD8ACCEF806FA00F837" box="[1213,1436,2001,2027]" italics="true" pageId="12" pageNumber="243">Dictyota dichotoma</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C494D26FF83FFDAA8B4FF4EFE02FF6F" box="[199,414,153,179]" class="Polypodiopsida" family="Dryopteridaceae" genus="Dictyopteris" higherTaxonomySource="CoL" kingdom="Plantae" order="Polypodiales" pageId="14" pageNumber="245" phylum="Tracheophyta" rank="species" species="lucida">
|
||
<emphasis id="B93DEAB7FF83FFDAA8B4FF4EFE02FF6F" box="[199,414,153,179]" italics="true" pageId="14" pageNumber="245">Dictyopteris lucida</emphasis>
|
||
</taxonomicName>
|
||
(21.9 and
|
||
<quantity id="4CB19B40FF83FFDAAA64FF4DFDFCFF6F" box="[535,608,153,179]" metricMagnitude="0" metricUnit="kg" metricValue="3.1" pageId="14" pageNumber="245" unit="kg" value="3.1">3.1 kg</quantity>
|
||
, respectively). Finally, the
|
||
<typeStatus id="54F28807FF83FFDAABFCFF4DFC6DFF6F" box="[911,1009,153,179]" pageId="14" pageNumber="245" type="holotype">holotype</typeStatus>
|
||
was collected from a rhodolith bed in the
|
||
<collectingCountry id="F35E7635FF83FFDAA8B3FF69FEB8FF0B" box="[192,292,189,215]" name="Spain" pageId="14" pageNumber="245">Menorca</collectingCountry>
|
||
Channel (
|
||
<figureCitation id="13722A20FF83FFDAA9E6FF69FE4AFF0B" box="[405,470,189,215]" captionStart="FIGURE 1" captionStartId="2.[151,250,1666,1691]" captionTargetBox="[151,1435,737,1642]" captionTargetId="figure@2.[151,1435,733,1647]" captionTargetPageId="2" captionText="FIGURE 1. Map showing the three sampling stations where all the individuals of Lebetus patzneri sp. nov. (stars) and L. guilleti newly recorded individuals (circles) from the Balearic Islands were collected. The holotype and the paratype of L. patzneri sp. nov. were collected from the sampling station in the Menorca Channel and off southeast Mallorca, respectively. The holotype, paratype and the two unpreserved specimens off south Menorca were collected at 72, 60 and 67 m, respectively. The 50, 100, 500 and 800 m isobaths are shown. Map drawn by F. Ordinas." figureDoi="http://doi.org/10.5281/zenodo.3567338" httpUri="https://zenodo.org/record/3567338/files/figure.png" pageId="14" pageNumber="245">Fig. 1</figureCitation>
|
||
). In that sample
|
||
<quantity id="4CB19B40FF83FFDAAAE2FF68FD53FF0B" box="[657,719,188,215]" metricMagnitude="1" metricUnit="kg" metricValue="5.6" pageId="14" pageNumber="245" unit="kg" value="56.0">56 kg</quantity>
|
||
of algae were collected from a swept area of
|
||
<quantity id="4CB19B40FF83FFDAACB5FF69FA90FF0B" box="[1222,1292,189,215]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="14" pageNumber="245" unit="m" value="400.0">400 m</quantity>
|
||
<superScript id="7C3C9BEDFF83FFDAAD7FFF68FA88FF16" attach="left" box="[1292,1300,188,202]" fontSize="6" pageId="14" pageNumber="245">2</superScript>
|
||
, with dominance of coralline red algae species, mainly
|
||
<emphasis id="B93DEAB7FF83FFDAAA0CFF36FD76FF27" box="[639,746,225,251]" italics="true" pageId="14" pageNumber="245">
|
||
<taxonomicName id="4C494D26FF83FFDAAA0CFF36FD7AFF27" authorityName="Foslie" authorityYear="1909" box="[639,742,225,251]" class="Florideophyceae" family="Corallinaceae" genus="Lithothamnion" higherTaxonomySource="GBIF" kingdom="Plantae" order="Corallinales" pageId="14" pageNumber="245" phylum="Rhodophyta" rank="species" species="valens">L. valens</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
unidentified crustose coralline algae and
|
||
<taxonomicName id="4C494D26FF83FFDAACC5FF35FAC9FF27" authorityName="Kutzing" authorityYear="1841" box="[1206,1365,225,251]" class="Florideophyceae" family="Corallinaceae" genus="Spongites" higherTaxonomySource="GBIF" kingdom="Plantae" order="Corallinales" pageId="14" pageNumber="245" phylum="Rhodophyta" rank="species" species="fruticulosus">
|
||
<emphasis id="B93DEAB7FF83FFDAACC5FF35FAC9FF27" box="[1206,1365,225,251]" italics="true" pageId="14" pageNumber="245">S. fruticulosus</emphasis>
|
||
</taxonomicName>
|
||
(22.6, 19.9 and
|
||
<quantity id="4CB19B40FF83FFDAA88CFED1FECBFEC3" box="[255,343,261,287]" metricMagnitude="1" metricUnit="kg" metricValue="1.24" pageId="14" pageNumber="245" unit="kg" value="12.4">12.4 kg</quantity>
|
||
, respectively). In all three sampling stations, the specimens were also accompanied by a large diversity of benthic animals (both sessile and mobile) with most abundant species including the decapods crustaceans
|
||
<taxonomicName id="4C494D26FF83FFDAA8E4FE99FEE3FEBB" baseAuthorityName="Pennant" baseAuthorityYear="1777" box="[151,383,333,359]" class="Malacostraca" family="Inachidae" genus="Inachus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="245" phylum="Arthropoda" rank="species" species="dorsettensis">
|
||
<emphasis id="B93DEAB7FF83FFDAA8E4FE99FEE3FEBB" box="[151,383,333,359]" italics="true" pageId="14" pageNumber="245">Inachus dorsettensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C494D26FF83FFDAA9FCFE99FDF4FEBB" authorityName="Roux" authorityYear="1830" box="[399,616,333,359]" class="Malacostraca" family="Inachidae" genus="Inachus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="245" phylum="Arthropoda" rank="species" species="thoracicus">
|
||
<emphasis id="B93DEAB7FF83FFDAA9FCFE99FDF4FEBB" box="[399,616,333,359]" italics="true" pageId="14" pageNumber="245">Inachus thoracicus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C494D26FF83FFDAAAD0FE9AFCF3FEBB" authorityName="Leach" authorityYear="1815" box="[675,879,333,359]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="245" phylum="Arthropoda" rank="species" species="prideaux">
|
||
<emphasis id="B93DEAB7FF83FFDAAAD0FE9AFCF3FEBB" box="[675,879,333,359]" italics="true" pageId="14" pageNumber="245">Pagurus prideaux</emphasis>
|
||
</taxonomicName>
|
||
, the teleost fish
|
||
<taxonomicName id="4C494D26FF83FFDAAC42FE9AFB7BFEBB" authorityName="Kovacic, Ordines & Schliewen" authorityYear="2017" box="[1073,1255,334,359]" class="Actinopterygii" family="Gobiidae" genus="Buenia" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="245" phylum="Chordata" rank="species" species="massutii">
|
||
<emphasis id="B93DEAB7FF83FFDAAC42FE9AFB7BFEBB" box="[1073,1255,334,359]" italics="true" pageId="14" pageNumber="245">Buenia massutii</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C494D26FF83FFDAAC84FE99FE9EFE57" baseAuthorityName="Pellegrin & Fage" baseAuthorityYear="1907" class="Actinopterygii" family="Gobiidae" genus="Odondebuenia" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="245" phylum="Chordata" rank="species" species="balearica">
|
||
<emphasis id="B93DEAB7FF83FFDAAC84FE99FE9EFE57" italics="true" pageId="14" pageNumber="245">Odondebuenia balearica</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C494D26FF83FFDAA961FEA5FDA4FE57" baseAuthorityName="Bonnaterre" baseAuthorityYear="1788" box="[274,568,369,395]" class="Actinopterygii" family="Gobiesocidae" genus="Diplecogaster" kingdom="Animalia" order="Gobiesociformes" pageId="14" pageNumber="245" phylum="Chordata" rank="species" species="bimaculata">
|
||
<emphasis id="B93DEAB7FF83FFDAA961FEA5FDA4FE57" box="[274,568,369,395]" italics="true" pageId="14" pageNumber="245">Diplecogaster bimaculata</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C494D26FF83FFDAAA01FEA5FCABFE57" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[626,823,369,395]" class="Actinopterygii" family="Serranidae" genus="Serranus" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="245" phylum="Chordata" rank="species" species="hepatus">
|
||
<emphasis id="B93DEAB7FF83FFDAAA01FEA5FCABFE57" box="[626,823,369,395]" italics="true" pageId="14" pageNumber="245">Serranus hepatus</emphasis>
|
||
</taxonomicName>
|
||
, the echinoderms
|
||
<taxonomicName id="4C494D26FF83FFDAAC79FEA5FAB1FE57" baseAuthorityName="Lamarck" baseAuthorityYear="1816" box="[1034,1325,369,395]" class="Echinoidea" family="Toxopneustidae" genus="Sphaerechinus" kingdom="Animalia" order="Camarodonta" pageId="14" pageNumber="245" phylum="Echinodermata" rank="species" species="granularis">
|
||
<emphasis id="B93DEAB7FF83FFDAAC79FEA5FAB1FE57" box="[1034,1325,369,395]" italics="true" pageId="14" pageNumber="245">Sphaerechinus granularis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C494D26FF83FFDAAD4EFEA5FF43FE73" authorityName="Forbes" authorityYear="1839" class="Ophiuroidea" family="Ophiuridae" genus="Ophiura" kingdom="Animalia" order="Ophiurida" pageId="14" pageNumber="245" phylum="Echinodermata" rank="species" species="albida">
|
||
<emphasis id="B93DEAB7FF83FFDAAD4EFEA5FF43FE73" italics="true" pageId="14" pageNumber="245">Ophiura albida</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C494D26FF83FFDAA966FE41FE61FE73" baseAuthorityName="Retzius" baseAuthorityYear="1783" box="[277,509,405,431]" class="Asteroidea" family="Echinasteridae" genus="Echinaster" kingdom="Animalia" order="Spinulosida" pageId="14" pageNumber="245" phylum="Echinodermata" rank="species" species="sepositus">
|
||
<emphasis id="B93DEAB7FF83FFDAA966FE41FE61FE73" box="[277,509,405,431]" italics="true" pageId="14" pageNumber="245">Echinaster sepositus</emphasis>
|
||
</taxonomicName>
|
||
, the annelids
|
||
<taxonomicName id="4C494D26FF83FFDAAAEBFE41FC18FE73" baseAuthorityName="O. F. Müller" baseAuthorityYear="1776" box="[664,900,405,431]" class="Polychaeta" family="Onuphidae" genus="Hyalinoecia" kingdom="Animalia" order="Eunicida" pageId="14" pageNumber="245" phylum="Annelida" rank="species" species="tubicola">
|
||
<emphasis id="B93DEAB7FF83FFDAAAEBFE41FC18FE73" box="[664,900,405,431]" italics="true" pageId="14" pageNumber="245">Hyalinoecia tubicola</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C494D26FF83FFDAABC8FE42FB0CFE73" baseAuthorityName="Savigny in Lamarck" baseAuthorityYear="1818" box="[955,1168,405,431]" class="Polychaeta" family="Aphroditidae" genus="Laetmonice" kingdom="Animalia" order="Phyllodocida" pageId="14" pageNumber="245" phylum="Annelida" rank="species" species="hystrix">
|
||
<emphasis id="B93DEAB7FF83FFDAABC8FE42FB0CFE73" box="[955,1168,405,431]" italics="true" pageId="14" pageNumber="245">Laetmonice hystrix</emphasis>
|
||
</taxonomicName>
|
||
, the ascidians
|
||
<taxonomicName id="4C494D26FF83FFDAAD45FE41FE8DFE0F" baseAuthorityName="Milne Edwards" baseAuthorityYear="1841" class="Ascidiacea" family="Polyclinidae" genus="Aplidium" kingdom="Animalia" order="Enterogona" pageId="14" pageNumber="245" phylum="Chordata" rank="species" species="nordmanni">
|
||
<emphasis id="B93DEAB7FF83FFDAAD45FE41FE8DFE0F" italics="true" pageId="14" pageNumber="245">Aplidium nordmanni</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C494D26FF83FFDAA96CFE6DFEFFFE0F" authorityName="Fleming" authorityYear="1822" box="[287,355,441,467]" class="Ascidiacea" family="Cionidae" genus="Ciona" kingdom="Animalia" order="Enterogona" pageId="14" pageNumber="245" phylum="Chordata" rank="genus">
|
||
<emphasis id="B93DEAB7FF83FFDAA96CFE6DFEFFFE0F" box="[287,355,441,467]" italics="true" pageId="14" pageNumber="245">Ciona</emphasis>
|
||
</taxonomicName>
|
||
spp. and
|
||
<taxonomicName id="4C494D26FF83FFDAA9BCFE6DFD18FE0F" box="[463,644,441,467]" class="Ascidiacea" family="Ascidiidae" genus="Ascidia" kingdom="Animalia" order="Enterogona" pageId="14" pageNumber="245" phylum="Chordata" rank="species" species="mentula">
|
||
<emphasis id="B93DEAB7FF83FFDAA9BCFE6DFD18FE0F" box="[463,644,441,467]" italics="true" pageId="14" pageNumber="245">Ascidia mentula</emphasis>
|
||
</taxonomicName>
|
||
, and several unidentified sponge species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |